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<document id="9B8755FB6B52D9FF4093BD21BFEFA278" ID-CLB-Dataset="32109" ID-DOI="http://dx.doi.org/10.3897/zookeys.697.14060" ID-GBIF-Dataset="586b8dd8-47f6-455e-98e5-c4f1ea6a0ea8" ID-PMC="PMC5673843" ID-Pensoft-Pub="1313-2970-697-21" ID-PubMed="29134014" ID-ZooBank="8F43E4946D4B40EF913944D53822BCE0" ModsDocAuthor="" ModsDocDate="2017" ModsDocID="1313-2970-697-21" ModsDocOrigin="ZooKeys 697" ModsDocTitle="On the identity of Leptoxistaeniata – a misapplied name for the threatened Painted Rocksnail (Cerithioidea, Pleuroceridae)" checkinTime="1505404177079" checkinUser="pensoft" docAuthor="Whelan, Nathan V., Johnson, Paul D., Garner, Jeffrey T. &amp; Strong, Ellen E." docDate="2017" docId="9687A6DA90CD4A52CFD8D8981D6C9678" docLanguage="en" docName="ZooKeys 697: 21-36" docOrigin="ZooKeys 697" docSource="http://dx.doi.org/10.3897/zookeys.697.14060" docTitle="Leptoxis coosaensis Lea 1861" docType="treatment" docVersion="6" lastPageNumber="27" masterDocId="6D3BFFB1B906FFB0572FC02BFFF1FF9E" masterDocTitle="On the identity of Leptoxistaeniata - a misapplied name for the threatened Painted Rocksnail (Cerithioidea, Pleuroceridae)" masterLastPageNumber="36" masterPageNumber="21" pageNumber="24" updateTime="1701377205082" updateUser="ExternalLinkService">
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<mods:title id="3A0BE3CA46AFE54ABB102879EB0B2204">On the identity of Leptoxistaeniata - a misapplied name for the threatened Painted Rocksnail (Cerithioidea, Pleuroceridae)</mods:title>
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<mods:namePart id="ADBD97DA7032B57FECDBC70C13F247AD">Whelan, Nathan V.</mods:namePart>
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<mods:namePart id="74E928F33389084B614AE2619476CDE2">Garner, Jeffrey T.</mods:namePart>
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<mods:namePart id="FFDFA5546ABA76A7FB2B5A1BE64856AC">Strong, Ellen E.</mods:namePart>
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<treatment id="9687A6DA90CD4A52CFD8D8981D6C9678" ID-GBIF-Taxon="134089512" LSID="urn:lsid:plazi:treatment:9687A6DA90CD4A52CFD8D8981D6C9678" httpUri="http://treatment.plazi.org/id/9687A6DA90CD4A52CFD8D8981D6C9678" lastPageId="6" lastPageNumber="27" pageId="3" pageNumber="24">
<subSubSection id="8BCA19152745E4F3F6584DEC643AFC07" pageId="3" pageNumber="24" type="nomenclature">
<paragraph id="B5043F177EAFFEEF815D6B1529F56C7F" pageId="3" pageNumber="24">
<taxonomicName id="C23DBFCAB8732C659407A66BC864556B" ID-CoL="3TZB3" authority="Lea, 1861" authorityName="Lea" authorityYear="1861" class="Gastropoda" family="Pleuroceridae" genus="Leptoxis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Leptoxis coosaensis" order="Neotaenioglossa" pageId="3" pageNumber="24" phylum="Mollusca" rank="species" species="coosaensis">Leptoxis coosaensis (Lea, 1861)</taxonomicName>
</paragraph>
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<paragraph id="1AE37FEB9174EA9CC07BEB3515AED846" pageId="3" pageNumber="24">
<taxonomicName id="25D49733C329BF4BA1512D4BBAD29677" class="Gastropoda" family="Pleuroceridae" genus="Anculosa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anculosa coosaensis" order="Neotaenioglossa" pageId="3" pageNumber="24" phylum="Mollusca" rank="species" species="coosaensis">Anculosa coosaensis</taxonomicName>
Lea, 1861: 54; 1863a: 257-258, pl. 35, fig. 65; 1863b: 79-80, pl. 35, fig. 65. Lectotype USNM 121295 (Graf, 2001); paralectotypes USNM 1456804 (1 spm), USNM 121294 (4 spms) and USNM 121296 (1 spm) leg. Showalter. &quot;Coosa River, Alabama.&quot;
</paragraph>
<paragraph id="AF1CD2F2B65A228EC2807F312A10A271" pageId="3" pageNumber="24">
<taxonomicName id="5860917E889BF666DB3DDDE57BEEE488" class="Gastropoda" family="Pleuroceridae" genus="Anculosa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anculosa aldrichi" order="Neotaenioglossa" pageId="3" pageNumber="24" phylum="Mollusca" rank="species" species="aldrichi">Anculosa aldrichi</taxonomicName>
Goodrich, 1922: 31, pl. 1, figs. 1, 2. Holotype UMMZ 10139, by original designation; paratype, MCZ 169987 (1 spm). &quot;Coosa River, near mouth of Yellowleaf [sic, Yellow Leaf] Creek, Chilton County, Alabama.&quot;
</paragraph>
<paragraph id="D9CC2A7351E55BDF334AA72E6F842240" lastPageId="4" lastPageNumber="25" pageId="3" pageNumber="24">
<taxonomicName id="CF5AA2084E981CC2BD684D0B83466058" class="Gastropoda" family="Pleuroceridae" genus="Anculosa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anculosa brevispira" order="Neotaenioglossa" pageId="3" pageNumber="24" phylum="Mollusca" rank="species" species="brevispira">Anculosa brevispira</taxonomicName>
Goodrich, 1922: 35, pl. 1, fig. 6. Holotype UMMZ 10144, by original designation; paratype MCZ 169991 (1 spm). &quot;Fort William Shoals,
<pageBreakToken id="31B71F6D05F5539C6F4BEF1643CC1031" pageId="4" pageNumber="25" start="start">Coosa</pageBreakToken>
River, Talladega County, Alabama&quot;; possible paratypes UF 18202 (7 spms). &quot;Coosa River&quot;.
</paragraph>
<paragraph id="781988C0DC0A7FF9665959E4B548B264" pageId="4" pageNumber="25">
<taxonomicName id="FABCEF6487A23470FEE16ECAB1FFDB94" class="Gastropoda" family="Pleuroceridae" genus="Anculosa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anculosa choccoloccoensis" order="Neotaenioglossa" pageId="4" pageNumber="25" phylum="Mollusca" rank="species" species="choccoloccoensis">Anculosa choccoloccoensis</taxonomicName>
Goodrich, 1922: 34, pl. 1, fig. 7. Holotype UMMZ 10145, by original designation; paratype MCZ 169989 (1 spm). &quot;Choccolocco Creek at Jackson Shoals, Talladega County, Alabama.&quot;
</paragraph>
<paragraph id="88DA1424BD27693A188918733F3CC187" pageId="4" pageNumber="25">
<taxonomicName id="3BE7B9B517B7B8263339FBB5D91BC469" class="Gastropoda" family="Pleuroceridae" genus="Anculosa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anculosa taeniata" order="Neotaenioglossa" pageId="4" pageNumber="25" phylum="Mollusca" rank="subspecies" species="taeniata" subspecies="lucida">Anculosa taeniata lucida</taxonomicName>
Goodrich, 1944: 42. Type material not located, potentially lost. &quot;Coosa [River] tributaries.&quot;
</paragraph>
<paragraph id="B49E47A2C6FEEED7F0551A2E1DA69975" pageId="4" pageNumber="25">
Other references:
<taxonomicName id="E45CFFDA78139CEB872FFAC8CAD4E22A" class="Gastropoda" family="Pleuroceridae" genus="Anculosa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anculosa coosaensis" order="Neotaenioglossa" pageId="4" pageNumber="25" phylum="Mollusca" rank="species" species="coosaensis">Anculosa coosaensis</taxonomicName>
-
<bibRefCitation id="62BA4011EFC7AF059459E601BE345268" author="Lea, I" journalOrPublisher="Journal of the Academy of Natural Sciences of Philadelphia" pageId="12" pageNumber="33" pagination="217 - 356" title="New Melanida of the United States." volume="5" year="1863">Lea 1863</bibRefCitation>
: 257-258, pl. 35, fig. 65;
<bibRefCitation id="A3F35160C3B5BBBF53DD40977A7400C1" author="Goodrich, C" journalOrPublisher="University of Michigan Museum of Zoology Miscellaneous Publication No" pageId="11" pageNumber="32" pagination="1 - 63" title="The Anculosae of the Alabama River drainage." volume="7" year="1922">Goodrich 1922</bibRefCitation>
: 28, pl. 1, figs. 13, 14.
</paragraph>
<paragraph id="4B1ED901F1A9D3225AC244D7CAF72DD5" pageId="4" pageNumber="25">
<taxonomicName id="789EA94C9D8A8250EBBDB2AE9B4950FC" class="Gastropoda" family="Pleuroceridae" genus="Leptoxis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Leptoxis taeniata" order="Neotaenioglossa" pageId="4" pageNumber="25" phylum="Mollusca" rank="species" species="taeniata">Leptoxis taeniata</taxonomicName>
-Haldeman 1848: 3, pl. 3 figs. 71-72;
<bibRefCitation id="44478ABF10B5E1957758BF3819C61007" author="Burch, JB" journalOrPublisher="Walkerana" pageId="11" pageNumber="32" pagination="1 - 215" title="North American freshwater snails, species list, ranges, and illustrations." url="http://molluskconservation.org/PUBLICATIONS/WALKERANA/Vol1/walkerana%20vol1%20no3%2081-216.PDF" volume="1" year="1980">Burch and Tottenham 1980</bibRefCitation>
: 156, figs. 484-486;
<bibRefCitation id="7EEEEEB9C2626646874EAC2B633BAF5B" author="Burch, JB" journalOrPublisher="Walkerana" pageId="11" pageNumber="32" pagination="217 - 365" title="North American freshwater snails, identification keys, generic synonymy, supplemental notes, glossary, references, index." url="http://molluskconservation.org/PUBLICATIONS/WALKERANA/Vol1/walkerana%20vol1%20no4%20217-366.PDF" volume="1" year="1982">Burch 1982</bibRefCitation>
: 43, figs. 484-486;
<bibRefCitation id="D264BCCBA60324A166B38F06681C656C" author="Lydeard, C" journalOrPublisher="Molecular Phylogenetics and Evolution" pageId="12" pageNumber="33" pagination="117 - 128" title="A molecular phylogeny of Mobile River drainage basin pleurocerid snails (Caenogastropoda: Cerithioidea)." url="https://doi.org/10.1006/mpev.1996.0377" volume="7" year="1997">Lydeard et al. 1997</bibRefCitation>
: 117-128;
<bibRefCitation id="F9E6F94AD2A4041E3E0BA1BDDF25E6B6" author="Lydeard, C" journalOrPublisher="Malacologia" pageId="12" pageNumber="33" pagination="183 - 193" title="Phylogenetic utility of the 5 ' - half of mitochondrial 16 S rDNA gene sequences for inferring relationships of Elimia (Cerithioidea: Pleuroceridae)." volume="39" year="1998">Lydeard et al. 1998</bibRefCitation>
: 183-193;
<bibRefCitation id="BAECDC156661A92B672D7F004508E823" author="Dillon, RT" journalOrPublisher="Malacologia" pageId="11" pageNumber="32" pagination="111 - 119" title="Divergence among Mobile Basin populations of the pleurocerid snail genus, Leptoxis, estimated by allozyme electrophoresis." volume="39" year="1998">Dillon and Lydeard 1998</bibRefCitation>
: 113-121, fig. 2;
<bibRefCitation id="46F62EC31853DAE5706CDF1692CBEB83" author="Holznagel, WE" journalOrPublisher="Journal of Molluscan Studies" pageId="12" pageNumber="33" pagination="233 - 257" title="A molecular phylogeny of North American Pleuroceridae (Gastropoda: Cerithioidea) based on mitochondrial 16 S rDNA sequences." url="https://doi.org/10.1093/mollus/66.2.233" volume="66" year="2000">Holznagel and Lydeard 2000</bibRefCitation>
: 233-257;
<bibRefCitation id="F845DA5A6FB00C9B731D70AF99DA644B" author="Lee, T" journalOrPublisher="Journal of Molluscan Studies" pageId="12" pageNumber="33" pagination="314 - 317" title="Crossing the continental divide: the Columbia drainage species Jugahemphili (Henderson, 1935) is a cryptic member of the eastern North American genus Elimia (Cerithioidea: Pleuroceridae)." url="https://doi.org/10.1093/mollus/eyl005" volume="72" year="2006">Lee et al. 2006</bibRefCitation>
: 314-317;
<bibRefCitation id="EA788D893100840B5E84D6C3B6B0CDA0" author="Strong, EE" journalOrPublisher="Zoologica Scripta" pageId="12" pageNumber="33" pagination="483 - 502" title="Morphological and molecular analysis of ' Melaniajacqueti' Dautzenberg and Fischer, 1906: from anonymous orphan to critical basal offshoot of the Semisulcospiridae (Gastropoda: Cerithioidea)." url="https://doi.org/10.1111/j.1463-6409.2008.00385.x" volume="38" year="2009">
Strong and
<normalizedToken id="7F85C44C29C397ACC01840177F0AAD22" originalValue="Köhler">Koehler</normalizedToken>
2009
</bibRefCitation>
: 483-502;
<bibRefCitation id="3009A463D281D5691826146FCB2C8A99" author="Tolley-Jordan, LR" journalOrPublisher="Aquatic Conservation: Marine and Freshwater Ecosystems" pageId="12" pageNumber="33" pagination="235 - 249" title="Effects of land-use change on a diverse pleurocerid snail assemblage." url="https://doi.org/10.1002/aqc.2474" volume="25" year="2015">Tolley-Jordan et al. 2015</bibRefCitation>
: 235-249;
<bibRefCitation id="A5B2BAA408ABAE30927DEFDBB184A9CF" author="Whelan, NV" journalOrPublisher="Journal of Molluscan Studies" pageId="12" pageNumber="33" pagination="85 - 95" title="Life-history traits and shell morphology in the genus Leptoxis Rafinesque, 1819 (Gastropoda: Cerithioidea: Pleuroceridae)." url="https://doi.org/10.1093/mollus/eyu058" volume="81" year="2015">Whelan et al. 2015</bibRefCitation>
: 85-95, fig. 4;
<bibRefCitation id="A4B29175BBD627A93B15152098E1E52E" author="Whelan, NV" journalOrPublisher="American Malacological Bulletin" pageId="12" pageNumber="33" pagination="221 - 226" title="Radular morphology of extinct pleurocerids (Gastropoda: Cerithioidea: Pleuroceridae)." url="https://doi.org/10.4003/006.033.0202" volume="33" year="2016">Whelan 2016</bibRefCitation>
: 221-226. [Not
<taxonomicName id="14F831B6F8091A4161020231FC9E7706" family="Pleuroceridae" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="L. taeniata" order="Neotaenioglossa" pageId="4" pageNumber="25" rank="species" species="taeniata">L. taeniata</taxonomicName>
of Conrad]
</paragraph>
<paragraph id="D7A6C9FA1DDF9F54880A8A82B08292B4" pageId="4" pageNumber="25">
<taxonomicName id="0BFAE3E3D4BB1C9F860F15B68075CA4E" class="Gastropoda" family="Pleuroceridae" genus="Anculosa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anculosa taeniata" order="Neotaenioglossa" pageId="4" pageNumber="25" phylum="Mollusca" rank="species" species="taeniata">Anculosa taeniata</taxonomicName>
-
<bibRefCitation id="0B1639E82091C14F8A6E356CD8F6006D" author="Tryon, GW" journalOrPublisher="Smithsonian Institution, Washington" pageId="12" pageNumber="33" title="Land and fresh-water shells of North America part IV. Streptomatidae (American melanians)." year="1873">Tryon 1873</bibRefCitation>
: 408-409, figs. 813-815. [Not
<taxonomicName id="4F1D41D4FF7F5FBB3A47CCFDC4AEB45F" family="Pleuroceridae" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="L. taeniata" order="Neotaenioglossa" pageId="4" pageNumber="25" rank="species" species="taeniata">L. taeniata</taxonomicName>
of Conrad]
</paragraph>
<paragraph id="5734C3A66E5301B545F4B721A799B51A" pageId="4" pageNumber="25">
<taxonomicName id="FF4AF741F46D49251298A3ED7EFD41DF" class="Gastropoda" family="Pleuroceridae" genus="Anculotus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anculotus taeniatus" order="Neotaenioglossa" pageId="4" pageNumber="25" phylum="Mollusca" rank="species" species="taeniatus">Anculotus taeniatus</taxonomicName>
-Reeve 1860: pl. 6, fig. 50. [Not
<taxonomicName id="D6A8143D39683D493DD213CE7F3174EC" family="Pleuroceridae" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="L. taeniata" order="Neotaenioglossa" pageId="4" pageNumber="25" rank="species" species="taeniata">L. taeniata</taxonomicName>
of Conrad]
</paragraph>
</subSubSection>
<subSubSection id="1308638A57D47D18D6CEA71D89B3C51C" lastPageId="6" lastPageNumber="27" pageId="4" pageNumber="25" type="materials_examined">
<paragraph id="4AD7819BE4333BB6610118D10027CBC3" pageId="4" pageNumber="25">Other material examined.</paragraph>
<paragraph id="4A53B2B69106A776F65AE715EE2E7606" lastPageId="6" lastPageNumber="27" pageId="4" pageNumber="25">
UMMZ 10144, Coosa River at Fort Williams Shoals, Talladega County, Alabama (~
<geoCoordinate id="2B177CFF54BA52DEB0C276473065DA41" direction="north" orientation="latitude" precision="5" value="33.1477">33.1477°N</geoCoordinate>
,
<geoCoordinate id="CAA7DE8F11494C556A0B06833E47B355" direction="west" orientation="longitude" precision="5" value="-86.4831">86.4831°W</geoCoordinate>
); UMMZ 10139, Coosa River near mouth of Yellow Leaf Creek, Chilton County, Alabama (~
<geoCoordinate id="0544DE614ABF4C92059E68C06867C95A" direction="north" orientation="latitude" precision="5" value="32.9566">32.9566°N</geoCoordinate>
,
<geoCoordinate id="2085112BF0AD97B3100EDC9D79E055F1" direction="west" orientation="longitude" precision="5" value="-86.5177">86.5177°W</geoCoordinate>
); UMMZ 10145, Choccolocco Creek at Jackson Shoals, Talladega County, Alabama (~
<geoCoordinate id="C13E75F252A2248EB8BD6CDA4C68E1BC" direction="north" orientation="latitude" precision="5" value="33.545">33.5450°N</geoCoordinate>
,
<geoCoordinate id="20C099E854097A98AD6D7ECC759D6604" direction="west" orientation="longitude" precision="5" value="-86.0896">86.0896°W</geoCoordinate>
); MCZ 169987, Coosa River near mouth of Yellow Leaf Creek, Chilton County, Alabama (~
<geoCoordinate id="46F6A5C78F2B0D83332E77E623F460A0" direction="north" orientation="latitude" precision="5" value="32.9566">32.9566°N</geoCoordinate>
,
<geoCoordinate id="4A66BDB0784D0FB49DB502325C53F325" direction="west" orientation="longitude" precision="5" value="-86.5177">86.5177°W</geoCoordinate>
); MCZ 169989, Choccolocco Creek at Jackson Shoals, Talladega County, Alabama (~
<geoCoordinate id="D71D5C99D1FDF4668615B8E009486651" direction="north" orientation="latitude" precision="5" value="33.545">33.5450°N</geoCoordinate>
,
<geoCoordinate id="7D77942C037F1AA661DA45D18CE9AAC7" direction="west" orientation="longitude" precision="5" value="-86.0896">86.0896°W</geoCoordinate>
); USNM 12068, Coosa River, Alabama; USNM 321181,
<normalizedToken id="E42464E91C079DE3288149F5B4ACC3BB" originalValue="Duncan’s">Duncan's</normalizedToken>
Riffle Coosa River, Chilton County (~
<geoCoordinate id="89CE5A3C22585A045B8C3923C657B001" direction="north" orientation="latitude" precision="5" value="32.8057">32.8057°N</geoCoordinate>
,
<geoCoordinate id="660D4CB5C0066A108D9423C5BBDEAA8E" direction="west" orientation="longitude" precision="5" value="-86.445">86.4450°W</geoCoordinate>
; USNM 321862,
<normalizedToken id="D02AFF0E8358C778535B9469BBF25B8E" originalValue="Higgin’s">Higgin's</normalizedToken>
Ferry Coosa River, Chilton County, Alabama (~
<geoCoordinate id="C628950ECCD85D960085BE145AC8A297" direction="north" orientation="latitude" precision="5" value="32.8056">32.8056°N</geoCoordinate>
,
<geoCoordinate id="2078D44820E7AAA50DC7236FFDC5E39C" direction="west" orientation="longitude" precision="5" value="-86.4448">86.4448°W</geoCoordinate>
); UF 82401, Peckerwood Shoals, Talladega County, Alabama (~
<geoCoordinate id="32F740687C39785107AB6BDDAAFE2683" direction="north" orientation="latitude" precision="5" value="33.1176">33.1176°N</geoCoordinate>
,
<geoCoordinate id="96665C790F062B4047E02CAB3ACC8EE3" direction="west" orientation="longitude" precision="5" value="-86.4728">86.4728°W</geoCoordinate>
); UF 416903, Talladega Creek near Nottingham, Talladega County, Alabama (~
<geoCoordinate id="A051F412DB1B69F80548A919D813D493" direction="north" orientation="latitude" precision="5" value="33.3614">33.3614°N</geoCoordinate>
,
<geoCoordinate id="14FD43617F6C11F2923ABC008B3D327D" direction="west" orientation="longitude" precision="5" value="-86.2237">86.2237°W</geoCoordinate>
); UF 82358, Coosa River at Fort Williams Shoals, Talladega County, Alabama (~
<geoCoordinate id="6A3874B43EC0D66FE421C387C0DFF3BE" direction="north" orientation="latitude" precision="5" value="33.1477">33.1477°N</geoCoordinate>
,
<geoCoordinate id="9C06DC9252878216857F17D3FCFEB7F8" direction="west" orientation="longitude" precision="5" value="-86.4831">86.4831°W</geoCoordinate>
); UF 81660, Coosa River at Fort Williams Shoals, Talladega County, Alabama (~
<geoCoordinate id="D4409D49DAD0144D535CD5C02BF9C7FC" direction="north" orientation="latitude" precision="5" value="33.1477">33.1477°N</geoCoordinate>
,
<geoCoordinate id="66099B0CFC693362A70C069A421613AA" direction="west" orientation="longitude" precision="5" value="-86.4831">86.4831°W</geoCoordinate>
); UF 82419, Coosa River at Butting Ram Shoals, Coosa County, Alabama (~
<geoCoordinate id="DE2FE98EFE1CAC6CCD636DE11AB6B217" direction="north" orientation="latitude" precision="5" value="32.9414">32.9414°N</geoCoordinate>
<geoCoordinate id="296F1C297611F9FAEC4D860AC82BFD02" direction="west" orientation="longitude" precision="5" value="-86.5159">86.5159°W</geoCoordinate>
); UF 413800, Coosa River at Lonigan Shoals, St. Clair County, Alabama (~
<geoCoordinate id="119C217AD85C0A7EB7C37546C530A07B" direction="north" orientation="latitude" precision="5" value="33.7627">33.7627°N</geoCoordinate>
,
<geoCoordinate id="92A89394F7A6BF829728034FB90A37DE" direction="west" orientation="longitude" precision="5" value="-86.0447">86.0447°W</geoCoordinate>
); UF 81085, Choccolocco Creek at Jackson Shoals, Alabama (
<geoCoordinate id="C896A5A948F9DE9E1C0DD936BF455108" direction="north" orientation="latitude" precision="5" value="33.545">33.5450°N</geoCoordinate>
,
<geoCoordinate id="774A183BFBBC6D2FEB5594386421EED6" direction="west" orientation="longitude" precision="5" value="-86.0896">86.0896°W</geoCoordinate>
); UF 18202, Coosa River; UF 416913, Coosa River at Wetumpka, Elmore County, Alabama (~
<geoCoordinate id="6D43DCD336C5AFDE79BE7496C697E3E6" direction="north" orientation="latitude" precision="5" value="32.5396">32.5396°N</geoCoordinate>
,
<geoCoordinate id="4233523B0B9846DA3CCAB16E91089E97" direction="west" orientation="longitude" precision="5" value="-86.2056">86.2056°W</geoCoordinate>
); UF 416911, Coosa River at Wetumpka, Elmore County, Alabama (~
<geoCoordinate id="3282423BF0C41FE23119BA9DB9923F56" direction="north" orientation="latitude" precision="5" value="32.5396">32.5396°N</geoCoordinate>
,
<geoCoordinate id="18AC09DAD46D3710D716F1A23B02122B" direction="west" orientation="longitude" precision="5" value="-86.2056">86.2056°W</geoCoordinate>
); UF 416908, Waxahatchee Creek 4 mi above mouth, Shelby County, Alabama (~
<geoCoordinate id="C58E952317ED5F670712A284376FC484" direction="north" orientation="latitude" precision="5" value="33.0335">33.0335°N</geoCoordinate>
,
<geoCoordinate id="388B177305A48C263DC445C13579F052" direction="west" orientation="longitude" precision="5" value="-86.5787">86.5787°W</geoCoordinate>
); UF 416920, Kellys [sic, Kelly] Creek 2 mi above mouth, St. Clair County, Alabama (~
<geoCoordinate id="78A0A93F91E853DE4227543A692ED55A" direction="north" orientation="latitude" precision="5" value="33.4348">33.4348°N</geoCoordinate>
,
<geoCoordinate id="0F53346B7818A64D45BF3628398959D5" direction="west" orientation="longitude" precision="5" value="-86.3539">86.3539°W</geoCoordinate>
); UF 416897,
<pageBreakToken id="45DE259440FAC203852CF7D7FB029CB5" pageId="5" pageNumber="26" start="start">Tallasahatchee</pageBreakToken>
[sic, Tallaseehatchee] Creek 2-3 mi east of Childersburg, Talladega County, Alabama (~
<geoCoordinate id="6690582104D4EDFAE047161473A2AB4F" direction="north" orientation="latitude" precision="5" value="33.2841">33.2841°N</geoCoordinate>
,
<geoCoordinate id="8A5B2D241E5C169E87B5A0D1DC7918CF" direction="west" orientation="longitude" precision="5" value="-86.3098">86.3098°W</geoCoordinate>
); UF 416917, Beeswax Creek, Shelby County, Alabama; NCSM 59354, Terrapin Creek 7 mi south of Centre, Cherokee County, Alabama (~
<geoCoordinate id="0F7813B35C0B314265B5FB7E126CFF3C" direction="north" orientation="latitude" precision="5" value="34.0639">34.0639°N</geoCoordinate>
,
<geoCoordinate id="1FEAF3EA3DFEAF32AB757888EF45E10A" direction="west" orientation="longitude" precision="5" value="-85.6146">85.6146°W</geoCoordinate>
); NCSM 59359, Tallahatchie [sic, Tallasseehatchee] Creek, Calhoun County, Alabama (
<geoCoordinate id="EE5799A5D5DC6CA4C4E6D4B2DC00022D" direction="north" orientation="latitude" precision="5" value="33.7785">33.7785°N</geoCoordinate>
,
<geoCoordinate id="115B14EB71C304300C53051FC71B214B" direction="west" orientation="longitude" precision="5" value="-85.9908">85.9908°W</geoCoordinate>
); NCSM 59358, Ohatchee Creek, Calhoun County, Alabama (
<geoCoordinate id="87AF47DC1A552404DF66DF65AEA62734" direction="north" orientation="latitude" precision="5" value="33.7801">33.7801°N</geoCoordinate>
,
<geoCoordinate id="66420B24F4EFDDE1368C249CEEDEE5A3" direction="west" orientation="longitude" precision="5" value="-85.9972">85.9972°W</geoCoordinate>
); USNM 504866, Coosa River, near Wilsonville, Shelby County, Alabama (~
<geoCoordinate id="712B91ECDB012AD51FBBB87AC67E3E38" direction="north" orientation="latitude" precision="5" value="33.2162">33.2162°N</geoCoordinate>
,
<geoCoordinate id="208AF658B7861BC2BDA40D6CC1A94AC4" direction="west" orientation="longitude" precision="5" value="-86.4645">86.4645°W</geoCoordinate>
); USNM 504867, Coosa River 1 mi from Wilsonville, Shelby County, Alabama (~
<geoCoordinate id="D24017827BAA6FA3AC410EB7FC7F5FCB" direction="north" orientation="latitude" precision="5" value="33.2162">33.2162°N</geoCoordinate>
,
<geoCoordinate id="299931FCBB8B8896EBCB11986E500253" direction="west" orientation="longitude" precision="5" value="-86.4645">86.4645°W</geoCoordinate>
); USNM 336408, Coosa River at The Bar, Chilton County, Alabama (~
<geoCoordinate id="DDCEDBD9441FF8AAB3139672AE7A29F8" direction="north" orientation="latitude" precision="5" value="32.7976">32.7976°N</geoCoordinate>
,
<geoCoordinate id="78734D303B441F75CA3FE2F4E3E36839" direction="west" orientation="longitude" precision="5" value="-86.4348">86.4348°W</geoCoordinate>
); USNM 1437765, Coosa River, Shelby County, Alabama (
<geoCoordinate id="C4C31E2F9AEE91674EEABE49D01DCE19" direction="north" orientation="latitude" precision="5" value="33.3744">33.3744°N</geoCoordinate>
,
<geoCoordinate id="F0D8E3AA187D41E402ACD850272EA8F4" direction="west" orientation="longitude" precision="5" value="-86.355">86.3550°W</geoCoordinate>
); USNM 1249597, Choccolocco Creek, Talladega County,
<pageBreakToken id="5D9665DDFC9EFE5BE0300FAD7078399F" pageId="6" pageNumber="27" start="start">Alabama</pageBreakToken>
(
<geoCoordinate id="A1BD5331C930A92784FFA4CEA4F56FA6" direction="north" orientation="latitude" precision="5" value="33.5445">33.5445°N</geoCoordinate>
,
<geoCoordinate id="7B6BD7AC14C85EE01990B3AE913E78EA" direction="west" orientation="longitude" precision="5" value="-86.0414">86.0414°W</geoCoordinate>
); USNM 1249600, Buxahatchee Creek, Shelby County, Alabama (
<geoCoordinate id="DDAABF74C742E29EDBECCF3DA36E5407" direction="north" orientation="latitude" precision="5" value="33.0727">33.0727°N</geoCoordinate>
,
<geoCoordinate id="D6D06CF752C08AF6819B697076E0DCBB" direction="west" orientation="longitude" precision="5" value="-86.6775">86.6775°W</geoCoordinate>
); USNM 1437762, Ohatchee Creek, Calhoun County, Alabama (
<geoCoordinate id="8D6D32DF928DD4750B91212C00D8636A" direction="north" orientation="latitude" precision="5" value="33.7795">33.7795°N</geoCoordinate>
,
<geoCoordinate id="A1D2C4B9A04B89520813688FB9268295" direction="west" orientation="longitude" precision="5" value="-86.0002">86.0002°W</geoCoordinate>
); USNM 1437763, Ohatchee Creek, Calhoun County, Alabama (
<geoCoordinate id="488065258C457AE045F3953F66667A90" direction="north" orientation="latitude" precision="5" value="33.7795">33.7795°N</geoCoordinate>
,
<geoCoordinate id="1E7F3A8409DDC5D01A36E3C5BEE72C81" direction="west" orientation="longitude" precision="5" value="-86.0002">86.0002°W</geoCoordinate>
); USNM 1437764, Ohatchee Creek, Calhoun County, Alabama (
<geoCoordinate id="F8917331612CDA2A991E36D94F320D9F" direction="north" orientation="latitude" precision="5" value="33.7795">33.7795°N</geoCoordinate>
,
<geoCoordinate id="2E50223A36B7C01263989BC7573FABB9" direction="west" orientation="longitude" precision="5" value="-86.0002">86.0002°W</geoCoordinate>
).
</paragraph>
</subSubSection>
<subSubSection id="BE4884737DFF90723B36C6C070B6B6E0" pageId="6" pageNumber="27" type="diagnosis">
<paragraph id="EAF4706F45B7E609131400ADF93972BE" pageId="6" pageNumber="27">Diagnosis.</paragraph>
<paragraph id="874E01E4363995C4BB0DC3019BEBAE63" pageId="6" pageNumber="27">Shell ovate, two to four whorls, spire often reduced to obsolete but sometimes elevated with obtuse apex. Aperture large, ovate, at least half the height of body whorl. Reddish brown spiral bands typically present, usually four in number, almost always interrupted. Columella often purple. Head-foot and mantle pigmented orange, mottled with black, with one transverse black band across middle of snout and one transverse black band across middle of head. Clutches small (&lt;6 eggs), with minimal organic and/or inorganic matter incorporated into external casings.</paragraph>
</subSubSection>
<subSubSection id="A00D2FCCDE58BFCFA5BC69DB6DD62FE5" pageId="6" pageNumber="27" type="distribution">
<paragraph id="3F35BFBC476C4E266FFAE4530E6171E6" pageId="6" pageNumber="27">Historical distribution.</paragraph>
<paragraph id="01D1D49E956D8F1C3066AE4E8C5DF953" pageId="6" pageNumber="27">Coosa River above the Fall Line from Wetumpka, Alabama, upstream to the confluence of Terrapin Creek and the Coosa River in Cherokee County, Alabama. Some large Coosa River tributaries including Choccolocco, Buxahatchee, Talladega, and Terrapin creeks.</paragraph>
</subSubSection>
<subSubSection id="C98DD5E81D27AC96FB532F5DE1ABDC69" pageId="6" pageNumber="27" type="distribution">
<paragraph id="4B33D858C1A4B9177C19C79AF69553FD" pageId="6" pageNumber="27">Current distribution.</paragraph>
<paragraph id="7B11F139A9A84B88C4C9138E5DC2700C" pageId="6" pageNumber="27">Four disjunct populations: Choccolocco Creek, Talladega County, Alabama; Buxahatchee and Watson creeks, Shelby County, Alabama; Ohatchee Creek, Calhoun County, Alabama; Logan Martin Dam tailwaters of the Coosa River, Shelby-Talladega counties, Alabama.</paragraph>
</subSubSection>
<subSubSection id="183E1D279E5CE2C6B547301639CE4887" pageId="6" pageNumber="27" type="remarks">
<paragraph id="D7C8A359A41E89F69AC80750FD8C4367" pageId="6" pageNumber="27">Remarks.</paragraph>
<paragraph id="F0BE1E6C9E69769DE8B2EF8CF7E5490D" pageId="6" pageNumber="27">
USNM 121295 (Fig. 3A) originates from the Lea collection, is from the published
<taxonomicName id="0A2793BA0FF099762BDF253DDC5DFFD5" lsidName="L. coosaensis" pageId="6" pageNumber="27" rank="species" species="coosaensis">L. coosaensis</taxonomicName>
type locality, leg. Showalter, and is the shell figured by
<bibRefCitation id="0CC1CB76189A5A3AEF13E90EE7E45FE8" author="Lea, I" journalOrPublisher="Journal of the Academy of Natural Sciences of Philadelphia" pageId="12" pageNumber="33" pagination="217 - 356" title="New Melanida of the United States." volume="5" year="1863">Lea (1863)</bibRefCitation>
, with the number 65 inked onto the apertural aspect of the body whorl corresponding to the figure number.
<bibRefCitation id="B3AD86A8F0D3FA67616B350A568F2423" author="Lea, I" journalOrPublisher="Journal of the Academy of Natural Sciences of Philadelphia" pageId="12" pageNumber="33" pagination="217 - 356" title="New Melanida of the United States." volume="5" year="1863">Lea (1863)</bibRefCitation>
indicated that he had six specimens, yet there are seven specimens distributed among the three simultaneously accessioned lots now registered as USNM 121294 (Fig. 3F-I), USNM 1456804 (see photographs on FigShare, https://doi.org/10.6084/m9.figshare.5084272.v1), and USNM 121296 (Fig. 3J). Consequently, either the specimen count in
<bibRefCitation id="783FEDEDF47BBA40FF04F63A7356EC4D" author="Lea, I" journalOrPublisher="Journal of the Academy of Natural Sciences of Philadelphia" pageId="12" pageNumber="33" pagination="217 - 356" title="New Melanida of the United States." volume="5" year="1863">Lea (1863)</bibRefCitation>
was in error, or one of the included shells was acquired subsequently and has no type status.
</paragraph>
<paragraph id="0E1A4B89A90A78A3E60825C3E24C9B7A" pageId="6" pageNumber="27">
<bibRefCitation id="A36B82249BAB8712ABEB61089DBBF374" author="Lea, I" journalOrPublisher="Proceedings of the Academy of Natural Sciences, Philadelphia" pageId="12" pageNumber="33" pagination="54 - 55" title="Descriptions of new species of Schizostoma, Anculosa, and Lithasia." volume="13" year="1861">Lea (1861)</bibRefCitation>
described
<taxonomicName id="02015C3B666DD4AAC7CE7FBB47CA8E78" class="Gastropoda" family="Pleuroceridae" genus="Anculosa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anculosa coosaensis" pageId="6" pageNumber="27" phylum="Mollusca" rank="species" species="coosaensis">Anculosa coosaensis</taxonomicName>
from the Coosa River and provided a brief description in Latin. In a subsequent extended description in English,
<bibRefCitation id="85463CA9A2CB03CFE14AB02E61D592E3" author="Lea, I" journalOrPublisher="Journal of the Academy of Natural Sciences of Philadelphia" pageId="12" pageNumber="33" pagination="217 - 356" title="New Melanida of the United States." volume="5" year="1863">Lea (1863)</bibRefCitation>
characterized the species as, &quot;smooth, obtusely conical, thick, dark horn-color, very much banded; spire elevated, obtuse at the apex; sutures very much impressed; whorls four, very much constricted below the sutures, the last large; aperture rounded, white, much banded within; columella thickened, incurved, dark purple; outer lip acute and expanded.&quot; In the remarks, he commented that the aperture is more than half the length of the shell and that the bands may be interrupted. This description, as well as the type material (Fig. 3
<normalizedToken id="DCB38474D8BD314B0FB210B9AC5D2F0E" originalValue="F–J">F-J</normalizedToken>
) match the current concept of the Painted Rocksnail (Fig. 2). Although formerly considered a synonym of
<taxonomicName id="16EEE7C306A864A683DC79717AD7D936" lsidName="L. taeniata" pageId="6" pageNumber="27" rank="species" species="taeniata">L. taeniata</taxonomicName>
,
<taxonomicName id="D3750AFD412A1CE4C2E421413E276BFC" lsidName="L. coosaensis" pageId="6" pageNumber="27" rank="species" species="coosaensis">L. coosaensis</taxonomicName>
is the oldest available name for the Painted Rocksnail.
</paragraph>
<paragraph id="1AF8F16122235812444EB65C51A60648" pageId="6" pageNumber="27">
We have been unable to locate type material of
<taxonomicName id="C3F5111DEA3506CE024E774F8E6AF553" class="Gastropoda" family="Pleuroceridae" genus="Anculosa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anculosa taeniata" pageId="6" pageNumber="27" phylum="Mollusca" rank="subspecies" species="taeniata" subspecies="lucida">Anculosa taeniata lucida</taxonomicName>
Goodrich, 1944. No holotype was designated, nor was a figure provided, but based on the original description and the type locality of tributaries of the Coosa River, we conclude that this entity does not merit recognition at the subspecies level and synonymize it with
<taxonomicName id="A5DDC54979D8C04F1B3FB3F1D05DD88E" lsidName="L. coosaensis" pageId="6" pageNumber="27" rank="species" species="coosaensis">L. coosaensis</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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