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<document id="9A51AC61EF2B5D8D3D0F198E80E9AF02" ID-DOI="10.5281/zenodo.5360916" ID-ISSN="2345-7600" ID-Zenodo-Dep="5360916" ID-ZooBank="5AC21789-E1C1-4B48-813D-667D7638893C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630235294695" checkinUser="valdenar" docAuthor="Uy, Christine Jewel C., Malicky, Hans &amp; Bae, Yeon Jae" docDate="2018" docId="862D87A3FF94FFBDFC0DABC1FD277AAB" docLanguage="en" docName="66rbz664-703.pdf" docOrigin="Raffles Bulletin of Zoology 66" docStyle="DocumentStyle:F2D34FF25DBF9998BB8DF6F24A5B097A.5:RafflesBZool.2014-2018.journal_article" docStyleId="F2D34FF25DBF9998BB8DF6F24A5B097A" docStyleName="RafflesBZool.2014-2018.journal_article" docStyleVersion="5" docTitle="Trichomacronema vietnamensis Uy &amp; Malicky &amp; Bae 2018, new species" docType="treatment" docVersion="3" lastPageNumber="699" masterDocId="7A14FFDBFFB5FF9EFF84AD57FF997C5E" masterDocTitle="Review of the filter-feeding caddisfly subfamily Macronematinae (Trichoptera: Hydropsychidae) in tropical Southeast Asia" masterLastPageNumber="703" masterPageNumber="664" pageNumber="697" updateTime="1699194071636" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
<mods:mods id="33B64DD899CFFC6AFCA737E05CC2111A" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="582A90E412F8A39B430AB1C8F55F51F0">Review of the filter-feeding caddisfly subfamily Macronematinae (Trichoptera: Hydropsychidae) in tropical Southeast Asia</mods:title>
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<mods:name id="D7F0F998FB3EA90D63D37C491FFECDB3" type="personal">
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<mods:namePart id="176976456E5883A1D76B4FA94BD0B327">Uy, Christine Jewel C.</mods:namePart>
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<mods:namePart id="B8FD6A8171B7BE9769BDACC1A7DE1AB3">Malicky, Hans</mods:namePart>
</mods:name>
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<mods:namePart id="0D94FF43701EBED3546ECAF1D413DB0F">Bae, Yeon Jae</mods:namePart>
</mods:name>
<mods:typeOfResource id="FB1D945390B513658C7DA9A7EC953F38">text</mods:typeOfResource>
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<mods:title id="B1320CB7218461270905D3669EA8BDCD">Raffles Bulletin of Zoology</mods:title>
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<mods:part id="F8CF0482F1ED69D9CF3416A87DE58D2E">
<mods:date id="8671A00CF8035EE9028DA984A0A27737">2018</mods:date>
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<mods:number id="7670753BE1952CDDB4FD2367FAEE79BD">2018-11-15</mods:number>
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<mods:number id="E6D3FE8DFF1140191ABFC5DF90DD80A6">66</mods:number>
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<mods:start id="F6B4AA131B7BB4BC5BA02A291FB9E20F">664</mods:start>
<mods:end id="983A831943167CC0F3200890269C4789">703</mods:end>
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<mods:classification id="EA2B6F5814D7702D1701C946BE879582">journal article</mods:classification>
<mods:identifier id="FC53F46DF63D7D38E04CBDC69F81F3EC" type="DOI">10.5281/zenodo.5360916</mods:identifier>
<mods:identifier id="C76A711A62ACA10AE4E2979F22CA4E72" type="ISSN">2345-7600</mods:identifier>
<mods:identifier id="90E198B975BC983441D7ACA07469A561" type="Zenodo-Dep">5360916</mods:identifier>
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</mods:mods>
<treatment id="862D87A3FF94FFBDFC0DABC1FD277AAB" LSID="urn:lsid:plazi:treatment:862D87A3FF94FFBDFC0DABC1FD277AAB" httpUri="http://treatment.plazi.org/id/862D87A3FF94FFBDFC0DABC1FD277AAB" lastPageId="35" lastPageNumber="699" pageId="33" pageNumber="697">
<subSubSection id="469E653EFF94FFBFFC0DABC1FAE67AF1" box="[905,1407,1686,1711]" pageId="33" pageNumber="697" type="nomenclature">
<paragraph id="0E3B36B5FF94FFBFFC0DABC1FAE67AF1" blockId="33.[905,1407,1686,1743]" box="[905,1407,1686,1711]" pageId="33" pageNumber="697">
<heading id="557381D9FF94FFBFFC0DABC1FAE67AF1" bold="true" box="[905,1407,1686,1711]" centered="true" fontSize="10" level="2" pageId="33" pageNumber="697" reason="2">
<emphasis id="3CF0EAA7FF94FFBFFC0DABC1FAE67AF1" bold="true" box="[905,1407,1686,1711]" pageId="33" pageNumber="697">
<taxonomicName id="C9844D36FF94FFBFFC0DABC1FB737AF1" authority="Uy &amp; Malicky &amp; Bae, 2018" authorityName="Uy &amp; Malicky &amp; Bae" authorityYear="2018" box="[905,1258,1686,1711]" class="Insecta" family="Hydropsychidae" genus="Trichomacronema" kingdom="Animalia" order="Trichoptera" pageId="33" pageNumber="697" phylum="Arthropoda" rank="species" species="vietnamensis" status="sp. nov.">
<emphasis id="3CF0EAA7FF94FFBFFC0DABC1FB737AF1" bold="true" box="[905,1258,1686,1711]" italics="true" pageId="33" pageNumber="697">Trichomacronema vietnamensis</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="27C357DCFF94FFBFFB7DABC0FAE67AF1" box="[1273,1407,1687,1711]" pageId="33" pageNumber="697" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="469E653EFF94FFBDFBA8ABE0FF627F0B" lastPageId="35" lastPageNumber="699" pageId="33" pageNumber="697" type="description">
<paragraph id="0E3B36B5FF94FFBFFBA8ABE0FB417A91" blockId="33.[905,1407,1686,1743]" box="[1068,1240,1719,1743]" pageId="33" pageNumber="697">
(
<figureCitation id="96BF2A30FF94FFBFFBB0ABE0FB487A91" box="[1076,1233,1719,1743]" captionStart="Figs" captionStartId="34.[113,157,1849,1871]" captionTargetBox="[111,1448,347,1832]" captionTargetId="figure-9@34.[97,1458,315,1842]" captionTargetPageId="34" captionText="Figs. 258–271. Trichomacronema paniae. 258, right forewing; Male genitalia: 259, lateral; 260, segment X dorsal; 261, Phallus lateral; 262, Phallus tip. Trichomacronema tamdao. 263, right forewing; Male genitalia: 264, lateral; 265, segment X dorsal; 266, Phallus lateral; 267, Phallus tip. Trichomacronema vietnamensis, new species. 268, right forewing; 269, right hind wing; Female genitalia: 270, lateral; 271, ventral. Scale: 258, 263, 268–269 = 2 mm; 259–262, 264–267, 270–271 = 0.02 mm." figureDoi="http://doi.org/10.5281/zenodo.5360961" httpUri="https://zenodo.org/record/5360961/files/figure.png" pageId="33" pageNumber="697">Figs. 268–271</figureCitation>
)
</paragraph>
<paragraph id="0E3B36B5FF94FFBFFCC7ABA0FB147B51" blockId="33.[835,1165,1783,1807]" box="[835,1165,1783,1807]" pageId="33" pageNumber="697">
<emphasis id="3CF0EAA7FF94FFBFFCC7ABA0FC577B51" bold="true" box="[835,974,1783,1807]" pageId="33" pageNumber="697">Description.</emphasis>
<emphasis id="3CF0EAA7FF94FFBFFC52ABA0FB8C7B51" box="[982,1045,1783,1807]" italics="true" pageId="33" pageNumber="697">Male.</emphasis>
Unknown.
</paragraph>
<paragraph id="0E3B36B5FF94FFBDFCC7AA60FE377E0B" blockId="33.[834,1475,1847,2095]" lastBlockId="35.[135,777,189,597]" lastPageId="35" lastPageNumber="699" pageId="33" pageNumber="697">
<emphasis id="3CF0EAA7FF94FFBFFCC7AA60FC067B11" bold="true" box="[835,927,1847,1871]" pageId="33" pageNumber="697">Female.</emphasis>
Antennae
<quantity id="C97C9B50FF94FFBFFB99AA60FBF57B10" box="[1053,1132,1847,1871]" metricMagnitude="-2" metricUnit="m" metricValue="2.6" pageId="33" pageNumber="697" unit="mm" value="26.0">26 mm</quantity>
in length. Antennal flagellum brown with darker striations. Head dark brown with five lighter setal warts; pair of large anterior setal warts occupying almost anterior half of head, very close to anteromesal setal wart; pair of posterior setal warts smaller. Forewing
<quantity id="C97C9B50FF94FFBFFAF3AAE0FA5B7B90" box="[1399,1474,1975,1999]" metricMagnitude="-2" metricUnit="m" metricValue="2.3" pageId="33" pageNumber="697" unit="mm" value="23.0">23 mm</quantity>
in length (
<figureCitation id="96BF2A30FF94FFBFFC33AA80FB817BB1" box="[951,1048,2007,2031]" captionStart="Figs" captionStartId="34.[113,157,1849,1871]" captionTargetBox="[111,1448,347,1832]" captionTargetId="figure-9@34.[97,1458,315,1842]" captionTargetPageId="34" captionText="Figs. 258–271. Trichomacronema paniae. 258, right forewing; Male genitalia: 259, lateral; 260, segment X dorsal; 261, Phallus lateral; 262, Phallus tip. Trichomacronema tamdao. 263, right forewing; Male genitalia: 264, lateral; 265, segment X dorsal; 266, Phallus lateral; 267, Phallus tip. Trichomacronema vietnamensis, new species. 268, right forewing; 269, right hind wing; Female genitalia: 270, lateral; 271, ventral. Scale: 258, 263, 268–269 = 2 mm; 259–262, 264–267, 270–271 = 0.02 mm." figureDoi="http://doi.org/10.5281/zenodo.5360961" httpUri="https://zenodo.org/record/5360961/files/figure.png" pageId="33" pageNumber="697">Fig. 268</figureCitation>
); veins Sc and R1 not joined; Sc vein ending in costa; R1 not ending in costa, obsolete; base of Rs obsolete, joined to R1 by crossvein; discoidal cell present; medial cell obvious; thyridial cell long; nygmas present in middle of thyridial cell and fork II; Cu1 and Cu 2 not joined; crossvein cu-a present. Hind wing
<quantity id="C97C9B50FF96FFBDFD84ADAAFDA77D4B" box="[512,574,253,277]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" pageId="35" pageNumber="699" unit="mm" value="9.0">9 mm</quantity>
in length; vein Sc joining R1, ending in costa; nygma present in fork II; median and thyridial cell absent. Wing pattern with alternating dark and brown longitudinal streak marks as shown in
<figureCitation id="96BF2A30FF96FFBDFD19AC0AFF2D7DCB" captionStart="Figs" captionStartId="34.[113,157,1849,1871]" captionTargetBox="[111,1448,347,1832]" captionTargetId="figure-9@34.[97,1458,315,1842]" captionTargetPageId="34" captionText="Figs. 258–271. Trichomacronema paniae. 258, right forewing; Male genitalia: 259, lateral; 260, segment X dorsal; 261, Phallus lateral; 262, Phallus tip. Trichomacronema tamdao. 263, right forewing; Male genitalia: 264, lateral; 265, segment X dorsal; 266, Phallus lateral; 267, Phallus tip. Trichomacronema vietnamensis, new species. 268, right forewing; 269, right hind wing; Female genitalia: 270, lateral; 271, ventral. Scale: 258, 263, 268–269 = 2 mm; 259–262, 264–267, 270–271 = 0.02 mm." figureDoi="http://doi.org/10.5281/zenodo.5360961" httpUri="https://zenodo.org/record/5360961/files/figure.png" pageId="35" pageNumber="699">Figs. 268, 269</figureCitation>
, more evident in forewing; presence of dark brown marking concentrated on half of anterior margin of median cell running towards base of M1 vein of forewing (
<figureCitation id="96BF2A30FF96FFBDFD58ACEAFF2D7DAB" captionStart="Figs" captionStartId="34.[113,157,1849,1871]" captionTargetBox="[111,1448,347,1832]" captionTargetId="figure-9@34.[97,1458,315,1842]" captionTargetPageId="34" captionText="Figs. 258–271. Trichomacronema paniae. 258, right forewing; Male genitalia: 259, lateral; 260, segment X dorsal; 261, Phallus lateral; 262, Phallus tip. Trichomacronema tamdao. 263, right forewing; Male genitalia: 264, lateral; 265, segment X dorsal; 266, Phallus lateral; 267, Phallus tip. Trichomacronema vietnamensis, new species. 268, right forewing; 269, right hind wing; Female genitalia: 270, lateral; 271, ventral. Scale: 258, 263, 268–269 = 2 mm; 259–262, 264–267, 270–271 = 0.02 mm." figureDoi="http://doi.org/10.5281/zenodo.5360961" httpUri="https://zenodo.org/record/5360961/files/figure.png" pageId="35" pageNumber="699">Fig. 268</figureCitation>
). Body large (about
<quantity id="C97C9B50FF96FFBDFE11AC8AFE7B7DAB" box="[405,482,477,501]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" pageId="35" pageNumber="699" unit="mm" value="12.0">12 mm</quantity>
in length) and dark brown in alcohol. Mesoscutum and mesoscutellum without evident markings. Legs brown, with dark brown mark at basal end of tibia; Tibial spurs 1.4.4.
</paragraph>
<caption id="5AFB663DFF97FFBCFFF5AA6EFC7D7BFB" ID-DOI="http://doi.org/10.5281/zenodo.5360961" ID-Zenodo-Dep="5360961" httpUri="https://zenodo.org/record/5360961/files/figure.png" pageId="34" pageNumber="698" startId="34.[113,157,1849,1871]" targetBox="[111,1448,347,1832]" targetPageId="34">
<paragraph id="0E3B36B5FF97FFBCFFF5AA6EFC7D7BFB" blockId="34.[113,1451,1849,1957]" pageId="34" pageNumber="698">
Figs. 258–271.
<taxonomicName id="C9844D36FF97FFBCFE8EAA6EFD9C7B11" authorityName="Malicky &amp; Chantaramongkol" authorityYear="1991" box="[266,517,1849,1871]" class="Insecta" family="Hydropsychidae" genus="Trichomacronema" kingdom="Animalia" order="Trichoptera" pageId="34" pageNumber="698" phylum="Arthropoda" rank="species" species="paniae">
<emphasis id="3CF0EAA7FF97FFBCFE8EAA6EFD9C7B11" box="[266,517,1849,1871]" italics="true" pageId="34" pageNumber="698">Trichomacronema paniae</emphasis>
</taxonomicName>
. 258, right forewing; Male genitalia: 259, lateral; 260, segment X dorsal; 261, Phallus lateral; 262, Phallus tip.
<taxonomicName id="C9844D36FF97FFBCFE92AA01FD8F7B35" authorityName="Malicky" authorityYear="1998" box="[278,534,1878,1899]" class="Insecta" family="Hydropsychidae" genus="Trichomacronema" kingdom="Animalia" order="Trichoptera" pageId="34" pageNumber="698" phylum="Arthropoda" rank="species" species="tamdao">
<emphasis id="3CF0EAA7FF97FFBCFE92AA01FD8F7B35" box="[278,534,1878,1899]" italics="true" pageId="34" pageNumber="698">Trichomacronema tamdao</emphasis>
</taxonomicName>
. 263, right forewing; Male genitalia: 264, lateral; 265, segment X dorsal; 266, Phallus lateral; 267, Phallus tip.
<emphasis id="3CF0EAA7FF97FFBCFE9DAA25FDCC7BD6" box="[281,597,1906,1928]" italics="true" pageId="34" pageNumber="698">
<taxonomicName id="C9844D36FF97FFBCFE9DAA25FDC97BD6" authority="Uy &amp; Malicky &amp; Bae, 2018" authorityName="Uy &amp; Malicky &amp; Bae" authorityYear="2018" box="[281,592,1906,1928]" class="Insecta" family="Hydropsychidae" genus="Trichomacronema" kingdom="Animalia" order="Trichoptera" pageId="34" pageNumber="698" phylum="Arthropoda" rank="species" species="vietnamensis" status="sp. nov.">Trichomacronema vietnamensis</taxonomicName>
,
</emphasis>
<taxonomicNameLabel id="27C357DCFF97FFBCFDD8AA24FD4D7BD6" box="[604,724,1906,1928]" pageId="34" pageNumber="698" rank="species">new species</taxonomicNameLabel>
. 268, right forewing; 269, right hind wing; Female genitalia: 270, lateral; 271, ventral. Scale: 258, 263, 268–269 = 2 mm; 259–262, 264–267, 270–271 = 0.02 mm.
</paragraph>
</caption>
<paragraph id="0E3B36B5FF96FFBDFF0CAF2AFF627F0B" blockId="35.[135,775,637,853]" pageId="35" pageNumber="699">
<emphasis id="3CF0EAA7FF96FFBDFF0CAF2AFECE7ECB" bold="true" box="[136,343,637,661]" pageId="35" pageNumber="699">Female genitalia.</emphasis>
Segment IX heavily sclerotised with row of long setae (
<figureCitation id="96BF2A30FF96FFBDFED1AFCAFE2C7EEB" box="[341,437,669,693]" captionStart="Figs" captionStartId="34.[113,157,1849,1871]" captionTargetBox="[111,1448,347,1832]" captionTargetId="figure-9@34.[97,1458,315,1842]" captionTargetPageId="34" captionText="Figs. 258–271. Trichomacronema paniae. 258, right forewing; Male genitalia: 259, lateral; 260, segment X dorsal; 261, Phallus lateral; 262, Phallus tip. Trichomacronema tamdao. 263, right forewing; Male genitalia: 264, lateral; 265, segment X dorsal; 266, Phallus lateral; 267, Phallus tip. Trichomacronema vietnamensis, new species. 268, right forewing; 269, right hind wing; Female genitalia: 270, lateral; 271, ventral. Scale: 258, 263, 268–269 = 2 mm; 259–262, 264–267, 270–271 = 0.02 mm." figureDoi="http://doi.org/10.5281/zenodo.5360961" httpUri="https://zenodo.org/record/5360961/files/figure.png" pageId="35" pageNumber="699">Fig. 270</figureCitation>
). Ventral membrane with thick walls especially at posterior portion, with shorter setation posteriorly and longer setae anteriorly (
<figureCitation id="96BF2A30FF96FFBDFDC2AF8AFD317EAB" box="[582,680,733,757]" captionStart="Figs" captionStartId="34.[113,157,1849,1871]" captionTargetBox="[111,1448,347,1832]" captionTargetId="figure-9@34.[97,1458,315,1842]" captionTargetPageId="34" captionText="Figs. 258–271. Trichomacronema paniae. 258, right forewing; Male genitalia: 259, lateral; 260, segment X dorsal; 261, Phallus lateral; 262, Phallus tip. Trichomacronema tamdao. 263, right forewing; Male genitalia: 264, lateral; 265, segment X dorsal; 266, Phallus lateral; 267, Phallus tip. Trichomacronema vietnamensis, new species. 268, right forewing; 269, right hind wing; Female genitalia: 270, lateral; 271, ventral. Scale: 258, 263, 268–269 = 2 mm; 259–262, 264–267, 270–271 = 0.02 mm." figureDoi="http://doi.org/10.5281/zenodo.5360961" httpUri="https://zenodo.org/record/5360961/files/figure.png" pageId="35" pageNumber="699">Fig. 271</figureCitation>
). Dorsal papillary lobe close to cerci; ventral papillary lobe shorter and larger with heavy setation, situated distantly from cercus (
<figureCitation id="96BF2A30FF96FFBDFF0BAE6AFF767F0B" box="[143,239,829,853]" captionStart="Figs" captionStartId="34.[113,157,1849,1871]" captionTargetBox="[111,1448,347,1832]" captionTargetId="figure-9@34.[97,1458,315,1842]" captionTargetPageId="34" captionText="Figs. 258–271. Trichomacronema paniae. 258, right forewing; Male genitalia: 259, lateral; 260, segment X dorsal; 261, Phallus lateral; 262, Phallus tip. Trichomacronema tamdao. 263, right forewing; Male genitalia: 264, lateral; 265, segment X dorsal; 266, Phallus lateral; 267, Phallus tip. Trichomacronema vietnamensis, new species. 268, right forewing; 269, right hind wing; Female genitalia: 270, lateral; 271, ventral. Scale: 258, 263, 268–269 = 2 mm; 259–262, 264–267, 270–271 = 0.02 mm." figureDoi="http://doi.org/10.5281/zenodo.5360961" httpUri="https://zenodo.org/record/5360961/files/figure.png" pageId="35" pageNumber="699">Fig. 270</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="469E653EFF96FFBDFF0CAE2AFD6A782B" pageId="35" pageNumber="699" type="materials_examined">
<paragraph id="0E3B36B5FF96FFBDFF0CAE2AFF517FAB" blockId="35.[136,775,893,1013]" pageId="35" pageNumber="699">
<emphasis id="3CF0EAA7FF96FFBDFF0CAE2AFE257FCB" bold="true" box="[136,444,893,917]" pageId="35" pageNumber="699">Type material examined.</emphasis>
<materialsCitation id="BEEC3CE8FF96FFBDFE43AE2AFF517FAB" collectingDate="2002-04-23" collectorName="D. H. Hoang &amp; Y. J. Bae" country="Vietnam" county="Sa Pa County" location="County Muong Hoa" pageId="35" pageNumber="699" specimenCount="1" stateProvince="Lao Cai" typeStatus="holotype">
<emphasis id="3CF0EAA7FF96FFBDFE43AE2AFDA57FCB" bold="true" box="[455,572,893,917]" pageId="35" pageNumber="699">
<typeStatus id="D13F8817FF96FFBDFE43AE2AFDAE7FCB" box="[455,567,893,917]" pageId="35" pageNumber="699" type="holotype">Holotype</typeStatus>
.
</emphasis>
<emphasis id="3CF0EAA7FF96FFBDFDCCAE29FD377FCB" box="[584,686,894,917]" italics="true" pageId="35" pageNumber="699">
<collectingCountry id="76937625FF96FFBDFDCCAE29FD307FCB" box="[584,681,894,917]" name="Vietnam" pageId="35" pageNumber="699">Vietnam</collectingCountry>
:
</emphasis>
female (KUEM0000002403),
<collectingRegion id="CC40F857FF96FFBDFE0EAEC9FDC07FEB" box="[394,601,925,949]" country="Vietnam" name="Lao Cai" pageId="35" pageNumber="699">Lao Cai Province</collectingRegion>
,
<collectingCounty id="E75A4E39FF96FFBDFDECAECAFC9F7FEB" box="[616,774,925,949]" pageId="35" pageNumber="699">
Sa Pa
<location id="0B5B606EFF96FFBDFD31AECAFC9F7FEB" LSID="urn:lsid:plazi:treatment:862D87A3FF94FFBDFC0DABC1FD277AAB:0B5B606EFF96FFBDFD31AECAFC9F7FEB" box="[693,774,925,949]" country="Vietnam" county="Sa Pa County" name="County Muong Hoa" pageId="35" pageNumber="699" stateProvince="Lao Cai">County</location>
</collectingCounty>
Muong Hoa, coll.
<collectorName id="A3715363FF96FFBDFED7AEE9FE417F88" box="[339,472,958,982]" pageId="35" pageNumber="699">D.H. Hoang</collectorName>
&amp;
<collectorName id="A3715363FF96FFBDFE7BAEE9FDC67F8B" box="[511,607,958,981]" pageId="35" pageNumber="699">Y.J. Bae</collectorName>
,
<date id="7A3A1075FF96FFBDFDE9AEEAFC9E7F8B" box="[621,775,957,981]" pageId="35" pageNumber="699" value="2002-04-23">
<collectingDate id="6A7EE99DFF96FFBDFDE9AEEAFC9E7F8B" box="[621,775,957,981]" pageId="35" pageNumber="699" value="2002-04-23">23 April 2002</collectingDate>
</date>
(KU).
</materialsCitation>
</paragraph>
<paragraph id="0E3B36B5FF96FFBDFF0CA94AFD6A782B" blockId="35.[136,775,1053,1142]" pageId="35" pageNumber="699">
<materialsCitation id="BEEC3CE8FF96FFBDFF0CA94AFD76782B" collectingDate="2002-04-23" collectorName="Sa Pa County Muong Hoa &amp; D. H. Hoang &amp; Y. J. Bae" country="Vietnam" county="Sa Pa County" location="County Muong Hoa" pageId="35" pageNumber="699" specimenCount="3" specimenCount-female="3" stateProvince="Lao Cai" typeStatus="paratype">
<emphasis id="3CF0EAA7FF96FFBDFF0CA94AFE92786B" bold="true" box="[136,267,1053,1077]" pageId="35" pageNumber="699">
<typeStatus id="D13F8817FF96FFBDFF0CA94AFE9F786B" box="[136,262,1053,1077]" pageId="35" pageNumber="699" type="paratype">Paratypes</typeStatus>
.
</emphasis>
<emphasis id="3CF0EAA7FF96FFBDFE93A949FE18786B" box="[279,385,1054,1077]" italics="true" pageId="35" pageNumber="699">
<collectingCountry id="76937625FF96FFBDFE93A949FEE3786B" box="[279,378,1054,1077]" name="Vietnam" pageId="35" pageNumber="699">Vietnam</collectingCountry>
:
</emphasis>
<specimenCount id="1882FD3CFF96FFBDFE09A94AFD99786B" box="[397,512,1053,1077]" pageId="35" pageNumber="699" type="female">3 females</specimenCount>
(KUEM0000002404– KUEM0000002406),
<collectingRegion id="CC40F857FF96FFBDFEE8A969FDBB780B" box="[364,546,1085,1109]" country="Vietnam" name="Lao Cai" pageId="35" pageNumber="699">Lao Cai Province</collectingRegion>
,
<collectorName id="A3715363FF96FFBDFDAFA96AFF2E782B" pageId="35" pageNumber="699">
<collectingCounty id="E75A4E39FF96FFBDFDAFA96AFD2F780B" box="[555,694,1085,1109]" pageId="35" pageNumber="699">
Sa Pa
<location id="0B5B606EFF96FFBDFDEDA96AFD2F780B" LSID="urn:lsid:plazi:treatment:862D87A3FF94FFBDFC0DABC1FD277AAB:0B5B606EFF96FFBDFDEDA96AFD2F780B" box="[617,694,1085,1109]" country="Vietnam" county="Sa Pa County" name="County Muong Hoa" pageId="35" pageNumber="699" stateProvince="Lao Cai">County</location>
</collectingCounty>
Muong Hoa
</collectorName>
, coll.
<collectorName id="A3715363FF96FFBDFF7FA909FEE77828" box="[251,382,1118,1142]" pageId="35" pageNumber="699">D.H. Hoang</collectorName>
&amp;
<collectorName id="A3715363FF96FFBDFE21A909FD9D782B" box="[421,516,1118,1141]" pageId="35" pageNumber="699">Y.J. Bae</collectorName>
,
<date id="7A3A1075FF96FFBDFD95A90AFD33782B" box="[529,682,1117,1141]" pageId="35" pageNumber="699" value="2002-04-23">
<collectingDate id="6A7EE99DFF96FFBDFD95A90AFD33782B" box="[529,682,1117,1141]" pageId="35" pageNumber="699" value="2002-04-23">23 April 2002</collectingDate>
</date>
(KU)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="469E653EFF96FFBDFF0CA9CAFE1F78EB" box="[136,390,1181,1205]" pageId="35" pageNumber="699" type="distribution">
<paragraph id="0E3B36B5FF96FFBDFF0CA9CAFE1F78EB" blockId="35.[136,390,1181,1205]" box="[136,390,1181,1205]" pageId="35" pageNumber="699">
<emphasis id="3CF0EAA7FF96FFBDFF0CA9CAFE8378EB" bold="true" box="[136,282,1181,1205]" pageId="35" pageNumber="699">Distribution.</emphasis>
<collectingCountry id="76937625FF96FFBDFEA7A9CAFE1B78EB" box="[291,386,1181,1205]" name="Vietnam" pageId="35" pageNumber="699">Vietnam</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="469E653EFF96FFBDFF0CA98AFE72794B" pageId="35" pageNumber="699" type="etymology">
<paragraph id="0E3B36B5FF96FFBDFF0CA98AFE72794B" blockId="35.[136,775,1245,1301]" pageId="35" pageNumber="699">
<emphasis id="3CF0EAA7FF96FFBDFF0CA98AFE9778AB" bold="true" box="[136,270,1245,1269]" pageId="35" pageNumber="699">Etymology.</emphasis>
The species epithet refers to
<collectingCountry id="76937625FF96FFBDFDECA98AFD5F78AB" box="[616,710,1245,1269]" name="Vietnam" pageId="35" pageNumber="699">Vietnam</collectingCountry>
from which the species was collected.
</paragraph>
</subSubSection>
<subSubSection id="469E653EFF96FFBDFF0CA86AFD277AAB" pageId="35" pageNumber="699" type="discussion">
<paragraph id="0E3B36B5FF96FFBDFF0CA86AFD277AAB" blockId="35.[135,776,1341,1781]" pageId="35" pageNumber="699">
<emphasis id="3CF0EAA7FF96FFBDFF0CA86AFF63790B" bold="true" box="[136,250,1341,1365]" pageId="35" pageNumber="699">Remarks.</emphasis>
This species is obviously distinct from other congeners. Its wing structure is similar to that of
<taxonomicName id="C9844D36FF96FFBDFD05A809FEB979CB" authority="Schmid, 1964" authorityName="Schmid" authorityYear="1964" class="Insecta" family="Hydropsychidae" genus="Trichomacronema" kingdom="Animalia" order="Trichoptera" pageId="35" pageNumber="699" phylum="Arthropoda" rank="species" species="shanorum">
<emphasis id="3CF0EAA7FF96FFBDFD05A809FC9C792B" box="[641,773,1373,1397]" italics="true" pageId="35" pageNumber="699">T. shanorum</emphasis>
<bibRefCitation id="6A154B44FF96FFBDFF03A82AFEB979CB" author="Schmid F" box="[135,288,1405,1429]" pageId="35" pageNumber="699" pagination="825 - 840" refId="ref27405" refString="Schmid F (1964) Quelques Trichopteres Asiatiques. Canadian Entomologist, 96: 825 - 840." type="journal article" year="1964">Schmid, 1964</bibRefCitation>
</taxonomicName>
, but its wing pattern is different. The radius and median forewing cells of
<taxonomicName id="C9844D36FF96FFBDFE5EA8C9FDFF79EB" authorityName="Schmid" authorityYear="1964" box="[474,614,1437,1461]" class="Insecta" family="Hydropsychidae" genus="Trichomacronema" kingdom="Animalia" order="Trichoptera" pageId="35" pageNumber="699" phylum="Arthropoda" rank="species" species="shanorum">
<emphasis id="3CF0EAA7FF96FFBDFE5EA8C9FDFF79EB" box="[474,614,1437,1461]" italics="true" pageId="35" pageNumber="699">T. shanorum</emphasis>
</taxonomicName>
possess white spots, whereas those of
<taxonomicName id="C9844D36FF96FFBDFE03A8E9FDB4798B" authorityName="Uy &amp; Malicky &amp; Bae" authorityYear="2018" box="[391,557,1470,1493]" class="Insecta" family="Hydropsychidae" genus="Trichomacronema" kingdom="Animalia" order="Trichoptera" pageId="35" pageNumber="699" phylum="Arthropoda" rank="species" species="vietnamensis">
<emphasis id="3CF0EAA7FF96FFBDFE03A8E9FDB4798B" box="[391,557,1470,1493]" italics="true" pageId="35" pageNumber="699">T. vietnamensis</emphasis>
</taxonomicName>
do not. The body of
<taxonomicName id="C9844D36FF96FFBDFF0CA889FE8879AB" authorityName="Schmid" authorityYear="1964" box="[136,273,1501,1525]" class="Insecta" family="Hydropsychidae" genus="Trichomacronema" kingdom="Animalia" order="Trichoptera" pageId="35" pageNumber="699" phylum="Arthropoda" rank="species" species="shanorum">
<emphasis id="3CF0EAA7FF96FFBDFF0CA889FE8879AB" box="[136,273,1501,1525]" italics="true" pageId="35" pageNumber="699">T. shanorum</emphasis>
</taxonomicName>
also possesses brown and dark-brown stripes, whereas that of
<taxonomicName id="C9844D36FF96FFBDFEC6A8A9FE687A4B" authorityName="Uy &amp; Malicky &amp; Bae" authorityYear="2018" box="[322,497,1534,1557]" class="Insecta" family="Hydropsychidae" genus="Trichomacronema" kingdom="Animalia" order="Trichoptera" pageId="35" pageNumber="699" phylum="Arthropoda" rank="species" species="vietnamensis">
<emphasis id="3CF0EAA7FF96FFBDFEC6A8A9FE687A4B" box="[322,497,1534,1557]" italics="true" pageId="35" pageNumber="699">T. vietnamensis</emphasis>
</taxonomicName>
is uniformly brown. As compared to the two species described from Southeast Asia,
<taxonomicName id="C9844D36FF96FFBDFF0CAB69FF6E7A0B" authorityName="Malicky" authorityYear="1998" box="[136,247,1597,1621]" class="Insecta" family="Hydropsychidae" genus="Trichomacronema" kingdom="Animalia" order="Trichoptera" pageId="35" pageNumber="699" phylum="Arthropoda" rank="species" species="tamdao">
<emphasis id="3CF0EAA7FF96FFBDFF0CAB69FF6E7A0B" box="[136,247,1597,1621]" italics="true" pageId="35" pageNumber="699">T. tamdao</emphasis>
</taxonomicName>
and
<emphasis id="3CF0EAA7FF96FFBDFEB6AB69FE397A0B" box="[306,416,1598,1621]" italics="true" pageId="35" pageNumber="699">
<taxonomicName id="C9844D36FF96FFBDFEB6AB69FE057A0B" authorityName="Malicky &amp; Chantaramongkol" authorityYear="1991" box="[306,412,1598,1621]" class="Insecta" family="Hydropsychidae" genus="Trichomacronema" kingdom="Animalia" order="Trichoptera" pageId="35" pageNumber="699" phylum="Arthropoda" rank="species" species="paniae">T. paniae</taxonomicName>
,
</emphasis>
this new species is much larger (whole body size was measured longitudinally from the head to the end of the forewing). Body size of
<taxonomicName id="C9844D36FF96FFBDFDCAAB29FD257ACB" authorityName="Malicky" authorityYear="1998" box="[590,700,1661,1685]" class="Insecta" family="Hydropsychidae" genus="Trichomacronema" kingdom="Animalia" order="Trichoptera" pageId="35" pageNumber="699" phylum="Arthropoda" rank="species" species="tamdao">
<emphasis id="3CF0EAA7FF96FFBDFDCAAB29FD257ACB" box="[590,700,1661,1685]" italics="true" pageId="35" pageNumber="699">T. tamdao</emphasis>
</taxonomicName>
and
<taxonomicName id="C9844D36FF96FFBDFD75AB29FF487AEB" authorityName="Malicky &amp; Chantaramongkol" authorityYear="1991" class="Insecta" family="Hydropsychidae" genus="Trichomacronema" kingdom="Animalia" order="Trichoptera" pageId="35" pageNumber="699" phylum="Arthropoda" rank="species" species="paniae">
<emphasis id="3CF0EAA7FF96FFBDFD75AB29FF487AEB" italics="true" pageId="35" pageNumber="699">T. paniae</emphasis>
</taxonomicName>
are just half the size of
<emphasis id="3CF0EAA7FF96FFBDFE6CABC9FD007AEB" box="[488,665,1694,1717]" italics="true" pageId="35" pageNumber="699">
<taxonomicName id="C9844D36FF96FFBDFE6CABC9FD0F7AEB" authorityName="Uy &amp; Malicky &amp; Bae" authorityYear="2018" box="[488,662,1694,1717]" class="Insecta" family="Hydropsychidae" genus="Trichomacronema" kingdom="Animalia" order="Trichoptera" pageId="35" pageNumber="699" phylum="Arthropoda" rank="species" species="vietnamensis">T. vietnamensis</taxonomicName>
.
</emphasis>
At a first glance,
<taxonomicName id="C9844D36FF96FFBDFF58ABE9FE1B7A8B" authorityName="Uy &amp; Malicky &amp; Bae" authorityYear="2018" box="[220,386,1726,1749]" class="Insecta" family="Hydropsychidae" genus="Trichomacronema" kingdom="Animalia" order="Trichoptera" pageId="35" pageNumber="699" phylum="Arthropoda" rank="species" species="vietnamensis">
<emphasis id="3CF0EAA7FF96FFBDFF58ABE9FE1B7A8B" box="[220,386,1726,1749]" italics="true" pageId="35" pageNumber="699">T. vietnamensis</emphasis>
</taxonomicName>
can be mistaken of a
<taxonomicName id="C9844D36FF96FFBDFDEBABE9FF4C7AAB" class="Insecta" family="Hydropsychidae" genus="Macrostemum" kingdom="Animalia" order="Trichoptera" pageId="35" pageNumber="699" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="3CF0EAA7FF96FFBDFDEBABE9FC917A8B" box="[623,776,1726,1749]" italics="true" pageId="35" pageNumber="699">Macrostemum</emphasis>
species
</taxonomicName>
because of the structure of head and thorax.
</paragraph>
</subSubSection>
</treatment>
</document>
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