647 lines
98 KiB
XML
647 lines
98 KiB
XML
<document id="F405E3A9E6959BBBD524B6B5FEBC9DF6" ID-CLB-Dataset="284910" ID-DOI="10.5852/ejt.2023.908.2337" ID-GBIF-Dataset="6156c195-5953-4af2-9999-a1a03fd0f87e" ID-ISSN="2118-9773" ID-Zenodo-Dep="10149298" ID-ZooBank="1728B5B0-A130-4370-9610-9DA9CEDF4717" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1700226465688" checkinUser="plazi" docAuthor="Antić, Dragan, Šević, Mirko, Barjadze, Shalva & Makarov, Slobodan" docDate="2023" docId="03E187FCFFD8697CFDC70C3DFDAFF9ED" docLanguage="en" docName="EJT.2023.908.39-76.pdf" docOrigin="European Journal of Taxonomy 908" docSource="https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2337/10171" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Acanthophorella valerii Antic 2023, sp. nov." docType="treatment" docUuid="B24D09EE-1BDA-4F40-A853-F3280C49DD7B" docUuidSource="ZooBank" docVersion="2" lastPageNumber="64" masterDocId="FFD8FF84FFC96965FFF20A17FFD4FFE7" masterDocTitle="Review of the genus Acanthophorella Antić & Makarov, 2016 (Diplopoda, Chordeumatida, Anthroleucosomatidae), with descriptions of three new species from the Caucasus" masterLastPageNumber="76" masterPageNumber="39" pageNumber="56" updateTime="1705069979865" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="DE9E3992C17F4C6F0619772E3326CC7B">Review of the genus Acanthophorella Antić & Makarov, 2016 (Diplopoda, Chordeumatida, Anthroleucosomatidae), with descriptions of three new species from the Caucasus</mods:title>
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<mods:namePart id="E6930A236096E32E33B9CF760DE7D0E2">Antić, Dragan</mods:namePart>
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<mods:affiliation id="B5B8A79115A851F6670328EB7B81E6CD">University of Belgrade – Faculty of Biology, Institute of Zoology, Studentski Trg 16, 11000 Belgrade, Serbia. & Serbian Biospeleological Society, Trg Dositeja Obradovića 2, 21000 Novi Sad, Serbia.</mods:affiliation>
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<mods:nameIdentifier id="7AAC0B9CEF3699C5F4A5F25F689DC840" type="email">dragan.antic@bio.bg.ac.rs</mods:nameIdentifier>
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<mods:namePart id="FFAE5558F8C22A4BE61EA1D2042AD748">Šević, Mirko</mods:namePart>
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<mods:affiliation id="DB7E846ABFAD36252C7B45BE7672FC22">University of Belgrade – Faculty of Biology, Institute of Zoology, Studentski Trg 16, 11000 Belgrade, Serbia. & Serbian Biospeleological Society, Trg Dositeja Obradovića 2, 21000 Novi Sad, Serbia.</mods:affiliation>
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<mods:namePart id="A5DE91A3DFA4E56B63F1432E827C17D7">Barjadze, Shalva</mods:namePart>
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<mods:affiliation id="3CD96F53FC182ADFE58161707A657DF1">Institute of Zoology, Ilia State University, Giorgi Tsereteli 3, 0162, Tbilisi, Georgia.</mods:affiliation>
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<mods:affiliation id="9923B4E7480B9C069C1127D4687A56A4">University of Belgrade – Faculty of Biology, Institute of Zoology, Studentski Trg 16, 11000 Belgrade, Serbia. & Serbian Biospeleological Society, Trg Dositeja Obradovića 2, 21000 Novi Sad, Serbia.</mods:affiliation>
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<mods:title id="B328E987F824F5BB93B2AB17904EF489">European Journal of Taxonomy</mods:title>
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<subSubSection id="C3526561FFD86974FDC70C3DFC2BF9A2" box="[565,1023,1578,1605]" pageId="17" pageNumber="56" type="nomenclature">
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<heading id="D0BF8186FFD86974FDC70C3DFC2BF9A2" box="[565,1023,1578,1605]" centered="true" fontSize="11" level="2" pageId="17" pageNumber="56" reason="2">
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<taxonomicName id="4C484D69FFD86974FDC70C3DFC49F9A2" ID-CoL="BW7HR" authority="Antic" authorityName="Antic" authorityYear="2023" box="[565,925,1578,1605]" class="Diplopoda" family="Anthroleucosomatidae" genus="Acanthophorella" kingdom="Animalia" order="Chordeumatida" pageId="17" pageNumber="56" phylum="Arthropoda" rank="species" species="valerii" status="sp. nov.">
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<emphasis id="B93CEAF8FFD86974FDC70C3DFC81F9A2" bold="true" box="[565,853,1578,1605]" italics="true" pageId="17" pageNumber="56">Acanthophorella valerii</emphasis>
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Antić
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<taxonomicNameLabel id="A20F5783FFD86974FC560C3CFC2BF9A2" box="[932,1023,1579,1605]" pageId="17" pageNumber="56" rank="species">sp. nov.</taxonomicNameLabel>
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urn:lsid:zoobank.org:act:
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<paragraph id="8BF736EAFFD86974FD880C66FC6DF96C" blockId="17.[358,1228,1578,1675]" box="[634,953,1649,1675]" pageId="17" pageNumber="56">
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<figureCitation id="13732A6FFFD86974FD880C66FD2BF96C" box="[634,767,1649,1675]" captionStart="Fig" captionStartId="1.[189,232,1857,1883]" captionTargetBox="[189,1398,420,1810]" captionTargetId="figure-57@1.[189,1398,420,1810]" captionTargetPageId="1" captionText="Fig. 1. Troglobiotic members of the genus Acanthophorella Antić & Makarov, 2016 in situ. A–B. Acanthophorella barjadzei Antić & Makarov, 2016, ♂ from Nikortsminda Sakinule Cave. C–D. Acanthophorella barjadzei Antić & Makarov, 2016, ♂ from Muradi Cave. E–F. Acanthophorella devi Antić sp. nov., types, ♂♂. G–H. Acanthophorella valerii Antić sp. nov., holotype, ♂ (NHMW MY10366). Photos by Dragan Antić." figureDoi="http://doi.org/10.5281/zenodo.10149300" httpUri="https://zenodo.org/record/10149300/files/figure.png" pageId="17" pageNumber="56">Figs 1G–H</figureCitation>
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,
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<figureCitation id="13732A6FFFD86974FCF90C66FC82F96C" box="[779,854,1649,1675]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStart-3="Fig" captionStart-4="Fig" captionStart-5="Fig" captionStartId-0="19.[189,232,1892,1918]" captionStartId-1="20.[189,232,1857,1883]" captionStartId-2="21.[189,232,1929,1955]" captionStartId-3="22.[189,232,1787,1813]" captionStartId-4="23.[189,232,1787,1813]" captionStartId-5="24.[191,234,1963,1989]" captionTargetBox-0="[223,1365,265,1845]" captionTargetBox-1="[250,1338,265,1811]" captionTargetBox-2="[189,1398,437,1883]" captionTargetBox-3="[273,1315,265,1742]" captionTargetBox-4="[259,1317,275,1707]" captionTargetBox-5="[189,1398,1153,1920]" captionTargetId-0="figure-11@19.[223,1365,265,1845]" captionTargetId-1="figure-17@20.[250,1338,265,1811]" captionTargetId-2="figure-43@21.[189,1398,437,1883]" captionTargetId-3="figure-16@22.[273,1315,265,1742]" captionTargetId-4="figure-11@23.[247,1341,265,1747]" captionTargetId-5="figure-343@24.[189,1398,1151,1920]" captionTargetPageId-0="19" captionTargetPageId-1="20" captionTargetPageId-2="21" captionTargetPageId-3="22" captionTargetPageId-4="23" captionTargetPageId-5="24" captionText-0="Fig. 11. Acanthophorella valeriiAntić sp. nov. A, C, E, F. Holotype, ♂ (NHMW MY10366). B. Paratype, ♀ (IZB). D. Paratype, ♂ (IZB). A. Habitus, lateral view. B. Head, lateral view. C. Head, lateral view. D. Anterior part of the body, anterior and ventral view. E. Midbody rings, lateral view. F. Rings 15 and 16, dorsal view. Scale bars: A, D–F = 1 mm; B–C = 0.5 mm." captionText-1="Fig. 12. Acanthophorella valerii Antić sp. nov., paratype, ♂ (IZB). A. Labrum, anterior view. B. Left field with ommatidia and organ of Tömösvary, lateral view. C. Left antennomere 7, showing bacilliform sensillum. D. Left field with ommatidia, anterior view. E. Ring 15, dorsal view. F. Right interior macrochaeta of ring 15, dorsal view. G. Rings 14–16, dorsolateral view. H. Rings 14 and 15, anterior view. Scale bars: A–B, F = 0.1 mm; C–D = 0.01 mm; E = 0.2 mm; G, H = 0.5 mm." captionText-2="Fig. 13. Acanthophorella valerii Antić sp. nov. A. Paratype, ♀ (IZB). B–F. Paratype, ♂ (IZB). A. Legpair 2, anterior view.B. Leg-pair 2, posterior view.C. Leg-pair 7, posterior view.D. Leg-pair 10, anterior view. E. Anterior gonopods, anterior view. F. Posterior gonopods, posterior view. Scale bars = 0.2 mm." captionText-3="Fig. 14. Acanthophorella valerii Antić sp. nov. A–F. Paratype, ♂ (IZB). G–I. Paratype, ♀ (IZB). A. Anterior gonopods, anterior view. B. Anterior gonopods, posterodistal view. C. Anterior gonopods, left tuft, posterodistal view. D. Anterior gonopods, lateral view. E. Anterior gonopods, distal view. F. Anterior gonopods, laterodistal view. G. Vulvae, distal view. H. Left vulvae, lateral view. I. Left vulvae, anterior view. Abbreviations: a = angiocoxite; ap = anterior process; b = bursa; cv = syncoxal vesicle; ll = lateral lamella; mp = medial part; o = operculum; s = sternum; sa = synangiocoxal base; sl = sternal lamella; tf = angiocoxal tuft. Scale bars: A–B, D–I = 0.1 mm; C = 0.02 mm." captionText-4="Fig. 15. Acanthophorella valerii Antić sp. nov. A–E. Paratype, ♂ (IZB). F–G. Paratype ♀ (IZB). A. Anterior gonopods, anterior view. B. Anterior gonopods, posterior view. C. Anterior gonopods, lateral view. D. Posterior gonopods, posterior view. E. Left posterior gonopod, lateral view. F. Left vulva, distal view. G. Left vulva, lateral view. Abbreviations: a = angiocoxite; ap = anterior process; b = bursa; c = colpocoxite; cv = syncoxal vesicle; ll = lateral lamella; mp = medial part; o = operculum; s = sternum; sa = synangiocoxal base; sl = sternal lamella; t = telopodite; tf = angiocoxal tuft. Drawings by DA (A–E) and MŠ (F–G). Scale bars = 0.2 mm." captionText-5="Fig. 16. Dead adult specimen of Acanthophorella valerii Antić sp. nov. under the stone on the bank of the cave stream. Photo by Dragan Antić." figureDoi-0="http://doi.org/10.5281/zenodo.10149322" figureDoi-1="http://doi.org/10.5281/zenodo.10149324" figureDoi-2="http://doi.org/10.5281/zenodo.10149326" figureDoi-3="http://doi.org/10.5281/zenodo.10149328" figureDoi-4="http://doi.org/10.5281/zenodo.10149333" figureDoi-5="http://doi.org/10.5281/zenodo.10149335" httpUri-0="https://zenodo.org/record/10149322/files/figure.png" httpUri-1="https://zenodo.org/record/10149324/files/figure.png" httpUri-2="https://zenodo.org/record/10149326/files/figure.png" httpUri-3="https://zenodo.org/record/10149328/files/figure.png" httpUri-4="https://zenodo.org/record/10149333/files/figure.png" httpUri-5="https://zenodo.org/record/10149335/files/figure.png" pageId="17" pageNumber="56">11–16</figureCitation>
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,
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<figureCitation id="13732A6FFFD86974FC900C66FC44F96C" box="[866,912,1649,1675]" captionStart="Fig" captionStartId="33.[189,232,1822,1848]" captionTargetBox="[202,1378,705,1770]" captionTargetId="figure-170@33.[189,1398,686,1780]" captionTargetPageId="33" captionText="Fig. 21. Anterior gonopods in the genera Acanthophorella Antić & Makarov, 2016 and PseudoflagellophorellaAntić & Makarov, 2016, anterior views. A. A. auritaAntić sp. nov. B. A. barjadzei Antić & Makarov, 2016. C. A. chegemi Antić & Makarov, 2016. D. A. devi Antić sp. nov. E. A. irystoni Antić & Makarov, 2016. F. A. valerii Antić sp. nov. G. P. eskovi Antić & Makarov, 2016. H. P. mirabilis Antić & Makarov, 2016. I. P. papilioformis Antić & Makarov, 2016. B–C, E, G–I after Antić & Makarov (2016). Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10149345" httpUri="https://zenodo.org/record/10149345/files/figure.png" pageId="17" pageNumber="56">21F</figureCitation>
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,
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<figureCitation id="13732A6FFFD86974FC6E0C66FC6DF96C" box="[924,953,1649,1675]" captionStart="Fig" captionStartId="35.[189,232,1998,2024]" captionTargetBox="[189,1398,1153,1959]" captionTargetId="figure-355@35.[189,1398,1150,1959]" captionTargetPageId="35" captionText="Fig. 22. Distribution of the genus Acanthophorella Antić & Makarov, 2016." figureDoi="http://doi.org/10.5281/zenodo.10149347" httpUri="https://zenodo.org/record/10149347/files/figure.png" pageId="17" pageNumber="56">22</figureCitation>
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</paragraph>
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<subSubSection id="C3526561FFD86977FF4F0CA5FAB2FEA1" lastPageId="18" lastPageNumber="57" pageId="17" pageNumber="56" type="diagnosis">
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<paragraph id="8BF736EAFFD86974FF4F0CA5FEE2F92B" blockId="17.[189,310,1714,1740]" box="[189,310,1714,1740]" pageId="17" pageNumber="56">
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<heading id="D0BF8186FFD86974FF4F0CA5FEE2F92B" bold="true" box="[189,310,1714,1740]" fontSize="11" level="3" pageId="17" pageNumber="56" reason="3">
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<emphasis id="B93CEAF8FFD86974FF4F0CA5FEE2F92B" bold="true" box="[189,310,1714,1740]" pageId="17" pageNumber="56">Diagnosis</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BF736EAFFD86977FF4F0CF6FAB2FEA1" blockId="17.[189,1399,1761,1999]" lastBlockId="18.[188,1399,265,327]" lastPageId="18" lastPageNumber="57" pageId="17" pageNumber="56">
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Distinguished from congeners and members of the genus
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<taxonomicName id="4C484D69FFD86974FCA20CF6FBB5F91C" authorityName="Antic & Makarov" authorityYear="2016" box="[848,1121,1761,1787]" class="Diplopoda" family="Anthroleucosomatidae" genus="Pseudoflagellophorella" kingdom="Animalia" order="Chordeumatida" pageId="17" pageNumber="56" phylum="Arthropoda" rank="genus">
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<emphasis id="B93CEAF8FFD86974FCA20CF6FBB5F91C" box="[848,1121,1761,1787]" italics="true" pageId="17" pageNumber="56">Pseudoflagellophorella</emphasis>
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</taxonomicName>
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, except for
|
||
<taxonomicName id="4C484D69FFD86974FB150CF6FAA1F91C" authorityName="Antic & Makarov" authorityYear="2016" box="[1255,1397,1761,1787]" class="Diplopoda" family="Anthroleucosomatidae" genus="Acanthophorella" kingdom="Animalia" order="Chordeumatida" pageId="17" pageNumber="56" phylum="Arthropoda" rank="species" species="barjadzei">
|
||
<emphasis id="B93CEAF8FFD86974FB150CF6FAA1F91C" box="[1255,1397,1761,1787]" italics="true" pageId="17" pageNumber="56">A. barjadzei</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C484D69FFD86974FF000D12FE92F8F9" authority="Antić & Šević & Barjadze & Makarov, 2023" authorityName="Antić & Šević & Barjadze & Makarov" authorityYear="2023" box="[242,326,1796,1823]" class="Diplopoda" family="Anthroleucosomatidae" genus="Acanthophorella" kingdom="Animalia" order="Chordeumatida" pageId="17" pageNumber="56" phylum="Arthropoda" rank="species" species="devi" status="sp. nov.">
|
||
<emphasis id="B93CEAF8FFD86974FF000D12FE92F8F9" box="[242,326,1796,1823]" italics="true" pageId="17" pageNumber="56">A. devi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A20F5783FFD86974FEA20D12FE64F8F8" box="[336,432,1797,1823]" pageId="17" pageNumber="56" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, by a whitish and unpigmented body (vs pigmented body in others). From both latter it differs by the presence of 10–13 black ommatidia (vs a mainly smaller number of transparent to pale brownish ommatidia in
|
||
<taxonomicName id="4C484D69FFD86974FDC60D5CFD11F882" authorityName="Antic & Makarov" authorityYear="2016" box="[564,709,1867,1893]" class="Diplopoda" family="Anthroleucosomatidae" genus="Acanthophorella" kingdom="Animalia" order="Chordeumatida" pageId="17" pageNumber="56" phylum="Arthropoda" rank="species" species="barjadzei">
|
||
<emphasis id="B93CEAF8FFD86974FDC60D5CFD11F882" box="[564,709,1867,1893]" italics="true" pageId="17" pageNumber="56">A. barjadzei</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C484D69FFD86974FCF30D5CFC86F882" authorityName="Antić & Šević & Barjadze & Makarov" authorityYear="2023" box="[769,850,1867,1893]" class="Diplopoda" family="Anthroleucosomatidae" genus="Acanthophorella" kingdom="Animalia" order="Chordeumatida" pageId="17" pageNumber="56" phylum="Arthropoda" rank="species" species="devi">
|
||
<emphasis id="B93CEAF8FFD86974FCF30D5CFC86F882" box="[769,850,1867,1893]" italics="true" pageId="17" pageNumber="56">A. devi</emphasis>
|
||
</taxonomicName>
|
||
). Additionally, from
|
||
<taxonomicName id="4C484D69FFD86974FBBD0D5CFB34F882" authorityName="Antic & Makarov" authorityYear="2016" box="[1103,1248,1867,1893]" class="Diplopoda" family="Anthroleucosomatidae" genus="Acanthophorella" kingdom="Animalia" order="Chordeumatida" pageId="17" pageNumber="56" phylum="Arthropoda" rank="species" species="barjadzei">
|
||
<emphasis id="B93CEAF8FFD86974FBBD0D5CFB34F882" box="[1103,1248,1867,1893]" italics="true" pageId="17" pageNumber="56">A. barjadzei</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C484D69FFD86974FAEC0D5CFABBF882" authorityName="Antić & Šević & Barjadze & Makarov" authorityYear="2023" box="[1310,1391,1867,1893]" class="Diplopoda" family="Anthroleucosomatidae" genus="Acanthophorella" kingdom="Animalia" order="Chordeumatida" pageId="17" pageNumber="56" phylum="Arthropoda" rank="species" species="devi">
|
||
<emphasis id="B93CEAF8FFD86974FAEC0D5CFABBF882" box="[1310,1391,1867,1893]" italics="true" pageId="17" pageNumber="56">A. devi</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<taxonomicName id="4C484D69FFD86974FF4F0D79FE02F86F" authority="Antić & Šević & Barjadze & Makarov, 2023" authorityName="Antić & Šević & Barjadze & Makarov" authorityYear="2023" box="[189,470,1902,1928]" class="Diplopoda" family="Anthroleucosomatidae" genus="Acanthophorella" kingdom="Animalia" order="Chordeumatida" pageId="17" pageNumber="56" phylum="Arthropoda" rank="species" species="valerii" status="sp. nov.">
|
||
<emphasis id="B93CEAF8FFD86974FF4F0D79FE02F86F" box="[189,470,1902,1928]" italics="true" pageId="17" pageNumber="56">Acanthophorella valerii</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A20F5783FFD86974FE2F0D79FDECF86F" box="[477,568,1902,1928]" pageId="17" pageNumber="56" rank="species">sp. nov.</taxonomicNameLabel>
|
||
differs in the general shape of the anterior gonopods, the angiocoxites of the posterior gonopods being short, poorly developed (vs angiocoxites well-developed and strong in
|
||
<taxonomicName id="4C484D69FFD86974FF4F0DA2FE9AF829" authorityName="Antic & Makarov" authorityYear="2016" box="[189,334,1972,1999]" class="Diplopoda" family="Anthroleucosomatidae" genus="Acanthophorella" kingdom="Animalia" order="Chordeumatida" pageId="17" pageNumber="56" phylum="Arthropoda" rank="species" species="barjadzei">
|
||
<emphasis id="B93CEAF8FFD86974FF4F0DA2FE9AF829" box="[189,334,1972,1999]" italics="true" pageId="17" pageNumber="56">A. barjadzei</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C484D69FFD86974FE7B0DA2FE0FF829" authorityName="Antić & Šević & Barjadze & Makarov" authorityYear="2023" box="[393,475,1972,1999]" class="Diplopoda" family="Anthroleucosomatidae" genus="Acanthophorella" kingdom="Animalia" order="Chordeumatida" pageId="17" pageNumber="56" phylum="Arthropoda" rank="species" species="devi">
|
||
<emphasis id="B93CEAF8FFD86974FE7B0DA2FE0FF829" box="[393,475,1972,1999]" italics="true" pageId="17" pageNumber="56">A. devi</emphasis>
|
||
</taxonomicName>
|
||
), the coxal processes of leg-pair 7 without lateral lobe (vs lateral lobe present in
|
||
<taxonomicName id="4C484D69FFDB6977FF280B1EFEBDFEC4" authorityName="Antic & Makarov" authorityYear="2016" box="[218,361,265,291]" class="Diplopoda" family="Anthroleucosomatidae" genus="Acanthophorella" kingdom="Animalia" order="Chordeumatida" pageId="18" pageNumber="57" phylum="Arthropoda" rank="species" species="barjadzei">
|
||
<emphasis id="B93CEAF8FFDB6977FF280B1EFEBDFEC4" box="[218,361,265,291]" italics="true" pageId="18" pageNumber="57">A. barjadzei</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C484D69FFDB6977FE520B1EFE25FEC4" authorityName="Antić & Šević & Barjadze & Makarov" authorityYear="2023" box="[416,497,265,291]" class="Diplopoda" family="Anthroleucosomatidae" genus="Acanthophorella" kingdom="Animalia" order="Chordeumatida" pageId="18" pageNumber="57" phylum="Arthropoda" rank="species" species="devi">
|
||
<emphasis id="B93CEAF8FFDB6977FE520B1EFE25FEC4" box="[416,497,265,291]" italics="true" pageId="18" pageNumber="57">A. devi</emphasis>
|
||
</taxonomicName>
|
||
) and leg-pair 10 with well-developed coxal processes (vs leg-pair 10 without or with strongly reduced coxal protrusions only in
|
||
<taxonomicName id="4C484D69FFDB6977FCE20B3AFCB5FEA1" authorityName="Antić & Šević & Barjadze & Makarov" authorityYear="2023" box="[784,865,300,327]" class="Diplopoda" family="Anthroleucosomatidae" genus="Acanthophorella" kingdom="Animalia" order="Chordeumatida" pageId="18" pageNumber="57" phylum="Arthropoda" rank="species" species="devi">
|
||
<emphasis id="B93CEAF8FFDB6977FCE20B3AFCB5FEA1" box="[784,865,300,327]" italics="true" pageId="18" pageNumber="57">A. devi</emphasis>
|
||
</taxonomicName>
|
||
, or with strong protrusions in
|
||
<taxonomicName id="4C484D69FFDB6977FB370B3AFA81FEA1" authorityName="Antic & Makarov" authorityYear="2016" box="[1221,1365,300,327]" class="Diplopoda" family="Anthroleucosomatidae" genus="Acanthophorella" kingdom="Animalia" order="Chordeumatida" pageId="18" pageNumber="57" phylum="Arthropoda" rank="species" species="barjadzei">
|
||
<emphasis id="B93CEAF8FFDB6977FB370B3AFA81FEA1" box="[1221,1365,300,327]" italics="true" pageId="18" pageNumber="57">A. barjadzei</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3526561FFDB6977FF4F0B72FCDBFD0C" pageId="18" pageNumber="57" type="etymology">
|
||
<paragraph id="8BF736EAFFDB6977FF4F0B72FE90FE98" blockId="18.[189,324,357,383]" box="[189,324,357,383]" pageId="18" pageNumber="57">
|
||
<heading id="D0BF8186FFDB6977FF4F0B72FE90FE98" bold="true" box="[189,324,357,383]" fontSize="11" level="3" pageId="18" pageNumber="57" reason="3">
|
||
<emphasis id="B93CEAF8FFDB6977FF4F0B72FE90FE98" bold="true" box="[189,324,357,383]" pageId="18" pageNumber="57">Etymology</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BF736EAFFDB6977FF4F0B84FCDBFD0C" blockId="18.[189,1400,403,747]" pageId="18" pageNumber="57">
|
||
The new species is named after Valeri Barbakadze, a Georgian speleologist, cave diver and rescuer. He was an excellent and safe driver, guide and photographer of the cave interior, landscapes and our work during the 2022 expedition to Georgian caves. An interesting thing that gives this name a special charm happened right in front of the
|
||
<typeStatus id="54F38848FFDB6977FDEC0BEAFD85FDF0" box="[542,593,509,535]" pageId="18" pageNumber="57">type</typeStatus>
|
||
locality of this species, the Usholta Cave. At one point, all members of the expedition came out of the cave, except Valeri, who was still photographing the interior. After a while, the youngest member of the expedition, Luka Barjadze, became worried about Valeri (or most likely about who would drive us back) and started calling him and shouting ‘Valeriiiii’. As there is a river running through the cave, Valeri could not hear the call, so the youngster persistently repeated ‘Valeriiiii... Valeriiiii’. ‘Valeriiiii’ could (still) be heard hundreds of metres from the cave entrance. The epithet, a name in the genitive case, is a patronym.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3526561FFDB6977FF4F091FFDD3FBF0" pageId="18" pageNumber="57" type="materials_examined">
|
||
<paragraph id="8BF736EAFFDB6977FF4F091FFD43FCC5" blockId="18.[189,663,774,803]" box="[189,663,774,803]" pageId="18" pageNumber="57">
|
||
<emphasis id="B93CEAF8FFDB6977FF4F091FFE7EFCC5" bold="true" box="[189,426,776,802]" pageId="18" pageNumber="57">Material examined</emphasis>
|
||
(2 ♂♂,
|
||
<specimenCount id="9D4EFD63FFDB6977FDFE091EFDECFCC4" box="[524,568,777,803]" count="1" pageId="18" pageNumber="57" type="female">1 ♀</specimenCount>
|
||
, 8 juvs)
|
||
</paragraph>
|
||
<paragraph id="8BF736EAFFDB6977FF180920FA8FFC70" blockId="18.[189,1399,823,920]" pageId="18" pageNumber="57">
|
||
<materialsCitation id="3B203CB7FFDB6977FF180920FA8FFC70" collectingDate="2022-07-25" collectionCode="NHMW" collectorName="D. Antic & E. Kiria & L. Shavadze & S. Barjadze" country="Georgia" elevation="1772" location="Racha karst massif" municipality="Oni Municipality" pageId="18" pageNumber="57" specimenCode="MY10366" specimenCount="1" specimenCount-male="1" stateProvince="Racha" typeStatus="holotype">
|
||
<emphasis id="B93CEAF8FFDB6977FF180920FE88FCB6" bold="true" box="[234,348,823,849]" pageId="18" pageNumber="57">
|
||
<typeStatus id="54F38848FFDB6977FF180920FE88FCB6" box="[234,348,823,849]" pageId="18" pageNumber="57" type="holotype">Holotype</typeStatus>
|
||
</emphasis>
|
||
<collectingCountry id="F35F767AFFDB6977FF4F094DFEEBFC93" box="[189,319,858,884]" name="Georgia" pageId="18" pageNumber="57">GEORGIA</collectingCountry>
|
||
•
|
||
<specimenCount id="9D4EFD63FFDB6977FE9E094FFE57FC93" box="[364,387,856,884]" count="1" pageId="18" pageNumber="57" type="male">♂</specimenCount>
|
||
;
|
||
<collectingRegion id="498CF808FFDB6977FE69094DFE33FC93" box="[411,487,858,884]" pageId="18" pageNumber="57">Racha</collectingRegion>
|
||
,
|
||
<collectingMunicipality id="6B93AC90FFDB6977FE0F094DFD07FC93" box="[509,723,858,884]" pageId="18" pageNumber="57">Oni Municipality</collectingMunicipality>
|
||
,
|
||
<location id="8E976031FFDB6977FD1A094DFC08FC93" LSID="urn:lsid:plazi:treatment:03E187FCFFD8697CFDC70C3DFDAFF9ED:8E976031FFDB6977FD1A094DFC08FC93" box="[744,988,858,884]" country="Georgia" municipality="Oni Municipality" name="Racha karst massif" pageId="18" pageNumber="57" stateProvince="Racha">Racha karst massif</location>
|
||
,
|
||
<location id="8E976031FFDB6977FC03094DFB64FC93" LSID="urn:lsid:plazi:treatment:03E187FCFFD8697CFDC70C3DFDAFF9ED:8E976031FFDB6977FC03094DFB64FC93" box="[1009,1200,858,884]" country="Georgia" municipality="Oni Municipality" name="Usholta village" pageId="18" pageNumber="57" stateProvince="Racha">Usholta village</location>
|
||
,
|
||
<location id="8E976031FFDB6977FB34094DFAA6FC93" LSID="urn:lsid:plazi:treatment:03E187FCFFD8697CFDC70C3DFDAFF9ED:8E976031FFDB6977FB34094DFAA6FC93" box="[1222,1394,858,884]" country="Georgia" municipality="Oni Municipality" name="Usholta Cave" pageId="18" pageNumber="57" stateProvince="Racha">Usholta Cave</location>
|
||
;
|
||
<elevation id="0065D1D9FFDB6977FF4F0969FE82FC70" box="[189,342,893,920]" metricMagnitude="3" metricUnit="m" metricValue="1.7719999999999998" pageId="18" pageNumber="57" unit="m" value="1772.0">
|
||
<quantity id="4CB09B0FFFDB6977FF4F0969FEC2FC7F" box="[189,278,894,920]" metricMagnitude="3" metricUnit="m" metricValue="1.7719999999999998" pageId="18" pageNumber="57" unit="m" value="1772.0">1772 m</quantity>
|
||
a.s.l.
|
||
</elevation>
|
||
;
|
||
<date id="FFF6102AFFDB6977FE91096AFE23FC7F" box="[355,503,893,920]" pageId="18" pageNumber="57" value="2022-07-25">
|
||
<collectingDate id="EFB2E9C2FFDB6977FE91096AFE23FC7F" box="[355,503,893,920]" pageId="18" pageNumber="57" value="2022-07-25">25 Jul. 2022</collectingDate>
|
||
</date>
|
||
;
|
||
<collectorName id="26BD533CFFDB6977FDF10969FDBDFC70" box="[515,617,893,920]" pageId="18" pageNumber="57">D. Antić</collectorName>
|
||
,
|
||
<collectorName id="26BD533CFFDB6977FD870969FD01FC70" box="[629,725,893,920]" pageId="18" pageNumber="57">E. Kiria</collectorName>
|
||
,
|
||
<collectorName id="26BD533CFFDB6977FD130969FCA5FC70" box="[737,881,893,920]" pageId="18" pageNumber="57">L. Shavadze</collectorName>
|
||
and
|
||
<collectorName id="26BD533CFFDB6977FC590969FBE6FC70" box="[939,1074,893,920]" pageId="18" pageNumber="57">S. Barjadze</collectorName>
|
||
leg.;
|
||
<collectionCode id="ED59AE2FFFDB6977FB860969FB00FC7F" box="[1140,1236,894,920]" collectionName="Austria, Wien, Naturhistorisches Museum Wien" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="18" pageNumber="57" type="Museum">NHMW</collectionCode>
|
||
<specimenCode id="DBEE9E91FFDB6977FB29096AFA83FC70" box="[1243,1367,893,919]" collectionCode="NHMW" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="18" pageNumber="57" type="Museum">MY10366</specimenCode>
|
||
.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="8BF736EAFFDB6977FF1809A1FDD3FBF0" blockId="18.[189,1399,950,1047]" pageId="18" pageNumber="57">
|
||
<materialsCitation id="3B203CB7FFDB6977FF1809A1FB8CFC13" collectingDate="2022-07-25" collectionCode="IZB" collectorName="D. Antic & E. Kiria & L. Shavadze & S. Barjadze" country="Georgia" elevation="1772" location="GEORGIA" municipality="Oni Municipality" pageId="18" pageNumber="57" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Racha" typeStatus="paratype">
|
||
<emphasis id="B93CEAF8FFDB6977FF1809A1FEB3FC37" bold="true" box="[234,359,950,976]" pageId="18" pageNumber="57">
|
||
<typeStatus id="54F38848FFDB6977FF1809A1FEB3FC37" box="[234,359,950,976]" pageId="18" pageNumber="57" type="paratype">Paratypes</typeStatus>
|
||
</emphasis>
|
||
<collectingCountry id="F35F767AFFDB6977FF4F09CEFEEBFC14" box="[189,319,985,1011]" name="Georgia" pageId="18" pageNumber="57">GEORGIA</collectingCountry>
|
||
•
|
||
<specimenCount id="9D4EFD63FFDB6977FEA709CEFEABFC14" box="[341,383,983,1011]" count="1" pageId="18" pageNumber="57" type="male">1 ♂</specimenCount>
|
||
,
|
||
<specimenCount id="9D4EFD63FFDB6977FE7E09CEFE62FC13" box="[396,438,985,1012]" count="1" pageId="18" pageNumber="57" type="female">1 ♀</specimenCount>
|
||
(used for SEM); same collection data as for holotype;
|
||
<collectionCode id="ED59AE2FFFDB6977FBDB09CDFB8CFC13" box="[1065,1112,986,1012]" collectionName="IZB" pageId="18" pageNumber="57">IZB</collectionCode>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="3B203CB7FFDB6977FB9C09CEFDD0FBF0" collectingDate="2022-07-25" collectionCode="IZISU" collectorName="D. Antic & E. Kiria & L. Shavadze & S. Barjadze" country="Georgia" elevation="1772" location="Racha karst massif" municipality="Oni Municipality" pageId="18" pageNumber="57" specimenCount="8" stateProvince="Racha" typeStatus="paratype">
|
||
<specimenCount id="9D4EFD63FFDB6977FB9C09CEFB63FC14" box="[1134,1207,985,1011]" pageId="18" pageNumber="57">8 juvs</specimenCount>
|
||
; same collection data as for holotype;
|
||
<collectionCode id="ED59AE2FFFDB6977FE4709EAFDD0FBF0" box="[437,516,1021,1047]" collectionName="IZISU" pageId="18" pageNumber="57">IZISU</collectionCode>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3526561FFDB697CFF4F0E22FC4FFDB1" lastPageId="25" lastPageNumber="64" pageId="18" pageNumber="57" type="description">
|
||
<paragraph id="8BF736EAFFDB6977FF4F0E22FE9AFBA8" blockId="18.[189,334,1077,1103]" box="[189,334,1077,1103]" pageId="18" pageNumber="57">
|
||
<heading id="D0BF8186FFDB6977FF4F0E22FE9AFBA8" bold="true" box="[189,334,1077,1103]" fontSize="11" level="3" pageId="18" pageNumber="57" reason="3">
|
||
<emphasis id="B93CEAF8FFDB6977FF4F0E22FE9AFBA8" bold="true" box="[189,334,1077,1103]" pageId="18" pageNumber="57">Description</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BF736EAFFDB6977FF4F0E73FB04FB23" blockId="18.[188,1399,1123,1221]" pageId="18" pageNumber="57">
|
||
<smallCapsWord id="8D11A036FFDB6977FF4F0E73FF39FB9B" baselines="1143,1143" box="[189,237,1124,1150]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Size" pageId="18" pageNumber="57">SIZE</smallCapsWord>
|
||
<smallCapsWord id="8D11A036FFDB6977FF0A0E7EFEF1FB9B" baselines="1143" box="[248,293,1129,1148]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="and" pageId="18" pageNumber="57">AND</smallCapsWord>
|
||
<smallCapsWord id="8D11A036FFDB6977FEDD0E7EFE5CFB9B" baselines="1143" box="[303,392,1129,1148]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="number" pageId="18" pageNumber="57">NUMBER</smallCapsWord>
|
||
<smallCapsWord id="8D11A036FFDB6977FE600E7EFE78FB9B" baselines="1143" box="[402,428,1129,1148]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="of" pageId="18" pageNumber="57">OF</smallCapsWord>
|
||
<smallCapsWord id="8D11A036FFDB6977FE440E7EFE25FB9B" baselines="1143" box="[438,497,1129,1148]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="body" pageId="18" pageNumber="57">BODY</smallCapsWord>
|
||
<smallCapsWord id="8D11A036FFDB6977FE090E7EFDEDFB9B" baselines="1143" box="[507,569,1129,1148]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="rings" pageId="18" pageNumber="57">RINGS</smallCapsWord>
|
||
. Body with 31 rings (including telson).
|
||
<typeStatus id="54F38848FFDB6977FBD00E74FB5BFB9A" box="[1058,1167,1123,1149]" pageId="18" pageNumber="57" type="holotype">Holotype</typeStatus>
|
||
male
|
||
<quantity id="4CB09B0FFFDB6977FB2E0E74FAE5FB99" box="[1244,1329,1123,1150]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="18" pageNumber="57" unit="mm" value="15.0">15 mm</quantity>
|
||
long, vertical diameter of the largest ring
|
||
<quantity id="4CB09B0FFFDB6977FDA80E90FD11FB46" box="[602,709,1159,1185]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="18" pageNumber="57" unit="mm" value="1.1">1.10 mm</quantity>
|
||
.
|
||
<typeStatus id="54F38848FFDB6977FD3D0E90FCE2FB46" box="[719,822,1159,1185]" pageId="18" pageNumber="57" type="paratype">Paratype</typeStatus>
|
||
male
|
||
<quantity id="4CB09B0FFFDB6977FC890E90FC19FB46" box="[891,973,1159,1185]" metricMagnitude="-2" metricUnit="m" metricValue="1.4" pageId="18" pageNumber="57" unit="mm" value="14.0">14 mm</quantity>
|
||
long, vertical diameter of the largest ring 1.0 mm.
|
||
<typeStatus id="54F38848FFDB6977FEA90EBDFE16FB23" box="[347,450,1194,1220]" pageId="18" pageNumber="57" type="paratype">Paratype</typeStatus>
|
||
female
|
||
<quantity id="4CB09B0FFFDB6977FDD20EBDFDA7FB23" box="[544,627,1194,1221]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="18" pageNumber="57" unit="mm" value="15.0">15 mm</quantity>
|
||
long, vertical diameter of the largest ring
|
||
<quantity id="4CB09B0FFFDB6977FB920EBDFB18FB23" box="[1120,1228,1194,1221]" metricMagnitude="-3" metricUnit="m" metricValue="1.25" pageId="18" pageNumber="57" unit="mm" value="1.25">1.25 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BF736EAFFDB6977FF4F0EFCFE96FACF" blockId="18.[189,1399,1259,1320]" pageId="18" pageNumber="57">
|
||
<smallCapsWord id="8D11A036FFDB6977FF4F0EFCFE9CFAE3" baselines="1278,1279" box="[189,328,1259,1285]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Coloration" pageId="18" pageNumber="57">COLORATION</smallCapsWord>
|
||
(
|
||
<figureCitation id="13732A6FFFDB6977FEAC0EFCFE3DFAE1" box="[350,489,1259,1286]" captionStart="Fig" captionStartId="1.[189,232,1857,1883]" captionTargetBox="[189,1398,420,1810]" captionTargetId="figure-57@1.[189,1398,420,1810]" captionTargetPageId="1" captionText="Fig. 1. Troglobiotic members of the genus Acanthophorella Antić & Makarov, 2016 in situ. A–B. Acanthophorella barjadzei Antić & Makarov, 2016, ♂ from Nikortsminda Sakinule Cave. C–D. Acanthophorella barjadzei Antić & Makarov, 2016, ♂ from Muradi Cave. E–F. Acanthophorella devi Antić sp. nov., types, ♂♂. G–H. Acanthophorella valerii Antić sp. nov., holotype, ♂ (NHMW MY10366). Photos by Dragan Antić." figureDoi="http://doi.org/10.5281/zenodo.10149300" httpUri="https://zenodo.org/record/10149300/files/figure.png" pageId="18" pageNumber="57">Figs 1G–H</figureCitation>
|
||
,
|
||
<figureCitation id="13732A6FFFDB6977FE0B0EFCFDCCFAE2" box="[505,536,1259,1285]" captionStart="Fig" captionStartId="19.[189,232,1892,1918]" captionTargetBox="[223,1365,265,1845]" captionTargetId="figure-11@19.[223,1365,265,1845]" captionTargetPageId="19" captionText="Fig. 11. Acanthophorella valeriiAntić sp. nov. A, C, E, F. Holotype, ♂ (NHMW MY10366). B. Paratype, ♀ (IZB). D. Paratype, ♂ (IZB). A. Habitus, lateral view. B. Head, lateral view. C. Head, lateral view. D. Anterior part of the body, anterior and ventral view. E. Midbody rings, lateral view. F. Rings 15 and 16, dorsal view. Scale bars: A, D–F = 1 mm; B–C = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10149322" httpUri="https://zenodo.org/record/10149322/files/figure.png" pageId="18" pageNumber="57">11</figureCitation>
|
||
). Living animals dirty white, with white legs and antennae and black ommatidia.
|
||
</paragraph>
|
||
<paragraph id="8BF736EAFFDB6977FF4F0F46FC60F92D" blockId="18.[187,1399,1360,1739]" pageId="18" pageNumber="57">
|
||
<smallCapsWord id="8D11A036FFDB6977FF4F0F46FF28FA8E" baselines="1380,1380" box="[189,252,1361,1387]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Head" pageId="18" pageNumber="57">HEAD</smallCapsWord>
|
||
(
|
||
<figureCitation id="13732A6FFFDB6977FEE60F47FE73FA8C" box="[276,423,1360,1387]" captionStart="Fig" captionStartId="19.[189,232,1892,1918]" captionTargetBox="[223,1365,265,1845]" captionTargetId="figure-11@19.[223,1365,265,1845]" captionTargetPageId="19" captionText="Fig. 11. Acanthophorella valeriiAntić sp. nov. A, C, E, F. Holotype, ♂ (NHMW MY10366). B. Paratype, ♀ (IZB). D. Paratype, ♂ (IZB). A. Habitus, lateral view. B. Head, lateral view. C. Head, lateral view. D. Anterior part of the body, anterior and ventral view. E. Midbody rings, lateral view. F. Rings 15 and 16, dorsal view. Scale bars: A, D–F = 1 mm; B–C = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10149322" httpUri="https://zenodo.org/record/10149322/files/figure.png" pageId="18" pageNumber="57">Figs 11B–D</figureCitation>
|
||
,
|
||
<figureCitation id="13732A6FFFDB6977FE4E0F47FDC7FA8C" box="[444,531,1360,1387]" captionStart="Fig" captionStartId="20.[189,232,1857,1883]" captionTargetBox="[250,1338,265,1811]" captionTargetId="figure-17@20.[250,1338,265,1811]" captionTargetPageId="20" captionText="Fig. 12. Acanthophorella valerii Antić sp. nov., paratype, ♂ (IZB). A. Labrum, anterior view. B. Left field with ommatidia and organ of Tömösvary, lateral view. C. Left antennomere 7, showing bacilliform sensillum. D. Left field with ommatidia, anterior view. E. Ring 15, dorsal view. F. Right interior macrochaeta of ring 15, dorsal view. G. Rings 14–16, dorsolateral view. H. Rings 14 and 15, anterior view. Scale bars: A–B, F = 0.1 mm; C–D = 0.01 mm; E = 0.2 mm; G, H = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10149324" httpUri="https://zenodo.org/record/10149324/files/figure.png" pageId="18" pageNumber="57">12A–D</figureCitation>
|
||
). Setose, roundly convex in females, in males with labral and frontal surfaces flat with convexity between and a pair of poorly developed lateral lobes, each below antennal sockets. Labrum with three medial teeth and 4+4 labral and 2+2 supralabral setae. Promentum triangular, without setae. Lamellae linguales with 8+8 setae. Stipites with ca 30 setae each. Antennae
|
||
<quantity id="4CB09B0FFFDB6977FB120FAEFAEEFA33" box="[1248,1338,1465,1492]" metricMagnitude="-3" metricUnit="m" metricValue="2.6" pageId="18" pageNumber="57" unit="mm" value="2.6">2.6 mm</quantity>
|
||
long in
|
||
<typeStatus id="54F38848FFDB6977FF2C0FCAFE97FA10" box="[222,323,1501,1527]" pageId="18" pageNumber="57" type="paratype">paratype</typeStatus>
|
||
male. Length of antennomeres (in mm): I (0.1), II (0.25), III (0.74), IV (0.34), V (0.7), VI (0.21), VII (0.21) and VIII (0.05). Length/breadth ratios of antennomeres I–VII: I (1), II (1.7), III (6.7), IV (3.1), V (4.6), VI (1.4) and VII (1.8). Antennomeres II, IV, V, VI and VII with one, three, one, four and one long sensillum trichoideum, respectively. Antennomere 7 with one rather bacilliform sensillum (sensillum basiconicum?) curved distad, located below sensillum trichodeum (
|
||
<figureCitation id="13732A6FFFDB6977FB0B0C7DFABDF963" box="[1273,1385,1642,1668]" captionStart="Fig" captionStartId="20.[189,232,1857,1883]" captionTargetBox="[250,1338,265,1811]" captionTargetId="figure-17@20.[250,1338,265,1811]" captionTargetPageId="20" captionText="Fig. 12. Acanthophorella valerii Antić sp. nov., paratype, ♂ (IZB). A. Labrum, anterior view. B. Left field with ommatidia and organ of Tömösvary, lateral view. C. Left antennomere 7, showing bacilliform sensillum. D. Left field with ommatidia, anterior view. E. Ring 15, dorsal view. F. Right interior macrochaeta of ring 15, dorsal view. G. Rings 14–16, dorsolateral view. H. Rings 14 and 15, anterior view. Scale bars: A–B, F = 0.1 mm; C–D = 0.01 mm; E = 0.2 mm; G, H = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10149324" httpUri="https://zenodo.org/record/10149324/files/figure.png" pageId="18" pageNumber="57">Fig. 12C</figureCitation>
|
||
). Lateral to antennal sockets, a group of papilliform outgrowths present. Number of ommatidia:
|
||
<quantity id="4CB09B0FFFDB6977FAFD0C9AFAA3F940" box="[1295,1399,1677,1703]" metricMagnitude="-1" metricUnit="m" metricValue="2.921" metricValueMax="3.302" metricValueMin="2.54" pageId="18" pageNumber="57" unit="in" value="11.5" valueMax="13.0" valueMin="10.0">10–13 in</quantity>
|
||
3–4 rows, arranged in elongated triangles (
|
||
<figureCitation id="13732A6FFFDB6977FD400CA7FC96F92D" box="[690,834,1712,1738]" captionStart="Fig" captionStartId="19.[189,232,1892,1918]" captionTargetBox="[223,1365,265,1845]" captionTargetId="figure-11@19.[223,1365,265,1845]" captionTargetPageId="19" captionText="Fig. 11. Acanthophorella valeriiAntić sp. nov. A, C, E, F. Holotype, ♂ (NHMW MY10366). B. Paratype, ♀ (IZB). D. Paratype, ♂ (IZB). A. Habitus, lateral view. B. Head, lateral view. C. Head, lateral view. D. Anterior part of the body, anterior and ventral view. E. Midbody rings, lateral view. F. Rings 15 and 16, dorsal view. Scale bars: A, D–F = 1 mm; B–C = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10149322" httpUri="https://zenodo.org/record/10149322/files/figure.png" pageId="18" pageNumber="57">Figs 11B–C</figureCitation>
|
||
,
|
||
<figureCitation id="13732A6FFFDB6977FCBC0CA7FC72F92D" box="[846,934,1712,1738]" captionStart="Fig" captionStartId="20.[189,232,1857,1883]" captionTargetBox="[250,1338,265,1811]" captionTargetId="figure-17@20.[250,1338,265,1811]" captionTargetPageId="20" captionText="Fig. 12. Acanthophorella valerii Antić sp. nov., paratype, ♂ (IZB). A. Labrum, anterior view. B. Left field with ommatidia and organ of Tömösvary, lateral view. C. Left antennomere 7, showing bacilliform sensillum. D. Left field with ommatidia, anterior view. E. Ring 15, dorsal view. F. Right interior macrochaeta of ring 15, dorsal view. G. Rings 14–16, dorsolateral view. H. Rings 14 and 15, anterior view. Scale bars: A–B, F = 0.1 mm; C–D = 0.01 mm; E = 0.2 mm; G, H = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10149324" httpUri="https://zenodo.org/record/10149324/files/figure.png" pageId="18" pageNumber="57">12B, D</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BF736EAFFDB6977FF4F0CE5FDEBF8C8" blockId="18.[189,1398,1777,1839]" pageId="18" pageNumber="57">
|
||
<smallCapsWord id="8D11A036FFDB6977FF4F0CE5FECFF8ED" baselines="1797,1797" box="[189,283,1778,1804]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Collum" pageId="18" pageNumber="57">COLLUM</smallCapsWord>
|
||
. Narrower than head, with six macrochaetae as all body rings. Anterior edge semi-circular, posterior margin gently concave.
|
||
</paragraph>
|
||
<paragraph id="8BF736EAFFDB6977FF4F0D40FD7CF850" blockId="18.[189,1399,1878,1975]" pageId="18" pageNumber="57">
|
||
<smallCapsWord id="8D11A036FFDB6977FF4F0D40FF2AF888" baselines="1898,1898" box="[189,254,1879,1905]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Body" pageId="18" pageNumber="57">BODY</smallCapsWord>
|
||
<smallCapsWord id="8D11A036FFDB6977FEF50D4BFE91F888" baselines="1898" box="[263,325,1884,1903]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="rings" pageId="18" pageNumber="57">RINGS</smallCapsWord>
|
||
(
|
||
<figureCitation id="13732A6FFFDB6977FEAB0D41FDDCF896" box="[345,520,1878,1905]" captionStart="Fig" captionStartId="19.[189,232,1892,1918]" captionTargetBox="[223,1365,265,1845]" captionTargetId="figure-11@19.[223,1365,265,1845]" captionTargetPageId="19" captionText="Fig. 11. Acanthophorella valeriiAntić sp. nov. A, C, E, F. Holotype, ♂ (NHMW MY10366). B. Paratype, ♀ (IZB). D. Paratype, ♂ (IZB). A. Habitus, lateral view. B. Head, lateral view. C. Head, lateral view. D. Anterior part of the body, anterior and ventral view. E. Midbody rings, lateral view. F. Rings 15 and 16, dorsal view. Scale bars: A, D–F = 1 mm; B–C = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10149322" httpUri="https://zenodo.org/record/10149322/files/figure.png" pageId="18" pageNumber="57">Figs 11A, E–F</figureCitation>
|
||
,
|
||
<figureCitation id="13732A6FFFDB6977FDE80D41FDA7F897" box="[538,627,1878,1905]" captionStart="Fig" captionStartId="20.[189,232,1857,1883]" captionTargetBox="[250,1338,265,1811]" captionTargetId="figure-17@20.[250,1338,265,1811]" captionTargetPageId="20" captionText="Fig. 12. Acanthophorella valerii Antić sp. nov., paratype, ♂ (IZB). A. Labrum, anterior view. B. Left field with ommatidia and organ of Tömösvary, lateral view. C. Left antennomere 7, showing bacilliform sensillum. D. Left field with ommatidia, anterior view. E. Ring 15, dorsal view. F. Right interior macrochaeta of ring 15, dorsal view. G. Rings 14–16, dorsolateral view. H. Rings 14 and 15, anterior view. Scale bars: A–B, F = 0.1 mm; C–D = 0.01 mm; E = 0.2 mm; G, H = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10149324" httpUri="https://zenodo.org/record/10149324/files/figure.png" pageId="18" pageNumber="57">12E–H</figureCitation>
|
||
). With well-developed lateral keels, anterior margins rounded in dorsal view. Macrochaetae long and rather trichoid (
|
||
<figureCitation id="13732A6FFFDB6977FCE90D6EFC57F874" box="[795,899,1913,1939]" captionStart="Fig" captionStartId="20.[189,232,1857,1883]" captionTargetBox="[250,1338,265,1811]" captionTargetId="figure-17@20.[250,1338,265,1811]" captionTargetPageId="20" captionText="Fig. 12. Acanthophorella valerii Antić sp. nov., paratype, ♂ (IZB). A. Labrum, anterior view. B. Left field with ommatidia and organ of Tömösvary, lateral view. C. Left antennomere 7, showing bacilliform sensillum. D. Left field with ommatidia, anterior view. E. Ring 15, dorsal view. F. Right interior macrochaeta of ring 15, dorsal view. G. Rings 14–16, dorsolateral view. H. Rings 14 and 15, anterior view. Scale bars: A–B, F = 0.1 mm; C–D = 0.01 mm; E = 0.2 mm; G, H = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10149324" httpUri="https://zenodo.org/record/10149324/files/figure.png" pageId="18" pageNumber="57">Fig. 12F</figureCitation>
|
||
). CIX (ring 15) = 0.6; MIX (ring 15) ~ 1.6; PIX (ring 15) = 0.5; MA (ring 15) ~ 100°.
|
||
</paragraph>
|
||
<caption id="DF376662FFDA6976FF4F0D73FC50F80E" ID-DOI="http://doi.org/10.5281/zenodo.10149322" ID-Zenodo-Dep="10149322" httpUri="https://zenodo.org/record/10149322/files/figure.png" pageId="19" pageNumber="58" startId="19.[189,232,1892,1918]" targetBox="[223,1365,265,1845]" targetPageId="19">
|
||
<paragraph id="8BF736EAFFDA6976FF4F0D73FC50F80E" blockId="19.[187,1399,1891,2025]" pageId="19" pageNumber="58">
|
||
<emphasis id="B93CEAF8FFDA6976FF4F0D73FEC2F898" bold="true" box="[189,278,1892,1919]" pageId="19" pageNumber="58">Fig. 11.</emphasis>
|
||
<taxonomicName id="4C484D69FFDA6976FEE90D72FDA1F898" authority="Antic" authorityName="Antic" authorityYear="2023" box="[283,629,1893,1919]" class="Diplopoda" family="Anthroleucosomatidae" genus="Acanthophorella" kingdom="Animalia" order="Chordeumatida" pageId="19" pageNumber="58" phylum="Arthropoda" rank="species" species="valerii" status="sp. nov.">
|
||
<emphasis id="B93CEAF8FFDA6976FEE90D72FDE5F898" box="[283,561,1893,1919]" italics="true" pageId="19" pageNumber="58">Acanthophorella valerii</emphasis>
|
||
Antić
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A20F5783FFDA6976FD880D72FD06F898" box="[634,722,1893,1919]" pageId="19" pageNumber="58" rank="species">sp. nov.</taxonomicNameLabel>
|
||
<emphasis id="B93CEAF8FFDA6976FD250D73FD38F899" bold="true" box="[727,748,1892,1918]" pageId="19" pageNumber="58">A</emphasis>
|
||
,
|
||
<emphasis id="B93CEAF8FFDA6976FD0A0D73FCD9F899" bold="true" box="[760,781,1892,1918]" pageId="19" pageNumber="58">C</emphasis>
|
||
,
|
||
<emphasis id="B93CEAF8FFDA6976FCEB0D72FCF9F898" bold="true" box="[793,813,1893,1919]" pageId="19" pageNumber="58">E</emphasis>
|
||
,
|
||
<emphasis id="B93CEAF8FFDA6976FCCB0D72FC9FF898" bold="true" box="[825,843,1893,1919]" pageId="19" pageNumber="58">F</emphasis>
|
||
. Holotype, ♂ (NHMW MY10366).
|
||
<emphasis id="B93CEAF8FFDA6976FB1B0D72FB29F898" bold="true" box="[1257,1277,1893,1919]" pageId="19" pageNumber="58">B</emphasis>
|
||
. Paratype, ♀ (IZB).
|
||
<emphasis id="B93CEAF8FFDA6976FEDD0D9FFE90F845" bold="true" box="[303,324,1928,1954]" pageId="19" pageNumber="58">D</emphasis>
|
||
. Paratype, ♂ (IZB).
|
||
<emphasis id="B93CEAF8FFDA6976FDCF0D9FFD86F845" bold="true" box="[573,594,1928,1954]" pageId="19" pageNumber="58">A</emphasis>
|
||
. Habitus, lateral view.
|
||
<emphasis id="B93CEAF8FFDA6976FC950D9FFCAFF845" bold="true" box="[871,891,1928,1954]" pageId="19" pageNumber="58">B</emphasis>
|
||
. Head, lateral view.
|
||
<emphasis id="B93CEAF8FFDA6976FB800D9FFB53F845" bold="true" box="[1138,1159,1928,1954]" pageId="19" pageNumber="58">C</emphasis>
|
||
. Head, lateral view.
|
||
<emphasis id="B93CEAF8FFDA6976FF4F0DBCFF06F822" bold="true" box="[189,210,1963,1989]" pageId="19" pageNumber="58">D</emphasis>
|
||
. Anterior part of the body, anterior and ventral view.
|
||
<emphasis id="B93CEAF8FFDA6976FCB70DBCFC8CF822" bold="true" box="[837,856,1963,1989]" pageId="19" pageNumber="58">E</emphasis>
|
||
. Midbody rings, lateral view.
|
||
<emphasis id="B93CEAF8FFDA6976FB490DBCFB19F822" bold="true" box="[1211,1229,1963,1989]" pageId="19" pageNumber="58">F</emphasis>
|
||
. Rings 15 and 16, dorsal view. Scale bars: A, D–F = 1 mm; B–C = 0.5 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF376662FFDD6971FF4F0D56FB5FF80E" ID-DOI="http://doi.org/10.5281/zenodo.10149324" ID-Zenodo-Dep="10149324" httpUri="https://zenodo.org/record/10149324/files/figure.png" pageId="20" pageNumber="59" startId="20.[189,232,1857,1883]" targetBox="[250,1338,265,1811]" targetPageId="20">
|
||
<paragraph id="8BF736EAFFDD6971FF4F0D56FB5FF80E" blockId="20.[188,1399,1855,2025]" pageId="20" pageNumber="59">
|
||
<emphasis id="B93CEAF8FFDD6971FF4F0D56FEC8F8BC" bold="true" box="[189,284,1857,1883]" pageId="20" pageNumber="59">Fig. 12.</emphasis>
|
||
<taxonomicName id="4C484D69FFDD6971FED70D56FD53F8BC" authority="Antic" authorityName="Antic" authorityYear="2023" box="[293,647,1857,1883]" class="Diplopoda" family="Anthroleucosomatidae" genus="Acanthophorella" kingdom="Animalia" order="Chordeumatida" pageId="20" pageNumber="59" phylum="Arthropoda" rank="species" species="valerii" status="sp. nov.">
|
||
<emphasis id="B93CEAF8FFDD6971FED70D56FDEBF8BC" box="[293,575,1857,1883]" italics="true" pageId="20" pageNumber="59">Acanthophorella valerii</emphasis>
|
||
Antić
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A20F5783FFDD6971FD620D55FD3BF8BB" box="[656,751,1858,1884]" pageId="20" pageNumber="59" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, paratype, ♂ (IZB).
|
||
<emphasis id="B93CEAF8FFDD6971FC110D56FC2CF8BC" bold="true" box="[995,1016,1857,1883]" pageId="20" pageNumber="59">A</emphasis>
|
||
. Labrum, anterior view.
|
||
<emphasis id="B93CEAF8FFDD6971FAD10D55FAE1F8BB" bold="true" box="[1315,1333,1858,1884]" pageId="20" pageNumber="59">B</emphasis>
|
||
. Left field with ommatidia and organ of Tömösvary, lateral view.
|
||
<emphasis id="B93CEAF8FFDD6971FC870D73FC5EF899" bold="true" box="[885,906,1892,1918]" pageId="20" pageNumber="59">C</emphasis>
|
||
. Left antennomere 7, showing bacilliform sensillum.
|
||
<emphasis id="B93CEAF8FFDD6971FEB30D9FFE82F845" bold="true" box="[321,342,1928,1954]" pageId="20" pageNumber="59">D</emphasis>
|
||
. Left field with ommatidia, anterior view.
|
||
<emphasis id="B93CEAF8FFDD6971FC9C0D9FFC56F845" bold="true" box="[878,898,1928,1954]" pageId="20" pageNumber="59">E</emphasis>
|
||
. Ring 15, dorsal view.
|
||
<emphasis id="B93CEAF8FFDD6971FB5E0D9FFB6AF845" bold="true" box="[1196,1214,1928,1954]" pageId="20" pageNumber="59">F</emphasis>
|
||
. Right interior macrochaeta of ring 15, dorsal view.
|
||
<emphasis id="B93CEAF8FFDD6971FD880DBCFD5BF822" bold="true" box="[634,655,1963,1989]" pageId="20" pageNumber="59">G</emphasis>
|
||
. Rings 14–16, dorsolateral view.
|
||
<emphasis id="B93CEAF8FFDD6971FBD00DBCFBE3F822" bold="true" box="[1058,1079,1963,1989]" pageId="20" pageNumber="59">H</emphasis>
|
||
. Rings 14 and 15, anterior view. Scale bars: A–B, F = 0.1 mm; C–D = 0.01 mm; E = 0.2 mm; G, H = 0.5 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BF736EAFFDC6970FF4F0B1EFC29FEA0" blockId="21.[189,1398,265,327]" pageId="21" pageNumber="60">
|
||
<smallCapsWord id="8D11A036FFDC6970FF4F0B1EFEC6FEC6" baselines="284,284" box="[189,274,265,291]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Telson" pageId="21" pageNumber="60">TELSON</smallCapsWord>
|
||
. Epiproct with a pair of spinnerets and 3+3 setae (1+1 paramedian, 2+2 marginal). Hypoproct with 1+1 distal setae. Paraprocts with 3+3 marginal setae in distal part.
|
||
</paragraph>
|
||
<caption id="DF376662FFDC6970FF4F0D9EFAA3F80D" ID-DOI="http://doi.org/10.5281/zenodo.10149326" ID-Zenodo-Dep="10149326" httpUri="https://zenodo.org/record/10149326/files/figure.png" pageId="21" pageNumber="60" startId="21.[189,232,1929,1955]" targetBox="[189,1398,437,1883]" targetPageId="21">
|
||
<paragraph id="8BF736EAFFDC6970FF4F0D9EFAA3F80D" blockId="21.[187,1399,1927,2026]" pageId="21" pageNumber="60">
|
||
<emphasis id="B93CEAF8FFDC6970FF4F0D9EFECEF844" bold="true" box="[189,282,1929,1955]" pageId="21" pageNumber="60">Fig. 13.</emphasis>
|
||
<taxonomicName id="4C484D69FFDC6970FED00D9EFD55F844" authority="Antic" authorityName="Antic" authorityYear="2023" box="[290,641,1929,1955]" class="Diplopoda" family="Anthroleucosomatidae" genus="Acanthophorella" kingdom="Animalia" order="Chordeumatida" pageId="21" pageNumber="60" phylum="Arthropoda" rank="species" species="valerii" status="sp. nov.">
|
||
<emphasis id="B93CEAF8FFDC6970FED00D9EFDEFF844" box="[290,571,1929,1955]" italics="true" pageId="21" pageNumber="60">Acanthophorella valerii</emphasis>
|
||
Antić
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A20F5783FFDC6970FD7B0D9DFD30F843" box="[649,740,1930,1956]" pageId="21" pageNumber="60" rank="species">sp. nov.</taxonomicNameLabel>
|
||
<emphasis id="B93CEAF8FFDC6970FD190D9EFCD4F844" bold="true" box="[747,768,1929,1955]" pageId="21" pageNumber="60">A</emphasis>
|
||
. Paratype, ♀ (IZB).
|
||
<emphasis id="B93CEAF8FFDC6970FC010D9EFBFCF844" bold="true" box="[1011,1064,1929,1956]" pageId="21" pageNumber="60">B–F</emphasis>
|
||
. Paratype, ♂ (IZB).
|
||
<emphasis id="B93CEAF8FFDC6970FAE90D9EFAE4F844" bold="true" box="[1307,1328,1929,1955]" pageId="21" pageNumber="60">A</emphasis>
|
||
. Legpair 2, anterior view.
|
||
<emphasis id="B93CEAF8FFDC6970FE5D0DBAFE17F820" bold="true" box="[431,451,1965,1991]" pageId="21" pageNumber="60">B</emphasis>
|
||
. Leg-pair 2, posterior view.
|
||
<emphasis id="B93CEAF8FFDC6970FCFA0DBBFCC9F821" bold="true" box="[776,797,1964,1990]" pageId="21" pageNumber="60">C</emphasis>
|
||
. Leg-pair 7, posterior view.
|
||
<emphasis id="B93CEAF8FFDC6970FB900DBAFBA3F820" bold="true" box="[1122,1143,1965,1991]" pageId="21" pageNumber="60">D</emphasis>
|
||
. Leg-pair 10, anterior view.
|
||
<emphasis id="B93CEAF8FFDC6970FEF10DC7FEC3F80D" bold="true" box="[259,279,2000,2026]" pageId="21" pageNumber="60">E</emphasis>
|
||
. Anterior gonopods, anterior view.
|
||
<emphasis id="B93CEAF8FFDC6970FD450DC7FD1DF80D" bold="true" box="[695,713,2000,2026]" pageId="21" pageNumber="60">F</emphasis>
|
||
. Posterior gonopods, posterior view. Scale bars = 0.2 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF376662FFDF6973FF4F0CECFB59F80E" ID-DOI="http://doi.org/10.5281/zenodo.10149328" ID-Zenodo-Dep="10149328" httpUri="https://zenodo.org/record/10149328/files/figure.png" pageId="22" pageNumber="61" startId="22.[189,232,1787,1813]" targetBox="[273,1315,265,1742]" targetPageId="22">
|
||
<paragraph id="8BF736EAFFDF6973FF4F0CECFB59F80E" blockId="22.[189,1399,1785,2025]" pageId="22" pageNumber="61">
|
||
<emphasis id="B93CEAF8FFDF6973FF4F0CECFECBF8F2" bold="true" box="[189,287,1787,1813]" pageId="22" pageNumber="61">Fig. 14.</emphasis>
|
||
<taxonomicName id="4C484D69FFDF6973FEDE0CECFD41F8F2" authority="Antic" authorityName="Antic" authorityYear="2023" box="[300,661,1787,1813]" class="Diplopoda" family="Anthroleucosomatidae" genus="Acanthophorella" kingdom="Animalia" order="Chordeumatida" pageId="22" pageNumber="61" phylum="Arthropoda" rank="species" species="valerii" status="sp. nov.">
|
||
<emphasis id="B93CEAF8FFDF6973FEDE0CECFD9DF8F2" box="[300,585,1787,1813]" italics="true" pageId="22" pageNumber="61">Acanthophorella valerii</emphasis>
|
||
Antić
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A20F5783FFDF6973FD500CEBFCD5F8F1" box="[674,769,1788,1814]" pageId="22" pageNumber="61" rank="species">sp. nov.</taxonomicNameLabel>
|
||
<emphasis id="B93CEAF8FFDF6973FCFF0CECFC97F8F2" bold="true" box="[781,835,1787,1814]" pageId="22" pageNumber="61">A–F</emphasis>
|
||
. Paratype, ♂ (IZB).
|
||
<emphasis id="B93CEAF8FFDF6973FBB90CECFBAFF8F2" bold="true" box="[1099,1147,1787,1814]" pageId="22" pageNumber="61">G–I</emphasis>
|
||
. Paratype, ♀ (IZB).
|
||
<emphasis id="B93CEAF8FFDF6973FF4F0D09FF06F8DF" bold="true" box="[189,210,1822,1848]" pageId="22" pageNumber="61">A</emphasis>
|
||
. Anterior gonopods, anterior view.
|
||
<emphasis id="B93CEAF8FFDF6973FD880D09FD58F8DF" bold="true" box="[634,652,1822,1848]" pageId="22" pageNumber="61">B</emphasis>
|
||
. Anterior gonopods, posterodistal view.
|
||
<emphasis id="B93CEAF8FFDF6973FB9D0D09FB57F8DF" bold="true" box="[1135,1155,1822,1848]" pageId="22" pageNumber="61">C</emphasis>
|
||
. Anterior gonopods, left tuft, posterodistal view.
|
||
<emphasis id="B93CEAF8FFDF6973FDEA0D55FDF9F8BB" bold="true" box="[536,557,1858,1884]" pageId="22" pageNumber="61">D</emphasis>
|
||
. Anterior gonopods, lateral view.
|
||
<emphasis id="B93CEAF8FFDF6973FC210D55FC33F8BB" bold="true" box="[979,999,1858,1884]" pageId="22" pageNumber="61">E</emphasis>
|
||
. Anterior gonopods, distal view.
|
||
<emphasis id="B93CEAF8FFDF6973FF4F0D72FF1BF898" bold="true" box="[189,207,1893,1919]" pageId="22" pageNumber="61">F</emphasis>
|
||
. Anterior gonopods, laterodistal view.
|
||
<emphasis id="B93CEAF8FFDF6973FD550D73FD6AF899" bold="true" box="[679,702,1892,1918]" pageId="22" pageNumber="61">G</emphasis>
|
||
. Vulvae, distal view.
|
||
<emphasis id="B93CEAF8FFDF6973FC350D72FC0AF898" bold="true" box="[967,990,1893,1919]" pageId="22" pageNumber="61">H</emphasis>
|
||
. Left vulvae, lateral view.
|
||
<emphasis id="B93CEAF8FFDF6973FADA0D72FAE7F898" bold="true" box="[1320,1331,1893,1919]" pageId="22" pageNumber="61">I</emphasis>
|
||
. Left vulvae, anterior view. Abbreviations: a = angiocoxite; ap = anterior process; b = bursa; cv = syncoxal vesicle; ll = lateral lamella; mp = medial part; o = operculum; s = sternum; sa = synangiocoxal base; sl = sternal lamella; tf = angiocoxal tuft. Scale bars: A–B, D–I = 0.1 mm; C = 0.02 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF376662FFDE6972FF4F0CECFD24F80E" ID-DOI="http://doi.org/10.5281/zenodo.10149333" ID-Zenodo-Dep="10149333" httpUri="https://zenodo.org/record/10149333/files/figure.png" pageId="23" pageNumber="62" startId="23.[189,232,1787,1813]" targetBox="[259,1317,275,1707]" targetPageId="23">
|
||
<paragraph id="8BF736EAFFDE6972FF4F0CECFD24F80E" blockId="23.[189,1399,1785,2027]" pageId="23" pageNumber="62">
|
||
<emphasis id="B93CEAF8FFDE6972FF4F0CECFECBF8F2" bold="true" box="[189,287,1787,1813]" pageId="23" pageNumber="62">Fig. 15.</emphasis>
|
||
<taxonomicName id="4C484D69FFDE6972FEDE0CECFD41F8F2" authority="Antic" authorityName="Antic" authorityYear="2023" box="[300,661,1787,1813]" class="Diplopoda" family="Anthroleucosomatidae" genus="Acanthophorella" kingdom="Animalia" order="Chordeumatida" pageId="23" pageNumber="62" phylum="Arthropoda" rank="species" species="valerii" status="sp. nov.">
|
||
<emphasis id="B93CEAF8FFDE6972FEDE0CECFD9DF8F2" box="[300,585,1787,1813]" italics="true" pageId="23" pageNumber="62">Acanthophorella valerii</emphasis>
|
||
Antić
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A20F5783FFDE6972FD530CEBFCD5F8F1" box="[673,769,1788,1814]" pageId="23" pageNumber="62" rank="species">sp. nov.</taxonomicNameLabel>
|
||
<emphasis id="B93CEAF8FFDE6972FCFF0CECFC91F8F2" bold="true" box="[781,837,1787,1814]" pageId="23" pageNumber="62">A–E</emphasis>
|
||
. Paratype, ♂ (IZB).
|
||
<emphasis id="B93CEAF8FFDE6972FBBE0CECFB57F8F2" bold="true" box="[1100,1155,1787,1813]" pageId="23" pageNumber="62">F–G</emphasis>
|
||
. Paratype ♀ (IZB).
|
||
<emphasis id="B93CEAF8FFDE6972FF4F0D09FF06F8DF" bold="true" box="[189,210,1822,1848]" pageId="23" pageNumber="62">A</emphasis>
|
||
. Anterior gonopods, anterior view.
|
||
<emphasis id="B93CEAF8FFDE6972FD7E0D09FD74F8DF" bold="true" box="[652,672,1822,1848]" pageId="23" pageNumber="62">B</emphasis>
|
||
. Anterior gonopods, posterior view.
|
||
<emphasis id="B93CEAF8FFDE6972FB940D09FBAFF8DF" bold="true" box="[1126,1147,1822,1848]" pageId="23" pageNumber="62">C</emphasis>
|
||
. Anterior gonopods, lateral view.
|
||
<emphasis id="B93CEAF8FFDE6972FEA80D55FEBBF8BB" bold="true" box="[346,367,1858,1884]" pageId="23" pageNumber="62">D</emphasis>
|
||
. Posterior gonopods, posterior view.
|
||
<emphasis id="B93CEAF8FFDE6972FCC50D55FC9FF8BB" bold="true" box="[823,843,1858,1884]" pageId="23" pageNumber="62">E</emphasis>
|
||
. Left posterior gonopod, lateral view.
|
||
<emphasis id="B93CEAF8FFDE6972FAD00D55FAE0F8BB" bold="true" box="[1314,1332,1858,1884]" pageId="23" pageNumber="62">F</emphasis>
|
||
. Left vulva, distal view.
|
||
<emphasis id="B93CEAF8FFDE6972FE650D73FE7AF899" bold="true" box="[407,430,1892,1918]" pageId="23" pageNumber="62">G</emphasis>
|
||
. Left vulva, lateral view. Abbreviations: a = angiocoxite; ap = anterior process; b = bursa; c = colpocoxite; cv = syncoxal vesicle; ll = lateral lamella; mp = medial part; o = operculum; s = sternum; sa = synangiocoxal base; sl = sternal lamella; t = telopodite; tf = angiocoxal tuft. Drawings by DA (A–E) and MŠ (F–G). Scale bars = 0.2 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BF736EAFFD1697DFF4F0B1EFF2BFEA0" blockId="24.[189,1398,265,327]" pageId="24" pageNumber="63">
|
||
<smallCapsWord id="8D11A036FFD1697DFF4F0B1EFEFAFEC6" baselines="284,284,284,284" box="[189,302,265,291]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Leg-pairs" pageId="24" pageNumber="63">LEG-PAIRS</smallCapsWord>
|
||
1
|
||
<smallCapsWord id="8D11A036FFD1697DFEB90B18FEACFEC5" baselines="285" box="[331,376,271,290]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="and" pageId="24" pageNumber="63">AND</smallCapsWord>
|
||
2. In both sexes with tarsal combs; femora, postfemora and tibiae with long and robust setae.
|
||
</paragraph>
|
||
<paragraph id="8BF736EAFFD1697DFF4F0B79FB5BFD99" blockId="24.[189,1399,366,639]" pageId="24" pageNumber="63">
|
||
<smallCapsWord id="8D11A036FFD1697DFF4F0B79FF2BFE61" baselines="385,385" box="[189,255,366,392]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Male" pageId="24" pageNumber="63">MALE</smallCapsWord>
|
||
<smallCapsWord id="8D11A036FFD1697DFEFA0B64FE8EFE61" baselines="385" box="[264,346,371,390]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="sexual" pageId="24" pageNumber="63">SEXUAL</smallCapsWord>
|
||
<smallCapsWord id="8D11A036FFD1697DFE910B64FE38FE61" baselines="385" box="[355,492,371,390]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="characters" pageId="24" pageNumber="63">CHARACTERS</smallCapsWord>
|
||
(
|
||
<figureCitation id="13732A6FFFD1697DFE0D0B79FD53FE6F" box="[511,647,366,392]" captionStart="Fig" captionStartId="1.[189,232,1857,1883]" captionTargetBox="[189,1398,420,1810]" captionTargetId="figure-57@1.[189,1398,420,1810]" captionTargetPageId="1" captionText="Fig. 1. Troglobiotic members of the genus Acanthophorella Antić & Makarov, 2016 in situ. A–B. Acanthophorella barjadzei Antić & Makarov, 2016, ♂ from Nikortsminda Sakinule Cave. C–D. Acanthophorella barjadzei Antić & Makarov, 2016, ♂ from Muradi Cave. E–F. Acanthophorella devi Antić sp. nov., types, ♂♂. G–H. Acanthophorella valerii Antić sp. nov., holotype, ♂ (NHMW MY10366). Photos by Dragan Antić." figureDoi="http://doi.org/10.5281/zenodo.10149300" httpUri="https://zenodo.org/record/10149300/files/figure.png" pageId="24" pageNumber="63">Figs 1G–H</figureCitation>
|
||
,
|
||
<figureCitation id="13732A6FFFD1697DFD610B79FD3DFE6F" box="[659,745,366,392]" captionStart="Fig" captionStartId="21.[189,232,1929,1955]" captionTargetBox="[189,1398,437,1883]" captionTargetId="figure-43@21.[189,1398,437,1883]" captionTargetPageId="21" captionText="Fig. 13. Acanthophorella valerii Antić sp. nov. A. Paratype, ♀ (IZB). B–F. Paratype, ♂ (IZB). A. Legpair 2, anterior view.B. Leg-pair 2, posterior view.C. Leg-pair 7, posterior view.D. Leg-pair 10, anterior view. E. Anterior gonopods, anterior view. F. Posterior gonopods, posterior view. Scale bars = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10149326" httpUri="https://zenodo.org/record/10149326/files/figure.png" pageId="24" pageNumber="63">13B–D</figureCitation>
|
||
). Gonopores mesally on coxae 2 (
|
||
<figureCitation id="13732A6FFFD1697DFB710B79FB24FE6F" box="[1155,1264,366,392]" captionStart="Fig" captionStartId="21.[189,232,1929,1955]" captionTargetBox="[189,1398,437,1883]" captionTargetId="figure-43@21.[189,1398,437,1883]" captionTargetPageId="21" captionText="Fig. 13. Acanthophorella valerii Antić sp. nov. A. Paratype, ♀ (IZB). B–F. Paratype, ♂ (IZB). A. Legpair 2, anterior view.B. Leg-pair 2, posterior view.C. Leg-pair 7, posterior view.D. Leg-pair 10, anterior view. E. Anterior gonopods, anterior view. F. Posterior gonopods, posterior view. Scale bars = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10149326" httpUri="https://zenodo.org/record/10149326/files/figure.png" pageId="24" pageNumber="63">Fig. 13B</figureCitation>
|
||
). Leg-pairs 3–7 enlarged, especially leg-pairs 3, 4 and 7 (
|
||
<figureCitation id="13732A6FFFD1697DFD140B86FCB8FE4C" box="[742,876,401,427]" captionStart="Fig" captionStartId="1.[189,232,1857,1883]" captionTargetBox="[189,1398,420,1810]" captionTargetId="figure-57@1.[189,1398,420,1810]" captionTargetPageId="1" captionText="Fig. 1. Troglobiotic members of the genus Acanthophorella Antić & Makarov, 2016 in situ. A–B. Acanthophorella barjadzei Antić & Makarov, 2016, ♂ from Nikortsminda Sakinule Cave. C–D. Acanthophorella barjadzei Antić & Makarov, 2016, ♂ from Muradi Cave. E–F. Acanthophorella devi Antić sp. nov., types, ♂♂. G–H. Acanthophorella valerii Antić sp. nov., holotype, ♂ (NHMW MY10366). Photos by Dragan Antić." figureDoi="http://doi.org/10.5281/zenodo.10149300" httpUri="https://zenodo.org/record/10149300/files/figure.png" pageId="24" pageNumber="63">Fig. 1G, H</figureCitation>
|
||
). Leg-pairs 3 and 4 very thick, each with a proximal lateral protrusion on prefemora; prefemora and femora strong, rectangular; tarsi shorter and thicker compared to other legs; femora, postfemora and tibiae each with a distoventral pad. Leg-pair 5 with a proximal, anterior, triangular, coxal protrusion. Leg-pair 6 without peculiarities. Leg-pair 7 robust; coxae with wide, well-developed, flattened posterior processes, covered with long setae anteriorly and each with a mesal tooth (
|
||
<figureCitation id="13732A6FFFD1697DFE170856FD84FDBC" box="[485,592,577,603]" captionStart="Fig" captionStartId="21.[189,232,1929,1955]" captionTargetBox="[189,1398,437,1883]" captionTargetId="figure-43@21.[189,1398,437,1883]" captionTargetPageId="21" captionText="Fig. 13. Acanthophorella valerii Antić sp. nov. A. Paratype, ♀ (IZB). B–F. Paratype, ♂ (IZB). A. Legpair 2, anterior view.B. Leg-pair 2, posterior view.C. Leg-pair 7, posterior view.D. Leg-pair 10, anterior view. E. Anterior gonopods, anterior view. F. Posterior gonopods, posterior view. Scale bars = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10149326" httpUri="https://zenodo.org/record/10149326/files/figure.png" pageId="24" pageNumber="63">Fig. 13C</figureCitation>
|
||
). Leg-pair 10 with coxal glands and well-developed coxal processes oriented posteriad (
|
||
<figureCitation id="13732A6FFFD1697DFE530873FDD9FD99" box="[417,525,612,638]" captionStart="Fig" captionStartId="21.[189,232,1929,1955]" captionTargetBox="[189,1398,437,1883]" captionTargetId="figure-43@21.[189,1398,437,1883]" captionTargetPageId="21" captionText="Fig. 13. Acanthophorella valerii Antić sp. nov. A. Paratype, ♀ (IZB). B–F. Paratype, ♂ (IZB). A. Legpair 2, anterior view.B. Leg-pair 2, posterior view.C. Leg-pair 7, posterior view.D. Leg-pair 10, anterior view. E. Anterior gonopods, anterior view. F. Posterior gonopods, posterior view. Scale bars = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10149326" httpUri="https://zenodo.org/record/10149326/files/figure.png" pageId="24" pageNumber="63">Fig. 13D</figureCitation>
|
||
). Leg-pair 11 with coxal glands, no other peculiarities.
|
||
</paragraph>
|
||
<paragraph id="8BF736EAFFD1697DFF4F08B1FC4DFC3E" blockId="24.[189,1399,677,986]" pageId="24" pageNumber="63">
|
||
<smallCapsWord id="8D11A036FFD1697DFF4F08B1FEF8FD59" baselines="697,697" box="[189,300,678,704]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Anterior" pageId="24" pageNumber="63">ANTERIOR</smallCapsWord>
|
||
<smallCapsWord id="8D11A036FFD1697DFEC508BCFE7CFD59" baselines="697" box="[311,424,683,702]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="gonopods" pageId="24" pageNumber="63">GONOPODS</smallCapsWord>
|
||
(
|
||
<figureCitation id="13732A6FFFD1697DFE4E08B2FDFEFD27" box="[444,554,677,704]" captionStart="Fig" captionStartId="21.[189,232,1929,1955]" captionTargetBox="[189,1398,437,1883]" captionTargetId="figure-43@21.[189,1398,437,1883]" captionTargetPageId="21" captionText="Fig. 13. Acanthophorella valerii Antić sp. nov. A. Paratype, ♀ (IZB). B–F. Paratype, ♂ (IZB). A. Legpair 2, anterior view.B. Leg-pair 2, posterior view.C. Leg-pair 7, posterior view.D. Leg-pair 10, anterior view. E. Anterior gonopods, anterior view. F. Posterior gonopods, posterior view. Scale bars = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10149326" httpUri="https://zenodo.org/record/10149326/files/figure.png" pageId="24" pageNumber="63">Figs 13E</figureCitation>
|
||
,
|
||
<figureCitation id="13732A6FFFD1697DFDCE08B1FD58FD27" box="[572,652,678,704]" captionStart="Fig" captionStartId="22.[189,232,1787,1813]" captionTargetBox="[273,1315,265,1742]" captionTargetId="figure-16@22.[273,1315,265,1742]" captionTargetPageId="22" captionText="Fig. 14. Acanthophorella valerii Antić sp. nov. A–F. Paratype, ♂ (IZB). G–I. Paratype, ♀ (IZB). A. Anterior gonopods, anterior view. B. Anterior gonopods, posterodistal view. C. Anterior gonopods, left tuft, posterodistal view. D. Anterior gonopods, lateral view. E. Anterior gonopods, distal view. F. Anterior gonopods, laterodistal view. G. Vulvae, distal view. H. Left vulvae, lateral view. I. Left vulvae, anterior view. Abbreviations: a = angiocoxite; ap = anterior process; b = bursa; cv = syncoxal vesicle; ll = lateral lamella; mp = medial part; o = operculum; s = sternum; sa = synangiocoxal base; sl = sternal lamella; tf = angiocoxal tuft. Scale bars: A–B, D–I = 0.1 mm; C = 0.02 mm." figureDoi="http://doi.org/10.5281/zenodo.10149328" httpUri="https://zenodo.org/record/10149328/files/figure.png" pageId="24" pageNumber="63">14A–F</figureCitation>
|
||
,
|
||
<figureCitation id="13732A6FFFD1697DFD6C08B2FD22FD27" box="[670,758,677,704]" captionStart="Fig" captionStartId="23.[189,232,1787,1813]" captionTargetBox="[259,1317,275,1707]" captionTargetId="figure-11@23.[247,1341,265,1747]" captionTargetPageId="23" captionText="Fig. 15. Acanthophorella valerii Antić sp. nov. A–E. Paratype, ♂ (IZB). F–G. Paratype ♀ (IZB). A. Anterior gonopods, anterior view. B. Anterior gonopods, posterior view. C. Anterior gonopods, lateral view. D. Posterior gonopods, posterior view. E. Left posterior gonopod, lateral view. F. Left vulva, distal view. G. Left vulva, lateral view. Abbreviations: a = angiocoxite; ap = anterior process; b = bursa; c = colpocoxite; cv = syncoxal vesicle; ll = lateral lamella; mp = medial part; o = operculum; s = sternum; sa = synangiocoxal base; sl = sternal lamella; t = telopodite; tf = angiocoxal tuft. Drawings by DA (A–E) and MŠ (F–G). Scale bars = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10149333" httpUri="https://zenodo.org/record/10149333/files/figure.png" pageId="24" pageNumber="63">15A–C</figureCitation>
|
||
,
|
||
<figureCitation id="13732A6FFFD1697DFCF408B1FCE7FD27" box="[774,819,678,704]" captionStart="Fig" captionStartId="33.[189,232,1822,1848]" captionTargetBox="[202,1378,705,1770]" captionTargetId="figure-170@33.[189,1398,686,1780]" captionTargetPageId="33" captionText="Fig. 21. Anterior gonopods in the genera Acanthophorella Antić & Makarov, 2016 and PseudoflagellophorellaAntić & Makarov, 2016, anterior views. A. A. auritaAntić sp. nov. B. A. barjadzei Antić & Makarov, 2016. C. A. chegemi Antić & Makarov, 2016. D. A. devi Antić sp. nov. E. A. irystoni Antić & Makarov, 2016. F. A. valerii Antić sp. nov. G. P. eskovi Antić & Makarov, 2016. H. P. mirabilis Antić & Makarov, 2016. I. P. papilioformis Antić & Makarov, 2016. B–C, E, G–I after Antić & Makarov (2016). Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10149345" httpUri="https://zenodo.org/record/10149345/files/figure.png" pageId="24" pageNumber="63">21F</figureCitation>
|
||
). Gonopodal sternum (s) wide, medially with a moderately developed and poorly fimbriate lamella (sl) on anterior side and a bilobed membranous part above. Angiocoxites (a) consisting of a medial part (mp), lateral lamellae (ll) and a synangiocoxal base with anterior processes (ap). Medial parts well-developed, ear-shaped, divided, but appressed to each other, shieldlike, distomesal margins strongly denticulate posteriorly; angiocoxites posteroproximally with a pair of tufts (tf) with longer and shorter hairlike outgrowths, with a spinelike process and a rounded lobe, covered by spiculiform outgrowths. Lateral lamellae low, wide and spoon-shaped in lateral view, with denticulate margins. Anterior processes (ap) tapering distad, acuminate, twice as high as lateral lamellae. A syncoxal vesicle (cv) present posteriorly.
|
||
</paragraph>
|
||
<paragraph id="8BF736EAFFD1697CFF4F0E16FBE5FEA1" blockId="24.[189,1399,1024,1086]" lastBlockId="25.[189,1399,265,326]" lastPageId="25" lastPageNumber="64" pageId="24" pageNumber="63">
|
||
<smallCapsWord id="8D11A036FFD1697DFF4F0E16FEE6FBFE" baselines="1044,1044" box="[189,306,1025,1051]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Posterior" pageId="24" pageNumber="63">POSTERIOR</smallCapsWord>
|
||
<smallCapsWord id="8D11A036FFD1697DFECF0E11FE7AFBFE" baselines="1044" box="[317,430,1030,1049]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="gonopods" pageId="24" pageNumber="63">GONOPODS</smallCapsWord>
|
||
(
|
||
<figureCitation id="13732A6FFFD1697DFE300E17FDFFFBFC" box="[450,555,1024,1051]" captionStart="Fig" captionStartId="21.[189,232,1929,1955]" captionTargetBox="[189,1398,437,1883]" captionTargetId="figure-43@21.[189,1398,437,1883]" captionTargetPageId="21" captionText="Fig. 13. Acanthophorella valerii Antić sp. nov. A. Paratype, ♀ (IZB). B–F. Paratype, ♂ (IZB). A. Legpair 2, anterior view.B. Leg-pair 2, posterior view.C. Leg-pair 7, posterior view.D. Leg-pair 10, anterior view. E. Anterior gonopods, anterior view. F. Posterior gonopods, posterior view. Scale bars = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10149326" httpUri="https://zenodo.org/record/10149326/files/figure.png" pageId="24" pageNumber="63">Figs 13F</figureCitation>
|
||
,
|
||
<figureCitation id="13732A6FFFD1697DFDCF0E17FD45FBFC" box="[573,657,1024,1051]" captionStart="Fig" captionStartId="23.[189,232,1787,1813]" captionTargetBox="[259,1317,275,1707]" captionTargetId="figure-11@23.[247,1341,265,1747]" captionTargetPageId="23" captionText="Fig. 15. Acanthophorella valerii Antić sp. nov. A–E. Paratype, ♂ (IZB). F–G. Paratype ♀ (IZB). A. Anterior gonopods, anterior view. B. Anterior gonopods, posterior view. C. Anterior gonopods, lateral view. D. Posterior gonopods, posterior view. E. Left posterior gonopod, lateral view. F. Left vulva, distal view. G. Left vulva, lateral view. Abbreviations: a = angiocoxite; ap = anterior process; b = bursa; c = colpocoxite; cv = syncoxal vesicle; ll = lateral lamella; mp = medial part; o = operculum; s = sternum; sa = synangiocoxal base; sl = sternal lamella; t = telopodite; tf = angiocoxal tuft. Drawings by DA (A–E) and MŠ (F–G). Scale bars = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10149333" httpUri="https://zenodo.org/record/10149333/files/figure.png" pageId="24" pageNumber="63">15D–E</figureCitation>
|
||
). Gonopodal sternum (s) wide, well-developed. Angiocoxites (a) positioned posteriorly, poorly developed, short, much smaller than colpocoxties (c). Colpocoxites strongly developed, tapering in anterior and posterior views, but wide and spoon-shaped in lateral view; margins denticulate. Telopodites (t) small, rounded, placed posterolaterally.
|
||
</paragraph>
|
||
<caption id="DF376662FFD1697DFF4D0DBCFD4CF80F" ID-DOI="http://doi.org/10.5281/zenodo.10149335" ID-Zenodo-Dep="10149335" httpUri="https://zenodo.org/record/10149335/files/figure.png" pageId="24" pageNumber="63" startId="24.[191,234,1963,1989]" targetBox="[189,1398,1153,1920]" targetPageId="24">
|
||
<paragraph id="8BF736EAFFD1697DFF4D0DBCFD4CF80F" blockId="24.[189,1398,1963,2025]" pageId="24" pageNumber="63">
|
||
<emphasis id="B93CEAF8FFD1697DFF4D0DBCFECFF822" bold="true" box="[191,283,1963,1989]" pageId="24" pageNumber="63">Fig. 16.</emphasis>
|
||
Dead adult specimen of
|
||
<taxonomicName id="4C484D69FFD1697DFDB60DBCFC77F822" authority="Antic" authorityName="Antic" authorityYear="2023" box="[580,931,1963,1989]" class="Diplopoda" family="Anthroleucosomatidae" genus="Acanthophorella" kingdom="Animalia" order="Chordeumatida" pageId="24" pageNumber="63" phylum="Arthropoda" rank="species" species="valerii" status="sp. nov.">
|
||
<emphasis id="B93CEAF8FFD1697DFDB60DBCFC89F822" box="[580,861,1963,1989]" italics="true" pageId="24" pageNumber="63">Acanthophorella valerii</emphasis>
|
||
Antić
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A20F5783FFD1697DFC5E0DBBFBD3F821" box="[940,1031,1964,1990]" pageId="24" pageNumber="63" rank="species">sp. nov.</taxonomicNameLabel>
|
||
under the stone on the bank of the cave stream. Photo by Dragan Antić.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BF736EAFFD0697CFF4F0B79FDC9FE4C" blockId="25.[189,1398,366,427]" pageId="25" pageNumber="64">
|
||
<smallCapsWord id="8D11A036FFD0697CFF4F0B79FF3FFE61" baselines="385,385" box="[189,235,366,392]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Leg" pageId="25" pageNumber="64">LEG</smallCapsWord>
|
||
<smallCapsWord id="8D11A036FFD0697CFF010B64FEF4FE61" baselines="385" box="[243,288,371,390]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="pair" pageId="25" pageNumber="64">PAIR</smallCapsWord>
|
||
2
|
||
<smallCapsWord id="8D11A036FFD0697CFEB20B64FE82FE61" baselines="385" box="[320,342,371,390]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="in" pageId="25" pageNumber="64">IN</smallCapsWord>
|
||
<smallCapsWord id="8D11A036FFD0697CFEAC0B64FE68FE61" baselines="385" box="[350,444,371,390]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="females" pageId="25" pageNumber="64">FEMALES</smallCapsWord>
|
||
(
|
||
<figureCitation id="13732A6FFFD0697CFE3D0B79FDE9FE6F" box="[463,573,366,392]" captionStart="Fig" captionStartId="21.[189,232,1929,1955]" captionTargetBox="[189,1398,437,1883]" captionTargetId="figure-43@21.[189,1398,437,1883]" captionTargetPageId="21" captionText="Fig. 13. Acanthophorella valerii Antić sp. nov. A. Paratype, ♀ (IZB). B–F. Paratype, ♂ (IZB). A. Legpair 2, anterior view.B. Leg-pair 2, posterior view.C. Leg-pair 7, posterior view.D. Leg-pair 10, anterior view. E. Anterior gonopods, anterior view. F. Posterior gonopods, posterior view. Scale bars = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10149326" httpUri="https://zenodo.org/record/10149326/files/figure.png" pageId="25" pageNumber="64">Fig. 13A</figureCitation>
|
||
). Coxae with poorly developed distomesal protrusions covered with a few small tubercles and setae.
|
||
</paragraph>
|
||
<paragraph id="8BF736EAFFD0697CFF4F0BC4FC4FFDB1" blockId="25.[188,1399,466,598]" pageId="25" pageNumber="64">
|
||
<smallCapsWord id="8D11A036FFD0697CFF4F0BC4FEC0FE0C" baselines="486,486" box="[189,276,467,493]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Vulvae" pageId="25" pageNumber="64">VULVAE</smallCapsWord>
|
||
(
|
||
<figureCitation id="13732A6FFFD0697CFED60BC5FE7DFE0A" box="[292,425,466,493]" captionStart="Fig" captionStartId="22.[189,232,1787,1813]" captionTargetBox="[273,1315,265,1742]" captionTargetId="figure-16@22.[273,1315,265,1742]" captionTargetPageId="22" captionText="Fig. 14. Acanthophorella valerii Antić sp. nov. A–F. Paratype, ♂ (IZB). G–I. Paratype, ♀ (IZB). A. Anterior gonopods, anterior view. B. Anterior gonopods, posterodistal view. C. Anterior gonopods, left tuft, posterodistal view. D. Anterior gonopods, lateral view. E. Anterior gonopods, distal view. F. Anterior gonopods, laterodistal view. G. Vulvae, distal view. H. Left vulvae, lateral view. I. Left vulvae, anterior view. Abbreviations: a = angiocoxite; ap = anterior process; b = bursa; cv = syncoxal vesicle; ll = lateral lamella; mp = medial part; o = operculum; s = sternum; sa = synangiocoxal base; sl = sternal lamella; tf = angiocoxal tuft. Scale bars: A–B, D–I = 0.1 mm; C = 0.02 mm." figureDoi="http://doi.org/10.5281/zenodo.10149328" httpUri="https://zenodo.org/record/10149328/files/figure.png" pageId="25" pageNumber="64">Figs 14G–I</figureCitation>
|
||
,
|
||
<figureCitation id="13732A6FFFD0697CFE440BC5FDDCFE0B" box="[438,520,466,493]" captionStart="Fig" captionStartId="23.[189,232,1787,1813]" captionTargetBox="[259,1317,275,1707]" captionTargetId="figure-11@23.[247,1341,265,1747]" captionTargetPageId="23" captionText="Fig. 15. Acanthophorella valerii Antić sp. nov. A–E. Paratype, ♂ (IZB). F–G. Paratype ♀ (IZB). A. Anterior gonopods, anterior view. B. Anterior gonopods, posterior view. C. Anterior gonopods, lateral view. D. Posterior gonopods, posterior view. E. Left posterior gonopod, lateral view. F. Left vulva, distal view. G. Left vulva, lateral view. Abbreviations: a = angiocoxite; ap = anterior process; b = bursa; c = colpocoxite; cv = syncoxal vesicle; ll = lateral lamella; mp = medial part; o = operculum; s = sternum; sa = synangiocoxal base; sl = sternal lamella; t = telopodite; tf = angiocoxal tuft. Drawings by DA (A–E) and MŠ (F–G). Scale bars = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10149333" httpUri="https://zenodo.org/record/10149333/files/figure.png" pageId="25" pageNumber="64">15F–G</figureCitation>
|
||
). Rounded, as long as wide. Operculum (o) well-developed, bilobed, lateral lobe with three, mesal with two setae. Bursa (b) with strongly thickened anteroproximal lips on which the operculum rests. Lateral valve with a rounded lobe carrying six setae, mesal valve with a rounded lobe carrying five setae with an additional seta below the lobe.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3526561FFD0697CFF4F086AFAA3FA98" pageId="25" pageNumber="64" type="biology_ecology">
|
||
<paragraph id="8BF736EAFFD0697CFF4F086AFE6BFD70" blockId="25.[189,447,637,663]" box="[189,447,637,663]" pageId="25" pageNumber="64">
|
||
<heading id="D0BF8186FFD0697CFF4F086AFE6BFD70" bold="true" box="[189,447,637,663]" fontSize="11" level="3" pageId="25" pageNumber="64" reason="3">
|
||
<emphasis id="B93CEAF8FFD0697CFF4F086AFE6BFD70" bold="true" box="[189,447,637,663]" pageId="25" pageNumber="64">Locality and ecology</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BF736EAFFD0697CFF4F08BBFBFFFCC8" blockId="25.[189,1400,684,816]" pageId="25" pageNumber="64">
|
||
The Usholta Cave, formed in the Lower Cretaceous limestones, is characterised by
|
||
<quantity id="4CB09B0FFFD0697CFB6208BBFB3DFD21" box="[1168,1257,684,710]" metricMagnitude="3" metricUnit="m" metricValue="2.2" pageId="25" pageNumber="64" unit="m" value="2200.0">2200 m</quantity>
|
||
of explored channels, with a permanent water flow in the main channel (
|
||
<bibRefCitation id="EFD94B1BFFD0697CFC7D08D8FB4CFD0E" author="Tatashidze Z. K. & Tsikarishvili K. D. & Jishkariani J. M." box="[911,1176,719,745]" pageId="25" pageNumber="64" refId="ref14980" refString="Tatashidze Z. K., Tsikarishvili K. D. & Jishkariani J. M. 2009. The Cadastre of the Karst Caves of Georgia. Petiti Publishing House, Tbilisi. [In Georgian.]" type="book" year="2009">
|
||
Tatashidze
|
||
<emphasis id="B93CEAF8FFD0697CFBE508C7FB87FD0E" box="[1047,1107,719,745]" italics="true" pageId="25" pageNumber="64">et al.</emphasis>
|
||
2009
|
||
</bibRefCitation>
|
||
). This part is poor in speleothems and is mainly characterised by a sandy and gravelly bottom. The deeper, side channels without permanent water flow are rich in speleothems and a clay substrate.
|
||
</paragraph>
|
||
<paragraph id="8BF736EAFFD0697CFF4F0959FC07FBDB" blockId="25.[188,1399,846,1084]" pageId="25" pageNumber="64">
|
||
Although many juveniles (only eight were collected) were observed in the dark and damp parts of the cave in all parts explored, exclusively on or in dead wood, only three living adults were found and all three under the stones in the main channel on the banks of the stream. Interestingly, about a dozen dead adults were also found under the stones in the same main channel (
|
||
<figureCitation id="13732A6FFFD0697CFC3E09A0FBF0FC36" box="[972,1060,951,977]" captionStart="Fig" captionStartId="24.[191,234,1963,1989]" captionTargetBox="[189,1398,1153,1920]" captionTargetId="figure-343@24.[189,1398,1151,1920]" captionTargetPageId="24" captionText="Fig. 16. Dead adult specimen of Acanthophorella valerii Antić sp. nov. under the stone on the bank of the cave stream. Photo by Dragan Antić." figureDoi="http://doi.org/10.5281/zenodo.10149335" httpUri="https://zenodo.org/record/10149335/files/figure.png" pageId="25" pageNumber="64">Fig. 16</figureCitation>
|
||
). This could mean that at the time of our visit (end of July) the life cycle of the adults was coming to an end and that it is possible that for some reason the adults go under the stones before they die, considering that all of them (both live and dead) were found exclusively under the stones near the stream.
|
||
</paragraph>
|
||
<paragraph id="8BF736EAFFD0697CFF4F0E4DFC08FAA0" blockId="25.[188,1400,1114,1351]" pageId="25" pageNumber="64">
|
||
Until now, this cave was very poorly studied in terms of its fauna, as only two common Palaearctic insect species from the families
|
||
<taxonomicName id="4C484D69FFD0697CFDC20E6AFCB3FB70" authority="Latreille, 1809" authorityName="Latreille" authorityYear="1809" box="[560,871,1149,1175]" class="Insecta" family="Formicidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="25" pageNumber="64" phylum="Arthropoda" rank="family">Formicidae Latreille, 1809</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C484D69FFD0697CFC6F0E6AFB1FFB7F" authority="Leach, 1815" authorityName="Leach" authorityYear="1815" box="[925,1227,1149,1176]" class="Insecta" family="Geometridae" kingdom="Animalia" order="Lepidoptera" pageId="25" pageNumber="64" phylum="Arthropoda" rank="family">Geometridae Leach, 1815</taxonomicName>
|
||
were recorded (
|
||
<bibRefCitation id="EFD94B1BFFD0697CFF310EB7FE6BFB5C" author="Barjadze S. & Murvanidze M. & Arabuli T. & Mumladze L. & Pkhakadze V. & Djanashvili R. & Salakaia M." box="[195,447,1184,1211]" pageId="25" pageNumber="64" refId="ref14737" refString="Barjadze S., Murvanidze M., Arabuli T., Mumladze L., Pkhakadze V., Djanashvili R. & Salakaia M. 2015. Annotated List of Invertebrates of the Georgian Karst Caves. Georgian Academic Book, Tbilisi." type="book" year="2015">
|
||
Barjadze
|
||
<emphasis id="B93CEAF8FFD0697CFECA0EB6FEA1FB5D" box="[312,373,1184,1210]" italics="true" pageId="25" pageNumber="64">et al.</emphasis>
|
||
2015
|
||
</bibRefCitation>
|
||
). Besides
|
||
<taxonomicName id="4C484D69FFD0697CFDC90EB7FD7FFB5D" authority="Antić & Šević & Barjadze & Makarov, 2023" authorityName="Antić & Šević & Barjadze & Makarov" authorityYear="2023" box="[571,683,1184,1210]" class="Diplopoda" family="Anthroleucosomatidae" genus="Acanthophorella" kingdom="Animalia" order="Chordeumatida" pageId="25" pageNumber="64" phylum="Arthropoda" rank="species" species="valerii" status="sp. nov.">
|
||
<emphasis id="B93CEAF8FFD0697CFDC90EB7FD7FFB5D" box="[571,683,1184,1210]" italics="true" pageId="25" pageNumber="64">A. valerii</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A20F5783FFD0697CFD400EB6FCC5FB5C" box="[690,785,1185,1211]" pageId="25" pageNumber="64" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, two other troglobionts from the genera
|
||
<taxonomicName id="4C484D69FFD0697CFB090EB7FE40FB3A" authority="Chamberlin, 1930" authorityName="Chamberlin" authorityYear="1930" class="Arachnida" family="Neobisiidae" genus="Neobisium" kingdom="Animalia" order="Pseudoscorpiones" pageId="25" pageNumber="64" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B93CEAF8FFD0697CFB090EB7FAACFB5D" box="[1275,1400,1184,1210]" italics="true" pageId="25" pageNumber="64">Neobisium</emphasis>
|
||
Chamberlin, 1930
|
||
</taxonomicName>
|
||
(
|
||
<taxonomicName id="4C484D69FFD0697CFE540ED4FDA8FB3A" authorityName="de Geer" authorityYear="1778" box="[422,636,1219,1245]" class="Arachnida" kingdom="Animalia" order="Pseudoscorpiones" pageId="25" pageNumber="64" phylum="Arthropoda" rank="order">Pseudoscorpiones</taxonomicName>
|
||
) and
|
||
<taxonomicName id="4C484D69FFD0697CFD300ED4FB0CFB3A" authority="Dolzhanski & Ljovuschkin, 1989" authorityName="Dolzhanski & Ljovuschkin" authorityYear="1989" box="[706,1240,1219,1245]" class="Insecta" family="Carabidae" genus="Inotrechus" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="64" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B93CEAF8FFD0697CFD300ED4FCEAFB3A" box="[706,830,1219,1245]" italics="true" pageId="25" pageNumber="64">Inotrechus</emphasis>
|
||
Dolzhanski & Ljovuschkin, 1989
|
||
</taxonomicName>
|
||
(
|
||
<taxonomicName id="4C484D69FFD0697CFB180ED4FABBFB3A" authorityName="Linnaeus" authorityYear="1758" box="[1258,1391,1219,1245]" class="Insecta" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="64" phylum="Arthropoda" rank="order">Coleoptera</taxonomicName>
|
||
) were found in the cave, as well as the troglophiles
|
||
<taxonomicName id="4C484D69FFD0697CFCF20EF0FB65FAE7" authority="Martynov, 1916" authorityName="Martynov" authorityYear="1916" box="[768,1201,1254,1281]" class="Insecta" family="Limnephilidae" genus="Micropterna" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="25" pageNumber="64" phylum="Arthropoda" rank="species" species="clavata">
|
||
<emphasis id="B93CEAF8FFD0697CFCF20EF0FC3BFAE7" box="[768,1007,1254,1280]" italics="true" pageId="25" pageNumber="64">Micropterna clavata</emphasis>
|
||
Martynov, 1916
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C484D69FFD0697CFB140EF1FDDEFAC3" authority="McLachlan, 1895" authorityName="McLachlan" authorityYear="1895" class="Insecta" family="Limnephilidae" genus="Stenophylax" kingdom="Animalia" order="Trichoptera" pageId="25" pageNumber="64" phylum="Arthropoda" rank="species" species="permistus">
|
||
<emphasis id="B93CEAF8FFD0697CFB140EF1FEFBFAC4" italics="true" pageId="25" pageNumber="64">Stenophylax permistus</emphasis>
|
||
McLachlan, 1895
|
||
</taxonomicName>
|
||
(both
|
||
<taxonomicName id="4C484D69FFD0697CFDA80F1DFD30FAC3" box="[602,740,1290,1316]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="25" pageNumber="64" phylum="Arthropoda" rank="order">Trichoptera</taxonomicName>
|
||
) and a species from the genus
|
||
<taxonomicName id="4C484D69FFD0697CFBAD0F1DFAA3FAC3" authority="Yosii, 1956" authorityName="Yosii" authorityYear="1956" box="[1119,1399,1289,1316]" class="Collembola" family="Tomoceridae" genus="Plutomurus" kingdom="Animalia" order="Entomobryomorpha" pageId="25" pageNumber="64" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B93CEAF8FFD0697CFBAD0F1DFB3CFAC3" box="[1119,1256,1290,1316]" italics="true" pageId="25" pageNumber="64">Plutomurus</emphasis>
|
||
Yosii, 1956
|
||
</taxonomicName>
|
||
(
|
||
<taxonomicName id="4C484D69FFD0697CFF340F3AFE80FAA0" authorityName="Lubbock J" authorityYear="1870" box="[198,340,1325,1351]" class="Collembola" kingdom="Animalia" pageId="25" pageNumber="64" phylum="Arthropoda" rank="class">Collembola</taxonomicName>
|
||
;
|
||
<specimenCount id="9D4EFD63FFD0697CFE920F3AFE25FAA0" box="[352,497,1325,1351]" count="1" pageId="25" pageNumber="64" type="juvenile">one juvenile</specimenCount>
|
||
can be spotted in
|
||
<figureCitation id="13732A6FFFD0697CFD370F3AFCCBFAA0" box="[709,799,1325,1351]" captionStart="Fig" captionStartId="1.[189,232,1857,1883]" captionTargetBox="[189,1398,420,1810]" captionTargetId="figure-57@1.[189,1398,420,1810]" captionTargetPageId="1" captionText="Fig. 1. Troglobiotic members of the genus Acanthophorella Antić & Makarov, 2016 in situ. A–B. Acanthophorella barjadzei Antić & Makarov, 2016, ♂ from Nikortsminda Sakinule Cave. C–D. Acanthophorella barjadzei Antić & Makarov, 2016, ♂ from Muradi Cave. E–F. Acanthophorella devi Antić sp. nov., types, ♂♂. G–H. Acanthophorella valerii Antić sp. nov., holotype, ♂ (NHMW MY10366). Photos by Dragan Antić." figureDoi="http://doi.org/10.5281/zenodo.10149300" httpUri="https://zenodo.org/record/10149300/files/figure.png" pageId="25" pageNumber="64">Fig. 1G</figureCitation>
|
||
) (unpubl. data).
|
||
</paragraph>
|
||
<paragraph id="8BF736EAFFD0697CFF4F0F72FAA3FA98" blockId="25.[189,1399,1381,1408]" box="[189,1399,1381,1408]" pageId="25" pageNumber="64">Based on the ecology, as well as the unpigmented body, this species can be considered as a neotroglobiont.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3526561FFD0697CFF4F0F89FDAFF9ED" pageId="25" pageNumber="64" type="distribution">
|
||
<paragraph id="8BF736EAFFD0697CFF4F0F89FE82FA5F" blockId="25.[189,342,1438,1464]" box="[189,342,1438,1464]" pageId="25" pageNumber="64">
|
||
<heading id="D0BF8186FFD0697CFF4F0F89FE82FA5F" bold="true" box="[189,342,1438,1464]" fontSize="11" level="3" pageId="25" pageNumber="64" reason="3">
|
||
<emphasis id="B93CEAF8FFD0697CFF4F0F89FE82FA5F" bold="true" box="[189,342,1438,1464]" pageId="25" pageNumber="64">Distribution</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BF736EAFFD0697CFF4F0FDAFDAFF9ED" blockId="25.[189,1399,1484,1546]" pageId="25" pageNumber="64">
|
||
A Georgian endemic known only from its
|
||
<typeStatus id="54F38848FFD0697CFD250FDAFCDEFA00" box="[727,778,1485,1511]" pageId="25" pageNumber="64">type</typeStatus>
|
||
locality, Usholta Cave, near Usholta village (Oni Municipality) (
|
||
<figureCitation id="13732A6FFFD0697CFE9C0FE7FE11F9ED" box="[366,453,1520,1546]" captionStart="Fig" captionStartId="35.[189,232,1998,2024]" captionTargetBox="[189,1398,1153,1959]" captionTargetId="figure-355@35.[189,1398,1150,1959]" captionTargetPageId="35" captionText="Fig. 22. Distribution of the genus Acanthophorella Antić & Makarov, 2016." figureDoi="http://doi.org/10.5281/zenodo.10149347" httpUri="https://zenodo.org/record/10149347/files/figure.png" pageId="25" pageNumber="64">Fig. 22</figureCitation>
|
||
, orange circle).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |