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<document id="D9746D77BE4BD1398A62FD95AD90D80B" ID-CLB-Dataset="22543" ID-DOI="10.11646/zootaxa.4657.2.12" ID-GBIF-Dataset="a4391bbb-b0ae-4ea4-91b1-c3d8660ca183" ID-ISSN="1175-5326" ID-Zenodo-Dep="3772293" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1588057189965" checkinUser="plazi" docAuthor="Wang, Lu-Yu, Zhang, Zhi-Sheng &amp; Peng, Xian-Jin" docDate="2019" docId="03AE7812FFC8A1560A8EF92E8A5F96DB" docLanguage="en" docName="zootaxa.4657.2.12.pdf" docOrigin="Zootaxa 4657 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Artoria weiweii Wang &amp; Zhang &amp; Peng 2019, sp. nov." docType="treatment" docVersion="6" lastPageNumber="393" masterDocId="FF97006AFFC8A1570A19FFB28E69957D" masterDocTitle="First record of Artoria Thorell, 1877 (Araneae: Lycosidae) from Malaysia, with the description of a new species" masterLastPageNumber="396" masterPageNumber="392" pageNumber="392" updateTime="1698837611729" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="B638D97EDAD4C088060FABFDDD4F5D7E">First record of Artoria Thorell, 1877 (Araneae: Lycosidae) from Malaysia, with the description of a new species</mods:title>
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<mods:namePart id="32834D640A3C140E23A6F96CA70AA180">Wang, Lu-Yu</mods:namePart>
<mods:affiliation id="EA98206FADE9442226A8949D3809DC15">Key Laboratory of Eco-environments in Three Gorges Reservoir Region (Ministry of Education), School of Life Sciences, Southwest University, Chongqing 400715, China.</mods:affiliation>
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<mods:namePart id="01D2664C4D0B9F531C8C68A74C73C2CB">Zhang, Zhi-Sheng</mods:namePart>
<mods:affiliation id="4104C32B3E687B83AC231CE6D69CCBC7">Key Laboratory of Eco-environments in Three Gorges Reservoir Region (Ministry of Education), School of Life Sciences, Southwest University, Chongqing 400715, China.</mods:affiliation>
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<mods:namePart id="C6C3B71DDAD193A8FD609D54F3C57420">Peng, Xian-Jin</mods:namePart>
<mods:affiliation id="E6329C6BDA708F5071CAD9ADCA0C66C6">College of Life Sciences, Hunan Normal University, Changsha 410081, Hunan, China</mods:affiliation>
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<treatment id="03AE7812FFC8A1560A8EF92E8A5F96DB" ID-DOI="http://doi.org/10.5281/zenodo.3798238" ID-GBIF-Taxon="163829492" ID-Zenodo-Dep="3798238" LSID="urn:lsid:plazi:treatment:03AE7812FFC8A1560A8EF92E8A5F96DB" httpUri="http://treatment.plazi.org/id/03AE7812FFC8A1560A8EF92E8A5F96DB" lastPageId="1" lastPageNumber="393" pageId="0" pageNumber="392">
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<paragraph id="8BB8C904FFC8A1570A8EF92E8C3C93CB" blockId="0.[151,1437,1549,2042]" box="[151,597,1692,1719]" pageId="0" pageNumber="392">
<emphasis id="B9731516FFC8A1570A8EF92E8C3C93CB" bold="true" box="[151,597,1692,1719]" pageId="0" pageNumber="392">
<taxonomicName id="4C07B287FFC8A1570A8EF92E8F3893CB" ID-CoL="95FCV" authority="Wang &amp; Zhang &amp; Peng, 2019" authorityName="Wang &amp; Zhang &amp; Peng" authorityYear="2019" box="[151,337,1692,1718]" class="Arachnida" family="Lycosidae" genus="Artoria" kingdom="Animalia" order="Araneae" pageId="0" pageNumber="392" phylum="Arthropoda" rank="species" species="weiweii" status="sp. nov.">
<emphasis id="B9731516FFC8A1570A8EF92E8F3893CB" bold="true" box="[151,337,1692,1718]" italics="true" pageId="0" pageNumber="392">Artoria weiweii</emphasis>
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<taxonomicNameLabel id="A240A86DFFC8A1570B4EF92F8FDC93CA" box="[343,437,1693,1719]" pageId="0" pageNumber="392" rank="species">sp. nov.</taxonomicNameLabel>
(
<figureCitation id="133CD581FFC8A1570BDCF92E8C4693CB" box="[453,559,1692,1718]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStartId-0="1.[151,244,1983,2005]" captionStartId-1="2.[151,244,1949,1971]" captionTargetBox-0="[160,1431,1128,1864]" captionTargetBox-1="[173,1415,483,1924]" captionTargetId-0="figure@1.[151,1436,1114,1878]" captionTargetId-1="figure@2.[173,1415,483,1924]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionText-0="FIGURE 1. Artoria weiweii sp. nov., holotype male (AB) and paratype female (CD). A. Left male pedipalp, ventral view; B. Same, retrolateral view; C. Epigyne, ventral view; D. Female internal genitalia, dorsal view." captionText-1="FIGURE 2. Artoria weiweii sp. nov., holotype male (A, CE) and paratype female (B, FG).A. Male habitus, dorsal view; B. Female habitus, dorsal view; C. Left male pedipalp, prolateral view; D. Same, ventral view; E. Same, retrolateral view; F. Epigyne, ventral view; G. Female internal genitalia, dorsal view." figureDoi-0="http://doi.org/10.5281/zenodo.3772295" figureDoi-1="http://doi.org/10.5281/zenodo.3772299" httpUri-0="https://zenodo.org/record/3772295/files/figure.png" httpUri-1="https://zenodo.org/record/3772299/files/figure.png" pageId="0" pageNumber="392">Figs 12</figureCitation>
,
<figureCitation id="133CD581FFC8A1570824F92E8C2493CB" box="[573,589,1692,1718]" captionStart="FIGURE 4" captionStartId="4.[151,244,1084,1106]" captionTargetBox="[216,1374,189,1057]" captionTargetId="figure@4.[211,1377,182,1063]" captionTargetPageId="4" captionText="FIGURE 4. Distribution records of Artoria in Malaysia." figureDoi="http://doi.org/10.5281/zenodo.3772303" httpUri="https://zenodo.org/record/3772303/files/figure.png" pageId="0" pageNumber="392">4</figureCitation>
)
</emphasis>
</paragraph>
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<paragraph id="8BB8C904FFC8A1570A8EF9708D6D925F" blockId="0.[151,1437,1549,2042]" pageId="0" pageNumber="392">
<emphasis id="B9731516FFC8A1570A8EF9708FEE93A7" bold="true" box="[151,391,1730,1754]" pageId="0" pageNumber="392">Type material. Male</emphasis>
<materialsCitation id="3B6FC359FFC8A1570B8BF9708B6A9383" ID-GBIF-Occurrence="2609011227" collectingDate="2015-10-16" collectorName="L. Y. Wang &amp; G. Q. Huang &amp; Z. Z. Gao" country="Malaysia" elevation="1583" latitude="5.812317" location="Penampang" longLatPrecision="1" longitude="116.34203" municipality="Miles Jalan Tambunan" pageId="0" pageNumber="392" specimenCount="1" stateProvince="Sabah" typeStatus="holotype">
<emphasis id="B9731516FFC8A1570B8BF9708F9D93A7" bold="true" box="[402,500,1730,1754]" pageId="0" pageNumber="392">
<typeStatus id="54BC77A6FFC8A1570B8BF9708F9D93A7" box="[402,500,1730,1754]" pageId="0" pageNumber="392" type="holotype">holotype</typeStatus>
</emphasis>
(SWUC-T-LY04-01),
<emphasis id="B9731516FFC8A15708E9F9708DE193A7" bold="true" box="[752,904,1730,1754]" pageId="0" pageNumber="392">
<collectingCountry id="F3108994FFC8A15708E9F9708DEA93A7" box="[752,899,1730,1754]" name="Malaysia" pageId="0" pageNumber="392">MALAYSIA</collectingCountry>
:
</emphasis>
<collectingRegion id="49C307E6FFC8A157098AF9708DBE93A7" box="[915,983,1730,1754]" country="Malaysia" name="Sabah" pageId="0" pageNumber="392">Sabah</collectingRegion>
, 33
<collectingMunicipality id="6BDC537EFFC8A1570E15F9708B6393A7" box="[1036,1290,1730,1754]" pageId="0" pageNumber="392">Miles Jalan Tambunan</collectingMunicipality>
,
<location id="8ED89FDFFFC8A1570F00F9708BF193A7" LSID="urn:lsid:plazi:treatment:03AE7812FFC8A1560A8EF92E8A5F96DB:8ED89FDFFFC8A1570F00F9708BF193A7" box="[1305,1432,1730,1754]" country="Malaysia" latitude="5.812317" longLatPrecision="1" longitude="116.34203" municipality="Miles Jalan Tambunan" name="Penampang" pageId="0" pageNumber="392" stateProvince="Sabah">Penampang</location>
,
<geoCoordinate id="EE33AFC3FFC8A1570A8EF9568F439380" box="[151,298,1764,1790]" degrees="05" direction="north" minutes="48.739" orientation="latitude" pageId="0" pageNumber="392" precision="1" value="5.812317">05°48.739 N</geoCoordinate>
,
<geoCoordinate id="EE33AFC3FFC8A1570B2EF9568FBD9380" box="[311,468,1764,1790]" degrees="116" direction="east" minutes="20.522" orientation="longitude" pageId="0" pageNumber="392" precision="1" value="116.34203">116°20.522 E</geoCoordinate>
, elev.
<quantity id="4CFF64E1FFC8A1570803F9548C069380" box="[538,623,1766,1790]" metricMagnitude="3" metricUnit="m" metricValue="1.583" pageId="0" pageNumber="392" unit="m" value="1583.0">
<elevation id="002A2E37FFC8A1570803F9548C069380" box="[538,623,1766,1790]" metricMagnitude="3" metricUnit="m" metricValue="1.583" pageId="0" pageNumber="392" unit="m" value="1583.0">1583 m</elevation>
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,
<date id="FFB9EFC4FFC8A1570865F9548D5F9383" box="[636,822,1766,1790]" pageId="0" pageNumber="392" value="2015-10-16">
<collectingDate id="EFFD162CFFC8A1570865F9548D5F9383" box="[636,822,1766,1790]" pageId="0" pageNumber="392" value="2015-10-16">16 October 2015</collectingDate>
</date>
,
<collectorName id="26F2ACD2FFC8A157095AF9548DD09383" box="[835,953,1766,1790]" pageId="0" pageNumber="392">L.Y. Wang</collectorName>
,
<collectorName id="26F2ACD2FFC8A15709DFF9548A209383" box="[966,1097,1766,1790]" pageId="0" pageNumber="392">G.Q. Huang</collectorName>
&amp;
<collectorName id="26F2ACD2FFC8A1570E69F9548ABB9383" box="[1136,1234,1766,1790]" pageId="0" pageNumber="392">Z.Z. Gao</collectorName>
leg.
</materialsCitation>
<materialsCitation id="3B6FC359FFC8A1570F12F9548D69925F" ID-GBIF-Occurrence="2609011229" collectingDate="2015-10-16" collectorName="L. Y. Wang &amp; G. Q. Huang &amp; Z. Z. Gao" country="Malaysia" elevation="1583" latitude="5.812317" location="Penampang" longLatPrecision="1" longitude="116.34203" municipality="Miles Jalan Tambunan" pageId="0" pageNumber="392" specimenCount="5" specimenCount-female="5" stateProvince="Sabah" typeStatus="paratype">
<emphasis id="B9731516FFC8A1570F12F9548BEF9383" bold="true" box="[1291,1414,1766,1790]" pageId="0" pageNumber="392">
<typeStatus id="54BC77A6FFC8A1570F12F9548BE89383" box="[1291,1409,1766,1790]" pageId="0" pageNumber="392" type="paratype">Paratypes</typeStatus>
:
</emphasis>
<specimenCount id="9D01028DFFC8A1570F96F9548E83925F" pageId="0" pageNumber="392" type="female">5 females</specimenCount>
(SWUC-T-LY04-02 ~ 06), same data as holotype
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C31D9A8FFFC8A1570ADEF89C8C6E9217" pageId="0" pageNumber="392" type="etymology">
<paragraph id="8BB8C904FFC8A1570ADEF89C8C6E9217" blockId="0.[151,1437,1549,2042]" pageId="0" pageNumber="392">
<emphasis id="B9731516FFC8A1570ADEF89C8F21923B" bold="true" box="[199,328,1838,1862]" pageId="0" pageNumber="392">Etymology.</emphasis>
The specific epithet honors Mr. Weiwei Zhang, a Chinese entomologist and photographer, who was of tremendous assistance in the field.
</paragraph>
</subSubSection>
<subSubSection id="C31D9A8FFFC8A1570ADEF8C48B129287" pageId="0" pageNumber="392" type="diagnosis">
<paragraph id="8BB8C904FFC8A1570ADEF8C48B129287" blockId="0.[151,1437,1549,2042]" pageId="0" pageNumber="392">
<emphasis id="B9731516FFC8A1570ADEF8C48F5592F3" bold="true" box="[199,316,1910,1934]" pageId="0" pageNumber="392">Diagnosis.</emphasis>
The male of this species can be easily distinguished from other
<taxonomicName id="4C07B287FFC8A15709F1F8C48A5E92F0" authorityName="Thorell" authorityYear="1877" box="[1000,1079,1910,1933]" class="Arachnida" family="Lycosidae" genus="Artoria" kingdom="Animalia" order="Araneae" pageId="0" pageNumber="392" phylum="Arthropoda" rank="genus">
<emphasis id="B9731516FFC8A15709F1F8C48A5E92F0" box="[1000,1079,1910,1933]" italics="true" pageId="0" pageNumber="392">Artoria</emphasis>
</taxonomicName>
species by the long and bifurcate median apophysis (
<figureCitation id="133CD581FFC8A1570B7BF8288FC992CF" box="[354,416,1946,1970]" captionStart="FIGURE 1" captionStartId="1.[151,244,1983,2005]" captionTargetBox="[160,1431,1128,1864]" captionTargetId="figure@1.[151,1436,1114,1878]" captionTargetPageId="1" captionText="FIGURE 1. Artoria weiweii sp. nov., holotype male (AB) and paratype female (CD). A. Left male pedipalp, ventral view; B. Same, retrolateral view; C. Epigyne, ventral view; D. Female internal genitalia, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3772295" httpUri="https://zenodo.org/record/3772295/files/figure.png" pageId="0" pageNumber="392">Figs 1</figureCitation>
AB, 2CE). The female of this species is similar to
<taxonomicName id="4C07B287FFC8A15709D3F8288A8B92CF" authority="(Simon, 1909)" baseAuthorityName="Simon" baseAuthorityYear="1909" box="[970,1250,1946,1970]" class="Arachnida" family="Lycosidae" genus="Artoria" kingdom="Animalia" order="Araneae" pageId="0" pageNumber="392" phylum="Arthropoda" rank="species" species="impedita">
<emphasis id="B9731516FFC8A15709D3F8288A2A92CF" box="[970,1091,1946,1970]" italics="true" pageId="0" pageNumber="392">A. impedita</emphasis>
(
<bibRefCitation id="EF96B4F5FFC8A1570E49F8288AB392CF" author="Simon, E." box="[1104,1242,1946,1970]" pageId="0" pageNumber="392" pagination="152 - 212" refId="ref2303" refString="Simon, E. (1909) Araneae. 2 e partie. In: Michaelsen, W. &amp; Hartmeyer, S. - A. F. (Eds.), Die Fauna Sudwest-Australiens. Jena 2 (13), 152 - 212." type="journal article" year="1909">Simon, 1909</bibRefCitation>
)
</taxonomicName>
(
<bibRefCitation id="EF96B4F5FFC8A1570EF7F8288EA892AB" author="Framenau, V. W." pageId="0" pageNumber="392" pagination="265 - 292" refId="ref2013" refString="Framenau, V. W. (2005) The wolf spider genus Artoria Thorell in Australia: new synonymies and generic transfers (Araneae: Lycosidae). Records of the Western Australian Museum, 22, 265 - 292. https: // doi. org / 10.18195 / issn. 0312 - 3162.22 (4). 2005.265 - 292" type="journal article" year="2005">Framenau 2005: 283</bibRefCitation>
, figs 9DF) from
<collectingRegion id="49C307E6FFC8A1570B91F80C8C2292AB" box="[392,587,1982,2006]" country="Australia" name="Western Australia" pageId="0" pageNumber="392">Western Australia</collectingRegion>
, in having partly elevated median apophysis, but differs from the latter by the small epigynal atrium, the large spermathecae and the narrow distance between copulatory ducts (
<figureCitation id="133CD581FFC8A1570EB2F8508A839287" box="[1195,1258,2018,2042]" captionStart="FIGURE 1" captionStartId="1.[151,244,1983,2005]" captionTargetBox="[160,1431,1128,1864]" captionTargetId="figure@1.[151,1436,1114,1878]" captionTargetPageId="1" captionText="FIGURE 1. Artoria weiweii sp. nov., holotype male (AB) and paratype female (CD). A. Left male pedipalp, ventral view; B. Same, retrolateral view; C. Epigyne, ventral view; D. Female internal genitalia, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3772295" httpUri="https://zenodo.org/record/3772295/files/figure.png" pageId="0" pageNumber="392">Figs 1</figureCitation>
CD, 2FG).
</paragraph>
</subSubSection>
<subSubSection id="C31D9A8FFFC9A1560ADEFF2B8C8B96FF" pageId="1" pageNumber="393" type="description">
<paragraph id="8BB8C904FFC9A1560ADEFF2B8AB894AF" blockId="1.[151,1437,153,934]" pageId="1" pageNumber="393">
<emphasis id="B9731516FFC9A1560ADEFF2B8F3B95CC" bold="true" box="[199,338,153,177]" pageId="1" pageNumber="393">Description.</emphasis>
Male
<typeStatus id="54BC77A6FFC9A1560B8FFF288F9A95CF" box="[406,499,154,178]" pageId="1" pageNumber="393" type="holotype">holotype</typeStatus>
(
<figureCitation id="133CD581FFC9A156081BFF288C3F95CF" box="[514,598,154,178]" captionStart="FIGURE 2" captionStartId="2.[151,244,1949,1971]" captionTargetBox="[173,1415,483,1924]" captionTargetId="figure@2.[173,1415,483,1924]" captionTargetPageId="2" captionText="FIGURE 2. Artoria weiweii sp. nov., holotype male (A, CE) and paratype female (B, FG).A. Male habitus, dorsal view; B. Female habitus, dorsal view; C. Left male pedipalp, prolateral view; D. Same, ventral view; E. Same, retrolateral view; F. Epigyne, ventral view; G. Female internal genitalia, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3772299" httpUri="https://zenodo.org/record/3772299/files/figure.png" pageId="1" pageNumber="393">Fig. 2A</figureCitation>
) total length 3.23. Prosoma 1.84 long, 1.21 wide; opisthosoma 1.39 long, 0.97 wide. Carapace grey-brown, medially lighter. Fovea longitudinal. Cervical groove indistinct. Radial furrows distinct. Eye sizes and interdistances: AME 0.07, ALE 0.06, PME 0.20, PLE 0.15; AMEAME 0.07, AMEALE 0.05, PMEPME 0.18, PMEPLE 0.19. Clypeus height 0.06. Chelicerae brown, with three teeth on each the pro- and retromargin. Labium yellow-brown, with dark base, longer than wide. Endites yellow-brown, longer than wide. Sternum yellow-brown and scutellate, with sparse brown setae. Legs yellow-brown, with brown pigmentation. Leg measurements: I 4.55 (1.22, 1.59, 1.04, 0.70); II 4.05 (1.10, 1.39, 0.93, 0.63); III 3.52 (0.97, 1.11, 0.94, 0.50); IV 5.30 (1.39, 1.64, 1.51, 0.76). Leg formula: 4123. Opisthosoma oval, brown, with an indistinct cardiac mark anteriorly and reddish longitudinal grain posteriorly; with some white setae near anal tubercle. Venter yellow-brown, with small, yellow-brown spinnerets.
</paragraph>
<paragraph id="8BB8C904FFC9A1560ADEFE6C8FD39743" blockId="1.[151,1437,153,934]" pageId="1" pageNumber="393">
Male pedipalp (
<figureCitation id="133CD581FFC9A1560B6AFE6C8FDD948B" box="[371,436,478,502]" captionStart="FIGURE 1" captionStartId="1.[151,244,1983,2005]" captionTargetBox="[160,1431,1128,1864]" captionTargetId="figure@1.[151,1436,1114,1878]" captionTargetPageId="1" captionText="FIGURE 1. Artoria weiweii sp. nov., holotype male (AB) and paratype female (CD). A. Left male pedipalp, ventral view; B. Same, retrolateral view; C. Epigyne, ventral view; D. Female internal genitalia, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3772295" httpUri="https://zenodo.org/record/3772295/files/figure.png" pageId="1" pageNumber="393">Figs 1</figureCitation>
AB, 2CE). Median apophysis two-lobed with thin prolateral and broad retrolateral lobes. Embolus largely hidden beneath the tegular margin and the strong, long and slightly rotated tegular apophysis. The basoembolic apophysis is large.
</paragraph>
<paragraph id="8BB8C904FFC9A1560ADEFDF88EBD978F" blockId="1.[151,1437,153,934]" pageId="1" pageNumber="393">
Female (SWUC-T-LY04-02,
<figureCitation id="133CD581FFC9A1560BE7FDF88C26971F" box="[510,591,586,610]" captionStart="FIGURE 2" captionStartId="2.[151,244,1949,1971]" captionTargetBox="[173,1415,483,1924]" captionTargetId="figure@2.[173,1415,483,1924]" captionTargetPageId="2" captionText="FIGURE 2. Artoria weiweii sp. nov., holotype male (A, CE) and paratype female (B, FG).A. Male habitus, dorsal view; B. Female habitus, dorsal view; C. Left male pedipalp, prolateral view; D. Same, ventral view; E. Same, retrolateral view; F. Epigyne, ventral view; G. Female internal genitalia, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3772299" httpUri="https://zenodo.org/record/3772299/files/figure.png" pageId="1" pageNumber="393">Fig. 2B</figureCitation>
) total length: 3.54. Prosoma 1.75 long, 1.13 wide; opisthosoma 1.74 long, 1.24 wide. Coloration as in the male. Eye sizes and interdistances: AME 0.08, ALE 0.06, PME 0.21, PLE 0.14; AMEAME 0.06, AMEALE 0.05, PMEPME 0.17, PMEPLE 0.19. Clypeus height 0.08. Leg measurements: I 3.91 (1.12, 1.35, 0.86, 0.58); II 3.36 (0.96, 1.15, 0.74, 0.51); III 2.96 (0.86, 0.95, 0.73, 0.42); IV 4.87 (1.39, 1.47, 1.32, 0.69). Leg formula: 4123.
</paragraph>
<paragraph id="8BB8C904FFC9A1560ADEFD4C8C8B96FF" blockId="1.[151,1437,153,934]" pageId="1" pageNumber="393">
Epigyne (
<figureCitation id="133CD581FFC9A1560B36FD4C8F06966B" box="[303,367,766,790]" captionStart="FIGURE 1" captionStartId="1.[151,244,1983,2005]" captionTargetBox="[160,1431,1128,1864]" captionTargetId="figure@1.[151,1436,1114,1878]" captionTargetPageId="1" captionText="FIGURE 1. Artoria weiweii sp. nov., holotype male (AB) and paratype female (CD). A. Left male pedipalp, ventral view; B. Same, retrolateral view; C. Epigyne, ventral view; D. Female internal genitalia, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3772295" httpUri="https://zenodo.org/record/3772295/files/figure.png" pageId="1" pageNumber="393">Figs 1</figureCitation>
CD, 2FG). Septum somewhat like the tail of a rattlesnake, inflated and with transversal furrows. Copulatory openings small, arch-like, beneath the lateral margin of septum. Stalk of spermatheca posteriorly located, twisty and close to each other. Head of spermatheca anteriorly located, large and ball-like. Fertilization ducts small, connected with the posterior margin of spermathecal head.
</paragraph>
</subSubSection>
<subSubSection id="C31D9A8FFFC9A1560ADEFC3F8A5F96DB" box="[199,1078,909,934]" pageId="1" pageNumber="393" type="distribution">
<paragraph id="8BB8C904FFC9A1560ADEFC3F8A5F96DB" blockId="1.[151,1437,153,934]" box="[199,1078,909,934]" pageId="1" pageNumber="393">
<emphasis id="B9731516FFC9A1560ADEFC3F8F3396D8" bold="true" box="[199,346,909,933]" pageId="1" pageNumber="393">Distribution.</emphasis>
Known only from the
<typeStatus id="54BC77A6FFC9A1560854FC3C8C1296DB" box="[589,635,910,934]" pageId="1" pageNumber="393">type</typeStatus>
locality in
<collectingRegion id="49C307E6FFC9A15608EAFC3C8D5E96DB" box="[755,823,910,934]" country="Malaysia" name="Sabah" pageId="1" pageNumber="393">Sabah</collectingRegion>
, East
<collectingCountry id="F3108994FFC9A156096EFC3C8DB096DB" box="[887,985,910,934]" name="Malaysia" pageId="1" pageNumber="393">Malaysia</collectingCountry>
(
<figureCitation id="133CD581FFC9A15609FEFC3C8A4096DB" box="[999,1065,910,934]" captionStart="FIGURE 4" captionStartId="4.[151,244,1084,1106]" captionTargetBox="[216,1374,189,1057]" captionTargetId="figure@4.[211,1377,182,1063]" captionTargetPageId="4" captionText="FIGURE 4. Distribution records of Artoria in Malaysia." figureDoi="http://doi.org/10.5281/zenodo.3772303" httpUri="https://zenodo.org/record/3772303/files/figure.png" pageId="1" pageNumber="393">Fig. 4</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>