treatments-xml/data/46/1B/A2/461BA2D26D18529E95FD84B0FB3156D6.xml

<document ID-DOI="http://dx.doi.org/10.3897/zookeys.914.46652" ID-GBIF-Dataset="2fcb00c0-aa8e-48ef-892e-32f99adda476" ID-PMC="PMC7046710" ID-Pensoft-Pub="1313-2970-914-81" ID-Pensoft-UUID="B946CC658E5E5292A35787308816FFF0" ID-PubMed="32132856" ID-ZooBank="91B11F13EF554E71B0D64491942912DC" ModsDocID="1313-2970-914-81" checkinTime="1582245055288" checkinUser="pensoft" docAuthor="Kluge, Nikita J., Godunko, Roman J. &amp; Svitok, Marek" docDate="2020" docId="461BA2D26D18529E95FD84B0FB3156D6" docLanguage="en" docName="ZooKeys 914: 81-125" docOrigin="ZooKeys 914" docSource="http://dx.doi.org/10.3897/zookeys.914.46652" docTitle="Centroptella Braasch &amp; Soldan 1980" docType="treatment" docVersion="4" id="B946CC658E5E5292A35787308816FFF0" lastPageNumber="81" masterDocId="B946CC658E5E5292A35787308816FFF0" masterDocTitle="Nomenclatural changes in Centroptella Braasch &amp; Soldan, 1980 (Ephemeroptera, Baetidae)" masterLastPageNumber="125" masterPageNumber="81" pageNumber="81" updateTime="1668168509386" updateUser="ExternalLinkService">
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<mods:title>Nomenclatural changes in Centroptella Braasch &amp; Soldan, 1980 (Ephemeroptera, Baetidae)</mods:title>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Kluge, Nikita J.</mods:namePart>
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<mods:namePart>Godunko, Roman J.</mods:namePart>
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<mods:namePart>Svitok, Marek</mods:namePart>
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<mods:date>2020</mods:date>
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<mods:number>914</mods:number>
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<treatment ID-GBIF-Taxon="162244680" LSID="urn:lsid:plazi:treatment:461BA2D26D18529E95FD84B0FB3156D6" httpUri="http://treatment.plazi.org/id/461BA2D26D18529E95FD84B0FB3156D6" lastPageNumber="81" pageId="0" pageNumber="81">
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<taxonomicName LSID="461BA2D2-6D18-529E-95FD-84B0FB3156D6" authority="Braasch &amp; Soldan, 1980" authorityName="Braasch &amp; Soldan" authorityYear="1980" class="Insecta" family="Baetidae" genus="Centroptella" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Centroptella" order="Ephemeroptera" pageId="0" pageNumber="81" phylum="Arthropoda" rank="genus">
Centroptella Braasch &amp; 
<normalizedToken originalValue="Soldán">Soldan</normalizedToken>
, 1980
</taxonomicName>
<figureCitation captionStart="Figures 1–15" captionStartId="F1" captionText="Figures 1 - 15. Australian Centroptella. 1 - 7 C. illiesi, tergalii I-VII 8 - 14 C. fustipalpus (paratype), tergalii I-VII 15 C. fustipalpus (holotype), right mandible." figureDoi="10.3897/zookeys.914.46652.figures1-15" httpUri="https://binary.pensoft.net/fig/381996" pageId="0" pageNumber="81">Figs 1-15</figureCitation>
<figureCitation captionStart="Figures 16–18" captionStartId="F2" captionText="Figures 16 - 18. Centroptella illiesi, tibiae of fore, middle and hind legs, view from anterior side (bases of long setae shown both on anterior and posterior sides)." figureDoi="10.3897/zookeys.914.46652.figures16-18" httpUri="https://binary.pensoft.net/fig/381997" pageId="0" pageNumber="81">, 16-18</figureCitation>
<figureCitation captionStart="Figures 19–39" captionStartId="F3" captionText="Figures 19 - 39. Asian Centroptella. 19 - 32 C. longisetosa: (19 - 24) tergalii I-VI of paratype of C. longisetosa (25) the same, apex costal rib of tergalius IV (26 - 32) tergalii I-VII of neotype of C. liebenauae (actual C. longisetosa) 33 - 39 C. ingridae, tergalii I-VII of sp. nov." figureDoi="10.3897/zookeys.914.46652.figures19-39" httpUri="https://binary.pensoft.net/fig/381998" pageId="0" pageNumber="81">, 19-39</figureCitation>
<figureCitation captionStart="Figures 40–48" captionStartId="F4" captionText="Figures 40 - 48. Centroptella longisetosa. 40 - 46 paratype of C. longisetosa: (40, 41) left and right mandibles (42) apex of maxilla (ds 1, ds 2, ds 3 dentisetae) (43) labrum (44) glossa and paraglossa (ventral view) (45) labium (at left ventral view, at right dorsal view; muscles shown by interrupted lines) (46) maxillary palp 47 the same, neotype of C. liebenauae (actually C. longisetosa) 48 the same, specimen from India." figureDoi="10.3897/zookeys.914.46652.figures40-48" httpUri="https://binary.pensoft.net/fig/381999" pageId="0" pageNumber="81">, 40-48</figureCitation>
<figureCitation captionStart="Figures 49–52" captionStartId="F5" captionText="Figures 49 - 52. Centroptella longisetosa. 49 - 51 paratype of C. longisetosa: tibiae of fore, middle and hind legs, view from anterior side (bases of long setae shown both on anterior and posterior sides) 52 neotype of C. liebenauae (actually C. longisetosa): tenth abdominal segment without caudalii, ventral view." figureDoi="10.3897/zookeys.914.46652.figures49-52" httpUri="https://binary.pensoft.net/fig/382000" pageId="0" pageNumber="81">, 49-52</figureCitation>
<figureCitation captionStart="Figures 53–59" captionStartId="F6" captionText="Figures 53 - 59. Larval exuviae of Centroptella longisetosa (neotype of C. liebenauae). 53 - 57 at the same magnification: (53) left half of pro- and mesonotum (54) thoracic pleura, left half of metanotum, fore- and hind legs (55) frons, antenna and labrum (56) tenth uromere and caudalii; (57) abdominal sterna and terga I-IX 58 margins of abdominal sterna IV-VII 59 cercus." figureDoi="10.3897/zookeys.914.46652.figures53-59" httpUri="https://binary.pensoft.net/fig/382001" pageId="0" pageNumber="81">, 53-59</figureCitation>
<figureCitation captionStart="Figures 60–76" captionStartId="F7" captionText="Figures 60 - 76. Larval exuviae of Centroptella longisetosa. 60 - 75 female, paratype of C. longisetosa: (60 - 68) fragments of abdominal terga II-IX (indicated by Roman numbers) (69) sternum VI (70 - 75) fragments of abdominal sterna IV-IX (indicated by Roman numbers) 76 male, neotype of C. liebenauae: fragment of abdominal sternum IX. Arrows on Figs 68 and 76 show median line." figureDoi="10.3897/zookeys.914.46652.figures60-76" httpUri="https://binary.pensoft.net/fig/382002" pageId="0" pageNumber="81">, 60-76</figureCitation>
<figureCitation captionStart="Figures 77–80" captionStartId="F8" captionText="Figures 77 - 80. Male genitalia of Centroptella longisetosa. 77 - 79 specimens from India: (77) genitalia of imago (78) subimaginal exuviae (79) subimaginal gonostyli crumpled under larval cuticle (80) paratype of C. liebenauae (possibly reared from neotype): genitalia of imago." figureDoi="10.3897/zookeys.914.46652.figures77-80" httpUri="https://binary.pensoft.net/fig/382003" pageId="0" pageNumber="81">, 77-80</figureCitation>
<figureCitation captionStart="Figures 81–82" captionStartId="F9" captionText="Figures 81 - 82. Egg extracted from larva - paratype of Centroptella longisetosa." figureDoi="10.3897/zookeys.914.46652.figures81-82" httpUri="https://binary.pensoft.net/fig/382004" pageId="0" pageNumber="81">, 81-82</figureCitation>
<figureCitation captionStart="Figures 83–89" captionStartId="F10" captionText="Figures 83 - 89. Centroptella ingridae sp. nov., larvae. 83 - 85 specimens from Thailand 86 - 89 paratypes of C. liebenauae (actually C. ingridae sp. nov.). Arrows show pair of protuberances on pronotum." figureDoi="10.3897/zookeys.914.46652.figures83-89" httpUri="https://binary.pensoft.net/fig/382005" pageId="0" pageNumber="81">, 83-89</figureCitation>
<figureCitation captionStart="Figures 90–93" captionStartId="F11" captionText="Figures 90 - 93. Centroptella ingridae sp. nov. (holotype). 90 - 92 tibia of fore, middle and hind leg, view from anterior side (bases of long setae shown both on anterior and posterior sides) 93 tenth abdominal segment without caudalii, ventral view." figureDoi="10.3897/zookeys.914.46652.figures90-93" httpUri="https://binary.pensoft.net/fig/382006" pageId="0" pageNumber="81">, 90-93</figureCitation>
<figureCitation captionStart="Figures 94–109" captionStartId="F12" captionText="Figures 94 - 109. Centroptella ingridae sp. nov. 94 - 102 fragments of abdominal terga I-IX (indicated by Roman numbers) 103 - 108 fragments of abdominal sterna IV-IX of male (indicated by Roman numbers) 109 fragment of abdominal sternum IX of female (94 - 101 and 103 - 108 male paratype of C. liebenauae; 102 and 109 female specimen from Thailand)." figureDoi="10.3897/zookeys.914.46652.figures94-109" httpUri="https://binary.pensoft.net/fig/382007" pageId="0" pageNumber="81">, 94-109</figureCitation>
<figureCitation captionStart="Figures 110–115" captionStartId="F13" captionText="Figures 110 - 115. Centroptella ingridae sp. nov. (paratype of C. liebenauae), SEM photos of larva. 110 clypeus 111 pronotum 112 - 113 abdominal tergum I 114 abdominal tergum III 115 cercus." figureDoi="10.3897/zookeys.914.46652.figures110-115" httpUri="https://binary.pensoft.net/fig/382008" pageId="0" pageNumber="81">, 110-115</figureCitation>
<figureCitation captionStart="Figures 116–122" captionStartId="F14" captionText="Figures 116 - 122. Centroptella ingridae sp. nov. (paratype of C. liebenauae), SEM photos of larva. 116 labrum 117 bifurcate setae on abdominal sternum IV 118 - 119 abdominal sterna IV-VI 120 - 121 paraprocts 122 flagellum of antenna." figureDoi="10.3897/zookeys.914.46652.figures116-122" httpUri="https://binary.pensoft.net/fig/382009" pageId="0" pageNumber="81">, 116-122</figureCitation>
<figureCitation captionStart="Figures 123–128" captionStartId="F15" captionText="Figures 123 - 128. Centroptella ingridae sp. nov. (specimens from Thailand). 123 - 124 labium 125 tergalius IV (cr costal rib; mr middle rib; ar anal rib) 126 - 127 larval claws; 128 egg." figureDoi="10.3897/zookeys.914.46652.figures123-128" httpUri="https://binary.pensoft.net/fig/382010" pageId="0" pageNumber="81">, 123-128</figureCitation>
<figureCitation captionStart="Figures 129–132" captionStartId="F16" captionText="Figures 129 - 132. Centroptella ingridae sp. nov. (specimen from Thailand), exuviae of larval cercus (lateral view). 129 - 131 focus on lateral side to show bases of secondary swimming setae; 132 focus on median side to show bases of primary swimming setae." figureDoi="10.3897/zookeys.914.46652.figures129-132" httpUri="https://binary.pensoft.net/fig/382011" pageId="0" pageNumber="81">, 129-132</figureCitation>
<figureCitation captionStart="Figures 133–137" captionStartId="F17" captionText="Figures 133 - 137. Centroptella ingridae sp. nov. (specimens from Thailand). 133 - 134 female imago 135 - 137 holotype (male): (135) head and thorax of imago (136) imaginal middle leg (137) subimaginal exuviae of tarsus." figureDoi="10.3897/zookeys.914.46652.figures133-137" httpUri="https://binary.pensoft.net/fig/382012" pageId="0" pageNumber="81">, 133-137</figureCitation>
<figureCitation captionStart="Figures 138–143" captionStartId="F18" captionText="Figures 138 - 143. Centroptella ingridae sp. nov. (specimens from Thailand). 138 abdominal sterna and terga of female imago 139 the same, male imago 140 - 143 genitalia of male imago (139 - 141 holotype)." figureDoi="10.3897/zookeys.914.46652.figures138-143" httpUri="https://binary.pensoft.net/fig/382013" pageId="0" pageNumber="81">, 138-143</figureCitation>
<figureCitation captionStart="Figures 144–148" captionStartId="F19" captionText="Figures 144 - 148. Centroptella ingridae sp. nov. (specimens from Thailand). 144 - 145 left and right mandibles 146 genitalia of male imago 147 their subimaginal exuviae 148 subimaginal gonostyli crumpled under larval cuticle (146 - 147 holotype)." figureDoi="10.3897/zookeys.914.46652.figures144-148" httpUri="https://binary.pensoft.net/fig/382014" pageId="0" pageNumber="81">, 144-148</figureCitation>
<figureCitation captionStart="Figures 149–152" captionStartId="F20" captionText="Figures 149 - 152. Centroptella of group inzingae - ingridae. Centroptella ingridae sp. nov. (holotype), fore wing 150 Centroptella inzingae (lectotype), genitalia (gv gonovectis us proximal margin of unistyliger) 151 - 152 Centroptella saxophila, gonovectes (specimens reared from larvae by N. Kluge in the Western Cape Province of South Africa)." figureDoi="10.3897/zookeys.914.46652.figures149-152" httpUri="https://binary.pensoft.net/fig/382015" pageId="0" pageNumber="81">, 149-152</figureCitation>
<figureCitation captionStart="Figure 153" captionStartId="F21" captionText="Figure 153. Centroptella colorata (holotype), last instar larva." figureDoi="10.3897/zookeys.914.46652.figure153" httpUri="https://binary.pensoft.net/fig/382016" pageId="0" pageNumber="81">, 153</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="81" type="reference_group">
<paragraph pageId="0" pageNumber="81">
= 
<taxonomicName authorityName="Waltz &amp; McCafferty" authorityYear="1987" class="Insecta" family="Baetidae" genus="Chopralla" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chopralla" order="Ephemeroptera" pageId="0" pageNumber="81" phylum="Arthropoda" rank="genus">Chopralla</taxonomicName>
Waltz &amp; McCafferty, 1987: 182, syn. nov.
</paragraph>
<paragraph pageId="0" pageNumber="81">
= 
<taxonomicName authorityName="Salles, Gattolliat &amp; Sartori" authorityYear="2015" class="Insecta" family="Baetidae" genus="Crassolus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Crassolus" order="Ephemeroptera" pageId="0" pageNumber="81" phylum="Arthropoda" rank="genus">Crassolus</taxonomicName>
Salles, Gattolliat &amp; Sartori, 2016: 104, syn. nov.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="81" type="type species">
<paragraph pageId="0" pageNumber="81">Type species.</paragraph>
<paragraph pageId="0" pageNumber="81">
<taxonomicName authorityName="Braasch &amp; Soldan" authorityYear="1980" class="Insecta" family="Baetidae" genus="Centroptella" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Centroptella longisetosa" order="Ephemeroptera" pageId="0" pageNumber="81" phylum="Arthropoda" rank="species" species="longisetosa">
<emphasis italics="true" pageId="0" pageNumber="81">Centroptella longisetosa</emphasis>
</taxonomicName>
Braasch &amp; 
<normalizedToken originalValue="Soldán">Soldan</normalizedToken>
, 1980.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="81" type="description">
<paragraph pageId="0" pageNumber="81">Systematic position and characters.</paragraph>
<paragraph pageId="0" pageNumber="81">
<taxonomicName authorityName="Braasch &amp; Soldan" authorityYear="1980" class="Insecta" family="Baetidae" genus="Centroptella" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Centroptella" order="Ephemeroptera" pageId="0" pageNumber="81" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="81">Centroptella</emphasis>
</taxonomicName>
is characterized by an unusual combination of characters: on one hand, it undoubtedly belongs to the holophyletic taxon Baetovectata Kluge &amp; Novikova, 2011, based on (1) presence of two marginal intercalaries in each space of wing (Fig. 
<figureCitation captionStart="Figures 149–152" captionStartId="F20" captionText="Figures 149 - 152. Centroptella of group inzingae - ingridae. Centroptella ingridae sp. nov. (holotype), fore wing 150 Centroptella inzingae (lectotype), genitalia (gv gonovectis us proximal margin of unistyliger) 151 - 152 Centroptella saxophila, gonovectes (specimens reared from larvae by N. Kluge in the Western Cape Province of South Africa)." figureDoi="10.3897/zookeys.914.46652.figures149-152" httpUri="https://binary.pensoft.net/fig/382015" pageId="0" pageNumber="81">149</figureCitation>
), (2) narrow and arched gonovectes of the penis (Figs 
<figureCitation captionStart="Figures 77–80" captionStartId="F8" captionText="Figures 77 - 80. Male genitalia of Centroptella longisetosa. 77 - 79 specimens from India: (77) genitalia of imago (78) subimaginal exuviae (79) subimaginal gonostyli crumpled under larval cuticle (80) paratype of C. liebenauae (possibly reared from neotype): genitalia of imago." figureDoi="10.3897/zookeys.914.46652.figures77-80" httpUri="https://binary.pensoft.net/fig/382003" pageId="0" pageNumber="81">77-80</figureCitation>
, 
<figureCitation captionStart="Figures 138–143" captionStartId="F18" captionText="Figures 138 - 143. Centroptella ingridae sp. nov. (specimens from Thailand). 138 abdominal sterna and terga of female imago 139 the same, male imago 140 - 143 genitalia of male imago (139 - 141 holotype)." figureDoi="10.3897/zookeys.914.46652.figures138-143" httpUri="https://binary.pensoft.net/fig/382013" pageId="0" pageNumber="81">141-143</figureCitation>
, 
<figureCitation captionStart="Figures 144–148" captionStartId="F19" captionText="Figures 144 - 148. Centroptella ingridae sp. nov. (specimens from Thailand). 144 - 145 left and right mandibles 146 genitalia of male imago 147 their subimaginal exuviae 148 subimaginal gonostyli crumpled under larval cuticle (146 - 147 holotype)." figureDoi="10.3897/zookeys.914.46652.figures144-148" httpUri="https://binary.pensoft.net/fig/382014" pageId="0" pageNumber="81">146</figureCitation>
, 150152) and (3) medially inclined subimaginal gonostyli when they are developing under the larval cuticle (Figs 
<figureCitation captionStart="Figures 77–80" captionStartId="F8" captionText="Figures 77 - 80. Male genitalia of Centroptella longisetosa. 77 - 79 specimens from India: (77) genitalia of imago (78) subimaginal exuviae (79) subimaginal gonostyli crumpled under larval cuticle (80) paratype of C. liebenauae (possibly reared from neotype): genitalia of imago." figureDoi="10.3897/zookeys.914.46652.figures77-80" httpUri="https://binary.pensoft.net/fig/382003" pageId="0" pageNumber="81">79</figureCitation>
, 
<figureCitation captionStart="Figures 144–148" captionStartId="F19" captionText="Figures 144 - 148. Centroptella ingridae sp. nov. (specimens from Thailand). 144 - 145 left and right mandibles 146 genitalia of male imago 147 their subimaginal exuviae 148 subimaginal gonostyli crumpled under larval cuticle (146 - 147 holotype)." figureDoi="10.3897/zookeys.914.46652.figures144-148" httpUri="https://binary.pensoft.net/fig/382014" pageId="0" pageNumber="81">148</figureCitation>
). The taxon Baetovectata belongs to the holophyletic taxon Anteropatellata Kluge, 1997, which is characterized by the presence of a patella-tibial suture on the forelegs of the larva and female imago and subimago. On the other hand, the leg structure of 
<taxonomicName authorityName="Braasch &amp; Soldan" authorityYear="1980" class="Insecta" family="Baetidae" genus="Centroptella" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Centroptella" order="Ephemeroptera" pageId="0" pageNumber="81" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="81">Centroptella</emphasis>
</taxonomicName>
does not conform with the characteristics of Anteropatellata. The larva of 
<taxonomicName authorityName="Braasch &amp; Soldan" authorityYear="1980" class="Insecta" family="Baetidae" genus="Centroptella" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Centroptella" order="Ephemeroptera" pageId="0" pageNumber="81" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="81">Centroptella</emphasis>
</taxonomicName>
has the structure of the tibia modified and different on each pair of legs, so that the patella-tibial suture is absent on forelegs and greatly shifted distally on the middle and hind legs; a row of long setae, which in some other taxa forms a transverse arc, in 
<taxonomicName authorityName="Braasch &amp; Soldan" authorityYear="1980" class="Insecta" family="Baetidae" genus="Centroptella" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Centroptella" order="Ephemeroptera" pageId="0" pageNumber="81" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="81">Centroptella</emphasis>
</taxonomicName>
is greatly stretched along the tibia, being different on the fore, middle and hind legs (Figs 
<figureCitation captionStart="Figures 16–18" captionStartId="F2" captionText="Figures 16 - 18. Centroptella illiesi, tibiae of fore, middle and hind legs, view from anterior side (bases of long setae shown both on anterior and posterior sides)." figureDoi="10.3897/zookeys.914.46652.figures16-18" httpUri="https://binary.pensoft.net/fig/381997" pageId="0" pageNumber="81">16-18</figureCitation>
, 
<figureCitation captionStart="Figures 49–52" captionStartId="F5" captionText="Figures 49 - 52. Centroptella longisetosa. 49 - 51 paratype of C. longisetosa: tibiae of fore, middle and hind legs, view from anterior side (bases of long setae shown both on anterior and posterior sides) 52 neotype of C. liebenauae (actually C. longisetosa): tenth abdominal segment without caudalii, ventral view." figureDoi="10.3897/zookeys.914.46652.figures49-52" httpUri="https://binary.pensoft.net/fig/382000" pageId="0" pageNumber="81">49-51</figureCitation>
, 
<figureCitation captionStart="Figures 90–93" captionStartId="F11" captionText="Figures 90 - 93. Centroptella ingridae sp. nov. (holotype). 90 - 92 tibia of fore, middle and hind leg, view from anterior side (bases of long setae shown both on anterior and posterior sides) 93 tenth abdominal segment without caudalii, ventral view." figureDoi="10.3897/zookeys.914.46652.figures90-93" httpUri="https://binary.pensoft.net/fig/382006" pageId="0" pageNumber="81">90-92</figureCitation>
); the female imago and subimago of 
<taxonomicName authorityName="Braasch &amp; Soldan" authorityYear="1980" class="Insecta" family="Baetidae" genus="Centroptella" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Centroptella" order="Ephemeroptera" pageId="0" pageNumber="81" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="81">Centroptella</emphasis>
</taxonomicName>
has the usual leg structure, with the patella-tibial suture not shifted distally, but without patella-tibial suture on forelegs. This leg structure is characteristic of the plesiomorphon Protopatellata Kluge &amp; Novikova, 2011 and has striking similarity with the Afrotropical taxon 
<taxonomicName authorityName="Gillies" authorityYear="1990" class="Insecta" family="Baetidae" genus="Potamocloeon" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Potamocloeon" order="Ephemeroptera" pageId="0" pageNumber="81" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="81">Potamocloeon</emphasis>
</taxonomicName>
Gillies, 1990 (= 
<taxonomicName authorityName="Lugo-Ortiz &amp; McCafferty" authorityYear="1997" class="Insecta" family="Baetidae" genus="Maliqua" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Maliqua" order="Ephemeroptera" pageId="0" pageNumber="81" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="81">Maliqua</emphasis>
</taxonomicName>
Lugo-Ortiz &amp; McCafferty, 1997), which undoubtedly belongs to Protopatellata and has no features of Baetovectata (
<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.4648.2.6" author="Kluge, NJ" journalOrPublisher="Zootaxa" pageId="0" pageNumber="81" pagination="299 - 317" refId="B15" refString="Kluge, NJ, 2019. Potamocloeon edentatum sp. nov. from Usambara Mountains in Tanzania (Ephemeroptera, Baetidae). Zootaxa 4648 (2): 299 - 317, DOI: https://doi.org/10.11646/zootaxa.4648.2.6" title="Potamocloeon edentatum sp. nov. from Usambara Mountains in Tanzania (Ephemeroptera, Baetidae)." url="https://doi.org/10.11646/zootaxa.4648.2.6" volume="4648" year="2019">Kluge 2019</bibRefCitation>
). The Neotropical genus 
<taxonomicName authorityName="Traver" authorityYear="1938" class="Insecta" family="Baetidae" genus="Cloeodes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cloeodes" order="Ephemeroptera" pageId="0" pageNumber="81" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="81">Cloeodes</emphasis>
</taxonomicName>
Traver, 1938, which some authors have confused with 
<taxonomicName authorityName="Braasch &amp; Soldan" authorityYear="1980" class="Insecta" family="Baetidae" genus="Centroptella" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Centroptella" order="Ephemeroptera" pageId="0" pageNumber="81" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="81">Centroptella</emphasis>
</taxonomicName>
(see above), has none of these features, and its larval and imaginal leg structure is typical for Anteropatellata (
<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.4319.1.5" author="Kluge, NJ" journalOrPublisher="Zootaxa" pageId="0" pageNumber="81" pagination="91 - 127" refId="B14" refString="Kluge, NJ, 2017. Contribution to the knowledge of Cloeodes Traver 1938 (Ephemeroptera, Baetidae). Zootaxa 4319: 91 - 127, DOI: https://doi.org/10.11646/zootaxa.4319.1.5" title="Contribution to the knowledge of Cloeodes Traver 1938 (Ephemeroptera, Baetidae)." url="https://doi.org/10.11646/zootaxa.4319.1.5" volume="4319" year="2017">Kluge 2017</bibRefCitation>
).
</paragraph>
<caption doi="10.3897/zookeys.914.46652.figures16-18" httpUri="https://binary.pensoft.net/fig/381997" pageId="0" pageNumber="81" start="Figures 16–18" startId="F2">
<paragraph pageId="0" pageNumber="81">
<emphasis bold="true" pageId="0" pageNumber="81">Figures 16-18.</emphasis>
<taxonomicName class="Insecta" family="Baetidae" genus="Centroptella" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Centroptella illiesi" order="Ephemeroptera" pageId="0" pageNumber="81" phylum="Arthropoda" rank="species" species="illiesi">
<emphasis italics="true" pageId="0" pageNumber="81">Centroptella illiesi</emphasis>
</taxonomicName>
, tibiae of fore, middle and hind legs, view from anterior side (bases of long setae shown both on anterior and posterior sides).
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="81">
Besides this paradoxical combination of baetovectatan and protopatellatan characters, 
<taxonomicName authorityName="Braasch &amp; Soldan" authorityYear="1980" class="Insecta" family="Baetidae" genus="Centroptella" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Centroptella" order="Ephemeroptera" pageId="0" pageNumber="81" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="81">Centroptella</emphasis>
</taxonomicName>
has an evident autapomorphy: secondary swimming setae on the outer sides of the larval cerci in the distal part of the cercus have oval transverse bases and form a regular row (Figs 
<figureCitation captionStart="Figures 53–59" captionStartId="F6" captionText="Figures 53 - 59. Larval exuviae of Centroptella longisetosa (neotype of C. liebenauae). 53 - 57 at the same magnification: (53) left half of pro- and mesonotum (54) thoracic pleura, left half of metanotum, fore- and hind legs (55) frons, antenna and labrum (56) tenth uromere and caudalii; (57) abdominal sterna and terga I-IX 58 margins of abdominal sterna IV-VII 59 cercus." figureDoi="10.3897/zookeys.914.46652.figures53-59" httpUri="https://binary.pensoft.net/fig/382001" pageId="0" pageNumber="81">59</figureCitation>
, 
<figureCitation captionStart="Figures 129–132" captionStartId="F16" captionText="Figures 129 - 132. Centroptella ingridae sp. nov. (specimen from Thailand), exuviae of larval cercus (lateral view). 129 - 131 focus on lateral side to show bases of secondary swimming setae; 132 focus on median side to show bases of primary swimming setae." figureDoi="10.3897/zookeys.914.46652.figures129-132" httpUri="https://binary.pensoft.net/fig/382011" pageId="0" pageNumber="81">129-131</figureCitation>
); in this respect, they resemble the primary swimming setae on the inner side of the cercus (Fig. 
<figureCitation captionStart="Figures 129–132" captionStartId="F16" captionText="Figures 129 - 132. Centroptella ingridae sp. nov. (specimen from Thailand), exuviae of larval cercus (lateral view). 129 - 131 focus on lateral side to show bases of secondary swimming setae; 132 focus on median side to show bases of primary swimming setae." figureDoi="10.3897/zookeys.914.46652.figures129-132" httpUri="https://binary.pensoft.net/fig/382011" pageId="0" pageNumber="81">132</figureCitation>
), but they are smaller and less densely arranged.
</paragraph>
<paragraph pageId="0" pageNumber="81">
Another peculiar character of 
<taxonomicName authorityName="Braasch &amp; Soldan" authorityYear="1980" class="Insecta" family="Baetidae" genus="Centroptella" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Centroptella" order="Ephemeroptera" pageId="0" pageNumber="81" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="81">Centroptella</emphasis>
</taxonomicName>
is the presence of a pair of spaced transverse rows of long bifurcate setae on certain abdominal sterna of the larva (Figs 
<figureCitation captionStart="Figures 53–59" captionStartId="F6" captionText="Figures 53 - 59. Larval exuviae of Centroptella longisetosa (neotype of C. liebenauae). 53 - 57 at the same magnification: (53) left half of pro- and mesonotum (54) thoracic pleura, left half of metanotum, fore- and hind legs (55) frons, antenna and labrum (56) tenth uromere and caudalii; (57) abdominal sterna and terga I-IX 58 margins of abdominal sterna IV-VII 59 cercus." figureDoi="10.3897/zookeys.914.46652.figures53-59" httpUri="https://binary.pensoft.net/fig/382001" pageId="0" pageNumber="81">58</figureCitation>
, 
<figureCitation captionStart="Figures 116–122" captionStartId="F14" captionText="Figures 116 - 122. Centroptella ingridae sp. nov. (paratype of C. liebenauae), SEM photos of larva. 116 labrum 117 bifurcate setae on abdominal sternum IV 118 - 119 abdominal sterna IV-VI 120 - 121 paraprocts 122 flagellum of antenna." figureDoi="10.3897/zookeys.914.46652.figures116-122" httpUri="https://binary.pensoft.net/fig/382009" pageId="0" pageNumber="81">117-119</figureCitation>
); in different species these setal rows are present on sterna II-VI or on part of them, at least on sterna IV-V. Identical setal rows are found in a few other, non-related taxa (e.g., 
<taxonomicName authorityName="Gillies" authorityYear="1990" class="Insecta" family="Baetidae" genus="Potamocloeon" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Potamocloeon" order="Ephemeroptera" pageId="0" pageNumber="81" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="81">Potamocloeon</emphasis>
</taxonomicName>
Gillies, 1990 and 
<taxonomicName authorityName="Traver" authorityYear="1938" class="Insecta" family="Baetidae" genus="Cloeodes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cloeodes" order="Ephemeroptera" pageId="0" pageNumber="81" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="81">Cloeodes</emphasis>
</taxonomicName>
Traver, 1938).
</paragraph>
</subSubSection>
</treatment>
</document>