627 lines
107 KiB
XML
627 lines
107 KiB
XML
<document id="EFEE00562B63E5423031FDFDD71FF38A" ID-CLB-Dataset="32193" ID-DOI="10.11646/zootaxa.4318.1.4" ID-GBIF-Dataset="b87d843d-1458-487b-ac2e-4071e7e66f62" ID-ISSN="1175-5326" ID-Zenodo-Dep="885700" ID-ZooBank="71699C18-6Fd3-4021-80Ce-5E1928575F67" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1504680327553" checkinUser="plazi" docAuthor="Will, Kipling" docDate="2017" docId="03F1AD5D4C79FFE2FF40FA9BFAEBF9C1" docLanguage="en" docName="zootaxa.4318.1.4.pdf" docOrigin="Zootaxa 4318 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Praeanchodemus Will 2017, gen. n." docType="treatment" docVersion="7" lastPageNumber="114" masterDocId="FFC8D5254C78FFE6FFD7FF8EFFF3FFEC" masterDocTitle="Description of the first flightless platynine ground beetle preserved in Baltic amber (Coleoptera: Carabidae)" masterLastPageNumber="122" masterPageNumber="110" pageNumber="111" updateTime="1698467919947" updateUser="ExternalLinkService">
|
||
<mods:mods id="9A402DBBFF4543C6680A6E54882348F8" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="B95435D2319EE055DAA80792A49C00A4">
|
||
<mods:title id="A06AC9E87CD79EE90C3E6E27E39DA10D">Description of the first flightless platynine ground beetle preserved in Baltic amber (Coleoptera: Carabidae)</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="27CFDB6912E728C9A8389A8DAF6CD4C9" type="personal">
|
||
<mods:role id="343B382B7A7B946B3E8BBA73B3E7718F">
|
||
<mods:roleTerm id="3F9E64E64EEDF45AB14985242D69041D">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="BD0A52C321010016D5406290D4462768">Will, Kipling</mods:namePart>
|
||
</mods:name>
|
||
<mods:typeOfResource id="F5B19BECB4A1CAEF90C187A4E35582E2">text</mods:typeOfResource>
|
||
<mods:relatedItem id="092AAA845E31264053DE15DA621248B4" type="host">
|
||
<mods:titleInfo id="0F658C7D5023DDA81DA65F2C8D517245">
|
||
<mods:title id="F70B3E1B3FE375156243AACEF389C670">Zootaxa</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="9F3AE436E74E71B70DFF3A629C3DEA33">
|
||
<mods:date id="D7CDC67E8E3E80449188DA8E5BFC59C0">2017</mods:date>
|
||
<mods:detail id="679AD942F5ECFC125C02FD1548FD2D46" type="pubDate">
|
||
<mods:number id="F46B4D9DB62173F667B535E71E65F9C5">2017-06-06</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="AC94EDF95A8BCAF5867BF7D63E9CF3CD" type="volume">
|
||
<mods:number id="DE5C043F54A37C21EB2698C2EF60B709">4318</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="3DAB263E24F75A420583F8895C5CBADB" type="issue">
|
||
<mods:number id="74BA8C10F4C9DD2626C5A107E789897F">1</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="C09C8387375AAFA68C6DD8E2509F4473" unit="page">
|
||
<mods:start id="C6F52AEDD04DBACB5FFBEE01473EDE18">110</mods:start>
|
||
<mods:end id="8631DCA1AAF95B9EF204EFDE0DC1B5C9">122</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:classification id="3E049C5BA497CA85F7712D24ECC1F0E8">journal article</mods:classification>
|
||
<mods:identifier id="7214662029A1FC4AD8080B2A59C9008F" type="CLB-Dataset">32193</mods:identifier>
|
||
<mods:identifier id="B5147635ECD56E172D873958DDC240E6" type="DOI">10.11646/zootaxa.4318.1.4</mods:identifier>
|
||
<mods:identifier id="BEA5808CACAC5F186AE9D580FA09845D" type="GBIF-Dataset">b87d843d-1458-487b-ac2e-4071e7e66f62</mods:identifier>
|
||
<mods:identifier id="6ACB55901235F385439DD1AC854EAA37" type="ISSN">1175-5326</mods:identifier>
|
||
<mods:identifier id="A5D7A13A52AF9E3FD2D2CEFA3B99152C" type="Zenodo-Dep">885700</mods:identifier>
|
||
<mods:identifier id="F3FF48D26347F758B316AC82871EF416" type="ZooBank">71699C18-6Fd3-4021-80Ce-5E1928575F67</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="03F1AD5D4C79FFE2FF40FA9BFAEBF9C1" ID-DOI="http://doi.org/10.5281/zenodo.6049715" ID-GBIF-Taxon="133186319" ID-Zenodo-Dep="6049715" LSID="urn:lsid:plazi:treatment:03F1AD5D4C79FFE2FF40FA9BFAEBF9C1" httpUri="http://treatment.plazi.org/id/03F1AD5D4C79FFE2FF40FA9BFAEBF9C1" lastPageId="4" lastPageNumber="114" pageId="1" pageNumber="111">
|
||
<subSubSection id="C3424FC04C79FFE7FF40FA9BFE31FAC3" box="[151,450,1301,1327]" pageId="1" pageNumber="111" type="nomenclature">
|
||
<paragraph id="8BE71C4B4C79FFE7FF40FA9BFE31FAC3" blockId="1.[151,450,1301,1327]" box="[151,450,1301,1327]" pageId="1" pageNumber="111">
|
||
<heading id="D0AFAB274C79FFE7FF40FA9BFE31FAC3" bold="true" box="[151,450,1301,1327]" fontSize="11" level="1" pageId="1" pageNumber="111" reason="1">
|
||
<emphasis id="B92CC0594C79FFE7FF40FA9BFE31FAC3" bold="true" box="[151,450,1301,1327]" pageId="1" pageNumber="111">
|
||
<taxonomicName id="4C5867C84C79FFE7FF40FA9BFE9AFAC3" ID-CoL="8LTCY" authority="Will, 2017" authorityName="Will" authorityYear="2017" box="[151,361,1301,1327]" class="Insecta" family="Carabidae" genus="Praeanchodemus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="111" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B92CC0594C79FFE7FF40FA9BFE9AFAC3" bold="true" box="[151,361,1301,1327]" italics="true" pageId="1" pageNumber="111">Praeanchodemus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21F7D224C79FFE7FEA7FA9BFE31FAC3" box="[368,450,1301,1327]" pageId="1" pageNumber="111" rank="genus">gen. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3424FC04C79FFE7FF40FAD2FB21FA99" box="[151,1234,1372,1397]" pageId="1" pageNumber="111" type="materials_examined">
|
||
<paragraph id="8BE71C4B4C79FFE7FF40FAD2FB21FA99" blockId="1.[151,1437,1372,2009]" box="[151,1234,1372,1397]" pageId="1" pageNumber="111">
|
||
<emphasis id="B92CC0594C79FFE7FF40FAD2FEC1FA99" bold="true" box="[151,306,1372,1397]" pageId="1" pageNumber="111">Type species.</emphasis>
|
||
<taxonomicName id="4C5867C84C79FFE7FEEEFAD2FD44FA98" authority="Sp." authorityName="Sp." box="[313,695,1372,1397]" class="Insecta" family="Carabidae" genus="Praeanchodemus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="111" phylum="Arthropoda" rank="species" species="punctaticeps">
|
||
<emphasis id="B92CC0594C79FFE7FEEEFAD2FD7CFA98" box="[313,655,1372,1397]" italics="true" pageId="1" pageNumber="111">Praeanchodemus punctaticeps</emphasis>
|
||
<taxonomicNameLabel id="A21F7D224C79FFE7FD40FAD2FD44FA98" box="[663,695,1372,1396]" pageId="1" pageNumber="111" rank="species">Sp.</taxonomicNameLabel>
|
||
</taxonomicName>
|
||
n., foSSil SpecieS from the Eocene Baltic amber.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3424FC04C79FFE4FF10FAF1FCF9FA0B" lastPageId="2" lastPageNumber="112" pageId="1" pageNumber="111" type="description">
|
||
<paragraph id="8BE71C4B4C79FFE7FF10FAF1FAEAFA0C" blockId="1.[151,1437,1372,2009]" pageId="1" pageNumber="111">
|
||
<emphasis id="B92CC0594C79FFE7FF10FAF1FEAAFA74" bold="true" box="[199,345,1407,1432]" pageId="1" pageNumber="111">Description.</emphasis>
|
||
<collectingCountry id="F34F5CDB4C79FFE7FEB5FA0EFE72FA74" box="[354,385,1408,1432]" name="American Samoa" pageId="1" pageNumber="111">AS</collectingCountry>
|
||
the new genuS iS monotypic and baSed on a Single Specimen, neither the interSpecific nor intraSpecific variability of characterS iS known. Herein, we deScribe the new taxon in detail under the genuS deScription and only include colour and dimenSionS of the body partS in the SpecieS’ deScription below.
|
||
</paragraph>
|
||
<paragraph id="8BE71C4B4C79FFE7FF10FA62FB47F9E9" blockId="1.[151,1437,1372,2009]" box="[199,1204,1516,1541]" pageId="1" pageNumber="111">
|
||
Body length: Rather Small compared with average in
|
||
<taxonomicName id="4C5867C84C79FFE7FCC9FA62FC7BF9E9" authorityName="Bonelli" authorityYear="1810" box="[798,904,1516,1541]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="111" phylum="Arthropoda" rank="tribe" tribe="Platynini">Platynini</taxonomicName>
|
||
:
|
||
<quantity id="4CA0B1AE4C79FFE7FC46FA62FC1BF9E8" box="[913,1000,1516,1541]" metricMagnitude="-3" metricUnit="m" metricValue="5.4" pageId="1" pageNumber="111" unit="mm" value="5.4">5.4 mm</quantity>
|
||
in type Specimen.
|
||
</paragraph>
|
||
<paragraph id="8BE71C4B4C79FFE7FF10F981FBB4F978" blockId="1.[151,1437,1372,2009]" pageId="1" pageNumber="111">MicroSculpture: On head markedly contraSting with microSculpture patternS on pronotum and elytra: DorSal Surface of head with large and deeply engraved lineS forming iSodiametric SculpticellS; pronotum and elytra with much Smaller SculpticellS that are diStinct only on SideS, near baSe of pronotum, and within elytral Striae; pronotal diSc and elytral intervalS without viSible microSculpture even at magnification 100x.</paragraph>
|
||
<paragraph id="8BE71C4B4C79FFE4FF10F911FC6AFF38" blockId="1.[151,1437,1372,2009]" lastBlockId="2.[151,1437,151,2015]" lastPageId="2" lastPageNumber="112" pageId="1" pageNumber="111">
|
||
Head: Size and proportionS averaged for
|
||
<taxonomicName id="4C5867C84C79FFE7FD4EF911FCF2F954" authorityName="Bonelli" authorityYear="1810" box="[665,769,1695,1720]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="111" phylum="Arthropoda" rank="tribe" tribe="Platynini">Platynini</taxonomicName>
|
||
, in dorSal view not conStricted towardS neck. DiSc convex, throughout with deep coarSe punctureS (
|
||
<figureCitation id="136300CE4C79FFE7FD8DF94AFD51F930" box="[602,674,1732,1757]" captionStart="FIGURES 1 – 4" captionStartId="6.[151,264,1939,1962]" captionTargetBox="[151,1436,195,1915]" captionTargetId="figure@6.[151,1436,195,1915]" captionTargetPageId="6" captionText="FIGURES 1 – 4. Praeanchodemus punctaticeps sp. n., light microscopic images of the holotype. Fig. 1, dorsal aspect. Fig. 2, Ventral aspect. Fig. 3, left lateral aspect. Fig. 4, general View with contours of the amber piece." httpUri="https://zenodo.org/record/885702/files/figure.png" pageId="1" pageNumber="111">FigS 1</figureCitation>
|
||
,
|
||
<figureCitation id="136300CE4C79FFE7FD66F94AFD32F931" box="[689,705,1732,1757]" captionStart="FIGURES 5 – 9" captionStartId="7.[151,264,1723,1746]" captionTargetBox="[151,1436,197,1702]" captionTargetId="figure@7.[151,1436,197,1702]" captionTargetPageId="7" captionText="FIGURES 5 – 9. Praeanchodemus punctaticeps sp. n., Volume rendering of the holotype. Fig. 5, anterior part of body, dorsal aspect; the arrows point to the insertions of the supraorbital setae, pronotal lateral and laterobasal seta, and parascutellar seta. Figs 6 – 7, Ventral aspect of head using different grayscale thresholds; the arrows point to the insertions of the setae at the mentum tooth and at the submentum. Fig. 8, anterior part of body, right dorsolateral aspect. Fig. 9, elytra, View from caudat; the arrows point to the insertions of the third discal seta and the subapical seta. Note in Figs. 8 – 9 that the fossilized exoskeleton is markedly dissociated from the inclusion wall (= the negatiVe imprint of the beetle body)." httpUri="https://zenodo.org/record/885704/files/figure.png" pageId="1" pageNumber="111">5</figureCitation>
|
||
,
|
||
<figureCitation id="136300CE4C79FFE7FD19F94AFD1EF930" box="[718,749,1732,1756]" captionStart="FIGURE 18" captionStartId="10.[151,250,2014,2036]" captionTargetBox="[363,1225,186,2000]" captionTargetId="figure@10.[362,1225,186,2000]" captionTargetPageId="10" captionText="FIGURE 18. Praeanchodemus punctaticeps sp. n., reconstruction of the external shape in dorsal View." httpUri="https://zenodo.org/record/885710/files/figure.png" pageId="1" pageNumber="111">18</figureCitation>
|
||
). MandibleS (
|
||
<figureCitation id="136300CE4C79FFE7FC51F94AFC1FF931" box="[902,1004,1732,1757]" captionStart="FIGURES 5 – 9" captionStartId="7.[151,264,1723,1746]" captionTargetBox="[151,1436,197,1702]" captionTargetId="figure@7.[151,1436,197,1702]" captionTargetPageId="7" captionText="FIGURES 5 – 9. Praeanchodemus punctaticeps sp. n., Volume rendering of the holotype. Fig. 5, anterior part of body, dorsal aspect; the arrows point to the insertions of the supraorbital setae, pronotal lateral and laterobasal seta, and parascutellar seta. Figs 6 – 7, Ventral aspect of head using different grayscale thresholds; the arrows point to the insertions of the setae at the mentum tooth and at the submentum. Fig. 8, anterior part of body, right dorsolateral aspect. Fig. 9, elytra, View from caudat; the arrows point to the insertions of the third discal seta and the subapical seta. Note in Figs. 8 – 9 that the fossilized exoskeleton is markedly dissociated from the inclusion wall (= the negatiVe imprint of the beetle body)." httpUri="https://zenodo.org/record/885704/files/figure.png" pageId="1" pageNumber="111">FigS 6–7</figureCitation>
|
||
) Slender, with Scrobe aSetoSe. Labrum 6-SetoSe, clypeuS with a Single Seta each Side. EyeS moderately large, moderately protruded (
|
||
<figureCitation id="136300CE4C79FFE7FB74F969FB14F8EC" box="[1187,1255,1767,1792]" captionStart="FIGURES 5 – 9" captionStartId="7.[151,264,1723,1746]" captionTargetBox="[151,1436,197,1702]" captionTargetId="figure@7.[151,1436,197,1702]" captionTargetPageId="7" captionText="FIGURES 5 – 9. Praeanchodemus punctaticeps sp. n., Volume rendering of the holotype. Fig. 5, anterior part of body, dorsal aspect; the arrows point to the insertions of the supraorbital setae, pronotal lateral and laterobasal seta, and parascutellar seta. Figs 6 – 7, Ventral aspect of head using different grayscale thresholds; the arrows point to the insertions of the setae at the mentum tooth and at the submentum. Fig. 8, anterior part of body, right dorsolateral aspect. Fig. 9, elytra, View from caudat; the arrows point to the insertions of the third discal seta and the subapical seta. Note in Figs. 8 – 9 that the fossilized exoskeleton is markedly dissociated from the inclusion wall (= the negatiVe imprint of the beetle body)." httpUri="https://zenodo.org/record/885704/files/figure.png" pageId="1" pageNumber="111">Fig. 5</figureCitation>
|
||
); templeS rather long (
|
||
<figureCitation id="136300CE4C79FFE7FF0DF882FEDDF8C9" box="[218,302,1804,1829]" captionStart="FIGURES 15 – 17" captionStartId="9.[151,264,1920,1943]" captionTargetBox="[265,1321,207,1895]" captionTargetId="figure@9.[265,1321,195,1901]" captionTargetPageId="9" captionText="FIGURES 15 – 17. Praeanchodemus punctaticeps sp. n., Volume rendering of the holotype. Figs 15, anterior part of body, right lateral aspect. Fig. 16, posterior part of body, right lateral aspect; the arrows point to the insertions of the setae of the elytral umbilical series. Fig. 17, seVenth abdominal segment and caudal parts of the female genital using different grayscale thresholds (17 a – b), and reconstructed shape of the gonocoxites (17 c). AbbreViations: acxt: apical gonocoxite; bcxt: basal gonocoxite." httpUri="https://zenodo.org/record/885708/files/figure.png" pageId="1" pageNumber="111">Fig. 15</figureCitation>
|
||
), 0.3 timeS of eye length, weakly wrinkled to neck; Supraorbital furrow indiStinct, two Supraorbital Setae preSent (
|
||
<figureCitation id="136300CE4C79FFE7FEE1F8A1FE88F8A4" box="[310,379,1839,1864]" captionStart="FIGURES 5 – 9" captionStartId="7.[151,264,1723,1746]" captionTargetBox="[151,1436,197,1702]" captionTargetId="figure@7.[151,1436,197,1702]" captionTargetPageId="7" captionText="FIGURES 5 – 9. Praeanchodemus punctaticeps sp. n., Volume rendering of the holotype. Fig. 5, anterior part of body, dorsal aspect; the arrows point to the insertions of the supraorbital setae, pronotal lateral and laterobasal seta, and parascutellar seta. Figs 6 – 7, Ventral aspect of head using different grayscale thresholds; the arrows point to the insertions of the setae at the mentum tooth and at the submentum. Fig. 8, anterior part of body, right dorsolateral aspect. Fig. 9, elytra, View from caudat; the arrows point to the insertions of the third discal seta and the subapical seta. Note in Figs. 8 – 9 that the fossilized exoskeleton is markedly dissociated from the inclusion wall (= the negatiVe imprint of the beetle body)." httpUri="https://zenodo.org/record/885704/files/figure.png" pageId="1" pageNumber="111">Fig. 5</figureCitation>
|
||
). Apical tooth of mentum Simple; Setae near mentum tooth Small, inSerted very cloSe together, immediately baSad of mentum tooth, Submentum with two pairS of lateral Setae, outer pair diStinctly Smaller (
|
||
<figureCitation id="136300CE4C79FFE7FABDF8DAFF37F87C" captionStart="FIGURES 5 – 9" captionStartId="7.[151,264,1723,1746]" captionTargetBox="[151,1436,197,1702]" captionTargetId="figure@7.[151,1436,197,1702]" captionTargetPageId="7" captionText="FIGURES 5 – 9. Praeanchodemus punctaticeps sp. n., Volume rendering of the holotype. Fig. 5, anterior part of body, dorsal aspect; the arrows point to the insertions of the supraorbital setae, pronotal lateral and laterobasal seta, and parascutellar seta. Figs 6 – 7, Ventral aspect of head using different grayscale thresholds; the arrows point to the insertions of the setae at the mentum tooth and at the submentum. Fig. 8, anterior part of body, right dorsolateral aspect. Fig. 9, elytra, View from caudat; the arrows point to the insertions of the third discal seta and the subapical seta. Note in Figs. 8 – 9 that the fossilized exoskeleton is markedly dissociated from the inclusion wall (= the negatiVe imprint of the beetle body)." httpUri="https://zenodo.org/record/885704/files/figure.png" pageId="1" pageNumber="111">FigS 6–7</figureCitation>
|
||
). StipeS with a large Seta externally near baSe; penultimate article of maxillary palpuS medially biSetoSe; ligular Sclerite with two Setae near apical margin. Antennae Slender, pubeScent from fourth antennomere onwardS, Scape not markedly enlarged, Same length aS third antennomere, with one Seta on dorSal Side; pedicel 0.6 timeS aS long aS Scape, with a primary apical Seta on ventral Side and a finer apical Seta each on dorSal and ventral SideS; third antennomere without Setae in addition to normal ring of apical Setae.
|
||
</paragraph>
|
||
<paragraph id="8BE71C4B4C7AFFE4FF10FF51FAADFD8C" blockId="2.[151,1437,151,2015]" pageId="2" pageNumber="112">
|
||
Prothorax: In dorSal view Subcordate-Subquadrate, moderately tranSverSe, broadeSt in middle, with baSal margin Slightly broader than apical margin; SideS evenly rounded in anterior 7/8 and Slightly concave near laterobaSal angleS (
|
||
<figureCitation id="136300CE4C7AFFE4FEA1FEA9FE4DFEAC" box="[374,446,295,320]" captionStart="FIGURES 5 – 9" captionStartId="7.[151,264,1723,1746]" captionTargetBox="[151,1436,197,1702]" captionTargetId="figure@7.[151,1436,197,1702]" captionTargetPageId="7" captionText="FIGURES 5 – 9. Praeanchodemus punctaticeps sp. n., Volume rendering of the holotype. Fig. 5, anterior part of body, dorsal aspect; the arrows point to the insertions of the supraorbital setae, pronotal lateral and laterobasal seta, and parascutellar seta. Figs 6 – 7, Ventral aspect of head using different grayscale thresholds; the arrows point to the insertions of the setae at the mentum tooth and at the submentum. Fig. 8, anterior part of body, right dorsolateral aspect. Fig. 9, elytra, View from caudat; the arrows point to the insertions of the third discal seta and the subapical seta. Note in Figs. 8 – 9 that the fossilized exoskeleton is markedly dissociated from the inclusion wall (= the negatiVe imprint of the beetle body)." httpUri="https://zenodo.org/record/885704/files/figure.png" pageId="2" pageNumber="112">Fig. 5</figureCitation>
|
||
). Anterior margin moderately concave with front angleS diStinctly protruded, Shortly rounded; baSal bead fine, interrupted in middle. PoSterior margin Straight in middle, with external Sixth markedly bent anteriad; laterobaSal angleS markedly Shifted anteriad, obtuSe, not protruded laterally; baSal bead broadly interrupted in middle. DiSc convex, Slightly but diStinctly rugoSe; baSe coarSely rugoSe, without punctureS. Median longitudinal impreSSion moderately deep in middle, diSappearing near apex and baSe; both anterior and poSterior tranSverSe impreSSionS indiStinct. Lateral gutter narrow throughout, very Slightly widened towardS baSe; laterobaSal depreSSionS rather Shallow. Both lateral and laterobaSal Setae preSent, with laterobaSal Setae located beSide lateral gutter at anterior 1/3 of pronotal length, and laterobaSal Setae located on margin (
|
||
<figureCitation id="136300CE4C7AFFE4FBAAFDAAFB3BFDD1" box="[1149,1224,548,573]" captionStart="FIGURES 5 – 9" captionStartId="7.[151,264,1723,1746]" captionTargetBox="[151,1436,197,1702]" captionTargetId="figure@7.[151,1436,197,1702]" captionTargetPageId="7" captionText="FIGURES 5 – 9. Praeanchodemus punctaticeps sp. n., Volume rendering of the holotype. Fig. 5, anterior part of body, dorsal aspect; the arrows point to the insertions of the supraorbital setae, pronotal lateral and laterobasal seta, and parascutellar seta. Figs 6 – 7, Ventral aspect of head using different grayscale thresholds; the arrows point to the insertions of the setae at the mentum tooth and at the submentum. Fig. 8, anterior part of body, right dorsolateral aspect. Fig. 9, elytra, View from caudat; the arrows point to the insertions of the third discal seta and the subapical seta. Note in Figs. 8 – 9 that the fossilized exoskeleton is markedly dissociated from the inclusion wall (= the negatiVe imprint of the beetle body)." httpUri="https://zenodo.org/record/885704/files/figure.png" pageId="2" pageNumber="112">Fig. 5</figureCitation>
|
||
). ProSternum and proepiSternum Smooth, proSternal proceSS not marginated (
|
||
<figureCitation id="136300CE4C7AFFE4FCFEFDC9FC8EFD8C" box="[809,893,583,608]" captionStart="FIGURES 10 – 14" captionStartId="8.[151,264,1842,1865]" captionTargetBox="[224,1362,192,1825]" captionTargetId="figure@8.[224,1362,192,1825]" captionTargetPageId="8" captionText="FIGURES 10 – 14. Praeanchodemus punctaticeps sp. n., Volume rendering of the holotype. Figs 10 – 11, elytra using different grayscale thresholds; the arrows point to the insertions of the discal setae (note that the fossilized left elytron is markedly dissociated from the inclusion wall). Fig. 12, left lateral aspect of pro-, meso-, and metathorax; the arrow points to the insertion of the seta at the mesocoxal ridge. Fig. 13, coxa, trochanter, and femoral base of hind legs; the arrows point to the insertion points of setae (see text for details). Fig. 14, abdominal sternites V-VII; the arrows point to the insertions of the two pairs of setae near apical margin of the seVenth sternite. AbbreViations: mep: metepisternum; mscx: mesocoxa; mtcx: metacoxa; pcx: procoxa." httpUri="https://zenodo.org/record/885706/files/figure.png" pageId="2" pageNumber="112">Fig. 12</figureCitation>
|
||
); anterior coxal cavitieS cloSed poSteriorly.
|
||
</paragraph>
|
||
<paragraph id="8BE71C4B4C7AFFE4FF10FDE2FE2DFC03" blockId="2.[151,1437,151,2015]" pageId="2" pageNumber="112">
|
||
Pterothorax: Elytra markedly convex on diSc (
|
||
<figureCitation id="136300CE4C7AFFE4FD04FDE2FCD5FD69" box="[723,806,620,645]" captionStart="FIGURES 15 – 17" captionStartId="9.[151,264,1920,1943]" captionTargetBox="[265,1321,207,1895]" captionTargetId="figure@9.[265,1321,195,1901]" captionTargetPageId="9" captionText="FIGURES 15 – 17. Praeanchodemus punctaticeps sp. n., Volume rendering of the holotype. Figs 15, anterior part of body, right lateral aspect. Fig. 16, posterior part of body, right lateral aspect; the arrows point to the insertions of the setae of the elytral umbilical series. Fig. 17, seVenth abdominal segment and caudal parts of the female genital using different grayscale thresholds (17 a – b), and reconstructed shape of the gonocoxites (17 c). AbbreViations: acxt: apical gonocoxite; bcxt: basal gonocoxite." httpUri="https://zenodo.org/record/885708/files/figure.png" pageId="2" pageNumber="112">Fig. 16</figureCitation>
|
||
), flattened towardS baSe, elongate-ovate in dorSal view, with ShoulderS moderately broad, humeral angleS and apex Shortly rounded (
|
||
<figureCitation id="136300CE4C7AFFE4FBD5FD01FB75FD44" box="[1026,1158,655,680]" captionStart="FIGURES 10 – 14" captionStartId="8.[151,264,1842,1865]" captionTargetBox="[224,1362,192,1825]" captionTargetId="figure@8.[224,1362,192,1825]" captionTargetPageId="8" captionText="FIGURES 10 – 14. Praeanchodemus punctaticeps sp. n., Volume rendering of the holotype. Figs 10 – 11, elytra using different grayscale thresholds; the arrows point to the insertions of the discal setae (note that the fossilized left elytron is markedly dissociated from the inclusion wall). Fig. 12, left lateral aspect of pro-, meso-, and metathorax; the arrow points to the insertion of the seta at the mesocoxal ridge. Fig. 13, coxa, trochanter, and femoral base of hind legs; the arrows point to the insertion points of setae (see text for details). Fig. 14, abdominal sternites V-VII; the arrows point to the insertions of the two pairs of setae near apical margin of the seVenth sternite. AbbreViations: mep: metepisternum; mscx: mesocoxa; mtcx: metacoxa; pcx: procoxa." httpUri="https://zenodo.org/record/885706/files/figure.png" pageId="2" pageNumber="112">FigS 10–11</figureCitation>
|
||
). BaSal margin concave, protruded towardS Scutellum and humeruS aS well. All Striae deeply impreSSed, impunctate (
|
||
<figureCitation id="136300CE4C7AFFE4FB74FD3AFB0BFD20" box="[1187,1272,692,717]" captionStart="FIGURES 10 – 14" captionStartId="8.[151,264,1842,1865]" captionTargetBox="[224,1362,192,1825]" captionTargetId="figure@8.[224,1362,192,1825]" captionTargetPageId="8" captionText="FIGURES 10 – 14. Praeanchodemus punctaticeps sp. n., Volume rendering of the holotype. Figs 10 – 11, elytra using different grayscale thresholds; the arrows point to the insertions of the discal setae (note that the fossilized left elytron is markedly dissociated from the inclusion wall). Fig. 12, left lateral aspect of pro-, meso-, and metathorax; the arrow points to the insertion of the seta at the mesocoxal ridge. Fig. 13, coxa, trochanter, and femoral base of hind legs; the arrows point to the insertion points of setae (see text for details). Fig. 14, abdominal sternites V-VII; the arrows point to the insertions of the two pairs of setae near apical margin of the seVenth sternite. AbbreViations: mep: metepisternum; mscx: mesocoxa; mtcx: metacoxa; pcx: procoxa." httpUri="https://zenodo.org/record/885706/files/figure.png" pageId="2" pageNumber="112">Fig. 10</figureCitation>
|
||
); paraScutellar Stria moderately long, not connected to firSt Stria; firSt Stria complete to baSe; intervalS moderately convex; fifth interval not impreSSed at apex. ParaScutellar Seta at baSe of firSt Striae, three diScal Seta on third interval, and Subapical Seta near end of Seventh Stria preSent (
|
||
<figureCitation id="136300CE4C7AFFE4FD62FC91FD0CFCD4" box="[693,767,799,824]" captionStart="FIGURES 5 – 9" captionStartId="7.[151,264,1723,1746]" captionTargetBox="[151,1436,197,1702]" captionTargetId="figure@7.[151,1436,197,1702]" captionTargetPageId="7" captionText="FIGURES 5 – 9. Praeanchodemus punctaticeps sp. n., Volume rendering of the holotype. Fig. 5, anterior part of body, dorsal aspect; the arrows point to the insertions of the supraorbital setae, pronotal lateral and laterobasal seta, and parascutellar seta. Figs 6 – 7, Ventral aspect of head using different grayscale thresholds; the arrows point to the insertions of the setae at the mentum tooth and at the submentum. Fig. 8, anterior part of body, right dorsolateral aspect. Fig. 9, elytra, View from caudat; the arrows point to the insertions of the third discal seta and the subapical seta. Note in Figs. 8 – 9 that the fossilized exoskeleton is markedly dissociated from the inclusion wall (= the negatiVe imprint of the beetle body)." httpUri="https://zenodo.org/record/885704/files/figure.png" pageId="2" pageNumber="112">FigS 9</figureCitation>
|
||
,
|
||
<figureCitation id="136300CE4C7AFFE4FCDCFCAEFCDBFCD4" box="[779,808,800,824]" captionStart="FIGURES 10 – 14" captionStartId="8.[151,264,1842,1865]" captionTargetBox="[224,1362,192,1825]" captionTargetId="figure@8.[224,1362,192,1825]" captionTargetPageId="8" captionText="FIGURES 10 – 14. Praeanchodemus punctaticeps sp. n., Volume rendering of the holotype. Figs 10 – 11, elytra using different grayscale thresholds; the arrows point to the insertions of the discal setae (note that the fossilized left elytron is markedly dissociated from the inclusion wall). Fig. 12, left lateral aspect of pro-, meso-, and metathorax; the arrow points to the insertion of the seta at the mesocoxal ridge. Fig. 13, coxa, trochanter, and femoral base of hind legs; the arrows point to the insertion points of setae (see text for details). Fig. 14, abdominal sternites V-VII; the arrows point to the insertions of the two pairs of setae near apical margin of the seVenth sternite. AbbreViations: mep: metepisternum; mscx: mesocoxa; mtcx: metacoxa; pcx: procoxa." httpUri="https://zenodo.org/record/885706/files/figure.png" pageId="2" pageNumber="112">11</figureCitation>
|
||
), with anterior diScal Seta located between anterior fifth to Sixth of elytral length, adjoined to third Stria, middle Seta located Slightly behind elytral middle, adjoined to third Stria, and poSterior Seta located at poSterior elytral Seventh, adjoined to Second Stria; diScal Setae not Set in foSSae. Umbilicate SerieS conSiStS of 15 Setae adjoining eighth Stria, with diStance between 6th and 7th Seta Slightly larger (
|
||
<figureCitation id="136300CE4C7AFFE4FF48FC3CFF00FC27" box="[159,243,946,971]" captionStart="FIGURES 15 – 17" captionStartId="9.[151,264,1920,1943]" captionTargetBox="[265,1321,207,1895]" captionTargetId="figure@9.[265,1321,195,1901]" captionTargetPageId="9" captionText="FIGURES 15 – 17. Praeanchodemus punctaticeps sp. n., Volume rendering of the holotype. Figs 15, anterior part of body, right lateral aspect. Fig. 16, posterior part of body, right lateral aspect; the arrows point to the insertions of the setae of the elytral umbilical series. Fig. 17, seVenth abdominal segment and caudal parts of the female genital using different grayscale thresholds (17 a – b), and reconstructed shape of the gonocoxites (17 c). AbbreViations: acxt: apical gonocoxite; bcxt: basal gonocoxite." httpUri="https://zenodo.org/record/885708/files/figure.png" pageId="2" pageNumber="112">Fig. 16</figureCitation>
|
||
). MetepiSternum Smooth, Short, with outer margin Slightly longer than anterior margin (
|
||
<figureCitation id="136300CE4C7AFFE4FB1AFC3CFAD7FC27" box="[1229,1316,946,971]" captionStart="FIGURES 10 – 14" captionStartId="8.[151,264,1842,1865]" captionTargetBox="[224,1362,192,1825]" captionTargetId="figure@8.[224,1362,192,1825]" captionTargetPageId="8" captionText="FIGURES 10 – 14. Praeanchodemus punctaticeps sp. n., Volume rendering of the holotype. Figs 10 – 11, elytra using different grayscale thresholds; the arrows point to the insertions of the discal setae (note that the fossilized left elytron is markedly dissociated from the inclusion wall). Fig. 12, left lateral aspect of pro-, meso-, and metathorax; the arrow points to the insertion of the seta at the mesocoxal ridge. Fig. 13, coxa, trochanter, and femoral base of hind legs; the arrows point to the insertion points of setae (see text for details). Fig. 14, abdominal sternites V-VII; the arrows point to the insertions of the two pairs of setae near apical margin of the seVenth sternite. AbbreViations: mep: metepisternum; mscx: mesocoxa; mtcx: metacoxa; pcx: procoxa." httpUri="https://zenodo.org/record/885706/files/figure.png" pageId="2" pageNumber="112">FigS 12</figureCitation>
|
||
,
|
||
<figureCitation id="136300CE4C7AFFE4FAE7FC3CFABCFC27" box="[1328,1359,946,971]" captionStart="FIGURES 15 – 17" captionStartId="9.[151,264,1920,1943]" captionTargetBox="[265,1321,207,1895]" captionTargetId="figure@9.[265,1321,195,1901]" captionTargetPageId="9" captionText="FIGURES 15 – 17. Praeanchodemus punctaticeps sp. n., Volume rendering of the holotype. Figs 15, anterior part of body, right lateral aspect. Fig. 16, posterior part of body, right lateral aspect; the arrows point to the insertions of the setae of the elytral umbilical series. Fig. 17, seVenth abdominal segment and caudal parts of the female genital using different grayscale thresholds (17 a – b), and reconstructed shape of the gonocoxites (17 c). AbbreViations: acxt: apical gonocoxite; bcxt: basal gonocoxite." httpUri="https://zenodo.org/record/885708/files/figure.png" pageId="2" pageNumber="112">16</figureCitation>
|
||
). Hind wingS reduced to Short StubS.
|
||
</paragraph>
|
||
<paragraph id="8BE71C4B4C7AFFE4FF10FC75FC81FADF" blockId="2.[151,1437,151,2015]" pageId="2" pageNumber="112">
|
||
LegS: Rather Short. All trochanterS with a Single Seta on external Surface. MeSocoxa with a Single ridge Seta (
|
||
<figureCitation id="136300CE4C7AFFE4FF48FB90FF0BFBDB" box="[159,248,1054,1079]" captionStart="FIGURES 10 – 14" captionStartId="8.[151,264,1842,1865]" captionTargetBox="[224,1362,192,1825]" captionTargetId="figure@8.[224,1362,192,1825]" captionTargetPageId="8" captionText="FIGURES 10 – 14. Praeanchodemus punctaticeps sp. n., Volume rendering of the holotype. Figs 10 – 11, elytra using different grayscale thresholds; the arrows point to the insertions of the discal setae (note that the fossilized left elytron is markedly dissociated from the inclusion wall). Fig. 12, left lateral aspect of pro-, meso-, and metathorax; the arrow points to the insertion of the seta at the mesocoxal ridge. Fig. 13, coxa, trochanter, and femoral base of hind legs; the arrows point to the insertion points of setae (see text for details). Fig. 14, abdominal sternites V-VII; the arrows point to the insertions of the two pairs of setae near apical margin of the seVenth sternite. AbbreViations: mep: metepisternum; mscx: mesocoxa; mtcx: metacoxa; pcx: procoxa." httpUri="https://zenodo.org/record/885706/files/figure.png" pageId="2" pageNumber="112">Fig. 12</figureCitation>
|
||
); metacoxa biSetoSe, without Seta near internal margin (
|
||
<figureCitation id="136300CE4C7AFFE4FC40FB90FC1CFBDB" box="[919,1007,1054,1079]" captionStart="FIGURES 10 – 14" captionStartId="8.[151,264,1842,1865]" captionTargetBox="[224,1362,192,1825]" captionTargetId="figure@8.[224,1362,192,1825]" captionTargetPageId="8" captionText="FIGURES 10 – 14. Praeanchodemus punctaticeps sp. n., Volume rendering of the holotype. Figs 10 – 11, elytra using different grayscale thresholds; the arrows point to the insertions of the discal setae (note that the fossilized left elytron is markedly dissociated from the inclusion wall). Fig. 12, left lateral aspect of pro-, meso-, and metathorax; the arrow points to the insertion of the seta at the mesocoxal ridge. Fig. 13, coxa, trochanter, and femoral base of hind legs; the arrows point to the insertion points of setae (see text for details). Fig. 14, abdominal sternites V-VII; the arrows point to the insertions of the two pairs of setae near apical margin of the seVenth sternite. AbbreViations: mep: metepisternum; mscx: mesocoxa; mtcx: metacoxa; pcx: procoxa." httpUri="https://zenodo.org/record/885706/files/figure.png" pageId="2" pageNumber="112">Fig. 13</figureCitation>
|
||
); anterior metacoxal SulcuS Straight. Profemur with a Single Seta on ventral Surface and with two rowS of Six Short Setae on anterior Surface; meSofemur with two Setae on ventral Surface and with two rowS of nine Short Setae on anterior Surface; metafemur with two Setae on ventral Surface (
|
||
<figureCitation id="136300CE4C7AFFE4FE67FB04FDF0FB4F" box="[432,515,1162,1187]" captionStart="FIGURES 10 – 14" captionStartId="8.[151,264,1842,1865]" captionTargetBox="[224,1362,192,1825]" captionTargetId="figure@8.[224,1362,192,1825]" captionTargetPageId="8" captionText="FIGURES 10 – 14. Praeanchodemus punctaticeps sp. n., Volume rendering of the holotype. Figs 10 – 11, elytra using different grayscale thresholds; the arrows point to the insertions of the discal setae (note that the fossilized left elytron is markedly dissociated from the inclusion wall). Fig. 12, left lateral aspect of pro-, meso-, and metathorax; the arrow points to the insertion of the seta at the mesocoxal ridge. Fig. 13, coxa, trochanter, and femoral base of hind legs; the arrows point to the insertion points of setae (see text for details). Fig. 14, abdominal sternites V-VII; the arrows point to the insertions of the two pairs of setae near apical margin of the seVenth sternite. AbbreViations: mep: metepisternum; mscx: mesocoxa; mtcx: metacoxa; pcx: procoxa." httpUri="https://zenodo.org/record/885706/files/figure.png" pageId="2" pageNumber="112">Fig. 13</figureCitation>
|
||
), with dorSal Surface Smooth even near apex. Protibia not modified, with external Surface Smooth. FirSt and Second tarSomereS each with a pair of dorSoapical Setae, latter lacking on diStal tarSomereS; fourth tarSomereS of all legS Simple, with ventro-apical lobeS very Short, with pair of lateroapical Setae preSent and without dorSoapical Setae; ventral SurfaceS of fifth tarSomereS Smooth; clawS Simple. PreSence of lateral longitudinal grooveS on dorSal SurfaceS of tarSomereS could not be confirmed.
|
||
</paragraph>
|
||
<paragraph id="8BE71C4B4C7AFFE4FF10FAB0FDE5FA97" blockId="2.[151,1437,151,2015]" pageId="2" pageNumber="112">
|
||
Abdomen: Abdominal SterniteS Smooth beSide normal Setation; V–VI with one, VII with two (female) pairS of Setae near apical margin (
|
||
<figureCitation id="136300CE4C7AFFE4FE62FAECFDFBFA97" box="[437,520,1378,1403]" captionStart="FIGURES 10 – 14" captionStartId="8.[151,264,1842,1865]" captionTargetBox="[224,1362,192,1825]" captionTargetId="figure@8.[224,1362,192,1825]" captionTargetPageId="8" captionText="FIGURES 10 – 14. Praeanchodemus punctaticeps sp. n., Volume rendering of the holotype. Figs 10 – 11, elytra using different grayscale thresholds; the arrows point to the insertions of the discal setae (note that the fossilized left elytron is markedly dissociated from the inclusion wall). Fig. 12, left lateral aspect of pro-, meso-, and metathorax; the arrow points to the insertion of the seta at the mesocoxal ridge. Fig. 13, coxa, trochanter, and femoral base of hind legs; the arrows point to the insertion points of setae (see text for details). Fig. 14, abdominal sternites V-VII; the arrows point to the insertions of the two pairs of setae near apical margin of the seVenth sternite. AbbreViations: mep: metepisternum; mscx: mesocoxa; mtcx: metacoxa; pcx: procoxa." httpUri="https://zenodo.org/record/885706/files/figure.png" pageId="2" pageNumber="112">Fig. 14</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BE71C4B4C7AFFE4FF10FA08FCF9FA0B" blockId="2.[151,1437,151,2015]" pageId="2" pageNumber="112">
|
||
Female genitalia: Form of gonocoxiteS aS typical of moSt
|
||
<taxonomicName id="4C5867C84C7AFFE4FCE9FA08FC53FA73" authorityName="Bonelli" authorityYear="1810" box="[830,928,1414,1439]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="112" phylum="Arthropoda" rank="tribe" tribe="Platynini">Platynini</taxonomicName>
|
||
(
|
||
<figureCitation id="136300CE4C7AFFE4FC67FA08FBF3FA73" box="[944,1024,1414,1439]" captionStart="FIGURES 15 – 17" captionStartId="9.[151,264,1920,1943]" captionTargetBox="[265,1321,207,1895]" captionTargetId="figure@9.[265,1321,195,1901]" captionTargetPageId="9" captionText="FIGURES 15 – 17. Praeanchodemus punctaticeps sp. n., Volume rendering of the holotype. Figs 15, anterior part of body, right lateral aspect. Fig. 16, posterior part of body, right lateral aspect; the arrows point to the insertions of the setae of the elytral umbilical series. Fig. 17, seVenth abdominal segment and caudal parts of the female genital using different grayscale thresholds (17 a – b), and reconstructed shape of the gonocoxites (17 c). AbbreViations: acxt: apical gonocoxite; bcxt: basal gonocoxite." httpUri="https://zenodo.org/record/885708/files/figure.png" pageId="2" pageNumber="112">Fig. 17</figureCitation>
|
||
); baSal gonocoxiteS moderately Stout, apical gonocoxiteS Slender cone-Shaped, very Slightly bent outwardly, with Subapical SetoSe organ preSent; chaetotaxy and internal genital tract could not be Seen nor viSualized.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3424FC04C7AFFE4FF10FA7CFEAEF99B" pageId="2" pageNumber="112" type="etymology">
|
||
<paragraph id="8BE71C4B4C7AFFE4FF10FA7CFEAEF99B" blockId="2.[151,1437,151,2015]" pageId="2" pageNumber="112">
|
||
<emphasis id="B92CC0594C7AFFE4FF10FA7CFEBDF9E7" bold="true" box="[199,334,1522,1547]" pageId="2" pageNumber="112">Etymology.</emphasis>
|
||
A compound word, baSed on the Latin prefix “prae-“ and the name of the platynine taxon
|
||
<taxonomicName id="4C5867C84C7AFFE4FF40F998FED5F9C3" box="[151,294,1558,1583]" class="Insecta" family="Curculionidae" genus="Anchodemus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="112" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B92CC0594C7AFFE4FF40F998FED5F9C3" box="[151,294,1558,1583]" italics="true" pageId="2" pageNumber="112">Anchodemus</emphasis>
|
||
</taxonomicName>
|
||
MotSchulSky, 1864. The name of the new genuS particularly referS to the externally Similar taxon
|
||
<taxonomicName id="4C5867C84C7AFFE4FF40F9B4FE13F9BF" authority="Habu, 1978" authorityName="Habu" authorityYear="1978" box="[151,480,1594,1619]" class="Insecta" family="Carabidae" genus="Paranchodemus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="112" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B92CC0594C7AFFE4FF40F9B4FEBEF9BF" box="[151,333,1594,1619]" italics="true" pageId="2" pageNumber="112">Paranchodemus</emphasis>
|
||
<bibRefCitation id="EFC961BA4C7AFFE4FE80F9B4FE13F9BF" author="Habu" box="[343,480,1594,1619]" pageId="2" pageNumber="112" refString="Habu, A. (1978) Fauna Japonica. Carabidae: Platynini (Insecta: Coleoptera). Keigaku Publishing, Tokyo, 447 pp., 36 pls." type="book" year="1978">Habu, 1978</bibRefCitation>
|
||
</taxonomicName>
|
||
, which waS deScribed aS SubgenuS of
|
||
<taxonomicName id="4C5867C84C7AFFE4FC48F9B4FBDDF9BF" box="[927,1070,1594,1619]" class="Insecta" family="Curculionidae" genus="Anchodemus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="112" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B92CC0594C7AFFE4FC48F9B4FBDDF9BF" box="[927,1070,1594,1619]" italics="true" pageId="2" pageNumber="112">Anchodemus</emphasis>
|
||
</taxonomicName>
|
||
and elevated to genuS level by
|
||
<bibRefCitation id="EFC961BA4C7AFFE4FF40F9D0FEABF99B" author="Liebherr" box="[151,344,1630,1655]" pageId="2" pageNumber="112" refString="Liebherr, J. K. (1989 a) ReView of the Palearctic genus Paranchodemus Habu (Coleoptera: Carabidae: Platynini). Pan-Pacific Entomologist, 65 (1), 1 - 11." type="journal article" year="1989" yearSuffix="a">Liebherr (1989a)</bibRefCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3424FC04C7AFFE5FF10F90CFD51FDD0" lastPageId="3" lastPageNumber="113" pageId="2" pageNumber="112" type="diagnosis">
|
||
<paragraph id="8BE71C4B4C7AFFE4FF10F90CFAB7F953" blockId="2.[151,1437,151,2015]" pageId="2" pageNumber="112">
|
||
<emphasis id="B92CC0594C7AFFE4FF10F90CFD56F977" bold="true" box="[199,677,1666,1691]" pageId="2" pageNumber="112">Differential diagnosis and relationships.</emphasis>
|
||
A Small repreSentative of
|
||
<taxonomicName id="4C5867C84C7AFFE4FC06F90CFBC5F977" authorityName="Bonelli" authorityYear="1810" box="[977,1078,1666,1691]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="112" phylum="Arthropoda" rank="tribe" tribe="Platynini">Platynini</taxonomicName>
|
||
with rather Short legS, which iS eaSily diStinguiShed from all other genera of that tribe by the following combination of external characterS:
|
||
</paragraph>
|
||
<paragraph id="8BE71C4B4C7AFFE4FF10F945FB5AF90F" blockId="2.[151,1437,151,2015]" box="[199,1193,1738,1763]" pageId="2" pageNumber="112">1) Setae near mentum tooth inSerted cloSe together, immediately baSad of mentum tooth.</paragraph>
|
||
<paragraph id="8BE71C4B4C7AFFE4FF10F961FCC0F8EB" blockId="2.[151,1437,151,2015]" box="[199,819,1774,1799]" pageId="2" pageNumber="112">2) DiSc of head throughout with deep coarSe punctureS.</paragraph>
|
||
<paragraph id="8BE71C4B4C7AFFE4FF10F89DFD51F8C7" blockId="2.[151,1437,151,2015]" box="[199,674,1810,1835]" pageId="2" pageNumber="112">3) Head lacking diStinct neck conStriction.</paragraph>
|
||
<paragraph id="8BE71C4B4C7AFFE4FF10F8B9FA60F8A3" blockId="2.[151,1437,151,2015]" box="[199,1427,1846,1871]" pageId="2" pageNumber="112">4) MicroSculpture patternS markedly contraSting on different body partS, with large and deeply engraved lineS</paragraph>
|
||
<paragraph id="8BE71C4B4C7AFFE4FF34F8D4FAA8F89F" blockId="2.[151,1437,151,2015]" box="[227,1371,1882,1907]" pageId="2" pageNumber="112">forming iSodiametric SculpticellS on head and very indiStinct SculpticellS on pronotal and elytral diScS.</paragraph>
|
||
<paragraph id="8BE71C4B4C7AFFE4FF10F8F0FBD4F87B" blockId="2.[151,1437,151,2015]" box="[199,1063,1918,1944]" pageId="2" pageNumber="112">5) Pronotal lateral gutter narrow throughout, laterobaSal depreSSionS Shallow.</paragraph>
|
||
<paragraph id="8BE71C4B4C7AFFE4FF10F82CFCCFF857" blockId="2.[151,1437,151,2015]" box="[199,828,1954,1979]" pageId="2" pageNumber="112">6) Metacoxa biSetoSe, without Seta near internal margin.</paragraph>
|
||
<paragraph id="8BE71C4B4C7AFFE4FF10F849FBAEF833" blockId="2.[151,1437,151,2015]" box="[199,1117,1990,2015]" pageId="2" pageNumber="112">7) Profemur with a Single Seta, meSofemur with only two Setae on ventral Surface.</paragraph>
|
||
<paragraph id="8BE71C4B4C7BFFE5FF10FF16FD43FF5C" blockId="3.[151,1437,151,2013]" box="[199,688,151,176]" pageId="3" pageNumber="113">8) Metafemur dorSally without apical Setae.</paragraph>
|
||
<paragraph id="8BE71C4B4C7BFFE5FF10FF32FCB9FF39" blockId="3.[151,1437,151,2013]" box="[199,842,188,213]" pageId="3" pageNumber="113">9) All legS with fourth tarSomere dorSoapical Setae lacking.</paragraph>
|
||
<paragraph id="8BE71C4B4C7BFFE5FF10FF6EFCA2FF14" blockId="3.[151,1437,151,2013]" box="[199,849,223,248]" pageId="3" pageNumber="113">10) All legS with fifth tarSomere Smooth on ventral Surface.</paragraph>
|
||
<paragraph id="8BE71C4B4C7BFFE5FF10FE8AFD51FDD0" blockId="3.[151,1437,151,2013]" pageId="3" pageNumber="113">
|
||
The poSition of mentum Setae iS conSidered an important indication for SpecieS-group relationShipS (
|
||
<bibRefCitation id="EFC961BA4C7BFFE5FAEAFE8AFF13FEAC" author="Schmidt" pageId="3" pageNumber="113" refString="Schmidt, J. (2001 a) Archicolpodes n. gen. - eine neue Laufkafergattung der Tribus Platynini aus China (Coleoptera, Carabidae). Entomologische Blatter, Keltern, 96, 211 - 218. [2000]" type="journal article" year="2001" yearSuffix="a">Schmidt 2001a</bibRefCitation>
|
||
). By far, moSt lineageS of platynine ground beetleS have a Single pair of Setae inSerted caudally of the deep, apical inciSionS of the mentum, laterally of the apical tooth, and thuS, the inSertion pointS of the mentum Setae are located rather diStant from each other (See
|
||
<bibRefCitation id="EFC961BA4C7BFFE5FDA6FEE1FCEDFE64" author="Schmidt" box="[625,798,367,392]" pageId="3" pageNumber="113" refString="Schmidt, J. (2001 a) Archicolpodes n. gen. - eine neue Laufkafergattung der Tribus Platynini aus China (Coleoptera, Carabidae). Entomologische Blatter, Keltern, 96, 211 - 218. [2000]" type="journal article" year="2001" yearSuffix="a">Schmidt 2001a</bibRefCitation>
|
||
:
|
||
<figureCitation id="136300CE4C7BFFE5FCF0FEE1FC55FE64" box="[807,934,367,392]" captionStart="FIGURES 10 – 14" captionStartId="8.[151,264,1842,1865]" captionTargetBox="[224,1362,192,1825]" captionTargetId="figure@8.[224,1362,192,1825]" captionTargetPageId="8" captionText="FIGURES 10 – 14. Praeanchodemus punctaticeps sp. n., Volume rendering of the holotype. Figs 10 – 11, elytra using different grayscale thresholds; the arrows point to the insertions of the discal setae (note that the fossilized left elytron is markedly dissociated from the inclusion wall). Fig. 12, left lateral aspect of pro-, meso-, and metathorax; the arrow points to the insertion of the seta at the mesocoxal ridge. Fig. 13, coxa, trochanter, and femoral base of hind legs; the arrows point to the insertion points of setae (see text for details). Fig. 14, abdominal sternites V-VII; the arrows point to the insertions of the two pairs of setae near apical margin of the seVenth sternite. AbbreViations: mep: metepisternum; mscx: mesocoxa; mtcx: metacoxa; pcx: procoxa." httpUri="https://zenodo.org/record/885706/files/figure.png" pageId="3" pageNumber="113">FigS 12, 14</figureCitation>
|
||
,
|
||
<figureCitation id="136300CE4C7BFFE5FC66FEE1FC23FE64" box="[945,976,367,392]" captionStart="FIGURES 15 – 17" captionStartId="9.[151,264,1920,1943]" captionTargetBox="[265,1321,207,1895]" captionTargetId="figure@9.[265,1321,195,1901]" captionTargetPageId="9" captionText="FIGURES 15 – 17. Praeanchodemus punctaticeps sp. n., Volume rendering of the holotype. Figs 15, anterior part of body, right lateral aspect. Fig. 16, posterior part of body, right lateral aspect; the arrows point to the insertions of the setae of the elytral umbilical series. Fig. 17, seVenth abdominal segment and caudal parts of the female genital using different grayscale thresholds (17 a – b), and reconstructed shape of the gonocoxites (17 c). AbbreViations: acxt: apical gonocoxite; bcxt: basal gonocoxite." httpUri="https://zenodo.org/record/885708/files/figure.png" pageId="3" pageNumber="113">16</figureCitation>
|
||
). In contraSt, the character State obServed in the foSSil
|
||
<taxonomicName id="4C5867C84C7BFFE5FEF0FE1AFE1AFE41" authority="Will, 2017" authorityName="Will" authorityYear="2017" box="[295,489,404,429]" class="Insecta" family="Carabidae" genus="Praeanchodemus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="113" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B92CC0594C7BFFE5FEF0FE1AFE1AFE41" box="[295,489,404,429]" italics="true" pageId="3" pageNumber="113">Praeanchodemus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21F7D224C7BFFE5FE25FE1BFDB3FE40" box="[498,576,405,429]" pageId="3" pageNumber="113" rank="genus">gen. n.</taxonomicNameLabel>
|
||
(character State 1) iS preSent in only Six terminal
|
||
<taxonomicName id="4C5867C84C7BFFE5FBA9FE1AFB10FE41" authorityName="Bonelli" authorityYear="1810" box="[1150,1251,404,429]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="113" phylum="Arthropoda" rank="tribe" tribe="Platynini">Platynini</taxonomicName>
|
||
lineageS, and iS conSidered apomorphic. However, given the current State of knowledge, there iS no evidence for cloSe relationShipS of theSe lineageS (
|
||
<bibRefCitation id="EFC961BA4C7BFFE5FEB1FE52FDFAFE18" author="Schmidt" box="[358,521,476,501]" pageId="3" pageNumber="113" refString="Schmidt, J. (2000) Ein Vorschlag zur Losung nomenklaturischer Probleme der Gattung Platynus Bonelli, 1810 (Coleoptera, Carabidae). Entomologische Blatter, Keltern, 96, 9 - 23." type="journal article" year="2000">Schmidt 2000</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFC961BA4C7BFFE5FDC1FE52FDACFE18" author="Schmidt" box="[534,607,476,500]" pageId="3" pageNumber="113" refString="Schmidt, J. (2001 a) Archicolpodes n. gen. - eine neue Laufkafergattung der Tribus Platynini aus China (Coleoptera, Carabidae). Entomologische Blatter, Keltern, 96, 211 - 218. [2000]" type="journal article" year="2001" yearSuffix="a">2001a</bibRefCitation>
|
||
). It iS thuS likely that the adjacent poSition of the mentum Setae evolved within the tribe two or more timeS independently. In the following, the Six relevant lineageS are diScuSSed in more detail and compared with the new foSSil taxon.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3424FC04C7BFFE2FF10FDC9FAEBF9C1" lastPageId="4" lastPageNumber="114" pageId="3" pageNumber="113" type="reference_group">
|
||
<paragraph id="8BE71C4B4C7BFFE5FF10FDC9FD95FBD9" blockId="3.[151,1437,151,2013]" pageId="3" pageNumber="113">
|
||
<taxonomicName id="4C5867C84C7BFFE5FF10FDC9FDCFFD8C" authority="MotSchulSky, 1850" authorityName="MotSchulSky" authorityYear="1850" box="[199,572,583,608]" class="Insecta" family="Carabidae" genus="Limodromus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="122" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B92CC0594C7BFFE5FF10FDC9FEA0FD8C" box="[199,339,583,608]" italics="true" pageId="3" pageNumber="113">Limodromus</emphasis>
|
||
MotSchulSky, 1850
|
||
</taxonomicName>
|
||
: Occurrence of thiS genuS in the foSSil fauna of the Eocene Baltic amber foreStS waS confirmed recently (
|
||
<bibRefCitation id="EFC961BA4C7BFFE5FDD0FDE2FD5EFD69" author="Schmidt" box="[519,685,620,645]" pageId="3" pageNumber="113" refString="Schmidt, J. (2015) On the Eocene age of Limodromus Motschulsky, 1850, with description of L. hoffeinsorum sp. n. from Baltic Amber (Coleoptera, Carabidae, Platynini). Zootaxa, 3974 (4), 573 - 581. https: // doi. org / 10.11646 / zootaxa. 3974.4.8" type="journal article" year="2015">Schmidt 2015</bibRefCitation>
|
||
). Monophyly of
|
||
<taxonomicName id="4C5867C84C7BFFE5FCB9FDE2FC08FD69" authorityName="MotSchulSky" authorityYear="1850" box="[878,1019,620,645]" class="Insecta" family="Carabidae" genus="Limodromus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="113" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B92CC0594C7BFFE5FCB9FDE2FC08FD69" box="[878,1019,620,645]" italics="true" pageId="3" pageNumber="113">Limodromus</emphasis>
|
||
</taxonomicName>
|
||
iS well Supported by highly derived morphology of the aedeagal median lobe that iS unique within
|
||
<taxonomicName id="4C5867C84C7BFFE5FC98FD01FC47FD44" ID-CoL="8KTWY" authorityName="Bonelli" authorityYear="1810" box="[847,948,655,680]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="113" phylum="Arthropoda" rank="tribe" tribe="Platynini">Platynini</taxonomicName>
|
||
(
|
||
<bibRefCitation id="EFC961BA4C7BFFE5FC15FD01FB90FD44" author="Schmidt" box="[962,1123,655,680]" pageId="3" pageNumber="113" refString="Schmidt, J. (2000) Ein Vorschlag zur Losung nomenklaturischer Probleme der Gattung Platynus Bonelli, 1810 (Coleoptera, Carabidae). Entomologische Blatter, Keltern, 96, 9 - 23." type="journal article" year="2000">Schmidt 2000</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFC961BA4C7BFFE5FBBAFD01FB59FD44" author="Schmidt" box="[1133,1194,655,680]" pageId="3" pageNumber="113" refString="Schmidt, J. (2005) Limodromus klausnitzeri sp. n., ein phylogeographisch interessanter Fund aus Nord-Vietnam, sowie Bestimmungsschlussel fur die Arten der Gattung Limodromus Motschulsky, 1850 (Coleoptera, Carabidae, Platynini). Entomologische Blatter, Keltern, 100 [2004], 181 - 190." type="journal article" year="2005">2005</bibRefCitation>
|
||
). BecauSe deScription of
|
||
<taxonomicName id="4C5867C84C7BFFE5FF60FD3AFE89FD21" ID-CoL="8LTCY" authority="Will, 2017" authorityName="Will" authorityYear="2017" box="[183,378,692,717]" class="Insecta" family="Carabidae" genus="Praeanchodemus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="113" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B92CC0594C7BFFE5FF60FD3AFE89FD21" box="[183,378,692,717]" italics="true" pageId="3" pageNumber="113">Praeanchodemus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21F7D224C7BFFE5FE55FD3BFE3CFD20" box="[386,463,693,717]" pageId="3" pageNumber="113" rank="genus">gen. n.</taxonomicNameLabel>
|
||
iS baSed on a female Specimen the relevant character StateS have not been examined. However, in external characterS, repreSentativeS of
|
||
<taxonomicName id="4C5867C84C7BFFE5FD3DFD59FC84FD1C" authorityName="MotSchulSky" authorityYear="1850" box="[746,887,727,752]" class="Insecta" family="Carabidae" genus="Limodromus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="113" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B92CC0594C7BFFE5FD3DFD59FC84FD1C" box="[746,887,727,752]" italics="true" pageId="3" pageNumber="113">Limodromus</emphasis>
|
||
</taxonomicName>
|
||
are eaSily diStinguiShed from
|
||
<taxonomicName id="4C5867C84C7BFFE5FB0DFD59FA6FFD1C" ID-CoL="8LTCY" authority="Will, 2017" authorityName="Will" authorityYear="2017" box="[1242,1436,727,752]" class="Insecta" family="Carabidae" genus="Praeanchodemus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="113" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B92CC0594C7BFFE5FB0DFD59FA6FFD1C" box="[1242,1436,727,752]" italics="true" pageId="3" pageNumber="113">Praeanchodemus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21F7D224C7BFFE5FF40FD73FF11FCF8" box="[151,226,765,789]" pageId="3" pageNumber="113" rank="genus">gen. n.</taxonomicNameLabel>
|
||
by the head, which iS diStinctly conStricted towardS neck, and by the pronotum, which iS large, cordate, and haS lateral gutter and laterobaSal grooveS very broadly developed. WhereaS in
|
||
<taxonomicName id="4C5867C84C7BFFE5FBF1FC91FB1AFCD4" ID-CoL="8LTCY" authority="Will, 2017" authorityName="Will" authorityYear="2017" box="[1062,1257,799,824]" class="Insecta" family="Carabidae" genus="Praeanchodemus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="113" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B92CC0594C7BFFE5FBF1FC91FB1AFCD4" box="[1062,1257,799,824]" italics="true" pageId="3" pageNumber="113">Praeanchodemus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21F7D224C7BFFE5FB24FCAEFAB1FCDB" box="[1267,1346,800,824]" pageId="3" pageNumber="113" rank="genus">gen. n.</taxonomicNameLabel>
|
||
a baSal conStriction of the head capSule iS lacking, the pronotal lateral gutter iS markedly Small and the laterobaSal depreSSionS are Shallow (See character StateS 3 and 5). In addition, patternS of micro- and macroSculpture of the dorSal Surface of the body are very differently developed in
|
||
<taxonomicName id="4C5867C84C7BFFE5FC96FC02FC3DFC49" authorityName="MotSchulSky" authorityYear="1850" box="[833,974,908,933]" class="Insecta" family="Carabidae" genus="Limodromus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="113" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B92CC0594C7BFFE5FC96FC02FC3DFC49" box="[833,974,908,933]" italics="true" pageId="3" pageNumber="113">Limodromus</emphasis>
|
||
</taxonomicName>
|
||
with reSpect to
|
||
<taxonomicName id="4C5867C84C7BFFE5FB52FC02FABBFC49" ID-CoL="8LTCY" authority="Will, 2017" authorityName="Will" authorityYear="2017" box="[1157,1352,908,933]" class="Insecta" family="Carabidae" genus="Praeanchodemus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="113" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B92CC0594C7BFFE5FB52FC02FABBFC49" box="[1157,1352,908,933]" italics="true" pageId="3" pageNumber="113">Praeanchodemus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21F7D224C7BFFE5FA98FC03FA6FFC48" box="[1359,1436,909,933]" pageId="3" pageNumber="113" rank="genus">gen. n.</taxonomicNameLabel>
|
||
(See character StateS 2 and 4) and, in
|
||
<taxonomicName id="4C5867C84C7BFFE5FDEFFC21FD37FC24" authorityName="MotSchulSky" authorityYear="1850" box="[568,708,943,968]" class="Insecta" family="Carabidae" genus="Limodromus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="113" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B92CC0594C7BFFE5FDEFFC21FD37FC24" box="[568,708,943,968]" italics="true" pageId="3" pageNumber="113">Limodromus</emphasis>
|
||
</taxonomicName>
|
||
SpecieS the metafemurS bear dorSoapical Setae that repreSent the pleSiomorphic character State, which are completely reduced in
|
||
<taxonomicName id="4C5867C84C7BFFE5FCA1FC5AFBCAFC01" ID-CoL="8LTCY" authority="Will, 2017" authorityName="Will" authorityYear="2017" box="[886,1081,980,1005]" class="Insecta" family="Carabidae" genus="Praeanchodemus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="113" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B92CC0594C7BFFE5FCA1FC5AFBCAFC01" box="[886,1081,980,1005]" italics="true" pageId="3" pageNumber="113">Praeanchodemus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21F7D224C7BFFE5FB95FC5BFB63FC00" box="[1090,1168,981,1005]" pageId="3" pageNumber="113" rank="genus">gen. n.</taxonomicNameLabel>
|
||
(See character State 8). ConSequently, due to the lack of apparent SynapomorphieS a cloSe relationShip of
|
||
<taxonomicName id="4C5867C84C7BFFE5FB74FC79FAC3FBFC" authorityName="MotSchulSky" authorityYear="1850" box="[1187,1328,1015,1040]" class="Insecta" family="Carabidae" genus="Limodromus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="113" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B92CC0594C7BFFE5FB74FC79FAC3FBFC" box="[1187,1328,1015,1040]" italics="true" pageId="3" pageNumber="113">Limodromus</emphasis>
|
||
</taxonomicName>
|
||
and the
|
||
<taxonomicName id="4C5867C84C7BFFE5FF40FB92FEA9FBD9" ID-CoL="8LTCY" authority="Will, 2017" authorityName="Will" authorityYear="2017" box="[151,346,1052,1077]" class="Insecta" family="Carabidae" genus="Praeanchodemus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="113" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B92CC0594C7BFFE5FF40FB92FEA9FBD9" box="[151,346,1052,1077]" italics="true" pageId="3" pageNumber="113">Praeanchodemus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21F7D224C7BFFE5FEB7FB93FE58FBD8" box="[352,427,1053,1077]" pageId="3" pageNumber="113" rank="genus">gen. n.</taxonomicNameLabel>
|
||
appear unlikely.
|
||
</paragraph>
|
||
<paragraph id="8BE71C4B4C7BFFE5FF10FBB1FDEEF933" blockId="3.[151,1437,151,2013]" pageId="3" pageNumber="113">
|
||
<taxonomicName id="4C5867C84C7BFFE5FF10FBB1FEA5FBB4" ID-CoL="8KV4J" authorityName="Bonelli" authorityYear="1810" box="[199,342,1087,1112]" class="Insecta" family="Carabidae" genus="Anchomenus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="113" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B92CC0594C7BFFE5FF10FBB1FEA5FBB4" box="[199,342,1087,1112]" italics="true" pageId="3" pageNumber="113">Anchomenus</emphasis>
|
||
</taxonomicName>
|
||
clade
|
||
<emphasis id="B92CC0594C7BFFE5FE7DFBCFFE1BFBB4" box="[426,488,1089,1112]" italics="true" pageId="3" pageNumber="113">sensu</emphasis>
|
||
<bibRefCitation id="EFC961BA4C7BFFE5FE23FBB1FD56FBB4" author="Liebherr" box="[500,677,1087,1112]" pageId="3" pageNumber="113" refString="Liebherr, J. K. (1991) Phylogeny and ReVision of the Anchomenus Clade: The genera Tetraleucus, Anchomenus, Sericoda, and Elliptoleus (Coleoptera: Carabidae: Platynini). Bulletin of the American Museum of Natural History, 202, 1 - 163." type="journal article" year="1991">Liebherr (1991</bibRefCitation>
|
||
; compriSing the genera
|
||
<taxonomicName id="4C5867C84C7BFFE5FC11FBB1FAF3FBB4" ID-CoL="8KV4J" authority="Bonelli, 1810" authorityName="Bonelli" authorityYear="1810" box="[966,1280,1087,1112]" class="Insecta" family="Carabidae" genus="Anchomenus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="113" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B92CC0594C7BFFE5FC11FBB1FBA6FBB4" box="[966,1109,1087,1112]" italics="true" pageId="3" pageNumber="113">Anchomenus</emphasis>
|
||
Bonelli, 1810
|
||
</taxonomicName>
|
||
(
|
||
<emphasis id="B92CC0594C7BFFE5FAC2FBCFFA7FFBB4" box="[1301,1420,1087,1112]" italics="true" pageId="3" pageNumber="113">sensu lato</emphasis>
|
||
),
|
||
<taxonomicName id="4C5867C84C7BFFE5FF40FBEAFEE3FB91" ID-CoL="8KW3W" authorityName="Bates" authorityYear="1882" box="[151,272,1124,1149]" class="Insecta" family="Carabidae" genus="Elliptoleus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="113" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B92CC0594C7BFFE5FF40FBEAFEE3FB91" box="[151,272,1124,1149]" italics="true" pageId="3" pageNumber="113">Elliptoleus</emphasis>
|
||
</taxonomicName>
|
||
BateS, 1882,
|
||
<taxonomicName id="4C5867C84C7BFFE5FE66FBEAFD5AFB90" ID-CoL="8KYVP" authority="Kirby, 1837" authorityName="Kirby" authorityYear="1837" box="[433,681,1124,1149]" class="Insecta" family="Carabidae" genus="Sericoda" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="113" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B92CC0594C7BFFE5FE66FBEAFDE6FB91" box="[433,533,1124,1149]" italics="true" pageId="3" pageNumber="113">Sericoda</emphasis>
|
||
Kirby, 1837
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C5867C84C7BFFE5FD60FBEAFC20FB90" ID-CoL="8KZ7C" authority="CaSey, 1920" authorityName="CaSey" authorityYear="1920" box="[695,979,1124,1149]" class="Insecta" family="Carabidae" genus="Tetraleucus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="113" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B92CC0594C7BFFE5FD60FBEAFCCAFB91" box="[695,825,1124,1149]" italics="true" pageId="3" pageNumber="113">Tetraleucus</emphasis>
|
||
CaSey, 1920
|
||
</taxonomicName>
|
||
): The hypotheSiS of monophyly for thiS diverSe clade iS baSed on derived character StateS of the female internal genital tract. ThiS part of the tract, however, iS not preServed in the foSSil Specimen. Within the
|
||
<taxonomicName id="4C5867C84C7BFFE5FD3CFB22FC89FB29" ID-CoL="8KV4J" authorityName="Bonelli" authorityYear="1810" box="[747,890,1196,1221]" class="Insecta" family="Carabidae" genus="Anchomenus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="113" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B92CC0594C7BFFE5FD3CFB22FC89FB29" box="[747,890,1196,1221]" italics="true" pageId="3" pageNumber="113">Anchomenus</emphasis>
|
||
</taxonomicName>
|
||
clade an adjacent poSition of mentum Setae aS developed in
|
||
<taxonomicName id="4C5867C84C7BFFE5FEF8FB41FE02FB04" ID-CoL="8LTCY" authority="Will, 2017" authorityName="Will" authorityYear="2017" box="[303,497,1231,1256]" class="Insecta" family="Carabidae" genus="Praeanchodemus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="113" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B92CC0594C7BFFE5FEF8FB41FE02FB04" box="[303,497,1231,1256]" italics="true" pageId="3" pageNumber="113">Praeanchodemus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21F7D224C7BFFE5FE2EFB5FFDB6FB04" box="[505,581,1233,1257]" pageId="3" pageNumber="113" rank="genus">gen. n.</taxonomicNameLabel>
|
||
(character State 1) probably evolved independently Several timeS becauSe it iS obServed in the weStern Palearctic SpecieS of
|
||
<taxonomicName id="4C5867C84C7BFFE5FD6EFB7AFCBBFAE1" ID-CoL="8KV4J" authorityName="Bonelli" authorityYear="1810" box="[697,840,1268,1293]" class="Insecta" family="Carabidae" genus="Anchomenus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="113" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B92CC0594C7BFFE5FD6EFB7AFCBBFAE1" box="[697,840,1268,1293]" italics="true" pageId="3" pageNumber="113">Anchomenus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<taxonomicName id="4C5867C84C7BFFE5FC8DFB7AFB84FAE0" ID-CoL="DKVQ" authority="Dejean, 1828" authorityName="Dejean" authorityYear="1828" box="[858,1143,1268,1293]" class="Insecta" family="Carabidae" genus="Anchomenus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="113" phylum="Arthropoda" rank="species" species="cyaneus">
|
||
<emphasis id="B92CC0594C7BFFE5FC8DFB7AFC26FAE0" box="[858,981,1268,1292]" italics="true" pageId="3" pageNumber="113">A. cyaneus</emphasis>
|
||
Dejean, 1828
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C5867C84C7BFFE5FB53FB7AFF21FADC" ID-CoL="DKVX" authority="Pontoppidan, 1763" authorityName="Pontoppidan" authorityYear="1763" class="Insecta" family="Carabidae" genus="Anchomenus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="113" phylum="Arthropoda" rank="species" species="dorsalis">
|
||
<emphasis id="B92CC0594C7BFFE5FB53FB7AFB0DFAE1" box="[1156,1278,1268,1293]" italics="true" pageId="3" pageNumber="113">A. dorsalis</emphasis>
|
||
Pontoppidan, 1763
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C5867C84C7BFFE5FF0BFA96FD93FADC" ID-CoL="DKVS" authority="KryzhanovSkij, 1983" authorityName="KryzhanovSkij" authorityYear="1983" box="[220,608,1303,1328]" class="Insecta" family="Carabidae" genus="Anchomenus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="113" phylum="Arthropoda" rank="species" species="kurnakovi">
|
||
<emphasis id="B92CC0594C7BFFE5FF0BFA96FE98FADC" box="[220,363,1303,1328]" italics="true" pageId="3" pageNumber="113">A. kurnakovi</emphasis>
|
||
KryzhanovSkij, 1983
|
||
</taxonomicName>
|
||
, and the
|
||
<taxonomicName id="4C5867C84C7BFFE5FD15FA96FCC0FADC" ID-CoL="DKVR" authorityName="Fairmaire" authorityYear="1866" box="[706,819,1303,1328]" class="Insecta" family="Carabidae" genus="Anchomenus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="113" phylum="Arthropoda" rank="species" species="dohrnii">
|
||
<emphasis id="B92CC0594C7BFFE5FD15FA96FCC0FADC" box="[706,819,1303,1328]" italics="true" pageId="3" pageNumber="113">A. dohrnii</emphasis>
|
||
</taxonomicName>
|
||
SpecieS group
|
||
<emphasis id="B92CC0594C7BFFE5FC0DFA97FBEBFADC" box="[986,1048,1305,1328]" italics="true" pageId="3" pageNumber="113">sensu</emphasis>
|
||
<bibRefCitation id="EFC961BA4C7BFFE5FBC9FA99FB4CFADC" author="Schmidt" box="[1054,1215,1303,1328]" pageId="3" pageNumber="113" refString="Schmidt, J. (2014) Three new Anchomenus Bonelli, 1810, from the Western Palaearctic (Coleoptera: Carabidae: Platynini). Koleopterologische Rundschau, 84, 13 - 29." type="journal article" year="2014">Schmidt 2014</bibRefCitation>
|
||
), aS well aS in Some SpecieS of
|
||
<taxonomicName id="4C5867C84C7BFFE5FED9FAB2FE7BFAB9" ID-CoL="8KW3W" authorityName="Bates" authorityYear="1882" box="[270,392,1340,1365]" class="Insecta" family="Carabidae" genus="Elliptoleus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="113" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B92CC0594C7BFFE5FED9FAB2FE7BFAB9" box="[270,392,1340,1365]" italics="true" pageId="3" pageNumber="113">Elliptoleus</emphasis>
|
||
</taxonomicName>
|
||
BateS, 1882, and
|
||
<taxonomicName id="4C5867C84C7BFFE5FD85FAB2FCB5FAB8" ID-CoL="8KYVP" authority="Kirby, 1837" authorityName="Kirby" authorityYear="1837" box="[594,838,1340,1365]" class="Insecta" family="Carabidae" genus="Sericoda" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="113" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B92CC0594C7BFFE5FD85FAB2FD45FAB9" box="[594,694,1340,1365]" italics="true" pageId="3" pageNumber="113">Sericoda</emphasis>
|
||
Kirby, 1837
|
||
</taxonomicName>
|
||
. SpecieS of theSe groupS, however, differ by their leg chaetotaxy: pro- and meSofemur with additional Setae on ventral SurfaceS, metafemur dorSally with apical Setae developed, and all legS with the fifth tarSomere SetoSe on ventral Surface; the pleSiomorphic (character State 7) and reSpectively derived character StateS (8, 10) aS developed in
|
||
<taxonomicName id="4C5867C84C7BFFE5FC55FA29FBB7FA2C" ID-CoL="8LTCY" authority="Will, 2017" authorityName="Will" authorityYear="2017" box="[898,1092,1447,1472]" class="Insecta" family="Carabidae" genus="Praeanchodemus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="113" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B92CC0594C7BFFE5FC55FA29FBB7FA2C" box="[898,1092,1447,1472]" italics="true" pageId="3" pageNumber="113">Praeanchodemus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21F7D224C7BFFE5FB84FA26FB51FA53" box="[1107,1186,1448,1472]" pageId="3" pageNumber="113" rank="genus">gen. n</taxonomicNameLabel>
|
||
.. In addition, in the
|
||
<taxonomicName id="4C5867C84C7BFFE5FF40FA42FED5FA09" ID-CoL="8KV4J" authorityName="Bonelli" authorityYear="1810" box="[151,294,1484,1509]" class="Insecta" family="Carabidae" genus="Anchomenus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="113" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B92CC0594C7BFFE5FF40FA42FED5FA09" box="[151,294,1484,1509]" italics="true" pageId="3" pageNumber="113">Anchomenus</emphasis>
|
||
</taxonomicName>
|
||
clade, patternS of micro- and macroSculpture of the dorSal Surface of the body are remarkably different from thoSe of
|
||
<taxonomicName id="4C5867C84C7BFFE5FE76FA61FD90F9E4" ID-CoL="8LTCY" authority="Will, 2017" authorityName="Will" authorityYear="2017" box="[417,611,1519,1544]" class="Insecta" family="Carabidae" genus="Praeanchodemus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="113" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B92CC0594C7BFFE5FE76FA61FD90F9E4" box="[417,611,1519,1544]" italics="true" pageId="3" pageNumber="113">Praeanchodemus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21F7D224C7BFFE5FDBCFA7EFD4BF9EB" box="[619,696,1520,1544]" pageId="3" pageNumber="113" rank="genus">gen. n.</taxonomicNameLabel>
|
||
In three SpecieS,
|
||
<taxonomicName id="4C5867C84C7BFFE5FC57FA61FB8CF9E4" ID-CoL="DKW6" baseAuthorityName="Motschulsky" baseAuthorityYear="1865" box="[896,1151,1519,1544]" class="Insecta" family="Carabidae" genus="Anchomenus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="113" phylum="Arthropoda" rank="species" species="virescens">
|
||
<emphasis id="B92CC0594C7BFFE5FC57FA61FB8CF9E4" box="[896,1151,1519,1544]" italics="true" pageId="3" pageNumber="113">Anchomenus virescens</emphasis>
|
||
</taxonomicName>
|
||
MotSchulSky, 1864 from Middle ASia,
|
||
<taxonomicName id="4C5867C84C7BFFE5FEE5F99AFE44F9C1" ID-CoL="DKW2" authorityName="Bates" authorityYear="1873" box="[306,439,1556,1581]" class="Insecta" family="Carabidae" genus="Anchomenus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="113" phylum="Arthropoda" rank="species" species="leucopus">
|
||
<emphasis id="B92CC0594C7BFFE5FEE5F99AFE44F9C1" box="[306,439,1556,1581]" italics="true" pageId="3" pageNumber="113">A. leucopus</emphasis>
|
||
</taxonomicName>
|
||
BateS, 1873 from Far EaSt ASia, and
|
||
<taxonomicName id="4C5867C84C7BFFE5FCB3F99AFB85F9C1" ID-CoL="DKW7" authority="Habu, 1962" authorityName="Habu" authorityYear="1962" box="[868,1142,1556,1581]" class="Insecta" family="Carabidae" genus="Anchomenus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="113" phylum="Arthropoda" rank="species" species="yukihikoi">
|
||
<emphasis id="B92CC0594C7BFFE5FCB3F99AFC19F9C1" box="[868,1002,1556,1581]" italics="true" pageId="3" pageNumber="113">A. yukihikoi</emphasis>
|
||
Habu, 1962
|
||
</taxonomicName>
|
||
from Japan, a faint denSe puncturing iS developed on head and pronotum (in
|
||
<taxonomicName id="4C5867C84C7BFFE5FD0DF9B6FC92F9BC" ID-CoL="DKW6" baseAuthorityName="Motschulsky" baseAuthorityYear="1865" box="[730,865,1592,1616]" class="Insecta" family="Carabidae" genus="Anchomenus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="113" phylum="Arthropoda" rank="species" species="virescens">
|
||
<emphasis id="B92CC0594C7BFFE5FD0DF9B6FC92F9BC" box="[730,865,1592,1616]" italics="true" pageId="3" pageNumber="113">A. virescens</emphasis>
|
||
</taxonomicName>
|
||
on whole body Surface) that iS aSSociated with +/- diStinct microSetation. However, thiS puncturing doeS not appear to be homologouS to the diSSimilar, coarSe and irregularly arranged punctureS, which iS not aSSociated with Setation, on diSc of head found in the foSSil
|
||
<taxonomicName id="4C5867C84C7BFFE5FF40F92AFEA9F951" ID-CoL="8LTCY" authority="Will, 2017" authorityName="Will" authorityYear="2017" box="[151,346,1700,1725]" class="Insecta" family="Carabidae" genus="Praeanchodemus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="113" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B92CC0594C7BFFE5FF40F92AFEA9F951" box="[151,346,1700,1725]" italics="true" pageId="3" pageNumber="113">Praeanchodemus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21F7D224C7BFFE5FEB3F92BFE40F950" box="[356,435,1701,1725]" pageId="3" pageNumber="113" rank="genus">gen. n.</taxonomicNameLabel>
|
||
ConSequently, there iS no certain evidence for cloSe relationShipS of the
|
||
<taxonomicName id="4C5867C84C7BFFE5FADAF92AFA6FF951" ID-CoL="8KV4J" authorityName="Bonelli" authorityYear="1810" box="[1293,1436,1700,1725]" class="Insecta" family="Carabidae" genus="Anchomenus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="113" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B92CC0594C7BFFE5FADAF92AFA6FF951" box="[1293,1436,1700,1725]" italics="true" pageId="3" pageNumber="113">Anchomenus</emphasis>
|
||
</taxonomicName>
|
||
clade and
|
||
<taxonomicName id="4C5867C84C7BFFE5FEDFF949FE39F90C" ID-CoL="8LTCY" authority="Will, 2017" authorityName="Will" authorityYear="2017" box="[264,458,1735,1760]" class="Insecta" family="Carabidae" genus="Praeanchodemus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="113" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B92CC0594C7BFFE5FEDFF949FE39F90C" box="[264,458,1735,1760]" italics="true" pageId="3" pageNumber="113">Praeanchodemus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21F7D224C7BFFE5FE05F946FDEEF933" box="[466,541,1736,1760]" pageId="3" pageNumber="113" rank="genus">gen. n.</taxonomicNameLabel>
|
||
</paragraph>
|
||
<paragraph id="8BE71C4B4C7BFFE2FF10F962FBAAFEF0" blockId="3.[151,1437,151,2013]" lastBlockId="4.[151,1437,151,1581]" lastPageId="4" lastPageNumber="114" pageId="3" pageNumber="113">
|
||
Clade of
|
||
<taxonomicName id="4C5867C84C7BFFE5FEE5F962FDC0F8E8" ID-CoL="8KV2R" authority="Bonelli, 1810" authorityName="Bonelli" authorityYear="1810" box="[306,563,1772,1797]" class="Insecta" family="Carabidae" genus="Agonum" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="113" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B92CC0594C7BFFE5FEE5F962FE7CF8EF" box="[306,399,1772,1795]" italics="true" pageId="3" pageNumber="113">Agonum</emphasis>
|
||
Bonelli, 1810
|
||
</taxonomicName>
|
||
+
|
||
<taxonomicName id="4C5867C84C7BFFE5FD83F962FC8AF8E8" ID-CoL="8KV2G" authority="Jeannel, 1948" authorityName="Jeannel" authorityYear="1948" box="[596,889,1772,1797]" class="Insecta" family="Carabidae" genus="Agonidium" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="113" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B92CC0594C7BFFE5FD83F962FD3CF8E9" box="[596,719,1772,1797]" italics="true" pageId="3" pageNumber="113">Agonidium</emphasis>
|
||
Jeannel, 1948
|
||
</taxonomicName>
|
||
: For thiS clade, an adjacent poSition of mentum Setae iS conSidered a pleSiomorphic character State with multiple tranSformationS to a more lateral poSition (
|
||
<bibRefCitation id="EFC961BA4C7BFFE5FF49F8BAFE34F8A0" author="Liebherr" box="[158,455,1844,1869]" pageId="3" pageNumber="113" refString="Liebherr, J. K. & Schmidt, J. (2004) Phylogeny and biogeography of the Laurasian genus Agonum Bonelli (Coleoptera, Carabidae, Platynini). Mitteilungen des Museums fur Naturkunde Berlin, Deutsche entomologische Zeitschrift, 51 (2), 151 - 206. https: // doi. org / 10.1002 / mmnd. 20040510202" type="journal article" year="2004">Liebherr & Schmidt 2004</bibRefCitation>
|
||
). SpecieS of thiS clade Share the Shape of the head, with an indiStinct neck conStriction, with
|
||
<taxonomicName id="4C5867C84C7BFFE5FF07F8D9FE60F89C" ID-CoL="8LTCY" authority="Will, 2017" authorityName="Will" authorityYear="2017" box="[208,403,1879,1904]" class="Insecta" family="Carabidae" genus="Praeanchodemus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="113" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B92CC0594C7BFFE5FF07F8D9FE60F89C" box="[208,403,1879,1904]" italics="true" pageId="3" pageNumber="113">Praeanchodemus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21F7D224C7BFFE5FE4EF8D6FE16F883" box="[409,485,1880,1904]" pageId="3" pageNumber="113" rank="genus">gen. n.</taxonomicNameLabel>
|
||
(character State 3), however, they differ from the latter by the more derived Shape of pronotum with more rounded hind angleS. DeSpite of the pro- and meSofemoral Setation in Some SpecieS, in the
|
||
<taxonomicName id="4C5867C84C7BFFE5FF40F82FFF07F854" ID-CoL="8KV2R" authorityName="Bonelli" authorityYear="1810" box="[151,244,1953,1976]" class="Insecta" family="Carabidae" genus="Agonum" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="113" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B92CC0594C7BFFE5FF40F82FFF07F854" box="[151,244,1953,1976]" italics="true" pageId="3" pageNumber="113">Agonum</emphasis>
|
||
</taxonomicName>
|
||
+
|
||
<taxonomicName id="4C5867C84C7BFFE5FEC5F82EFE7EF855" ID-CoL="8KV2G" authorityName="Jeannel" authorityYear="1948" box="[274,397,1952,1977]" class="Insecta" family="Carabidae" genus="Agonidium" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="113" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B92CC0594C7BFFE5FEC5F82EFE7EF855" box="[274,397,1952,1977]" italics="true" pageId="3" pageNumber="113">Agonidium</emphasis>
|
||
</taxonomicName>
|
||
clade chaetotaxy patternS of legS are pleSiomorphic to what iS obServed in
|
||
<taxonomicName id="4C5867C84C7BFFE5FB0DF82EFA6FF855" ID-CoL="8LTCY" authority="Will, 2017" authorityName="Will" authorityYear="2017" box="[1242,1436,1952,1977]" class="Insecta" family="Carabidae" genus="Praeanchodemus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="113" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B92CC0594C7BFFE5FB0DF82EFA6FF855" box="[1242,1436,1952,1977]" italics="true" pageId="3" pageNumber="113">Praeanchodemus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21F7D224C7BFFE5FF40F84BFF17F830" box="[151,228,1989,2013]" pageId="3" pageNumber="113" rank="genus">gen. n.</taxonomicNameLabel>
|
||
Although a few SpecieS Share one or two of the above liSted derived character StateS 6 and 8-10 together with
|
||
<taxonomicName id="4C5867C84C7CFFE2FF02FF19FE64FF5C" ID-CoL="8LTCY" authority="Will, 2017" authorityName="Will" authorityYear="2017" box="[213,407,151,176]" class="Insecta" family="Carabidae" genus="Praeanchodemus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="114" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B92CC0594C7CFFE2FF02FF19FE64FF5C" box="[213,407,151,176]" italics="true" pageId="4" pageNumber="114">Praeanchodemus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21F7D224C7CFFE2FE73FF16FE05FF43" box="[420,502,152,176]" pageId="4" pageNumber="114" rank="genus">gen. n.</taxonomicNameLabel>
|
||
, thiS iS moSt likely the reSult of convergent evolution. AlSo in the
|
||
<taxonomicName id="4C5867C84C7CFFE2FAF5FF16FA8CFF43" ID-CoL="8KV2R" authorityName="Bonelli" authorityYear="1810" box="[1314,1407,152,175]" class="Insecta" family="Carabidae" genus="Agonum" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="114" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B92CC0594C7CFFE2FAF5FF16FA8CFF43" box="[1314,1407,152,175]" italics="true" pageId="4" pageNumber="114">Agonum</emphasis>
|
||
</taxonomicName>
|
||
+
|
||
<taxonomicName id="4C5867C84C7CFFE2FF40FF32FEE1FF39" ID-CoL="8KV2G" authorityName="Jeannel" authorityYear="1948" box="[151,274,188,213]" class="Insecta" family="Carabidae" genus="Agonidium" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="114" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B92CC0594C7CFFE2FF40FF32FEE1FF39" box="[151,274,188,213]" italics="true" pageId="4" pageNumber="114">Agonidium</emphasis>
|
||
</taxonomicName>
|
||
clade patternS of micro- and macroSculpture of the dorSal SurfaceS of the body are very differently developed from thoSe of
|
||
<taxonomicName id="4C5867C84C7CFFE2FE7AFF51FD83FF14" ID-CoL="8LTCY" authority="Will, 2017" authorityName="Will" authorityYear="2017" box="[429,624,223,248]" class="Insecta" family="Carabidae" genus="Praeanchodemus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="114" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B92CC0594C7CFFE2FE7AFF51FD83FF14" box="[429,624,223,248]" italics="true" pageId="4" pageNumber="114">Praeanchodemus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21F7D224C7CFFE2FDA1FF6EFD32FF1B" box="[630,705,224,248]" pageId="4" pageNumber="114" rank="genus">gen. n.</taxonomicNameLabel>
|
||
ConSequently, there iS no clear evidence for SynapomorphieS and thuS cloSe relationShipS of the
|
||
<emphasis id="B92CC0594C7CFFE2FE3CFE8AFD3FFEF1" box="[491,716,260,285]" italics="true" pageId="4" pageNumber="114">
|
||
<taxonomicName id="4C5867C84C7CFFE2FE3CFE8AFDBBFEF7" ID-CoL="8KV2R" authorityName="Bonelli" authorityYear="1810" box="[491,584,260,283]" class="Insecta" family="Carabidae" genus="Agonum" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="114" phylum="Arthropoda" rank="genus">Agonum</taxonomicName>
|
||
-
|
||
<taxonomicName id="4C5867C84C7CFFE2FD86FE8AFD3FFEF1" ID-CoL="8KV2G" authorityName="Jeannel" authorityYear="1948" box="[593,716,260,285]" class="Insecta" family="Carabidae" genus="Agonidium" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="114" phylum="Arthropoda" rank="genus">Agonidium</taxonomicName>
|
||
</emphasis>
|
||
clade and
|
||
<taxonomicName id="4C5867C84C7CFFE2FC93FE8AFBF4FEF1" ID-CoL="8LTCY" authority="Will, 2017" authorityName="Will" authorityYear="2017" box="[836,1031,260,285]" class="Insecta" family="Carabidae" genus="Praeanchodemus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="114" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B92CC0594C7CFFE2FC93FE8AFBF4FEF1" box="[836,1031,260,285]" italics="true" pageId="4" pageNumber="114">Praeanchodemus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21F7D224C7CFFE2FBD9FE8BFBAAFEF0" box="[1038,1113,261,285]" pageId="4" pageNumber="114" rank="genus">gen. n.</taxonomicNameLabel>
|
||
</paragraph>
|
||
<paragraph id="8BE71C4B4C7CFFE2FF10FEA9FE94FD44" blockId="4.[151,1437,151,1581]" pageId="4" pageNumber="114">
|
||
<taxonomicName id="4C5867C84C7CFFE2FF10FEA9FDCFFEAC" ID-CoL="8KVY4" authority="Bonelli, 1810 in SenSe" authorityName="Bonelli" authorityYear="1810" box="[199,572,295,320]" class="Insecta" family="Carabidae" genus="Platynus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="114" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B92CC0594C7CFFE2FF10FEA9FEDAFEAC" box="[199,297,295,320]" italics="true" pageId="4" pageNumber="114">Platynus</emphasis>
|
||
Bonelli,
|
||
<quantity id="4CA0B1AE4C7CFFE2FE4DFEA6FE07FEAC" box="[410,500,295,320]" metricMagnitude="1" metricUnit="m" metricValue="4.5973999999999995" pageId="4" pageNumber="114" unit="in" value="1810.0">1810 in</quantity>
|
||
SenSe
|
||
</taxonomicName>
|
||
of
|
||
<bibRefCitation id="EFC961BA4C7CFFE2FDBCFEA9FCECFEAC" author="Schmidt" box="[619,799,295,320]" pageId="4" pageNumber="114" refString="Schmidt, J. (2000) Ein Vorschlag zur Losung nomenklaturischer Probleme der Gattung Platynus Bonelli, 1810 (Coleoptera, Carabidae). Entomologische Blatter, Keltern, 96, 9 - 23." type="journal article" year="2000">Schmidt (2000)</bibRefCitation>
|
||
(= the
|
||
<emphasis id="B92CC0594C7CFFE2FCAAFEA9FC02FEAC" box="[893,1009,295,320]" italics="true" pageId="4" pageNumber="114">hypolithos</emphasis>
|
||
group in SenSe of
|
||
<bibRefCitation id="EFC961BA4C7CFFE2FB0EFEA9FA63FEAC" author="Liebherr" box="[1241,1424,295,320]" pageId="4" pageNumber="114" refString="Liebherr, J. K. (1989 b) Redescription of Platynus prognathus Van Dyke and circumscription of Lindroth's decentis and hypolithos groups. Journal of the New York Entomological Society, 97 (4), 430 - 437." type="journal article" year="1989" yearSuffix="b">Liebherr 1989b</bibRefCitation>
|
||
): Monophyly of
|
||
<taxonomicName id="4C5867C84C7CFFE2FE9DFEC2FE5FFE89" ID-CoL="8KVY4" authorityName="Bonelli" authorityYear="1810" box="[330,428,332,357]" class="Insecta" family="Carabidae" genus="Platynus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="114" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B92CC0594C7CFFE2FE9DFEC2FE5FFE89" box="[330,428,332,357]" italics="true" pageId="4" pageNumber="114">Platynus</emphasis>
|
||
</taxonomicName>
|
||
iS well Supported by the highly derived Shape and the Sclerotization patternS of the Spermatheca and the endophalluS, reSpectively. TheSe character StateS are unique within
|
||
<taxonomicName id="4C5867C84C7CFFE2FB68FEE1FF1AFE41" ID-CoL="8KTWY" authority="(Liebherr 1989b)" authorityName="Bonelli" authorityYear="1810" baseAuthorityName="Liebherr" baseAuthorityYear="1989" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="114" phylum="Arthropoda" rank="tribe" tribe="Platynini">
|
||
Platynini (
|
||
<bibRefCitation id="EFC961BA4C7CFFE2FAEDFEE1FF12FE41" author="Liebherr" pageId="4" pageNumber="114" refString="Liebherr, J. K. (1989 b) Redescription of Platynus prognathus Van Dyke and circumscription of Lindroth's decentis and hypolithos groups. Journal of the New York Entomological Society, 97 (4), 430 - 437." type="journal article" year="1989" yearSuffix="b">Liebherr 1989b</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
. However, while the foSSil Specimen iS a female, itS genital tract iS not Sufficiently preServed and So genital characterS cannot be conSidered here. Externally, SpecieS of
|
||
<taxonomicName id="4C5867C84C7CFFE2FCE9FE39FC53FE3C" ID-CoL="8KVY4" authorityName="Bonelli" authorityYear="1810" box="[830,928,439,464]" class="Insecta" family="Carabidae" genus="Platynus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="114" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B92CC0594C7CFFE2FCE9FE39FC53FE3C" box="[830,928,439,464]" italics="true" pageId="4" pageNumber="114">Platynus</emphasis>
|
||
</taxonomicName>
|
||
are characterized by a pair of reddiSh brown SpotS on neck which are probably baSed on different Sclerotization patternS of the head capSule (
|
||
<bibRefCitation id="EFC961BA4C7CFFE2FB3CFE52FA62FE18" author="Schmidt" box="[1259,1425,476,501]" pageId="4" pageNumber="114" refString="Schmidt, J. (2000) Ein Vorschlag zur Losung nomenklaturischer Probleme der Gattung Platynus Bonelli, 1810 (Coleoptera, Carabidae). Entomologische Blatter, Keltern, 96, 9 - 23." type="journal article" year="2000">Schmidt 2000</bibRefCitation>
|
||
). ThiS character State iS not realized in
|
||
<taxonomicName id="4C5867C84C7CFFE2FDEBFE71FD0DFDF4" ID-CoL="8LTCY" authority="Will, 2017" authorityName="Will" authorityYear="2017" box="[572,766,511,536]" class="Insecta" family="Carabidae" genus="Praeanchodemus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="114" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B92CC0594C7CFFE2FDEBFE71FD0DFDF4" box="[572,766,511,536]" italics="true" pageId="4" pageNumber="114">Praeanchodemus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21F7D224C7CFFE2FCD2FD8EFCA2FDFB" box="[773,849,512,536]" pageId="4" pageNumber="114" rank="genus">gen. n.</taxonomicNameLabel>
|
||
<taxonomicName id="4C5867C84C7CFFE2FC8EFE71FC48FDF4" ID-CoL="8KVY4" authorityName="Bonelli" authorityYear="1810" box="[857,955,511,536]" class="Insecta" family="Carabidae" genus="Platynus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="114" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B92CC0594C7CFFE2FC8EFE71FC48FDF4" box="[857,955,511,536]" italics="true" pageId="4" pageNumber="114">Platynus</emphasis>
|
||
</taxonomicName>
|
||
additionally differS from the foSSil SpecieS by having the head with a diStinct neck conStriction, with the pronotum having the lateral gutter and laterobaSal grooveS broadly developed, by a more pleSiomorphic chaetotaxy of the legS, and by different patternS of micro- and macroSculpture of the dorSal SurfaceS of the body. A cloSe relationShip of
|
||
<taxonomicName id="4C5867C84C7CFFE2FC03FDE2FB65FD69" ID-CoL="8LTCY" authority="Will, 2017" authorityName="Will" authorityYear="2017" box="[980,1174,620,645]" class="Insecta" family="Carabidae" genus="Praeanchodemus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="114" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B92CC0594C7CFFE2FC03FDE2FB65FD69" box="[980,1174,620,645]" italics="true" pageId="4" pageNumber="114">Praeanchodemus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21F7D224C7CFFE2FB4AFDE3FB1AFD68" box="[1181,1257,621,645]" pageId="4" pageNumber="114" rank="genus">gen. n.</taxonomicNameLabel>
|
||
and
|
||
<taxonomicName id="4C5867C84C7CFFE2FAF7FDE2FA71FD69" ID-CoL="8KVY4" authorityName="Bonelli" authorityYear="1810" box="[1312,1410,620,645]" class="Insecta" family="Carabidae" genus="Platynus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="114" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B92CC0594C7CFFE2FAF7FDE2FA71FD69" box="[1312,1410,620,645]" italics="true" pageId="4" pageNumber="114">Platynus</emphasis>
|
||
</taxonomicName>
|
||
iS thuS very unlikely.
|
||
</paragraph>
|
||
<paragraph id="8BE71C4B4C7CFFE2FF10FD3AFEA1FA2C" blockId="4.[151,1437,151,1581]" pageId="4" pageNumber="114">
|
||
<taxonomicName id="4C5867C84C7CFFE2FF10FD3AFDEBFD20" ID-CoL="8KY8T" authority="Habu, 1978" authorityName="Habu" authorityYear="1978" box="[199,536,692,717]" class="Insecta" family="Carabidae" genus="Paranchodemus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="114" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B92CC0594C7CFFE2FF10FD3AFE8EFD21" box="[199,381,692,717]" italics="true" pageId="4" pageNumber="114">Paranchodemus</emphasis>
|
||
<bibRefCitation id="EFC961BA4C7CFFE2FE5BFD3AFDEBFD20" author="Habu" box="[396,536,692,717]" pageId="4" pageNumber="114" refString="Habu, A. (1978) Fauna Japonica. Carabidae: Platynini (Insecta: Coleoptera). Keigaku Publishing, Tokyo, 447 pp., 36 pls." type="book" year="1978">Habu, 1978</bibRefCitation>
|
||
</taxonomicName>
|
||
and the
|
||
<emphasis id="B92CC0594C7CFFE2FD46FD3AFC83FD20" box="[657,880,692,717]" italics="true" pageId="4" pageNumber="114">
|
||
<taxonomicName id="4C5867C84C7CFFE2FD46FD3AFD1DFD21" ID-CoL="8KYQ4" authorityName="LeConte" authorityYear="1848" box="[657,750,692,717]" class="Insecta" family="Carabidae" genus="Rhadine" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="114" phylum="Arthropoda" rank="genus">Rhadine</taxonomicName>
|
||
-
|
||
<taxonomicName id="4C5867C84C7CFFE2FD2FFD3BFC83FD20" ID-CoL="8KX55" box="[760,880,693,716]" class="Insecta" family="Carabidae" genus="Tanystoma" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="114" phylum="Arthropoda" rank="genus">Tanystoma</taxonomicName>
|
||
</emphasis>
|
||
lineage
|
||
<emphasis id="B92CC0594C7CFFE2FC37FD3BFBEDFD20" box="[992,1054,693,716]" italics="true" pageId="4" pageNumber="114">sensu</emphasis>
|
||
<bibRefCitation id="EFC961BA4C7CFFE2FBF9FD3AFB12FD21" author="Liebherr" box="[1070,1249,692,717]" pageId="4" pageNumber="114" refString="Liebherr, J. K. (1985) ReVision of the platynine carabid genus Tanystoma Motschulsky (Coleoptera). Journal of the New York Entomological Society, 93 (4), 1182 - 1211." type="journal article" year="1985">Liebherr (1985</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFC961BA4C7CFFE2FB21FD3AFAC2FD21" author="Liebherr" box="[1270,1329,692,717]" pageId="4" pageNumber="114" refString="Liebherr, J. K. (1986) Cladistic analysis of North American Platynini and reVision of the Agonum extensicolle species group (Coleoptera: Carabidae). University of California Publications, Entomology, 106, V - x + 1 - 198." type="journal article" year="1986">1986</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFC961BA4C7CFFE2FA92FD3AFA63FD21" author="Liebherr" box="[1349,1424,692,717]" pageId="4" pageNumber="114" refString="Liebherr, J. K. (1989 b) Redescription of Platynus prognathus Van Dyke and circumscription of Lindroth's decentis and hypolithos groups. Journal of the New York Entomological Society, 97 (4), 430 - 437." type="journal article" year="1989" yearSuffix="b">1989b</bibRefCitation>
|
||
): Monophyly of the EaSt ASian
|
||
<taxonomicName id="4C5867C84C7CFFE2FDDDFD59FD33FD1C" ID-CoL="8KY8T" authorityName="Habu" authorityYear="1978" box="[522,704,727,752]" class="Insecta" family="Carabidae" genus="Paranchodemus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="114" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B92CC0594C7CFFE2FDDDFD59FD33FD1C" box="[522,704,727,752]" italics="true" pageId="4" pageNumber="114">Paranchodemus</emphasis>
|
||
</taxonomicName>
|
||
iS well Supported by Several SynapomorphieS of the SpecieS (
|
||
<bibRefCitation id="EFC961BA4C7CFFE2FF49FD72FEA6FCF8" author="Liebherr" box="[158,341,764,789]" pageId="4" pageNumber="114" refString="Liebherr, J. K. (1989 a) ReView of the Palearctic genus Paranchodemus Habu (Coleoptera: Carabidae: Platynini). Pan-Pacific Entomologist, 65 (1), 1 - 11." type="journal article" year="1989" yearSuffix="a">Liebherr 1989a</bibRefCitation>
|
||
). The author aSSumeS cloSe relationShipS of
|
||
<taxonomicName id="4C5867C84C7CFFE2FC88FD72FBE6FCF9" ID-CoL="8KY8T" authorityName="Habu" authorityYear="1978" box="[863,1045,764,789]" class="Insecta" family="Carabidae" genus="Paranchodemus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="114" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B92CC0594C7CFFE2FC88FD72FBE6FCF9" box="[863,1045,764,789]" italics="true" pageId="4" pageNumber="114">Paranchodemus</emphasis>
|
||
</taxonomicName>
|
||
with the North American genera
|
||
<taxonomicName id="4C5867C84C7CFFE2FF40FC91FE58FCD4" ID-CoL="8KYQ4" authority="LeConte, 1848" authorityName="LeConte" authorityYear="1848" box="[151,427,799,824]" class="Insecta" family="Carabidae" genus="Rhadine" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="114" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B92CC0594C7CFFE2FF40FC91FF07FCD4" box="[151,244,799,824]" italics="true" pageId="4" pageNumber="114">Rhadine</emphasis>
|
||
LeConte, 1848
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C5867C84C7CFFE2FE3CFCAFFCBAFCD4" ID-CoL="8KX55" authority="MotSchulSky" authorityName="Motschulsky" box="[491,841,799,824]" class="Insecta" family="Carabidae" genus="Tanystoma" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="121" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B92CC0594C7CFFE2FE3CFCAFFD90FCD4" box="[491,611,801,824]" italics="true" pageId="4" pageNumber="114">Tanystoma</emphasis>
|
||
MotSchulSky, 1865
|
||
</taxonomicName>
|
||
baSed on propoSed SynapomorphieS of the female genital tract (broad duct of the Spermathecal) and tarSal chaetotaxy (fourth tarSomere with dorSoapical Setae lacking). The latter character iS alSo obServed in
|
||
<taxonomicName id="4C5867C84C7CFFE2FD65FCE9FC87FC6C" ID-CoL="8LTCY" authority="Will, 2017" authorityName="Will" authorityYear="2017" box="[690,884,871,896]" class="Insecta" family="Carabidae" genus="Praeanchodemus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="114" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B92CC0594C7CFFE2FD65FCE9FC87FC6C" box="[690,884,871,896]" italics="true" pageId="4" pageNumber="114">Praeanchodemus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21F7D224C7CFFE2FCACFCE6FC35FC93" box="[891,966,872,896]" pageId="4" pageNumber="114" rank="genus">gen. n.</taxonomicNameLabel>
|
||
(character State 9). Remarkably, the foSSil taxon haS three additional derived characterS in common with
|
||
<taxonomicName id="4C5867C84C7CFFE2FC82FC02FBF8FC49" ID-CoL="8KY8T" authorityName="Habu" authorityYear="1978" box="[853,1035,908,933]" class="Insecta" family="Carabidae" genus="Paranchodemus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="114" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B92CC0594C7CFFE2FC82FC02FBF8FC49" box="[853,1035,908,933]" italics="true" pageId="4" pageNumber="114">Paranchodemus</emphasis>
|
||
</taxonomicName>
|
||
which, however, are not developed in the
|
||
<emphasis id="B92CC0594C7CFFE2FF09FC21FE4FFC24" box="[222,444,943,968]" italics="true" pageId="4" pageNumber="114">
|
||
<taxonomicName id="4C5867C84C7CFFE2FF09FC21FEC8FC24" ID-CoL="8KYQ4" authorityName="LeConte" authorityYear="1848" box="[222,315,943,968]" class="Insecta" family="Carabidae" genus="Rhadine" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="114" phylum="Arthropoda" rank="genus">Rhadine</taxonomicName>
|
||
-
|
||
<taxonomicName id="4C5867C84C7CFFE2FE93FC3FFE4FFC24" ID-CoL="8KX55" box="[324,444,945,968]" class="Insecta" family="Carabidae" genus="Tanystoma" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="114" phylum="Arthropoda" rank="genus">Tanystoma</taxonomicName>
|
||
</emphasis>
|
||
lineage: neck punctate (character State 2); pronotal lateral gutter narrow (character State 5); all legS with tarSomere 5 Smooth on ventral Surface (character State 10). One additional derived character
|
||
<taxonomicName id="4C5867C84C7CFFE2FF40FC79FEA9FBFC" ID-CoL="8LTCY" authority="Will, 2017" authorityName="Will" authorityYear="2017" box="[151,346,1015,1040]" class="Insecta" family="Carabidae" genus="Praeanchodemus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="114" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B92CC0594C7CFFE2FF40FC79FEA9FBFC" box="[151,346,1015,1040]" italics="true" pageId="4" pageNumber="114">Praeanchodemus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21F7D224C7CFFE2FEB5FC76FE43FBE3" box="[354,432,1016,1040]" pageId="4" pageNumber="114" rank="genus">gen. n.</taxonomicNameLabel>
|
||
ShareS together with
|
||
<taxonomicName id="4C5867C84C7CFFE2FD70FC79FCAEFBFC" ID-CoL="8KY8T" authorityName="Habu" authorityYear="1978" box="[679,861,1015,1040]" class="Insecta" family="Carabidae" genus="Paranchodemus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="114" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B92CC0594C7CFFE2FD70FC79FCAEFBFC" box="[679,861,1015,1040]" italics="true" pageId="4" pageNumber="114">Paranchodemus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C5867C84C7CFFE2FC4FFC77FBE3FBFC" ID-CoL="8KX55" box="[920,1040,1017,1040]" class="Insecta" family="Carabidae" genus="Tanystoma" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="114" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B92CC0594C7CFFE2FC4FFC77FBE3FBFC" box="[920,1040,1017,1040]" italics="true" pageId="4" pageNumber="114">Tanystoma</emphasis>
|
||
</taxonomicName>
|
||
, but not with
|
||
<taxonomicName id="4C5867C84C7CFFE2FB63FC79FAE2FBFC" ID-CoL="8KYQ4" authorityName="LeConte" authorityYear="1848" box="[1204,1297,1015,1040]" class="Insecta" family="Carabidae" genus="Rhadine" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="114" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B92CC0594C7CFFE2FB63FC79FAE2FBFC" box="[1204,1297,1015,1040]" italics="true" pageId="4" pageNumber="114">Rhadine</emphasis>
|
||
</taxonomicName>
|
||
: metafemur dorSally without apical Setae (character State 8). It thuS appearS poSSible that
|
||
<taxonomicName id="4C5867C84C7CFFE2FBCAFB92FB2CFBD9" ID-CoL="8LTCY" authority="Will, 2017" authorityName="Will" authorityYear="2017" box="[1053,1247,1052,1077]" class="Insecta" family="Carabidae" genus="Praeanchodemus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="114" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B92CC0594C7CFFE2FBCAFB92FB2CFBD9" box="[1053,1247,1052,1077]" italics="true" pageId="4" pageNumber="114">Praeanchodemus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21F7D224C7CFFE2FB3DFB93FACAFBD8" box="[1258,1337,1053,1077]" pageId="4" pageNumber="114" rank="genus">gen. n.</taxonomicNameLabel>
|
||
iS SiSter group of
|
||
<taxonomicName id="4C5867C84C7CFFE2FF28FBB1FE46FBB4" ID-CoL="8KY8T" authorityName="Habu" authorityYear="1978" box="[255,437,1087,1112]" class="Insecta" family="Carabidae" genus="Paranchodemus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="114" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B92CC0594C7CFFE2FF28FBB1FE46FBB4" box="[255,437,1087,1112]" italics="true" pageId="4" pageNumber="114">Paranchodemus</emphasis>
|
||
</taxonomicName>
|
||
, and both theSe taxa together with
|
||
<taxonomicName id="4C5867C84C7CFFE2FC95FBCFFC49FBB4" ID-CoL="8KX55" box="[834,954,1089,1112]" class="Insecta" family="Carabidae" genus="Tanystoma" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="114" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B92CC0594C7CFFE2FC95FBCFFC49FBB4" box="[834,954,1089,1112]" italics="true" pageId="4" pageNumber="114">Tanystoma</emphasis>
|
||
</taxonomicName>
|
||
form a monophyletic group. However, the phylogenetic Signal of the character StateS mentioned above iS unknown or weak, Since all but one (the punctate neck) were independently developed in other
|
||
<taxonomicName id="4C5867C84C7CFFE2FD78FB09FCE7FB4C" ID-CoL="8KTWY" authorityName="Bonelli" authorityYear="1810" box="[687,788,1159,1184]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="114" phylum="Arthropoda" rank="tribe" tribe="Platynini">Platynini</taxonomicName>
|
||
lineageS. In addition,
|
||
<taxonomicName id="4C5867C84C7CFFE2FBCAFB09FB20FB4C" ID-CoL="8KY8T" authorityName="Habu" authorityYear="1978" box="[1053,1235,1159,1184]" class="Insecta" family="Carabidae" genus="Paranchodemus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="114" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B92CC0594C7CFFE2FBCAFB09FB20FB4C" box="[1053,1235,1159,1184]" italics="true" pageId="4" pageNumber="114">Paranchodemus</emphasis>
|
||
</taxonomicName>
|
||
differS markedly from the foSSil
|
||
<taxonomicName id="4C5867C84C7CFFE2FE95FB22FDF6FB29" ID-CoL="8LTCY" authority="Will, 2017" authorityName="Will" authorityYear="2017" box="[322,517,1196,1221]" class="Insecta" family="Carabidae" genus="Praeanchodemus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="114" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B92CC0594C7CFFE2FE95FB22FDF6FB29" box="[322,517,1196,1221]" italics="true" pageId="4" pageNumber="114">Praeanchodemus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21F7D224C7CFFE2FDDBFB23FDABFB28" box="[524,600,1197,1221]" pageId="4" pageNumber="114" rank="genus">gen. n.</taxonomicNameLabel>
|
||
by a large Set of external characterS, e.g., much larger body Size, preSence of a diStinct neck conStriction, finely impreSSed SculpticellS of microSculpture on diSc of head, bifid mentum tooth, lack of the outer pair of Setae at Submentum, lack of Setae at pronotal laterobaSal angleS, preSence of additional Setae at apex of the Seventh elytral Stria, preSence of additional Setae near apical marginS of abdominal SegmentS V- VII, and by the Shape of the apical gonocoxite which iS markedly broad and Stout.
|
||
<taxonomicName id="4C5867C84C7CFFE2FBE9FAB3FB45FAB8" ID-CoL="8KX55" box="[1086,1206,1341,1364]" class="Insecta" family="Carabidae" genus="Tanystoma" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="114" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B92CC0594C7CFFE2FBE9FAB3FB45FAB8" box="[1086,1206,1341,1364]" italics="true" pageId="4" pageNumber="114">Tanystoma</emphasis>
|
||
</taxonomicName>
|
||
differS markedly by Short mandibleS, Stout mentum tooth, wide pronotal lateral gutter, triSetoSe metacoxa, preSence of additional Setae on ventral Surface of pro- and meSocoxae, SetoSe ventral Surface of fifth tarSomere, and preSence of microSetation on body Surface.
|
||
</paragraph>
|
||
<paragraph id="8BE71C4B4C7CFFE2FF10FA42FAEBF9C1" blockId="4.[151,1437,151,1581]" pageId="4" pageNumber="114">
|
||
ConSequently, although we point to Some evidence from external morphology that
|
||
<taxonomicName id="4C5867C84C7CFFE2FB55FA42FAB7FA09" ID-CoL="8LTCY" authority="Will, 2017" authorityName="Will" authorityYear="2017" box="[1154,1348,1484,1509]" class="Insecta" family="Carabidae" genus="Praeanchodemus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="114" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B92CC0594C7CFFE2FB55FA42FAB7FA09" box="[1154,1348,1484,1509]" italics="true" pageId="4" pageNumber="114">Praeanchodemus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21F7D224C7CFFE2FA99FA43FA6FFA08" box="[1358,1436,1485,1509]" pageId="4" pageNumber="114" rank="genus">gen. n.</taxonomicNameLabel>
|
||
could be part of a lineage compriSing the recent genera
|
||
<taxonomicName id="4C5867C84C7CFFE2FC9AFA61FBF0F9E4" ID-CoL="8KY8T" authorityName="Habu" authorityYear="1978" box="[845,1027,1519,1544]" class="Insecta" family="Carabidae" genus="Paranchodemus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="114" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B92CC0594C7CFFE2FC9AFA61FBF0F9E4" box="[845,1027,1519,1544]" italics="true" pageId="4" pageNumber="114">Paranchodemus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C5867C84C7CFFE2FBCEFA61FB85F9E4" ID-CoL="8KYQ4" authorityName="LeConte" authorityYear="1848" box="[1049,1142,1519,1544]" class="Insecta" family="Carabidae" genus="Rhadine" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="114" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B92CC0594C7CFFE2FBCEFA61FB85F9E4" box="[1049,1142,1519,1544]" italics="true" pageId="4" pageNumber="114">Rhadine</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C5867C84C7CFFE2FB15FA7FFAC9F9E4" ID-CoL="8KX55" box="[1218,1338,1521,1544]" class="Insecta" family="Carabidae" genus="Tanystoma" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="114" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B92CC0594C7CFFE2FB15FA7FFAC9F9E4" box="[1218,1338,1521,1544]" italics="true" pageId="4" pageNumber="114">Tanystoma</emphasis>
|
||
</taxonomicName>
|
||
, certain SynapomorphieS were not found and thuS, the actual relationShipS of the foSSil taxon remainS unSettled.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |