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<document id="386F0C44911CE8A4E472F5855A037D4D" ID-DOI="10.5281/zenodo.5360916" ID-ISSN="2345-7600" ID-Zenodo-Dep="5360916" ID-ZooBank="5AC21789-E1C1-4B48-813D-667D7638893C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630235294695" checkinUser="valdenar" docAuthor="Uy, Christine Jewel C., Malicky, Hans &amp; Bae, Yeon Jae" docDate="2018" docId="862D87A3FFA8FF83FF5AA8F6FC077BC4" docLanguage="en" docName="66rbz664-703.pdf" docOrigin="Raffles Bulletin of Zoology 66" docStyle="DocumentStyle:F2D34FF25DBF9998BB8DF6F24A5B097A.5:RafflesBZool.2014-2018.journal_article" docStyleId="F2D34FF25DBF9998BB8DF6F24A5B097A" docStyleName="RafflesBZool.2014-2018.journal_article" docStyleVersion="5" docTitle="Polymorphanisus scutellatus Banks 1939" docType="treatment" docVersion="4" lastPageNumber="693" masterDocId="7A14FFDBFFB5FF9EFF84AD57FF997C5E" masterDocTitle="Review of the filter-feeding caddisfly subfamily Macronematinae (Trichoptera: Hydropsychidae) in tropical Southeast Asia" masterLastPageNumber="703" masterPageNumber="664" pageNumber="693" updateTime="1699194071636" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="B8E8525BB9FF1AF110A266D03CA79D6B">Review of the filter-feeding caddisfly subfamily Macronematinae (Trichoptera: Hydropsychidae) in tropical Southeast Asia</mods:title>
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<mods:namePart id="015F4730EFDDB89529F764801DA3499A">Uy, Christine Jewel C.</mods:namePart>
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<mods:namePart id="8948E646F42A5F4ACAE56B092B779E58">Malicky, Hans</mods:namePart>
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<mods:namePart id="ACA8C81761967D5DC1D7C0E53B522AAC">Bae, Yeon Jae</mods:namePart>
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<treatment id="862D87A3FFA8FF83FF5AA8F6FC077BC4" ID-DOI="http://doi.org/10.5281/zenodo.5457041" ID-Zenodo-Dep="5457041" LSID="urn:lsid:plazi:treatment:862D87A3FFA8FF83FF5AA8F6FC077BC4" httpUri="http://treatment.plazi.org/id/862D87A3FFA8FF83FF5AA8F6FC077BC4" lastPageNumber="693" pageId="29" pageNumber="693">
<subSubSection id="469E653EFFA8FF83FF5AA8F6FD2879E7" box="[222,689,1441,1466]" pageId="29" pageNumber="693" type="nomenclature">
<paragraph id="0E3B36B5FFA8FF83FF5AA8F6FD2879E7" blockId="29.[222,689,1441,1498]" box="[222,689,1441,1466]" pageId="29" pageNumber="693">
<heading id="557381D9FFA8FF83FF5AA8F6FD2879E7" bold="true" box="[222,689,1441,1466]" centered="true" fontSize="10" level="2" pageId="29" pageNumber="693" reason="2">
<taxonomicName id="C9844D36FFA8FF83FF5AA8F6FD2879E7" ID-CoL="4LBLP" authority="Banks, 1939" authorityName="Banks" authorityYear="1939" box="[222,689,1441,1466]" class="Insecta" family="Hydropsychidae" genus="Polymorphanisus" kingdom="Animalia" order="Trichoptera" pageId="29" pageNumber="693" phylum="Arthropoda" rank="species" species="scutellatus">
<emphasis id="3CF0EAA7FFA8FF83FF5AA8F6FD2879E7" bold="true" box="[222,689,1441,1466]" pageId="29" pageNumber="693">
<emphasis id="3CF0EAA7FFA8FF83FF5AA8F6FD8579E7" bold="true" box="[222,540,1441,1465]" italics="true" pageId="29" pageNumber="693">Polymorphanisus scutellatus</emphasis>
<bibRefCitation id="6A154B44FFA8FF83FDA1A8F5FD2879E7" author="Banks N" box="[549,689,1441,1466]" pageId="29" pageNumber="693" pagination="52 - 61" refId="ref25480" refString="Banks N (1939) Notes and descriptions of Oriental Oestropsychinae (Trichoptera). Psyche, 46: 52 - 61." type="journal article" year="1939">Banks, 1939</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="469E653EFFA8FF83FE87A896FD147984" box="[259,653,1473,1498]" pageId="29" pageNumber="693" type="description">
<paragraph id="0E3B36B5FFA8FF83FE87A896FD147984" blockId="29.[222,689,1441,1498]" box="[259,653,1473,1498]" pageId="29" pageNumber="693">
(
<figureCitation id="96BF2A30FFA8FF83FE8EA896FE337984" box="[266,426,1473,1498]" captionStart="Figs" captionStartId="25.[136,180,1964,1986]" captionTargetBox="[141,1475,231,1948]" captionTargetId="figure-8@25.[127,1483,211,1957]" captionTargetPageId="25" captionText="Figs. 183192. Polymorphanisus thorax. 183, astictus, ocularis male and female thorax; 16, fuscus male thorax; 184185, fuscus female thorax; 186, muluensis male and female thorax; 187, nigricornis male and female thorax; 188, quadripunctatus male and female thorax; 189, semperi male and female thorax; 190, scutellatus male thorax; 191, scutellatus female thorax; 192, unipunctus male and female thorax. Scale = 2 mm (189 redrawn from Barnard, 1980)." figureDoi="http://doi.org/10.5281/zenodo.5360953" httpUri="https://zenodo.org/record/5360953/files/figure.png" pageId="29" pageNumber="693">Figs. 190, 191</figureCitation>
,
<figureCitation id="96BF2A30FFA8FF83FE33A895FE7B7984" box="[439,482,1474,1498]" captionStart="Figs" captionStartId="27.[136,180,1351,1373]" captionTargetBox="[133,1475,171,1405]" captionTargetId="figure-8@27.[133,1475,171,1405]" captionTargetPageId="27" captionText="Figs. 193201. Polymorphanisus male right fore- and hind wings. 193, astictus; 194, fuscus; 195, muluensis; 196, nigricornis; 197, ocularis; 198, quadripunctatus; 199, semperi; 200, scutellatus; 201, unipunctus. Scale = 5 mm (199 redrawn from Barnard, 1980)." figureDoi="http://doi.org/10.5281/zenodo.5360955" httpUri="https://zenodo.org/record/5360955/files/figure.png" pageId="29" pageNumber="693">200</figureCitation>
, 220222, 233)
</paragraph>
</subSubSection>
<subSubSection id="469E653EFFA8FF83FF0CAB57FE4E7A32" pageId="29" pageNumber="693" type="reference_group">
<paragraph id="0E3B36B5FFA8FF83FF0CAB57FEDF7A6C" blockId="29.[136,775,1536,1644]" pageId="29" pageNumber="693">
<treatmentCitationGroup id="2E94119BFFA8FF83FF0CAB57FEDF7A6C" pageId="29" pageNumber="693">
<treatmentCitation id="8F2510A4FFA8FF83FF0CAB57FDD27A48" author="Banks N" box="[136,587,1536,1558]" page="55" pageId="29" pageNumber="693" year="1939">
<taxonomicName id="C9844D36FFA8FF83FF0CAB57FDD27A48" ID-CoL="4LBLP" authority="Banks, 1939: 55" authorityName="Banks" authorityPageNumber="55" authorityYear="1939" box="[136,587,1536,1558]" class="Insecta" family="Hydropsychidae" genus="Polymorphanisus" kingdom="Animalia" order="Trichoptera" pageId="29" pageNumber="693" phylum="Arthropoda" rank="species" species="scutellatus">
<emphasis id="3CF0EAA7FFA8FF83FF0CAB57FE397A4B" box="[136,416,1536,1557]" italics="true" pageId="29" pageNumber="693">Polymorphanisus scutellatus</emphasis>
<materialsCitation id="BEEC3CE8FFA8FF83FE2DAB57FDD27A48" box="[425,587,1536,1558]" collectionCode="MCZ" pageId="29" pageNumber="693" specimenCount="1" typeStatus="holotype">
<bibRefCitation id="6A154B44FFA8FF83FE2DAB57FDD27A48" author="Banks N" box="[425,587,1536,1558]" pageId="29" pageNumber="693" pagination="52 - 61" refId="ref25480" refString="Banks N (1939) Notes and descriptions of Oriental Oestropsychinae (Trichoptera). Psyche, 46: 52 - 61." type="journal article" year="1939">Banks, 1939: 55</bibRefCitation>
</materialsCitation>
</taxonomicName>
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[
<typeStatus id="D13F8817FFA8FF83FDDEAB57FD2F7A48" box="[602,694,1536,1558]" pageId="29" pageNumber="693" type="holotype">Holotype</typeStatus>
female; Borneo; MCZ].
</treatmentCitationGroup>
</paragraph>
<paragraph id="0E3B36B5FFA8FF83FF0CAB6DFE4E7A32" blockId="29.[136,775,1536,1644]" pageId="29" pageNumber="693">
<treatmentCitationGroup id="2E94119BFFA8FF83FF0CAB6DFE4E7A32" pageId="29" pageNumber="693">
<taxonomicName id="C9844D36FFA8FF83FF0CAB6DFE427A0E" authority="Banks" authorityName="Banks" box="[136,475,1594,1616]" class="Insecta" family="Hydropsychidae" genus="Polymorphanisus" kingdom="Animalia" order="Trichoptera" pageId="29" pageNumber="693" phylum="Arthropoda" rank="species" species="scutellaris">
<emphasis id="3CF0EAA7FFA8FF83FF0CAB6DFE0E7A11" box="[136,407,1594,1615]" italics="true" pageId="29" pageNumber="693">Polymorphanisus scutellaris</emphasis>
Banks
</taxonomicName>
;
<treatmentCitation id="8F2510A4FFA8FF83FE67AB6DFD3C7A11" author="Kimmins DE" box="[483,677,1593,1616]" page="399" pageId="29" pageNumber="693" year="1955">
<bibRefCitation id="6A154B44FFA8FF83FE67AB6DFD3C7A11" author="Kimmins DE" box="[483,677,1593,1616]" pageId="29" pageNumber="693" pagination="374 - 442" refId="ref26171" refString="Kimmins DE (1955) Results of the Oxford University expedition to Sarawak, 1932. Order Trichoptera. The Sarawak Museum Journal, 6: 374 - 442." type="journal article" year="1955">Kimmins, 1955: 399</bibRefCitation>
</treatmentCitation>
(incorrect spelling of
<taxonomicName id="C9844D36FFA8FF83FE9BAB01FE557A32" ID-CoL="4LBLP" authority="Banks" authorityName="Banks" authorityYear="1939" box="[287,460,1622,1644]" class="Insecta" family="Hydropsychidae" genus="Polymorphanisus" kingdom="Animalia" order="Trichoptera" pageId="29" pageNumber="693" phylum="Arthropoda" rank="species" species="scutellatus">
<emphasis id="3CF0EAA7FFA8FF83FE9BAB01FE1F7A35" box="[287,390,1622,1643]" italics="true" pageId="29" pageNumber="693">scutellatus</emphasis>
Banks
</taxonomicName>
).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="469E653EFFA8FF83FF0CABC3FB1C79A4" pageId="29" pageNumber="693" type="diagnosis">
<paragraph id="0E3B36B5FFA8FF83FF0CABC3FDF97B73" blockId="29.[136,775,1684,1837]" pageId="29" pageNumber="693">
<emphasis id="3CF0EAA7FFA8FF83FF0CABC3FE9B7AF2" bold="true" box="[136,258,1684,1708]" pageId="29" pageNumber="693">Diagnosis.</emphasis>
<emphasis id="3CF0EAA7FFA8FF83FE89ABC2FED47AF3" box="[269,333,1685,1709]" italics="true" pageId="29" pageNumber="693">Male.</emphasis>
Antennae
<quantity id="C97C9B50FFA8FF83FE49ABC3FD847AF2" box="[461,541,1684,1708]" metricMagnitude="-2" metricUnit="m" metricValue="5.0" pageId="29" pageNumber="693" unit="mm" value="50.0">50 mm</quantity>
in length. Antennal flagellum brown with darker striations. Forewing
<quantity id="C97C9B50FFA8FF83FD3CABE2FC9F7A92" box="[696,774,1717,1741]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" pageId="29" pageNumber="693" unit="mm" value="20.0">20 mm</quantity>
and hind wing
<quantity id="C97C9B50FFA8FF83FEB0AB82FE1D7AB2" box="[308,388,1749,1773]" metricMagnitude="-2" metricUnit="m" metricValue="1.4" pageId="29" pageNumber="693" unit="mm" value="14.0">14 mm</quantity>
in length (
<figureCitation id="96BF2A30FFA8FF83FD80AB82FDFE7AB3" box="[516,615,1749,1773]" captionStart="Figs" captionStartId="27.[136,180,1351,1373]" captionTargetBox="[133,1475,171,1405]" captionTargetId="figure-8@27.[133,1475,171,1405]" captionTargetPageId="27" captionText="Figs. 193201. Polymorphanisus male right fore- and hind wings. 193, astictus; 194, fuscus; 195, muluensis; 196, nigricornis; 197, ocularis; 198, quadripunctatus; 199, semperi; 200, scutellatus; 201, unipunctus. Scale = 5 mm (199 redrawn from Barnard, 1980)." figureDoi="http://doi.org/10.5281/zenodo.5360955" httpUri="https://zenodo.org/record/5360955/files/figure.png" pageId="29" pageNumber="693">Fig. 200</figureCitation>
). Mesoscutum without evident markings. Mesoscutellum with two oval markings as in
<figureCitation id="96BF2A30FFA8FF83FEB4AA42FE177B73" box="[304,398,1813,1837]" captionStart="Figs" captionStartId="25.[136,180,1964,1986]" captionTargetBox="[141,1475,231,1948]" captionTargetId="figure-8@25.[127,1483,211,1957]" captionTargetPageId="25" captionText="Figs. 183192. Polymorphanisus thorax. 183, astictus, ocularis male and female thorax; 16, fuscus male thorax; 184185, fuscus female thorax; 186, muluensis male and female thorax; 187, nigricornis male and female thorax; 188, quadripunctatus male and female thorax; 189, semperi male and female thorax; 190, scutellatus male thorax; 191, scutellatus female thorax; 192, unipunctus male and female thorax. Scale = 2 mm (189 redrawn from Barnard, 1980)." figureDoi="http://doi.org/10.5281/zenodo.5360953" httpUri="https://zenodo.org/record/5360953/files/figure.png" pageId="29" pageNumber="693">Fig. 190</figureCitation>
. Tibial spurs 1.3.2.
</paragraph>
<paragraph id="0E3B36B5FFA8FF83FF0CAA02FDF47BB3" blockId="29.[135,775,1876,2029]" pageId="29" pageNumber="693">
<emphasis id="3CF0EAA7FFA8FF83FF0CAA02FEB47B32" bold="true" box="[136,301,1876,1901]" pageId="29" pageNumber="693">Male genitalia.</emphasis>
Inferior appendages two-segmented, harpago as thick as coxopodite (Fig. 220). Phallus with wide opening in apex (Figs. 221, 222). Segment IX squarish, curving apicodorsad (Fig, 220). In dorsal view, segment X bifid with each half consisting of two lobes (
<figureCitation id="96BF2A30FFA8FF83FD85AA82FDF87BB3" box="[513,609,2005,2029]" captionStart="Figs" captionStartId="29.[136,180,1249,1271]" captionTargetBox="[143,1473,192,1234]" captionTargetId="figure-256@29.[122,1491,173,1244]" captionTargetPageId="29" captionText="Figs. 226234. Polymorphanisus segment X of male genitalia. 226, astictus; 227, fuscus; 228, muluensis; 229, nigricornis; 230, ocularis; 231, quadripunctatus; 232, semperi; 233, scutellatus; 234, unipunctus. Scale = 0.5 mm (232 redrawn from Barnard, 1980)." figureDoi="http://doi.org/10.5281/zenodo.5360957" httpUri="https://zenodo.org/record/5360957/files/figure.png" pageId="29" pageNumber="693">Fig. 233</figureCitation>
).
</paragraph>
<paragraph id="0E3B36B5FFA8FF83FCC7A815FB1C79A4" blockId="29.[835,1476,1345,1530]" pageId="29" pageNumber="693">
<emphasis id="3CF0EAA7FFA8FF83FCC7A815FC067904" bold="true" box="[835,927,1346,1370]" pageId="29" pageNumber="693">Female.</emphasis>
Antennae
<quantity id="C97C9B50FFA8FF83FB99A816FBF57907" box="[1053,1132,1345,1369]" metricMagnitude="-2" metricUnit="m" metricValue="3.5" pageId="29" pageNumber="693" unit="mm" value="35.0">35 mm</quantity>
in length. Antennal flagellum same colouration as male. Forewing and hind wing length same as male. Mesoscutum without evident markings. Mesoscutellum with two larger dark spots covering almost whole area and in some individuals looking like lung shape (
<figureCitation id="96BF2A30FFA8FF83FCC8A8B6FC3279A4" box="[844,939,1505,1530]" captionStart="Figs" captionStartId="25.[136,180,1964,1986]" captionTargetBox="[141,1475,231,1948]" captionTargetId="figure-8@25.[127,1483,211,1957]" captionTargetPageId="25" captionText="Figs. 183192. Polymorphanisus thorax. 183, astictus, ocularis male and female thorax; 16, fuscus male thorax; 184185, fuscus female thorax; 186, muluensis male and female thorax; 187, nigricornis male and female thorax; 188, quadripunctatus male and female thorax; 189, semperi male and female thorax; 190, scutellatus male thorax; 191, scutellatus female thorax; 192, unipunctus male and female thorax. Scale = 2 mm (189 redrawn from Barnard, 1980)." figureDoi="http://doi.org/10.5281/zenodo.5360953" httpUri="https://zenodo.org/record/5360953/files/figure.png" pageId="29" pageNumber="693">Fig. 191</figureCitation>
). Tibial spurs 1.3.2.
</paragraph>
</subSubSection>
<subSubSection id="469E653EFFA8FF83FCC7AB76FAFA7A04" pageId="29" pageNumber="693" type="materials_examined">
<paragraph id="0E3B36B5FFA8FF83FCC7AB76FAFA7A04" blockId="29.[835,1474,1569,1626]" pageId="29" pageNumber="693">
<emphasis id="3CF0EAA7FFA8FF83FCC7AB76FBB17A67" bold="true" box="[835,1064,1569,1593]" pageId="29" pageNumber="693">Material examined.</emphasis>
<materialsCitation id="BEEC3CE8FFA8FF83FBB6AB76FAC77A04" collectingDate="1994-03-06" collectorName="Aek Tarum &amp; H. Malicky" country="Indonesia" location="Indonesia" pageId="29" pageNumber="693" specimenCount="4" specimenCount-female="2" specimenCount-male="2">
<emphasis id="3CF0EAA7FFA8FF83FBB6AB76FB3F7A67" box="[1074,1190,1569,1593]" italics="true" pageId="29" pageNumber="693">
<collectingCountry id="76937625FFA8FF83FBB6AB76FB397A67" box="[1074,1184,1569,1593]" name="Indonesia" pageId="29" pageNumber="693">Indonesia</collectingCountry>
:
</emphasis>
<specimenCount id="1882FD3CFFA8FF83FB34AB75FA937A67" box="[1200,1290,1569,1594]" pageId="29" pageNumber="693" type="male">2 males</specimenCount>
,
<specimenCount id="1882FD3CFFA8FF83FA9CAB75FA1E7A67" box="[1304,1415,1569,1594]" pageId="29" pageNumber="693" type="female">2 females</specimenCount>
,
<collectorName id="A3715363FFA8FF83FA11AB76FC147A07" pageId="29" pageNumber="693">Aek Tarum</collectorName>
, coll.
<collectorName id="A3715363FFA8FF83FC55AB15FBC97A07" box="[977,1104,1601,1625]" pageId="29" pageNumber="693">H. Malicky</collectorName>
,
<date id="7A3A1075FFA8FF83FBD8AB16FB6C7A04" box="[1116,1269,1601,1626]" pageId="29" pageNumber="693" value="1994-03-06">
<collectingDate id="6A7EE99DFFA8FF83FBD8AB16FB6C7A04" box="[1116,1269,1601,1626]" pageId="29" pageNumber="693" value="1994-03-06">6 March 1994</collectingDate>
</date>
(HMPC)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="469E653EFFA8FF83FCC7ABD6FAD37AC7" box="[835,1354,1665,1690]" pageId="29" pageNumber="693" type="distribution">
<paragraph id="0E3B36B5FFA8FF83FCC7ABD6FAD37AC7" blockId="29.[835,1354,1665,1690]" box="[835,1354,1665,1690]" pageId="29" pageNumber="693">
<emphasis id="3CF0EAA7FFA8FF83FCC7ABD6FC4F7AC7" bold="true" box="[835,982,1665,1689]" pageId="29" pageNumber="693">Distribution.</emphasis>
Borneo,
<collectingRegion id="CC40F857FFA8FF83FBB8ABD5FB017AC4" box="[1084,1176,1666,1690]" country="Indonesia" name="Sumatera Utara" pageId="29" pageNumber="693">Sumatra</collectingRegion>
,
<collectingRegion id="CC40F857FFA8FF83FB20ABD5FB4F7AC7" box="[1188,1238,1666,1689]" country="Indonesia" name="Jawa Timur" pageId="29" pageNumber="693">Java</collectingRegion>
,
<collectingRegion id="CC40F857FFA8FF83FB67ABD6FADE7AC7" box="[1251,1351,1665,1689]" country="Indonesia" name="Sulawesi Utara" pageId="29" pageNumber="693">Sulawesi</collectingRegion>
.
</paragraph>
</subSubSection>
<subSubSection id="469E653EFFA8FF83FCC7AB95FC077BC4" pageId="29" pageNumber="693" type="discussion">
<paragraph id="0E3B36B5FFA8FF83FCC7AB95FC077BC4" blockId="29.[834,1475,1729,1946]" pageId="29" pageNumber="693">
<emphasis id="3CF0EAA7FFA8FF83FCC7AB95FC287A84" bold="true" box="[835,945,1730,1754]" pageId="29" pageNumber="693">Remarks.</emphasis>
<bibRefCitation id="6A154B44FFA8FF83FC32AB96FBCE7A84" author="Barnard PC" box="[950,1111,1729,1754]" pageId="29" pageNumber="693" pagination="59 - 106" refId="ref25534" refString="Barnard PC (1980) A revision of the Old World Polymorphanisini (Trichoptera: Hydropsychidae). Bulletin of the British Museum (Natural History), Entomology, 41 (2): 59 - 106." type="journal article" year="1980">Barnard (1980)</bibRefCitation>
only observed
<specimenCount id="1882FD3CFFA8FF83FB70AB95FAD47A84" box="[1268,1357,1730,1754]" pageId="29" pageNumber="693" type="generic" typeStatus="type">one type</specimenCount>
of thoracic markings for this species (
<figureCitation id="96BF2A30FFA8FF83FBD9ABB6FB227AA4" box="[1117,1211,1761,1786]" captionStart="Figs" captionStartId="25.[136,180,1964,1986]" captionTargetBox="[141,1475,231,1948]" captionTargetId="figure-8@25.[127,1483,211,1957]" captionTargetPageId="25" captionText="Figs. 183192. Polymorphanisus thorax. 183, astictus, ocularis male and female thorax; 16, fuscus male thorax; 184185, fuscus female thorax; 186, muluensis male and female thorax; 187, nigricornis male and female thorax; 188, quadripunctatus male and female thorax; 189, semperi male and female thorax; 190, scutellatus male thorax; 191, scutellatus female thorax; 192, unipunctus male and female thorax. Scale = 2 mm (189 redrawn from Barnard, 1980)." figureDoi="http://doi.org/10.5281/zenodo.5360953" httpUri="https://zenodo.org/record/5360953/files/figure.png" pageId="29" pageNumber="693">Fig. 190</figureCitation>
), whereas
<bibRefCitation id="6A154B44FFA8FF83FAAAABB6FA587AA7" author="Ulmer G" box="[1326,1473,1761,1785]" pageId="29" pageNumber="693" pagination="1 - 528" refId="ref27648" refString="Ulmer G (1951) Kocherfliegen (Trichopteren) von den Sunda-Inseln (Teil I). Archiv fur Hydrobiologie. Supplement, 19: 1 - 528." type="journal article" year="1951">Ulmer (1951)</bibRefCitation>
reported two variations. The specimens examined here exhibited
<specimenCount id="1882FD3CFFA8FF83FC2FAA75FB897B64" box="[939,1040,1826,1850]" pageId="29" pageNumber="693" type="generic" typeStatus="types">two types</specimenCount>
of thoracic markings, and the males were different from the females (
<figureCitation id="96BF2A30FFA8FF83FBF4AA16FA887B04" box="[1136,1297,1857,1882]" captionStart="Figs" captionStartId="25.[136,180,1964,1986]" captionTargetBox="[141,1475,231,1948]" captionTargetId="figure-8@25.[127,1483,211,1957]" captionTargetPageId="25" captionText="Figs. 183192. Polymorphanisus thorax. 183, astictus, ocularis male and female thorax; 16, fuscus male thorax; 184185, fuscus female thorax; 186, muluensis male and female thorax; 187, nigricornis male and female thorax; 188, quadripunctatus male and female thorax; 189, semperi male and female thorax; 190, scutellatus male thorax; 191, scutellatus female thorax; 192, unipunctus male and female thorax. Scale = 2 mm (189 redrawn from Barnard, 1980)." figureDoi="http://doi.org/10.5281/zenodo.5360953" httpUri="https://zenodo.org/record/5360953/files/figure.png" pageId="29" pageNumber="693">Figs. 190, 191</figureCitation>
). The examined specimens possessed tibial spur formulas of 2.3.2, 1.3.3, or 1.3.3.
</paragraph>
</subSubSection>
</treatment>
</document>