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<mods:title id="B64C818C634B8539DBF6808E6C007012">A review of New World Malaxa (Hemiptera: Fulgoroidea: Delphacidae)</mods:title>
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<treatment id="038ACF05B343FF92FF1D6C0AFA3C8E84" ID-DOI="http://doi.org/10.5281/zenodo.6485102" ID-GBIF-Taxon="150835297" ID-Zenodo-Dep="6485102" LSID="urn:lsid:plazi:treatment:038ACF05B343FF92FF1D6C0AFA3C8E84" httpUri="http://treatment.plazi.org/id/038ACF05B343FF92FF1D6C0AFA3C8E84" lastPageId="3" lastPageNumber="514" pageId="2" pageNumber="513">
<subSubSection id="C3392D98B343FF93FF1D6C0AFBEA8B29" box="[151,1098,1813,1839]" pageId="2" pageNumber="513" type="nomenclature">
<paragraph id="8B9C7E13B343FF93FF1D6C0AFBEA8B29" blockId="2.[151,1098,1813,1839]" box="[151,1098,1813,1839]" pageId="2" pageNumber="513">
<heading id="D0D4C97FB343FF93FF1D6C0AFBEA8B29" bold="true" box="[151,1098,1813,1839]" fontSize="11" level="1" pageId="2" pageNumber="513" reason="1">
<emphasis id="B957A201B343FF93FF1D6C0AFBEA8B29" bold="true" box="[151,1098,1813,1839]" pageId="2" pageNumber="513">
Key to Genera of New World
<taxonomicName id="4C230590B343FF93FD806C0AFD408B29" authorityName="Muir" authorityYear="1915" box="[522,736,1813,1839]" class="Insecta" family="Delphacidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="513" phylum="Arthropoda" rank="tribe" tribe="Tropidocephalini">Tropidocephalini</taxonomicName>
(revised from
<bibRefCitation id="EFB203E2B343FF93FC1C6C0AFB9D8B29" box="[918,1085,1813,1839]" pageId="2" pageNumber="513" refId="ref10790" refString="Bartlett, C. R. (2009) A new genus of New World Tropidocephalini (Hemiptera: Delphacidae: Delphacinae), with the description of two new species. Entomological News, 120 (4), 387 - 396. https: // doi. org / 10.3157 / 021.120.0407" type="journal article">Bartlett 2009</bibRefCitation>
).
</emphasis>
</heading>
</paragraph>
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<subSubSection id="C3392D98B343FF92FF1D6C44FA3C8E84" lastPageId="3" lastPageNumber="514" pageId="2" pageNumber="513" type="key">
<keyStep id="30D766B6B343FF93FF1D6C44FA3B8BCF" pageId="2" pageNumber="513">
<paragraph id="8B9C7E13B343FF93FF1D6C44FA3C8B88" blockId="2.[151,1436,1883,2022]" pageId="2" pageNumber="513">
<keyLead id="30D2DD26B343FF93FF1D6C44FA3C8B88" pageId="2" pageNumber="513">
1 Body strongly dorsoventrally flattened (
<figureCitation id="13186296B343FF93FDEB6C44FD098B77" box="[609,681,1883,1905]" captionStart="FIGURES 3034" captionStartId="12.[151,264,1113,1135]" captionTargetBox="[460,1127,185,1086]" captionTargetId="figure@12.[460,1127,185,1092]" captionTargetPageId="12" captionText="FIGURES 3034. New World Tropidocephalini genera; dorsal view of head and thorax (3032), frontal view (33), lateral view of head and thorax (34); 30, Macrocorupha sp. (probably new species, Colombia); 31, Columbisoga saracura (holotype); 32, Columbiana sp. (probably new species, Colombia); 3334 Procidelphax dejecta (paratype)." httpUri="https://zenodo.org/record/1301494/files/figure.png" pageId="2" pageNumber="513">Fig. 34</figureCitation>
); frons rather square, median carinae of frons forked ventrally near lower margin of eyes (
<figureCitation id="13186296B343FF93FEDD6C67FE3D8B88" box="[343,413,1912,1934]" captionStart="FIGURES 3537" captionStartId="13.[151,264,1796,1818]" captionTargetBox="[270,1314,565,1738]" captionTargetId="figure@13.[260,1327,553,1774]" captionTargetPageId="13" captionText="FIGURES 3537. Forewing, line art, leading margin ventrad; 35, Malaxa acutipennis; 36, Lamaxa occidentalis; 37, Xalama microstyla." httpUri="https://zenodo.org/record/1301496/files/figure.png" pageId="2" pageNumber="513">Fig. 35</figureCitation>
)..................................................................
<taxonomicName id="4C230590B343FF93FB476C66FA3C8B88" ID-CoL="6W9B" authority="Bartlett" authorityName="Bartlett" box="[1229,1436,1912,1934]" class="Insecta" family="Delphacidae" genus="Procidelphax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="513" phylum="Arthropoda" rank="genus">
<emphasis id="B957A201B343FF93FB476C66FAED8B88" box="[1229,1357,1913,1934]" italics="true" pageId="2" pageNumber="513">Procidelphax</emphasis>
Bartlett
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B9C7E13B343FF93FF1D6C89FA3B8BCF" blockId="2.[151,1436,1883,2022]" pageId="2" pageNumber="513">
<keyLead id="30D2DD26B343FF93FF1D6C89FA3B8BCF" pageId="2" pageNumber="513">
- Body not flattened (e.g.,
<figureCitation id="13186296B343FF93FE456C8AFDB58BAD" box="[463,533,1941,1963]" captionStart="FIGURES 1518" captionStartId="8.[151,264,1996,2018]" captionTargetBox="[151,1436,1281,1854]" captionTargetId="figure@8.[151,1436,1271,1974]" captionTargetPageId="8" captionText="FIGURES 1518. Lamaxa gracilis (holotype); 15, habitus, dorsal view; 16, habitus, lateral view; 17 frontal view; 18, labels." httpUri="https://zenodo.org/record/1301488/files/figure.png" pageId="2" pageNumber="513">Fig. 16</figureCitation>
), frons rectangular, median carinae not forked (e.g.,
<figureCitation id="13186296B343FF93FB806C8AFBF08BAD" box="[1034,1104,1941,1963]" captionStart="FIGURES 1518" captionStartId="8.[151,264,1996,2018]" captionTargetBox="[151,1436,1281,1854]" captionTargetId="figure@8.[151,1436,1271,1974]" captionTargetPageId="8" captionText="FIGURES 1518. Lamaxa gracilis (holotype); 15, habitus, dorsal view; 16, habitus, lateral view; 17 frontal view; 18, labels." httpUri="https://zenodo.org/record/1301488/files/figure.png" pageId="2" pageNumber="513">Fig. 17</figureCitation>
), except dorsally near fastigium in some species......................................................................................... 2
</keyLead>
</paragraph>
</keyStep>
<paragraph id="8B9C7E13B343FF92FF1D6CCFFA3B8CCC" blockId="2.[151,1436,1883,2022]" lastBlockId="3.[151,1436,151,642]" lastPageId="3" lastPageNumber="514" pageId="2" pageNumber="513">
<keyStep id="30D766B6B343FF92FF1D6CCFFA3B8CCC" lastPageId="3" lastPageNumber="514" pageId="2" pageNumber="513">
<keyLead id="30D2DD26B343FF92FF1D6CCFFA3B8CCC" lastPageId="3" lastPageNumber="514" pageId="2" pageNumber="513">
2 Vertex much (ca. 1.5x) longer than broad, rounded anteriorly in dorsal view (
<figureCitation id="13186296B343FF93FC496CCFFBAB8BE0" box="[963,1035,2000,2022]" captionStart="FIGURES 3034" captionStartId="12.[151,264,1113,1135]" captionTargetBox="[460,1127,185,1086]" captionTargetId="figure@12.[460,1127,185,1092]" captionTargetPageId="12" captionText="FIGURES 3034. New World Tropidocephalini genera; dorsal view of head and thorax (3032), frontal view (33), lateral view of head and thorax (34); 30, Macrocorupha sp. (probably new species, Colombia); 31, Columbisoga saracura (holotype); 32, Columbiana sp. (probably new species, Colombia); 3334 Procidelphax dejecta (paratype)." httpUri="https://zenodo.org/record/1301494/files/figure.png" pageId="2" pageNumber="513">Fig. 30</figureCitation>
); median carina of vertex unbranched.......................................................................................
<taxonomicName id="4C230590B342FF92FB5C6B88FA3C8CAB" ID-CoL="5JBP" authority="Muir" authorityName="Muir" authorityYear="1926" box="[1238,1436,151,173]" class="Insecta" family="Delphacidae" genus="Macrocorupha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="514" phylum="Arthropoda" rank="genus">
<emphasis id="B957A201B342FF92FB5C6B88FAC58CAA" box="[1238,1381,151,172]" italics="true" pageId="3" pageNumber="514">Macrocorupha</emphasis>
Muir
</taxonomicName>
- Vertex shorter (
<figureCitation id="13186296B342FF92FEFF6BABFE4A8CCC" box="[373,490,180,202]" captionStart="FIGURES 3034" captionStartId="12.[151,264,1113,1135]" captionTargetBox="[460,1127,185,1086]" captionTargetId="figure@12.[460,1127,185,1092]" captionTargetPageId="12" captionText="FIGURES 3034. New World Tropidocephalini genera; dorsal view of head and thorax (3032), frontal view (33), lateral view of head and thorax (34); 30, Macrocorupha sp. (probably new species, Colombia); 31, Columbisoga saracura (holotype); 32, Columbiana sp. (probably new species, Colombia); 3334 Procidelphax dejecta (paratype)." httpUri="https://zenodo.org/record/1301494/files/figure.png" pageId="3" pageNumber="514">Figs. 3132</figureCitation>
), more truncate anteriorly; median carina of vertex variable.............................. 3
</keyLead>
</keyStep>
</paragraph>
<keyStep id="30D766B6B342FF92FF1D6BCEFA3B8D39" pageId="3" pageNumber="514">
<paragraph id="8B9C7E13B342FF92FF1D6BCEFA3C8D03" blockId="3.[151,1436,151,642]" pageId="3" pageNumber="514">
<keyLead id="30D2DD26B342FF92FF1D6BCEFA3C8D03" pageId="3" pageNumber="514">
3 Antennae very long (
<figureCitation id="13186296B342FF92FE3A6BCEFD878CE1" box="[432,551,209,231]" captionStart="FIGURES 714" captionStartId="7.[151,264,1401,1423]" captionTargetBox="[151,1436,481,1377]" captionTargetId="figure@7.[151,1436,481,1380]" captionTargetPageId="7" captionText="FIGURES 714. Lamaxa occidentalis (79, 1314 paratype male, Napo River, Ecuador; 1012 La Selva Biological Station Costa Rica); 7, habitus dorsal view; 8, habitus lateral view; 9, frontal view; 10 habitus dorsal view; 11, habitus lateral view; 12, frontal view; 13, pygofer, caudal view (embedded in balsam); 14, aedeagus, connectives, gonostyli and anal tube, left lateral view (embedded in balsam)." httpUri="https://zenodo.org/record/1301486/files/figure.png" pageId="3" pageNumber="514">Figs. 10, 12</figureCitation>
), exceeding posterior margin of mesothorax; both segments long (much longer than wide) with segment I nearly ½ length segment II........................................................
<taxonomicName id="4C230590B342FF92FA976BF0FACA8D02" authority="Bartlett &amp; Kennedy, 2018" authorityName="Bartlett &amp; Kennedy" authorityYear="2018" box="[1309,1386,239,260]" class="Liliopsida" family="Poaceae" genus="Lamaxa" higherTaxonomySource="GBIF" kingdom="Plantae" order="Poales" pageId="3" pageNumber="516" phylum="Tracheophyta" rank="genus" status="gen. nov.">
<emphasis id="B957A201B342FF92FA976BF0FACA8D02" box="[1309,1386,239,260]" italics="true" pageId="3" pageNumber="514">Lamaxa</emphasis>
</taxonomicName>
<emphasis id="B957A201B342FF92FAFA6BF0FA3C8D03" bold="true" box="[1392,1436,239,261]" pageId="3" pageNumber="514">n. g.</emphasis>
</keyLead>
</paragraph>
<paragraph id="8B9C7E13B342FF92FF1D6A12FA3B8D39" blockId="3.[151,1436,151,642]" pageId="3" pageNumber="514">
<keyLead id="30D2DD26B342FF92FF1D6A12FA3B8D39" pageId="3" pageNumber="514">
- Antennae not as long (
<figureCitation id="13186296B342FF92FE326A13FD888D24" box="[440,552,268,290]" captionStart="FIGURES 1924" captionStartId="10.[151,264,1286,1308]" captionTargetBox="[151,1436,193,1264]" captionTargetId="figure@10.[151,1436,193,1265]" captionTargetPageId="10" captionText="FIGURES 1924. Xalama microstyla (1922 holotype; 2324 Peru); 19, habitus, dorsal view; 20, frontal view; 21, habitus lateral view; 22, labels; 23, habitus, dorsal view; 24, frontal view." httpUri="https://zenodo.org/record/1301490/files/figure.png" pageId="3" pageNumber="514">Figs 1920</figureCitation>
), not exceeding mesothorax; generally segment I somewhat longer than wide and 1/3 or less length of II......................................................................................... 4
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30D766B6B342FF92FF1D6A58FA3B8DD4" pageId="3" pageNumber="514">
<paragraph id="8B9C7E13B342FF92FF1D6A58FA3C8D91" blockId="3.[151,1436,151,642]" pageId="3" pageNumber="514">
<keyLead id="30D2DD26B342FF92FF1D6A58FA3C8D91" pageId="3" pageNumber="514">
4 Lateral carinae of pronotum reaching hind margin (
<figureCitation id="13186296B342FF92FD5A6A58FCB88D5B" box="[720,792,327,349]" captionStart="FIGURES 3034" captionStartId="12.[151,264,1113,1135]" captionTargetBox="[460,1127,185,1086]" captionTargetId="figure@12.[460,1127,185,1092]" captionTargetPageId="12" captionText="FIGURES 3034. New World Tropidocephalini genera; dorsal view of head and thorax (3032), frontal view (33), lateral view of head and thorax (34); 30, Macrocorupha sp. (probably new species, Colombia); 31, Columbisoga saracura (holotype); 32, Columbiana sp. (probably new species, Colombia); 3334 Procidelphax dejecta (paratype)." httpUri="https://zenodo.org/record/1301494/files/figure.png" pageId="3" pageNumber="514">Fig. 31</figureCitation>
); the Y-carina of vertex not distinct, sometimes forming a rounded areolet at apex of the median carina; anal tube with or without processes (a heterogeneous taxon, often strongly marked on body and wings).......................................................................
<taxonomicName id="4C230590B342FF92FB6C6A9DFA3C8D91" ID-CoL="3SNB" authority="Muir" authorityName="Muir. Bartlett &amp; Kunz" authorityYear="2015" box="[1254,1436,385,407]" class="Insecta" family="Delphacidae" genus="Columbisoga" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="514" phylum="Arthropoda" rank="genus">
<emphasis id="B957A201B342FF92FB6C6A9DFAC68D91" box="[1254,1382,386,407]" italics="true" pageId="3" pageNumber="514">Columbisoga</emphasis>
Muir
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B9C7E13B342FF92FF1D6A80FA3B8DD4" blockId="3.[151,1436,151,642]" pageId="3" pageNumber="514">
<keyLead id="30D2DD26B342FF92FF1D6A80FA3B8DD4" pageId="3" pageNumber="514">
- Lateral carinae of pronotum not attaining hind margin (
<figureCitation id="13186296B342FF92FD736A80FCE28DB3" box="[761,834,415,437]" captionStart="FIGURES 3034" captionStartId="12.[151,264,1113,1135]" captionTargetBox="[460,1127,185,1086]" captionTargetId="figure@12.[460,1127,185,1092]" captionTargetPageId="12" captionText="FIGURES 3034. New World Tropidocephalini genera; dorsal view of head and thorax (3032), frontal view (33), lateral view of head and thorax (34); 30, Macrocorupha sp. (probably new species, Colombia); 31, Columbisoga saracura (holotype); 32, Columbiana sp. (probably new species, Colombia); 3334 Procidelphax dejecta (paratype)." httpUri="https://zenodo.org/record/1301494/files/figure.png" pageId="3" pageNumber="514">Fig. 32</figureCitation>
); the Y-carinae of vertex distinct, rounded areolet not present; anal tube without processes............................................................................. 5
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30D766B6B342FF92FF1D6AC6FA3C8E84" pageId="3" pageNumber="514">
<paragraph id="8B9C7E13B342FF92FF1D6AC6FA3C8E2C" blockId="3.[151,1436,151,642]" pageId="3" pageNumber="514">
<keyLead id="30D2DD26B342FF92FF1D6AC6FA3C8E2C" pageId="3" pageNumber="514">
5 Wings clear and unmarked (except fuscous at wing base in
<taxonomicName id="4C230590B342FF92FC926AC5FCCE8DE9" box="[792,878,474,495]" class="Insecta" family="Delphacidae" genus="Columbisoga" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="514" phylum="Arthropoda" rank="species" species="caresi">
<emphasis id="B957A201B342FF92FC926AC5FCCE8DE9" box="[792,878,474,495]" italics="true" pageId="3" pageNumber="514">C. caresi</emphasis>
</taxonomicName>
); head and body uniformly colored (
<figureCitation id="13186296B342FF92FB476AC6FAB58DE9" box="[1229,1301,473,495]" captionStart="FIGURES 3034" captionStartId="12.[151,264,1113,1135]" captionTargetBox="[460,1127,185,1086]" captionTargetId="figure@12.[460,1127,185,1092]" captionTargetPageId="12" captionText="FIGURES 3034. New World Tropidocephalini genera; dorsal view of head and thorax (3032), frontal view (33), lateral view of head and thorax (34); 30, Macrocorupha sp. (probably new species, Colombia); 31, Columbisoga saracura (holotype); 32, Columbiana sp. (probably new species, Colombia); 3334 Procidelphax dejecta (paratype)." httpUri="https://zenodo.org/record/1301494/files/figure.png" pageId="3" pageNumber="514">Fig. 32</figureCitation>
); ventral margin of pygofer with broad forked process (also with a pair of lateral projections in the type species
<taxonomicName id="4C230590B342FF92FB206AE8FB5E8E0A" box="[1194,1278,503,524]" class="Insecta" family="Delphacidae" genus="Columbisoga" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="514" phylum="Arthropoda" rank="species" species="lloydi">
<emphasis id="B957A201B342FF92FB206AE8FB5E8E0A" box="[1194,1278,503,524]" italics="true" pageId="3" pageNumber="514">C. lloydi</emphasis>
</taxonomicName>
); aedeagus caudally directed or somewhat twisted, not strongly curved ventrad..................................
<taxonomicName id="4C230590B342FF92FB64690AFA3C8E2C" ID-CoL="3SMX" authority="Muir" authorityName="Muir" authorityYear="1919" box="[1262,1436,532,554]" class="Insecta" family="Delphacidae" genus="Columbiana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="514" phylum="Arthropoda" rank="genus">
<emphasis id="B957A201B342FF92FB64690AFAC48E2C" box="[1262,1380,533,554]" italics="true" pageId="3" pageNumber="514">Columbiana</emphasis>
Muir
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B9C7E13B342FF92FF1D692DFA3C8E84" blockId="3.[151,1436,151,642]" pageId="3" pageNumber="514">
<keyLead id="30D2DD26B342FF92FF1D692DFA3C8E84" pageId="3" pageNumber="514">
- Wings clear, strongly marked with fuscous (
<figureCitation id="13186296B342FF92FD0F692EFD6B8E41" box="[645,715,561,583]" captionStart="FIGURES 1924" captionStartId="10.[151,264,1286,1308]" captionTargetBox="[151,1436,193,1264]" captionTargetId="figure@10.[151,1436,193,1265]" captionTargetPageId="10" captionText="FIGURES 1924. Xalama microstyla (1922 holotype; 2324 Peru); 19, habitus, dorsal view; 20, frontal view; 21, habitus lateral view; 22, labels; 23, habitus, dorsal view; 24, frontal view." httpUri="https://zenodo.org/record/1301490/files/figure.png" pageId="3" pageNumber="514">Fig. 21</figureCitation>
); body dark, marked with pale (especially on head;
<figureCitation id="13186296B342FF92FB32692EFB5F8E41" box="[1208,1279,561,583]" captionStart="FIGURES 1924" captionStartId="10.[151,264,1286,1308]" captionTargetBox="[151,1436,193,1264]" captionTargetId="figure@10.[151,1436,193,1265]" captionTargetPageId="10" captionText="FIGURES 1924. Xalama microstyla (1922 holotype; 2324 Peru); 19, habitus, dorsal view; 20, frontal view; 21, habitus lateral view; 22, labels; 23, habitus, dorsal view; 24, frontal view." httpUri="https://zenodo.org/record/1301490/files/figure.png" pageId="3" pageNumber="514">Fig. 23</figureCitation>
); ventral margin of pygofer with broad scoop-like projection (not apically forked;
<figureCitation id="13186296B342FF92FCD26950FC3E8E63" box="[856,926,591,613]" captionStart="FIGURES 2529" captionStartId="11.[151,264,1882,1904]" captionTargetBox="[226,1350,805,1854]" captionTargetId="figure@11.[226,1360,805,1861]" captionTargetPageId="11" captionText="FIGURES 2529. Male genitalia of Lamaxa occidentalis (2526) and Xalama microstyla (2729); 25, pygofer, semiventral view (note shape of process on ventral margin of pygofer opening); 26, pygofer, left lateral view; 27, pygofer, left lateral view; 28 pygofer, near caudal view; 29, pygofer, ventral view (note shape of process on ventral margin of pygofer opening)." httpUri="https://zenodo.org/record/1301492/files/figure.png" pageId="3" pageNumber="514">Fig. 27</figureCitation>
) and a pair of lateral teeth; aedeagus strongly downcurved.....................................................................................
<taxonomicName id="4C230590B342FF92FA966972FACA8E84" authority="Bartlett &amp; Kennedy, 2018" authorityName="Bartlett &amp; Kennedy" authorityYear="2018" box="[1308,1386,621,642]" class="Liliopsida" family="Poaceae" genus="Xalama" higherTaxonomySource="GBIF" kingdom="Plantae" order="Poales" pageId="3" pageNumber="520" phylum="Tracheophyta" rank="genus" status="gen. nov.">
<emphasis id="B957A201B342FF92FA966972FACA8E84" box="[1308,1386,621,642]" italics="true" pageId="3" pageNumber="514">Xalama</emphasis>
</taxonomicName>
<emphasis id="B957A201B342FF92FAFA6973FA3C8E84" bold="true" box="[1392,1436,620,642]" pageId="3" pageNumber="514">n. g.</emphasis>
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
</treatment>
</document>