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<document id="4D466A96B1B90F464C991DDD39E64A2A" ID-DOI="10.5281/zenodo.157951" ID-GBIF-Dataset="5336457f-a8a5-4110-8153-fac75a584ba1" ID-ISSN="1175­5326" ID-Zenodo-Dep="157951" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461016276046" checkinUser="plazi" docAuthor="Heraty, John" docDate="2004" docId="D54A87F0FF857645FEB3F97B4C95BBE3" docLanguage="en" docName="zt00497.pdf" docOrigin="Zootaxa 497" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Gollumiella ochreata Heraty, 2004, new species" docType="treatment" docVersion="9" lastPageNumber="8" masterDocId="2973FF88FF807642FFBBFFB94812BA65" masterDocTitle="Three new species of Gollumiella Hedqvist (Hymenoptera: Eucharitidae)" masterLastPageNumber="10" masterPageNumber="1" pageNumber="6" updateTime="1698568833848" updateUser="plazi">
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<mods:title id="8578DE6ACDD2B1D62AC5A1D2FF8EA97D">Three new species of Gollumiella Hedqvist (Hymenoptera: Eucharitidae)</mods:title>
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<mods:date id="CF1A300233230119F478007D6F9E4653">2004</mods:date>
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<treatment id="D54A87F0FF857645FEB3F97B4C95BBE3" ID-DOI="http://doi.org/10.5281/zenodo.6269421" ID-GBIF-Taxon="119601622" ID-Zenodo-Dep="6269421" LSID="urn:lsid:plazi:treatment:D54A87F0FF857645FEB3F97B4C95BBE3" httpUri="http://treatment.plazi.org/id/D54A87F0FF857645FEB3F97B4C95BBE3" lastPageId="7" lastPageNumber="8" pageId="5" pageNumber="6">
<subSubSection id="15F9656DFF857647FEB3F97B4B13BCBE" box="[264,769,1729,1756]" pageId="5" pageNumber="6" type="nomenclature">
<paragraph id="5D5C36E6FF857647FEB3F97B4B13BCBE" blockId="5.[264,769,1729,1756]" box="[264,769,1729,1756]" pageId="5" pageNumber="6">
<heading id="0614818AFF857647FEB3F97B4B13BCBE" bold="true" box="[264,769,1729,1756]" fontSize="11" level="1" pageId="5" pageNumber="6" reason="1">
<emphasis id="6F97EAF4FF857647FEB3F97B4B13BCBE" bold="true" box="[264,769,1729,1756]" pageId="5" pageNumber="6">
<taxonomicName id="9AE34D65FF857647FEB3F97B4A1ABCB9" ID-CoL="3GRVZ" box="[264,520,1730,1756]" class="Insecta" family="Eucharitidae" genus="Gollumiella" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="ochreata" status="sp. nov.">
<emphasis id="6F97EAF4FF857647FEB3F97B4A1ABCB9" bold="true" box="[264,520,1730,1756]" italics="true" pageId="5" pageNumber="6">Gollumiella ochreata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="74A4578FFF857647FDB4F9784A8DBCBE" box="[527,671,1729,1755]" pageId="5" pageNumber="6" rank="species">new species</taxonomicNameLabel>
(
<figureCitation id="C5D82A63FF857647FD14F9784AEABCBE" box="[687,760,1729,1755]" captionStart="FIGURES 1 2. 1" captionStartId="2.[264,383,1546,1570]" captionTargetBox="[274,1309,1001,1508]" captionTargetId="figure@2.[264,1323,997,1519]" captionTargetPageId="2" captionText="FIGURES 1 2. 1, Gollumiella buffingtoni, habitus of holotype female, inset is dorsolateral view of petiole; arrow points to pale apical coloration of petiole. 2, Gollumiella ochreata, habitus of holotype male. Scale bar is 0.5 mm." httpUri="https://zenodo.org/record/168981/files/figure.png" pageId="5" pageNumber="6">Fig. 2</figureCitation>
)
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="15F9656DFF857644FEB3F8B54BE9B843" lastPageId="6" lastPageNumber="7" pageId="5" pageNumber="6" type="diagnosis">
<paragraph id="5D5C36E6FF857644FEB3F8B54BE9B843" blockId="5.[264,1322,1804,1870]" lastBlockId="6.[264,1323,284,1870]" lastPageId="6" lastPageNumber="7" pageId="5" pageNumber="6">
<emphasis id="6F97EAF4FF857647FEB3F8B54992BD43" bold="true" box="[264,384,1804,1830]" pageId="5" pageNumber="6">Diagnosis</emphasis>
. This species keys to
<taxonomicName id="9AE34D65FF857647FD3AF8B44B57BD43" box="[641,837,1805,1830]" class="Insecta" family="Eucharitidae" genus="Gollumiella" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="longipetiolata">
<emphasis id="6F97EAF4FF857647FD3AF8B44B57BD43" box="[641,837,1805,1830]" italics="true" pageId="5" pageNumber="6">G. longipetiolata</emphasis>
</taxonomicName>
in
<bibRefCitation id="39724B17FF857647FCD1F8B54C01BD43" author="Heraty" box="[874,1043,1804,1830]" pageId="5" pageNumber="9" refString="Heraty, J. M. (1992) Revision of the genera Gollumiella Hedqvist and Anorasema Boucek (Hymenoptera: Eucharitidae). Invertebrate Taxonomy, 6, 583 - 604." type="journal article" year="1992">Heraty (1992)</bibRefCitation>
.
<taxonomicName id="9AE34D65FF857647FBA5F8B44CB7BD43" box="[1054,1189,1805,1830]" class="Insecta" family="Eucharitidae" genus="Gollumiella" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="ochreata">
<emphasis id="6F97EAF4FF857647FBA5F8B44CB7BD43" box="[1054,1189,1805,1830]" italics="true" pageId="5" pageNumber="6">G. ochreata</emphasis>
</taxonomicName>
can be distinguished from all other species by the combination of dark brown coxae, yellowish white legs, basally striate and completely yellow petiole, and an elongate scape exceeding the ventral margin of the median ocellus, but not the dorsal margin. As well, the fore wing is 2.362.38 X as long as broad, as compared to
<taxonomicName id="9AE34D65FF867644FC91FED54BFCBBE0" box="[810,1006,364,389]" class="Insecta" family="Eucharitidae" genus="Gollumiella" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="longipetiolata">
<emphasis id="6F97EAF4FF867644FC91FED54BFCBBE0" box="[810,1006,364,389]" italics="true" pageId="6" pageNumber="7">G. longipetiolata</emphasis>
</taxonomicName>
, in which it is greater than 2.4 X as long as broad, and more than 2.5 X in the same gardens where the
<typeStatus id="82588844FF867644FB3FFE2D4CF8BBCB" box="[1156,1258,404,430]" pageId="6" pageNumber="7" type="holotype">holotype</typeStatus>
of
<taxonomicName id="9AE34D65FF867644FAA9FE2D497DBBB0" class="Insecta" family="Eucharitidae" genus="Gollumiella" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="ochreata">
<emphasis id="6F97EAF4FF867644FAA9FE2D497DBBB0" italics="true" pageId="6" pageNumber="7">G. ochreata</emphasis>
</taxonomicName>
was collected.
<taxonomicName id="9AE34D65FF867644FD93FE054AACBBB0" box="[552,702,444,469]" class="Insecta" family="Eucharitidae" genus="Gollumiella" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="antennata">
<emphasis id="6F97EAF4FF867644FD93FE054AACBBB0" box="[552,702,444,469]" italics="true" pageId="6" pageNumber="7">G. antennata</emphasis>
</taxonomicName>
also has a yellow petiole, but differs by having yellow coxae, smooth and curved petiole, nearly circular head in frontal view, and a short scape that reaches only 0.8 X the distance to the median ocellus.
</paragraph>
</subSubSection>
<subSubSection id="15F9656DFF867644FE83FD8A4A0EBC6B" pageId="6" pageNumber="7" type="description">
<paragraph id="5D5C36E6FF867644FE83FD8A49AAB8FB" blockId="6.[264,1323,284,1870]" pageId="6" pageNumber="7">
<emphasis id="6F97EAF4FF867644FE83FD8A496CB828" bold="true" box="[312,382,563,589]" pageId="6" pageNumber="7">Male.</emphasis>
Length
<quantity id="9A1B9B03FF867644FE52FD8D4A49B82B" box="[489,603,564,590]" metricMagnitude="-3" metricUnit="m" metricValue="2.17" pageId="6" pageNumber="7" unit="mm" value="2.17">2.17 mm</quantity>
. Dark brown; antennae, legs excluding coxae, petiole, and mandibles except for brown outer edge, pale yellow (
<figureCitation id="C5D82A63FF867644FCC1FDE54BD0B813" box="[890,962,604,630]" captionStart="FIGURES 1 2. 1" captionStartId="2.[264,383,1546,1570]" captionTargetBox="[274,1309,1001,1508]" captionTargetId="figure@2.[264,1323,997,1519]" captionTargetPageId="2" captionText="FIGURES 1 2. 1, Gollumiella buffingtoni, habitus of holotype female, inset is dorsolateral view of petiole; arrow points to pale apical coloration of petiole. 2, Gollumiella ochreata, habitus of holotype male. Scale bar is 0.5 mm." httpUri="https://zenodo.org/record/168981/files/figure.png" pageId="6" pageNumber="7">Fig. 2</figureCitation>
); distal tarsomere dark brown. Wings hyaline.
</paragraph>
<paragraph id="5D5C36E6FF867644FE83FD154AF6BE4B" blockId="6.[264,1323,284,1870]" pageId="6" pageNumber="7">Head subtriangular, 1.23 X as broad as high. POL 0.750.85 X OOL. Face rounded and smooth with scattered fine setae, genae slightly convex in frontal view; occiput glabrous, dorsal angle with occiput abrupt and without sculpture. Clypeal margin truncate and with scattered fine setae. Eyes separated by 1.51.6 X their height. Malar space 0.70 X height of eye. Labrum with 4 long digits. Antenna 11 segmented; scape exceeding ventral margin of median ocellus, but length reaching only 0.880.96 X distance between torulus and upper margin of median ocellus, scape slightly expanded medioventrally; 8 funicular segments, FL1 and FL2 separated, combined length 2.32.6 X as long as broad, FL1 0.6 0.8 X as long as FL2; flagellum 0.87 X height of head, setae of flagellum long, dense and semi­erect; clava tapering to acute point.</paragraph>
<paragraph id="5D5C36E6FF867644FE83FB854A71BFF3" blockId="6.[264,1323,284,1870]" pageId="6" pageNumber="7">
Mesosoma with rugose­alveolate sculpture, scutoscutellar sulcus deeply impressed and strongly transverse­carinate; scutellum as long as broad, frenal carina prominent and bisinuate. Propleuron with 45 transverse carinae. Coxae smooth and semi­globose, hind coxa 1.92.0 X as long as broad; hind femur bare laterally, tibia sparsely setose. Fore wing 2.362.38 X as long as broad, 2.883.03 X length of mesothorax; speculum closed basally by broad band of dense setae, basal area mostly pilose; venation distinct (
<figureCitation id="C5D82A63FF867644FBD2FABD4CA3BF7B" box="[1129,1201,1284,1310]" captionStart="FIGURES 1 2. 1" captionStartId="2.[264,383,1546,1570]" captionTargetBox="[274,1309,1001,1508]" captionTargetId="figure@2.[264,1323,997,1519]" captionTargetPageId="2" captionText="FIGURES 1 2. 1, Gollumiella buffingtoni, habitus of holotype female, inset is dorsolateral view of petiole; arrow points to pale apical coloration of petiole. 2, Gollumiella ochreata, habitus of holotype male. Scale bar is 0.5 mm." httpUri="https://zenodo.org/record/168981/files/figure.png" pageId="6" pageNumber="7">Fig. 2</figureCitation>
); submarginal vein dorsally with few minute setae basally, but otherwise bare; stigmal vein 2.02.5 X as long as broad and equal in width to apical group of sensilla; postmarginal vein 3.6 4.0 X as long as stigmal vein.
