280 lines
36 KiB
XML
280 lines
36 KiB
XML
<document id="1114B5FECB0388BD25F4DEB836A752BA" ID-DOI="10.11646/zootaxa.5369.2.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="10146170" ID-ZooBank="42563452-CF2B-47A6-B0B0-485AC4F320BD" IM.bibliography_approvedBy="diego" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="diego" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="diego" IM.taxonomicNames_approvedBy="diego" IM.treatments_approvedBy="diego" checkinTime="1700185584045" checkinUser="plazi" docAuthor="Shavrin, Alexey V." docDate="2023" docId="038DBF6E5428CC4FFF06F8FBFF14FDB8" docLanguage="en" docName="zootaxa.5369.2.1.pdf" docOrigin="Zootaxa 5369 (2)" docSource="https://www.mapress.com/zt/article/download/zootaxa.5369.2.1/52237" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Omalium bilobum Shavrin 2023, sp. n." docType="treatment" docVersion="1" lastPageNumber="167" masterDocId="FFB4C7165425CC5FFF91FFD8FFD8FF82" masterDocTitle="A revision of the genus Omalium Gravenhorst, 1802 (Coleoptera: Staphylinidae: Omaliinae) of the Himalayan region" masterLastPageNumber="206" masterPageNumber="151" pageNumber="164" updateTime="1700578467175" updateUser="diego" zenodo-license-document="CLOSED">
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<mods:title id="C1331DD06AB66FBFC26CAB3C5C947171">A revision of the genus Omalium Gravenhorst, 1802 (Coleoptera: Staphylinidae: Omaliinae) of the Himalayan region</mods:title>
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<mods:namePart id="5082913EDC935B32F2D7831B03FEEE3C">Shavrin, Alexey V.</mods:namePart>
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<mods:title id="6E102229CF01B87BC80784FA79783837">Zootaxa</mods:title>
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<mods:part id="DC8F49776F0565F7A4D798699DF93454">
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<mods:date id="D18B561089FF1E894FC09DB07F1E9126">2023</mods:date>
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<mods:number id="BC000E58BBEC396EB8DCD79010F95F2A">2023-11-09</mods:number>
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<mods:number id="B8C4887F06DD42432587D0A09177C015">5369</mods:number>
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<mods:url id="98293FE84A565A3FE9817AC05D0419CF">https://www.mapress.com/zt/article/download/zootaxa.5369.2.1/52237</mods:url>
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<mods:classification id="32E06702A9C138771DC19613B31D6D43">journal article</mods:classification>
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<mods:identifier id="537BE12FE4DB91B74E8667668145D90A" type="DOI">10.11646/zootaxa.5369.2.1</mods:identifier>
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<mods:identifier id="8B0E891FD0736AFBC82190CD54FA327B" type="ISSN">1175-5326</mods:identifier>
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<treatment id="038DBF6E5428CC4FFF06F8FBFF14FDB8" LSID="urn:lsid:plazi:treatment:038DBF6E5428CC4FFF06F8FBFF14FDB8" httpUri="http://treatment.plazi.org/id/038DBF6E5428CC4FFF06F8FBFF14FDB8" lastPageId="16" lastPageNumber="167" pageId="13" pageNumber="164">
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<subSubSection id="C33E5DF35428CC52FF06F8FBFDEBF8BC" box="[151,563,1827,1854]" pageId="13" pageNumber="164" type="nomenclature">
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<paragraph id="8B9B0E785428CC52FF06F8FBFDEBF8BC" blockId="13.[151,563,1827,1890]" box="[151,563,1827,1854]" pageId="13" pageNumber="164">
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<heading id="D0D3B9145428CC52FF06F8FBFDEBF8BC" bold="true" box="[151,563,1827,1854]" fontSize="11" level="1" pageId="13" pageNumber="164" reason="1">
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<emphasis id="B950D26A5428CC52FF06F8FBFDEBF8BC" bold="true" box="[151,563,1827,1854]" pageId="13" pageNumber="164">
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3.2.2.2.2.
