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<document id="A9D26198D658C3898D39CFAB1D410D5E" ID-DOI="10.5252/geodiversitas2024v46a6" ID-ISSN="1638-9395" ID-Zenodo-Dep="11106598" ID-ZooBank="urn:lsid:zoobank.org:pub:6ACF6A79-9149-4781-808D-478668673EB6" IM.bibliography_approvedBy="juliana" IM.illustrations_approvedBy="juliana" IM.materialsCitations_approvedBy="juliana" IM.metadata_approvedBy="juliana" IM.tables_approvedBy="juliana" IM.taxonomicNames_approvedBy="juliana" IM.treatments_approvedBy="juliana" checkinTime="1714640714399" checkinUser="plazi" docAuthor="Scavezzoni, Isaure, Fischer, Valentin, Johnson, Michela M. &amp; Jouve, Stéphane" docDate="2024" docId="038A56761D11FFDAFF269551FBD25005" docLanguage="en" docName="geodiversitas.2024.46.6.pdf" docOrigin="Geodiversitas 46 (6)" docSource="https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/g2024v46a6.pdf" docStyle="DocumentStyle:F830B10FF475E64C1F1601E3B32DDC00.4:Geodiversitas.2018-.journal_article" docStyleId="F830B10FF475E64C1F1601E3B32DDC00" docStyleName="Geodiversitas.2018-.journal_article" docStyleVersion="4" docTitle="Cricosaurus bambergensis Sachs et al. 2019" docType="treatment" docVersion="4" lastPageNumber="178" masterDocId="FFB32E0E1D3CFFF7FFA29243FFC5576E" masterDocTitle="Form and function of the pelvic girdle of Thalattosuchia and Dyrosauridae (Crocodyliformes)" masterLastPageNumber="326" masterPageNumber="135" pageNumber="178" updateTime="1714680973700" updateUser="ExternalLinkService" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0">
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<mods:title id="18AB514AB12F77ECBB4C44335C744328">Form and function of the pelvic girdle of Thalattosuchia and Dyrosauridae (Crocodyliformes)</mods:title>
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<mods:namePart id="CB4231880028B53AFCD76B6E214BDBE0">Scavezzoni, Isaure</mods:namePart>
<mods:affiliation id="9223683669FFA085A9D0A19F1B1C6D4C">Universite de Liège, Evolution and Diversity Dynamics Lab, All. du Six Août 14, 4000 Liège (Belgique) isaure. scavezzoni @ gmail. com v. fischer @ uliege. be</mods:affiliation>
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<mods:namePart id="D1E1B18F0BB89C76978F940EEE080D9D">Fischer, Valentin</mods:namePart>
<mods:affiliation id="33613522C7E03942858E5B56B519316C">Universite de Liège, Evolution and Diversity Dynamics Lab, All. du Six Août 14, 4000 Liège (Belgique) isaure. scavezzoni @ gmail. com v. fischer @ uliege. be</mods:affiliation>
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<mods:namePart id="72B4C81159D3D45FF6AC26DF1E6D638C">Johnson, Michela M.</mods:namePart>
<mods:affiliation id="7B99512B4C7DBAE25A3663C7BB3703D1">Department of Palaeontology, Staatliches Museum für Naturkunde Stuttgart, Museum am LÖwentor, Rosenstein 1, 70191 Stuttgart (Germany) michela. johnson @ smns-bw. de</mods:affiliation>
<mods:nameIdentifier id="3ED0352B76475CA371969AD1BC375CFF" type="email">michela.johnson@smns-bw.de</mods:nameIdentifier>
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<mods:namePart id="0C4989A5B7DC851AA7CAC29403F60D6D">Jouve, Stéphane</mods:namePart>
<mods:affiliation id="71DF0B6B2721EA49178838518E72CA92">Sorbonne Universite, BUPMC - Pôle Collections, Tour Zamansky, 15 étage, bureau 1513, 4 place Jussieu, 75252 Paris Cedex 05 (France) stephane. jouve @ sorbonne-universite. fr</mods:affiliation>
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<mods:title id="27ED397BD73795FE8FCFB62B2C636C52">Geodiversitas</mods:title>
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<mods:date id="8F87D279630FB59CD97A34D062AAA681">2024</mods:date>
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<treatment id="038A56761D11FFDAFF269551FBD25005" ID-DOI="http://doi.org/10.5281/zenodo.11105970" ID-Zenodo-Dep="11105970" LSID="urn:lsid:plazi:treatment:038A56761D11FFDAFF269551FBD25005" httpUri="http://treatment.plazi.org/id/038A56761D11FFDAFF269551FBD25005" lastPageNumber="178" pageId="45" pageNumber="178">
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<paragraph id="8B9CE7601D11FFDAFF269551FE815022" blockId="45.[132,706,1810,1900]" pageId="45" pageNumber="178">
<taxonomicName id="4C239CE31D11FFDAFF269551FE815022" authority="(SACHS, YOUNG, ABEL &amp; MALLISON, 2019)" authorityName="Sachs et al." authorityYear="2019" baseAuthorityName="Sachs, Young, Abel &amp; Mallison" baseAuthorityYear="2019" class="Reptilia" family="Metriorhynchidae" genus="Cricosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="45" pageNumber="287" phylum="Chordata" rank="species" species="bambergensis">
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</emphasis>
(
<bibRefCitation id="EFB29A911D11FFDAFE1E9551FEF95022" author="SACHS S. &amp; YOUNG M. T. &amp; ABEL P. &amp; MALLISON H." pageId="45" pageNumber="178" pagination="343 - 356" refId="ref115725" refString="SACHS S., YOUNG M. T., ABEL P. &amp; MALLISON H. 2019. - A new species of the metriorhynchid crocodylomorph Cricosaurus from the Upper Jurassic of southern Germany. Acta Palaeontologica Polonica 64 (2): 343 - 356. https: // doi. org / 10.4202 / app. 00541.2018" type="journal article" year="2019">
<smallCapsWord id="8D7A71BC1D11FFDAFE1E9551FDC75044" baselines="1829,1829" box="[444,514,1810,1836]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Sachs" pageId="45" pageNumber="178">SACHS</smallCapsWord>
,
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,
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&amp;
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, 2019
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)
</taxonomicName>
</paragraph>
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<subSubSection id="C339B4EB1D11FFDAFF269511FBD25005" pageId="45" pageNumber="178" type="description">
<paragraph id="8B9CE7601D11FFDAFF269511FE365002" blockId="45.[132,706,1810,1900]" box="[132,499,1873,1900]" pageId="45" pageNumber="178">
For measurements, see
<tableCitation id="C6A1D2DB1D11FFDAFEDF9512FE295002" box="[381,492,1873,1900]" captionStart-0="TABLE" captionStart-1="TABLE" captionStart-2="TABLE" captionStartId-0="19.[61,72,462,479]" captionStartId-1="25.[61,72,457,474]" captionStartId-2="29.[61,72,511,528]" captionTargetPageId-0="19" captionTargetPageId-1="25" captionTargetPageId-2="29" captionText-0="TABLE 7. — List of measurements in mm of crocodyliformes ilia. Abbreviations: Ha, anterior height; Hv, ventral length; Hd, dorsal length; PST.l, postacetabular process anteroposterior length; PST.h, postacetabular process dorsoventral height; PRE.l, preacetabular process anteroposterior length; PRE.h, preacetabular process dorsoventral height; ACE.h, acetabulum dorsoventral height; ACE.w, acetabulum width at mid-height; GAP.w, base length of acetabular perforation; GAP.h, max height of acetabular perforation; PUB.l, pubic peduncle anteroposterior length; PUB.h, pubic peduncle dorsoventral height; ISC.l, ischial peduncle anteroposterior length; ISC.h, Ischial peduncle dorsoventral height." captionText-1="TABLE 8. — List of measurements in mm of crocodyliformes ischia. When measurements are missing on the first ischium, it takes measurements from next ischium. NHMUK PV R 4763 shaft reconstructed.Abbreviations: Hd, total height from posterior peduncle to tip of posterior process; PP.h, height of posterior process tip; AP.h, base height of anterior process; AP.w, total length of anterior process; BL.l, total length of ischial blade; GAP.w, base length of acetabular perforation; GAP.h, max height of acetabular perforation;PPed.Ll, anteroposterior length of posterior peduncle taken laterally;PPed.Wa, mediolateral width of posterior peduncle taken anteriorly; BR.l, total length of peduncle bridge; APed.w, mediolateral width of anterior peduncle; APed.h, dorsoventral height of anterior peduncle; Arch.l, anterior arch length from anterior peduncle to anterior process; Arch.h, maximal depth between base arch and bone margin; S, length from proximal extremity to maximal constriction; C, anteroposterior width of constriction." captionText-2="TABLE 9. — List of measurements in mm of crocodyliformes pubes. When measurements are missing on the first bone, it takes measurements from the next one. SMNS 81608 has been reconstructed. Abbreviations: H, total proximodistal height; Wh, mediolateral width at mid length; Ldia, length of diaphysis; Lbl, freehand length of distal blade up to diaphysis; Dbl, distance between both medial and lateral extremities of distal blade; Wprox, anteroposterior width of proximal peduncle; Lprox, mediolateral length of proximal peduncle; Shaft, proximodistal length of shaft; Neck, proximodistal length of neck; C, anteroposterior width of constriction; Arch.l, anterior arch length from peduncle to pubic diaphysis; Arch.h, maximal depth between base arch and bone margin; Arch.d, distance between proximal base of arch and position of max depth along the arch." httpUri-0="http://table.plazi.org/id/DF5CB7E81D2FFFE4FF9F938DFDDF554D" httpUri-1="http://table.plazi.org/id/DF5CB7E81D25FFEEFF9F938AFAB1555A" httpUri-2="http://table.plazi.org/id/DF5CB7E81D21FFEAFF9F93BCF8FE553A" pageId="45" pageNumber="178" tableUuid-0="DF5CB7E81D2FFFE4FF9F938DFDDF554D" tableUuid-1="DF5CB7E81D25FFEEFF9F938AFAB1555A" tableUuid-2="DF5CB7E81D21FFEAFF9F93BCF8FE553A">Tables 7-9</tableCitation>
.
</paragraph>
<paragraph id="8B9CE7601D11FFDAFF2695D2FF1050C5" blockId="45.[132,213,1937,1963]" box="[132,213,1937,1963]" pageId="45" pageNumber="178">
<heading id="D0D4500C1D11FFDAFF2695D2FF1050C5" box="[132,213,1937,1963]" fontSize="11" level="2" pageId="45" pageNumber="178" reason="8">
<emphasis id="B9573B721D11FFDAFF2695D2FF1050C5" box="[132,213,1937,1963]" italics="true" pageId="45" pageNumber="178">Ischium</emphasis>
</heading>
</paragraph>
<paragraph id="8B9CE7601D11FFDAFF2695F2FB3A55BE" blockId="45.[132,774,1969,2028]" lastBlockId="45.[812,1456,215,1039]" pageId="45" pageNumber="178">
Like other thalattosuchians, the ischium of
<taxonomicName id="4C239CE31D11FFDAFD2495F1FED75085" authorityName="Sachs et al." authorityYear="2019" baseAuthorityName="Sachs, Young, Abel &amp; Mallison" baseAuthorityYear="2019" class="Reptilia" family="Metriorhynchidae" genus="Cricosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="45" pageNumber="178" phylum="Chordata" rank="species" species="bambergensis">
<emphasis id="B9573B721D11FFDAFD2495F1FED75085" italics="true" pageId="45" pageNumber="178">Cricosaurus bambergensis</emphasis>
</taxonomicName>
(
<figureCitation id="1318FBE51D11FFDAFE849592FEBD5085" box="[294,376,2001,2027]" captionStart="FIG" captionStartId="46.[132,143,1819,1836]" captionTargetBox="[141,1456,181,1794]" captionTargetId="figure-68@46.[141,1447,269,1777]" captionTargetPageId="46" captionText="FIG. 21. — Pelvic and thoracic girdle elements of Cricosaurus bambergensis (Sachs, Young, Abel &amp; Mallison, 2019) (Sachs et al. 2019), NKMB-P-Watt14/274 (holotype): A, overview of pelvic girdle and hindlimb; B, overview of thoracic girdle and forelimb; C, detail of pelvic girdle. Modified from Sachs et al. (2019). Scale bars: 1 cm." figureDoi="http://doi.org/10.5281/zenodo.11106644" httpUri="https://zenodo.org/record/11106644/files/figure.png" pageId="45" pageNumber="178">Fig. 21</figureCitation>
) possesses well-developed anterior and posterior processes (e.g.
<emphasis id="B9573B721D11FFDAFBD89294FA86579F" box="[1146,1347,215,241]" italics="true" pageId="45" pageNumber="178">
<taxonomicName id="4C239CE31D11FFDAFBD89294FAC6579F" box="[1146,1283,215,241]" class="Reptilia" family="Metriorhynchidae" genus="Pelagosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="45" pageNumber="178" phylum="Chordata" rank="genus">Pelagosaurus</taxonomicName>
typus
</emphasis>
, ‘
<taxonomicName id="4C239CE31D11FFDAFAFE929BFACF567C" authority="NHMUK PV R" authorityName=", NHMUK PV R" baseAuthorityName="Eudes-Deslongchamps" baseAuthorityYear="1867" class="Reptilia" family="Metriorhynchidae" genus="Metriorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="45" pageNumber="178" phylum="Chordata" rank="species" species="brachyrhynchus">
<emphasis id="B9573B721D11FFDAFAFE929BFC4B567F" italics="true" pageId="45" pageNumber="178">Metriorhynchus</emphasis>
’
<emphasis id="B9573B721D11FFDAFC3D92B4FB8D567F" box="[927,1096,247,273]" italics="true" pageId="45" pageNumber="178">brachyrhynchus</emphasis>
NHMUK PV R
</taxonomicName>
4763,
<taxonomicName id="4C239CE31D11FFDAFAFC92B4FBDB565F" authority="(Blainville, 1853)" authorityName="(Blainville)" authorityYear="1852" baseAuthorityName="Blainville" baseAuthorityYear="1853" class="Reptilia" family="Metriorhynchidae" genus="Thalattosuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="45" pageNumber="178" phylum="Chordata" rank="species" species="superciliosus">
<emphasis id="B9573B721D11FFDAFAFC92B4FBDB565F" italics="true" pageId="45" pageNumber="178">Thalattosuchus superciliosus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C239CE31D11FFDAFB939354FADD565F" baseAuthorityName="Fraas" baseAuthorityYear="1901" box="[1073,1304,279,305]" class="Reptilia" family="Metriorhynchidae" genus="Cricosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="45" pageNumber="178" phylum="Chordata" rank="species" species="suevicus">
<emphasis id="B9573B721D11FFDAFB939354FADD565F" box="[1073,1304,279,305]" italics="true" pageId="45" pageNumber="178">Cricosaurus suevicus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C239CE31D11FFDAFA899354FC4B563F" authorityName="SMNS" baseAuthorityName="Plieninger" baseAuthorityYear="1846" class="Reptilia" family="Metriorhynchidae" genus="Dakosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="45" pageNumber="178" phylum="Chordata" rank="species" species="maximus">
<emphasis id="B9573B721D11FFDAFA899354FC4B563F" italics="true" pageId="45" pageNumber="178">Dakosaurus maximus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C239CE31D11FFDAFC39937BFB57563F" baseAuthorityName="Wilkinson, Young &amp; Benton" baseAuthorityYear="2008" box="[923,1170,312,337]" class="Reptilia" family="Metriorhynchidae" genus="Torvoneustes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="45" pageNumber="178" phylum="Chordata" rank="species" species="carpenteri">
<emphasis id="B9573B721D11FFDAFC39937BFB57563F" box="[923,1170,312,337]" italics="true" pageId="45" pageNumber="178">Torvoneustes carpenteri</emphasis>
</taxonomicName>
,
<taxonomicName id="4C239CE31D11FFDAFB029374FA6D563F" authority="(Jager, 1828)" authorityName="(Jager)" baseAuthorityName="Jager" baseAuthorityYear="1828" box="[1184,1448,311,337]" class="Reptilia" family="Teleosauridae" genus="Macrospondylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="45" pageNumber="178" phylum="Chordata" rank="species" species="bollensis">
<emphasis id="B9573B721D11FFDAFB029374FA6D563F" box="[1184,1448,311,337]" italics="true" pageId="45" pageNumber="178">Macrospondylus bollensis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C239CE31D11FFDAFC8F9314FBFD561F" authority="(Andrews, 1909)" authorityName="(Andrews)" baseAuthorityName="Andrews" baseAuthorityYear="1909" box="[813,1080,343,369]" class="Reptilia" family="Teleosauridae" genus="Lemmysuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="45" pageNumber="178" phylum="Chordata" rank="species" species="obtusidens">
<emphasis id="B9573B721D11FFDAFC8F9314FBFD561F" box="[813,1080,343,369]" italics="true" pageId="45" pageNumber="178">Lemmysuchus obtusidens</emphasis>
</taxonomicName>
). The anterior process is thinner than the posterior one but only slightly. Moreover, it does not appear sharp unlike for example in ‘
<taxonomicName id="4C239CE31D11FFDAFA8993D5FB0156BF" authority="NHMUK PV R" authorityName=", NHMUK PV R" baseAuthorityName="Eudes-Deslongchamps" baseAuthorityYear="1867" class="Reptilia" family="Metriorhynchidae" genus="Metriorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="45" pageNumber="178" phylum="Chordata" rank="species" species="brachyrhynchus">
<emphasis id="B9573B721D11FFDAFA8993D5FC9E56BE" italics="true" pageId="45" pageNumber="178">Metriorhynchus</emphasis>
’
<emphasis id="B9573B721D11FFDAFCCB93F5FBCB56BE" box="[873,1038,438,464]" italics="true" pageId="45" pageNumber="178">brachyrhynchus</emphasis>
NHMUK PV R
</taxonomicName>
4763,
<taxonomicName id="4C239CE31D11FFDAFAB293F5FB75569F" authority="(Blainville, 1853)" authorityName="(Blainville)" authorityYear="1852" baseAuthorityName="Blainville" baseAuthorityYear="1853" class="Reptilia" family="Metriorhynchidae" genus="Thalattosuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="45" pageNumber="178" phylum="Chordata" rank="species" species="superciliosus">
<emphasis id="B9573B721D11FFDAFAB293F5FC75569E" italics="true" pageId="45" pageNumber="178">Thalattosuchus superciliosus</emphasis>
NHMUK PV R 2054
</taxonomicName>
,
<taxonomicName id="4C239CE31D11FFDAFB1D9395FA62569E" authorityName="SMNS" baseAuthorityName="Plieninger" baseAuthorityYear="1846" box="[1215,1447,470,496]" class="Reptilia" family="Metriorhynchidae" genus="Dakosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="45" pageNumber="178" phylum="Chordata" rank="species" species="maximus">
<emphasis id="B9573B721D11FFDAFB1D9395FA62569E" box="[1215,1447,470,496]" italics="true" pageId="45" pageNumber="178">Dakosaurus maximus</emphasis>
</taxonomicName>
,
<emphasis id="B9573B721D11FFDAFC8F93B5FC3D557E" box="[813,1016,502,528]" italics="true" pageId="45" pageNumber="178">
<taxonomicName id="4C239CE31D11FFDAFC8F93B5FC72557E" box="[813,951,502,528]" class="Reptilia" family="Metriorhynchidae" genus="Pelagosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="45" pageNumber="178" phylum="Chordata" rank="genus">Pelagosaurus</taxonomicName>
typus
</emphasis>
,
<taxonomicName id="4C239CE31D11FFDAFBAB93B4FACC557E" baseAuthorityName="Wilkinson, Young &amp; Benton" baseAuthorityYear="2008" box="[1033,1289,503,528]" class="Reptilia" family="Metriorhynchidae" genus="Torvoneustes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="45" pageNumber="178" phylum="Chordata" rank="species" species="carpenteri">
<emphasis id="B9573B721D11FFDAFBAB93B4FACC557E" box="[1033,1289,503,528]" italics="true" pageId="45" pageNumber="178">Torvoneustes carpenteri</emphasis>
</taxonomicName>
,
<taxonomicName id="4C239CE31D11FFDAFAB993B5FC59555E" authority="(Andrews, 1909)" authorityName="(Andrews)" baseAuthorityName="Andrews" baseAuthorityYear="1909" class="Reptilia" family="Teleosauridae" genus="Lemmysuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="45" pageNumber="178" phylum="Chordata" rank="species" species="obtusidens">
<emphasis id="B9573B721D11FFDAFAB993B5FC59555E" italics="true" pageId="45" pageNumber="178">Lemmysuchus obtusidens</emphasis>
</taxonomicName>
, or
<taxonomicName id="4C239CE31D11FFDAFC6F9055FB1D555E" authority="(Jager, 1828)" authorityName="(Jager)" baseAuthorityName="Jager" baseAuthorityYear="1828" box="[973,1240,534,560]" class="Reptilia" family="Teleosauridae" genus="Macrospondylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="45" pageNumber="178" phylum="Chordata" rank="species" species="bollensis">
<emphasis id="B9573B721D11FFDAFC6F9055FB1D555E" box="[973,1240,534,560]" italics="true" pageId="45" pageNumber="178">Macrospondylus bollensis</emphasis>
</taxonomicName>
. The distal margin of the ischium of
<taxonomicName id="4C239CE31D11FFDAFC5C9075FAD1553E" authorityName="Sachs et al." authorityYear="2019" baseAuthorityName="Sachs, Young, Abel &amp; Mallison" baseAuthorityYear="2019" box="[1022,1300,566,592]" class="Reptilia" family="Metriorhynchidae" genus="Cricosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="45" pageNumber="178" phylum="Chordata" rank="species" species="bambergensis">
<emphasis id="B9573B721D11FFDAFC5C9075FAD1553E" box="[1022,1300,566,592]" italics="true" pageId="45" pageNumber="178">Cricosaurus bambergensis</emphasis>
</taxonomicName>
uniting both processes is straight as for many other thalattosuchians (e.g.
<emphasis id="B9573B721D11FFDAFCCA9035FBF655FE" box="[872,1075,630,656]" italics="true" pageId="45" pageNumber="178">
<taxonomicName id="4C239CE31D11FFDAFCCA9035FC3755FE" box="[872,1010,630,656]" class="Reptilia" family="Metriorhynchidae" genus="Pelagosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="45" pageNumber="178" phylum="Chordata" rank="genus">Pelagosaurus</taxonomicName>
typus
</emphasis>
, ‘
<taxonomicName id="4C239CE31D11FFDAFBEE9035FC1E55DE" authority="NHMUK PV R" authorityName=", NHMUK PV R" baseAuthorityName="Eudes-Deslongchamps" baseAuthorityYear="1867" class="Reptilia" family="Metriorhynchidae" genus="Metriorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="45" pageNumber="178" phylum="Chordata" rank="species" species="brachyrhynchus">
<emphasis id="B9573B721D11FFDAFBEE9035FB3255FE" box="[1100,1271,630,656]" italics="true" pageId="45" pageNumber="178">Metriorhynchus</emphasis>
’
<emphasis id="B9573B721D11FFDAFAAA9035FA6A55FE" box="[1288,1455,630,656]" italics="true" pageId="45" pageNumber="178">brachyrhynchus</emphasis>
NHMUK PV R
</taxonomicName>
4763,
<taxonomicName id="4C239CE31D11FFDAFB8590D5FAC855C1" authorityName="SMNS" baseAuthorityName="Plieninger" baseAuthorityYear="1846" box="[1063,1293,662,688]" class="Reptilia" family="Metriorhynchidae" genus="Dakosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="45" pageNumber="178" phylum="Chordata" rank="species" species="maximus">
<emphasis id="B9573B721D11FFDAFB8590D5FAC855C1" box="[1063,1293,662,688]" italics="true" pageId="45" pageNumber="178">Dakosaurus maximus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C239CE31D11FFDAFABE90D5FC5955A1" authority="(Andrews, 1909)" authorityName="(Andrews)" baseAuthorityName="Andrews" baseAuthorityYear="1909" class="Reptilia" family="Teleosauridae" genus="Lemmysuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="45" pageNumber="178" phylum="Chordata" rank="species" species="obtusidens">
<emphasis id="B9573B721D11FFDAFABE90D5FC5955A1" italics="true" pageId="45" pageNumber="178">Lemmysuchus obtusidens</emphasis>
</taxonomicName>
,
<taxonomicName id="4C239CE31D11FFDAFC0990F6FB7255A1" authority="(Jager, 1828)" authorityName="(Jager)" baseAuthorityName="Jager" baseAuthorityYear="1828" box="[939,1207,693,719]" class="Reptilia" family="Teleosauridae" genus="Macrospondylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="45" pageNumber="178" phylum="Chordata" rank="species" species="bollensis">
<emphasis id="B9573B721D11FFDAFC0990F6FB7255A1" box="[939,1207,693,719]" italics="true" pageId="45" pageNumber="178">Macrospondylus bollensis</emphasis>
</taxonomicName>
, etc.).
</paragraph>
<paragraph id="8B9CE7601D11FFDAFCE19096FB885361" blockId="45.[812,1456,215,1039]" pageId="45" pageNumber="178">
The anterior and posterior margins of the ischium of
<taxonomicName id="4C239CE31D11FFDAFC8F90B5FB855461" authorityName="Sachs et al." authorityYear="2019" baseAuthorityName="Sachs, Young, Abel &amp; Mallison" baseAuthorityYear="2019" box="[813,1088,757,784]" class="Reptilia" family="Metriorhynchidae" genus="Cricosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="45" pageNumber="178" phylum="Chordata" rank="species" species="bambergensis">
<emphasis id="B9573B721D11FFDAFC8F90B5FB855461" box="[813,1088,757,784]" italics="true" pageId="45" pageNumber="178">Cricosaurus bambergensis</emphasis>
</taxonomicName>
are markedly concave as in
<taxonomicName id="4C239CE31D11FFDAFA2190B5FC2C5441" baseAuthorityName="Fraas" baseAuthorityYear="1901" class="Reptilia" family="Metriorhynchidae" genus="Cricosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="45" pageNumber="178" phylum="Chordata" rank="species" species="suevicus">
<emphasis id="B9573B721D11FFDAFA2190B5FC2C5441" italics="true" pageId="45" pageNumber="178">Cricosaurus suevicus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C239CE31D11FFDAFB859156FAD15441" authorityName="SMNS" baseAuthorityName="Plieninger" baseAuthorityYear="1846" box="[1063,1300,789,815]" class="Reptilia" family="Metriorhynchidae" genus="Dakosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="45" pageNumber="178" phylum="Chordata" rank="species" species="maximus">
<emphasis id="B9573B721D11FFDAFB859156FAD15441" box="[1063,1300,789,815]" italics="true" pageId="45" pageNumber="178">Dakosaurus maximus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C239CE31D11FFDAFA849155FC5C5421" baseAuthorityName="Wilkinson, Young &amp; Benton" baseAuthorityYear="2008" class="Reptilia" family="Metriorhynchidae" genus="Torvoneustes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="45" pageNumber="178" phylum="Chordata" rank="species" species="carpenteri">
<emphasis id="B9573B721D11FFDAFA849155FC5C5421" italics="true" pageId="45" pageNumber="178">Torvoneustes carpenteri</emphasis>
</taxonomicName>
, with the anterior margin displaying the greatest intensity. Dorsally, the anterior margin of the ischium leads to the anterior peduncle, which appears to be short and rounded as in other metriorhynchoids (e.g. ‘
<taxonomicName id="4C239CE31D11FFDAFAFE91D5FB2654A1" authority="NHMUK PV R" authorityName=", NHMUK PV R" baseAuthorityName="Eudes-Deslongchamps" baseAuthorityYear="1867" class="Reptilia" family="Metriorhynchidae" genus="Metriorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="45" pageNumber="178" phylum="Chordata" rank="species" species="brachyrhynchus">
<emphasis id="B9573B721D11FFDAFAFE91D5FC4C54A1" italics="true" pageId="45" pageNumber="178">Metriorhynchus</emphasis>
’
<emphasis id="B9573B721D11FFDAFC3791F6FBF354A1" box="[917,1078,949,975]" italics="true" pageId="45" pageNumber="178">brachyrhynchus</emphasis>
NHMUK PV R
</taxonomicName>
4763,
<taxonomicName id="4C239CE31D11FFDAFA8E91F6FC525480" baseAuthorityName="Wilkinson, Young &amp; Benton" baseAuthorityYear="2008" class="Reptilia" family="Metriorhynchidae" genus="Torvoneustes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="45" pageNumber="178" phylum="Chordata" rank="species" species="carpenteri">
<emphasis id="B9573B721D11FFDAFA8E91F6FC525480" italics="true" pageId="45" pageNumber="178">Torvoneustes carpenteri</emphasis>
</taxonomicName>
,
<taxonomicName id="4C239CE31D11FFDAFC019197FB005480" authority="(Blainville, 1853)" authorityName="(Blainville)" authorityYear="1852" baseAuthorityName="Blainville" baseAuthorityYear="1853" box="[931,1221,980,1006]" class="Reptilia" family="Metriorhynchidae" genus="Thalattosuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="45" pageNumber="178" phylum="Chordata" rank="species" species="superciliosus">
<emphasis id="B9573B721D11FFDAFC019197FB005480" box="[931,1221,980,1006]" italics="true" pageId="45" pageNumber="178">Thalattosuchus superciliosus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C239CE31D11FFDAFB719196FA6D5480" baseAuthorityName="Fraas" baseAuthorityYear="1901" box="[1235,1448,981,1007]" class="Reptilia" family="Metriorhynchidae" genus="Cricosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="45" pageNumber="178" phylum="Chordata" rank="species" species="suevicus">
<emphasis id="B9573B721D11FFDAFB719196FA6D5480" box="[1235,1448,981,1007]" italics="true" pageId="45" pageNumber="178">Cricosaurus suevicus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C239CE31D11FFDAFC8F91B6FBC05360" authorityName="NHMUK PV R" authorityYear="1230" box="[813,1029,1013,1039]" class="Reptilia" family="Metriorhynchidae" genus="Geosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="45" pageNumber="178" phylum="Chordata" rank="species" species="giganteus">
<emphasis id="B9573B721D11FFDAFC8F91B6FBC05360" box="[813,1029,1013,1039]" italics="true" pageId="45" pageNumber="178">Geosaurus giganteus</emphasis>
</taxonomicName>
, etc.).
</paragraph>
<paragraph id="8B9CE7601D11FFDAFC8F9677FCAD5320" blockId="45.[813,872,1076,1102]" box="[813,872,1076,1102]" pageId="45" pageNumber="178">
<heading id="D0D4500C1D11FFDAFC8F9677FCAD5320" box="[813,872,1076,1102]" fontSize="11" level="2" pageId="45" pageNumber="178" reason="8">
<emphasis id="B9573B721D11FFDAFC8F9677FCAD5320" box="[813,872,1076,1102]" italics="true" pageId="45" pageNumber="178">Pubis</emphasis>
</heading>
</paragraph>
<paragraph id="8B9CE7601D11FFDAFC8F9617FBD25005" blockId="45.[811,1456,1108,1899]" pageId="45" pageNumber="178">
The pubis of
<taxonomicName id="4C239CE31D11FFDAFC629616FB175300" authorityName="Sachs et al." authorityYear="2019" baseAuthorityName="Sachs, Young, Abel &amp; Mallison" baseAuthorityYear="2019" box="[960,1234,1108,1135]" class="Reptilia" family="Metriorhynchidae" genus="Cricosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="45" pageNumber="178" phylum="Chordata" rank="species" species="bambergensis">
<emphasis id="B9573B721D11FFDAFC629616FB175300" box="[960,1234,1108,1135]" italics="true" pageId="45" pageNumber="178">Cricosaurus bambergensis</emphasis>
</taxonomicName>
(
<figureCitation id="1318FBE51D11FFDAFB469617FAF65300" box="[1252,1331,1108,1134]" captionStart="FIG" captionStartId="46.[132,143,1819,1836]" captionTargetBox="[141,1456,181,1794]" captionTargetId="figure-68@46.[141,1447,269,1777]" captionTargetPageId="46" captionText="FIG. 21. — Pelvic and thoracic girdle elements of Cricosaurus bambergensis (Sachs, Young, Abel &amp; Mallison, 2019) (Sachs et al. 2019), NKMB-P-Watt14/274 (holotype): A, overview of pelvic girdle and hindlimb; B, overview of thoracic girdle and forelimb; C, detail of pelvic girdle. Modified from Sachs et al. (2019). Scale bars: 1 cm." figureDoi="http://doi.org/10.5281/zenodo.11106644" httpUri="https://zenodo.org/record/11106644/files/figure.png" pageId="45" pageNumber="178">Fig. 21</figureCitation>
) resembles those of other geosaurines and rhacheosaurines (i.e.
<taxonomicName id="4C239CE31D11FFDAFA209637FC2453C0" baseAuthorityName="Fraas" baseAuthorityYear="1901" class="Reptilia" family="Metriorhynchidae" genus="Cricosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="45" pageNumber="178" phylum="Chordata" rank="species" species="suevicus">
<emphasis id="B9573B721D11FFDAFA209637FC2453C0" italics="true" pageId="45" pageNumber="178">Cricosaurus suevicus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C239CE31D11FFDAFC4D96D7FAC053C0" authorityName="Sachs et al." authorityYear="2021" baseAuthorityName="Sachs, Young, Abel &amp; Mallison" baseAuthorityYear="2021" box="[1007,1285,1172,1198]" class="Reptilia" family="Metriorhynchidae" genus="Cricosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="45" pageNumber="178" phylum="Chordata" rank="species" species="albersdoerferi">
<emphasis id="B9573B721D11FFDAFC4D96D7FAC053C0" box="[1007,1285,1172,1198]" italics="true" pageId="45" pageNumber="178">Cricosaurus albersdoerferi</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C239CE31D11FFDAFAE396D7FC4953A0" authorityName="NHMUK PV R" authorityYear="1230" class="Reptilia" family="Metriorhynchidae" genus="Geosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="45" pageNumber="178" phylum="Chordata" rank="species" species="giganteus">
<emphasis id="B9573B721D11FFDAFAE396D7FC4953A0" italics="true" pageId="45" pageNumber="178">Geosaurus giganteus</emphasis>
</taxonomicName>
) in possessing a relatively reduced pubic symphysis. However, it differs from
<taxonomicName id="4C239CE31D11FFDAFBE29697FAD25383" authorityName="NHMUK PV R" authorityYear="1230" box="[1088,1303,1236,1262]" class="Reptilia" family="Metriorhynchidae" genus="Geosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="45" pageNumber="178" phylum="Chordata" rank="species" species="giganteus">
<emphasis id="B9573B721D11FFDAFBE29697FAD25383" box="[1088,1303,1236,1262]" italics="true" pageId="45" pageNumber="178">Geosaurus giganteus</emphasis>
</taxonomicName>
in displaying a mediolaterally larger pubic apron (i.e. greater than twice the size of the pubic peduncle, as in other
<taxonomicName id="4C239CE31D11FFDAFB5F9757FC9E5223" pageId="45" pageNumber="178">
<emphasis id="B9573B721D11FFDAFB5F9757FABE5240" box="[1277,1403,1300,1326]" italics="true" pageId="45" pageNumber="178">Cricosaurus</emphasis>
species
</taxonomicName>
). The pubic blade of
<taxonomicName id="4C239CE31D11FFDAFBF19777FAAF5223" authorityName="Sachs et al." authorityYear="2019" baseAuthorityName="Sachs, Young, Abel &amp; Mallison" baseAuthorityYear="2019" box="[1107,1386,1331,1358]" class="Reptilia" family="Metriorhynchidae" genus="Cricosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="45" pageNumber="178" phylum="Chordata" rank="species" species="bambergensis">
<emphasis id="B9573B721D11FFDAFBF19777FAAF5223" box="[1107,1386,1331,1358]" italics="true" pageId="45" pageNumber="178">Cricosaurus bambergensis</emphasis>
</taxonomicName>
looks like an axe head as its distal margin is entirely convex, and as the margins leading to it are strongly concave (i.e. the anterior and posterior margins of the pubis). Still, the pubic apron of
<taxonomicName id="4C239CE31D11FFDAFC3397F0FB6652A3" authorityName="Sachs et al." authorityYear="2019" baseAuthorityName="Sachs, Young, Abel &amp; Mallison" baseAuthorityYear="2019" box="[913,1187,1459,1485]" class="Reptilia" family="Metriorhynchidae" genus="Cricosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="45" pageNumber="178" phylum="Chordata" rank="species" species="bambergensis">
<emphasis id="B9573B721D11FFDAFC3397F0FB6652A3" box="[913,1187,1459,1485]" italics="true" pageId="45" pageNumber="178">Cricosaurus bambergensis</emphasis>
</taxonomicName>
is slightly asymmetrical with its medial half being larger than the lateral one, not unlike other
<taxonomicName id="4C239CE31D11FFDAFC6197B0FB505163" box="[963,1173,1523,1549]" pageId="45" pageNumber="178">
<emphasis id="B9573B721D11FFDAFC6197B0FB845163" box="[963,1089,1523,1549]" italics="true" pageId="45" pageNumber="178">Cricosaurus</emphasis>
species
</taxonomicName>
for which the effect was more pronounced (
<taxonomicName id="4C239CE31D11FFDAFBA09450FB1D5142" baseAuthorityName="Fraas" baseAuthorityYear="1901" box="[1026,1240,1555,1581]" class="Reptilia" family="Metriorhynchidae" genus="Cricosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="45" pageNumber="178" phylum="Chordata" rank="species" species="suevicus">
<emphasis id="B9573B721D11FFDAFBA09450FB1D5142" box="[1026,1240,1555,1581]" italics="true" pageId="45" pageNumber="178">Cricosaurus suevicus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C239CE31D11FFDAFB449450FCB85122" authorityName="Sachs et al." authorityYear="2021" baseAuthorityName="Sachs, Young, Abel &amp; Mallison" baseAuthorityYear="2021" class="Reptilia" family="Metriorhynchidae" genus="Cricosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="45" pageNumber="178" phylum="Chordata" rank="species" species="albersdoerferi">
<emphasis id="B9573B721D11FFDAFB449450FCB85122" italics="true" pageId="45" pageNumber="178">Cricosaurus albersdoerferi</emphasis>
</taxonomicName>
). In
<taxonomicName id="4C239CE31D11FFDAFC129470FB7E5122" authorityName="Sachs et al." authorityYear="2019" baseAuthorityName="Sachs, Young, Abel &amp; Mallison" baseAuthorityYear="2019" box="[944,1211,1586,1613]" class="Reptilia" family="Metriorhynchidae" genus="Cricosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="45" pageNumber="178" phylum="Chordata" rank="species" species="bambergensis">
<emphasis id="B9573B721D11FFDAFC129470FB7E5122" box="[944,1211,1586,1613]" italics="true" pageId="45" pageNumber="178">Cricosaurus bambergensis</emphasis>
</taxonomicName>
, the junction between the distal margin and the lateral margin is sharper than the opposite corner. Like
<taxonomicName id="4C239CE31D11FFDAFBEA9430FAE451E2" baseAuthorityName="Fraas" baseAuthorityYear="1901" box="[1096,1313,1651,1677]" class="Reptilia" family="Metriorhynchidae" genus="Cricosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="45" pageNumber="178" phylum="Chordata" rank="species" species="suevicus">
<emphasis id="B9573B721D11FFDAFBEA9430FAE451E2" box="[1096,1313,1651,1677]" italics="true" pageId="45" pageNumber="178">Cricosaurus suevicus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C239CE31D11FFDAFA939431FC4B51C2" authorityName="SMNS" baseAuthorityName="Plieninger" baseAuthorityYear="1846" class="Reptilia" family="Metriorhynchidae" genus="Dakosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="45" pageNumber="178" phylum="Chordata" rank="species" species="maximus">
<emphasis id="B9573B721D11FFDAFA939431FC4B51C2" italics="true" pageId="45" pageNumber="178">Dakosaurus maximus</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C239CE31D11FFDAFC7C94D0FB7351C2" authorityName="NHMUK PV R" authorityYear="1230" box="[990,1206,1683,1709]" class="Reptilia" family="Metriorhynchidae" genus="Geosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="45" pageNumber="178" phylum="Chordata" rank="species" species="giganteus">
<emphasis id="B9573B721D11FFDAFC7C94D0FB7351C2" box="[990,1206,1683,1709]" italics="true" pageId="45" pageNumber="178">Geosaurus giganteus</emphasis>
</taxonomicName>
, the thinnest portion of the shaft of
<taxonomicName id="4C239CE31D11FFDAFC7E94F1FB3451A2" authorityName="Sachs et al." authorityYear="2019" baseAuthorityName="Sachs, Young, Abel &amp; Mallison" baseAuthorityYear="2019" box="[988,1265,1714,1740]" class="Reptilia" family="Metriorhynchidae" genus="Cricosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="45" pageNumber="178" phylum="Chordata" rank="species" species="bambergensis">
<emphasis id="B9573B721D11FFDAFC7E94F1FB3451A2" box="[988,1265,1714,1740]" italics="true" pageId="45" pageNumber="178">Cricosaurus bambergensis</emphasis>
</taxonomicName>
is smaller mediolaterally than the pubic peduncle. The articular surface of the pubic peduncle of
<taxonomicName id="4C239CE31D11FFDAFB9C94B1FA885062" authorityName="Sachs et al." authorityYear="2019" baseAuthorityName="Sachs, Young, Abel &amp; Mallison" baseAuthorityYear="2019" box="[1086,1357,1778,1804]" class="Reptilia" family="Metriorhynchidae" genus="Cricosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="45" pageNumber="178" phylum="Chordata" rank="species" species="bambergensis">
<emphasis id="B9573B721D11FFDAFB9C94B1FA885062" box="[1086,1357,1778,1804]" italics="true" pageId="45" pageNumber="178">Cricosaurus bambergensis</emphasis>
</taxonomicName>
presents a slightly convex outline. Its extremities form rounded corners laterally and medially where they meets with the margins of the shaft.
</paragraph>
</subSubSection>
</treatment>
</document>
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