</paragraph>
<paragraph id="5D5C36E6FF867644FE83FA1D4C2DBF83" blockId="6.[264,1323,284,1870]" pageId="6" pageNumber="7">Petiole 2.12.4 X as long as hind coxa, straight in profile, irregularly longitudinally striate but with distal fifth mostly smooth. Aedeagus broadly rounded.</paragraph>
<paragraph id="5D5C36E6FF867644FE83FA4D4A0EBC6B" blockId="6.[264,1323,284,1870]" box="[312,540,1524,1550]" pageId="6" pageNumber="7">
<emphasis id="6F97EAF4FF867644FE83FA4D4981BC6B" bold="true" box="[312,403,1524,1550]" pageId="6" pageNumber="7">Female</emphasis>
. Unknown.
</paragraph>
</subSubSection>
<subSubSection id="15F9656DFF867644FE83F9A54ADFBC53" box="[312,717,1564,1590]" pageId="6" pageNumber="7" type="distribution">
<paragraph id="5D5C36E6FF867644FE83F9A54ADFBC53" blockId="6.[264,1323,284,1870]" box="[312,717,1564,1590]" pageId="6" pageNumber="7">
<emphasis id="6F97EAF4FF867644FE83F9A549CABC53" bold="true" box="[312,472,1564,1590]" pageId="6" pageNumber="7">Distribution.</emphasis>
<collectingCountry id="25F47676FF867644FE5CF9A54A40BC53" box="[487,594,1564,1590]" name="Malaysia" pageId="6" pageNumber="7">Malaysia</collectingCountry>
(
<collectingCountry id="25F47676FF867644FDD9F9A54AD3BC53" box="[610,705,1564,1590]" name="Malaysia" pageId="6" pageNumber="7">Malaya</collectingCountry>
).
</paragraph>
</subSubSection>
<subSubSection id="15F9656DFF867645FE83F9FD4B73BB3B" lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7" type="materials_examined">
<paragraph id="5D5C36E6FF867644FE83F9FD4A2BBCB3" blockId="6.[264,1323,284,1870]" pageId="6" pageNumber="7">
<emphasis id="6F97EAF4FF867644FE83F9FD49FCBC3B" bold="true" box="[312,494,1604,1630]" pageId="6" pageNumber="7">
<typeStatus id="82588844FF867644FE83F9FD4961BC3B" box="[312,371,1604,1630]" pageId="6" pageNumber="7">Type</typeStatus>
Material.
</emphasis>
<typeStatus id="82588844FF867644FE43F9FD4A7ABC3B" box="[504,616,1604,1630]" pageId="6" pageNumber="7" type="holotype">Holotype</typeStatus>
, ɗ, “
<collectingCountry id="25F47676FF867644FD1BF9FD4B01BC3B" box="[672,787,1604,1630]" name="Malaysia" pageId="6" pageNumber="7">Malaysia</collectingCountry>
: Selangor, K.L., Univ.
<collectingCountry id="25F47676FF867644FBA4F9FD4C68BC3B" box="[1055,1146,1604,1630]" name="Malaysia" pageId="6" pageNumber="7">Malaya</collectingCountry>
, Rimba Ilmu
<geoCoordinate id="38D75021FF867644FEB3F9D5498ABCE3" box="[264,408,1644,1670]" direction="north" orientation="latitude" pageId="6" pageNumber="7" precision="15" value="3.128611">03°0743”N</geoCoordinate>
<geoCoordinate id="38D75021FF867644FE19F9D54A2DBCE3" box="[418,575,1644,1670]" direction="east" orientation="longitude" pageId="6" pageNumber="7" precision="15" value="101.65806">101°3929”E</geoCoordinate>
,
<date id="295D1026FF867644FDF5F9D54AD9BCE3" box="[590,715,1644,1670]" pageId="6" pageNumber="7" value="2001-09-17">17.ix.2001</date>
J. Heraty,
<quantity id="9A1B9B03FF867644FCEAF9D54B9ABCE3" box="[849,904,1644,1670]" metricMagnitude="1" metricUnit="m" metricValue="7.5" pageId="6" pageNumber="7" unit="m" value="75.0">75m</quantity>
, sweep bot. gard. H01­045 /
<collectionCode id="3BF2AE23FF867644FB50F9D54D39BCE3" LSID="urn:lsid:biocol.org:col:15536" box="[1259,1323,1644,1670]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15536" name="Department of Natural Resources, Environment, The Arts and Sport" pageId="6" pageNumber="7">DNA</collectionCode>
voucher D# 426,
<collectionCode id="3BF2AE23FF867644FE69F92D4A00BCCB" box="[466,530,1684,1710]" pageId="6" pageNumber="7">UCR</collectionCode>
, J.M. Heraty”, deposited in
<collectionCode id="3BF2AE23FF867644FCECF92D4BA2BCCB" LSID="urn:lsid:biocol.org:col:34871" box="[855,944,1684,1710]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="6" pageNumber="7">USNM</collectionCode>
.
<emphasis id="6F97EAF4FF867644FC01F92D4C37BCCB" bold="true" box="[954,1061,1684,1710]" italics="true" pageId="6" pageNumber="7">
<typeStatus id="82588844FF867644FC01F92D4C37BCCB" box="[954,1061,1684,1710]" pageId="6" pageNumber="7" type="paratype">Paratype</typeStatus>
</emphasis>
: from same collection as
<typeStatus id="82588844FF867644FE93F905499CBCB3" box="[296,398,1724,1750]" pageId="6" pageNumber="7" type="holotype">holotype</typeStatus>
(1 ɗ,
<collectionCode id="3BF2AE23FF867644FE63F9054A3FBCB3" box="[472,557,1724,1750]" httpUri="http://grbio.org/cool/m2jc-wmeq" name="University of California, Riverside" pageId="6" pageNumber="7">UCRC</collectionCode>
);
</paragraph>
<paragraph id="5D5C36E6FF867645FE83F95D4B73BB3B" blockId="6.[264,1323,284,1870]" lastBlockId="7.[264,1320,284,390]" lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7">
<emphasis id="6F97EAF4FF867644FE83F95D4A4ABC9B" bold="true" box="[312,600,1764,1790]" pageId="6" pageNumber="7">Molecular information</emphasis>
. Three gene regions were sequenced for 1 individual. Data for the ribosomal transcript regions 28S­D2&amp;D3 and 18S­E23 are deposited on GenBank as assession numbers
<accessionNumber id="42B0AB05FF867644FDB0F88D4A98BD2B" box="[523,650,1844,1870]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/AY552269" pageId="6" pageNumber="7">AY552269</accessionNumber>
and
<accessionNumber id="42B0AB05FF867644FD7EF88D4B5ABD2B" box="[709,840,1844,1870]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/AY552192" pageId="6" pageNumber="7">AY552192</accessionNumber>
. Differs from
<taxonomicName id="9AE34D65FF867644FC4BF88D4CA7BD28" box="[1008,1205,1844,1869]" class="Insecta" family="Eucharitidae" genus="Gollumiella" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="longipetiolata">
<emphasis id="6F97EAF4FF867644FC4BF88D4CA7BD28" box="[1008,1205,1844,1869]" italics="true" pageId="6" pageNumber="7">G. longipetiolata</emphasis>
</taxonomicName>
that were collected sympatrically by 3 nucleotide differences (28S­D2:2; 28S­D3:1), and from
<taxonomicName id="9AE34D65FF877645FAA9FEA5499ABB38" class="Insecta" family="Eucharitidae" genus="Gollumiella" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="buffingtoni">
<emphasis id="6F97EAF4FF877645FAA9FEA5499ABB38" italics="true" pageId="7" pageNumber="8">G. buffingtoni</emphasis>
</taxonomicName>
by 9 differences (28S­D2:7; 28S­D3:2).
</paragraph>
</subSubSection>
<subSubSection id="15F9656DFF877645FE83FED24C95BBE3" box="[312,1159,363,390]" pageId="7" pageNumber="8" type="etymology">
<paragraph id="5D5C36E6FF877645FE83FED24C95BBE3" blockId="7.[264,1320,284,390]" box="[312,1159,363,390]" pageId="7" pageNumber="8">
<emphasis id="6F97EAF4FF877645FE83FED249D6BBE0" bold="true" box="[312,452,363,389]" pageId="7" pageNumber="8">Etymology.</emphasis>
From the Greek
<emphasis id="6F97EAF4FF877645FD37FED54AC9BBE0" box="[652,731,364,389]" italics="true" pageId="7" pageNumber="8">ochros</emphasis>
, referring to the pale yellow petiole.
</paragraph>
</subSubSection>
</treatment>
</document>