587 lines
75 KiB
XML
587 lines
75 KiB
XML
<document id="1ED8EE6F4A1E5748CCD7A1A19887354B" ID-CLB-Dataset="54091" ID-DOI="10.5852/ejt.2023.866.2107" ID-GBIF-Dataset="92a4f6eb-0d6c-4aa2-8290-a7d2f19a3f34" ID-ISSN="2118-9773" ID-Zenodo-Dep="7867353" ID-ZooBank="28B62104-CA03-481A-B229-D697148D4DE0" IM.bibliography_approvedBy="diego" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="diego" IM.treatments_approvedBy="diego" checkinTime="1682511971143" checkinUser="felipe" docAuthor="Miller, Jonathan P., Delicado, Diana, García-Guerrero, Fernando, Khalloufi, Noureddine & Ramos, Marian A." docDate="2023" docId="5C7987C4FFCEFFCAECAEFDA2348DFE90" docLanguage="en" docName="ejt-866_1-63.pdf" docOrigin="European Journal of Taxonomy 866 (1)" docSource="http://zoobank.org/28b62104-ca03-481a-b229-d697148d4de0" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Mercuria veronicae Miller, Khalloufi & Delicado 2023, sp. nov." docType="treatment" docUuid="986C0653-973B-4C3D-9A4A-C6A216321EBE" docUuidSource="ZooBank" docVersion="9" lastPageNumber="52" masterDocId="A040FFBCFFE0FFF9ED6DFF9E352BFFFA" masterDocTitle="Morphology and taxonomic assessment of eight genetic clades of Mercuria Boeters, 1971 (Caenogastropoda, Hydrobiidae), with the description of five new species" masterLastPageNumber="63" masterPageNumber="1" pageNumber="47" updateTime="1699514902494" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="C508165C4EB6BF8388C9F9C582450079">Morphology and taxonomic assessment of eight genetic clades of Mercuria Boeters, 1971 (Caenogastropoda, Hydrobiidae), with the description of five new species</mods:title>
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<mods:namePart id="69EB5E0DB5DA3EE3719A64BAF8EA0AF7">Miller, Jonathan P.</mods:namePart>
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<mods:affiliation id="42CBA524205DF858F0BC9648DCF377FE">Museo Nacional de Ciencias Naturales (MNCN-CSIC), José Gutiérrez Abascal 2, 28006 Madrid, Spain. Justus Liebig University Giessen, Department of Animal Ecology & Systematics, Heinrich-Buff-Ring 26 - 32 IFZD, 35392 Giessen, Germany. University of Carthage, Faculty of Sciences of Bizerte, LR 01 ES 14 Environmental Biomonitoring Laboratory, 7021 Jarzouna, Tunisia. Deceased 3 March 2023.</mods:affiliation>
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<mods:nameIdentifier id="30D933FB75BC81F7AF80622496EA59E6" type="email">jonathanmiller@mncn.csic.es</mods:nameIdentifier>
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<mods:namePart id="7FBFA5EAE245003B657F905893F5B138">Delicado, Diana</mods:namePart>
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<mods:namePart id="7F701119D6C618B7631A0E823BB67686">García-Guerrero, Fernando</mods:namePart>
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<mods:affiliation id="470A055547B3E663E48777B5D3ABB066">Museo Nacional de Ciencias Naturales (MNCN-CSIC), José Gutiérrez Abascal 2, 28006 Madrid, Spain. Justus Liebig University Giessen, Department of Animal Ecology & Systematics, Heinrich-Buff-Ring 26 - 32 IFZD, 35392 Giessen, Germany. University of Carthage, Faculty of Sciences of Bizerte, LR 01 ES 14 Environmental Biomonitoring Laboratory, 7021 Jarzouna, Tunisia. Deceased 3 March 2023.</mods:affiliation>
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<mods:namePart id="54CE4DB3C1E50DB5D149852FE6354614">Khalloufi, Noureddine</mods:namePart>
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<mods:title id="53570AB8F231A9B8048A2BB5B7C6948A">European Journal of Taxonomy</mods:title>
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<treatment id="5C7987C4FFCEFFCAECAEFDA2348DFE90" ID-DOI="http://doi.org/10.5281/zenodo.7891743" ID-GBIF-Taxon="209579762" ID-Zenodo-Dep="7891743" LSID="urn:lsid:zoobank.org:act:986C0653-973B-4C3D-9A4A-C6A216321EBE" httpUri="http://treatment.plazi.org/id/5C7987C4FFCEFFCAECAEFDA2348DFE90" lastPageId="51" lastPageNumber="52" pageId="46" pageNumber="47">
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<heading id="8F2781BEFFCEFFD7ECAEFDA2315BFDAC" box="[451,1136,572,598]" centered="true" fontSize="11" level="2" pageId="46" pageNumber="47" reason="2">
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<taxonomicName id="13D04D51FFCEFFD7ECAEFDA23124FDAC" ID-CoL="72221103-0251-44d9-9a0a-3bcb78a81425" authority="Miller, Khalloufi & Delicado" authorityName="Miller, Khalloufi & Delicado" authorityYear="2023" box="[451,1039,572,598]" class="Gastropoda" family="Hydrobiidae" genus="Mercuria" kingdom="Animalia" order="Littorinimorpha" pageId="46" pageNumber="47" phylum="Mollusca" rank="species" species="veronicae" status="sp. nov.">
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<emphasis id="E6A4EAC0FFCEFFD7ECAEFDA23799FDAC" bold="true" box="[451,690,572,598]" italics="true" pageId="46" pageNumber="47">Mercuria veronicae</emphasis>
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Miller, Khalloufi & Delicado
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<taxonomicNameLabel id="FD9757BBFFCEFFD7E97BFDA2315BFDAC" box="[1046,1136,572,598]" pageId="46" pageNumber="47" rank="species">sp. nov.</taxonomicNameLabel>
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<subSubSection id="9CCA6559FFCEFFD7EC01FDC13126FD66" pageId="46" pageNumber="47" type="description">
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urn:lsid:zoobank.org:act:
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<paragraph id="D46F36D2FFCEFFD7EF4AFD1C3126FD66" blockId="46.[364,1223,572,669]" box="[551,1037,642,669]" pageId="46" pageNumber="47">
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<figureCitation id="4CEB2A57FFCEFFD7EF4AFD1C3785FD66" box="[551,686,642,668]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStartId-0="47.[189,232,1884,1910]" captionStartId-1="48.[188,231,1928,1954]" captionStartId-2="49.[189,232,1928,1954]" captionTargetBox-0="[189,1398,269,1844]" captionTargetBox-1="[189,1398,790,1886]" captionTargetBox-2="[189,1397,900,1884]" captionTargetId-0="figure-11@47.[189,1398,269,1844]" captionTargetId-1="figure-174@48.[189,1398,790,1887]" captionTargetId-2="figure-236@49.[189,1398,900,1885]" captionTargetPageId-0="47" captionTargetPageId-1="48" captionTargetPageId-2="49" captionText-0="Fig. 29. Shells and operculum ofMercuria veronicaeMiller, Khalloufi & Delicado sp. nov.A–B. Holotype (MNCN 15.05/200124H), frontal and lateral views. C–D. Paratypes. C. MNCN 15.05/200124P. D. UGSB 17271. A–D, G–H. El Waha Spring, Oasis Waterfall, Tamerza, Tozeur. E–F. Lekbir Spring, Big Waterfall, Tamerza, Tozeur. G. Operculum, inner side. H. Operculum, outer side." captionText-1="Fig. 30. Radula of the Mercuria veronicae Miller, Khalloufi & Delicado sp. nov. from El Waha Spring, Tozeur. A. General view of the radular ribbon. B. Overview of radular teeth rows. C. Detailed view of the central teeth. D. Detailed view of the outer marginal teeth." captionText-2="Fig. 31. Anatomy of the Mercuria veronicae Miller, Khalloufi & Delicado sp. nov. from Spring El Waha, Tozeur, Tunisia. A. Head. B. Ctenidium and osphradium. C. Stomach. D. Head of a male. E. Penis. F. Prostate gland. G. Head with perioesophageal ring attached. H. Perioesophageal ring." figureDoi-0="http://doi.org/10.5281/zenodo.7867427" figureDoi-1="http://doi.org/10.5281/zenodo.7867429" figureDoi-2="http://doi.org/10.5281/zenodo.7867431" httpUri-0="https://zenodo.org/record/7867427/files/figure.png" httpUri-1="https://zenodo.org/record/7867429/files/figure.png" httpUri-2="https://zenodo.org/record/7867431/files/figure.png" pageId="46" pageNumber="47">Figs 29–31</figureCitation>
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; Supp. file 2: Tables S14–S18
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</paragraph>
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<subSubSection id="9CCA6559FFCEFFD7EDD0FD5D3074FC47" pageId="46" pageNumber="47" type="diagnosis">
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<paragraph id="D46F36D2FFCEFFD7EDD0FD5D341DFD27" blockId="46.[189,310,707,733]" box="[189,310,707,733]" pageId="46" pageNumber="47">
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<heading id="8F2781BEFFCEFFD7EDD0FD5D341DFD27" bold="true" box="[189,310,707,733]" fontSize="11" level="3" pageId="46" pageNumber="47" reason="3">
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<emphasis id="E6A4EAC0FFCEFFD7EDD0FD5D341DFD27" bold="true" box="[189,310,707,733]" pageId="46" pageNumber="47">Diagnosis</emphasis>
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<paragraph id="D46F36D2FFCEFFD7EDD0FD6C3074FC47" blockId="46.[187,1400,754,957]" pageId="46" pageNumber="47">Shell ovate-conic; aperture obliquely ovate; periostracum whitish to pale grey; central radular tooth formula (5)4-C-4(5)/1-1; female genitalia with a bursa copulatrix pyriform to elongate, ca 2 times as long as wide; seminal receptacle elongate; penis darkly pigmented; penial appendix slightly pigmented, ovate, shorter than or equal in length to the distal end of the penis, medially positioned on the inner edge of the penis; nervous system pigmented, elongate (mean RPG ratio = 0.55); cerebral ganglia approximately equal in size; pleuro-supraoesophageal connective ca 11 times as long as pleuro-suboesophageal one.</paragraph>
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</subSubSection>
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<paragraph id="D46F36D2FFCEFFD7EDD0FC7A346FFC04" blockId="46.[189,324,996,1022]" box="[189,324,996,1022]" pageId="46" pageNumber="47">
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<heading id="8F2781BEFFCEFFD7EDD0FC7A346FFC04" bold="true" box="[189,324,996,1022]" fontSize="11" level="3" pageId="46" pageNumber="47" reason="3">
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<emphasis id="E6A4EAC0FFCEFFD7EDD0FC7A346FFC04" bold="true" box="[189,324,996,1022]" pageId="46" pageNumber="47">Etymology</emphasis>
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</heading>
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</paragraph>
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<paragraph id="D46F36D2FFCEFFD7EDD0FB8C3630FBD6" blockId="46.[189,795,1042,1068]" box="[189,795,1042,1068]" pageId="46" pageNumber="47">After Veronica Ruiz, the first author’s grandmother.</paragraph>
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</subSubSection>
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<subSubSection id="9CCA6559FFCEFFD7EDD0FBCA305CF928" pageId="46" pageNumber="47" type="materials_examined">
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<heading id="8F2781BEFFCEFFD7EDD0FBCA3447FB97" bold="true" box="[189,364,1107,1134]" fontSize="11" level="3" pageId="46" pageNumber="47" reason="3">
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<emphasis id="E6A4EAC0FFCEFFD7EDD0FBCA3447FB97" bold="true" box="[189,364,1107,1134]" pageId="46" pageNumber="47">Type material</emphasis>
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</heading>
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<materialsCitation id="64B83C8FFFCEFFD7ED87FB1C34F7FB19" ID-GBIF-Occurrence="4098914328" collectionCode="MNCN" country="Tunisia" location="Oasis Waterfall. El Waha Spring" municipality="Tamerza" pageId="46" pageNumber="47" specimenCode="MNCN 15.05/200124H" specimenCount="1" stateProvince="Tozeur" typeStatus="holotype">
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<emphasis id="E6A4EAC0FFCEFFD7ED87FB1C3477FB66" bold="true" box="[234,348,1154,1180]" pageId="46" pageNumber="47">
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<typeStatus id="0B6B8870FFCEFFD7ED87FB1C3477FB66" box="[234,348,1154,1180]" pageId="46" pageNumber="47" type="holotype">Holotype</typeStatus>
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</emphasis>
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<collectingCountry id="ACC77642FFCEFFD7EDD0FB3B341AFB45" box="[189,305,1189,1215]" name="Tunisia" pageId="46" pageNumber="47">TUNISIA</collectingCountry>
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• sex unknown (preserved in ethanol 80%);
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<collectingRegion id="1614F830FFCEFFD7EE41FB383654FB3A" box="[812,895,1190,1216]" country="Tunisia" name="Tozeur" pageId="46" pageNumber="47">Tozeur</collectingRegion>
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,
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<collectingMunicipality id="340BACA8FFCEFFD7EEE5FB3836DBFB3A" box="[904,1008,1190,1216]" pageId="46" pageNumber="47">Tamerza</collectingMunicipality>
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,
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<location id="D10F6009FFCEFFD7EE97FB3B3059FB45" LSID="urn:lsid:plazi:treatment:5C7987C4FFCEFFCAECAEFDA2348DFE90:D10F6009FFCEFFD7EE97FB3B3059FB45" box="[1018,1394,1189,1215]" country="Tunisia" municipality="Tamerza" name="Oasis Waterfall. El Waha Spring" pageId="46" pageNumber="47" stateProvince="Tozeur">Oasis Waterfall. El Waha Spring</location>
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;
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<specimenCode id="84769EA9FFCEFFD7EDD0FB5734FCFB19" box="[189,471,1224,1251]" pageId="46" pageNumber="47">MNCN 15.05/200124H</specimenCode>
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.
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</materialsCitation>
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|
||
<materialsCitation id="64B83C8FFFCEFFD7ED87FA943445FA91" ID-GBIF-Occurrence="4098914357" collectionCode="MNCN" country="Tunisia" location="TUNISIA" municipality="Tamerza" pageId="46" pageNumber="47" specimenCode="MNCN 15.05/200124P" specimenCount="3" stateProvince="Tozeur" typeStatus="paratype">
|
||
<emphasis id="E6A4EAC0FFCEFFD7ED87FA94344CFADE" bold="true" box="[234,359,1290,1316]" pageId="46" pageNumber="47">
|
||
<typeStatus id="0B6B8870FFCEFFD7ED87FA94344CFADE" box="[234,359,1290,1316]" pageId="46" pageNumber="47" type="paratype">Paratypes</typeStatus>
|
||
</emphasis>
|
||
<collectingCountry id="ACC77642FFCEFFD7EDD0FAB3341AFABD" box="[189,305,1325,1351]" name="Tunisia" pageId="46" pageNumber="47">TUNISIA</collectingCountry>
|
||
•
|
||
<specimenCount id="C2D6FD5BFFCEFFD7EC37FAB33493FAB2" box="[346,440,1325,1352]" pageId="46" pageNumber="47">3 specs</specimenCount>
|
||
(preserved in ethanol 80%); same collection data as for holotype;
|
||
<specimenCode id="84769EA9FFCEFFD7E873FAB33445FA91" pageId="46" pageNumber="47">MNCN 15.05/200124P</specimenCode>
|
||
</materialsCitation>
|
||
<materialsCitation id="64B83C8FFFCEFFD7EC18FACF363DFA74" ID-GBIF-Occurrence="4098914433" collectionCode="UGSB" country="Tunisia" location="Oasis Waterfall. El Waha Spring" municipality="Tamerza" pageId="46" pageNumber="47" specimenCode="UGSB 17271" specimenCount="15" stateProvince="Tozeur" typeStatus="paratype">
|
||
•
|
||
<specimenCount id="C2D6FD5BFFCEFFD7ECE8FACE34C2FA91" box="[389,489,1360,1387]" pageId="46" pageNumber="47">15 specs</specimenCount>
|
||
(preserved in ethanol 80%, 6 dissected and 1 processed for DNA sequencing); same collection data as for holotype;
|
||
<specimenCode id="84769EA9FFCEFFD7EF1EFAEA363DFA74" box="[627,790,1396,1422]" collectionCode="UGSB" pageId="46" pageNumber="47">UGSB 17271</specimenCode>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="D46F36D2FFCEFFD7EDD0FA2B371FFA35" blockId="46.[189,564,1461,1487]" box="[189,564,1461,1487]" pageId="46" pageNumber="47">
|
||
<heading id="8F2781BEFFCEFFD7EDD0FA2B371FFA35" bold="true" box="[189,564,1461,1487]" fontSize="11" level="3" pageId="46" pageNumber="47" reason="3">
|
||
<emphasis id="E6A4EAC0FFCEFFD7EDD0FA2B371FFA35" bold="true" box="[189,564,1461,1487]" pageId="46" pageNumber="47">Additional material examined</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="D46F36D2FFCEFFD7EDD0FA7A36ACF9DB" blockId="46.[189,1398,1507,1569]" pageId="46" pageNumber="47">
|
||
<materialsCitation id="64B83C8FFFCEFFD7EDD0FA7A31B7FA04" ID-GBIF-Occurrence="4098914432" box="[189,1180,1507,1534]" collectionCode="UGSB" country="Tunisia" location="Big Waterfall, Lekbir Spring" municipality="Tamerza" pageId="46" pageNumber="47" specimenCode="UGSB 17274" specimenCount="15" stateProvince="Tozeur">
|
||
<collectingCountry id="ACC77642FFCEFFD7EDD0FA7A341AFA04" box="[189,305,1508,1534]" name="Tunisia" pageId="46" pageNumber="47">TUNISIA</collectingCountry>
|
||
•
|
||
<specimenCount id="C2D6FD5BFFCEFFD7EC21FA7D349DFA04" box="[332,438,1507,1534]" pageId="46" pageNumber="47">15 specs</specimenCount>
|
||
;
|
||
<collectingRegion id="1614F830FFCEFFD7ECAEFA7A373DFA04" box="[451,534,1508,1534]" country="Tunisia" name="Tozeur" pageId="46" pageNumber="47">Tozeur</collectingRegion>
|
||
,
|
||
<collectingMunicipality id="340BACA8FFCEFFD7EF4FFA7A37A1FA04" box="[546,650,1508,1534]" pageId="46" pageNumber="47">Tamerza</collectingMunicipality>
|
||
,
|
||
<location id="D10F6009FFCEFFD7EFFAFA7A36C4FA04" LSID="urn:lsid:plazi:treatment:5C7987C4FFCEFFCAECAEFDA2348DFE90:D10F6009FFCEFFD7EFFAFA7A36C4FA04" box="[663,1007,1508,1534]" country="Tunisia" municipality="Tamerza" name="Big Waterfall, Lekbir Spring" pageId="46" pageNumber="47" stateProvince="Tozeur">Big Waterfall, Lekbir Spring</location>
|
||
;
|
||
<specimenCode id="84769EA9FFCEFFD7EE91FA7A31B7FA04" box="[1020,1180,1508,1534]" collectionCode="UGSB" pageId="46" pageNumber="47">UGSB 17274</specimenCode>
|
||
</materialsCitation>
|
||
<materialsCitation id="64B83C8FFFCEFFD7E9C8FA7A36ACF9DB" ID-GBIF-Occurrence="4098914434" collectionCode="UGSB" location="Echbika Waterfall, Echbicka Spring" municipality="Echbika" pageId="46" pageNumber="47" specimenCode="UGSB 17276" specimenCount="4" stateProvince="Tozeur">
|
||
•
|
||
<specimenCount id="C2D6FD5BFFCEFFD7E9D5FA7A3039FA04" box="[1208,1298,1508,1534]" pageId="46" pageNumber="47">4 specs</specimenCount>
|
||
;
|
||
<collectingRegion id="1614F830FFCEFFD7E872FA7A3059FA04" box="[1311,1394,1508,1534]" country="Tunisia" name="Tozeur" pageId="46" pageNumber="47">Tozeur</collectingRegion>
|
||
,
|
||
<collectingMunicipality id="340BACA8FFCEFFD7EDD0F999340BF9DB" box="[189,288,1543,1569]" pageId="46" pageNumber="47">Echbika</collectingMunicipality>
|
||
,
|
||
<location id="D10F6009FFCEFFD7EC41F99937FFF9DB" LSID="urn:lsid:plazi:treatment:5C7987C4FFCEFFCAECAEFDA2348DFE90:D10F6009FFCEFFD7EC41F99937FFF9DB" box="[300,724,1543,1569]" municipality="Echbika" name="Echbika Waterfall, Echbicka Spring" pageId="46" pageNumber="47" stateProvince="Tozeur">Echbika Waterfall, Echbicka Spring</location>
|
||
;
|
||
<specimenCode id="84769EA9FFCEFFD7EF8DF99936A8F9DB" box="[736,899,1543,1569]" collectionCode="UGSB" pageId="46" pageNumber="47">UGSB 17276</specimenCode>
|
||
.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="D46F36D2FFCEFFD7EDD0F9D636EFF998" blockId="46.[189,964,1608,1635]" box="[189,964,1608,1635]" pageId="46" pageNumber="47">Additional locality information provided in Supp. file 1: Table S1.</paragraph>
|
||
<paragraph id="D46F36D2FFCEFFD7EDD0F9143476F959" blockId="46.[189,349,1673,1700]" box="[189,349,1673,1700]" pageId="46" pageNumber="47">
|
||
<heading id="8F2781BEFFCEFFD7EDD0F9143476F959" bold="true" box="[189,349,1673,1700]" fontSize="11" level="3" pageId="46" pageNumber="47" reason="3">
|
||
<emphasis id="E6A4EAC0FFCEFFD7EDD0F9143476F959" bold="true" box="[189,349,1673,1700]" pageId="46" pageNumber="47">Type locality</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="D46F36D2FFCEFFD7EDD0F926305CF928" blockId="46.[189,1399,1720,1747]" box="[189,1399,1720,1747]" pageId="46" pageNumber="47">
|
||
El Waha Spring, Oasis Waterfall, Tamerza,
|
||
<collectingRegion id="1614F830FFCEFFD7EFDBF9273622F929" box="[694,777,1721,1747]" country="Tunisia" name="Tozeur" pageId="46" pageNumber="47">Tozeur</collectingRegion>
|
||
,
|
||
<collectingCountry id="ACC77642FFCEFFD7EE7EF9263646F928" box="[787,877,1720,1746]" name="Tunisia" pageId="46" pageNumber="47">Tunisia</collectingCountry>
|
||
,
|
||
<geoCoordinate id="B1E45015FFCEFFD7EE15F9263134F929" box="[888,1055,1720,1747]" degrees="34.381933" direction="north" orientation="latitude" pageId="46" pageNumber="47" precision="1" value="34.38193">34.381933° N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="B1E45015FFCEFFD7E947F9263194F929" box="[1066,1215,1720,1747]" degrees="7.933450" direction="east" orientation="longitude" pageId="46" pageNumber="47" precision="1" value="7.93345">7.933450° E</geoCoordinate>
|
||
,
|
||
<quantity id="13289B37FFCEFFD7E9A6F9263012F929" box="[1227,1337,1720,1747]" metricMagnitude="2" metricUnit="m" metricValue="2.7867" pageId="46" pageNumber="47" unit="m" value="278.67">278.67 m</quantity>
|
||
a.s.l.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="9CCA6559FFCEFFCBEDD0F967377CFD66" lastPageId="50" lastPageNumber="51" pageId="46" pageNumber="47" type="description">
|
||
<paragraph id="D46F36D2FFCEFFD7EDD0F9673465F8E9" blockId="46.[189,334,1785,1811]" box="[189,334,1785,1811]" pageId="46" pageNumber="47">
|
||
<heading id="8F2781BEFFCEFFD7EDD0F9673465F8E9" bold="true" box="[189,334,1785,1811]" fontSize="11" level="3" pageId="46" pageNumber="47" reason="3">
|
||
<emphasis id="E6A4EAC0FFCEFFD7EDD0F9673465F8E9" bold="true" box="[189,334,1785,1811]" pageId="46" pageNumber="47">Description</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="D46F36D2FFCEFFD7EDD0F8B73010F89C" blockId="46.[189,1399,1832,1894]" pageId="46" pageNumber="47">
|
||
<smallCapsWord id="D289A00EFFCEFFD7EDD0F8B73445F8BB" baselines="1852,1852" box="[189,366,1833,1859]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Measurements" pageId="46" pageNumber="47">MEASUREMENTS</smallCapsWord>
|
||
.
|
||
<typeStatus id="0B6B8870FFCEFFD7EC13F8B634DBF8B8" box="[382,496,1832,1858]" pageId="46" pageNumber="47" type="holotype">Holotype</typeStatus>
|
||
: SL =
|
||
<quantity id="13289B37FFCEFFD7EF51F8B63783F8B9" box="[572,680,1832,1859]" metricMagnitude="-3" metricUnit="m" metricValue="4.87" pageId="46" pageNumber="47" unit="mm" value="4.87">4.87 mm</quantity>
|
||
, SW =
|
||
<quantity id="13289B37FFCEFFD7EE6DF8B63647F8B9" box="[768,876,1832,1859]" metricMagnitude="-3" metricUnit="m" metricValue="3.56" pageId="46" pageNumber="47" unit="mm" value="3.56">3.56 mm</quantity>
|
||
, SL/SW = 1.36, AH =
|
||
<quantity id="13289B37FFCEFFD7E914F8B631CDF8B9" box="[1145,1254,1832,1859]" metricMagnitude="-3" metricUnit="m" metricValue="2.03" pageId="46" pageNumber="47" unit="mm" value="2.03">2.03 mm</quantity>
|
||
, SL-LBW =
|
||
<quantity id="13289B37FFCEFFD7EDD3F8D53402F89C" box="[190,297,1867,1894]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="46" pageNumber="47" unit="mm" value="1.4">1.40 mm</quantity>
|
||
, WBW =
|
||
<quantity id="13289B37FFCEFFD7ECF3F8D53722F89C" box="[414,521,1867,1894]" metricMagnitude="-3" metricUnit="m" metricValue="2.96" pageId="46" pageNumber="47" unit="mm" value="2.96">2.96 mm</quantity>
|
||
, AL = XX mm, AW =
|
||
<quantity id="13289B37FFCEFFD7EE7FF8D53656F89C" box="[786,893,1867,1894]" metricMagnitude="-3" metricUnit="m" metricValue="2.06" pageId="46" pageNumber="47" unit="mm" value="2.06">2.06 mm</quantity>
|
||
, WPW =
|
||
<quantity id="13289B37FFCEFFD7EE82F8D53171F89C" box="[1007,1114,1867,1894]" metricMagnitude="-3" metricUnit="m" metricValue="1.67" pageId="46" pageNumber="47" unit="mm" value="1.67">1.67 mm</quantity>
|
||
, WAW =
|
||
<quantity id="13289B37FFCEFFD7E9A6F8D5301DF89C" box="[1227,1334,1867,1894]" metricMagnitude="-4" metricUnit="m" metricValue="8.5" pageId="46" pageNumber="47" unit="mm" value="0.85">0.85 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="D46F36D2FFCEFFC9EDD0F8133742FE76" blockId="46.[189,1399,1933,1995]" lastBlockId="48.[189,1399,265,397]" lastPageId="48" lastPageNumber="49" pageId="46" pageNumber="47">
|
||
<smallCapsWord id="D289A00EFFCEFFD7EDD0F8133429F85D" baselines="1952,1953" box="[189,258,1933,1959]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Shell" pageId="46" pageNumber="47">SHELL</smallCapsWord>
|
||
. Ovate-conic, whorls 4–5, height
|
||
<quantity id="13289B37FFCEFFD7EFC7F8133666F85D" box="[682,845,1933,1960]" metricMagnitude="-3" metricUnit="m" metricValue="4.275" metricValueMax="5.3" metricValueMin="3.25" pageId="46" pageNumber="47" unit="mm" value="4.275" valueMax="5.3" valueMin="3.25">3.25–5.3 mm</quantity>
|
||
, width
|
||
<quantity id="13289B37FFCEFFD7EEC1F8133103F85D" box="[940,1064,1933,1960]" metricMagnitude="-3" metricUnit="m" metricValue="3.25" metricValueMax="4.0" metricValueMin="2.5" pageId="46" pageNumber="47" unit="mm" value="3.25" valueMax="4.0" valueMin="2.5">2.5–4 mm</quantity>
|
||
(
|
||
<figureCitation id="4CEB2A57FFCEFFD7E950F81331E5F85D" box="[1085,1230,1933,1959]" captionStart="Fig" captionStartId="47.[189,232,1884,1910]" captionTargetBox="[189,1398,269,1844]" captionTargetId="figure-11@47.[189,1398,269,1844]" captionTargetPageId="47" captionText="Fig. 29. Shells and operculum ofMercuria veronicaeMiller, Khalloufi & Delicado sp. nov.A–B. Holotype (MNCN 15.05/200124H), frontal and lateral views. C–D. Paratypes. C. MNCN 15.05/200124P. D. UGSB 17271. A–D, G–H. El Waha Spring, Oasis Waterfall, Tamerza, Tozeur. E–F. Lekbir Spring, Big Waterfall, Tamerza, Tozeur. G. Operculum, inner side. H. Operculum, outer side." figureDoi="http://doi.org/10.5281/zenodo.7867427" httpUri="https://zenodo.org/record/7867427/files/figure.png" pageId="46" pageNumber="47">Fig. 29A–F</figureCitation>
|
||
; Supp. file 2: Table S14); periostracum whitish to pale grey; protoconch of 1.5 whorls, ca 250 µm wide, nucleus ca 150 µm wide; teleoconch whorls slightly convex, separated by a deep suture; body whorl large, convex, occupying about two-thirds of the total shell length; aperture obliquely broad ovate, complete; inner lip thicker than outer lip; aperture margin straight; inner lip touching the body whorl; umbilicus narrow, not covered by the inner lip.
|
||
</paragraph>
|
||
<caption id="80AF665AFFCFFFD6EDD0F8C2318DF81A" ID-DOI="http://doi.org/10.5281/zenodo.7867427" ID-Zenodo-Dep="7867427" httpUri="https://zenodo.org/record/7867427/files/figure.png" pageId="47" pageNumber="48" startId="47.[189,232,1884,1910]" targetBox="[189,1398,269,1844]" targetPageId="47">
|
||
<paragraph id="D46F36D2FFCFFFD6EDD0F8C2318DF81A" blockId="47.[186,1399,1884,2016]" pageId="47" pageNumber="48">
|
||
<emphasis id="E6A4EAC0FFCFFFD6EDD0F8C2343DF88C" bold="true" box="[189,278,1884,1910]" pageId="47" pageNumber="48">Fig. 29.</emphasis>
|
||
Shells and operculum of
|
||
<taxonomicName id="13D04D51FFCFFFD6EF5DF8C3314FF88C" authority="Miller, Khalloufi & Delicado" authorityName="Miller, Khalloufi & Delicado" authorityYear="2023" box="[560,1124,1884,1910]" class="Gastropoda" family="Hydrobiidae" genus="Mercuria" kingdom="Animalia" order="Littorinimorpha" pageId="47" pageNumber="48" phylum="Mollusca" rank="species" species="veronicae" status="sp. nov.">
|
||
<emphasis id="E6A4EAC0FFCFFFD6EF5DF8C3363EF88C" box="[560,789,1885,1910]" italics="true" pageId="47" pageNumber="48">Mercuria veronicae</emphasis>
|
||
Miller, Khalloufi & Delicado
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="FD9757BBFFCFFFD6E906F8C331E8F88D" box="[1131,1219,1885,1911]" pageId="47" pageNumber="48" rank="species">sp. nov.</taxonomicNameLabel>
|
||
<emphasis id="E6A4EAC0FFCFFFD6E9ABF8C231D5F88C" bold="true" box="[1222,1278,1884,1910]" pageId="47" pageNumber="48">A–B</emphasis>
|
||
. Holotype (MNCN 15.05/200124H), frontal and lateral views.
|
||
<emphasis id="E6A4EAC0FFCFFFD6EE3EF8E136A7F863" bold="true" box="[851,908,1919,1945]" pageId="47" pageNumber="48">C–D</emphasis>
|
||
. Paratypes.
|
||
<emphasis id="E6A4EAC0FFCFFFD6E941F8E1316AF863" bold="true" box="[1068,1089,1919,1945]" pageId="47" pageNumber="48">C</emphasis>
|
||
. MNCN 15.05/200124P.
|
||
<emphasis id="E6A4EAC0FFCFFFD6EDD0F83D35F9F847" bold="true" box="[189,210,1955,1981]" pageId="47" pageNumber="48">D</emphasis>
|
||
. UGSB 17271.
|
||
<emphasis id="E6A4EAC0FFCFFFD6ECFFF83C373DF846" bold="true" box="[402,534,1954,1980]" pageId="47" pageNumber="48">A–D, G–H</emphasis>
|
||
. El Waha Spring, Oasis Waterfall, Tamerza, Tozeur.
|
||
<emphasis id="E6A4EAC0FFCFFFD6E9EAF83D3191F847" bold="true" box="[1159,1210,1955,1981]" pageId="47" pageNumber="48">E–F</emphasis>
|
||
. Lekbir Spring, Big Waterfall, Tamerza, Tozeur.
|
||
<emphasis id="E6A4EAC0FFCFFFD6EF54F85B377BF825" bold="true" box="[569,592,1989,2015]" pageId="47" pageNumber="48">G</emphasis>
|
||
. Operculum, inner side.
|
||
<emphasis id="E6A4EAC0FFCFFFD6EE1EF85836A1F81A" bold="true" box="[883,906,1990,2016]" pageId="47" pageNumber="48">H</emphasis>
|
||
. Operculum, outer side.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="D46F36D2FFD0FFC9EDD0FE2A367BFE0B" blockId="48.[189,1398,436,498]" pageId="48" pageNumber="49">
|
||
<smallCapsWord id="D289A00EFFD0FFC9EDD0FE2A3468FE34" baselines="455,456" box="[189,323,436,462]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Operculum" pageId="48" pageNumber="49">OPERCULUM</smallCapsWord>
|
||
. As for the genus, orange to brown, sometimes yellowish, about two whorls; muscle attachment oval, located near the nucleus (
|
||
<figureCitation id="4CEB2A57FFD0FFC9EFDCFE493668FE0B" box="[689,835,471,497]" captionStart="Fig" captionStartId="47.[189,232,1884,1910]" captionTargetBox="[189,1398,269,1844]" captionTargetId="figure-11@47.[189,1398,269,1844]" captionTargetPageId="47" captionText="Fig. 29. Shells and operculum ofMercuria veronicaeMiller, Khalloufi & Delicado sp. nov.A–B. Holotype (MNCN 15.05/200124H), frontal and lateral views. C–D. Paratypes. C. MNCN 15.05/200124P. D. UGSB 17271. A–D, G–H. El Waha Spring, Oasis Waterfall, Tamerza, Tozeur. E–F. Lekbir Spring, Big Waterfall, Tamerza, Tozeur. G. Operculum, inner side. H. Operculum, outer side." figureDoi="http://doi.org/10.5281/zenodo.7867427" httpUri="https://zenodo.org/record/7867427/files/figure.png" pageId="48" pageNumber="49">Fig. 29G–H</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="D46F36D2FFD0FFC8EDD0FD873037FEBC" blockId="48.[189,1401,536,704]" lastBlockId="49.[189,1398,265,327]" lastPageId="49" lastPageNumber="50" pageId="48" pageNumber="49">
|
||
<smallCapsWord id="D289A00EFFD0FFC9EDD0FD873432FDC9" baselines="556,556" box="[189,281,537,563]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Radula" pageId="48" pageNumber="49">RADULA</smallCapsWord>
|
||
. Length intermediate, ca 750 µm long (35% of total shell length), containing about 60 rows of teeth (
|
||
<figureCitation id="4CEB2A57FFD0FFC9EC65FDA23453FDAC" box="[264,376,572,598]" captionStart="Fig" captionStartId="48.[188,231,1928,1954]" captionTargetBox="[189,1398,790,1886]" captionTargetId="figure-174@48.[189,1398,790,1887]" captionTargetPageId="48" captionText="Fig. 30. Radula of the Mercuria veronicae Miller, Khalloufi & Delicado sp. nov. from El Waha Spring, Tozeur. A. General view of the radular ribbon. B. Overview of radular teeth rows. C. Detailed view of the central teeth. D. Detailed view of the outer marginal teeth." figureDoi="http://doi.org/10.5281/zenodo.7867429" httpUri="https://zenodo.org/record/7867429/files/figure.png" pageId="48" pageNumber="49">Fig. 30A</figureCitation>
|
||
). Central tooth formula (5)4-C-4(5)/1-1, central cusp V shaped, cutting edge concave (
|
||
<figureCitation id="4CEB2A57FFD0FFC9EDA5FDC13472FD83" box="[200,345,607,633]" captionStart="Fig" captionStartId="48.[188,231,1928,1954]" captionTargetBox="[189,1398,790,1886]" captionTargetId="figure-174@48.[189,1398,790,1887]" captionTargetPageId="48" captionText="Fig. 30. Radula of the Mercuria veronicae Miller, Khalloufi & Delicado sp. nov. from El Waha Spring, Tozeur. A. General view of the radular ribbon. B. Overview of radular teeth rows. C. Detailed view of the central teeth. D. Detailed view of the outer marginal teeth." figureDoi="http://doi.org/10.5281/zenodo.7867429" httpUri="https://zenodo.org/record/7867429/files/figure.png" pageId="48" pageNumber="49">Fig. 30B–C</figureCitation>
|
||
). Lateral tooth formula (3)4-C-4(3), central cusp V shaped and slightly longer than the central tooth one. Inner marginal teeth with 11–15 cusps; outer marginal teeth with 21–25 cusps (
|
||
<figureCitation id="4CEB2A57FFD0FFC9EDABFD3B3418FD3A" box="[198,307,677,704]" captionStart="Fig" captionStartId="48.[188,231,1928,1954]" captionTargetBox="[189,1398,790,1886]" captionTargetId="figure-174@48.[189,1398,790,1887]" captionTargetPageId="48" captionText="Fig. 30. Radula of the Mercuria veronicae Miller, Khalloufi & Delicado sp. nov. from El Waha Spring, Tozeur. A. General view of the radular ribbon. B. Overview of radular teeth rows. C. Detailed view of the central teeth. D. Detailed view of the outer marginal teeth." figureDoi="http://doi.org/10.5281/zenodo.7867429" httpUri="https://zenodo.org/record/7867429/files/figure.png" pageId="48" pageNumber="49">Fig. 30D</figureCitation>
|
||
). Radular data were collected from the following specimens: UGSB 17271 – El Waha Spring, Oasis Waterfall, Tamerza,
|
||
<collectingRegion id="1614F830FFD1FFC8EF6CFE97377FFED9" box="[513,596,265,291]" country="Tunisia" name="Tozeur" pageId="49" pageNumber="50">Tozeur</collectingRegion>
|
||
,
|
||
<collectingCountry id="ACC77642FFD1FFC8EF0EFE973795FED9" box="[611,702,265,291]" name="Tunisia" pageId="49" pageNumber="50">Tunisia</collectingCountry>
|
||
; UGSB 17274 – Lekbir Spring, Big Waterfall, Tamerza,
|
||
<collectingRegion id="1614F830FFD1FFC8EDD0FEB3343BFEBD" box="[189,272,301,327]" country="Tunisia" name="Tozeur" pageId="49" pageNumber="50">Tozeur</collectingRegion>
|
||
,
|
||
<collectingCountry id="ACC77642FFD1FFC8EC76FEB2345DFEBC" box="[283,374,300,326]" name="Tunisia" pageId="49" pageNumber="50">Tunisia</collectingCountry>
|
||
; UGSB 17276 – Echbicka Spring, Echbika Waterfall, Echbika,
|
||
<collectingRegion id="1614F830FFD1FFC8E932FEB33199FEBD" box="[1119,1202,301,327]" country="Tunisia" name="Tozeur" pageId="49" pageNumber="50">Tozeur</collectingRegion>
|
||
,
|
||
<collectingCountry id="ACC77642FFD1FFC8E9D0FEB23033FEBC" box="[1213,1304,300,326]" name="Tunisia" pageId="49" pageNumber="50">Tunisia</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
<caption id="80AF665AFFD0FFC9EDD1F81636BFF812" ID-DOI="http://doi.org/10.5281/zenodo.7867429" ID-Zenodo-Dep="7867429" httpUri="https://zenodo.org/record/7867429/files/figure.png" pageId="48" pageNumber="49" startId="48.[188,231,1928,1954]" targetBox="[189,1398,790,1886]" targetPageId="48">
|
||
<paragraph id="D46F36D2FFD0FFC9EDD1F81636BFF812" blockId="48.[188,1398,1928,2025]" pageId="48" pageNumber="49">
|
||
<emphasis id="E6A4EAC0FFD0FFC9EDD1F8163433F858" bold="true" box="[188,280,1928,1954]" pageId="48" pageNumber="49">Fig. 30.</emphasis>
|
||
Radula of the
|
||
<taxonomicName id="13D04D51FFD0FFC9ECAAF8173127F858" authority="Miller, Khalloufi & Delicado" authorityName="Miller, Khalloufi & Delicado" authorityYear="2023" box="[455,1036,1928,1954]" class="Gastropoda" family="Hydrobiidae" genus="Mercuria" kingdom="Animalia" order="Littorinimorpha" pageId="48" pageNumber="49" phylum="Mollusca" rank="species" species="veronicae" status="sp. nov.">
|
||
<emphasis id="E6A4EAC0FFD0FFC9ECAAF8173784F858" box="[455,687,1929,1954]" italics="true" pageId="48" pageNumber="49">Mercuria veronicae</emphasis>
|
||
Miller, Khalloufi & Delicado
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="FD9757BBFFD0FFC9E979F8163145F858" box="[1044,1134,1928,1954]" pageId="48" pageNumber="49" rank="species">sp. nov.</taxonomicNameLabel>
|
||
from El Waha Spring, Tozeur.
|
||
<emphasis id="E6A4EAC0FFD0FFC9EC71F835341AF83F" bold="true" box="[284,305,1963,1989]" pageId="48" pageNumber="49">A</emphasis>
|
||
. General view of the radular ribbon.
|
||
<emphasis id="E6A4EAC0FFD0FFC9EF87F83537D6F83F" bold="true" box="[746,765,1963,1989]" pageId="48" pageNumber="49">B</emphasis>
|
||
. Overview of radular teeth rows.
|
||
<emphasis id="E6A4EAC0FFD0FFC9E9E3F8353188F83F" bold="true" box="[1166,1187,1963,1989]" pageId="48" pageNumber="49">C</emphasis>
|
||
. Detailed view of the central teeth.
|
||
<emphasis id="E6A4EAC0FFD0FFC9ECEAF85034B7F812" bold="true" box="[391,412,1998,2024]" pageId="48" pageNumber="49">D</emphasis>
|
||
. Detailed view of the outer marginal teeth.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="D46F36D2FFD1FFC8EDD0FEF03414FD84" blockId="49.[189,1401,366,638]" pageId="49" pageNumber="50">
|
||
<smallCapsWord id="D289A00EFFD1FFC8EDD0FEF03471FE72" baselines="385,385" box="[189,346,366,392]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Pigmentation" pageId="49" pageNumber="50">PIGMENTATION</smallCapsWord>
|
||
<smallCapsWord id="D289A00EFFD1FFC8EC09FEED34BAFE7C" baselines="385" box="[356,401,371,390]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="and" pageId="49" pageNumber="50">AND</smallCapsWord>
|
||
<smallCapsWord id="D289A00EFFD1FFC8ECF6FEED3729FE7C" baselines="385" box="[411,514,371,390]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="anatomy" pageId="49" pageNumber="50">ANATOMY</smallCapsWord>
|
||
. Animal darkly pigmented (
|
||
<figureCitation id="4CEB2A57FFD1FFC8EE38FEF036C1FE72" box="[853,1002,366,392]" captionStart="Fig" captionStartId="49.[189,232,1928,1954]" captionTargetBox="[189,1397,900,1884]" captionTargetId="figure-236@49.[189,1398,900,1885]" captionTargetPageId="49" captionText="Fig. 31. Anatomy of the Mercuria veronicae Miller, Khalloufi & Delicado sp. nov. from Spring El Waha, Tozeur, Tunisia. A. Head. B. Ctenidium and osphradium. C. Stomach. D. Head of a male. E. Penis. F. Prostate gland. G. Head with perioesophageal ring attached. H. Perioesophageal ring." figureDoi="http://doi.org/10.5281/zenodo.7867431" httpUri="https://zenodo.org/record/7867431/files/figure.png" pageId="49" pageNumber="50">Fig. 31A, D</figureCitation>
|
||
); head and tentacles dark brown, pigmentation lighter on eye lobes, snout and neck; snout about as long as wide, approximately parallel-sided, with medium distal lobation. Ctenidium occupying almost the total length of the pallial cavity; 28– 33 gill filaments; filaments broad, triangular, fused at the base by an epithelium (
|
||
<figureCitation id="4CEB2A57FFD1FFC8E90BFE4931FAFE0B" box="[1126,1233,471,497]" captionStart="Fig" captionStartId="49.[189,232,1928,1954]" captionTargetBox="[189,1397,900,1884]" captionTargetId="figure-236@49.[189,1398,900,1885]" captionTargetPageId="49" captionText="Fig. 31. Anatomy of the Mercuria veronicae Miller, Khalloufi & Delicado sp. nov. from Spring El Waha, Tozeur, Tunisia. A. Head. B. Ctenidium and osphradium. C. Stomach. D. Head of a male. E. Penis. F. Prostate gland. G. Head with perioesophageal ring attached. H. Perioesophageal ring." figureDoi="http://doi.org/10.5281/zenodo.7867431" httpUri="https://zenodo.org/record/7867431/files/figure.png" pageId="49" pageNumber="50">Fig. 31B</figureCitation>
|
||
). Osphradium elongate, more than 3 times as long as wide, positioned opposite middle of ctenidium. Stomach almost as long as wide with two chambers almost equal in size (Supp. file 2: Table S16); style sac longer than wide, with the unpigmented intestine surrounding its distal part and continuing on as a straight rectum (
|
||
<figureCitation id="4CEB2A57FFD1FFC8EDABFDFA3419FD84" box="[198,306,612,638]" captionStart="Fig" captionStartId="49.[189,232,1928,1954]" captionTargetBox="[189,1397,900,1884]" captionTargetId="figure-236@49.[189,1398,900,1885]" captionTargetPageId="49" captionText="Fig. 31. Anatomy of the Mercuria veronicae Miller, Khalloufi & Delicado sp. nov. from Spring El Waha, Tozeur, Tunisia. A. Head. B. Ctenidium and osphradium. C. Stomach. D. Head of a male. E. Penis. F. Prostate gland. G. Head with perioesophageal ring attached. H. Perioesophageal ring." figureDoi="http://doi.org/10.5281/zenodo.7867431" httpUri="https://zenodo.org/record/7867431/files/figure.png" pageId="49" pageNumber="50">Fig. 31C</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="D46F36D2FFD1FFCBEDD0FD38366CFEBC" blockId="49.[189,1400,677,810]" lastBlockId="50.[189,1399,265,327]" lastPageId="50" lastPageNumber="51" pageId="49" pageNumber="50">
|
||
<smallCapsWord id="D289A00EFFD1FFC8EDD0FD3835D4FD3A" baselines="697,697" box="[189,255,678,704]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Male" pageId="49" pageNumber="50">MALE</smallCapsWord>
|
||
<smallCapsWord id="D289A00EFFD1FFC8EC64FD35345CFD44" baselines="697" box="[265,375,683,702]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="genitalia" pageId="49" pageNumber="50">GENITALIA</smallCapsWord>
|
||
. Prostate gland bean-shaped, about 2 times as long as wide (Supp. file 2: Table S18). Pallial vas deferens emerge close to both the anterior edge of the prostate and the external margin of the penis (
|
||
<figureCitation id="4CEB2A57FFD1FFC8EC57FD72348FFCFC" box="[314,420,748,774]" captionStart="Fig" captionStartId="49.[189,232,1928,1954]" captionTargetBox="[189,1397,900,1884]" captionTargetId="figure-236@49.[189,1398,900,1885]" captionTargetPageId="49" captionText="Fig. 31. Anatomy of the Mercuria veronicae Miller, Khalloufi & Delicado sp. nov. from Spring El Waha, Tozeur, Tunisia. A. Head. B. Ctenidium and osphradium. C. Stomach. D. Head of a male. E. Penis. F. Prostate gland. G. Head with perioesophageal ring attached. H. Perioesophageal ring." figureDoi="http://doi.org/10.5281/zenodo.7867431" httpUri="https://zenodo.org/record/7867431/files/figure.png" pageId="49" pageNumber="50">Fig. 31F</figureCitation>
|
||
). Penis darkly pigmented, gradually tapering, attached to the neck behind the right eye, distal end of the penis broad, triangular; penial appendix pyriform, shorter than or about to the same length as the distal end of the penis, slightly pigmented at the junction with the penis, medially positioned on the inner edge of the penis (
|
||
<figureCitation id="4CEB2A57FFD2FFCBEFC6FEB23611FEBC" box="[683,826,300,326]" captionStart="Fig" captionStartId="49.[189,232,1928,1954]" captionTargetBox="[189,1397,900,1884]" captionTargetId="figure-236@49.[189,1398,900,1885]" captionTargetPageId="49" captionText="Fig. 31. Anatomy of the Mercuria veronicae Miller, Khalloufi & Delicado sp. nov. from Spring El Waha, Tozeur, Tunisia. A. Head. B. Ctenidium and osphradium. C. Stomach. D. Head of a male. E. Penis. F. Prostate gland. G. Head with perioesophageal ring attached. H. Perioesophageal ring." figureDoi="http://doi.org/10.5281/zenodo.7867431" httpUri="https://zenodo.org/record/7867431/files/figure.png" pageId="50" pageNumber="51">Fig. 31D–E</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="80AF665AFFD1FFC8EDD0F81631E1F812" ID-DOI="http://doi.org/10.5281/zenodo.7867431" ID-Zenodo-Dep="7867431" httpUri="https://zenodo.org/record/7867431/files/figure.png" pageId="49" pageNumber="50" startId="49.[189,232,1928,1954]" targetBox="[189,1397,900,1884]" targetPageId="49">
|
||
<paragraph id="D46F36D2FFD1FFC8EDD0F81631E1F812" blockId="49.[188,1399,1928,2025]" pageId="49" pageNumber="50">
|
||
<emphasis id="E6A4EAC0FFD1FFC8EDD0F8163433F858" bold="true" box="[189,280,1928,1954]" pageId="49" pageNumber="50">Fig. 31.</emphasis>
|
||
Anatomy of the
|
||
<taxonomicName id="13D04D51FFD1FFC8ECBBF817313FF858" authority="Miller, Khalloufi & Delicado" authorityName="Miller, Khalloufi & Delicado" authorityYear="2023" box="[470,1044,1928,1954]" class="Gastropoda" family="Hydrobiidae" genus="Mercuria" kingdom="Animalia" order="Littorinimorpha" pageId="49" pageNumber="50" phylum="Mollusca" rank="species" species="veronicae" status="sp. nov.">
|
||
<emphasis id="E6A4EAC0FFD1FFC8ECBBF8173796F858" box="[470,701,1929,1954]" italics="true" pageId="49" pageNumber="50">Mercuria veronicae</emphasis>
|
||
Miller, Khalloufi & Delicado
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="FD9757BBFFD1FFC8E977F816315EF858" box="[1050,1141,1928,1954]" pageId="49" pageNumber="50" rank="species">sp. nov.</taxonomicNameLabel>
|
||
from Spring El Waha, Tozeur, Tunisia.
|
||
<emphasis id="E6A4EAC0FFD1FFC8ECE4F83534B7F83F" bold="true" box="[393,412,1963,1989]" pageId="49" pageNumber="50">A</emphasis>
|
||
. Head.
|
||
<emphasis id="E6A4EAC0FFD1FFC8EC91F835373BF83F" bold="true" box="[508,528,1963,1989]" pageId="49" pageNumber="50">B</emphasis>
|
||
. Ctenidium and osphradium.
|
||
<emphasis id="E6A4EAC0FFD1FFC8EE15F83536A6F83F" bold="true" box="[888,909,1963,1989]" pageId="49" pageNumber="50">C</emphasis>
|
||
. Stomach.
|
||
<emphasis id="E6A4EAC0FFD1FFC8E97AF8353107F83F" bold="true" box="[1047,1068,1963,1989]" pageId="49" pageNumber="50">D</emphasis>
|
||
. Head of a male.
|
||
<emphasis id="E6A4EAC0FFD1FFC8E867F8353035F83F" bold="true" box="[1290,1310,1963,1989]" pageId="49" pageNumber="50">E</emphasis>
|
||
. Penis.
|
||
<emphasis id="E6A4EAC0FFD1FFC8EDD0F85035E4F812" bold="true" box="[189,207,1998,2024]" pageId="49" pageNumber="50">F</emphasis>
|
||
. Prostate gland.
|
||
<emphasis id="E6A4EAC0FFD1FFC8ECFEF8503481F812" bold="true" box="[403,426,1998,2024]" pageId="49" pageNumber="50">G</emphasis>
|
||
. Head with perioesophageal ring attached.
|
||
<emphasis id="E6A4EAC0FFD1FFC8EECBF8503696F812" bold="true" box="[934,957,1998,2024]" pageId="49" pageNumber="50">H</emphasis>
|
||
. Perioesophageal ring.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="D46F36D2FFD2FFCBEDD0FEF0345AFDEE" blockId="50.[188,1399,365,532]" pageId="50" pageNumber="51">
|
||
<smallCapsWord id="D289A00EFFD2FFCBEDD0FEF0343FFE72" baselines="385,385" box="[189,276,366,392]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Female" pageId="50" pageNumber="51">FEMALE</smallCapsWord>
|
||
<smallCapsWord id="D289A00EFFD2FFCBEC4DFEED34A5FE7C" baselines="385" box="[288,398,371,390]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="genitalia" pageId="50" pageNumber="51">GENITALIA</smallCapsWord>
|
||
. Glandular oviduct 2.5 times as long as wide, albumen gland longer than capsule gland; bursa copulatrix elongate, ca 2 times as long as wide (Supp. file 2: Table S17); bursal duct shorter than bursa copulatrix; renal oviduct unpigmented, highly coiled with three loops; seminal receptacle elongate, with a short duct, positioned on the distal part of the renal oviduct just above the junction with the bursal duct.
|
||
</paragraph>
|
||
<paragraph id="D46F36D2FFD2FFCBEDD0FDA2377CFD66" blockId="50.[189,1400,572,669]" pageId="50" pageNumber="51">
|
||
<smallCapsWord id="D289A00EFFD2FFCBEDD0FDA23408FDAC" baselines="591,591" box="[189,291,572,598]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Nervous" pageId="50" pageNumber="51">NERVOUS</smallCapsWord>
|
||
<smallCapsWord id="D289A00EFFD2FFCBEC40FDDF3454FDAE" baselines="591" box="[301,383,577,596]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="system" pageId="50" pageNumber="51">SYSTEM</smallCapsWord>
|
||
. Pigmented, elongate (mean RPG ratio = 0.55; see Supp. file 2: Table S15); cerebral ganglia approximately equal in size; pleuro-supraoesophageal connective ca 11 times as long as pleuro-suboesophageal one (
|
||
<figureCitation id="4CEB2A57FFD2FFCBECD5FD1C3761FD66" box="[440,586,642,668]" captionStart="Fig" captionStartId="49.[189,232,1928,1954]" captionTargetBox="[189,1397,900,1884]" captionTargetId="figure-236@49.[189,1398,900,1885]" captionTargetPageId="49" captionText="Fig. 31. Anatomy of the Mercuria veronicae Miller, Khalloufi & Delicado sp. nov. from Spring El Waha, Tozeur, Tunisia. A. Head. B. Ctenidium and osphradium. C. Stomach. D. Head of a male. E. Penis. F. Prostate gland. G. Head with perioesophageal ring attached. H. Perioesophageal ring." figureDoi="http://doi.org/10.5281/zenodo.7867431" httpUri="https://zenodo.org/record/7867431/files/figure.png" pageId="50" pageNumber="51">Fig. 31G–H</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="9CCA6559FFD2FFCBEDD0FD5A3484FB97" pageId="50" pageNumber="51" type="biology_ecology">
|
||
<paragraph id="D46F36D2FFD2FFCBEDD0FD5A34D9FD27" blockId="50.[189,498,707,734]" box="[189,498,707,734]" pageId="50" pageNumber="51">
|
||
<heading id="8F2781BEFFD2FFCBEDD0FD5A34D9FD27" bold="true" box="[189,498,707,734]" fontSize="11" level="3" pageId="50" pageNumber="51" reason="3">
|
||
<emphasis id="E6A4EAC0FFD2FFCBEDD0FD5A34D9FD27" bold="true" box="[189,498,707,734]" pageId="50" pageNumber="51">Ecology and distribution</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="D46F36D2FFD2FFCBEDD0FD6C3484FB97" blockId="50.[188,1400,754,1133]" pageId="50" pageNumber="51">
|
||
The studied samples of this species come from three localities in the
|
||
<collectingRegion id="1614F830FFD2FFCBEE8FFD6D31A3FCF6" box="[994,1160,754,781]" country="Tunisia" name="Tozeur" pageId="50" pageNumber="51">Tozeur region</collectingRegion>
|
||
, north of the Chotts, but its distribution extends to other regions around the Chotts el Gharsa and Djeride (
|
||
<collectingRegion id="1614F830FFD2FFCBE9A3FC8B303CFCD5" box="[1230,1303,789,815]" country="Tunisia" name="Gafsa" pageId="50" pageNumber="51">Gafsa</collectingRegion>
|
||
,
|
||
<collectingRegion id="1614F830FFD2FFCBE84BFC8B3059FCD5" box="[1318,1394,789,815]" country="Tunisia" name="Kebili" pageId="50" pageNumber="51">Kebili</collectingRegion>
|
||
, Nefzaoua and
|
||
<collectingRegion id="1614F830FFD2FFCBEC08FCA73498FCAE" box="[357,435,825,852]" country="Tunisia" name="Gabes" pageId="50" pageNumber="51">Gabes</collectingRegion>
|
||
). It occurs in streams, springs and irrigation ditches of the oasis areas. This species is present at a high density (more than 3000 ind./m
|
||
<superScript id="23A59B9AFFD2FFCBEE7AFCC5360BFC93" attach="left" box="[791,800,859,873]" fontSize="6" pageId="50" pageNumber="51">2</superScript>
|
||
) on the edge of the streams, where the flow is low and the substrate muddy, and attached to aquatic plants. The main streams of Tamarza (Oued Frid) and Echbika (Oued Echbika) are highly saline (conductivity 8667 and 3945 µS/cm, respectively) and flow over an alluvial and clayey-sandy substrate of the Plio-Quaternary, with a predominance of sulphate-sodium, gypsum and phosphorites (
|
||
<bibRefCitation id="B0414B23FFD2FFCBEFBDFC773676FBF9" author="Abidi B." box="[720,861,1001,1027]" pageId="50" pageNumber="51" refId="ref27634" refString="Abidi B. 2007. Caracterisation Hydrogeologique, Geochimique et Isotopique des Systemes Aquiferes du Synclinal de Tamerza et de la Plaine de Chott El Gharsa (Sud Ouest Tunisien). PhD thesis, University of Sfax, Tunisia." type="book" year="2007">Abidi 2007</bibRefCitation>
|
||
). Co-occurring species are
|
||
<taxonomicName id="13D04D51FFD2FFCBE9C9FC773044FBF9" baseAuthorityName="O. F. Muller" baseAuthorityYear="1774" box="[1188,1391,1001,1027]" class="Gastropoda" family="Lymnaeidae" genus="Galba" kingdom="Animalia" pageId="50" pageNumber="51" phylum="Mollusca" rank="species" species="truncatula">
|
||
<emphasis id="E6A4EAC0FFD2FFCBE9C9FC773044FBF9" box="[1188,1391,1001,1027]" italics="true" pageId="50" pageNumber="51">Galba truncatula</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="13D04D51FFD2FFCBEDD0FB92374AFBDD" authority="(Draparnaud, 1805)" baseAuthorityName="Draparnaud" baseAuthorityYear="1805" box="[189,609,1036,1063]" class="Gastropoda" family="Physidae" genus="Physella" kingdom="Animalia" pageId="50" pageNumber="51" phylum="Mollusca" rank="species" species="acuta">
|
||
<emphasis id="E6A4EAC0FFD2FFCBEDD0FB923447FBDC" box="[189,364,1036,1062]" italics="true" pageId="50" pageNumber="51">Physella acuta</emphasis>
|
||
(
|
||
<bibRefCitation id="B0414B23FFD2FFCBECEDFB923772FBDD" author="Draparnaud J. P. R." box="[384,601,1036,1063]" pageId="50" pageNumber="51" refId="ref29362" refString="Draparnaud J. P. R. 1805. Histoire naturelle des Mollusques terrestres et fluviatiles de la France. Chez Louis Colas, Paris. https: // doi. org / 10.5962 / bhl. title. 12856" type="book" year="1805">Draparnaud, 1805</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
(in low abundance),
|
||
<taxonomicName id="13D04D51FFD2FFCBEE0CFB92317BFBDC" box="[865,1104,1036,1062]" class="Gastropoda" family="Cochliopidae" genus="Eupaludestrina" kingdom="Animalia" order="Littorinimorpha" pageId="50" pageNumber="51" phylum="Mollusca" rank="species" species="undetermined">
|
||
<emphasis id="E6A4EAC0FFD2FFCBEE0CFB92313DFBDC" box="[865,1046,1036,1062]" italics="true" pageId="50" pageNumber="51">Eupaludestrina</emphasis>
|
||
spp.
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="13D04D51FFD2FFCBE90DFB923480FBB3" authority="(O. F. Muller, 1774)" baseAuthorityName="O. F. Muller" baseAuthorityYear="1774" class="Gastropoda" family="Thiaridae" genus="Melanoides" kingdom="Animalia" pageId="50" pageNumber="51" phylum="Mollusca" rank="species" species="tuberculata">
|
||
<emphasis id="E6A4EAC0FFD2FFCBE90DFB923053FBDC" box="[1120,1400,1036,1062]" italics="true" pageId="50" pageNumber="51">Melanoides tuberculata</emphasis>
|
||
(O.F. Müller, 1774)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="13D04D51FFD2FFCBECD4FBB1367CFBB3" authority="Pallary, 1916" authorityName="Pallary" authorityYear="1916" box="[441,855,1071,1097]" class="Gastropoda" family="Melanopsidae" genus="Melanopsis" kingdom="Animalia" order="Neogastropoda" pageId="50" pageNumber="51" phylum="Mollusca" rank="species" species="douvillei">
|
||
<emphasis id="E6A4EAC0FFD2FFCBECD4FBB1379AFBB3" box="[441,689,1071,1097]" italics="true" pageId="50" pageNumber="51">Melanopsis douvillei</emphasis>
|
||
Pallary, 1916
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="13D04D51FFD2FFCBEEFFFBB13053FBB0" authority="(Linnaeus, 1758)" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[914,1400,1071,1098]" class="Gastropoda" family="Melanopsidae" genus="Melanopsis" kingdom="Animalia" order="Neogastropoda" pageId="50" pageNumber="51" phylum="Mollusca" rank="species" species="praemorsa">
|
||
<emphasis id="E6A4EAC0FFD2FFCBEEFFFBB13189FBB3" box="[914,1186,1071,1097]" italics="true" pageId="50" pageNumber="51">Melanopsis praemorsa</emphasis>
|
||
(Linnaeus, 1758)
|
||
</taxonomicName>
|
||
(in high abundance).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="9CCA6559FFD2FFCAEDD0FB0A348DFE90" lastPageId="51" lastPageNumber="52" pageId="50" pageNumber="51" type="discussion">
|
||
<paragraph id="D46F36D2FFD2FFCBEDD0FB0A3404FB54" blockId="50.[189,303,1172,1198]" box="[189,303,1172,1198]" pageId="50" pageNumber="51">
|
||
<heading id="8F2781BEFFD2FFCBEDD0FB0A3404FB54" bold="true" box="[189,303,1172,1198]" fontSize="11" level="3" pageId="50" pageNumber="51" reason="3">
|
||
<emphasis id="E6A4EAC0FFD2FFCBEDD0FB0A3404FB54" bold="true" box="[189,303,1172,1198]" pageId="50" pageNumber="51">Remarks</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="D46F36D2FFD2FFCBEDD0FB5C3010FA76" blockId="50.[188,1401,1218,1421]" pageId="50" pageNumber="51">
|
||
Intraspecies morphological differences were observed mainly in the penial features of males from the same population (
|
||
<figureCitation id="4CEB2A57FFD2FFCBECFAFB7B3701FB05" box="[407,554,1253,1279]" captionStart="Fig" captionStartId="49.[189,232,1928,1954]" captionTargetBox="[189,1397,900,1884]" captionTargetId="figure-236@49.[189,1398,900,1885]" captionTargetPageId="49" captionText="Fig. 31. Anatomy of the Mercuria veronicae Miller, Khalloufi & Delicado sp. nov. from Spring El Waha, Tozeur, Tunisia. A. Head. B. Ctenidium and osphradium. C. Stomach. D. Head of a male. E. Penis. F. Prostate gland. G. Head with perioesophageal ring attached. H. Perioesophageal ring." figureDoi="http://doi.org/10.5281/zenodo.7867431" httpUri="https://zenodo.org/record/7867431/files/figure.png" pageId="50" pageNumber="51">Fig. 31D–E</figureCitation>
|
||
). Some individuals have a much-atrophied penis. Such variation may be parasite-induced, similar to the case reported in another species of
|
||
<taxonomicName id="13D04D51FFD2FFCBE979FA97342EFABC" authority="(Boulaassafer et al. 2018)" baseAuthorityName="Boulaassafer" baseAuthorityYear="2018" class="Gastropoda" family="Hydrobiidae" genus="Mercuria" kingdom="Animalia" order="Littorinimorpha" pageId="50" pageNumber="51" phylum="Mollusca" rank="genus">
|
||
<emphasis id="E6A4EAC0FFD2FFCBE979FA9731A9FAD8" box="[1044,1154,1289,1314]" italics="true" pageId="50" pageNumber="51">Mercuria</emphasis>
|
||
(
|
||
<bibRefCitation id="B0414B23FFD2FFCBE9F9FA9635D6FABC" author="Boulaassafer K. & Ghamizi M. & Delicado D." pageId="50" pageNumber="51" pagination="95 - 128" refId="ref28648" refString="Boulaassafer K., Ghamizi M. & Delicado D. 2018. The genus Mercuria Boeters, 1971 in Morocco: first molecular phylogeny of the genus and description of two new species (Caenogastropoda, Truncatelloidea, Hydrobiidae). ZooKeys 782: 95 - 128. https: // doi. org / 10.3897 / zookeys. 779.26797" type="journal article" year="2018">
|
||
Boulaassafer
|
||
<emphasis id="E6A4EAC0FFD2FFCBE855FA973044FAD8" box="[1336,1391,1288,1314]" italics="true" pageId="50" pageNumber="51">et al</emphasis>
|
||
. 2018
|
||
</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
. Parasitized females also presented an atrophied pallial oviduct, making it difficult to conduct a complete anatomical study. The pigmentation of the penis is also very variable, ranging from very slight to pronounced dark pigmentation at the junction between the appendix and distal end of the penis.
|
||
</paragraph>
|
||
<paragraph id="D46F36D2FFD2FFCBEDD0FA2A3029F86D" blockId="50.[188,1399,1460,1944]" pageId="50" pageNumber="51">
|
||
<taxonomicName id="13D04D51FFD2FFCBEDD0FA2A3482FA37" authority="Miller & Delicado & García-Guerrero & Khalloufi & Ramos, 2023" authorityName="Miller & Delicado & García-Guerrero & Khalloufi & Ramos" authorityYear="2023" box="[189,425,1460,1485]" class="Gastropoda" family="Hydrobiidae" genus="Mercuria" kingdom="Animalia" order="Littorinimorpha" pageId="50" pageNumber="51" phylum="Mollusca" rank="species" species="veronicae" status="sp. nov.">
|
||
<emphasis id="E6A4EAC0FFD2FFCBEDD0FA2A3482FA37" box="[189,425,1460,1485]" italics="true" pageId="50" pageNumber="51">Mercuria veronicae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="FD9757BBFFD2FFCBECD8FA2A3738FA34" box="[437,531,1460,1486]" pageId="50" pageNumber="51" rank="species">sp. nov.</taxonomicNameLabel>
|
||
differs from the phylogenetically closely related species
|
||
<taxonomicName id="13D04D51FFD2FFCBE9B9FA2A305DFA34" authority="Miller & Delicado & García-Guerrero & Khalloufi & Ramos, 2023" authorityName="Miller & Delicado & García-Guerrero & Khalloufi & Ramos" authorityYear="2023" box="[1236,1398,1460,1486]" class="Gastropoda" family="Hydrobiidae" genus="Mercuria" kingdom="Animalia" order="Littorinimorpha" pageId="50" pageNumber="51" phylum="Mollusca" rank="species" species="lupiaensis" status="sp. nov.">
|
||
<emphasis id="E6A4EAC0FFD2FFCBE9B9FA2A305DFA34" box="[1236,1398,1460,1486]" italics="true" pageId="50" pageNumber="51">M. lupiaensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="FD9757BBFFD2FFCBEDD0FA49343DFA0B" box="[189,278,1495,1521]" pageId="50" pageNumber="51" rank="species">sp. nov.</taxonomicNameLabel>
|
||
and
|
||
<taxonomicName id="13D04D51FFD2FFCBEC26FA4634CDFA0B" baseAuthorityName="Paladilhe" baseAuthorityYear="1869" box="[331,486,1495,1521]" class="Gastropoda" family="Hydrobiidae" genus="Mercuria" kingdom="Animalia" order="Littorinimorpha" pageId="50" pageNumber="51" phylum="Mollusca" rank="species" species="melitensis">
|
||
<emphasis id="E6A4EAC0FFD2FFCBEC26FA4634CDFA0B" box="[331,486,1495,1521]" italics="true" pageId="50" pageNumber="51">M. melitensis</emphasis>
|
||
</taxonomicName>
|
||
by having a high-spired shell and a more elongated bursa copulatrix compared with
|
||
<taxonomicName id="13D04D51FFD2FFCBED95FA6534BDF9EE" authorityName="Miller & Delicado & García-Guerrero & Khalloufi & Ramos" authorityYear="2023" box="[248,406,1530,1556]" class="Gastropoda" family="Hydrobiidae" genus="Mercuria" kingdom="Animalia" order="Littorinimorpha" pageId="50" pageNumber="51" phylum="Mollusca" rank="species" species="lupiaensis">
|
||
<emphasis id="E6A4EAC0FFD2FFCBED95FA6534BDF9EE" box="[248,406,1530,1556]" italics="true" pageId="50" pageNumber="51">M. lupiaensis</emphasis>
|
||
</taxonomicName>
|
||
and thinner inner and outer shell lips compared with the characteristically thick ones of
|
||
<taxonomicName id="13D04D51FFD2FFCBEDB1F9803777F9C2" authority="(Gloer et al. 2015)" baseAuthorityName="Gloer" baseAuthorityYear="2015" box="[220,604,1565,1592]" class="Gastropoda" family="Hydrobiidae" genus="Mercuria" kingdom="Animalia" order="Littorinimorpha" pageId="50" pageNumber="51" phylum="Mollusca" rank="species" species="melitensis">
|
||
<emphasis id="E6A4EAC0FFD2FFCBEDB1F9803453F9CD" box="[220,376,1565,1591]" italics="true" pageId="50" pageNumber="51">M. melitensis</emphasis>
|
||
(
|
||
<bibRefCitation id="B0414B23FFD2FFCBECEAF9833778F9C2" author="Gloer P. & Boeters H. D. & Walther F." box="[391,595,1565,1592]" pageId="50" pageNumber="51" pagination="279 - 291" refId="ref29897" refString="Gloer P., Boeters H. D. & Walther F. 2015. Species of the genus Mercuria Boeters, 1971 (Caenogastropoda: Truncatelloidea: Hydrobiidae) from the European Mediterranean region, Morocco and Madeira, with descriptions of new species. Folia Malacologica 23: 279 - 291. https: // doi. org / 10.12657 / folmal. 023.024" type="journal article" year="2015">
|
||
Glöer
|
||
<emphasis id="E6A4EAC0FFD2FFCBECBFF9803727F9CD" box="[466,524,1565,1591]" italics="true" pageId="50" pageNumber="51">et al.</emphasis>
|
||
2015
|
||
</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
.
|
||
<taxonomicName id="13D04D51FFD2FFCBEF08F9803666F9CD" authorityName="Miller & Delicado & García-Guerrero & Khalloufi & Ramos" authorityYear="2023" box="[613,845,1566,1591]" class="Gastropoda" family="Hydrobiidae" genus="Mercuria" kingdom="Animalia" order="Littorinimorpha" pageId="50" pageNumber="51" phylum="Mollusca" rank="species" species="veronicae">
|
||
<emphasis id="E6A4EAC0FFD2FFCBEF08F9803666F9CD" box="[613,845,1566,1591]" italics="true" pageId="50" pageNumber="51">Mercuria veronicae</emphasis>
|
||
</taxonomicName>
|
||
also has a long, triangular and more pigmented penial appendix compared with the short (to very short), ovate and slightly pigmented or unpigmented one in
|
||
<emphasis id="E6A4EAC0FFD2FFCBEC78F9FA3494F984" box="[277,447,1636,1662]" italics="true" pageId="50" pageNumber="51">
|
||
<taxonomicName id="13D04D51FFD2FFCBEC78F9FA3490F984" authorityName="Miller & Delicado & García-Guerrero & Khalloufi & Ramos" authorityYear="2023" box="[277,443,1636,1662]" class="Gastropoda" family="Hydrobiidae" genus="Mercuria" kingdom="Animalia" order="Littorinimorpha" pageId="50" pageNumber="51" phylum="Mollusca" rank="species" species="lupiaensis">M. lupiaensis</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
In
|
||
<taxonomicName id="13D04D51FFD2FFCBEC80F9FA37A5F984" box="[493,654,1636,1662]" class="Gastropoda" family="Hydrobiidae" genus="Mercuria" kingdom="Animalia" order="Littorinimorpha" pageId="50" pageNumber="51" phylum="Mollusca" rank="species" species="melitensis">
|
||
<emphasis id="E6A4EAC0FFD2FFCBEC80F9FA37A5F984" box="[493,654,1636,1662]" italics="true" pageId="50" pageNumber="51">M. melitensis</emphasis>
|
||
</taxonomicName>
|
||
, the penial appendix is only as long as the distal end of the penis. Compared with other geographically proximate species,
|
||
<taxonomicName id="13D04D51FFD2FFCBEEF0F916311CF95B" authorityName="Miller & Delicado & García-Guerrero & Khalloufi & Ramos" authorityYear="2023" box="[925,1079,1672,1697]" class="Gastropoda" family="Hydrobiidae" genus="Mercuria" kingdom="Animalia" order="Littorinimorpha" pageId="50" pageNumber="51" phylum="Mollusca" rank="species" species="veronicae">
|
||
<emphasis id="E6A4EAC0FFD2FFCBEEF0F916311CF95B" box="[925,1079,1672,1697]" italics="true" pageId="50" pageNumber="51">M. veronicae</emphasis>
|
||
</taxonomicName>
|
||
resembles
|
||
<taxonomicName id="13D04D51FFD2FFCBE9D7F9163480F93F" authority="(Bourguignat, 1862)" baseAuthorityName="Bourguignat" baseAuthorityYear="1862" class="Gastropoda" family="Hydrobiidae" genus="Mercuria" kingdom="Animalia" order="Littorinimorpha" pageId="50" pageNumber="51" phylum="Mollusca" rank="species" species="pycnocheilia">
|
||
<emphasis id="E6A4EAC0FFD2FFCBE9D7F916305CF95B" box="[1210,1399,1671,1697]" italics="true" pageId="50" pageNumber="51">M. pycnocheilia</emphasis>
|
||
(Bourguignat, 1862)
|
||
</taxonomicName>
|
||
in shell shape, although the latter presents a thicker inner shell lip (see
|
||
<bibRefCitation id="B0414B23FFD2FFCBE99CF93435D2F91D" author="Gloer P. & Bouzid S. & Boeters H. D." pageId="50" pageNumber="51" pagination="1 - 22" refId="ref29824" refString="Gloer P., Bouzid S. & Boeters H. D. 2010. Revision of the genera Pseudamnicola Paulucci 1878 and Mercuria Boeters 1971 from Algeria with particular emphasis on museum collections (Gastropoda: Prosobranchia: Hydrobiidae). Archiv fur Molluskenkunde 139: 1 - 22. https: // doi. org / 10.1127 / arch. moll / 1869 - 0963 / 139 / 001 - 022" type="journal article" year="2010">
|
||
Glöer
|
||
<emphasis id="E6A4EAC0FFD2FFCBE851F935305CF93E" box="[1340,1399,1706,1732]" italics="true" pageId="50" pageNumber="51">et al.</emphasis>
|
||
2010
|
||
</bibRefCitation>
|
||
) and the former, has smaller-sized shells. In addition,
|
||
<bibRefCitation id="B0414B23FFD2FFCBEE11F9533172F91D" author="Gloer P. & Bouzid S. & Boeters H. D." box="[892,1113,1741,1767]" pageId="50" pageNumber="51" pagination="1 - 22" refId="ref29824" refString="Gloer P., Bouzid S. & Boeters H. D. 2010. Revision of the genera Pseudamnicola Paulucci 1878 and Mercuria Boeters 1971 from Algeria with particular emphasis on museum collections (Gastropoda: Prosobranchia: Hydrobiidae). Archiv fur Molluskenkunde 139: 1 - 22. https: // doi. org / 10.1127 / arch. moll / 1869 - 0963 / 139 / 001 - 022" type="journal article" year="2010">
|
||
Glöer
|
||
<emphasis id="E6A4EAC0FFD2FFCBEEA5F950312FF91D" box="[968,1028,1741,1767]" italics="true" pageId="50" pageNumber="51">et al.</emphasis>
|
||
(2010)
|
||
</bibRefCitation>
|
||
and
|
||
<bibRefCitation id="B0414B23FFD2FFCBE9F9F9533006F91D" author="Gloer P." box="[1172,1325,1741,1767]" pageId="50" pageNumber="51" refId="ref29792" refString="Gloer P. 2019. The Freshwater Gastropods of the West-Paleartics, Vol. 1: Fresh- and Brackish Waters except Springs and Subterranean Snails. Biodiversity Research Lab, Hetlingen." type="book" year="2019">Glöer (2019)</bibRefCitation>
|
||
noted the acute apex of
|
||
<taxonomicName id="13D04D51FFD2FFCBECE8F96F361CF8F1" authority="(Bourguignat, 1862)" baseAuthorityName="Bourguignat" baseAuthorityYear="1862" box="[389,823,1776,1803]" class="Gastropoda" family="Hydrobiidae" genus="Mercuria" kingdom="Animalia" order="Littorinimorpha" pageId="50" pageNumber="51" phylum="Mollusca" rank="species" species="pycnocheilia">
|
||
<emphasis id="E6A4EAC0FFD2FFCBECE8F96F376BF8F0" box="[389,576,1776,1802]" italics="true" pageId="50" pageNumber="51">M. pycnocheilia</emphasis>
|
||
(Bourguignat, 1862)
|
||
</taxonomicName>
|
||
, which is not present in
|
||
<emphasis id="E6A4EAC0FFD2FFCBE926F96F31C0F8F0" box="[1099,1259,1777,1802]" italics="true" pageId="50" pageNumber="51">
|
||
<taxonomicName id="13D04D51FFD2FFCBE926F96F31CDF8F0" authorityName="Miller & Delicado & García-Guerrero & Khalloufi & Ramos" authorityYear="2023" box="[1099,1254,1777,1802]" class="Gastropoda" family="Hydrobiidae" genus="Mercuria" kingdom="Animalia" order="Littorinimorpha" pageId="50" pageNumber="51" phylum="Mollusca" rank="species" species="veronicae">M. veronicae</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
Shell shape features also distinguish the new species from
|
||
<taxonomicName id="13D04D51FFD2FFCBEF8EF88A3008F8D4" authority="(Letourneux & Bourguignat, 1887)" baseAuthorityName="Letourneux & Bourguignat" baseAuthorityYear="1887" box="[739,1315,1812,1838]" class="Gastropoda" family="Hydrobiidae" genus="Mercuria" kingdom="Animalia" order="Littorinimorpha" pageId="50" pageNumber="51" phylum="Mollusca" rank="species" species="globulina">
|
||
<emphasis id="E6A4EAC0FFD2FFCBEF8EF88A3651F8D4" box="[739,890,1812,1838]" italics="true" pageId="50" pageNumber="51">M. globulina</emphasis>
|
||
(
|
||
<bibRefCitation id="B0414B23FFD2FFCBEEE6F88A3031F8D4" author="Letourneux A. & Bourguignat J. R." box="[907,1306,1812,1838]" pageId="50" pageNumber="51" refId="ref30232" refString="Letourneux A. & Bourguignat J. R. 1887. Prodrome de la Malacologie terrestre et fluviatile de la Tunisie. Imprimerie nationale, Paris. https: // doi. org / 10.5962 / bhl. title. 132280" type="book" year="1887">Letourneux & Bourguignat, 1887</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, which has a larger, more slender shell, and
|
||
<taxonomicName id="13D04D51FFD2FFCBEF09F8A631BCF8AB" authority="Gloer, Bouzid & Boeters, 2010" authorityName="Gloer, Bouzid & Boeters" authorityYear="2010" box="[612,1175,1847,1873]" class="Gastropoda" family="Hydrobiidae" genus="Mercuria" kingdom="Animalia" order="Littorinimorpha" pageId="50" pageNumber="51" phylum="Mollusca" rank="species" species="bourguignati">
|
||
<emphasis id="E6A4EAC0FFD2FFCBEF09F8A63609F8AB" box="[612,802,1847,1873]" italics="true" pageId="50" pageNumber="51">M. bourguignati</emphasis>
|
||
<bibRefCitation id="B0414B23FFD2FFCBEE44F8A931BCF8AB" author="Gloer P. & Bouzid S. & Boeters H. D." box="[809,1175,1847,1873]" pageId="50" pageNumber="51" pagination="1 - 22" refId="ref29824" refString="Gloer P., Bouzid S. & Boeters H. D. 2010. Revision of the genera Pseudamnicola Paulucci 1878 and Mercuria Boeters 1971 from Algeria with particular emphasis on museum collections (Gastropoda: Prosobranchia: Hydrobiidae). Archiv fur Molluskenkunde 139: 1 - 22. https: // doi. org / 10.1127 / arch. moll / 1869 - 0963 / 139 / 001 - 022" type="journal article" year="2010">Glöer, Bouzid & Boeters, 2010</bibRefCitation>
|
||
</taxonomicName>
|
||
, which has a larger shell and smaller aperture (about half the shell length in
|
||
<taxonomicName id="13D04D51FFD2FFCBEE29F8C5312AF88E" authorityName="Gloer, Bouzid & Boeters" authorityYear="2010" box="[836,1025,1882,1908]" class="Gastropoda" family="Hydrobiidae" genus="Mercuria" kingdom="Animalia" order="Littorinimorpha" pageId="50" pageNumber="51" phylum="Mollusca" rank="species" species="bourguignati">
|
||
<emphasis id="E6A4EAC0FFD2FFCBEE29F8C5312AF88E" box="[836,1025,1882,1908]" italics="true" pageId="50" pageNumber="51">M. bourguignati</emphasis>
|
||
</taxonomicName>
|
||
vs about three quarters the shell length in
|
||
<taxonomicName id="13D04D51FFD2FFCBEC41F8E034EDF86D" authorityName="Miller & Delicado & García-Guerrero & Khalloufi & Ramos" authorityYear="2023" box="[300,454,1918,1943]" class="Gastropoda" family="Hydrobiidae" genus="Mercuria" kingdom="Animalia" order="Littorinimorpha" pageId="50" pageNumber="51" phylum="Mollusca" rank="species" species="veronicae">
|
||
<emphasis id="E6A4EAC0FFD2FFCBEC41F8E034EDF86D" box="[300,454,1918,1943]" italics="true" pageId="50" pageNumber="51">M. veronicae</emphasis>
|
||
</taxonomicName>
|
||
). The anatomy of these geographically proximate species is unknown.
|
||
</paragraph>
|
||
<paragraph id="D46F36D2FFD3FFCAEDD0FE97348DFE90" blockId="51.[189,1399,265,362]" pageId="51" pageNumber="52">
|
||
According to our sequence divergence estimates of COI,
|
||
<taxonomicName id="13D04D51FFD3FFCAEEE8FE94310FFED9" authority="Miller & Delicado & García-Guerrero & Khalloufi & Ramos, 2023" authorityName="Miller & Delicado & García-Guerrero & Khalloufi & Ramos" authorityYear="2023" box="[901,1060,266,291]" class="Gastropoda" family="Hydrobiidae" genus="Mercuria" kingdom="Animalia" order="Littorinimorpha" pageId="51" pageNumber="52" phylum="Mollusca" rank="species" species="veronicae" status="sp. nov.">
|
||
<emphasis id="E6A4EAC0FFD3FFCAEEE8FE94310FFED9" box="[901,1060,266,291]" italics="true" pageId="51" pageNumber="52">M. veronicae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="FD9757BBFFD3FFCAE95DFE9431BBFEDE" box="[1072,1168,266,292]" pageId="51" pageNumber="52" rank="species">sp. nov.</taxonomicNameLabel>
|
||
is most similar to
|
||
<taxonomicName id="13D04D51FFD3FFCAEDD0FEB33477FEBC" authority="Miller & Delicado & García-Guerrero & Khalloufi & Ramos, 2023" authorityName="Miller & Delicado & García-Guerrero & Khalloufi & Ramos" authorityYear="2023" box="[189,348,300,326]" class="Gastropoda" family="Hydrobiidae" genus="Mercuria" kingdom="Animalia" order="Littorinimorpha" pageId="51" pageNumber="52" phylum="Mollusca" rank="species" species="lupiaensis" status="sp. nov.">
|
||
<emphasis id="E6A4EAC0FFD3FFCAEDD0FEB33477FEBC" box="[189,348,300,326]" italics="true" pageId="51" pageNumber="52">M. lupiaensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="FD9757BBFFD3FFCAEC0BFEB334EAFEBD" box="[358,449,301,327]" pageId="51" pageNumber="52" rank="species">sp. nov.</taxonomicNameLabel>
|
||
(average divergence of 3.9%) and diverges most with
|
||
<taxonomicName id="13D04D51FFD3FFCAE920FEB331DCFEBC" authorityName="Boulaassafer, Ghamizi & Delicado" authorityYear="2018" box="[1101,1271,300,326]" class="Gastropoda" family="Hydrobiidae" genus="Mercuria" kingdom="Animalia" order="Littorinimorpha" pageId="51" pageNumber="52" phylum="Mollusca" rank="species" species="tensiftensis">
|
||
<emphasis id="E6A4EAC0FFD3FFCAE920FEB331DCFEBC" box="[1101,1271,300,326]" italics="true" pageId="51" pageNumber="52">M. tensiftensis</emphasis>
|
||
</taxonomicName>
|
||
(by 8.4%) (
|
||
<bibRefCitation id="B0414B23FFD3FFCAEDA8FED134B2FE90" author="Miller J. P. & Delicado D. & Garcia-Guerrero F. & Ramos M. A." box="[197,409,335,362]" pageId="51" pageNumber="52" pagination="107524" refId="ref30353" refString="Miller J. P., Delicado D., Garcia-Guerrero F. & Ramos M. A. 2022. Recurrent founder-event speciation across the Mediterranean likely shaped the species diversity and geographic distribution of the freshwater snail genus Mercuria Boeters, 1971 (Caenogastropoda: Hydrobiidae). Molecular Phylogenetics and Evolution 173: 107524. https: // doi. org / 10.1016 / j. ympev. 2022.107524" type="journal article" year="2022">
|
||
Miller
|
||
<emphasis id="E6A4EAC0FFD3FFCAEC7AFECE3479FE93" box="[279,338,335,361]" italics="true" pageId="51" pageNumber="52">et al.</emphasis>
|
||
2022
|
||
</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |