205 lines
32 KiB
XML
205 lines
32 KiB
XML
<document id="C6C8E4487282D9DB2997410B7D8D8888" ID-DOI="10.5281/zenodo.8108977" ID-ISSN="1536-9307" ID-Zenodo-Dep="8108977" ID-ZooBank="87F75781-5547-4D0F-8DA3-8B64E41B3879" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1688389712455" checkinUser="felipe" docAuthor="Kawai, Kazusa, Unnahachote, Thornthan, Suttisatid, Yossaphon & Tang, Victoria" docDate="2023" docId="D23287FA6B70FFCDBE94FCFEFD73E84F" docLanguage="en" docName="Euscorpius.373.1-25.pdf" docOrigin="Euscorpius 373" docStyle="DocumentStyle:5E96F6379CE20B0E8E4F9E7DF8E1C0DF.2:Euscorpius.2020-.journal_article" docStyleId="5E96F6379CE20B0E8E4F9E7DF8E1C0DF" docStyleName="Euscorpius.2020-.journal_article" docStyleVersion="2" docTitle="Heterometrus laoticus Couzijn 1981" docType="treatment" docVersion="3" lastPageNumber="15" masterDocId="2E0BFF826B60FFDDBE49FF9FFFFCE054" masterDocTitle="А Review of Heterometrus in Thailand (Scorpiones: Scorpionidae)" masterLastPageNumber="25" masterPageNumber="1" pageNumber="15" updateTime="1699531369301" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:title id="C4349C2C7966C5CACC7380C5C5D44406">А Review of Heterometrus in Thailand (Scorpiones: Scorpionidae)</mods:title>
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<mods:namePart id="AB5B856BEBDE2ECF874023266D226D0E">Kawai, Kazusa</mods:namePart>
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<mods:namePart id="601AF7526AB42BD48432CC051331D21F">Unnahachote, Thornthan</mods:namePart>
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<mods:namePart id="6A7A5FB91E44540AC52DDA559D1F6D66">Suttisatid, Yossaphon</mods:namePart>
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<mods:namePart id="8EC3375DF8DA33CAF00E37BD8BDC9DE9">Tang, Victoria</mods:namePart>
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<mods:title id="2843F27E8444FFBB539478D1D0DEA34A">Euscorpius</mods:title>
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<mods:part id="8BF1105C30789666343362BD1B96B808">
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<mods:date id="19F8C2081B5A4462D324143DE462D50B">2023</mods:date>
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<mods:number id="0D0D4673E06124FF053CA24CE3CFC697">373</mods:number>
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<mods:identifier id="2EC1346A87504A2BA603D52A7D2DFD29" type="DOI">10.5281/zenodo.8108977</mods:identifier>
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<heading id="016C81806B70FFCDBE94FCFEFD6BE32F" box="[221,663,864,891]" centered="true" fontSize="11" level="2" pageId="16" pageNumber="15" reason="2">
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<taxonomicName id="9D9B4D6F6B70FFCDBE94FCFEFD6BE32F" ID-CoL="6M4WW" ID-ENA="217256" authority="Couzijn, 1981" authorityName="Couzijn" authorityYear="1981" box="[221,663,864,891]" class="Arachnida" family="Scorpionidae" genus="Heterometrus" kingdom="Animalia" order="Scorpiones" pageId="16" pageNumber="15" phylum="Arthropoda" rank="species" species="laoticus">
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<emphasis id="68EFEAFE6B70FFCDBE94FCFEFE16E32F" bold="true" box="[221,490,864,891]" italics="true" pageId="16" pageNumber="15">Heterometrus laoticus</emphasis>
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<bibRefCitation id="3E0A4B1D6B70FFCDBFB8FCFEFD6BE32F" author="COUZIJN" box="[497,663,865,891]" pageId="16" pageNumber="15" pagination="1 - 196" refId="ref11657" refString="COUZIJN, H. W. C. 1981. Revision of the genus Heterometrus. Zoologische Verhandelingen, 184: 1 - 196." type="journal article" year="1981">Couzijn, 1981</bibRefCitation>
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</taxonomicName>
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<subSubSection id="128165676B70FFCDBE3CFC1CFD78E38E" pageId="16" pageNumber="15" type="description">
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<paragraph id="5A2436EC6B70FFCDBE3CFC1CFCFFE3EE" blockId="16.[113,771,864,986]" pageId="16" pageNumber="15">
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(
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<figureCitation id="C2A02A696B70FFCDBE35FC1CFEE5E3CF" box="[124,281,899,923]" captionStart="Figures 4–11" captionStartId="5.[116,193,1930,1952]" captionTargetBox="[133,1447,263,1857]" captionTargetId="figure-82@5.[113,1474,234,1905]" captionTargetPageId="5" captionText="Figures 4–11: Comparison of Heterometrus spp. in Thailand, metasoma and telson in dorsal aspect. Figures 4–5. H. cimrmani, male from Trang (4) and female from Nakhon Si Thammarat (5), THNHM. Figures 6–7. H. minotaurus, male (6) and female from Surat Thani (7), THNHM. Figures 8–9. H. silenus, male from Chanthaburi (8) and female from Chonburi (9), TUPC. Figures 10–11. H. laoticus, male (10) and female from Lamphun (11), TUPC." figureDoi="http://doi.org/10.5281/zenodo.8108984" httpUri="https://zenodo.org/record/8108984/files/figure.png" pageId="16" pageNumber="15">Figures 10–11</figureCitation>
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,
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<figureCitation id="C2A02A696B70FFCDBF6DFC1CFE95E3CF" box="[292,361,899,923]" captionStart="Figures 12–19" captionStartId="6.[115,192,1927,1949]" captionTargetBox="[124,1469,261,1873]" captionTargetId="figure-87@6.[113,1474,234,1915]" captionTargetPageId="6" captionText="Figures 12–19: Comparison of Heterometrus spp. in Thailand, metasoma and telson in ventral aspect. Figures 12–13. H. cimrmani, male from Trang (12) and female from Nakhon Si Thammarat (13), THNHM. Figures 14–15. H. minotaurus, male (14) and female from Surat Thani (15), THNHM. Figures 16–17. H. silenus, male from Chanthaburi (16) and female from Chonburi (17), TUPC. Figures 18–19. H. laoticus, male (18) and female from Lamphun (19), TUPC." figureDoi="http://doi.org/10.5281/zenodo.8108986" httpUri="https://zenodo.org/record/8108986/files/figure.png" pageId="16" pageNumber="15">18–19</figureCitation>
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,
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<figureCitation id="C2A02A696B70FFCDBF3DFC1DFE45E3CE" box="[372,441,898,922]" captionStart="Figures 20–29" captionStartId="7.[115,192,993,1015]" captionTargetBox="[118,1460,249,936]" captionTargetId="figure-433@7.[113,1474,234,982]" captionTargetPageId="7" captionText="Figures 20–29: Comparison of Heterometrus spp. in Thailand, prosoma in dorsal aspect. Figures 20–21. H. cimrmani, male from Trang (20) and female from Nakhon Si Thammarat (21), THNHM. Figures 22–23. H. minotaurus, male (22) and female from Surat Thani (23), THNHM. Figures 24–25. H. silenus, male from Chanthaburi (24) and female from Chonburi (25), TUPC. Figures 26–27. H. laoticus, male (26) and female from Lamphun (27), TUPC. Figures 28–29. H. spinifer, males from Yala (28) and Narathiwat (29), TUPC." figureDoi="http://doi.org/10.5281/zenodo.8108988" httpUri="https://zenodo.org/record/8108988/files/figure.png" pageId="16" pageNumber="15">26–27</figureCitation>
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,
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<figureCitation id="C2A02A696B70FFCDBF8DFC1DFDF5E3CE" box="[452,521,898,922]" captionStart="Figures 30–39" captionStartId="8.[113,190,1686,1708]" captionTargetBox="[147,1464,246,1661]" captionTargetId="figure-206@8.[115,1476,234,1687]" captionTargetPageId="8" captionText="Figures 30–39: Comparison of Heterometrus spp. in Thailand, chela in dorsal aspect (trichobothria indicated by red circles). Figures 30–31. H. cimrmani, male from Trang (30) and female from Nakhon Si Thammarat (31), THNHM. Figures 32–33. H. minotaurus, male (32) and female from Surat Thani (33), THNHM. Figures 34–35. H. silenus, male from Chanthaburi (34) and female from Chonburi (35), TUPC. Figures 36–37. H. laoticus, male (36) and female from Lamphun (37), TUPC. Figures 28–29. H. spinifer, males from Yala (38) and Narathiwat (39), TUPC." figureDoi="http://doi.org/10.5281/zenodo.8108990" httpUri="https://zenodo.org/record/8108990/files/figure.png" pageId="16" pageNumber="15">36–37</figureCitation>
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,
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<figureCitation id="C2A02A696B70FFCDBC5DFC1DFDA5E3CE" box="[532,601,898,922]" captionStart="Figures 40–49" captionStartId="9.[113,190,1519,1541]" captionTargetBox="[119,1440,244,1498]" captionTargetId="figure-248@9.[113,1474,234,1522]" captionTargetPageId="9" captionText="Figures 40–49: Comparison of Heterometrus spp. in Thailand, chela in external aspect (trichobothria indicated by red circles). Figures 40–41. H. cimrmani, male from Trang (40) and female from Nakhon Si Thammarat (41), THNHM. Figures 42–43. H. minotaurus, male (42) and female from Surat Thani (43), THNHM. Figures 44–45. H. silenus, male from Chanthaburi (44) and female from Chonburi (45), TUPC. Figures 46–47. H. laoticus, male (46) and female from Lamphun (47), TUPC. Figures 48–49. H. spinifer, males from Yala (48) and Narathiwat (49), TUPC." figureDoi="http://doi.org/10.5281/zenodo.8108992" httpUri="https://zenodo.org/record/8108992/files/figure.png" pageId="16" pageNumber="15">46–47</figureCitation>
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,
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<figureCitation id="C2A02A696B70FFCDBC2DFC1DFD55E3CE" box="[612,681,898,922]" captionStart="Figures 50–57" captionStartId="10.[115,192,1986,2008]" captionTargetBox="[120,1462,220,1967]" captionTargetId="figure-90@10.[113,1474,208,1985]" captionTargetPageId="10" captionText="Figures 50–57: Comparison of Heterometrus spp. in Thailand, chela in ventral aspect (trichobothria indicated by red circles). Figures 50–51. H. cimrmani, male from Trang (50) and female from Nakhon Si Thammarat (51), THNHM. Figures 52–53. H. minotaurus, male (52) and female from Surat Thani (53), THNHM. Figures 54–55. H. silenus, male from Chanthaburi (54) and female from Chonburi (55), TUPC. Figures 56–57. H. laoticus, male (56) and female from Lamphun (57), TUPC." figureDoi="http://doi.org/10.5281/zenodo.8108994" httpUri="https://zenodo.org/record/8108994/files/figure.png" pageId="16" pageNumber="15">56–57</figureCitation>
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,
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<figureCitation id="C2A02A696B70FFCDBCFDFC1DFD04E3CE" box="[692,760,898,922]" captionStart="Figures 72–75" captionStartId="18.[113,190,2026,2048]" captionTargetBox="[135,1451,229,2009]" captionTargetId="figure-46@18.[116,1477,217,2014]" captionTargetPageId="18" captionText="Figures 72–75: Heterometrus laoticus, in dorsal (72, 74) and ventral (73, 75) aspects; scale bar = 10 mm. Figures 72–73. Male from Lamphun, TUPC. Figures 74–75. Female from Lamphun, TUPC." figureDoi="http://doi.org/10.5281/zenodo.8109008" httpUri="https://zenodo.org/record/8109008/files/figure.png" pageId="16" pageNumber="15">72–75</figureCitation>
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)
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<emphasis id="68EFEAFE6B70FFCDBE38FC3DFCFFE3EE" bold="true" box="[113,771,930,954]" pageId="16" pageNumber="15">http://zoobank.org/urn:lsid:zoobank.org:act:36ECBBCC-</emphasis>
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</paragraph>
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<emphasis id="68EFEAFE6B70FFCDBEB8FC5DFD78E38E" bold="true" box="[241,644,962,986]" pageId="16" pageNumber="15">A638-4D5E-AA78-6B9AAC2F626F</emphasis>
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</paragraph>
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</subSubSection>
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<subSubSection id="128165676B70FFCDBE38FB9CFEBAE42F" pageId="16" pageNumber="15" type="reference_group">
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<paragraph id="5A2436EC6B70FFCDBE38FB9CFEBAE42F" blockId="16.[113,771,1027,1147]" pageId="16" pageNumber="15">
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<treatmentCitation id="DB3A10FD6B70FFCDBE38FB9CFDCFE44F" author="COUZIJN" box="[113,563,1027,1051]" page="94" pageId="16" pageNumber="15" year="1981">
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<taxonomicName id="9D9B4D6F6B70FFCDBE38FB9CFDCFE44F" ID-CoL="6M4WW" authority="Couzijn, 1981: 94" authorityName="Couzijn" authorityPageNumber="94" authorityYear="1981" box="[113,563,1027,1051]" class="Arachnida" family="Scorpionidae" genus="Heterometrus" kingdom="Animalia" order="Scorpiones" pageId="16" pageNumber="15" phylum="Arthropoda" rank="species" species="laoticus">
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<emphasis id="68EFEAFE6B70FFCDBE38FB9CFE9DE44F" box="[113,353,1027,1051]" italics="true" pageId="16" pageNumber="15">Heterometrus laoticus</emphasis>
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<bibRefCitation id="3E0A4B1D6B70FFCDBF25FB9CFDCFE44F" author="COUZIJN" box="[364,563,1027,1051]" pageId="16" pageNumber="15" pagination="1 - 196" refId="ref11657" refString="COUZIJN, H. W. C. 1981. Revision of the genus Heterometrus. Zoologische Verhandelingen, 184: 1 - 196." type="journal article" year="1981">Couzijn, 1981: 94</bibRefCitation>
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(part);
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<treatmentCitation id="DB3A10FD6B70FFCDBCC3FB9CFE73E46E" author="PRENDINI" httpUri="http://treatment.plazi.org/id/5E6CB374FF056DFDFF1F656BFB55DDB9" page="245" pageId="16" pageNumber="15" year="2020">
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<bibRefCitation id="3E0A4B1D6B70FFCDBCC3FB9CFE73E46E" author="PRENDINI" pageId="16" pageNumber="15" refId="ref12183" refString="PRENDINI, L. & S. F. LORIA. 2020. Systematic revision of the Asian forest scorpions (Heterometrinae Simon, 1879), revised suprageneric classification of Scorpionidae Latreille, 1802, and revalidation of Rugodentidae Bastawade et al., 2005. Bulletin of the American Museum of Natural History, 442, 480 pp." type="book" year="2020">Prendini & Loria, 2020: 245–252</bibRefCitation>
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</treatmentCitation>
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, figs. 7F, 8F, 10, 24A, B, 38A, B, 51A–D, 70A, 71A, 72A, 157, 169–173, table 2 (reference list until 2020).
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</treatmentCitationGroup>
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<materialsCitation id="EAF33CB16B70FFCDBE38FB3CFEBBE4EF" box="[113,327,1187,1211]" collectionCode="LOCALITY, AND" pageId="16" pageNumber="15" specimenCount="1" typeStatus="holotype">
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<typeStatus id="8520884E6B70FFCDBE38FB3CFF5BE4EE" box="[113,167,1187,1210]" pageId="16" pageNumber="15">TYPE</typeStatus>
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<collectionCode id="3C8AAE296B70FFCDBEE7FB3CFEBBE4EF" box="[174,327,1187,1211]" pageId="16" pageNumber="15">LOCALITY AND</collectionCode>
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<materialsCitation id="EAF33CB16B70FFCDBF04FB3CFD7CE4EE" box="[333,640,1187,1211]" collectionCode="DEPOSITORY" country="Laos" location="Neis" pageId="16" pageNumber="15" specimenCount="1" typeStatus="holotype">
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<typeStatus id="8520884E6B70FFCDBF04FB3CFE83E4EE" box="[333,383,1187,1210]" pageId="16" pageNumber="15">TYPE</typeStatus>
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<collectionCode id="3C8AAE296B70FFCDBFCCFB3CFDFEE4EF" box="[389,514,1187,1211]" pageId="16" pageNumber="15">DEPOSITORY</collectionCode>
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.
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<collectingCountry id="228C767C6B70FFCDBC42FB3CFDBDE4EE" box="[523,577,1187,1210]" name="Laos" pageId="16" pageNumber="15">Laos</collectingCountry>
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,
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<location id="5F4460376B70FFCDBC05FB3CFD7CE4EE" LSID="urn:lsid:plazi:treatment:D23287FA6B70FFCDBE94FCFEFD73E84F:5F4460376B70FFCDBC05FB3CFD7CE4EE" box="[588,640,1187,1210]" country="Laos" name="Neis" pageId="16" pageNumber="15">Neïs</location>
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</materialsCitation>
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;
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<materialsCitation id="EAF33CB16B70FFCDBCC2FB3CFF56E48F" collectionCode="MNHN" country="Laos" pageId="16" pageNumber="15" specimenCode="RS 0030" specimenCount="1" typeStatus="holotype">
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<collectionCode id="3C8AAE296B70FFCDBCC2FB3CFD20E4EE" box="[651,732,1187,1210]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="16" pageNumber="15">MNHN</collectionCode>
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<specimenCode id="0A3D9E976B70FFCDBCABFB3CFF56E48F" collectionCode="MNHN" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="16" pageNumber="15">RS 0030</specimenCode>
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</materialsCitation>
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.
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</paragraph>
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<paragraph id="5A2436EC6B70FFCDBE38FA9CFE18E56F" blockId="16.[113,771,1282,1339]" pageId="16" pageNumber="15">
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<materialsCitation id="EAF33CB16B70FFCDBE38FA9CFE1CE56F" collectingDate="2019-05" collectionCode="TUPC" country="Thailand" location="Mae Tha District" pageId="16" pageNumber="15" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Lamphun">
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MATERIAL EXAMINED.
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|
||
<collectingCountry id="228C767C6B70FFCDBF10FA9DFE3DE54E" box="[345,449,1282,1306]" name="Thailand" pageId="16" pageNumber="15">Thailand</collectingCountry>
|
||
</emphasis>
|
||
,
|
||
<emphasis id="68EFEAFE6B70FFCDBF87FA9CFD64E54E" box="[462,664,1283,1307]" italics="true" pageId="16" pageNumber="15">
|
||
<collectingRegion id="985FF80E6B70FFCDBF87FA9CFD64E54E" box="[462,664,1283,1307]" country="Thailand" name="Lamphun" pageId="16" pageNumber="15">Lamphun Province</collectingRegion>
|
||
</emphasis>
|
||
,
|
||
<location id="5F4460376B70FFCDBCECFA9CFF38E56F" LSID="urn:lsid:plazi:treatment:D23287FA6B70FFCDBE94FCFEFD73E84F:5F4460376B70FFCDBCECFA9CFF38E56F" country="Thailand" name="Mae Tha District" pageId="16" pageNumber="15" stateProvince="Lamphun">Mae Tha District</location>
|
||
,
|
||
<date id="2E25102C6B70FFCDBE86FABCFEC2E56F" box="[207,318,1315,1339]" pageId="16" pageNumber="15" value="2019-05">
|
||
<collectingDate id="3E61E9C46B70FFCDBE86FABCFEC2E56F" box="[207,318,1315,1339]" pageId="16" pageNumber="15" value="2019-05">May 2019</collectingDate>
|
||
</date>
|
||
,
|
||
<specimenCount id="4C9DFD656B70FFCDBF00FABCFE93E56F" box="[329,367,1315,1339]" pageId="16" pageNumber="15" type="male">1♂</specimenCount>
|
||
<specimenCount id="4C9DFD656B70FFCDBF26FABCFE71E56F" box="[367,397,1315,1339]" pageId="16" pageNumber="15" type="female">1♀</specimenCount>
|
||
,
|
||
<collectionCode id="3C8AAE296B70FFCDBFD1FABCFE1CE56F" box="[408,480,1315,1339]" pageId="16" pageNumber="15">TUPC</collectionCode>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="128165676B70FFCDBE38FAFCFD40E7CF" pageId="16" pageNumber="15" type="diagnosis">
|
||
<paragraph id="5A2436EC6B70FFCDBE38FAFCFD40E7CF" blockId="16.[113,772,1379,1947]" pageId="16" pageNumber="15">
|
||
DIAGNOSIS (modified from
|
||
<bibRefCitation id="3E0A4B1D6B70FFCDBFC1FAFCFDE2E52F" author="KOVARIK" box="[392,542,1379,1403]" pageId="16" pageNumber="15" pagination="1 - 60" refId="ref11828" refString="KOVARIK, F. 2004. A review of the genus Heterometrus Ehrenberg, 1828, with descriptions of seven new species (Scorpiones, Scorpionidae). Euscorpius, 15: 1 - 60." type="journal article" year="2004">Kovařík, 2004</bibRefCitation>
|
||
and
|
||
<bibRefCitation id="3E0A4B1D6B70FFCDBC04FAFCFF56E5CF" author="PRENDINI" pageId="16" pageNumber="15" refId="ref12183" refString="PRENDINI, L. & S. F. LORIA. 2020. Systematic revision of the Asian forest scorpions (Heterometrinae Simon, 1879), revised suprageneric classification of Scorpionidae Latreille, 1802, and revalidation of Rugodentidae Bastawade et al., 2005. Bulletin of the American Museum of Natural History, 442, 480 pp." type="book" year="2020">Prendini & Loria, 2020</bibRefCitation>
|
||
). Total length of adults
|
||
<quantity id="9D639B096B70FFCDBFE2FA1DFDD7E5CE" box="[427,555,1410,1434]" metricMagnitude="-1" metricUnit="m" metricValue="1.075" metricValueMax="1.25" metricValueMin="0.9" pageId="16" pageNumber="15" unit="mm" value="107.5" valueMax="125.0" valueMin="90.0">90–125 mm</quantity>
|
||
(male
|
||
<typeStatus id="8520884E6B70FFCDBC30FA1CFD28E5CF" box="[633,724,1411,1435]" pageId="16" pageNumber="15" type="holotype">holotype</typeStatus>
|
||
<quantity id="9D639B096B70FFCDBC94FA1CFF61E5EE" metricMagnitude="-1" metricUnit="m" metricValue="1.17" pageId="16" pageNumber="15" unit="mm" value="117.0">117 mm</quantity>
|
||
;
|
||
<bibRefCitation id="3E0A4B1D6B70FFCDBEE7FA3CFEB7E5EF" author="COUZIJN" box="[174,331,1443,1467]" pageId="16" pageNumber="15" pagination="1 - 196" refId="ref11657" refString="COUZIJN, H. W. C. 1981. Revision of the genus Heterometrus. Zoologische Verhandelingen, 184: 1 - 196." type="journal article" year="1981">Couzijn, 1981</bibRefCitation>
|
||
). Base color of adults uniformly black, cuticular surface weakly to moderately lustrous (
|
||
<figureCitation id="C2A02A696B70FFCDBCD3FA5CFF73E5AE" captionStart="Figures 72–75" captionStartId="18.[113,190,2026,2048]" captionTargetBox="[135,1451,229,2009]" captionTargetId="figure-46@18.[116,1477,217,2014]" captionTargetPageId="18" captionText="Figures 72–75: Heterometrus laoticus, in dorsal (72, 74) and ventral (73, 75) aspects; scale bar = 10 mm. Figures 72–73. Male from Lamphun, TUPC. Figures 74–75. Female from Lamphun, TUPC." figureDoi="http://doi.org/10.5281/zenodo.8109008" httpUri="https://zenodo.org/record/8109008/files/figure.png" pageId="16" pageNumber="15">Figs. 72– 75</figureCitation>
|
||
). Carapace surface essentially smooth, with few granules scattered on lateral surfaces but absent in anteromedian area; dorsal profile triangular (resembles isosceles trapezoid) (
|
||
<figureCitation id="C2A02A696B70FFCDBC86F9BCFF4BE60E" captionStart="Figures 20–29" captionStartId="7.[115,192,993,1015]" captionTargetBox="[118,1460,249,936]" captionTargetId="figure-433@7.[113,1474,234,982]" captionTargetPageId="7" captionText="Figures 20–29: Comparison of Heterometrus spp. in Thailand, prosoma in dorsal aspect. Figures 20–21. H. cimrmani, male from Trang (20) and female from Nakhon Si Thammarat (21), THNHM. Figures 22–23. H. minotaurus, male (22) and female from Surat Thani (23), THNHM. Figures 24–25. H. silenus, male from Chanthaburi (24) and female from Chonburi (25), TUPC. Figures 26–27. H. laoticus, male (26) and female from Lamphun (27), TUPC. Figures 28–29. H. spinifer, males from Yala (28) and Narathiwat (29), TUPC." figureDoi="http://doi.org/10.5281/zenodo.8108988" httpUri="https://zenodo.org/record/8108988/files/figure.png" pageId="16" pageNumber="15">Figs. 26–27</figureCitation>
|
||
). Tergite surface in both sexes smooth with few granules (
|
||
<figureCitation id="C2A02A696B70FFCDBE30F9FCFF02E62E" box="[121,254,1634,1659]" captionStart="Figures 72–75" captionStartId="18.[113,190,2026,2048]" captionTargetBox="[135,1451,229,2009]" captionTargetId="figure-46@18.[116,1477,217,2014]" captionTargetPageId="18" captionText="Figures 72–75: Heterometrus laoticus, in dorsal (72, 74) and ventral (73, 75) aspects; scale bar = 10 mm. Figures 72–73. Male from Lamphun, TUPC. Figures 74–75. Female from Lamphun, TUPC." figureDoi="http://doi.org/10.5281/zenodo.8109008" httpUri="https://zenodo.org/record/8109008/files/figure.png" pageId="16" pageNumber="15">Figs. 72–75</figureCitation>
|
||
). PTC
|
||
<quantity id="9D639B096B70FFCDBF07F9FDFE52E62F" box="[334,430,1634,1659]" metricMagnitude="-1" metricUnit="m" metricValue="4.318" metricValueMax="4.826" metricValueMin="3.81" pageId="16" pageNumber="15" unit="in" value="17.0" valueMax="19.0" valueMin="15.0">15–19 in</quantity>
|
||
both sexes. Pedipalp relatively stout and short in both sexes among congeners (
|
||
<figureCitation id="C2A02A696B70FFCDBC3CF91CFD04E6CE" box="[629,760,1666,1691]" captionStart="Figures 72–75" captionStartId="18.[113,190,2026,2048]" captionTargetBox="[135,1451,229,2009]" captionTargetId="figure-46@18.[116,1477,217,2014]" captionTargetPageId="18" captionText="Figures 72–75: Heterometrus laoticus, in dorsal (72, 74) and ventral (73, 75) aspects; scale bar = 10 mm. Figures 72–73. Male from Lamphun, TUPC. Figures 74–75. Female from Lamphun, TUPC." figureDoi="http://doi.org/10.5281/zenodo.8109008" httpUri="https://zenodo.org/record/8109008/files/figure.png" pageId="16" pageNumber="15">Figs. 72–75</figureCitation>
|
||
); chelae of both sexes robust and short, ChL/W:
|
||
<specimenCount id="4C9DFD656B70FFCDBC13F93CFD91E6EF" box="[602,621,1699,1723]" pageId="16" pageNumber="15" type="male">♂</specimenCount>
|
||
<specimenCount id="4C9DFD656B70FFCDBC24F93CFD86E6EF" box="[621,634,1699,1723]" pageId="16" pageNumber="15" type="female">♀</specimenCount>
|
||
2.0–2.3; FL/ CL:
|
||
<specimenCount id="4C9DFD656B70FFCDBEE9F95CFF4BE68E" box="[160,183,1731,1754]" pageId="16" pageNumber="15" type="male">♂</specimenCount>
|
||
<specimenCount id="4C9DFD656B70FFCDBEF1F95CFF34E68E" box="[184,200,1731,1754]" pageId="16" pageNumber="15" type="female">♀</specimenCount>
|
||
<0.74. Dorsal surface of chelal manus smooth in both sexes (
|
||
<figureCitation id="C2A02A696B70FFCDBEF5F97CFEBDE6AE" box="[188,321,1762,1787]" captionStart="Figures 30–39" captionStartId="8.[113,190,1686,1708]" captionTargetBox="[147,1464,246,1661]" captionTargetId="figure-206@8.[115,1476,234,1687]" captionTargetPageId="8" captionText="Figures 30–39: Comparison of Heterometrus spp. in Thailand, chela in dorsal aspect (trichobothria indicated by red circles). Figures 30–31. H. cimrmani, male from Trang (30) and female from Nakhon Si Thammarat (31), THNHM. Figures 32–33. H. minotaurus, male (32) and female from Surat Thani (33), THNHM. Figures 34–35. H. silenus, male from Chanthaburi (34) and female from Chonburi (35), TUPC. Figures 36–37. H. laoticus, male (36) and female from Lamphun (37), TUPC. Figures 28–29. H. spinifer, males from Yala (38) and Narathiwat (39), TUPC." figureDoi="http://doi.org/10.5281/zenodo.8108990" httpUri="https://zenodo.org/record/8108990/files/figure.png" pageId="16" pageNumber="15">Figs. 36–37</figureCitation>
|
||
); prodorsal surface scattered with minute spiniform granules (
|
||
<figureCitation id="C2A02A696B70FFCDBF1BF89CFE26E74E" box="[338,474,1794,1819]" captionStart="Figures 50–57" captionStartId="10.[115,192,1986,2008]" captionTargetBox="[120,1462,220,1967]" captionTargetId="figure-90@10.[113,1474,208,1985]" captionTargetPageId="10" captionText="Figures 50–57: Comparison of Heterometrus spp. in Thailand, chela in ventral aspect (trichobothria indicated by red circles). Figures 50–51. H. cimrmani, male from Trang (50) and female from Nakhon Si Thammarat (51), THNHM. Figures 52–53. H. minotaurus, male (52) and female from Surat Thani (53), THNHM. Figures 54–55. H. silenus, male from Chanthaburi (54) and female from Chonburi (55), TUPC. Figures 56–57. H. laoticus, male (56) and female from Lamphun (57), TUPC." figureDoi="http://doi.org/10.5281/zenodo.8108994" httpUri="https://zenodo.org/record/8108994/files/figure.png" pageId="16" pageNumber="15">Figs. 56–57</figureCitation>
|
||
); fixed finger shorter than manus (
|
||
<figureCitation id="C2A02A696B70FFCDBE8AF8BCFEBEE76E" box="[195,322,1826,1851]" captionStart="Figures 40–49" captionStartId="9.[113,190,1519,1541]" captionTargetBox="[119,1440,244,1498]" captionTargetId="figure-248@9.[113,1474,234,1522]" captionTargetPageId="9" captionText="Figures 40–49: Comparison of Heterometrus spp. in Thailand, chela in external aspect (trichobothria indicated by red circles). Figures 40–41. H. cimrmani, male from Trang (40) and female from Nakhon Si Thammarat (41), THNHM. Figures 42–43. H. minotaurus, male (42) and female from Surat Thani (43), THNHM. Figures 44–45. H. silenus, male from Chanthaburi (44) and female from Chonburi (45), TUPC. Figures 46–47. H. laoticus, male (46) and female from Lamphun (47), TUPC. Figures 48–49. H. spinifer, males from Yala (48) and Narathiwat (49), TUPC." figureDoi="http://doi.org/10.5281/zenodo.8108992" httpUri="https://zenodo.org/record/8108992/files/figure.png" pageId="16" pageNumber="15">Figs. 46–47</figureCitation>
|
||
). Proximal lobe on cutting edge of movable finger lacks pronounced sexual dimorphism, but that of the males is slightly larger than females (
|
||
<figureCitation id="C2A02A696B70FFCDBC40F8FCFD70E72E" box="[521,652,1890,1915]" captionStart="Figures 40–49" captionStartId="9.[113,190,1519,1541]" captionTargetBox="[119,1440,244,1498]" captionTargetId="figure-248@9.[113,1474,234,1522]" captionTargetPageId="9" captionText="Figures 40–49: Comparison of Heterometrus spp. in Thailand, chela in external aspect (trichobothria indicated by red circles). Figures 40–41. H. cimrmani, male from Trang (40) and female from Nakhon Si Thammarat (41), THNHM. Figures 42–43. H. minotaurus, male (42) and female from Surat Thani (43), THNHM. Figures 44–45. H. silenus, male from Chanthaburi (44) and female from Chonburi (45), TUPC. Figures 46–47. H. laoticus, male (46) and female from Lamphun (47), TUPC. Figures 48–49. H. spinifer, males from Yala (48) and Narathiwat (49), TUPC." figureDoi="http://doi.org/10.5281/zenodo.8108992" httpUri="https://zenodo.org/record/8108992/files/figure.png" pageId="16" pageNumber="15">Figs. 46–47</figureCitation>
|
||
). Telson in adults reddish black to pitch black (
|
||
<figureCitation id="C2A02A696B70FFCDBFAAF81CFD9CE7CF" box="[483,608,1923,1947]" captionStart="Figures 4–11" captionStartId="5.[116,193,1930,1952]" captionTargetBox="[133,1447,263,1857]" captionTargetId="figure-82@5.[113,1474,234,1905]" captionTargetPageId="5" captionText="Figures 4–11: Comparison of Heterometrus spp. in Thailand, metasoma and telson in dorsal aspect. Figures 4–5. H. cimrmani, male from Trang (4) and female from Nakhon Si Thammarat (5), THNHM. Figures 6–7. H. minotaurus, male (6) and female from Surat Thani (7), THNHM. Figures 8–9. H. silenus, male from Chanthaburi (8) and female from Chonburi (9), TUPC. Figures 10–11. H. laoticus, male (10) and female from Lamphun (11), TUPC." figureDoi="http://doi.org/10.5281/zenodo.8108984" httpUri="https://zenodo.org/record/8108984/files/figure.png" pageId="16" pageNumber="15">Figs. 10–11</figureCitation>
|
||
,
|
||
<figureCitation id="C2A02A696B70FFCDBC22F81CFD4CE7CF" box="[619,688,1923,1947]" captionStart="Figures 12–19" captionStartId="6.[115,192,1927,1949]" captionTargetBox="[124,1469,261,1873]" captionTargetId="figure-87@6.[113,1474,234,1915]" captionTargetPageId="6" captionText="Figures 12–19: Comparison of Heterometrus spp. in Thailand, metasoma and telson in ventral aspect. Figures 12–13. H. cimrmani, male from Trang (12) and female from Nakhon Si Thammarat (13), THNHM. Figures 14–15. H. minotaurus, male (14) and female from Surat Thani (15), THNHM. Figures 16–17. H. silenus, male from Chanthaburi (16) and female from Chonburi (17), TUPC. Figures 18–19. H. laoticus, male (18) and female from Lamphun (19), TUPC." figureDoi="http://doi.org/10.5281/zenodo.8108986" httpUri="https://zenodo.org/record/8108986/files/figure.png" pageId="16" pageNumber="15">18–19</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="128165676B70FFCDBE38F85CFD73E84F" pageId="16" pageNumber="15" type="distribution">
|
||
<paragraph id="5A2436EC6B70FFCDBE38F85CFD73E84F" blockId="16.[113,771,1987,2075]" pageId="16" pageNumber="15">
|
||
DISTRIBUTION. This species is widely distributed in northern, central, and western
|
||
<collectingCountry id="228C767C6B70FFCDBF1BF87CFE48E7AF" box="[338,436,2019,2043]" name="Thailand" pageId="16" pageNumber="15">Thailand</collectingCountry>
|
||
. This species is found in
|
||
<collectingCountry id="228C767C6B70FFCDBC81F87CFD03E7AE" box="[712,767,2019,2042]" name="Laos" pageId="16" pageNumber="15">Laos</collectingCountry>
|
||
; the record from
|
||
<collectingCountry id="228C767C6B70FFCDBF57F79CFE70E84F" box="[286,396,2051,2075]" name="Cambodia" pageId="16" pageNumber="15">Cambodia</collectingCountry>
|
||
is doubtful (see below).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |