167 lines
14 KiB
XML
167 lines
14 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.127.1748" ID-GBIF-Dataset="5c4f147a-0e68-4abd-8b74-d15416f61f4d" ID-PMC="PMC3175127" ID-Pensoft-Pub="1313-2970-127-1" ID-Pensoft-UUID="FFF2FFC95E2C4223FFFFFFBCFFDAFF8F" ID-PubMed="21998544" ID-Zenodo-Dep="577174" ID-ZooBank="D170BE13535B467899997BA431A1AD0A" ModsDocAuthor="" ModsDocDate="2011" ModsDocID="1313-2970-127-1" ModsDocOrigin="ZooKeys 127" ModsDocTitle="Halocoryza Alluaud 1919, sea-side beetles of the Indian, Atlantic (sensu lato), and Pacific Oceans: a generic synopsis and description of a remarkable new species from Baja California Sur, México (Coleoptera, Carabidae, Scaritini, Clivinina)" checkinTime="1451249931271" checkinUser="pensoft" docAuthor="Erwin, Terry L." docDate="2011" docId="34D432D8C5E5AAD4819BA188DA30D1A8" docLanguage="en" docName="ZooKeys 127: 1-13" docOrigin="ZooKeys 127" docSource="http://dx.doi.org/10.3897/zookeys.127.1748" docTitle="Halocoryza whiteheadiana Erwin 2011, sp. n." docType="treatment" docUuid="FDB58B5D-FFB1-442F-BBEA-EBD82BC958F8" docUuidSource="ZooBank" docVersion="5" id="FFF2FFC95E2C4223FFFFFFBCFFDAFF8F" lastPageNumber="8" masterDocId="FFF2FFC95E2C4223FFFFFFBCFFDAFF8F" masterDocTitle="Halocoryza Alluaud 1919, sea-side beetles of the Indian, Atlantic (sensu lato), and Pacific Oceans: a generic synopsis and description of a remarkable new species from Baja California Sur, Mexico (Coleoptera, Carabidae, Scaritini, Clivinina)" masterLastPageNumber="13" masterPageNumber="1" pageNumber="6" updateTime="1668152055969" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Halocoryza Alluaud 1919, sea-side beetles of the Indian, Atlantic (sensu lato), and Pacific Oceans: a generic synopsis and description of a remarkable new species from Baja California Sur, Mexico (Coleoptera, Carabidae, Scaritini, Clivinina)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Erwin, Terry L.</mods:namePart>
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<mods:affiliation>Hyper-diversity Group, Department of Entomology, MRC- 187, National Museum of Natural History, Smithsonian Institution, Washington, P. O. Box 37012, DC 20013 - 7012, USA</mods:affiliation>
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<mods:nameIdentifier type="email">erwint@si.edu</mods:nameIdentifier>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2011</mods:date>
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<mods:detail type="volume">
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<mods:number>127</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>13</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.127.1748</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.127.1748</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-127-1</mods:identifier>
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<mods:identifier type="ZooBank">D170BE13535B467899997BA431A1AD0A</mods:identifier>
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<mods:identifier type="Pensoft-UUID">FFF2FFC95E2C4223FFFFFFBCFFDAFF8F</mods:identifier>
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<mods:identifier type="Zenodo-Dep">577174</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="152030634" LSID="urn:lsid:zoobank.org:act:FDB58B5D-FFB1-442F-BBEA-EBD82BC958F8" httpUri="http://treatment.plazi.org/id/34D432D8C5E5AAD4819BA188DA30D1A8" lastPageId="7" lastPageNumber="8" pageId="5" pageNumber="6">
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<subSubSection pageId="5" pageNumber="6" type="nomenclature">
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<paragraph pageId="5" pageNumber="6">
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<taxonomicName LSID="34D432D8-C5E5-AAD4-819B-A188DA30D1A8" authority="Erwin" authorityName="Erwin" authorityYear="2011" class="Insecta" family="Carabidae" genus="Halocoryza" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Halocoryza whiteheadiana" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="whiteheadiana" status="sp. n.">Halocoryza whiteheadiana Erwin</taxonomicName>
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<taxonomicNameLabel pageId="5" pageNumber="6">sp. n.</taxonomicNameLabel>
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<figureCitation captionStart="Figure 3" captionStartId="F2" captionText="Figure 3. Color image, showing habitus of Halocoryza whiteheadiana sp. n., 3 a, left lateral aspect, 3 b, dorsal aspect, ABL = 2.9 mm; Holotype: Isla Del Carmen, BJ, Mexico." httpUri="https://binary.pensoft.net/fig/17266" pageId="5" pageNumber="6">Figs 3a, 3b</figureCitation>
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<figureCitation captionStart="Figure 4–8" captionStartId="F3" captionText="Figure 4 - 8. 4 Line drawing, showing habitus of Halocoryza acapulcana Whitehead, 1966, dorsal aspect, ABL = 2.4 mm; Acapulco, Mexico; modified from Whitehead 1966 5 Line drawing, showing habitus of Halocoryza arenaria (Darlington, 1939), dorsal aspect, ABL = 2.9 mm; Rio Piedras, Puerto Rico; modified from Whitehead, 1966 6 Line drawing, showing habitus of Halocoryza whiteheadiana sp. n., dorsal aspect, ABL = 2.9 mm; ABL = 2.8 mm, ADP 127139; Isla Del Carmen, Mexico 7 Line drawing, showing male genitalia, median lobe, of Halocoryza acapulcana Whitehead, 1966, left lateral aspect, 0.41 mm (dorsal margin of basal lobe to apex); Acapulco, Mexico; modified from Whitehead 1966 8 Line drawing, showing male genitalia, median lobe, Halocoryza arenaria (Darlington, 1939), left lateral aspect with internal sac everted, 0.41 mm (dorsal margin of basal lobe to apex); Rio Piedras, Puerto Rico; modified from Whitehead, 1966." httpUri="https://binary.pensoft.net/fig/17267" pageId="5" pageNumber="6">6</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="5" pageNumber="6" type="common name">
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<paragraph pageId="5" pageNumber="6">Common name.</paragraph>
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<paragraph pageId="5" pageNumber="6">
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<emphasis bold="true" pageId="5" pageNumber="6">
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<normalizedToken originalValue="Whitehead’s">Whitehead's</normalizedToken>
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Saline Catarrh Beetle
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</emphasis>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="5" pageNumber="6" type="holotype. méxico">
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<paragraph pageId="5" pageNumber="6">
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Holotype.
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<normalizedToken originalValue="México">Mexico</normalizedToken>
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.
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</paragraph>
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<paragraph pageId="5" pageNumber="6">
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Baja California Sur, Isla Carmen, north end, sea level, approximately "
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<geoCoordinate degrees="26.05" direction="north" orientation="latitude" precision="555" value="26.05">26.05°N</geoCoordinate>
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,
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<geoCoordinate degrees="111.1" direction="west" orientation="longitude" precision="5555" value="-111.1">111.1°W</geoCoordinate>
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," 18-19 July 1984 (S.E. Miller) (NMNH: ADP127139, female).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="5" pageNumber="6" type="derivation of specific epithet">
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<paragraph pageId="5" pageNumber="6">Derivation of specific epithet.</paragraph>
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<paragraph pageId="5" pageNumber="6">
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The epithet
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<normalizedToken originalValue="“whiteheadiana”">
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"
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<emphasis italics="true" pageId="5" pageNumber="6">whiteheadiana</emphasis>
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"
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</normalizedToken>
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is an eponym, based on the family name of Donald R. Whitehead†, who had a profound interest in the species of this genus and its adelphotaxon,
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<taxonomicName authorityName="Putzeys" authorityYear="1846" class="Insecta" family="Carabidae" genus="Schizogenius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="5" pageNumber="6">Schizogenius</emphasis>
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</taxonomicName>
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Putzeys, during his relatively short career.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="5" pageNumber="6" type="proposed english vernacular name">
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<paragraph pageId="5" pageNumber="6">Proposed English vernacular name.</paragraph>
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<paragraph pageId="5" pageNumber="6">
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<normalizedToken originalValue="Whitehead’s">Whitehead's</normalizedToken>
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Saline Catarrh Beetle.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="5" pageNumber="6" type="diagnosis">
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<paragraph pageId="5" pageNumber="6">Diagnosis.</paragraph>
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<paragraph pageId="5" pageNumber="6">
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With the attributes of the genus described by
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<bibRefCitation author="Whitehead, DR" journalOrPublisher="Psyche" pageId="8" pageNumber="9" pagination="217 - 228" refId="B19" refString="Whitehead, DR, 1966. A review of Halocoryza Alluaud, with notes on its relationship to Schizogenius Putzeys (Coleoptera, Carabidae). Psyche 73: 217 - 228" title="A review of Halocoryza Alluaud, with notes on its relationship to Schizogenius Putzeys (Coleoptera, Carabidae)." volume="73" year="1966">Whitehead (1966)</bibRefCitation>
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and large sized for the genus. Adults rufotestaceous and shiny throughout; shallow microsculpture only in sulci. Occiput five-carinate each side with one medial and two lateral carinae on frons. Clypeus with 3 prominent tubercles; lateral margins prominently lobate. Elytron with 10 setae in interval 3, close to interneur 2.
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="6" lastPageNumber="7" pageId="5" pageNumber="6" type="description">
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<paragraph pageId="5" pageNumber="6">Description.</paragraph>
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<paragraph lastPageId="6" lastPageNumber="7" pageId="5" pageNumber="6">
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(Fig. 3).
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<emphasis italics="true" pageId="5" pageNumber="6">Size</emphasis>
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: Very small, ABL = 2.8 mm, SBL = 2.52 mm, EW = 0.69 mm, LP = 0.647mm, WP = 0.622mm, LE = 1.476mm.
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<emphasis italics="true" pageId="5" pageNumber="6">Color</emphasis>
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: Rufotestaceous throughout.
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<emphasis italics="true" pageId="5" pageNumber="6">Luster</emphasis>
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:Shiny throughout.
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<emphasis italics="true" pageId="5" pageNumber="6">Head:</emphasis>
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Labrum slightly emarginate apically, six-setose. Frons markedly tri-tuberculate apically, laterally markedly lobate, lobes nearly vertical, bicarinate basally, carinae set on an angle, widest basally. Eyes slightly convex; gena short and flat. Occiput five-carinate each side; rim above eye also carinate.
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<emphasis italics="true" pageId="5" pageNumber="6">Prothorax:</emphasis>
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Markedly convex, moderately longer than broad (W/L: 0.96), narrowed basally anteriad of posterior lateral pore; surface smooth, five-sulcate; lateral sulci 2/3 length of pronotum, extended to posterior lateral pore, paramedian sulci sigmoid shaped, not reaching base, median sulcus deep, extended from near apex to base of pronotum, slightly crossing anterior transverse sulcus; anterior and posterior lateral setae present.
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<emphasis italics="true" pageId="5" pageNumber="6">Pterothorax:</emphasis>
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Elytra markedly convex (W/L: 0.46), intervals markedly convex, intervals 3, 5, and 7 each with a serial row of setiferous punctures, 10 such in interval 3.
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<emphasis italics="true" pageId="6" pageNumber="7">
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<pageBreakToken pageId="6" pageNumber="7" start="start">Legs</pageBreakToken>
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:
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</emphasis>
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Normal in female.
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<emphasis italics="true" pageId="6" pageNumber="7">Abdomen</emphasis>
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: Abdominal sterna III to VI of female with normal ambulatory setae, VII with a pair of setae each side.
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<emphasis italics="true" pageId="6" pageNumber="7">Male genitalia:</emphasis>
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Unknown.
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<emphasis italics="true" pageId="6" pageNumber="7">Female genitalia</emphasis>
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: Not studied.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="6" pageNumber="7" type="dispersal potential">
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<paragraph pageId="6" pageNumber="7">Dispersal potential.</paragraph>
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<paragraph pageId="6" pageNumber="7">
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These beetles, as represented by the holotype, are macropterous and are capable of flight; they are slow runners, strong burrowers. However, both the Caribbean and Indian Ocean species are wing-polymorphic and perhaps with additional specimens,
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<taxonomicName authorityName="Erwin" authorityYear="2011" class="Insecta" family="Carabidae" genus="Halocoryza" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="whiteheadiana">
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<emphasis italics="true" pageId="6" pageNumber="7">Halocoryza whiteheadiana</emphasis>
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</taxonomicName>
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may prove to be the same.
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7" type="way of life">
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<paragraph pageId="6" pageNumber="7">Way of life.</paragraph>
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<paragraph lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7">
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Adults of other
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<taxonomicName authorityName="Alluaud" authorityYear="1919" class="Insecta" family="Carabidae" genus="Halocoryza" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="6" pageNumber="7">Halocoryza</emphasis>
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</taxonomicName>
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species are found on coralline sands in the intertidal zone of open beaches and among mangroves; a larva of the Indian Ocean species of this genus was found under dry seaweed just above the high-water mark on
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<pageBreakToken pageId="7" pageNumber="8" start="start">coralline</pageBreakToken>
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sands. The single known specimen of
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<taxonomicName authorityName="Erwin" authorityYear="2011" class="Insecta" family="Carabidae" genus="Halocoryza" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="whiteheadiana">
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<emphasis italics="true" pageId="7" pageNumber="8">Halocoryza whiteheadiana</emphasis>
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</taxonomicName>
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was collected at UV light on a sandy beach on the north shore of Isla Carmen, Baja California Sur. Adults of
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<taxonomicName authorityName="Erwin" authorityYear="2011" class="Insecta" family="Carabidae" genus="Halocoryza" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="whiteheadiana">
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<emphasis italics="true" pageId="7" pageNumber="8">Halocoryza whiteheadiana</emphasis>
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</taxonomicName>
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are likely nocturnal halobiont predators, as are members of the other known species of this genus.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="7" pageNumber="8" type="other specimens examined">
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<paragraph pageId="7" pageNumber="8">Other specimens examined.</paragraph>
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<paragraph pageId="7" pageNumber="8">None.</paragraph>
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</subSubSection>
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</treatment>
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</document> |