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<taxonomicName id="4C2475FB5428CC52FE9CF8FBFE31F8BC" authority="Shavrin, 2023" authorityName="Shavrin" authorityYear="2023" box="[269,489,1827,1854]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="164" phylum="Arthropoda" rank="species" species="bilobum" status="sp. nov.">
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<emphasis id="B950D26A5428CC52FE9CF8FBFE31F8BC" bold="true" box="[269,489,1827,1854]" italics="true" pageId="13" pageNumber="164">Omalium bilobum</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A2636F115428CC52FE60F8FCFDEBF8BC" box="[497,563,1828,1854]" pageId="13" pageNumber="164" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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||
</heading>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C33E5DF35428CC52FF06F89FFEA3F8E0" box="[151,379,1863,1890]" pageId="13" pageNumber="164" type="description">
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<paragraph id="8B9B0E785428CC52FF06F89FFEA3F8E0" blockId="13.[151,563,1827,1890]" box="[151,379,1863,1890]" pageId="13" pageNumber="164">
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(
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<figureCitation id="131F12FD5428CC52FF0EF89FFF2EF8E0" box="[159,246,1863,1890]" captionStart="FIGURES 8–18" captionStartId="6.[151,264,1896,1920]" captionTargetBox="[154,1433,564,1870]" captionTargetId="figure-184@6.[151,1436,561,1873]" captionTargetPageId="6" captionText="FIGURES 8–18. Female accessory sclerite (Figs 8, 10–16, 18) and spermatheca (Figs 9, 17) of Omalium: 8–9—O. acutangulum, 10—O. amicorum, 11—O. amplissimum, 12—O. bilobum, 13—O. alticola, 14—O.beesoni, 15—O. altivagans, 16–17—O. marginatum, 18—O. scapulare. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10146176" httpUri="https://zenodo.org/record/10146176/files/figure.png" pageId="13" pageNumber="164">Figs 12</figureCitation>
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,
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<figureCitation id="131F12FD5428CC52FE93F89FFEF8F8E0" box="[258,288,1863,1890]" captionStart="FIGURES 24–26" captionStartId="11.[151,264,1490,1514]" captionTargetBox="[164,1420,463,1452]" captionTargetId="figure-64@11.[151,1435,453,1467]" captionTargetPageId="11" captionText="FIGURES 24–26. Habitus of Omalium: 24—O. amplissimum (paratype), 25—O. bilobum (paratype), 26—O. altivagans (Kashmir). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10146184" httpUri="https://zenodo.org/record/10146184/files/figure.png" pageId="13" pageNumber="164">25</figureCitation>
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,
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<figureCitation id="131F12FD5428CC52FEBDF890FEABF8E0" box="[300,371,1864,1890]" captionStart-0="FIGURES 30–32" captionStart-1="FIGURE 33" captionStartId-0="14.[151,264,1598,1622]" captionStartId-1="15.[151,250,1334,1358]" captionTargetBox-0="[154,1433,672,1570]" captionTargetBox-1="[151,1436,417,1310]" captionTargetId-0="figure-236@14.[151,1436,669,1575]" captionTargetId-1="figure-126@15.[151,1436,417,1310]" captionTargetPageId-0="14" captionTargetPageId-1="15" captionText-0="FIGURES 30–32. Aedeagus of Omalium bilobum (holotype): 30—aedeagus, parameral view, 31—aedeagus, lateral view, 32—median lobe, lateral view. Scale bar: 0.1 mm." captionText-1="FIGURE 33. Distribution of Omalium in the Himalayan region: O. bilobum sp. n. (circle), O. marginatum (squares), O. altivagans sp. n. (triangle), O. humerale (diamond)." figureDoi-0="http://doi.org/10.5281/zenodo.10146188" figureDoi-1="http://doi.org/10.5281/zenodo.10146190" httpUri-0="https://zenodo.org/record/10146188/files/figure.png" httpUri-1="https://zenodo.org/record/10146190/files/figure.png" pageId="13" pageNumber="164">30–33</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C33E5DF35428CC51FF06F848FC90FF57" lastPageId="14" lastPageNumber="165" pageId="13" pageNumber="164" type="materials_examined">
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<paragraph id="8B9B0E785428CC52FF06F848FD67F870" blockId="13.[151,1437,1935,2034]" pageId="13" pageNumber="164">
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<emphasis id="B950D26A5428CC52FF06F848FE9DF82B" bold="true" box="[151,325,1935,1962]" pageId="13" pageNumber="164">Type material.</emphasis>
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<materialsCitation id="3B4C04255428CC52FEDAF857FC7CF84F" collectorName="Lobl & Smetana'" country="Nepal" elevation="2250" location="Kuwapani" municipality="Foret" pageId="13" pageNumber="164" specimenCount="1" stateProvince="Kosi" typeStatus="holotype">
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<typeStatus id="549FB0DA5428CC52FEDAF857FE6BF82B" box="[331,435,1935,1961]" pageId="13" pageNumber="164" type="holotype">Holotype</typeStatus>
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♁ (dissected; apical antennomere of left antenna missing): ‘E.
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<collectingCountry id="F3334EE85428CC52FBE3F848FB16F828" box="[1138,1230,1936,1962]" name="Nepal" pageId="13" pageNumber="164">NEPAL</collectingCountry>
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:
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<collectingRegion id="49E0C09A5428CC52FB49F848FAC1F828" box="[1240,1305,1936,1962]" country="Nepal" name="Kosi" pageId="13" pageNumber="164">KOSI</collectingRegion>
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<collectingMunicipality id="6BFF94025428CC52FABDF848FABFF828" box="[1324,1383,1936,1962]" pageId="13" pageNumber="164">Forêt</collectingMunicipality>
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N-E
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<location id="8EFB58A35428CC52FF06F86BFED2F84F" LSID="urn:lsid:plazi:treatment:038DBF6E5428CC4FFF06F8FBFF14FDB8:8EFB58A35428CC52FF06F86BFED2F84F" box="[151,266,1971,1997]" country="Nepal" municipality="Foret" name="Kuwapani" pageId="13" pageNumber="164" stateProvince="Kosi">Kuwapani</location>
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<quantity id="4CDCA39D5428CC52FE8EF86BFEAEF84C" box="[287,374,1971,1998]" metricMagnitude="3" metricUnit="m" metricValue="2.25" pageId="13" pageNumber="164" unit="m" value="2250.0">
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<elevation id="0009E94B5428CC52FE8EF86BFEAEF84C" box="[287,374,1971,1998]" metricMagnitude="3" metricUnit="m" metricValue="2.25" pageId="13" pageNumber="164" unit="m" value="2250.0">2250 m</elevation>
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</quantity>
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, 24.IV.[19]84 | [I.]
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<collectorName id="26D16BAE5428CC52FDC1F86BFD5DF84F" box="[592,645,1971,1997]" pageId="13" pageNumber="164">Löbl</collectorName>
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- [A.]
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<collectorName id="26D16BAE5428CC52FD43F86CFCE2F84C" box="[722,826,1972,1998]" pageId="13" pageNumber="164">Smetana’</collectorName>
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<printed
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</materialsCitation>
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>, ‘
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<typeStatus id="549FB0DA5428CC52FC53F86CFB83F84C" box="[962,1115,1972,1998]" pageId="13" pageNumber="164" type="holotype">HOLOTYPE</typeStatus>
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<taxonomicName id="4C2475FB5428CC52FBE1F86BFB00F84F" authorityName="Gravenhorst" authorityYear="1802" box="[1136,1240,1971,1997]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="164" phylum="Arthropoda" rank="genus">
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<emphasis id="B950D26A5428CC52FBE1F86BFB00F84F" box="[1136,1240,1971,1997]" italics="true" pageId="13" pageNumber="164">Omalium</emphasis>
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</taxonomicName>
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<taxonomicName id="4C2475FB5428CC52FB7CF86BFA91F84F" authority="Shavrin, 2023" authorityName="Shavrin" authorityYear="2023" box="[1261,1353,1971,1997]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="164" phylum="Arthropoda" rank="species" species="bilobum" status="sp. nov.">
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<emphasis id="B950D26A5428CC52FB7CF86BFA91F84F" box="[1261,1353,1971,1997]" italics="true" pageId="13" pageNumber="164">bilobum</emphasis>
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</taxonomicName>
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<emphasis id="B950D26A5428CC52FAC1F86CFA48F84C" bold="true" box="[1360,1424,1972,1998]" pageId="13" pageNumber="164">
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<taxonomicNameLabel id="A2636F115428CC52FAC1F86CFA48F84C" box="[1360,1424,1972,1998]" pageId="13" pageNumber="164" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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| Shavrin A.V. des. 2023’ <red, printed> (MHNG).
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</paragraph>
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<paragraph id="8B9B0E78542BCC51FF56FF4FFC90FF57" blockId="14.[151,1437,150,645]" pageId="14" pageNumber="165">
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<materialsCitation id="3B4C0425542BCC51FF56FF4FFB34FF33" box="[199,1260,150,177]" collectorName="Lobl & Smetana'" country="Nepal" elevation="2250" location="Kuwapani" municipality="Foret" pageId="14" pageNumber="165" specimenCount="1" specimenCount-female="1" stateProvince="Kosi" typeStatus="paratype">
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<typeStatus id="549FB0DA542BCC51FF56FF4FFEF1FF33" box="[199,297,151,177]" pageId="14" pageNumber="165" type="paratype">Paratype</typeStatus>
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<specimenCount id="9D22C5F1542BCC51FEA3FF4FFE9FFF33" box="[306,327,151,177]" count="1" pageId="14" pageNumber="165" type="female">♀</specimenCount>
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(
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<figureCitation id="131F12FD542BCC51FEC8FF4FFE77FF33" box="[345,431,150,177]" captionStart="FIGURES 24–26" captionStartId="11.[151,264,1490,1514]" captionTargetBox="[164,1420,463,1452]" captionTargetId="figure-64@11.[151,1435,453,1467]" captionTargetPageId="11" captionText="FIGURES 24–26. Habitus of Omalium: 24—O. amplissimum (paratype), 25—O. bilobum (paratype), 26—O. altivagans (Kashmir). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10146184" httpUri="https://zenodo.org/record/10146184/files/figure.png" pageId="14" pageNumber="165">Fig. 25</figureCitation>
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; dissected): same data as the holotype, with additional red printed label
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</materialsCitation>
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: ‘
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<typeStatus id="549FB0DA542BCC51FB6CFF4FFA55FF33" box="[1277,1421,151,177]" pageId="14" pageNumber="165" type="paratype">PARATYPE</typeStatus>
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<taxonomicName id="4C2475FB542BCC51FF06FF63FF27FF57" authorityName="Gravenhorst" authorityYear="1802" box="[151,255,187,213]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="165" phylum="Arthropoda" rank="genus">
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<emphasis id="B950D26A542BCC51FF06FF63FF27FF57" box="[151,255,187,213]" italics="true" pageId="14" pageNumber="165">Omalium</emphasis>
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</taxonomicName>
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<taxonomicName id="4C2475FB542BCC51FE82FF63FEB7FF57" authority="Shavrin, 2023" authorityName="Shavrin" authorityYear="2023" box="[275,367,187,213]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="165" phylum="Arthropoda" rank="species" species="bilobum" status="sp. nov.">
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<emphasis id="B950D26A542BCC51FE82FF63FEB7FF57" box="[275,367,187,213]" italics="true" pageId="14" pageNumber="165">bilobum</emphasis>
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</taxonomicName>
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<emphasis id="B950D26A542BCC51FEE7FF63FE6DFF57" bold="true" box="[374,437,187,213]" pageId="14" pageNumber="165">
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<taxonomicNameLabel id="A2636F11542BCC51FEE7FF63FE6DFF57" box="[374,437,187,213]" pageId="14" pageNumber="165" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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| Shavrin A.V. des. 2023’ (MHNG).
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C33E5DF3542BCC50FF56FF06FEC2F86B" lastPageId="15" lastPageNumber="166" pageId="14" pageNumber="165" type="description">
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||
<paragraph id="8B9B0E78542BCC51FF56FF06FA59FEC3" blockId="14.[151,1437,150,645]" pageId="14" pageNumber="165">
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<emphasis id="B950D26A542BCC51FF56FF06FE81FF7A" bold="true" box="[199,345,222,248]" pageId="14" pageNumber="165">Description.</emphasis>
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Measurements (n=2): HW: 0.47–0.51; HL: 0.37; OL: 0.15; TL: 0.07–0.09; AL (
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<typeStatus id="549FB0DA542BCC51FB76FF07FA88FF7B" box="[1255,1360,223,249]" pageId="14" pageNumber="165" type="holotype">holotype</typeStatus>
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): 0.93; PL: 0.37–0.40; PWmax: 0.60–0.62; PWmin: 0.56–0.58; ESL: 0.77–0.83; EW: 0.84–0.89; MTbL (
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<typeStatus id="549FB0DA542BCC51FB76FEDBFA88FE9F" box="[1255,1360,259,285]" pageId="14" pageNumber="165" type="holotype">holotype</typeStatus>
|
||
): 0.46; MTrL (
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<typeStatus id="549FB0DA542BCC51FF79FEFFFE89FEC3" box="[232,337,295,321]" pageId="14" pageNumber="165" type="holotype">holotype</typeStatus>
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||
): 0.17 (MTrL 1–4: 0.07; MTrL 5: 0.10); AW: 0.80–0.86; AedL: 0.45; BL: 2.80 (
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<typeStatus id="549FB0DA542BCC51FB5CFEFFFAECFEC3" box="[1229,1332,295,321]" pageId="14" pageNumber="165" type="holotype">holotype</typeStatus>
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||
)–2.97.
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</paragraph>
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<paragraph id="8B9B0E78542BCC51FF56FE93FABEFD07" blockId="14.[151,1437,150,645]" pageId="14" pageNumber="165">
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||
Habitus as in
|
||
<figureCitation id="131F12FD542BCC51FEF0FE93FE6CFEE7" box="[353,436,330,357]" captionStart="FIGURES 24–26" captionStartId="11.[151,264,1490,1514]" captionTargetBox="[164,1420,463,1452]" captionTargetId="figure-64@11.[151,1435,453,1467]" captionTargetPageId="11" captionText="FIGURES 24–26. Habitus of Omalium: 24—O. amplissimum (paratype), 25—O. bilobum (paratype), 26—O. altivagans (Kashmir). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10146184" httpUri="https://zenodo.org/record/10146184/files/figure.png" pageId="14" pageNumber="165">Fig. 25</figureCitation>
|
||
. Body reddish-brown, with distinctly darker head and abdomen, and slightly paler lateral portions of pronotum and middle portions of elytra; antennomeres 6–11 brown; mouthparts, antennomeres 1–5 and legs yellowish. Punctation of head dense and moderately deep, distinctly sparser on clypeus and denser, slightly larger and deeper in middle and infraorbital portions, interstices between punctures in middle as broad as two to three nearest punctures; nuchal constriction with moderately sparse and large punctation; punctation of pronotum dense, markedly larger than that on head, slightly sparser on lateral and mediobasal portions; punctation of elytra dense, slightly coarser and deeper than that on pronotum, denser and finer in parascutellar portion and sparser in middle portion of each elytron. Head without microsculpture except of anterior portion of clypeus with distinct transverse microreticulation; pronotum and elytra without sculpture; scutellum with fine isodiametric meshes.
|
||
</paragraph>
|
||
<caption id="DF5B5EF0542BCC51FF06F9E6FD4DF9F8" ID-DOI="http://doi.org/10.5281/zenodo.10146188" ID-Zenodo-Dep="10146188" httpUri="https://zenodo.org/record/10146188/files/figure.png" pageId="14" pageNumber="165" startId="14.[151,264,1598,1622]" targetBox="[154,1433,672,1570]" targetPageId="14" targetType="figure">
|
||
<paragraph id="8B9B0E78542BCC51FF06F9E6FD4DF9F8" blockId="14.[151,1436,1598,1658]" pageId="14" pageNumber="165">
|
||
<emphasis id="B950D26A542BCC51FF06F9E6FE80F9D4" bold="true" box="[151,344,1598,1622]" pageId="14" pageNumber="165">FIGURES 30–32.</emphasis>
|
||
Aedeagus of
|
||
<taxonomicName id="4C2475FB542BCC51FE7FF9E6FD72F9D4" authorityName="Shavrin" authorityYear="2023" box="[494,682,1598,1622]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="165" phylum="Arthropoda" rank="species" species="bilobum">
|
||
<emphasis id="B950D26A542BCC51FE7FF9E6FD72F9D4" box="[494,682,1598,1622]" italics="true" pageId="14" pageNumber="165">Omalium bilobum</emphasis>
|
||
</taxonomicName>
|
||
(holotype): 30—aedeagus, parameral view, 31—aedeagus, lateral view, 32—median lobe, lateral view. Scale bar: 0.1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B9B0E78542BCC50FF56F914FB30FE9F" blockId="14.[151,1437,1740,2019]" lastBlockId="15.[151,1436,150,393]" lastPageId="15" lastPageNumber="166" pageId="14" pageNumber="165">Head 1.2–1.3 times as broad as long, with moderately broad clypeus, and slightly convex supra-antennal elevations, posteriolateral margins of clypeus stretching posteriad toward level of middle length of eyes. Anteriomedian depressions moderately shallow, reaching level of anterior margin of eyes. Middle portion between eyes and infraorbital portions slightly elevated. Surface of head between anteocellar foveae with slightly elevated interspaces between punctures; surface on basal parts of infraorbital portions with longitudinal elevations between punctures; lateral portions between posterior margins of eyes and anteocellar fovea with three relatively long and slightly elevated wrinkles.Anteocellar foveae deep, sublinear, reaching level of middle length of eyes. Temples about twice shorter than longitudianl length of eyes. Distance between ocelli slightly less than twice as long as distance between ocellus and posterior margin of eye. Antennomeres 5–10 progressively broadened apicad; antennomere 2 slightly more than twice as long as scape, 3 about as long as and slightly narrower than 2, 4 very small, distinctly longer and shorter than 3, 5 insignificantly broader than 4, 6 slightly broader than 5, 7 distinctly broader than 6, 8 distinctly broader than 7, 9–10 slightly broader than 8, apical antennomere 1.3 times as long as 10.</paragraph>
|
||
<paragraph id="8B9B0E78542ACC50FF56FEFFFEFAFE0B" blockId="15.[151,1436,150,393]" pageId="15" pageNumber="166">Pronotum 1.5–1.6 times as broad as long, 1.2 times as broad as head, widest slightly above middle, with slightly protruded apical angles. Anterior margin widely rounded. Surface of disc with two distinct and wide longitudinal depressions.</paragraph>
|
||
<caption id="DF5B5EF0542ACC50FF06FAEEFD7FFAF0" ID-DOI="http://doi.org/10.5281/zenodo.10146190" ID-Zenodo-Dep="10146190" httpUri="https://zenodo.org/record/10146190/files/figure.png" pageId="15" pageNumber="166" startId="15.[151,250,1334,1358]" targetBox="[151,1436,417,1310]" targetPageId="15" targetType="figure">
|
||
<paragraph id="8B9B0E78542ACC50FF06FAEEFD7FFAF0" blockId="15.[151,1437,1334,1395]" pageId="15" pageNumber="166">
|
||
<emphasis id="B950D26A542ACC50FF06FAEEFEFCFACC" bold="true" box="[151,292,1334,1358]" pageId="15" pageNumber="166">FIGURE 33.</emphasis>
|
||
Distribution of
|
||
<taxonomicName id="4C2475FB542ACC50FE43FAEEFDE8FACC" authorityName="Gravenhorst" authorityYear="1802" box="[466,560,1334,1358]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="166" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B950D26A542ACC50FE43FAEEFDE8FACC" box="[466,560,1334,1358]" italics="true" pageId="15" pageNumber="166">Omalium</emphasis>
|
||
</taxonomicName>
|
||
in the Himalayan region:
|
||
<taxonomicName id="4C2475FB542ACC50FCDCFAEFFC1BFACC" authority="Shavrin, 2023" authorityName="Shavrin" authorityYear="2023" box="[845,963,1334,1358]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="166" phylum="Arthropoda" rank="species" species="bilobum" status="sp. n.">
|
||
<emphasis id="B950D26A542ACC50FCDCFAEFFC1BFACC" box="[845,963,1334,1358]" italics="true" pageId="15" pageNumber="166">O. bilobum</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B950D26A542ACC50FC5CFAEFFBD2FACD" bold="true" box="[973,1034,1335,1359]" pageId="15" pageNumber="166">
|
||
<taxonomicNameLabel id="A2636F11542ACC50FC5CFAEFFBD2FACD" box="[973,1034,1335,1359]" pageId="15" pageNumber="166" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(circle),
|
||
<taxonomicName id="4C2475FB542ACC50FBFFFAEFFAD4FACC" authorityName="Cameron" authorityYear="1941" box="[1134,1292,1335,1358]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="166" phylum="Arthropoda" rank="species" species="marginatum">
|
||
<emphasis id="B950D26A542ACC50FBFFFAEFFAD4FACC" box="[1134,1292,1335,1358]" italics="true" pageId="15" pageNumber="166">O. marginatum</emphasis>
|
||
</taxonomicName>
|
||
(squares),
|
||
<taxonomicName id="4C2475FB542ACC50FA15FAEFFED9FAF0" authority="Shavrin, 2023" authorityName="Shavrin" authorityYear="2023" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="166" phylum="Arthropoda" rank="species" species="altivagans" status="sp. n.">
|
||
<emphasis id="B950D26A542ACC50FA15FAEFFED9FAF0" italics="true" pageId="15" pageNumber="166">O. altivagans</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B950D26A542ACC50FE96FA83FE98FAF1" bold="true" box="[263,320,1371,1395]" pageId="15" pageNumber="166">
|
||
<taxonomicNameLabel id="A2636F11542ACC50FE96FA83FE98FAF1" box="[263,320,1371,1395]" pageId="15" pageNumber="166" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(triangle),
|
||
<taxonomicName id="4C2475FB542ACC50FE20FA83FDE8FAF0" authorityName="Cameron" authorityYear="1924" box="[433,560,1370,1394]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="166" phylum="Arthropoda" rank="species" species="humerale">
|
||
<emphasis id="B950D26A542ACC50FE20FA83FDE8FAF0" box="[433,560,1370,1394]" italics="true" pageId="15" pageNumber="166">O. humerale</emphasis>
|
||
</taxonomicName>
|
||
(diamond).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B9B0E78542ACC50FF56FA6FFBC1FA77" blockId="15.[151,1437,1463,2025]" pageId="15" pageNumber="166">Elytra slightly broader than long, twice as long as pronotum, indistinctly broadened in posterior portion. Mediobasal surface of each elytron with diagonal elevations between punctures.</paragraph>
|
||
<paragraph id="8B9B0E78542ACC50FF56FA26FC87F99A" blockId="15.[151,1437,1463,2025]" box="[199,863,1534,1561]" pageId="15" pageNumber="166">Metatarsi distinctly more than twice shorter than metatibia.</paragraph>
|
||
<paragraph id="8B9B0E78542ACC50FF56F9FAFDABF9BE" blockId="15.[151,1437,1463,2025]" box="[199,627,1570,1596]" pageId="15" pageNumber="166">Abdomen slightly broader than elytra.</paragraph>
|
||
<paragraph id="8B9B0E78542ACC50FF56F99EFDE9F8FC" blockId="15.[151,1437,1463,2025]" pageId="15" pageNumber="166">
|
||
Male. Antennomeres 9–10 strongly transverse. Posterior margin of abdominal tergite VIII straight. Posterior margin of abdominal sternite VIII rounded. Aedeagus wide and short, with very wide basal portion, gradually narrowed toward middle, with moderately short median lobe, from widest basal portion strongly narrowed toward middle and from middle broadened apicad toward apex with relatively wide excision in middle of apical margin; mediolateral portion with two elongate sclerotized curved accessory plates connected with middle part of median lobe and inner sides of parameres; parameres moderately wide, curved, slightly longer than apex of median lobe, with rounded apices and with three long apical setae; internal sac moderately short and wide (
|
||
<figureCitation id="131F12FD542ACC50FB0DF8C4FB37F8B4" box="[1180,1263,1820,1846]" captionStart="FIGURES 30–32" captionStartId="14.[151,264,1598,1622]" captionTargetBox="[154,1433,672,1570]" captionTargetId="figure-236@14.[151,1436,669,1575]" captionTargetPageId="14" captionText="FIGURES 30–32. Aedeagus of Omalium bilobum (holotype): 30—aedeagus, parameral view, 31—aedeagus, lateral view, 32—median lobe, lateral view. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10146188" httpUri="https://zenodo.org/record/10146188/files/figure.png" pageId="15" pageNumber="166">Fig. 30</figureCitation>
|
||
). Lateral aspect of the aedeagus as in
|
||
<figureCitation id="131F12FD542ACC50FE19F898FE02F8D8" box="[392,474,1856,1882]" captionStart="FIGURES 30–32" captionStartId="14.[151,264,1598,1622]" captionTargetBox="[154,1433,672,1570]" captionTargetId="figure-236@14.[151,1436,669,1575]" captionTargetPageId="14" captionText="FIGURES 30–32. Aedeagus of Omalium bilobum (holotype): 30—aedeagus, parameral view, 31—aedeagus, lateral view, 32—median lobe, lateral view. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10146188" httpUri="https://zenodo.org/record/10146188/files/figure.png" pageId="15" pageNumber="166">Fig. 31</figureCitation>
|
||
; apical portion of median lobe strongly narrowed toward apex, with slightly crenulate apical part of inner margin (
|
||
<figureCitation id="131F12FD542ACC50FE40F8BCFDFDF8FC" box="[465,549,1892,1918]" captionStart="FIGURES 30–32" captionStartId="14.[151,264,1598,1622]" captionTargetBox="[154,1433,672,1570]" captionTargetId="figure-236@14.[151,1436,669,1575]" captionTargetPageId="14" captionText="FIGURES 30–32. Aedeagus of Omalium bilobum (holotype): 30—aedeagus, parameral view, 31—aedeagus, lateral view, 32—median lobe, lateral view. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10146188" httpUri="https://zenodo.org/record/10146188/files/figure.png" pageId="15" pageNumber="166">Fig. 32</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9B0E78542ACC50FF56F85FFEC2F86B" blockId="15.[151,1437,1463,2025]" pageId="15" pageNumber="166">
|
||
Female. Antennomeres 9–10 slightly transverse. Posterior margins of abdominal tergite VIII and sternite VIII rounded. Accessory sclerite with wide basal portion and moderately wide median part (
|
||
<figureCitation id="131F12FD542ACC50FBE7F873FB12F847" box="[1142,1226,1963,1989]" captionStart="FIGURES 8–18" captionStartId="6.[151,264,1896,1920]" captionTargetBox="[154,1433,564,1870]" captionTargetId="figure-184@6.[151,1436,561,1873]" captionTargetPageId="6" captionText="FIGURES 8–18. Female accessory sclerite (Figs 8, 10–16, 18) and spermatheca (Figs 9, 17) of Omalium: 8–9—O. acutangulum, 10—O. amicorum, 11—O. amplissimum, 12—O. bilobum, 13—O. alticola, 14—O.beesoni, 15—O. altivagans, 16–17—O. marginatum, 18—O. scapulare. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10146176" httpUri="https://zenodo.org/record/10146176/files/figure.png" pageId="15" pageNumber="166">Fig. 12</figureCitation>
|
||
). Spermatheca not recognized.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33E5DF35435CC4FFF56FF4EFD60FE05" pageId="16" pageNumber="167" type="diagnosis">
|
||
<paragraph id="8B9B0E785435CC4FFF56FF4EFD60FE05" blockId="16.[151,1436,150,570]" pageId="16" pageNumber="167">
|
||
<emphasis id="B950D26A5435CC4FFF56FF4EFE6BFF33" bold="true" box="[199,435,150,177]" pageId="16" pageNumber="167">Comparative notes.</emphasis>
|
||
Based on the general shape of the forebody and aedeagus,
|
||
<taxonomicName id="4C2475FB5435CC4FFBCEFF4FFB38FF33" authority="Shavrin, 2023" authorityName="Shavrin" authorityYear="2023" box="[1119,1248,151,177]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="167" phylum="Arthropoda" rank="species" species="bilobum" status="sp. n.">
|
||
<emphasis id="B950D26A5435CC4FFBCEFF4FFBABFF33" box="[1119,1139,151,177]" italics="true" pageId="16" pageNumber="167">O</emphasis>
|
||
.
|
||
<emphasis id="B950D26A5435CC4FFB15FF4FFB38FF33" box="[1156,1248,151,177]" italics="true" pageId="16" pageNumber="167">bilobum</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B950D26A5435CC4FFB78FF4FFAF2FF33" bold="true" box="[1257,1322,151,177]" pageId="16" pageNumber="167">
|
||
<taxonomicNameLabel id="A2636F115435CC4FFB78FF4FFAF2FF33" box="[1257,1322,151,177]" pageId="16" pageNumber="167" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is similar to
|
||
<taxonomicName id="4C2475FB5435CC4FFF27FF63FEB7FF56" authority="Shavrin, 2023" authorityName="Shavrin" authorityYear="2023" box="[182,367,186,213]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="167" phylum="Arthropoda" rank="species" species="amplissimum" status="sp. n.">
|
||
<emphasis id="B950D26A5435CC4FFF27FF63FF12FF57" box="[182,202,187,213]" italics="true" pageId="16" pageNumber="167">O</emphasis>
|
||
.
|
||
<emphasis id="B950D26A5435CC4FFF4AFF62FEB7FF56" box="[219,367,186,212]" italics="true" pageId="16" pageNumber="167">amplissimum</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B950D26A5435CC4FFEE9FF63FE62FF57" bold="true" box="[376,442,187,213]" pageId="16" pageNumber="167">
|
||
<taxonomicNameLabel id="A2636F115435CC4FFEE9FF63FE62FF57" box="[376,442,187,213]" pageId="16" pageNumber="167" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, from which it can be distinguished by narrower antennomeres 7–9, finer sculpture of the head and the pronotum, the shape of the pronotum widest slightly above middle, with apical angles slightly protruded anteriad, the slightly broader and shorter elytra, the shape of the apical margin of the median lobe with distinct wide apical excision, shorter and narrower parameres, slightly longer than apex of the median lobe, and the morphology of the internal sac.
|
||
<taxonomicName id="4C2475FB5435CC4FFE69FE91FD1AFEE1" authority="Shavrin, 2023" authorityName="Shavrin" authorityYear="2023" box="[504,706,329,355]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="167" phylum="Arthropoda" rank="species" species="bilobum" status="sp. nov.">
|
||
<emphasis id="B950D26A5435CC4FFE69FE91FD1AFEE1" box="[504,706,329,355]" italics="true" pageId="16" pageNumber="167">Omalium bilobum</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B950D26A5435CC4FFD59FE91FCDFFEE1" bold="true" box="[712,775,329,355]" pageId="16" pageNumber="167">
|
||
<taxonomicNameLabel id="A2636F115435CC4FFD59FE91FCDFFEE1" box="[712,775,329,355]" pageId="16" pageNumber="167" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
differs from all known species of the genus by the presence of the wide excision on apex of the median lobe.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33E5DF35435CC4FFF56FE49FA4DFE29" box="[199,1429,401,427]" pageId="16" pageNumber="167" type="distribution">
|
||
<paragraph id="8B9B0E785435CC4FFF56FE49FA4DFE29" blockId="16.[151,1436,150,570]" box="[199,1429,401,427]" pageId="16" pageNumber="167">
|
||
<emphasis id="B950D26A5435CC4FFF56FE49FEB9FE29" bold="true" box="[199,353,401,427]" pageId="16" pageNumber="167">Distribution.</emphasis>
|
||
<taxonomicName id="4C2475FB5435CC4FFEF6FE49FDEBFE29" authority="Shavrin, 2023" authorityName="Shavrin" authorityYear="2023" box="[359,563,401,427]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="167" phylum="Arthropoda" rank="species" species="bilobum" status="sp. nov.">
|
||
<emphasis id="B950D26A5435CC4FFEF6FE49FDEBFE29" box="[359,563,401,427]" italics="true" pageId="16" pageNumber="167">Omalium bilobum</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B950D26A5435CC4FFDA8FE49FDA1FE29" bold="true" box="[569,633,401,427]" pageId="16" pageNumber="167">
|
||
<taxonomicNameLabel id="A2636F115435CC4FFDA8FE49FDA1FE29" box="[569,633,401,427]" pageId="16" pageNumber="167" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is known only from the
|
||
<typeStatus id="549FB0DA5435CC4FFC1DFE49FC64FE29" box="[908,956,401,427]" pageId="16" pageNumber="167">type</typeStatus>
|
||
locality in eastern part of
|
||
<collectingCountry id="F3334EE85435CC4FFB73FE49FAFDFE29" box="[1250,1317,401,427]" name="Nepal" pageId="16" pageNumber="167">Nepal</collectingCountry>
|
||
(
|
||
<figureCitation id="131F12FD5435CC4FFAA5FE49FA5FFE29" box="[1332,1415,401,427]" captionStart="FIGURE 33" captionStartId="15.[151,250,1334,1358]" captionTargetBox="[151,1436,417,1310]" captionTargetId="figure-126@15.[151,1436,417,1310]" captionTargetPageId="15" captionText="FIGURE 33. Distribution of Omalium in the Himalayan region: O. bilobum sp. n. (circle), O. marginatum (squares), O. altivagans sp. n. (triangle), O. humerale (diamond)." figureDoi="http://doi.org/10.5281/zenodo.10146190" httpUri="https://zenodo.org/record/10146190/files/figure.png" pageId="16" pageNumber="167">Fig. 33</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33E5DF35435CC4FFF56FE6DFC8AFE70" pageId="16" pageNumber="167" type="etymology">
|
||
<paragraph id="8B9B0E785435CC4FFF56FE6DFC8AFE70" blockId="16.[151,1436,150,570]" pageId="16" pageNumber="167">
|
||
<emphasis id="B950D26A5435CC4FFF56FE6DFE96FE4D" bold="true" box="[199,334,437,463]" pageId="16" pageNumber="167">Etymology.</emphasis>
|
||
The specific epithet is the Latin adjective
|
||
<emphasis id="B950D26A5435CC4FFCBBFE6CFCA4FE4C" box="[810,892,436,462]" italics="true" pageId="16" pageNumber="167">bilobus</emphasis>
|
||
, -
|
||
<emphasis id="B950D26A5435CC4FFC02FE6DFC79FE4C" box="[915,929,437,462]" italics="true" pageId="16" pageNumber="167">a</emphasis>
|
||
, -
|
||
<emphasis id="B950D26A5435CC4FFC28FE6DFC03FE4C" box="[953,987,437,462]" italics="true" pageId="16" pageNumber="167">um</emphasis>
|
||
(with two lobes). It alludes to the shape of th apical portion of the median lobe diveded into two lobes.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33E5DF35435CC4FFF56FE24FF14FDB8" pageId="16" pageNumber="167" type="biology_ecology">
|
||
<paragraph id="8B9B0E785435CC4FFF56FE24FF14FDB8" blockId="16.[151,1436,150,570]" pageId="16" pageNumber="167">
|
||
<emphasis id="B950D26A5435CC4FFF56FE24FE93FD94" bold="true" box="[199,331,508,534]" pageId="16" pageNumber="167">Bionomics.</emphasis>
|
||
Specimens were taken by sifting rotten branches and moss below a dead tree at elevation
|
||
<quantity id="4CDCA39D5435CC4FFAD6FE24FA44FD95" box="[1351,1436,508,535]" metricMagnitude="3" metricUnit="m" metricValue="2.25" pageId="16" pageNumber="167" unit="m" value="2250.0">2250 m</quantity>
|
||
a.s.l.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |