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<document ID-DOI="http://dx.doi.org/10.3897/zse.98.73162" ID-Pensoft-Pub="1860-0743-1-165" ID-Pensoft-UUID="FAA9F860168D569CAF6376719661F100" ID-ZooBank="EE314DBE46504C50B1D2953FDD47F876" ModsDocID="1860-0743-98-1-165" checkinTime="1654341530938" checkinUser="pensoft" docAuthor="Wang, Jian, Qi, Shuo, Dai, Ke-Yuan, Lyu, Zhi-Tong, Zeng, Zhao-Chi, Chen, Hong-Hui, Li, Yuan-Qiu, Zhao, Yong-You, Wang, Yun-Ze &amp; Wang, Ying-Yong" docDate="2022" docId="053962FC528B50BD97E46EBE1E8F38D7" docLanguage="en" docName="ZoosystEvol 98(1): 165-180" docOrigin="Zoosystematics and Evolution 98 (1)" docPubDate="2022-06-03" docSource="http://dx.doi.org/10.3897/zse.98.73162" docTitle="Leptobrachella shimentaina J. Wang, Z. - T. Lyu &amp; Y. - Y. Wang 2022, sp. nov." docType="treatment" docUuid="CF6EEF64-F539-415B-AD11-FA19C56BC3D5" docUuidSource="ZooBank" docVersion="2" id="FAA9F860168D569CAF6376719661F100" lastPageNumber="165" masterDocId="FAA9F860168D569CAF6376719661F100" masterDocTitle="A new Leptobrachella species (Anura, Megophryidae) from South China, with comments on the taxonomic status of L. chishuiensis and L. purpurus" masterLastPageNumber="180" masterPageNumber="165" pageNumber="165" updateTime="1654341832556" updateUser="ExternalLinkService">
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<mods:title>A new Leptobrachella species (Anura, Megophryidae) from South China, with comments on the taxonomic status of L. chishuiensis and L. purpurus</mods:title>
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<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Wang, Jian</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-4249-7767</mods:nameIdentifier>
<mods:affiliation>State Key Laboratory of Biocontrol, Sun Yat-sen University, Guangzhou 510275, China &amp; The Museum of Biology, School of Life Sciences, Sun Yat-sen University, Guangzhou 510275, China</mods:affiliation>
</mods:name>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Qi, Shuo</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-2924-6093</mods:nameIdentifier>
<mods:affiliation>State Key Laboratory of Biocontrol, Sun Yat-sen University, Guangzhou 510275, China</mods:affiliation>
</mods:name>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Dai, Ke-Yuan</mods:namePart>
<mods:affiliation>School of Ecology, Sun Yat-sen University, Guangzhou 510275, China</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Lyu, Zhi-Tong</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-4891-3859</mods:nameIdentifier>
<mods:affiliation>State Key Laboratory of Biocontrol, Sun Yat-sen University, Guangzhou 510275, China</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Zeng, Zhao-Chi</mods:namePart>
<mods:affiliation>State Key Laboratory of Biocontrol, Sun Yat-sen University, Guangzhou 510275, China &amp; The Museum of Biology, School of Life Sciences, Sun Yat-sen University, Guangzhou 510275, China</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Chen, Hong-Hui</mods:namePart>
<mods:affiliation>State Key Laboratory of Biocontrol, Sun Yat-sen University, Guangzhou 510275, China &amp; The Museum of Biology, School of Life Sciences, Sun Yat-sen University, Guangzhou 510275, China</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Li, Yuan-Qiu</mods:namePart>
<mods:affiliation>School of Ecology, Sun Yat-sen University, Guangzhou 510275, China</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Zhao, Yong-You</mods:namePart>
<mods:affiliation>School of Ecology, Sun Yat-sen University, Guangzhou 510275, China</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Wang, Yun-Ze</mods:namePart>
<mods:affiliation>Shuanghuamu Biological Technology Co., Ltd, Shenzhen 518000, China</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Wang, Ying-Yong</mods:namePart>
<mods:affiliation>State Key Laboratory of Biocontrol, Sun Yat-sen University, Guangzhou 510275, China</mods:affiliation>
<mods:nameIdentifier type="email">wangyy@mail.sysu.edu.cn</mods:nameIdentifier>
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<mods:date>2022</mods:date>
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<mods:number>2022-06-03</mods:number>
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<mods:number>98</mods:number>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zse.98.73162</mods:identifier>
<mods:identifier type="Pensoft-Pub">1860-0743-1-165</mods:identifier>
<mods:identifier type="ZooBank">EE314DBE46504C50B1D2953FDD47F876</mods:identifier>
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<treatment LSID="urn:lsid:zoobank.org:act:CF6EEF64-F539-415B-AD11-FA19C56BC3D5" httpUri="http://treatment.plazi.org/id/053962FC528B50BD97E46EBE1E8F38D7" lastPageNumber="165" pageId="0" pageNumber="165">
<subSubSection pageId="0" pageNumber="165" type="nomenclature">
<paragraph pageId="0" pageNumber="165">
<taxonomicName LSID="http://zoobank.org/CF6EEF64-F539-415B-AD11-FA19C56BC3D5" authority="J. Wang, Z. - T. Lyu &amp; Y. - Y. Wang" authorityName="J. Wang, Z. - T. Lyu &amp; Y. - Y. Wang" authorityYear="2022" class="Amphibia" family="Megophryidae" genus="Leptobrachella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Leptobrachella shimentaina" order="Anura" pageId="0" pageNumber="165" phylum="Chordata" rank="species" species="shimentaina" status="sp. nov.">Leptobrachella shimentaina J. Wang, Z.-T. Lyu &amp; Y.-Y. Wang</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="165">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="165" type="description">
<paragraph pageId="0" pageNumber="165">
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Morphological features of the holotype of Leptobrachella shimentaina sp. nov., SYS a 007684 in life: A. Dorsal view; B. Dorsolateral view; C. Ventral view; D. Character of eye and temporal region; E. Ventral view of hand; F. Ventral view of foot." figureDoi="10.3897/zse.98.73162.figure3" httpUri="https://binary.pensoft.net/fig/692037" pageId="0" pageNumber="165">Shimentai Leaf Litter Toad (in English) / Shi Men Tai Zhang Tu Chan (石门台掌突蟾in Chinese) Figs 3</figureCitation>
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. The holotype of Leptobrachella shimentaina sp. nov., SYS a 007684 in preservative." figureDoi="10.3897/zse.98.73162.figure4" httpUri="https://binary.pensoft.net/fig/692038" pageId="0" pageNumber="165">, 4</figureCitation>
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Morphological features of the paratypes of Leptobrachella shimentaina sp. nov.: A 1. A female (SYS a 004711 in situ; ventral view of the same specimen's hand [A 2] and foot [A 3]; B. Male SYS a 007685 in situ; C. Male SYS a 007686 in situ." figureDoi="10.3897/zse.98.73162.figure5" httpUri="https://binary.pensoft.net/fig/692039" pageId="0" pageNumber="165">, 5</figureCitation>
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Morphological features of a paraype (SYS a 007687) of Leptobrachella shimentaina sp. nov. in life." figureDoi="10.3897/zse.98.73162.figure6" httpUri="https://binary.pensoft.net/fig/692040" pageId="0" pageNumber="165">, 6</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="165" type="chresonymy">
<paragraph pageId="0" pageNumber="165">Chresonymy.</paragraph>
<paragraph pageId="0" pageNumber="165">
&quot;
<taxonomicName authorityName="Dubois" authorityYear="1980" class="Amphibia" family="Megophryidae" genus="Leptolalax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Leptolalax" order="Anura" pageId="0" pageNumber="165" phylum="Chordata" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="165">Leptolalax</emphasis>
</taxonomicName>
sp.6&quot; (
<bibRefCitation DOI="https://doi.org/10.1016/j.ympev.2018.02.020" author="Chen, JM" journalOrPublisher="Molecular Phylogenetics and Evolution" pageId="0" pageNumber="165" pagination="162 - 171" refId="B7" refString="Chen, JM, Poyarkov Jr, NJ, Suwannapoom, C, Lathrop, A, Wu, YH, Zhou, WW, Yuan, ZY, Jin, JQ, Chen, HM, Liu, HQ, Nguyen, TQ, Nguyen, SN, Duong, TV, Eto, K, Nishikawa, K, Matsui, M, Orlov, NL, Stuart, BL, Brown, RM, Rowley, JJL, Murphy, RW, Wang, YY, Che, J, 2018. Large-scale phylogenetic analyses provide insights into unrecognized diversity and historical biogeography of Asian leaf-litter frogs, genus Leptolalax (Anura: Megophryidae). Molecular Phylogenetics and Evolution 124: 162 - 171, DOI: https://doi.org/10.1016/j.ympev.2018.02.020" title="Large-scale phylogenetic analyses provide insights into unrecognized diversity and historical biogeography of Asian leaf-litter frogs, genus Leptolalax (Anura: Megophryidae)." url="https://doi.org/10.1016/j.ympev.2018.02.020" volume="124" year="2018">Chen et al. 2018</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="165" type="type material">
<paragraph pageId="0" pageNumber="165">Type material.</paragraph>
<paragraph pageId="0" pageNumber="165">
<emphasis bold="true" italics="true" pageId="0" pageNumber="165">Holotype</emphasis>
♂. SYS a007684, from Jiuzhou Yizhan (
<geoCoordinate degrees="24" direction="north" minutes="23" orientation="latitude" precision="1" seconds="38.01" value="24.393892">24°23'38.01&quot;N</geoCoordinate>
,
<geoCoordinate degrees="113" direction="east" minutes="06" orientation="longitude" precision="1" seconds="35.38" value="113.109825">113°06'35.38&quot;E</geoCoordinate>
; ca. 300 m a.s.l.), Shimentai Nature Reserve, Yingde City, Guangdong Province, China, collected by Jian Wang and Yong-You Zhao on 23 April 2019.
</paragraph>
<caption doi="10.3897/zse.98.73162.figure3" httpUri="https://binary.pensoft.net/fig/692037" pageId="0" pageNumber="165" start="Figure 3" startId="F3">
<paragraph pageId="0" pageNumber="165">
<emphasis bold="true" pageId="0" pageNumber="165">Figure 3.</emphasis>
Morphological features of the holotype of
<taxonomicName authorityName="J. Wang, Z. - T. Lyu &amp; Y. - Y. Wang" authorityYear="2022" class="Amphibia" family="Megophryidae" genus="Leptobrachella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Leptobrachella shimentaina" order="Anura" pageId="0" pageNumber="165" phylum="Chordata" rank="species" species="shimentaina">
<emphasis italics="true" pageId="0" pageNumber="165">Leptobrachella shimentaina</emphasis>
</taxonomicName>
sp. nov., SYS a007684 in life:
<emphasis bold="true" pageId="0" pageNumber="165">A.</emphasis>
Dorsal view;
<emphasis bold="true" pageId="0" pageNumber="165">B.</emphasis>
Dorsolateral view;
<emphasis bold="true" pageId="0" pageNumber="165">C.</emphasis>
Ventral view;
<emphasis bold="true" pageId="0" pageNumber="165">D.</emphasis>
Character of eye and temporal region;
<emphasis bold="true" pageId="0" pageNumber="165">E.</emphasis>
Ventral view of hand;
<emphasis bold="true" pageId="0" pageNumber="165">F.</emphasis>
Ventral view of foot.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="165">
<emphasis bold="true" italics="true" pageId="0" pageNumber="165">Paratypes</emphasis>
(n = 7). 5 ♂: SYS a007683/ CIB116079, same collection place as holotype; SYS a007685-7686, collected by Yu-Long Li and Hong-Hui Chen on 23 April 2019 from Shijiao (
<geoCoordinate degrees="24" direction="north" minutes="26" orientation="latitude" precision="1" seconds="35.31" value="24.44314">24°26'35.31&quot;N</geoCoordinate>
,
<geoCoordinate degrees="113" direction="east" minutes="09" orientation="longitude" precision="1" seconds="21.55" value="113.15599">113°09'21.55&quot;E</geoCoordinate>
; ca 400 m a.s.l.), Shimentai Nature Reserve; SYS a007687, collected by Jian Wang and Yong-You Zhao on 24 April 2019 from Longtankeng (
<geoCoordinate degrees="24" direction="north" minutes="26" orientation="latitude" precision="1" seconds="31.38" value="24.442049">24°26'31.38&quot;N</geoCoordinate>
,
<geoCoordinate degrees="113" direction="east" minutes="15" orientation="longitude" precision="1" seconds="28.81" value="113.258">113°15'28.81&quot;E</geoCoordinate>
; ca 310 m a.s.l.), Shimentai Nature Reserve; SYS a008329, collected by Jian Wang on 27 April 2020 from Luokeng Nature Reserve (
<geoCoordinate degrees="24" direction="north" minutes="31" orientation="latitude" precision="1" seconds="42.3" value="24.528416">24°31'42.3&quot;N</geoCoordinate>
,
<geoCoordinate degrees="113" direction="east" minutes="15" orientation="longitude" precision="1" seconds="54.3" value="113.26508">113°15'54.3&quot;E</geoCoordinate>
; ca 590 m a.s.l.), Shaoguan City, Guangdong Province, China. 2 ♀: SYS a004711-4712, collected by Ying-Yong Wang, Jian Wang, and Zhi-Tong Lyu on 27 April 2016 from Hengshitang (
<geoCoordinate degrees="24" direction="north" minutes="26" orientation="latitude" precision="1" seconds="14.92" value="24.437477">24°26'14.92&quot;N</geoCoordinate>
,
<geoCoordinate degrees="113" direction="east" minutes="17" orientation="longitude" precision="1" seconds="50.48" value="113.297356">113°17'50.48&quot;E</geoCoordinate>
; ca 380 m a.s.l.), Shimentai Nature Reserve.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="165" type="etymology">
<paragraph pageId="0" pageNumber="165">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="165">
The specific epithet &quot;
<emphasis italics="true" pageId="0" pageNumber="165">shimentaina</emphasis>
&quot; is an adjective derived from
<normalizedToken originalValue="“shimentai”">&quot;shimentai&quot;</normalizedToken>
, referring to the type locality of the new species, Shimentai Nature Reserve, &quot;
<emphasis italics="true" pageId="0" pageNumber="165">ina</emphasis>
&quot; is used as a feminine suffix which indicates the relationship of position.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="165" type="diagnosis">
<paragraph pageId="0" pageNumber="165">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="165">(1) small body size [SVL 26.4-28.9 mm in six adult males, SVL 30.1 and 30.7 mm in two adult females], (2) iris bicolored, upper half coppery orange and lower half silver, (3) tympanum distinct, (4) supratympanic line black, (5) fingers unwebbed, with distinct lateral fringes in males [absence in females], (6) toes with rudimentary webbing, lateral fringes wide in males [narrow in females], (7) longitudinal ridges under toes continuous, with constrictions at interphalangeal articulations, (8) heels slightly overlapping when adpressed, tibial-tarsal articulation reaching mid-orbit, (9) relative lengths of fingers I = II = IV &lt;III, and toe I &lt;II &lt;III = V &lt;IV, (10) dorsal surface shagreened and granular, lacking enlarged warts, with some granules forming short longitudinal folds, (11) dorsum grayish brown to yellowish brown, with small light orange granules and distinct darker brown scattered markings with irregular light orange pigmentation, (12) flanks with several dark spots, (13) ventral surface grayish pink, with distinct hazy brown speckling on chest and ventrolateral flanks, (14) densely-distributed, small, raised pectoral and abdominal tubercles present; (15) dorsal surfaces of forelimbs and digits with dark transverse bars.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="165" type="description">
<paragraph pageId="0" pageNumber="165">Description of holotype.</paragraph>
<paragraph pageId="0" pageNumber="165">Adult male. Body size small, SVL in 28.6 mm. Head length slightly larger than head width, HDW/HDL 0.91; snout slightly protruding, projecting slightly beyond margin of lower jaw; nostril closer to snout than eye; canthus rostralis gently rounded; loreal region slightly concave; interorbital space flat, internarial distance larger than interorbital distance, IND/IOD 1.10; pineal ocellus absent; pupil vertical; snout longer than eye diameter, SNT/EYE 1.19; tympanum distinct, rounded, diameter smaller than that of eye and larger than tympanum-eye distance, TMP/EYE 0.41, TEY/TMP 0.48; upper margin of tympanum in contact with supratympanic ridge; vomerine teeth absent; a single vocal sac; vocal sac openings slit-like, paired, located posterolaterally on floor of mouth, close to margins of mandible; tongue deeply notched posteriorly; supratympanic ridge distinct, extending from posterior corner of eye to supra-axillary gland.</paragraph>
<paragraph pageId="0" pageNumber="165">Tips of fingers rounded, slightly swollen; relative finger lengths I = II = IV &lt;III; nuptial pad absent; subarticular tubercles absent; inner palmar tubercle large, rounded, distinctly separated from outer palmar tubercle; outer palmar tubercle small, rounded; fingers lacking interdigital webbing, with distinct lateral fringes. Tips of toes rounded, slightly swollen; relative toe length I &lt;II &lt;III = V &lt;IV; longitudinal ridges under toes continuous, with constrictions at interphalangeal articulations; inner metatarsal tubercle large, oval; outer metatarsal tubercle absent; interdigital webbing between toes rudimentary; wide lateral fringes present on all toes. Tibia 47% of snout-vent length; tibiotarsal articulation reaches to middle of eye; heels slightly overlapping when thighs are appressed at right angles with respect to body.</paragraph>
<paragraph pageId="0" pageNumber="165">Dorsal surface shagreened and granular, lacking enlarged tubercles or warts, some granules forming short longitudinal folds; ventral skin smooth, densely-distributed small raised scapular and abdominal tubercles present; pectoral and femoral glands oval; both larger in diameter than tip of fingers; femoral glands larger in diameter than tip of toes; femoral gland situated on posteroventral surface of thigh, closer to knee than to vent; supra-axillary gland raised. Ventrolateral gland distinctly visible, forming an longitudinal uncontinuous series.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="165" type="coloration of holotype in life">
<paragraph pageId="0" pageNumber="165">Coloration of holotype in life.</paragraph>
<paragraph pageId="0" pageNumber="165">Dorsal background color yellowish-brown, with small orange granules, distinct dark brown markings, and rounded spots, and scattered with irregular light grayish-brown pigmentation. A dark brown inverted triangular marking present between anterior corners of eyes, connecting to dark brown W-shaped marking in interorbital region. This W-shaped marking is in connection to the other W-shaped marking on occipital region. Tympanum dark brown, lower margin grayish yellow. Supratympanic line black. Dorsal surfaces of body and limbs with small orange granules; a pair of dark brown vertical bars present under eyes; transverse dark brown bars on dorsal surfaces of distal limbs and digits; distinct dark brown blotches on flanks; surfaces of elbows and upper arms coppery orange, without dark bars.</paragraph>
<paragraph pageId="0" pageNumber="165">Surface of throat, chest, and belly grayish pink, with distinct hazy brown speckling on chest and ventrolateral flanks, without black spots; surface of chin and lower lip brown with grayish white spots and patches; ventral surface of limbs brown with pink hues. Supra-axillary gland coppery orange, pectoral glands grayish white, femoral glands coppery orange, ventrolateral glands brown. Iris bicolored, upper half coppery orange, lower half silver.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="165" type="coloration of holotype in preservative">
<paragraph pageId="0" pageNumber="165">Coloration of holotype in preservative</paragraph>
<paragraph pageId="0" pageNumber="165">
<emphasis bold="true" pageId="0" pageNumber="165">
(Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. The holotype of Leptobrachella shimentaina sp. nov., SYS a 007684 in preservative." figureDoi="10.3897/zse.98.73162.figure4" httpUri="https://binary.pensoft.net/fig/692038" pageId="0" pageNumber="165">4</figureCitation>
).
</emphasis>
Dorsal background color grayish brown, scattered with irregular light grayish-brown pigmentation. All markings, bars and spots become more distinct. Tympanum dark brown, lower margin gray.
</paragraph>
<caption doi="10.3897/zse.98.73162.figure4" httpUri="https://binary.pensoft.net/fig/692038" pageId="0" pageNumber="165" start="Figure 4" startId="F4">
<paragraph pageId="0" pageNumber="165">
<emphasis bold="true" pageId="0" pageNumber="165">Figure 4.</emphasis>
The holotype of
<taxonomicName authorityName="J. Wang, Z. - T. Lyu &amp; Y. - Y. Wang" authorityYear="2022" class="Amphibia" family="Megophryidae" genus="Leptobrachella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Leptobrachella shimentaina" order="Anura" pageId="0" pageNumber="165" phylum="Chordata" rank="species" species="shimentaina">
<emphasis italics="true" pageId="0" pageNumber="165">Leptobrachella shimentaina</emphasis>
</taxonomicName>
sp. nov., SYS a007684 in preservative.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="165">Ventral surface yellowish brown; speckling on chest and ventrolateral flanks become more distinct; ventral surface of limbs dark brown. Supra-axillary, pectoral, and ventrolateral glands greyish yellow.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="165" type="variation">
<paragraph pageId="0" pageNumber="165">Variation.</paragraph>
<paragraph pageId="0" pageNumber="165">
Measurements and body proportions are listed in Table
<tableCitation captionStart="Table 2" captionStartId="T2" captionText="Table 2. Measurements (minimum-maximum (mean + / - SD); in mm), and body proportions of Leptobrachella shimentaina sp. nov." httpUri="http://table.plazi.org/id/C371DC13290F2BA002DED3C82E2AE120" pageId="0" pageNumber="165" tableUuid="C371DC13290F2BA002DED3C82E2AE120">2</tableCitation>
. All male paratypes from Shimentai Nature Reserve (SYS a007683/ CIB116079, SYS a007685 (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Morphological features of the paratypes of Leptobrachella shimentaina sp. nov.: A 1. A female (SYS a 004711 in situ; ventral view of the same specimen's hand [A 2] and foot [A 3]; B. Male SYS a 007685 in situ; C. Male SYS a 007686 in situ." figureDoi="10.3897/zse.98.73162.figure5" httpUri="https://binary.pensoft.net/fig/692039" pageId="0" pageNumber="165">5B</figureCitation>
), 7686 (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Morphological features of the paratypes of Leptobrachella shimentaina sp. nov.: A 1. A female (SYS a 004711 in situ; ventral view of the same specimen's hand [A 2] and foot [A 3]; B. Male SYS a 007685 in situ; C. Male SYS a 007686 in situ." figureDoi="10.3897/zse.98.73162.figure5" httpUri="https://binary.pensoft.net/fig/692039" pageId="0" pageNumber="165">5C</figureCitation>
), 7687) show very similar morphological characters to holotype SYS a007684; paratype SYS a008329 (Fig.
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Morphological features of a paraype (SYS a 007687) of Leptobrachella shimentaina sp. nov. in life." figureDoi="10.3897/zse.98.73162.figure6" httpUri="https://binary.pensoft.net/fig/692040" pageId="0" pageNumber="165">6</figureCitation>
) from Luokeng Nature Reserve shows a darker coloration on dorsum, and brighter coloration on ventrum. Moreover, female paratypes, SYS a004711 (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Morphological features of the paratypes of Leptobrachella shimentaina sp. nov.: A 1. A female (SYS a 004711 in situ; ventral view of the same specimen's hand [A 2] and foot [A 3]; B. Male SYS a 007685 in situ; C. Male SYS a 007686 in situ." figureDoi="10.3897/zse.98.73162.figure5" httpUri="https://binary.pensoft.net/fig/692039" pageId="0" pageNumber="165">5A</figureCitation>
) and SYS a004712 show lighter background coloration and more obvious dark brown marking orange tubercles on back.
</paragraph>
<caption ID-Table-UUID="C371DC13290F2BA002DED3C82E2AE120" httpUri="http://table.plazi.org/id/C371DC13290F2BA002DED3C82E2AE120" pageId="0" pageNumber="165" start="Table 2" startId="T2">
<paragraph pageId="0" pageNumber="165">
<emphasis bold="true" pageId="0" pageNumber="165">Table 2.</emphasis>
Measurements (minimum-maximum (mean
<normalizedToken originalValue="±">+/-</normalizedToken>
SD); in mm), and body proportions of
<taxonomicName authorityName="J. Wang, Z. - T. Lyu &amp; Y. - Y. Wang" authorityYear="2022" class="Amphibia" family="Megophryidae" genus="Leptobrachella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Leptobrachella shimentaina" order="Anura" pageId="0" pageNumber="165" phylum="Chordata" rank="species" species="shimentaina">
<emphasis italics="true" pageId="0" pageNumber="165">Leptobrachella shimentaina</emphasis>
</taxonomicName>
sp. nov.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="165">
<table pageId="0" pageNumber="165">
<tr pageId="0" pageNumber="165">
<th colspan="1" pageId="0" pageNumber="165" rowspan="1">Vocher</th>
<th colspan="1" pageId="0" pageNumber="165" rowspan="1">SYS a007683</th>
<th colspan="1" pageId="0" pageNumber="165" rowspan="1">SYS a007684</th>
<th colspan="1" pageId="0" pageNumber="165" rowspan="1">SYS a007685</th>
<th colspan="1" pageId="0" pageNumber="165" rowspan="1">SYS a007686</th>
<th colspan="1" pageId="0" pageNumber="165" rowspan="1">SYS a007687</th>
<th colspan="1" pageId="0" pageNumber="165" rowspan="1">SYS a008329</th>
<th colspan="1" pageId="0" pageNumber="165" rowspan="1">Range</th>
<th colspan="1" pageId="0" pageNumber="165" rowspan="1">Vocher</th>
<th colspan="1" pageId="0" pageNumber="165" rowspan="1">SYS a004711</th>
<th colspan="1" pageId="0" pageNumber="165" rowspan="1">SYS a004712</th>
<th colspan="1" pageId="0" pageNumber="165" rowspan="1">Range</th>
</tr>
<tr pageId="0" pageNumber="165">
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="165">Sex</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">Male</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">Male</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">Male</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">Male</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">Male</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">Male</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">Males (n = 6)</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="165">Sex</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">Female</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">Female</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">Females (n = 2)</td>
</tr>
<tr pageId="0" pageNumber="165">
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="165">SVL</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">28.9</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">28.6</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">28.4</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">28.7</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">28.4</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">26.4</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">
26.4-28.9 (28.2
<normalizedToken originalValue="±">+/-</normalizedToken>
0.9)
</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="165">SVL</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">30.1</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">30.7</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">
30.1-30.7 (30.4
<normalizedToken originalValue="±">+/-</normalizedToken>
0.4)
</td>
</tr>
<tr pageId="0" pageNumber="165">
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="165">HDL</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">11.7</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">10.7</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">10.8</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">10.9</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">10.8</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">9.8</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">
9.8-11.7 (10.8
<normalizedToken originalValue="±">+/-</normalizedToken>
0.6)
</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="165">HDL</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">10.1</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">10.4</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">
10.1-10.4 (10.3
<normalizedToken originalValue="±">+/-</normalizedToken>
0.2)
</td>
</tr>
<tr pageId="0" pageNumber="165">
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="165">HDW</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">9.7</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">9.7</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">9.7</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">9.7</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">9.7</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">9.2</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">
9.2-9.7 (9.6
<normalizedToken originalValue="±">+/-</normalizedToken>
0.2)
</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="165">HDW</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">9.7</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">9.9</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">
9.7-9.9 (9.8
<normalizedToken originalValue="±">+/-</normalizedToken>
0.2)
</td>
</tr>
<tr pageId="0" pageNumber="165">
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="165">SNT</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">4.3</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">4.2</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">4.1</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">4.3</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">4.2</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">3.3</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">
3.3-4.3 (4.1
<normalizedToken originalValue="±">+/-</normalizedToken>
0.4)
</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="165">SNT</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">4.0</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">4.3</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">
4.0-4.3 (4.1
<normalizedToken originalValue="±">+/-</normalizedToken>
0.2)
</td>
</tr>
<tr pageId="0" pageNumber="165">
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="165">IND</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">2.9</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">2.7</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">2.9</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">2.8</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">2.7</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">2.9</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">
2.7-2.9 (2.8
<normalizedToken originalValue="±">+/-</normalizedToken>
0.1)
</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="165">IND</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">2.7</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">2.8</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">
2.7-2.8 (2.7
<normalizedToken originalValue="±">+/-</normalizedToken>
0.1)
</td>
</tr>
<tr pageId="0" pageNumber="165">
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="165">IOD</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">2.6</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">2.5</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">2.5</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">2.5</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">2.5</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">2.6</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">
2.5-2.6 (2.5
<normalizedToken originalValue="±">+/-</normalizedToken>
0.1)
</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="165">IOD</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">2.7</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">2.8</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">
2.7-2.8 (2.7
<normalizedToken originalValue="±">+/-</normalizedToken>
0.0)
</td>
</tr>
<tr pageId="0" pageNumber="165">
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="165">EYE</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">3.5</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">3.5</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">3.4</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">3.4</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">3.4</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">3.3</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">
3.3-3.5 (3.4
<normalizedToken originalValue="±">+/-</normalizedToken>
0.1)
</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="165">EYE</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">3.5</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">3.6</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">
3.5-3.6 (3.5
<normalizedToken originalValue="±">+/-</normalizedToken>
0.1)
</td>
</tr>
<tr pageId="0" pageNumber="165">
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="165">TMP</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">1.5</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">1.5</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">1.5</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">1.5</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">1.5</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">1.5</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">
1.5-1.5 (1.5
<normalizedToken originalValue="±">+/-</normalizedToken>
0.0)
</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="165">TMP</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">1.5</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">1.8</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">
1.5-1.8 (1.7
<normalizedToken originalValue="±">+/-</normalizedToken>
0.2)
</td>
</tr>
<tr pageId="0" pageNumber="165">
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="165">TEY</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">0.7</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">0.7</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">0.7</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">0.7</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">0.7</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">0.6</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">
0.6-0.7 (0.7
<normalizedToken originalValue="±">+/-</normalizedToken>
0.0)
</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="165">TEY</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">0.6</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">0.9</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">
0.6-0.9 (0.8
<normalizedToken originalValue="±">+/-</normalizedToken>
0.2)
</td>
</tr>
<tr pageId="0" pageNumber="165">
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="165">ML</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">7.4</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">7.2</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">7.2</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">7.4</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">7.3</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">6.9</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">
6.9-7.4 (7.2
<normalizedToken originalValue="±">+/-</normalizedToken>
0.2)
</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="165">ML</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">7.2</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">7.8</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">
7.2-7.8 (7.5
<normalizedToken originalValue="±">+/-</normalizedToken>
0.4)
</td>
</tr>
<tr pageId="0" pageNumber="165">
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="165">LAHL</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">14.4</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">14.0</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">14.4</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">14.5</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">14.4</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">12.5</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">
12.5-14.5 (14.0
<normalizedToken originalValue="±">+/-</normalizedToken>
0.8)
</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="165">LAHL</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">14.1</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">15.0</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">
14.1-15.0 (14.6
<normalizedToken originalValue="±">+/-</normalizedToken>
0.6)
</td>
</tr>
<tr pageId="0" pageNumber="165">
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="165">PL</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">12.6</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">12.6</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">13.0</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">12.2</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">12.6</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">11.9</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">
11.9-13.0 (12.5
<normalizedToken originalValue="±">+/-</normalizedToken>
0.4)
</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="165">PL</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">13.1</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">12.5</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">
12.5-13.1 (12.8
<normalizedToken originalValue="±">+/-</normalizedToken>
0.4)
</td>
</tr>
<tr pageId="0" pageNumber="165">
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="165">TIB</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">14.2</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">13.3</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">13.3</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">13.3</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">13.4</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">12.4</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">
12.4-14.2 (13.3
<normalizedToken originalValue="±">+/-</normalizedToken>
0.6)
</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="165">TIB</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">14.1</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">14.6</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">
14.1-14.6 (14.3
<normalizedToken originalValue="±">+/-</normalizedToken>
0.4)
</td>
</tr>
<tr pageId="0" pageNumber="165">
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="165">HLL</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">45.1</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">44.5</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">44.5</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">45.1</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">44.9</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">39.61</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">
39.6-45.1 (43.9
<normalizedToken originalValue="±">+/-</normalizedToken>
2.1)
</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="165">HLL</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">44.4</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">45.6</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">
44.4-45.6 (45.0
<normalizedToken originalValue="±">+/-</normalizedToken>
0.9)
</td>
</tr>
<tr pageId="0" pageNumber="165">
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="165">HDL/SVL</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">0.40</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">0.37</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">0.38</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">0.38</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">0.38</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">0.37</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">
0.37-0.40 (0.38
<normalizedToken originalValue="±">+/-</normalizedToken>
0.01)
</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="165">HDL/SVL</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">0.34</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">0.34</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">
0.34-0.34 (0.34
<normalizedToken originalValue="±">+/-</normalizedToken>
0.00)
</td>
</tr>
<tr pageId="0" pageNumber="165">
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="165">HDW/SVL</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">0.34</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">0.34</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">0.34</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">0.34</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">0.34</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">0.35</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">
0.34-0.35 (0.34
<normalizedToken originalValue="±">+/-</normalizedToken>
0.00)
</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="165">HDW/SVL</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">0.32</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">0.32</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">
0.32-0.32 (0.32
<normalizedToken originalValue="±">+/-</normalizedToken>
0.00)
</td>
</tr>
<tr pageId="0" pageNumber="165">
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="165">HDW/HDL</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">0.83</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">0.91</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">0.90</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">0.89</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">0.90</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">0.94</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">
0.83-0.94 (0.90
<normalizedToken originalValue="±">+/-</normalizedToken>
0.03)
</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="165">HDW/HDL</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">0.95</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">0.95</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">
0.95-0.95 (0.95
<normalizedToken originalValue="±">+/-</normalizedToken>
0.00)
</td>
</tr>
<tr pageId="0" pageNumber="165">
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="165">SNT/HDL</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">0.37</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">0.39</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">0.38</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">0.39</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">0.39</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">0.34</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">
0.34-0.39 (0.38
<normalizedToken originalValue="±">+/-</normalizedToken>
0.02)
</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="165">SNT/HDL</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">0.39</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">0.41</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">
0.39-0.41 (0.40
<normalizedToken originalValue="±">+/-</normalizedToken>
0.02)
</td>
</tr>
<tr pageId="0" pageNumber="165">
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="165">IND/HDW</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">0.30</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">0.28</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">0.29</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">0.28</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">0.28</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">0.31</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">
0.28-0.31 (0.29
<normalizedToken originalValue="±">+/-</normalizedToken>
0.01)
</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="165">IND/HDW</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">0.28</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">0.28</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">
0.28-0.28 (0.28
<normalizedToken originalValue="±">+/-</normalizedToken>
0.01)
</td>
</tr>
<tr pageId="0" pageNumber="165">
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="165">IOD/HDW</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">0.27</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">0.25</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">0.26</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">0.25</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">0.25</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">0.29</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">
0.25-0.29 (0.26
<normalizedToken originalValue="±">+/-</normalizedToken>
0.01)
</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="165">IOD/HDW</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">0.28</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">0.28</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">
0.28-0.28 (0.28
<normalizedToken originalValue="±">+/-</normalizedToken>
0.00)
</td>
</tr>
<tr pageId="0" pageNumber="165">
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="165">EYE/HDL</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">0.30</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">0.33</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">0.31</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">0.31</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">0.32</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">0.34</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">
0.30-0.34 (0.32
<normalizedToken originalValue="±">+/-</normalizedToken>
0.01)
</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="165">EYE/HDL</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">0.34</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">0.35</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">
0.34-0.35 (0.34
<normalizedToken originalValue="±">+/-</normalizedToken>
0.00)
</td>
</tr>
<tr pageId="0" pageNumber="165">
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="165">TMP/EYE</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">0.42</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">0.41</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">0.43</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">0.43</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">0.43</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">0.45</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">
0.41-0.45 (0.43
<normalizedToken originalValue="±">+/-</normalizedToken>
0.01)
</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="165">TMP/EYE</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">0.44</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">0.50</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">
0.44-0.50 (0.47
<normalizedToken originalValue="±">+/-</normalizedToken>
0.04)
</td>
</tr>
<tr pageId="0" pageNumber="165">
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="165">ML/SVL</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">0.25</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">0.25</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">0.25</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">0.26</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">0.26</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">0.26</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">
0.25-0.26 (0.26
<normalizedToken originalValue="±">+/-</normalizedToken>
0.00)
</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="165">ML/SVL</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">0.24</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">0.25</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">
0.24-0.25 (0.25
<normalizedToken originalValue="±">+/-</normalizedToken>
0.01)
</td>
</tr>
<tr pageId="0" pageNumber="165">
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="165">LAHL/SVL</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">0.50</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">0.49</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">0.51</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">0.50</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">0.51</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">0.47</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">
0.47-0.51 (0.50
<normalizedToken originalValue="±">+/-</normalizedToken>
0.01)
</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="165">LAHL/SVL</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">0.47</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">0.49</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">
0.47-0.49 (0.48
<normalizedToken originalValue="±">+/-</normalizedToken>
0.01)
</td>
</tr>
<tr pageId="0" pageNumber="165">
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="165">PL/SVL</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">0.43</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">0.44</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">0.46</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">0.42</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">0.44</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">0.45</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">
0.42-0.46 (0.44
<normalizedToken originalValue="±">+/-</normalizedToken>
0.01)
</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="165">PL/SVL</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">0.43</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">0.41</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">
0.41-0.43 (0.42
<normalizedToken originalValue="±">+/-</normalizedToken>
0.02)
</td>
</tr>
<tr pageId="0" pageNumber="165">
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="165">TIB/SVL</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">0.49</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">0.47</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">0.47</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">0.46</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">0.47</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">0.47</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">
0.46-0.49 (0.47
<normalizedToken originalValue="±">+/-</normalizedToken>
0.01)
</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="165">TIB/SVL</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">0.47</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">0.47</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">
0.47-0.47 (0.47
<normalizedToken originalValue="±">+/-</normalizedToken>
0.01)
</td>
</tr>
<tr pageId="0" pageNumber="165">
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="165">HLL/SVL</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">1.56</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">1.55</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">1.57</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">1.57</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">1.58</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">1.50</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">
1.50-1.58 (1.55
<normalizedToken originalValue="±">+/-</normalizedToken>
0.03)
</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="165">HLL/SVL</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">1.47</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">1.48</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">
1.47-1.48 (1.48
<normalizedToken originalValue="±">+/-</normalizedToken>
0.01)
</td>
</tr>
</table>
</paragraph>
<caption doi="10.3897/zse.98.73162.figure5" httpUri="https://binary.pensoft.net/fig/692039" pageId="0" pageNumber="165" start="Figure 5" startId="F5">
<paragraph pageId="0" pageNumber="165">
<emphasis bold="true" pageId="0" pageNumber="165">Figure 5.</emphasis>
Morphological features of the paratypes of
<taxonomicName authorityName="J. Wang, Z. - T. Lyu &amp; Y. - Y. Wang" authorityYear="2022" class="Amphibia" family="Megophryidae" genus="Leptobrachella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Leptobrachella shimentaina" order="Anura" pageId="0" pageNumber="165" phylum="Chordata" rank="species" species="shimentaina">
<emphasis italics="true" pageId="0" pageNumber="165">Leptobrachella shimentaina</emphasis>
</taxonomicName>
sp. nov.:
<emphasis bold="true" pageId="0" pageNumber="165">A1.</emphasis>
A female (SYS a004711 in situ; ventral view of the same
<normalizedToken originalValue="specimens">specimen's</normalizedToken>
hand [A2] and foot [A3];
<emphasis bold="true" pageId="0" pageNumber="165">B.</emphasis>
Male SYS a007685 in situ;
<emphasis bold="true" pageId="0" pageNumber="165">C.</emphasis>
Male SYS a007686 in situ.
</paragraph>
</caption>
<caption doi="10.3897/zse.98.73162.figure6" httpUri="https://binary.pensoft.net/fig/692040" pageId="0" pageNumber="165" start="Figure 6" startId="F6">
<paragraph pageId="0" pageNumber="165">
<emphasis bold="true" pageId="0" pageNumber="165">Figure 6.</emphasis>
Morphological features of a paraype (SYS a007687) of
<taxonomicName authorityName="J. Wang, Z. - T. Lyu &amp; Y. - Y. Wang" authorityYear="2022" class="Amphibia" family="Megophryidae" genus="Leptobrachella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Leptobrachella shimentaina" order="Anura" pageId="0" pageNumber="165" phylum="Chordata" rank="species" species="shimentaina">
<emphasis italics="true" pageId="0" pageNumber="165">Leptobrachella shimentaina</emphasis>
</taxonomicName>
sp. nov. in life.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="165" type="distribution">
<paragraph pageId="0" pageNumber="165">Distribution and ecology.</paragraph>
<paragraph pageId="0" pageNumber="165">
<taxonomicName authorityName="J. Wang, Z. - T. Lyu &amp; Y. - Y. Wang" authorityYear="2022" class="Amphibia" family="Megophryidae" genus="Leptobrachella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Leptobrachella shimentaina" order="Anura" pageId="0" pageNumber="165" phylum="Chordata" rank="species" species="shimentaina">
<emphasis italics="true" pageId="0" pageNumber="165">Leptobrachella shimentaina</emphasis>
</taxonomicName>
sp. nov. is known from its type locality Shimentai Nature Reserve, Yingde City, and the adjacent Luokeng Nature Reserve, Shaoguan City, Guangdong, China (Fig.
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Localities of Leptobrachella shimentaina sp. nov. (samples ID 1 - 10, Shimentai Nature Reserve, Guangdong, China; sample ID 11, Luokeng Nature Reserve, Guangdong, China); Leptobrachella bijie (samples ID 38 - 40, Zhaozishan Nature Reserve, Guizhou, China; sample ID 41, Chishui, Guizhou, China; samples ID 42 - 43, Huagaoxi Nature Reserve, Guizhou, China); Leptobrachella chishuiensis (samples ID 44 - 46, Chishui, Guizhou, China); Leptobrachella alpina (samples ID 50 - 51, Mt Huangcaoling, Yunnan, China; sample ID 52, Pu'er, Yunnan, China); and Leptobrachella purpurus (samples ID 53 - 54, Yingjiang, Yunnan, China). Numbers correspond to the ID numbers in Suppl. material 1: Table S 1." figureDoi="10.3897/zse.98.73162.figure1" httpUri="https://binary.pensoft.net/fig/692035" pageId="0" pageNumber="165">1</figureCitation>
; ca. 300-600 m a.s.l.). In the Hengshitang Protection Station of the Shimentai Nature Reserve, the new species is sympatric with
<taxonomicName genus="L." lsidName="L. mangshanensis" pageId="0" pageNumber="165" rank="species" species="mangshanensis">
<emphasis italics="true" pageId="0" pageNumber="165">L. mangshanensis</emphasis>
</taxonomicName>
. The new species inhabits primitive clear-water rocky streams surrounded by broad-leaved forest in karst landforms or granite landforms. From April to June, males were observed calling while perched on the rocks or under rocks in flowing streams, and females possessed mature oocytes.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="165" type="comparisons">
<paragraph pageId="0" pageNumber="165">Comparisons.</paragraph>
<paragraph pageId="0" pageNumber="165">
In our phylogenetic analyses (Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Bayesian inference tree derived from partial DNA sequences of the mitochondrial 16 S r RNA gene. Numbers before slashes indicate Bayesian posterior probabilities and numbers after slashes are bootstrap support for maximum likelihood (1000 replicates) analyses. The number at the branch terminal corresponds to the ID in Suppl. material 1: TableS 1." figureDoi="10.3897/zse.98.73162.figure2" httpUri="https://binary.pensoft.net/fig/692036" pageId="0" pageNumber="165">2</figureCitation>
, Table
<tableCitation captionStart="Table 3" captionStartId="T3" captionText="Table 3. Comparisons of selected diagnostic characters for the new species and its phylogenetically related congeners." httpUri="http://table.plazi.org/id/DDA4DD2B6338427D7057DF55FBCAD192" pageId="0" pageNumber="165" tableUuid="DDA4DD2B6338427D7057DF55FBCAD192">3</tableCitation>
)
<taxonomicName authorityName="J. Wang, Z. - T. Lyu &amp; Y. - Y. Wang" authorityYear="2022" class="Amphibia" family="Megophryidae" genus="Leptobrachella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Leptobrachella shimentaina" order="Anura" pageId="0" pageNumber="165" phylum="Chordata" rank="species" species="shimentaina">
<emphasis italics="true" pageId="0" pageNumber="165">Leptobrachella shimentaina</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="165">sp. nov.</emphasis>
clusters with
<taxonomicName genus="L." lsidName="L. bashaensis" pageId="0" pageNumber="165" rank="species" species="bashaensis">
<emphasis italics="true" pageId="0" pageNumber="165">L. bashaensis</emphasis>
</taxonomicName>
,
<taxonomicName genus="L." lsidName="L. laui" pageId="0" pageNumber="165" rank="species" species="laui">
<emphasis italics="true" pageId="0" pageNumber="165">L. laui</emphasis>
</taxonomicName>
,
<taxonomicName genus="L." lsidName="L. liui" pageId="0" pageNumber="165" rank="species" species="liui">
<emphasis italics="true" pageId="0" pageNumber="165">L. liui</emphasis>
</taxonomicName>
,
<taxonomicName genus="L." lsidName="L. mangshanensis" pageId="0" pageNumber="165" rank="species" species="mangshanensis">
<emphasis italics="true" pageId="0" pageNumber="165">L. mangshanensis</emphasis>
</taxonomicName>
,
<taxonomicName genus="L." lsidName="L. maoershanensis" pageId="0" pageNumber="165" rank="species" species="maoershanensis">
<emphasis italics="true" pageId="0" pageNumber="165">L. maoershanensis</emphasis>
</taxonomicName>
, and
<taxonomicName genus="L." lsidName="L. yunkaiensis" pageId="0" pageNumber="165" rank="species" species="yunkaiensis">
<emphasis italics="true" pageId="0" pageNumber="165">L. yunkaiensis</emphasis>
</taxonomicName>
, thus, these six species constitute the most important comparisons, for the purpose of diagnosis and recognition of the new species. In having wide fringes along lateral surfaces of toes in males, it can be distinguished from
<taxonomicName genus="L." lsidName="L. bashaensis" pageId="0" pageNumber="165" rank="species" species="bashaensis">
<emphasis italics="true" pageId="0" pageNumber="165">L. bashaensis</emphasis>
</taxonomicName>
,
<taxonomicName genus="L." lsidName="L. mangshanensis" pageId="0" pageNumber="165" rank="species" species="mangshanensis">
<emphasis italics="true" pageId="0" pageNumber="165">L. mangshanensis</emphasis>
</taxonomicName>
and
<taxonomicName genus="L." lsidName="L. maoershanensis" pageId="0" pageNumber="165" rank="species" species="maoershanensis">
<emphasis italics="true" pageId="0" pageNumber="165">L. maoershanensis</emphasis>
</taxonomicName>
(vs. lateral fringes narrow); by ventral coloration, grayish pink with distinct hazy brown speckling on chest and ventrolateral flanks; it can be distinguished from
<taxonomicName genus="L." lsidName="L. bashaensis" pageId="0" pageNumber="165" rank="species" species="bashaensis">
<emphasis italics="true" pageId="0" pageNumber="165">L. bashaensis</emphasis>
</taxonomicName>
(vs. creamy-white chest and belly with irregular black spots),
<taxonomicName genus="L." lsidName="L. laui" pageId="0" pageNumber="165" rank="species" species="laui">
<emphasis italics="true" pageId="0" pageNumber="165">L. laui</emphasis>
</taxonomicName>
(vs. creamy white with dark brown dusting on ventrolateral flanks),
<taxonomicName genus="L." lsidName="L. liui" pageId="0" pageNumber="165" rank="species" species="liui">
<emphasis italics="true" pageId="0" pageNumber="165">L. liui</emphasis>
</taxonomicName>
(vs. creamy white with dark brown spots on chest and ventrolateral flanks),
<taxonomicName genus="L." lsidName="L. maoershanensis" pageId="0" pageNumber="165" rank="species" species="maoershanensis">
<emphasis italics="true" pageId="0" pageNumber="165">L. maoershanensis</emphasis>
</taxonomicName>
(vs. creamy white chest and belly with irregular black spots), and
<taxonomicName genus="L." lsidName="L. yunkaiensis" pageId="0" pageNumber="165" rank="species" species="yunkaiensis">
<emphasis italics="true" pageId="0" pageNumber="165">L. yunkaiensis</emphasis>
</taxonomicName>
(vs. belly pink with distinct or indistinct speckling); by having dorsal skin with round granular tubercles and glandular folds, the new species can be further distinguished from
<taxonomicName genus="L." lsidName="L. laui" pageId="0" pageNumber="165" rank="species" species="laui">
<emphasis italics="true" pageId="0" pageNumber="165">L. laui</emphasis>
</taxonomicName>
(vs. dorsal skin lacking glandular folds),
<taxonomicName genus="L." lsidName="L. mangshanensis" pageId="0" pageNumber="165" rank="species" species="mangshanensis">
<emphasis italics="true" pageId="0" pageNumber="165">L. mangshanensis</emphasis>
</taxonomicName>
(vs. dorsal skin almost smooth with tiny transparent spines), and
<taxonomicName genus="L." lsidName="L. yunkaiensis" pageId="0" pageNumber="165" rank="species" species="yunkaiensis">
<emphasis italics="true" pageId="0" pageNumber="165">L. yunkaiensis</emphasis>
</taxonomicName>
(vs. dorsal skin shagreened with short skin ridges and raised warts); by dense small raised tubercles present on surface of chest and both sides of abdomen; it can be further distinguished from
<taxonomicName genus="L." lsidName="L. mangshanensis" pageId="0" pageNumber="165" rank="species" species="mangshanensis">
<emphasis italics="true" pageId="0" pageNumber="165">L. mangshanensis</emphasis>
</taxonomicName>
,
<taxonomicName genus="L." lsidName="L. liui" pageId="0" pageNumber="165" rank="species" species="liui">
<emphasis italics="true" pageId="0" pageNumber="165">L. liui</emphasis>
</taxonomicName>
,
<taxonomicName genus="L." lsidName="L. yunkaiensis" pageId="0" pageNumber="165" rank="species" species="yunkaiensis">
<emphasis italics="true" pageId="0" pageNumber="165">L. yunkaiensis</emphasis>
</taxonomicName>
,
<taxonomicName genus="L." lsidName="L. maoershanensis" pageId="0" pageNumber="165" rank="species" species="maoershanensis">
<emphasis italics="true" pageId="0" pageNumber="165">L. maoershanensis</emphasis>
</taxonomicName>
and
<taxonomicName genus="L." lsidName="L. laui" pageId="0" pageNumber="165" rank="species" species="laui">
<emphasis italics="true" pageId="0" pageNumber="165">L. laui</emphasis>
</taxonomicName>
(vs. ventral surface smooth); by having longitudinal ridges under toes continuous, with constrictions at interphalangeal articulations, it can be distinguished from
<taxonomicName genus="L." lsidName="L. laui" pageId="0" pageNumber="165" rank="species" species="laui">
<emphasis italics="true" pageId="0" pageNumber="165">L. laui</emphasis>
</taxonomicName>
(vs. longitudinal ridges without constrictions at interphalangeal articulations),
<taxonomicName genus="L." lsidName="L. liui" pageId="0" pageNumber="165" rank="species" species="liui">
<emphasis italics="true" pageId="0" pageNumber="165">L. liui</emphasis>
</taxonomicName>
(vs. longitudinal ridges interrupted at interphalangeal articulations),
<taxonomicName genus="L." lsidName="L. maoershanensis" pageId="0" pageNumber="165" rank="species" species="maoershanensis">
<emphasis italics="true" pageId="0" pageNumber="165">L. maoershanensis</emphasis>
</taxonomicName>
(vs. longitudinal ridges indistinct and not interrupted at interphalangeal articulations), and
<taxonomicName genus="L." lsidName="L. yunkaiensis" pageId="0" pageNumber="165" rank="species" species="yunkaiensis">
<emphasis italics="true" pageId="0" pageNumber="165">L. yunkaiensis</emphasis>
</taxonomicName>
(vs. longitudinal ridges without constrictions at interphalangeal articulations).
</paragraph>
<caption ID-Table-UUID="DDA4DD2B6338427D7057DF55FBCAD192" httpUri="http://table.plazi.org/id/DDA4DD2B6338427D7057DF55FBCAD192" pageId="0" pageNumber="165" start="Table 3" startId="T3">
<paragraph pageId="0" pageNumber="165">
<emphasis bold="true" pageId="0" pageNumber="165">Table 3.</emphasis>
Comparisons of selected diagnostic characters for the new species and its phylogenetically related congeners.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="165">
<table pageId="0" pageNumber="165">
<tr pageId="0" pageNumber="165">
<th colspan="1" pageId="0" pageNumber="165" rowspan="1">
<taxonomicName authorityName="Smith" authorityYear="1925" class="Amphibia" family="Megophryidae" genus="Leptobrachella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Leptobrachella" order="Anura" pageId="0" pageNumber="165" phylum="Chordata" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="165">Leptobrachella</emphasis>
</taxonomicName>
spp.
</th>
<th colspan="1" pageId="0" pageNumber="165" rowspan="1">Male SVL (mm)</th>
<th colspan="1" pageId="0" pageNumber="165" rowspan="1">Toes webbing</th>
<th colspan="1" pageId="0" pageNumber="165" rowspan="1">Fringes on toes</th>
<th colspan="1" pageId="0" pageNumber="165" rowspan="1">Longitudinal ridges under toes (interrupt + / not interrupt -)</th>
<th colspan="1" pageId="0" pageNumber="165" rowspan="1">Ventral coloration</th>
<th colspan="1" pageId="0" pageNumber="165" rowspan="1">Dorsal skin texture</th>
</tr>
<tr pageId="0" pageNumber="165">
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">
<taxonomicName genus="L." lsidName="L. shimentaina" pageId="0" pageNumber="165" rank="species" species="shimentaina">
<emphasis italics="true" pageId="0" pageNumber="165">L. shimentaina</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="165">sp. nov.</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">26.4-28.9</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">Rudimentary</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">Wide in males</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">Grayish pink with distinct hazy brown speckling on chest and ventrolateral flanks</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">Round granular tubercles with glandular folds</td>
</tr>
<tr pageId="0" pageNumber="165">
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">
<taxonomicName genus="L." lsidName="L. laui" pageId="0" pageNumber="165" rank="species" species="laui">
<emphasis italics="true" pageId="0" pageNumber="165">L. laui</emphasis>
</taxonomicName>
</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">24.8-26.7</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">Rudimentary</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">Wide</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">Creamy white with dark brown dusting on ventrolateral flanks</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">Round granular tubercles</td>
</tr>
<tr pageId="0" pageNumber="165">
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">
<taxonomicName genus="L." lsidName="L. liui" pageId="0" pageNumber="165" rank="species" species="liui">
<emphasis italics="true" pageId="0" pageNumber="165">L. liui</emphasis>
</taxonomicName>
</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">23.0-28.7</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">Rudimentary</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">Wide</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">+</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">Gray white to creamy white with dark brown spots on chest and ventrolateral flanks</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">Shagreened with numerous small tubercles, round warts and sparse short skin ridges</td>
</tr>
<tr pageId="0" pageNumber="165">
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">
<taxonomicName genus="L." lsidName="L. mangshanensis" pageId="0" pageNumber="165" rank="species" species="mangshanensis">
<emphasis italics="true" pageId="0" pageNumber="165">L. mangshanensis</emphasis>
</taxonomicName>
</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">22.2-27.8</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">Rudimentary</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">Narrow</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">Creamy white belly, scattered with white speckles</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">Almost smooth with tiny transparent spines, small tubercles and sparse short skin ridges</td>
</tr>
<tr pageId="0" pageNumber="165">
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">
<taxonomicName genus="L." lsidName="L. maoershanensis" pageId="0" pageNumber="165" rank="species" species="maoershanensis">
<emphasis italics="true" pageId="0" pageNumber="165">L. maoershanensis</emphasis>
</taxonomicName>
</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">25.2-30.4</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">Rudimentary</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">Narrow</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">Creamy white chest and belly with irregular black spots</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">With longitudinal folds</td>
</tr>
<tr pageId="0" pageNumber="165">
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">
<taxonomicName genus="L." lsidName="L. bashaensis" pageId="0" pageNumber="165" rank="species" species="bashaensis">
<emphasis italics="true" pageId="0" pageNumber="165">L. bashaensis</emphasis>
</taxonomicName>
</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">22.9-25.6</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">Rudimentary</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">Narrow</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">Creamy-white chest and belly with irregular black spots</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">Slightly shagreened with small tubercles</td>
</tr>
<tr pageId="0" pageNumber="165">
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">
<taxonomicName genus="L." lsidName="L. yunkaiensis" pageId="0" pageNumber="165" rank="species" species="yunkaiensis">
<emphasis italics="true" pageId="0" pageNumber="165">L. yunkaiensis</emphasis>
</taxonomicName>
</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">25.9-29.3</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">Rudimentary</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">Wide</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">Belly pink with distinct or indistinct speckles</td>
<td colspan="1" pageId="0" pageNumber="165" rowspan="1">Shagreened with short skin ridges and raised warts</td>
</tr>
</table>
</paragraph>
<paragraph pageId="0" pageNumber="165">
Compared with the 26 known congeners of the genus
<taxonomicName authorityName="Smith" authorityYear="1925" class="Amphibia" family="Megophryidae" genus="Leptobrachella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Leptobrachella" order="Anura" pageId="0" pageNumber="165" phylum="Chordata" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="165">Leptobrachella</emphasis>
</taxonomicName>
occurring south of the Isthmus of Kra, by the presence of supra-axillary and ventrolateral glands,
<taxonomicName genus="L." lsidName="L. shimentaina" pageId="0" pageNumber="165" rank="species" species="shimentaina">
<emphasis italics="true" pageId="0" pageNumber="165">L. shimentaina</emphasis>
</taxonomicName>
sp. nov. can easily be distinguished from
<taxonomicName genus="L." lsidName="L. arayai" pageId="0" pageNumber="165" rank="species" species="arayai">
<emphasis italics="true" pageId="0" pageNumber="165">L. arayai</emphasis>
</taxonomicName>
,
<taxonomicName genus="L." lsidName="L. dringi" pageId="0" pageNumber="165" rank="species" species="dringi">
<emphasis italics="true" pageId="0" pageNumber="165">L. dringi</emphasis>
</taxonomicName>
,
<taxonomicName genus="L." lsidName="L. fritinniens" pageId="0" pageNumber="165" rank="species" species="fritinniens">
<emphasis italics="true" pageId="0" pageNumber="165">L. fritinniens</emphasis>
</taxonomicName>
,
<taxonomicName genus="L." lsidName="L. gracilis" pageId="0" pageNumber="165" rank="species" species="gracilis">
<emphasis italics="true" pageId="0" pageNumber="165">L. gracilis</emphasis>
</taxonomicName>
,
<taxonomicName genus="L." lsidName="L. hamidi" pageId="0" pageNumber="165" rank="species" species="hamidi">
<emphasis italics="true" pageId="0" pageNumber="165">L. hamidi</emphasis>
</taxonomicName>
,
<taxonomicName genus="L." lsidName="L. heteropus" pageId="0" pageNumber="165" rank="species" species="heteropus">
<emphasis italics="true" pageId="0" pageNumber="165">L. heteropus</emphasis>
</taxonomicName>
,
<taxonomicName genus="L." lsidName="L. kajangensis" pageId="0" pageNumber="165" rank="species" species="kajangensis">
<emphasis italics="true" pageId="0" pageNumber="165">L. kajangensis</emphasis>
</taxonomicName>
,
<taxonomicName genus="L." lsidName="L. kecil" pageId="0" pageNumber="165" rank="species" species="kecil">
<emphasis italics="true" pageId="0" pageNumber="165">L. kecil</emphasis>
</taxonomicName>
,
<taxonomicName genus="L." lsidName="L. marmorata" pageId="0" pageNumber="165" rank="species" species="marmorata">
<emphasis italics="true" pageId="0" pageNumber="165">L. marmorata</emphasis>
</taxonomicName>
,
<taxonomicName genus="L." lsidName="L. melanoleuca" pageId="0" pageNumber="165" rank="species" species="melanoleuca">
<emphasis italics="true" pageId="0" pageNumber="165">L. melanoleuca</emphasis>
</taxonomicName>
,
<taxonomicName genus="L." lsidName="L. maura" pageId="0" pageNumber="165" rank="species" species="maura">
<emphasis italics="true" pageId="0" pageNumber="165">L. maura</emphasis>
</taxonomicName>
,
<taxonomicName genus="L." lsidName="L. picta" pageId="0" pageNumber="165" rank="species" species="picta">
<emphasis italics="true" pageId="0" pageNumber="165">L. picta</emphasis>
</taxonomicName>
,
<taxonomicName genus="L." lsidName="L. platycephala" pageId="0" pageNumber="165" rank="species" species="platycephala">
<emphasis italics="true" pageId="0" pageNumber="165">L. platycephala</emphasis>
</taxonomicName>
,
<taxonomicName genus="L." lsidName="L. sabahmontana" pageId="0" pageNumber="165" rank="species" species="sabahmontana">
<emphasis italics="true" pageId="0" pageNumber="165">L. sabahmontana</emphasis>
</taxonomicName>
, and
<taxonomicName genus="L." lsidName="L. sola" pageId="0" pageNumber="165" rank="species" species="sola">
<emphasis italics="true" pageId="0" pageNumber="165">L. sola</emphasis>
</taxonomicName>
, all of which lack supra-axillary and ventrolateral glands; and by its distinctly larger male body size, SVL 26.4-28.9 mm,
<taxonomicName genus="L." lsidName="L. shimentaina" pageId="0" pageNumber="165" rank="species" species="shimentaina">
<emphasis italics="true" pageId="0" pageNumber="165">L. shimentaina</emphasis>
</taxonomicName>
sp. nov. differs from the smaller
<taxonomicName genus="L." lsidName="L. baluensis" pageId="0" pageNumber="165" rank="species" species="baluensis">
<emphasis italics="true" pageId="0" pageNumber="165">L. baluensis</emphasis>
</taxonomicName>
(14.9-15.9),
<taxonomicName genus="L." lsidName="L. brevicrus" pageId="0" pageNumber="165" rank="species" species="brevicrus">
<emphasis italics="true" pageId="0" pageNumber="165">L. brevicrus</emphasis>
</taxonomicName>
(17.1-17.8),
<taxonomicName genus="L." lsidName="L. bondangensis" pageId="0" pageNumber="165" rank="species" species="bondangensis">
<emphasis italics="true" pageId="0" pageNumber="165">L. bondangensis</emphasis>
</taxonomicName>
(17.8),
<taxonomicName genus="L." lsidName="L. fusca" pageId="0" pageNumber="165" rank="species" species="fusca">
<emphasis italics="true" pageId="0" pageNumber="165">L. fusca</emphasis>
</taxonomicName>
(16.3),
<taxonomicName genus="L." lsidName="L. itiokai" pageId="0" pageNumber="165" rank="species" species="itiokai">
<emphasis italics="true" pageId="0" pageNumber="165">L. itiokai</emphasis>
</taxonomicName>
(15.2-16.7s),
<taxonomicName genus="L." lsidName="L. juliandringi" pageId="0" pageNumber="165" rank="species" species="juliandringi">
<emphasis italics="true" pageId="0" pageNumber="165">L. juliandringi</emphasis>
</taxonomicName>
(17.0-17.2),
<taxonomicName genus="L." lsidName="L. mjobergi" pageId="0" pageNumber="165" rank="species" species="mjobergi">
<emphasis italics="true" pageId="0" pageNumber="165">L. mjobergi</emphasis>
</taxonomicName>
(15.7-19.0),
<taxonomicName genus="L." lsidName="L. natunae" pageId="0" pageNumber="165" rank="species" species="natunae">
<emphasis italics="true" pageId="0" pageNumber="165">L. natunae</emphasis>
</taxonomicName>
(17.6),
<taxonomicName genus="L." lsidName="L. parva" pageId="0" pageNumber="165" rank="species" species="parva">
<emphasis italics="true" pageId="0" pageNumber="165">L. parva</emphasis>
</taxonomicName>
(15.0-16.9), and
<taxonomicName genus="L." lsidName="L. palmata" pageId="0" pageNumber="165" rank="species" species="palmata">
<emphasis italics="true" pageId="0" pageNumber="165">L. palmata</emphasis>
</taxonomicName>
(14.4-16.8); the female of
<taxonomicName genus="L." lsidName="L. serasanae" pageId="0" pageNumber="165" rank="species" species="serasanae">
<emphasis italics="true" pageId="0" pageNumber="165">L. serasanae</emphasis>
</taxonomicName>
(16.9), is also smaller than
<taxonomicName genus="L." lsidName="L. shimentaina" pageId="0" pageNumber="165" rank="species" species="shimentaina">
<emphasis italics="true" pageId="0" pageNumber="165">L. shimentaina</emphasis>
</taxonomicName>
sp. nov. (females 30.1-30.7).
</paragraph>
<paragraph pageId="0" pageNumber="165">
For the remaining 60 members of the genus
<taxonomicName authorityName="Smith" authorityYear="1925" class="Amphibia" family="Megophryidae" genus="Leptobrachella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Leptobrachella" order="Anura" pageId="0" pageNumber="165" phylum="Chordata" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="165">Leptobrachella</emphasis>
</taxonomicName>
, males
<taxonomicName genus="L." lsidName="L. shimentaina" pageId="0" pageNumber="165" rank="species" species="shimentaina">
<emphasis italics="true" pageId="0" pageNumber="165">L. shimentaina</emphasis>
</taxonomicName>
sp. nov. (SVL 26.4-28.9 mm) differs from males of the larger
<taxonomicName genus="L." lsidName="L. chishuiensis" pageId="0" pageNumber="165" rank="species" species="chishuiensis">
<emphasis italics="true" pageId="0" pageNumber="165">L. chishuiensis</emphasis>
</taxonomicName>
(30.8-33.4),
<taxonomicName genus="L." lsidName="L. damingshanensis" pageId="0" pageNumber="165" rank="species" species="damingshanensis">
<emphasis italics="true" pageId="0" pageNumber="165">L. damingshanensis</emphasis>
</taxonomicName>
(33.6-34.4),
<taxonomicName genus="L." lsidName="L. eos" pageId="0" pageNumber="165" rank="species" species="eos">
<emphasis italics="true" pageId="0" pageNumber="165">L. eos</emphasis>
</taxonomicName>
(33.1-34.7),
<taxonomicName genus="L." lsidName="L. graminicola" pageId="0" pageNumber="165" rank="species" species="graminicola">
<emphasis italics="true" pageId="0" pageNumber="165">L. graminicola</emphasis>
</taxonomicName>
(23.1-24.6),
<taxonomicName genus="L." lsidName="L. neangi" pageId="0" pageNumber="165" rank="species" species="neangi">
<emphasis italics="true" pageId="0" pageNumber="165">L. neangi</emphasis>
</taxonomicName>
(30.9),
<taxonomicName genus="L." lsidName="L. nahangensis" pageId="0" pageNumber="165" rank="species" species="nahangensis">
<emphasis italics="true" pageId="0" pageNumber="165">L. nahangensis</emphasis>
</taxonomicName>
(40.8),
<taxonomicName genus="L." lsidName="L. pyrrhops" pageId="0" pageNumber="165" rank="species" species="pyrrhops">
<emphasis italics="true" pageId="0" pageNumber="165">L. pyrrhops</emphasis>
</taxonomicName>
(30.8-34.3),
<taxonomicName genus="L." lsidName="L. sungi" pageId="0" pageNumber="165" rank="species" species="sungi">
<emphasis italics="true" pageId="0" pageNumber="165">L. sungi</emphasis>
</taxonomicName>
(48.3-52.7),
<taxonomicName genus="L." lsidName="L. tamdil" pageId="0" pageNumber="165" rank="species" species="tamdil">
<emphasis italics="true" pageId="0" pageNumber="165">L. tamdil</emphasis>
</taxonomicName>
(32.3), and
<taxonomicName genus="L." lsidName="L. zhangyapingi" pageId="0" pageNumber="165" rank="species" species="zhangyapingi">
<emphasis italics="true" pageId="0" pageNumber="165">L. zhangyapingi</emphasis>
</taxonomicName>
(45.8-52.5); and from the smaller
<taxonomicName genus="L." lsidName="L. aspera" pageId="0" pageNumber="165" rank="species" species="aspera">
<emphasis italics="true" pageId="0" pageNumber="165">L. aspera</emphasis>
</taxonomicName>
(22.4),
<taxonomicName genus="L." lsidName="L. applebyi" pageId="0" pageNumber="165" rank="species" species="applebyi">
<emphasis italics="true" pageId="0" pageNumber="165">L. applebyi</emphasis>
</taxonomicName>
(19.6-22.3),
<taxonomicName genus="L." lsidName="L. ardens" pageId="0" pageNumber="165" rank="species" species="ardens">
<emphasis italics="true" pageId="0" pageNumber="165">L. ardens</emphasis>
</taxonomicName>
(21.3-24.7),
<taxonomicName genus="L." lsidName="L. bidoupensis" pageId="0" pageNumber="165" rank="species" species="bidoupensis">
<emphasis italics="true" pageId="0" pageNumber="165">L. bidoupensis</emphasis>
</taxonomicName>
(18.5-25.4),
<taxonomicName genus="L." lsidName="L. feii" pageId="0" pageNumber="165" rank="species" species="feii">
<emphasis italics="true" pageId="0" pageNumber="165">L. feii</emphasis>
</taxonomicName>
(21.5-22.8),
<taxonomicName genus="L." lsidName="L. melica" pageId="0" pageNumber="165" rank="species" species="melica">
<emphasis italics="true" pageId="0" pageNumber="165">L. melica</emphasis>
</taxonomicName>
(19.5-22.7),
<taxonomicName genus="L." lsidName="L. murphyi" pageId="0" pageNumber="165" rank="species" species="murphyi">
<emphasis italics="true" pageId="0" pageNumber="165">L. murphyi</emphasis>
</taxonomicName>
(23.2-24.9),
<taxonomicName genus="L." lsidName="L. niveimontis" pageId="0" pageNumber="165" rank="species" species="niveimontis">
<emphasis italics="true" pageId="0" pageNumber="165">L. niveimontis</emphasis>
</taxonomicName>
(22.5-23.6),
<taxonomicName genus="L." lsidName="L. pluvialis" pageId="0" pageNumber="165" rank="species" species="pluvialis">
<emphasis italics="true" pageId="0" pageNumber="165">L. pluvialis</emphasis>
</taxonomicName>
(21.3-22.3), and
<taxonomicName genus="L." lsidName="L. rowleyae" pageId="0" pageNumber="165" rank="species" species="rowleyae">
<emphasis italics="true" pageId="0" pageNumber="165">L. rowleyae</emphasis>
</taxonomicName>
(23.4-25.4). Through its possession of toes with rudimentary webbing and with wide lateral fringes in males, the new species can be distinguished from
<taxonomicName genus="L." lsidName="L. bijie" pageId="0" pageNumber="165" rank="species" species="bijie">
<emphasis italics="true" pageId="0" pageNumber="165">L. bijie</emphasis>
</taxonomicName>
,
<taxonomicName genus="L." lsidName="L. dorsospina" pageId="0" pageNumber="165" rank="species" species="dorsospina">
<emphasis italics="true" pageId="0" pageNumber="165">L. dorsospina</emphasis>
</taxonomicName>
,
<taxonomicName genus="L." lsidName="L. flaviglandulosa" pageId="0" pageNumber="165" rank="species" species="flaviglandulosa">
<emphasis italics="true" pageId="0" pageNumber="165">L. flaviglandulosa</emphasis>
</taxonomicName>
,
<taxonomicName genus="L." lsidName="L. jinshaensis" pageId="0" pageNumber="165" rank="species" species="jinshaensis">
<emphasis italics="true" pageId="0" pageNumber="165">L. jinshaensis</emphasis>
</taxonomicName>
,
<taxonomicName genus="L." lsidName="L. purpuraventra" pageId="0" pageNumber="165" rank="species" species="purpuraventra">
<emphasis italics="true" pageId="0" pageNumber="165">L. purpuraventra</emphasis>
</taxonomicName>
,
<taxonomicName genus="L." lsidName="L. puhoatensis" pageId="0" pageNumber="165" rank="species" species="puhoatensis">
<emphasis italics="true" pageId="0" pageNumber="165">L. puhoatensis</emphasis>
</taxonomicName>
,
<taxonomicName genus="L." lsidName="L. shangsiensis" pageId="0" pageNumber="165" rank="species" species="shangsiensis">
<emphasis italics="true" pageId="0" pageNumber="165">L. shangsiensis</emphasis>
</taxonomicName>
,
<taxonomicName genus="L." lsidName="L. suiyangensis" pageId="0" pageNumber="165" rank="species" species="suiyangensis">
<emphasis italics="true" pageId="0" pageNumber="165">L. suiyangensis</emphasis>
</taxonomicName>
,
<taxonomicName genus="L." lsidName="L. tengchongensis" pageId="0" pageNumber="165" rank="species" species="tengchongensis">
<emphasis italics="true" pageId="0" pageNumber="165">L. tengchongensis</emphasis>
</taxonomicName>
,
<taxonomicName genus="L." lsidName="L. wuhuangmontis" pageId="0" pageNumber="165" rank="species" species="wuhuangmontis">
<emphasis italics="true" pageId="0" pageNumber="165">L. wuhuangmontis</emphasis>
</taxonomicName>
and
<taxonomicName genus="L." lsidName="L. yeae" pageId="0" pageNumber="165" rank="species" species="yeae">
<emphasis italics="true" pageId="0" pageNumber="165">L. yeae</emphasis>
</taxonomicName>
(lateral fringes narrow),
<taxonomicName genus="L." lsidName="L. bourreti" pageId="0" pageNumber="165" rank="species" species="bourreti">
<emphasis italics="true" pageId="0" pageNumber="165">L. bourreti</emphasis>
</taxonomicName>
,
<taxonomicName genus="L." lsidName="L. fuliginosa" pageId="0" pageNumber="165" rank="species" species="fuliginosa">
<emphasis italics="true" pageId="0" pageNumber="165">L. fuliginosa</emphasis>
</taxonomicName>
(lateral fringes weak),
<taxonomicName genus="L." lsidName="L. jinshaensis" pageId="0" pageNumber="165" rank="species" species="jinshaensis">
<emphasis italics="true" pageId="0" pageNumber="165">L. jinshaensis</emphasis>
</taxonomicName>
,
<taxonomicName genus="L." lsidName="L. kalonensis" pageId="0" pageNumber="165" rank="species" species="kalonensis">
<emphasis italics="true" pageId="0" pageNumber="165">L. kalonensis</emphasis>
</taxonomicName>
,
<taxonomicName genus="L." lsidName="L. maculosa" pageId="0" pageNumber="165" rank="species" species="maculosa">
<emphasis italics="true" pageId="0" pageNumber="165">L. maculosa</emphasis>
</taxonomicName>
,
<taxonomicName genus="L." lsidName="L. oshanensis" pageId="0" pageNumber="165" rank="species" species="oshanensis">
<emphasis italics="true" pageId="0" pageNumber="165">L. oshanensis</emphasis>
</taxonomicName>
,
<taxonomicName genus="L." lsidName="L. shiwandashan" pageId="0" pageNumber="165" rank="species" species="shiwandashan">
<emphasis italics="true" pageId="0" pageNumber="165">L. shiwandashan</emphasis>
</taxonomicName>
,
<taxonomicName genus="L." lsidName="L. tadungensis" pageId="0" pageNumber="165" rank="species" species="tadungensis">
<emphasis italics="true" pageId="0" pageNumber="165">L. tadungensis</emphasis>
</taxonomicName>
,
<taxonomicName genus="L." lsidName="L. ventripunctata" pageId="0" pageNumber="165" rank="species" species="ventripunctata">
<emphasis italics="true" pageId="0" pageNumber="165">L. ventripunctata</emphasis>
</taxonomicName>
(no webbing or lateral fringes),
<taxonomicName genus="L." lsidName="L. lateralis" pageId="0" pageNumber="165" rank="species" species="lateralis">
<emphasis italics="true" pageId="0" pageNumber="165">L. lateralis</emphasis>
</taxonomicName>
,
<taxonomicName genus="L." lsidName="L. namdongensis" pageId="0" pageNumber="165" rank="species" species="namdongensis">
<emphasis italics="true" pageId="0" pageNumber="165">L. namdongensis</emphasis>
</taxonomicName>
,
<taxonomicName genus="L." lsidName="L. macrops" pageId="0" pageNumber="165" rank="species" species="macrops">
<emphasis italics="true" pageId="0" pageNumber="165">L. macrops</emphasis>
</taxonomicName>
,
<taxonomicName genus="L." lsidName="L. minima" pageId="0" pageNumber="165" rank="species" species="minima">
<emphasis italics="true" pageId="0" pageNumber="165">L. minima</emphasis>
</taxonomicName>
,
<taxonomicName genus="L." lsidName="L. nyx" pageId="0" pageNumber="165" rank="species" species="nyx">
<emphasis italics="true" pageId="0" pageNumber="165">L. nyx</emphasis>
</taxonomicName>
(no lateral fringes), and
<taxonomicName genus="L." lsidName="L. pelodytoides" pageId="0" pageNumber="165" rank="species" species="pelodytoides">
<emphasis italics="true" pageId="0" pageNumber="165">L. pelodytoides</emphasis>
</taxonomicName>
(extensive webbing and narrow lateral fringes), and additionally from
<taxonomicName genus="L." lsidName="L. alpina" pageId="0" pageNumber="165" rank="species" species="alpina">
<emphasis italics="true" pageId="0" pageNumber="165">L. alpina</emphasis>
</taxonomicName>
,
<taxonomicName genus="L." lsidName="L. khasiorum" pageId="0" pageNumber="165" rank="species" species="khasiorum">
<emphasis italics="true" pageId="0" pageNumber="165">L. khasiorum</emphasis>
</taxonomicName>
,
<taxonomicName genus="L." lsidName="L. nokrekensis" pageId="0" pageNumber="165" rank="species" species="nokrekensis">
<emphasis italics="true" pageId="0" pageNumber="165">L. nokrekensis</emphasis>
</taxonomicName>
,
<taxonomicName genus="L." lsidName="L. yingjiangensis" pageId="0" pageNumber="165" rank="species" species="yingjiangensis">
<emphasis italics="true" pageId="0" pageNumber="165">L. yingjiangensis</emphasis>
</taxonomicName>
(ventral coloration creamy white) and
<taxonomicName genus="L." lsidName="L. purpurus" pageId="0" pageNumber="165" rank="species" species="purpurus">
<emphasis italics="true" pageId="0" pageNumber="165">L. purpurus</emphasis>
</taxonomicName>
(ventral coloration dull white) by having greyish pink ventral coloration. By having black spots on flanks, it further differs from
<taxonomicName genus="L." lsidName="L. aerea" pageId="0" pageNumber="165" rank="species" species="aerea">
<emphasis italics="true" pageId="0" pageNumber="165">L. aerea</emphasis>
</taxonomicName>
,
<taxonomicName genus="L." lsidName="L. botsfordi" pageId="0" pageNumber="165" rank="species" species="botsfordi">
<emphasis italics="true" pageId="0" pageNumber="165">L. botsfordi</emphasis>
</taxonomicName>
,
<taxonomicName genus="L." lsidName="L. crocea" pageId="0" pageNumber="165" rank="species" species="crocea">
<emphasis italics="true" pageId="0" pageNumber="165">L. crocea</emphasis>
</taxonomicName>
,
<taxonomicName genus="L." lsidName="L. firthi" pageId="0" pageNumber="165" rank="species" species="firthi">
<emphasis italics="true" pageId="0" pageNumber="165">L. firthi</emphasis>
</taxonomicName>
,
<taxonomicName genus="L." lsidName="L. isos" pageId="0" pageNumber="165" rank="species" species="isos">
<emphasis italics="true" pageId="0" pageNumber="165">L. isos</emphasis>
</taxonomicName>
,
<taxonomicName genus="L." lsidName="L. pallida" pageId="0" pageNumber="165" rank="species" species="pallida">
<emphasis italics="true" pageId="0" pageNumber="165">L. pallida</emphasis>
</taxonomicName>
,
<taxonomicName genus="L." lsidName="L. petrops" pageId="0" pageNumber="165" rank="species" species="petrops">
<emphasis italics="true" pageId="0" pageNumber="165">L. petrops</emphasis>
</taxonomicName>
, and
<taxonomicName genus="L." lsidName="L. tuberosa" pageId="0" pageNumber="165" rank="species" species="tuberosa">
<emphasis italics="true" pageId="0" pageNumber="165">L. tuberosa</emphasis>
</taxonomicName>
, all of which lack black spots on the flanks.
</paragraph>
</subSubSection>
</treatment>
</document>