265 lines
29 KiB
XML
265 lines
29 KiB
XML
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<mods:title id="853B33837CDB9790A3EA39A4BEE793AF">Seven new species of the earthworm genus Metaphire Sims & Easton, 1972 from Thailand (Clitellata: Megascolecidae)</mods:title>
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<mods:namePart id="64DAA9311A1B7CA6B461FBB982D5D23D">Bantaowong, Ueangfa</mods:namePart>
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<mods:namePart id="68FD93A23DD07BB8084B56AEBB0F414A">Chanabun, Ratmanee</mods:namePart>
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<mods:namePart id="B1EE672B7D7DE526876085C411FE24A7">James, Samuel W.</mods:namePart>
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<mods:namePart id="782E89ECFAE7AF75C241C425EEB25F2D">Panha, Somsak</mods:namePart>
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<mods:title id="879E80B78C4FB3FCB63B25E9A6B68894">Zootaxa</mods:title>
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<mods:date id="CCA42124A620CC107A28031E3E62200E">2016</mods:date>
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<mods:number id="97A834EB89AE2C9B3E171A14A4EF5442">4117</mods:number>
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<treatment id="5C1D87F3FF824976EDDA7294FC00FC67" ID-DOI="http://doi.org/10.5281/zenodo.5621297" ID-GBIF-Taxon="120733315" ID-Zenodo-Dep="5621297" LSID="urn:lsid:plazi:treatment:5C1D87F3FF824976EDDA7294FC00FC67" httpUri="http://treatment.plazi.org/id/5C1D87F3FF824976EDDA7294FC00FC67" lastPageId="5" lastPageNumber="68" pageId="4" pageNumber="67">
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<subSubSection id="9CAE656EFF824977EDDA7294FEE2FE45" pageId="4" pageNumber="67" type="nomenclature">
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<paragraph id="D40B36E5FF824977EDDA7294FDE4FE24" blockId="4.[151,622,404,463]" box="[151,622,404,430]" pageId="4" pageNumber="67">
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<heading id="8F438189FF824977EDDA7294FDE4FE24" bold="true" box="[151,622,404,430]" fontSize="11" level="1" pageId="4" pageNumber="67" reason="1">
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<taxonomicName id="13B44D66FF824977EDDA7294FDE4FE24" authority="Beddard, 1900" authorityName="Beddard" authorityYear="1900" box="[151,622,404,430]" class="Clitellata" family="Megascolecidae" genus="Metaphire" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="4" pageNumber="67" phylum="Annelida" rank="species" species="pulauensis">
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<emphasis id="E6C0EAF7FF824977EDDA7294FDE4FE24" bold="true" box="[151,622,404,430]" pageId="4" pageNumber="67">
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<emphasis id="E6C0EAF7FF824977EDDA7294FE14FE24" bold="true" box="[151,414,404,430]" italics="true" pageId="4" pageNumber="67">Metaphire pulauensis</emphasis>
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(
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<bibRefCitation id="B0254B14FF824977ECE17294FDECFE24" author="Beddard" box="[428,614,404,430]" pageId="4" pageNumber="67" refString="Beddard, F. E. (1900) On the earthworms collected during the " Skeat Expedition " to the Malay Peninsula, 1899 - 1900. Proceedings of the Zoological Society of London, 1900, 891 - 911." type="proceedings paper" year="1900">Beddard, 1900</bibRefCitation>
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)
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||
</emphasis>
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||
</taxonomicName>
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</heading>
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</paragraph>
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<paragraph id="D40B36E5FF824977EDDA72B6FEE2FE45" blockId="4.[151,622,404,463]" box="[151,360,438,463]" pageId="4" pageNumber="67">
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(
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||
<figureCitation id="4C8F2A60FF824977EDD372B6FF75FE45" box="[158,255,438,463]" captionStart="FIGURE 3" captionStartId="4.[151,250,1766,1788]" captionTargetBox="[155,1427,654,1744]" captionTargetId="figure@4.[151,1435,646,1745]" captionTargetPageId="4" captionText="FIGURE 3. External and internal morphology of Metaphire pulauensis (Beddard, 1900), lectotype (NHMUK 1924.3.1.214). A. External ventral view. B. Internal dorsal view. C. Spermatheca; arrowhead: location of spermathecal pore." httpUri="https://zenodo.org/record/259124/files/figure.png" pageId="4" pageNumber="67">Figure 3</figureCitation>
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,
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<tableCitation id="9936035EFF824977EC4172B6FED7FE45" box="[268,349,438,463]" captionStart="TABLE 1" captionStartId="6.[151,239,1427,1450]" captionTargetBox="[159,1307,1509,2017]" captionTargetPageId="6" captionText="TABLE 1. Comparison of morphological characters in the Metaphire pulauensis species group. The comma is used to separate body length and width, and setal counts." httpUri="http://table.plazi.org/id/80CB666DFF804975EDDA7693FD29FA40" pageId="4" pageNumber="67" tableUuid="80CB666DFF804975EDDA7693FD29FA40">Table 1</tableCitation>
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)
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</paragraph>
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||
</subSubSection>
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<subSubSection id="9CAE656EFF824977EDDA72FFFD7DFDD8" pageId="4" pageNumber="67" type="reference_group">
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<paragraph id="D40B36E5FF824977EDDA72FFFD20FD9E" blockId="4.[151,858,510,594]" box="[151,682,510,532]" pageId="4" pageNumber="67">
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<treatmentCitationGroup id="F4A411CBFF824977EDDA72FFFD20FD9E" box="[151,682,510,532]" pageId="4" pageNumber="67">
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<taxonomicName id="13B44D66FF824977EDDA72FFFDBAFD9E" authority="Beddard, 1900: 904" authorityName="Beddard" authorityPageNumber="904" authorityYear="1900" box="[151,560,510,532]" class="Clitellata" family="Megascolecidae" genus="Amyntas" kingdom="Animalia" order="Opisthopora" pageId="4" pageNumber="67" phylum="Annelida" rank="species" species="pulauensis">
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<emphasis id="E6C0EAF7FF824977EDDA72FFFEEBFD9E" box="[151,353,510,532]" italics="true" pageId="4" pageNumber="67">Amyntas pulauensis</emphasis>
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<treatmentCitation id="551510F4FF824977EC2A72FEFDBAFD9E" author="Beddard" box="[359,560,510,532]" page="904" pageId="4" pageNumber="67" year="1900">
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<bibRefCitation id="B0254B14FF824977EC2A72FEFE75FD9E" author="Beddard" box="[359,511,510,532]" pageId="4" pageNumber="67" refString="Beddard, F. E. (1900) On the earthworms collected during the " Skeat Expedition " to the Malay Peninsula, 1899 - 1900. Proceedings of the Zoological Society of London, 1900, 891 - 911." type="proceedings paper" year="1900">Beddard, 1900</bibRefCitation>
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: 904
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</treatmentCitation>
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||
</taxonomicName>
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||
–905,
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<figureCitation id="4C8F2A60FF824977EF2372FEFD2DFD9E" box="[622,679,510,532]" captionStart="FIGURE 5" captionStartId="7.[151,250,1752,1774]" captionTargetBox="[162,1432,629,1723]" captionTargetId="figure@7.[150,1436,623,1730]" captionTargetPageId="7" captionText="FIGURE 5. External and internal morphology of Metaphire trangensis sp. n., holotype (CUMZ 3375). A. External ventral view. B. Internal dorsal view. C. Spermatheca; arrowhead: location of spermathecal pore." httpUri="https://zenodo.org/record/259126/files/figure.png" pageId="4" pageNumber="67">Fig.5</figureCitation>
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||
.
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||
</treatmentCitationGroup>
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||
</paragraph>
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<paragraph id="D40B36E5FF824977EDDA711DFD7DFDD8" blockId="4.[151,858,510,594]" pageId="4" pageNumber="67">
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<treatmentCitationGroup id="F4A411CBFF824977EDDA711DFD7DFDD8" pageId="4" pageNumber="67">
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<taxonomicName id="13B44D66FF824977EDDA711DFE68FDB9" authority="Beddard" authorityName="Beddard" box="[151,482,541,563]" class="Clitellata" family="Megascolecidae" genus="Pheretima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="4" pageNumber="67" phylum="Annelida" rank="species" species="pulauensis">
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||
<emphasis id="E6C0EAF7FF824977EDDA711DFEF8FDB9" box="[151,370,541,563]" italics="true" pageId="4" pageNumber="67">Pheretima pulauensis</emphasis>
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(Beddard)
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</taxonomicName>
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||
.
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<treatmentCitation id="551510F4FF824977ECA6711DFD46FDB9" author="Stephenson" box="[491,716,541,563]" page="233" pageId="4" pageNumber="67" year="1932">
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||
<bibRefCitation id="B0254B14FF824977ECA6711DFD16FDB9" author="Stephenson" box="[491,668,541,563]" pageId="4" pageNumber="67" refString="Stephenson, J. (1932) A re-examination of certain Oligochaeta from the Malay Peninsula (Skeat Expedition, 1899 - 1900). The Annuals and Magazine of Natural History, 9, 201 - 240. http: // dx. doi. org / 10.1080 / 00222933208673494" type="journal article" year="1932">Stephenson 1932</bibRefCitation>
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||
: 233
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||
</treatmentCitation>
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||
–236,
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||
<figureCitation id="4C8F2A60FF824977EE46711DFCDCFDB9" box="[779,854,541,563]" captionStart="FIGURE 10" captionStartId="16.[151,250,1559,1581]" captionTargetBox="[157,1430,453,1523]" captionTargetId="figure@16.[150,1437,447,1535]" captionTargetPageId="16" captionText="FIGURE 10. External and internal morphology of Metaphire saxicalcis sp. n., holotype (CUMZ 3392). A. External ventral view. B. Internal dorsal view. C. Spermatheca; arrowhead: location of spermathecal pore." httpUri="https://zenodo.org/record/259131/files/figure.png" pageId="4" pageNumber="67">Fig. 10</figureCitation>
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.
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<taxonomicName id="13B44D66FF824977EDDA713CFE68FDD8" authority="Beddard" authorityName="Beddard" box="[151,482,572,594]" class="Clitellata" family="Megascolecidae" genus="Metaphire" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="4" pageNumber="67" phylum="Annelida" rank="species" species="pulauensis">
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<emphasis id="E6C0EAF7FF824977EDDA713CFEF8FDD8" box="[151,370,572,594]" italics="true" pageId="4" pageNumber="67">Metaphire pulauensis</emphasis>
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(Beddard)
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</taxonomicName>
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.
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<treatmentCitation id="551510F4FF824977ECA6713CFD79FDD8" author="Sims" box="[491,755,572,594]" page="239" pageId="4" pageNumber="67" year="1972">
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||
<bibRefCitation id="B0254B14FF824977ECA6713CFD4AFDD8" author="Sims" box="[491,704,572,594]" pageId="4" pageNumber="67" refString="Sims, R. W. & Easton, E. G. (1972) A numerical revision of the earthworm genus Pheretima (Megascolecidae: Oligochaeta) with the recognition of new genera and an appendix on the earthworms collected by the Royal Society North Borneo Expedition. Biological Journal of the Linnean Society, 4, 169 - 268. http: // dx. doi. org / 10.1111 / j. 1095 - 8312.1972. tb 00694. x" type="journal article" year="1972">Sims & Easton 1972</bibRefCitation>
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||
: 239
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</treatmentCitation>
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||
.
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||
</treatmentCitationGroup>
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||
</paragraph>
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||
</subSubSection>
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||
<caption id="80CB666DFF824977EDDA75E6FB62F891" httpUri="https://zenodo.org/record/259124/files/figure.png" pageId="4" pageNumber="67" targetBox="[155,1427,654,1744]" targetPageId="4">
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<paragraph id="D40B36E5FF824977EDDA75E6FB62F891" blockId="4.[151,1437,1766,1819]" pageId="4" pageNumber="67">
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||
<emphasis id="E6C0EAF7FF824977EDDA75E6FE9EF976" bold="true" box="[151,276,1766,1788]" pageId="4" pageNumber="67">FIGURE 3.</emphasis>
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External and internal morphology of
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<taxonomicName id="13B44D66FF824977EFD675E6FBA2F976" authority="Beddard, 1900" authorityName="Beddard" authorityYear="1900" box="[667,1064,1766,1788]" class="Clitellata" family="Megascolecidae" genus="Metaphire" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="4" pageNumber="67" phylum="Annelida" rank="species" species="pulauensis">
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||
<emphasis id="E6C0EAF7FF824977EFD675E6FCFDF976" box="[667,887,1766,1788]" italics="true" pageId="4" pageNumber="67">Metaphire pulauensis</emphasis>
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||
(Beddard, 1900)
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||
</taxonomicName>
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||
, lectotype (NHMUK 1924.3.1.214).
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||
<emphasis id="E6C0EAF7FF824977EDDA7405FF25F890" bold="true" box="[151,175,1797,1818]" pageId="4" pageNumber="67">A.</emphasis>
|
||
External ventral view.
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||
<emphasis id="E6C0EAF7FF824977ECD67406FE38F891" bold="true" box="[411,434,1798,1819]" pageId="4" pageNumber="67">B.</emphasis>
|
||
Internal dorsal view.
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||
<emphasis id="E6C0EAF7FF824977EFC07405FD2CF891" bold="true" box="[653,678,1797,1819]" pageId="4" pageNumber="67">C.</emphasis>
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||
Spermatheca; arrowhead: location of spermathecal pore.
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||
</paragraph>
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||
</caption>
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||
<subSubSection id="9CAE656EFF824976EDDA7442FC44FF5F" lastPageId="5" lastPageNumber="68" pageId="4" pageNumber="67" type="materials_examined">
|
||
<paragraph id="D40B36E5FF824976EDDA7442FC44FF5F" blockId="4.[151,1436,1858,2027]" lastBlockId="5.[151,1437,151,1005]" lastPageId="5" lastPageNumber="68" pageId="4" pageNumber="67">
|
||
<emphasis id="E6C0EAF7FF824977EDDA7442FE0CF8D1" bold="true" box="[151,390,1858,1883]" pageId="4" pageNumber="67">Material examined.</emphasis>
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||
The specimen that contained all the important morphological features and matched the original description is designated herein as the
|
||
<typeStatus id="0B0F8847FF824977EF967466FCCCF8F5" box="[731,838,1894,1919]" pageId="4" pageNumber="67" type="lectotype">lectotype</typeStatus>
|
||
:
|
||
<collectionCode id="B2A5AE20FF824977EE157468FC48F8F5" LSID="urn:lsid:biocol.org:col:34665" box="[856,962,1896,1919]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="4" pageNumber="67">NHMUK</collectionCode>
|
||
1924.3.1.214.
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||
<typeStatus id="0B0F8847FF824977E9357466FAAAF8F5" box="[1144,1312,1894,1919]" pageId="4" pageNumber="67" type="paralectotype">Paralectotypes</typeStatus>
|
||
:
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||
<collectionCode id="B2A5AE20FF824977E87F7468FA16F8F5" LSID="urn:lsid:biocol.org:col:34665" box="[1330,1436,1896,1919]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="4" pageNumber="67">NHMUK</collectionCode>
|
||
1924.3.1.215,
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||
<specimenCount id="C2B2FD6CFF824977EC75748BFE03F829" box="[312,393,1930,1955]" pageId="4" pageNumber="67" type="adult">1 adult</specimenCount>
|
||
.
|
||
<collectionCode id="B2A5AE20FF824977ECD9748BFE74F828" LSID="urn:lsid:biocol.org:col:34665" box="[404,510,1931,1954]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="4" pageNumber="67">NHMUK</collectionCode>
|
||
1904.10.5.1002,
|
||
<specimenCount id="C2B2FD6CFF824977EF8F748BFC99F829" box="[706,787,1930,1955]" pageId="4" pageNumber="67" type="adult">1 adult</specimenCount>
|
||
; same locality as
|
||
<typeStatus id="0B0F8847FF824977EE94748AFBCEF829" box="[985,1092,1930,1955]" pageId="4" pageNumber="67" type="lectotype">lectotype</typeStatus>
|
||
. The exact location of the
|
||
<typeStatus id="0B0F8847FF824977E821748BFA16F829" box="[1388,1436,1931,1955]" pageId="4" pageNumber="67">type</typeStatus>
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||
locality is unknown. The locality of all
|
||
<taxonomicName id="13B44D66FF824977EF3D74B0FC9BF84D" box="[624,785,1967,1991]" class="Clitellata" family="Megascolecidae" genus="Metaphire" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="4" pageNumber="67" phylum="Annelida" rank="species" species="pulauensis">
|
||
<emphasis id="E6C0EAF7FF824977EF3D74B0FC9BF84D" box="[624,785,1967,1991]" italics="true" pageId="4" pageNumber="67">M. pulauensis</emphasis>
|
||
</taxonomicName>
|
||
specimens in the Natural History Museum, London, is labelled as '
|
||
<collectingCountry id="ACA37675FF824977EC5574D2FED0F861" box="[280,346,2002,2027]" name="Iceland" pageId="4" pageNumber="67">Pulau</collectingCountry>
|
||
Bidan, Kedah, Malay Peninsula' while
|
||
<bibRefCitation id="B0254B14FF824977EE5474D2FB89F861" author="Beddard" box="[793,1027,2002,2027]" pageId="4" pageNumber="67" refString="Beddard, F. E. (1900) On the earthworms collected during the " Skeat Expedition " to the Malay Peninsula, 1899 - 1900. Proceedings of the Zoological Society of London, 1900, 891 - 911." type="proceedings paper" year="1900">Beddard (1900: 904)</bibRefCitation>
|
||
gives the locality as '
|
||
<collectingCountry id="ACA37675FF824977E9B974D2FAB3F861" box="[1268,1337,2002,2027]" name="Iceland" pageId="4" pageNumber="67">Pulau</collectingCountry>
|
||
, Bidang, Kelah, and Aring'.
|
||
<collectingCountry id="ACA37675FF834976EC3B7397FE3CFF3A" box="[374,438,151,176]" name="Iceland" pageId="5" pageNumber="68">Pulau</collectingCountry>
|
||
means "island" in Malay. There is a small island named "Bidan" north of the Pinang Island in Kedah State, northern
|
||
<collectingCountry id="ACA37675FF834976ECB773BCFDEEFF5F" box="[506,612,188,213]" name="Malaysia" pageId="5" pageNumber="68">Malaysia</collectingCountry>
|
||
, which may be the
|
||
<typeStatus id="0B0F8847FF834976EE7173BDFCE6FF5F" box="[828,876,189,213]" pageId="5" pageNumber="68">type</typeStatus>
|
||
locality.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="9CAE656EFF834976ED8A73DFFA17FE02" pageId="5" pageNumber="68" type="diagnosis">
|
||
<paragraph id="D40B36E5FF834976ED8A73DFFA17FE02" blockId="5.[151,1437,151,1005]" pageId="5" pageNumber="68">
|
||
<emphasis id="E6C0EAF7FF834976ED8A73DFFECBFF72" bold="true" box="[199,321,223,248]" pageId="5" pageNumber="68">Diagnosis.</emphasis>
|
||
Medium-sized, length
|
||
<quantity id="134C9B00FF834976EF0973DFFD53FF7D" box="[580,729,223,248]" metricMagnitude="-1" metricUnit="m" metricValue="1.54" metricValueMax="1.65" metricValueMin="1.43" pageId="5" pageNumber="68" unit="mm" value="154.0" valueMax="165.0" valueMin="143.0">143–165 mm</quantity>
|
||
with 104–110 segments. First dorsal pore in 12/13. Male pores on XVIII, secondary openings puckered radially around margins. Genital papillae aggregated together to form slightly transversely oval patches, mid-ventral, pre-setal of VII, VIII, IX, XVII and XVIII. Spermathecal pores paired at 5/6–8/9, spermathecal ampulla elongated, sac-shaped, diverticulum twisted into short loops. No nephridia on the spermathecal ducts. Holandric, intestinal caeca simple. Prostate glands large, genital marking glands sessile.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="9CAE656EFF834976ED8A7294FC00FC67" pageId="5" pageNumber="68" type="discussion">
|
||
<paragraph id="D40B36E5FF834976ED8A7294FD19FC9F" blockId="5.[151,1437,151,1005]" pageId="5" pageNumber="68">
|
||
<emphasis id="E6C0EAF7FF834976ED8A7294FEB1FE27" bold="true" box="[199,315,404,429]" pageId="5" pageNumber="68">Remarks.</emphasis>
|
||
The original description (
|
||
<bibRefCitation id="B0254B14FF834976EF2A7294FC80FE26" author="Beddard" box="[615,778,404,429]" pageId="5" pageNumber="68" refString="Beddard, F. E. (1900) On the earthworms collected during the " Skeat Expedition " to the Malay Peninsula, 1899 - 1900. Proceedings of the Zoological Society of London, 1900, 891 - 911." type="proceedings paper" year="1900">Beddard 1900</bibRefCitation>
|
||
) was based on an unspecified number of individuals that had been collected by Mr. R. Evans of Oxford, during the Skeat Expedition to the Malay Peninsula in
|
||
<date id="A00A1025FF834976E85B72B8FA12FE5A" box="[1302,1432,440,464]" pageId="5" pageNumber="68" value="1899" valueMax="1900">1899–1900</date>
|
||
;
|
||
<typeStatus id="0B0F8847FF834976EDDA72DDFF58FE7F" box="[151,210,477,501]" pageId="5" pageNumber="68">types</typeStatus>
|
||
were not designated in the original publication. Stephenson, reinvestigating the original material, states the following: 'Two specimens, both already dissected, in a jar labelled "
|
||
<typeStatus id="0B0F8847FF834976EEF07100FB82FD92" box="[957,1032,512,536]" pageId="5" pageNumber="68">Types</typeStatus>
|
||
";
|
||
<collectingCountry id="ACA37675FF834976E96C72FFFBEBFD92" box="[1057,1121,511,536]" name="Iceland" pageId="5" pageNumber="68">Pulau</collectingCountry>
|
||
Bidan, Kedah State; Skeat Expedition;
|
||
<date id="A00A1025FF834976EC687125FE5AFDB6" box="[293,464,548,572]" pageId="5" pageNumber="68" value="1899-11-23">Nov. 23, 1899</date>
|
||
. A third specimen in another jar, provenance given as Malay Peninsula … I have examined the two specimens in the first jar, both externally and internally, and the external characters of the third specimen also. The basis of the description is the larger of the two "
|
||
<typeStatus id="0B0F8847FF834976EEDA716CFC46FD0E" box="[919,972,620,644]" pageId="5" pageNumber="68">type</typeStatus>
|
||
" specimens' (
|
||
<bibRefCitation id="B0254B14FF834976E92B716CFAECFD0E" author="Stephenson" box="[1126,1382,620,645]" pageId="5" pageNumber="68" refString="Stephenson, J. (1932) A re-examination of certain Oligochaeta from the Malay Peninsula (Skeat Expedition, 1899 - 1900). The Annuals and Magazine of Natural History, 9, 201 - 240. http: // dx. doi. org / 10.1080 / 00222933208673494" type="journal article" year="1932">Stephenson 1932: 233</bibRefCitation>
|
||
). No
|
||
<typeStatus id="0B0F8847FF834976EDDA718FFF74FD22" box="[151,254,655,680]" pageId="5" pageNumber="68" type="lectotype">lectotype</typeStatus>
|
||
was designated by Stephenson. We consider all three specimens to belong to the
|
||
<typeStatus id="0B0F8847FF834976E9D07190FB7EFD22" box="[1181,1268,656,680]" pageId="5" pageNumber="68" type="syntype">syntype</typeStatus>
|
||
series, and we designate the specimen that contains the important morphological features and matches the original description best as the
|
||
<typeStatus id="0B0F8847FF834976EC5471D7FEF5FD7A" box="[281,383,727,752]" pageId="5" pageNumber="68" type="lectotype">lectotype</typeStatus>
|
||
(
|
||
<figureCitation id="4C8F2A60FF834976ECDC71D7FE5CFD7A" box="[401,470,727,752]" captionStart="FIGURE 3" captionStartId="4.[151,250,1766,1788]" captionTargetBox="[155,1427,654,1744]" captionTargetId="figure@4.[151,1435,646,1745]" captionTargetPageId="4" captionText="FIGURE 3. External and internal morphology of Metaphire pulauensis (Beddard, 1900), lectotype (NHMUK 1924.3.1.214). A. External ventral view. B. Internal dorsal view. C. Spermatheca; arrowhead: location of spermathecal pore." httpUri="https://zenodo.org/record/259124/files/figure.png" pageId="5" pageNumber="68">Fig. 3</figureCitation>
|
||
), and the other two as
|
||
<typeStatus id="0B0F8847FF834976EFAC71D7FC0CFD7A" box="[737,902,727,752]" pageId="5" pageNumber="68" type="paralectotype">paralectotypes</typeStatus>
|
||
. Both original description (
|
||
<bibRefCitation id="B0254B14FF834976E98D71D7FAE9FD7A" author="Beddard" box="[1216,1379,727,752]" pageId="5" pageNumber="68" refString="Beddard, F. E. (1900) On the earthworms collected during the " Skeat Expedition " to the Malay Peninsula, 1899 - 1900. Proceedings of the Zoological Society of London, 1900, 891 - 911." type="proceedings paper" year="1900">Beddard 1900</bibRefCitation>
|
||
) and redescription (
|
||
<bibRefCitation id="B0254B14FF834976EC7A71FCFE70FC9E" author="Stephenson" box="[311,506,764,789]" pageId="5" pageNumber="68" refString="Stephenson, J. (1932) A re-examination of certain Oligochaeta from the Malay Peninsula (Skeat Expedition, 1899 - 1900). The Annuals and Magazine of Natural History, 9, 201 - 240. http: // dx. doi. org / 10.1080 / 00222933208673494" type="journal article" year="1932">Stephenson 1932</bibRefCitation>
|
||
) are detailed.
|
||
</paragraph>
|
||
<paragraph id="D40B36E5FF834976ED8A701FFAA0FC2E" blockId="5.[151,1437,151,1005]" pageId="5" pageNumber="68">
|
||
The
|
||
<typeStatus id="0B0F8847FF834976EDB77020FEA0FCB2" box="[250,298,800,824]" pageId="5" pageNumber="68">type</typeStatus>
|
||
locality of
|
||
<taxonomicName id="13B44D66FF834976ECE17021FDC0FCB2" box="[428,586,800,824]" class="Clitellata" family="Megascolecidae" genus="Metaphire" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="5" pageNumber="68" phylum="Annelida" rank="species" species="pulauensis">
|
||
<emphasis id="E6C0EAF7FF834976ECE17021FDC0FCB2" box="[428,586,800,824]" italics="true" pageId="5" pageNumber="68">M. pulauensis</emphasis>
|
||
</taxonomicName>
|
||
in northern
|
||
<collectingCountry id="ACA37675FF834976EF98701FFCB6FCB2" box="[725,828,799,824]" name="Malaysia" pageId="5" pageNumber="68">Malaysia</collectingCountry>
|
||
is quite close to the
|
||
<typeStatus id="0B0F8847FF834976E96B7020FBDCFCB2" box="[1062,1110,800,824]" pageId="5" pageNumber="68">type</typeStatus>
|
||
locality of
|
||
<taxonomicName id="13B44D66FF834976E9947021FA11FCB2" box="[1241,1435,800,824]" class="Clitellata" family="Megascolecidae" genus="Metaphire" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="5" pageNumber="68" phylum="Annelida" rank="species" species="songkhlaensis" status="sp. nov.">
|
||
<emphasis id="E6C0EAF7FF834976E9947021FA11FCB2" box="[1241,1435,800,824]" italics="true" pageId="5" pageNumber="68">M. songkhlaensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="FDF3578CFF834976EDDA7044FF5CFCD7" box="[151,214,836,861]" pageId="5" pageNumber="68" rank="species">
|
||
<emphasis id="E6C0EAF7FF834976EDDA7044FF5CFCD7" bold="true" box="[151,214,836,861]" pageId="5" pageNumber="68">sp. n.</emphasis>
|
||
</taxonomicNameLabel>
|
||
in southern
|
||
<collectingCountry id="ACA37675FF834976EC137044FE4FFCD7" box="[350,453,836,861]" name="Thailand" pageId="5" pageNumber="68">Thailand</collectingCountry>
|
||
. However,
|
||
<taxonomicName id="13B44D66FF834976EF0F7045FD54FCD6" box="[578,734,836,860]" class="Clitellata" family="Megascolecidae" genus="Metaphire" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="5" pageNumber="68" phylum="Annelida" rank="species" species="pulauensis">
|
||
<emphasis id="E6C0EAF7FF834976EF0F7045FD54FCD6" box="[578,734,836,860]" italics="true" pageId="5" pageNumber="68">M. pulauensis</emphasis>
|
||
</taxonomicName>
|
||
can be easily distinguished from
|
||
<taxonomicName id="13B44D66FF834976E91E7045FA9EFCD6" box="[1107,1300,836,860]" class="Clitellata" family="Megascolecidae" genus="Metaphire" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="5" pageNumber="68" phylum="Annelida" rank="species" species="songkhlaensis">
|
||
<emphasis id="E6C0EAF7FF834976E91E7045FA9EFCD6" box="[1107,1300,836,860]" italics="true" pageId="5" pageNumber="68">M. songkhlaensis</emphasis>
|
||
</taxonomicName>
|
||
by the male field puckered radially around the margins and genital markings on segments VII, VIII, IX, XVII and XVIII, while
|
||
<taxonomicName id="13B44D66FF834976EDDA708DFED3FC2E" box="[151,345,908,932]" class="Clitellata" family="Megascolecidae" genus="Metaphire" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="5" pageNumber="68" phylum="Annelida" rank="species" species="songkhlaensis" status="sp. nov.">
|
||
<emphasis id="E6C0EAF7FF834976EDDA708DFED3FC2E" box="[151,345,908,932]" italics="true" pageId="5" pageNumber="68">M. songkhlaensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="FDF3578CFF834976EC2D708CFE15FC2F" box="[352,415,908,933]" pageId="5" pageNumber="68" rank="species">
|
||
<emphasis id="E6C0EAF7FF834976EC2D708CFE15FC2F" bold="true" box="[352,415,908,933]" pageId="5" pageNumber="68">sp. n.</emphasis>
|
||
</taxonomicNameLabel>
|
||
has prominent male field and no genital markings on segments XVII and XVIII.
|
||
</paragraph>
|
||
<paragraph id="D40B36E5FF834976ED8A70AFFC00FC67" blockId="5.[151,1437,151,1005]" pageId="5" pageNumber="68">
|
||
This species is apparently known only from the
|
||
<typeStatus id="0B0F8847FF834976EFBA70B0FCADFC42" box="[759,807,944,968]" pageId="5" pageNumber="68">type</typeStatus>
|
||
locality, but the mentioning of "Aring" in the original description may indicate that the species was also found elsewhere.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |