336 lines
33 KiB
XML
336 lines
33 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.898.47118" ID-GBIF-Dataset="6a9aaa1b-b903-4472-865d-7edce422a2dc" ID-PMC="PMC6917629" ID-Pensoft-Pub="1313-2970-898-1" ID-Pensoft-UUID="278CAE686DE65920AF5B0FE8BA1BBDC1" ID-PubMed="31866732" ID-ZooBank="7B407C809E644F5995C2B3229CF78C6B" ModsDocID="1313-2970-898-1" checkinTime="1575997091264" checkinUser="pensoft" docAuthor="Godunko, Roman J., Neumann, Christian & Staniczek, Arnold H." docDate="2019" docId="EC67D71F662D5B5DBF37B9A705B11950" docLanguage="en" docName="ZooKeys 898: 1-26" docOrigin="ZooKeys 898" docSource="http://dx.doi.org/10.3897/zookeys.898.47118" docTitle="Siphloplecton studemannae Godunko, Neumann & Staniczek, 2019, sp. nov." docType="treatment" docUuid="2D1C8631-DAFF-4B84-B10D-7D7FAB3E5334" docUuidSource="ZooBank" docVersion="5" id="278CAE686DE65920AF5B0FE8BA1BBDC1" lastPageNumber="1" masterDocId="278CAE686DE65920AF5B0FE8BA1BBDC1" masterDocTitle="Revision of fossil Metretopodidae (Insecta, Ephemeroptera) in Baltic amber - Part 4: Description of two new species of Siphloplecton Clemens, 1915, with notes on the new S. jaegeri species group and with key to fossil male adults of Siphloplecton" masterLastPageNumber="26" masterPageNumber="1" pageNumber="1" updateTime="1668168222060" updateUser="ExternalLinkService">
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<mods:title>Revision of fossil Metretopodidae (Insecta, Ephemeroptera) in Baltic amber - Part 4: Description of two new species of Siphloplecton Clemens, 1915, with notes on the new S. jaegeri species group and with key to fossil male adults of Siphloplecton</mods:title>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Godunko, Roman J.</mods:namePart>
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<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Neumann, Christian</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Staniczek, Arnold H.</mods:namePart>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2019</mods:date>
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<mods:detail type="volume">
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<mods:number>898</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>26</mods:end>
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<mods:url>http://dx.doi.org/10.3897/zookeys.898.47118</mods:url>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.898.47118</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-898-1</mods:identifier>
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<mods:identifier type="ZooBank">7B407C809E644F5995C2B3229CF78C6B</mods:identifier>
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<mods:identifier type="Pensoft-UUID">278CAE686DE65920AF5B0FE8BA1BBDC1</mods:identifier>
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<treatment ID-GBIF-Taxon="160689324" LSID="urn:lsid:zoobank.org:act:2D1C8631-DAFF-4B84-B10D-7D7FAB3E5334" httpUri="http://treatment.plazi.org/id/EC67D71F662D5B5DBF37B9A705B11950" lastPageNumber="1" pageId="0" pageNumber="1">
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<subSubSection pageId="0" pageNumber="1" type="nomenclature">
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName LSID="EC67D71F-662D-5B5D-BF37-B9A705B11950" class="Insecta" family="Metretopodidae" genus="Siphloplecton" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Siphloplecton studemannae" order="Ephemeroptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="studemannae">Siphloplecton studemannae</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
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<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Siphloplecton studemannae sp. nov., SMNS, BB- 2626, holotype, female imago (photographs) A general dorsal view B general ventral view C left forewing in dorsal view D wing base and cubital field of right forewing in ventral view E left hind wing in ventral view F right hind leg in medial view G claws of hind legs H abdominal segments VII-X in ventral view." figureDoi="10.3897/zookeys.898.47118.figure8" httpUri="https://binary.pensoft.net/fig/363724" pageId="0" pageNumber="1" tableDoi="10.3897/zookeys.898.47118.figure8">Figures 8</figureCitation>
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,
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<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Siphloplecton studemannae sp. nov., SMNS, BB- 2626, holotype, female imago (line drawing): abdomen and part of left forewing in ventral view." figureDoi="10.3897/zookeys.898.47118.figure9" httpUri="https://binary.pensoft.net/fig/363725" pageId="0" pageNumber="1" tableDoi="10.3897/zookeys.898.47118.figure9">9</figureCitation>
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,
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<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Siphloplecton studemannae sp. nov., MNHK, MP / 1626, paratype, female imago (photographs) A general ventral view B left forewing in dorsal view C right hind wing in ventral view D abdominal segments VII-X in ventral view." figureDoi="10.3897/zookeys.898.47118.figure10" httpUri="https://binary.pensoft.net/fig/363726" pageId="0" pageNumber="1" tableDoi="10.3897/zookeys.898.47118.figure10">10</figureCitation>
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,
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<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Siphloplecton studemannae sp. nov., MNHK, MP / 1626, paratype, female imago (line drawings) A fragments of left fore- and hind wings, dorsal view B abdominal segments VII-X in ventrolateral view." figureDoi="10.3897/zookeys.898.47118.figure11" httpUri="https://binary.pensoft.net/fig/363727" pageId="0" pageNumber="1" tableDoi="10.3897/zookeys.898.47118.figure11">11</figureCitation>
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<tableCitation captionStartId="T1" httpUri="http://table.plazi.org/id/23F53EE7FB0FC4D865035A6D181A4855" pageId="0" pageNumber="1" tableUuid="23F53EE7FB0FC4D865035A6D181A4855">; Table 1</tableCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="materials_examined">
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<paragraph pageId="0" pageNumber="1">Material examined.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" pageId="0" pageNumber="1">
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<emphasis italics="true" pageId="0" pageNumber="1">Holotype</emphasis>
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.
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</emphasis>
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Female imago in Baltic amber (Eocene), SMNS
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<normalizedToken originalValue="BB–">BB-</normalizedToken>
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2626. Well preserved specimen in clear amber, well visible in dorsoventral aspect. Right foreleg lost; hind wings (especially left wing) twisted. Head and thorax covered by
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<normalizedToken originalValue="“Verlumung”">"Verlumung"</normalizedToken>
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. Additionally, piece of amber with numerous cracks, thus details of thoracic terga invisible, and thorax only partly visible from ventral side. For measurements see Table
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<tableCitation captionStartId="T1" httpUri="http://table.plazi.org/id/23F53EE7FB0FC4D865035A6D181A4855" pageId="0" pageNumber="1" tableUuid="23F53EE7FB0FC4D865035A6D181A4855">1</tableCitation>
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.
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" pageId="0" pageNumber="1">
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<emphasis italics="true" pageId="0" pageNumber="1">Paratype</emphasis>
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.
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</emphasis>
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Female imago in Baltic amber (Eocene), MNHK, MP/1626. Partly damaged specimen, visible in ventral aspect. View on body hampered by resin influxes, numerous cracks, and considerable
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<normalizedToken originalValue="“Verlumung”">"Verlumung"</normalizedToken>
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, so thoracic sutures are hardly visible. Head lost. Pronotum damaged, its structure invisible. Distal part of mesonotum, metanotum and abdominal segments I−IV dorsally covered by plant tissue. Right forewing twisted along its length; left forewing lacks its basal part and distal end. Right foreleg, right middle leg, left middle tarsus, and both hind legs lost. Cerci only partly preserved. For measurements see Table
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<tableCitation captionStartId="T1" httpUri="http://table.plazi.org/id/23F53EE7FB0FC4D865035A6D181A4855" pageId="0" pageNumber="1" tableUuid="23F53EE7FB0FC4D865035A6D181A4855">1</tableCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="description">
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<paragraph pageId="0" pageNumber="1">Description of holotype.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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General colour of body pale, yellow to yellowish-brown. Ventral side of body slightly darker than dorsal side. Ocelli well preserved. Eyes medially separated, but approximated (
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<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Siphloplecton studemannae sp. nov., SMNS, BB- 2626, holotype, female imago (photographs) A general dorsal view B general ventral view C left forewing in dorsal view D wing base and cubital field of right forewing in ventral view E left hind wing in ventral view F right hind leg in medial view G claws of hind legs H abdominal segments VII-X in ventral view." figureDoi="10.3897/zookeys.898.47118.figure8" httpUri="https://binary.pensoft.net/fig/363724" pageId="0" pageNumber="1" tableDoi="10.3897/zookeys.898.47118.figure8">Fig. 8A</figureCitation>
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). Distance between eyes 0.22
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<normalizedToken originalValue="×">x</normalizedToken>
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of head width. Antennae complete, slightly longer than head, pale.
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</paragraph>
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<caption doi="10.3897/zookeys.898.47118.figure8" httpUri="https://binary.pensoft.net/fig/363724" pageId="0" pageNumber="1" start="Figure 8" startId="F8">
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" pageId="0" pageNumber="1">Figure 8.</emphasis>
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<taxonomicName class="Insecta" family="Metretopodidae" genus="Siphloplecton" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Siphloplecton studemannae" order="Ephemeroptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="studemannae">
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<emphasis italics="true" pageId="0" pageNumber="1">Siphloplecton studemannae</emphasis>
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</taxonomicName>
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sp. nov., SMNS,
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<normalizedToken originalValue="BB–">BB-</normalizedToken>
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2626, holotype, female imago (photographs)
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<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
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general dorsal view
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<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
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general ventral view
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<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
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left forewing in dorsal view
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<emphasis bold="true" pageId="0" pageNumber="1">D</emphasis>
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wing base and cubital field of right forewing in ventral view
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<emphasis bold="true" pageId="0" pageNumber="1">E</emphasis>
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left hind wing in ventral view
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<emphasis bold="true" pageId="0" pageNumber="1">F</emphasis>
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right hind leg in medial view
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<emphasis bold="true" pageId="0" pageNumber="1">G</emphasis>
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claws of hind legs
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<emphasis bold="true" pageId="0" pageNumber="1">H</emphasis>
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abdominal segments
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<normalizedToken originalValue="VII–X">VII-X</normalizedToken>
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in ventral view.
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</paragraph>
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</caption>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Thorax</emphasis>
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ventrally yellowish coloured by
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<normalizedToken originalValue="“Verlumung”">"Verlumung"</normalizedToken>
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, mainly in posterior part. Furcasternal protuberances of mesothorax contiguous (
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<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Siphloplecton studemannae sp. nov., SMNS, BB- 2626, holotype, female imago (photographs) A general dorsal view B general ventral view C left forewing in dorsal view D wing base and cubital field of right forewing in ventral view E left hind wing in ventral view F right hind leg in medial view G claws of hind legs H abdominal segments VII-X in ventral view." figureDoi="10.3897/zookeys.898.47118.figure8" httpUri="https://binary.pensoft.net/fig/363724" pageId="0" pageNumber="1" tableDoi="10.3897/zookeys.898.47118.figure8">Fig. 8B</figureCitation>
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); lateral aspect of thorax not visible.
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Wings</emphasis>
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translucent, hyaline, not pigmented. Wing venation well visible only from ventral side; basal part of wings covered by numerous cracks from dorsal side. Cubital field of both forewings with one pair of intercalary veins (iCu2, iCu3) towards CuP and one additional vein (iCu1) near CuA (
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<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Siphloplecton studemannae sp. nov., SMNS, BB- 2626, holotype, female imago (photographs) A general dorsal view B general ventral view C left forewing in dorsal view D wing base and cubital field of right forewing in ventral view E left hind wing in ventral view F right hind leg in medial view G claws of hind legs H abdominal segments VII-X in ventral view." figureDoi="10.3897/zookeys.898.47118.figure8" httpUri="https://binary.pensoft.net/fig/363724" pageId="0" pageNumber="1" tableDoi="10.3897/zookeys.898.47118.figure8">
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Figs 8
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<normalizedToken originalValue="C–D">C-D</normalizedToken>
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</figureCitation>
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,
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<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Siphloplecton studemannae sp. nov., SMNS, BB- 2626, holotype, female imago (line drawing): abdomen and part of left forewing in ventral view." figureDoi="10.3897/zookeys.898.47118.figure9" httpUri="https://binary.pensoft.net/fig/363725" pageId="0" pageNumber="1" tableDoi="10.3897/zookeys.898.47118.figure9">9</figureCitation>
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). Right hind wing hardly visible, twisted; left hind wing completely preserved with three pairs of triads, 0.34
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<normalizedToken originalValue="×">x</normalizedToken>
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of right forewing length. Costal process bluntly pointed apically and small (
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<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Siphloplecton studemannae sp. nov., SMNS, BB- 2626, holotype, female imago (photographs) A general dorsal view B general ventral view C left forewing in dorsal view D wing base and cubital field of right forewing in ventral view E left hind wing in ventral view F right hind leg in medial view G claws of hind legs H abdominal segments VII-X in ventral view." figureDoi="10.3897/zookeys.898.47118.figure8" httpUri="https://binary.pensoft.net/fig/363724" pageId="0" pageNumber="1" tableDoi="10.3897/zookeys.898.47118.figure8">Fig. 8E</figureCitation>
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).
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Legs</emphasis>
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relatively well preserved, except forelegs (right is lost; left incomplete). Measurements of leg segments in Table
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<tableCitation captionStartId="T1" httpUri="http://table.plazi.org/id/23F53EE7FB0FC4D865035A6D181A4855" pageId="0" pageNumber="1" tableUuid="23F53EE7FB0FC4D865035A6D181A4855">1</tableCitation>
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. Tibiae of middle and hind legs each with trace of tibiopatellar suture; first tarsomere of middle and hind legs fused with tibia (
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<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Siphloplecton studemannae sp. nov., SMNS, BB- 2626, holotype, female imago (photographs) A general dorsal view B general ventral view C left forewing in dorsal view D wing base and cubital field of right forewing in ventral view E left hind wing in ventral view F right hind leg in medial view G claws of hind legs H abdominal segments VII-X in ventral view." figureDoi="10.3897/zookeys.898.47118.figure8" httpUri="https://binary.pensoft.net/fig/363724" pageId="0" pageNumber="1" tableDoi="10.3897/zookeys.898.47118.figure8">Fig. 8F</figureCitation>
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). Tarsi with 5 tarsomeres; tarsal claws dissimilar: one hooked and one blunt (
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<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Siphloplecton studemannae sp. nov., SMNS, BB- 2626, holotype, female imago (photographs) A general dorsal view B general ventral view C left forewing in dorsal view D wing base and cubital field of right forewing in ventral view E left hind wing in ventral view F right hind leg in medial view G claws of hind legs H abdominal segments VII-X in ventral view." figureDoi="10.3897/zookeys.898.47118.figure8" httpUri="https://binary.pensoft.net/fig/363724" pageId="0" pageNumber="1" tableDoi="10.3897/zookeys.898.47118.figure8">Fig. 8G</figureCitation>
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).
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Abdominal</emphasis>
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segments completely preserved, pale; terga
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<normalizedToken originalValue="VIII–X">VIII-X</normalizedToken>
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slightly darker than remaining terga. Abdominal sterna slightly covered by
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<normalizedToken originalValue="“Verlumung”">"Verlumung"</normalizedToken>
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. Subgenital plate relatively broad, 2.00
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<normalizedToken originalValue="×">x</normalizedToken>
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as wide as long, convex and rounded apically. Subanal plate not elongated, narrow, with pointed tip (
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<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Siphloplecton studemannae sp. nov., SMNS, BB- 2626, holotype, female imago (photographs) A general dorsal view B general ventral view C left forewing in dorsal view D wing base and cubital field of right forewing in ventral view E left hind wing in ventral view F right hind leg in medial view G claws of hind legs H abdominal segments VII-X in ventral view." figureDoi="10.3897/zookeys.898.47118.figure8" httpUri="https://binary.pensoft.net/fig/363724" pageId="0" pageNumber="1" tableDoi="10.3897/zookeys.898.47118.figure8">Figs 8H</figureCitation>
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,
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<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Siphloplecton studemannae sp. nov., SMNS, BB- 2626, holotype, female imago (line drawing): abdomen and part of left forewing in ventral view." figureDoi="10.3897/zookeys.898.47118.figure9" httpUri="https://binary.pensoft.net/fig/363725" pageId="0" pageNumber="1" tableDoi="10.3897/zookeys.898.47118.figure9">9</figureCitation>
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). Paracercus poorly visible, vestigial, with at least 4 visible segments; cerci dark, partly preserved.
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</paragraph>
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<caption doi="10.3897/zookeys.898.47118.figure9" httpUri="https://binary.pensoft.net/fig/363725" pageId="0" pageNumber="1" start="Figure 9" startId="F9">
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" pageId="0" pageNumber="1">Figure 9.</emphasis>
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<taxonomicName class="Insecta" family="Metretopodidae" genus="Siphloplecton" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Siphloplecton studemannae" order="Ephemeroptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="studemannae">
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<emphasis italics="true" pageId="0" pageNumber="1">Siphloplecton studemannae</emphasis>
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</taxonomicName>
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sp. nov., SMNS,
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<normalizedToken originalValue="BB–">BB-</normalizedToken>
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2626, holotype, female imago (line drawing): abdomen and part of left forewing in ventral view.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="description">
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<paragraph pageId="0" pageNumber="1">Description of paratype.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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Body colouration light, yellowish-brown to brown. Irregular brown, dirty brown to black maculation over body, especially on forewings and legs (
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<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Siphloplecton studemannae sp. nov., MNHK, MP / 1626, paratype, female imago (photographs) A general ventral view B left forewing in dorsal view C right hind wing in ventral view D abdominal segments VII-X in ventral view." figureDoi="10.3897/zookeys.898.47118.figure10" httpUri="https://binary.pensoft.net/fig/363726" pageId="0" pageNumber="1" tableDoi="10.3897/zookeys.898.47118.figure10">Fig. 10A</figureCitation>
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).
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</paragraph>
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<caption doi="10.3897/zookeys.898.47118.figure10" httpUri="https://binary.pensoft.net/fig/363726" pageId="0" pageNumber="1" start="Figure 10" startId="F10">
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" pageId="0" pageNumber="1">Figure 10.</emphasis>
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<taxonomicName class="Insecta" family="Metretopodidae" genus="Siphloplecton" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Siphloplecton studemannae" order="Ephemeroptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="studemannae">
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<emphasis italics="true" pageId="0" pageNumber="1">Siphloplecton studemannae</emphasis>
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</taxonomicName>
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sp. nov., MNHK, MP/1626, paratype, female imago (photographs)
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<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
|
||
general ventral view
|
||
<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
|
||
left forewing in dorsal view
|
||
<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
|
||
right hind wing in ventral view
|
||
<emphasis bold="true" pageId="0" pageNumber="1">D</emphasis>
|
||
abdominal segments
|
||
<normalizedToken originalValue="VII–X">VII-X</normalizedToken>
|
||
in ventral view.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
Details of mesonotum hardly visible; mesonotal suture typical for
|
||
<taxonomicName class="Insecta" family="Metretopodidae" genus="Siphloplecton" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Siphloplecton" order="Ephemeroptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Siphloplecton</emphasis>
|
||
</taxonomicName>
|
||
, original lateroparapsidal suture colouration invisible due to resin influx. Furcasternal protuberances contiguous, distinctly brown; other parts of mesosternum paler.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Pterostigma</emphasis>
|
||
with anastomosed crossveins. Cubital field of right forewing well preserved; iCu1 basally connected to CuA and iCu2 by crossveins, 2 additional CuA-iCu1 crossveins present. Pair of intercalaries iCu2 and iCu3 basally connected to each other; further crossveins connect to CuA and CuP (
|
||
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Siphloplecton studemannae sp. nov., MNHK, MP / 1626, paratype, female imago (photographs) A general ventral view B left forewing in dorsal view C right hind wing in ventral view D abdominal segments VII-X in ventral view." figureDoi="10.3897/zookeys.898.47118.figure10" httpUri="https://binary.pensoft.net/fig/363726" pageId="0" pageNumber="1" tableDoi="10.3897/zookeys.898.47118.figure10">Figs 10B</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Siphloplecton studemannae sp. nov., MNHK, MP / 1626, paratype, female imago (line drawings) A fragments of left fore- and hind wings, dorsal view B abdominal segments VII-X in ventrolateral view." figureDoi="10.3897/zookeys.898.47118.figure11" httpUri="https://binary.pensoft.net/fig/363727" pageId="0" pageNumber="1" tableDoi="10.3897/zookeys.898.47118.figure11">11A</figureCitation>
|
||
). Hind wings relatively long, approximately 0.40
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
of length of preserved right forewing; costal processes hardly visible. Hind wings with triads RS, MA and MP; costal process (visible only on right hind wing) small and blunt (
|
||
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Siphloplecton studemannae sp. nov., MNHK, MP / 1626, paratype, female imago (photographs) A general ventral view B left forewing in dorsal view C right hind wing in ventral view D abdominal segments VII-X in ventral view." figureDoi="10.3897/zookeys.898.47118.figure10" httpUri="https://binary.pensoft.net/fig/363726" pageId="0" pageNumber="1" tableDoi="10.3897/zookeys.898.47118.figure10">Fig. 10C</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption doi="10.3897/zookeys.898.47118.figure11" httpUri="https://binary.pensoft.net/fig/363727" pageId="0" pageNumber="1" start="Figure 11" startId="F11">
|
||
<paragraph pageId="0" pageNumber="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="1">Figure 11.</emphasis>
|
||
<taxonomicName class="Insecta" family="Metretopodidae" genus="Siphloplecton" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Siphloplecton studemannae" order="Ephemeroptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="studemannae">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Siphloplecton studemannae</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., MNHK, MP/1626, paratype, female imago (line drawings)
|
||
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
|
||
fragments of left fore- and hind wings, dorsal view
|
||
<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
|
||
abdominal segments
|
||
<normalizedToken originalValue="VII–X">VII-X</normalizedToken>
|
||
in ventrolateral view.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Left</emphasis>
|
||
foreleg intensely brown to blackish distally; middle legs paler, yellow to yellowish-brown. Preserved foreclaw dissimilar (one claw hooked, one claw blunt).
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
Most of
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">abdominal</emphasis>
|
||
segments ventrally covered with
|
||
<normalizedToken originalValue="“Verlumung”">"Verlumung"</normalizedToken>
|
||
. Preserved sterna paler than terga. Subgenital plate not elongate, approximately 1.80
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
wide as long, widely rounded apically. Subanal plate not elongate, pointed at tip (
|
||
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Siphloplecton studemannae sp. nov., MNHK, MP / 1626, paratype, female imago (photographs) A general ventral view B left forewing in dorsal view C right hind wing in ventral view D abdominal segments VII-X in ventral view." figureDoi="10.3897/zookeys.898.47118.figure10" httpUri="https://binary.pensoft.net/fig/363726" pageId="0" pageNumber="1" tableDoi="10.3897/zookeys.898.47118.figure10">Figs 10D</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Siphloplecton studemannae sp. nov., MNHK, MP / 1626, paratype, female imago (line drawings) A fragments of left fore- and hind wings, dorsal view B abdominal segments VII-X in ventrolateral view." figureDoi="10.3897/zookeys.898.47118.figure11" httpUri="https://binary.pensoft.net/fig/363727" pageId="0" pageNumber="1" tableDoi="10.3897/zookeys.898.47118.figure11">11B</figureCitation>
|
||
). Paracercus 5-segmented.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="1" type="comments">
|
||
<paragraph pageId="0" pageNumber="1">Comments.</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
We allocate
|
||
<taxonomicName class="Insecta" family="Metretopodidae" genus="Siphloplecton" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Siphloplecton studemannae" order="Ephemeroptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="studemannae">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Siphloplecton studemannae</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. within the
|
||
<taxonomicName lsidName="S. jaegeri" pageId="0" pageNumber="1" rank="species" species="jaegeri">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">S. jaegeri</emphasis>
|
||
</taxonomicName>
|
||
species group based on the presence of three distinctive intercalaries (grouped in one pair and one additional intercalary vein) in the cubital field of both forewings. The same arrangement of intercalary veins was described and figured for the holotype of
|
||
<taxonomicName lsidName="S. jaegeri" pageId="0" pageNumber="1" rank="species" species="jaegeri">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">S. jaegeri</emphasis>
|
||
</taxonomicName>
|
||
by
|
||
<bibRefCitation author="Staniczek, AH" journalOrPublisher="Palaeodiversity" pageId="0" pageNumber="1" pagination="57 - 87" refId="B6" refString="Staniczek, AH, Godunko, RJ, 2012. Revision of fossil Metretopodidae (Insecta: Ephemeroptera) in Baltic amber. - Part 1: Type specimens of hitherto described fossil species of Siphloplecton Clemens, 1915 and Metretopus Eaton, 1901, with description of four new fossil species of Siphloplecton. Palaeodiversity 5: 57 - 87" title="Revision of fossil Metretopodidae (Insecta: Ephemeroptera) in Baltic amber. - Part 1: Type specimens of hitherto described fossil species of Siphloplecton Clemens, 1915 and Metretopus Eaton, 1901, with description of four new fossil species of Siphloplecton." volume="5" year="2012">Staniczek and Godunko (2012</bibRefCitation>
|
||
: 73, 74, fig. 10b). Other diagnostic characters of this species group, namely the presence of sharply pointed setae at the outer margin of the foretibia, cannot be observed in the type specimens of
|
||
<taxonomicName lsidName="S. studemannae" pageId="0" pageNumber="1" rank="species" species="studemannae">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">S. studemannae</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. So it cannot be excluded that this character is present in the females of this fossil species, similarly to the different occurrence of this character in the two sexes of extant species (
|
||
<bibRefCitation author="Berner, L" pageId="0" pageNumber="1" refId="B1" refString="Berner, L, 1978. A review of the family Metretopodidae. Transactions of the American Entomological Society 104: 91−137." title="A review of the family Metretopodidae. Transactions of the American Entomological Society 104: 91 − 137." year="1978">Berner 1978</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
The new species can be characterized by the presence of the following characters: (1) eyes separated, but close-set; (2) cubital field of forewings with one pair of intercalaries and one intercalary vein connected with CuA; (3) subgenital plate relatively broad with width/length ratio 1.80−2.00. This combination of characters definitely separates the described females of
|
||
<taxonomicName lsidName="S. studemannae" pageId="0" pageNumber="1" rank="species" species="studemannae">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">S. studemannae</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. from all other previously known Recent and fossil taxa.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
<taxonomicName class="Insecta" family="Metretopodidae" genus="Siphloplecton" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Siphloplecton studemannae" order="Ephemeroptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="studemannae">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Siphloplecton studemannae</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. is currently the only species of the genus
|
||
<taxonomicName class="Insecta" family="Metretopodidae" genus="Siphloplecton" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Siphloplecton" order="Ephemeroptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Siphloplecton</emphasis>
|
||
</taxonomicName>
|
||
described from female specimens that can also be attributed to a certain species group. This is only possible due to the unique arrangement of cubital intercalaries, which is characteristic for the
|
||
<taxonomicName lsidName="S. jaegeri" pageId="0" pageNumber="1" rank="species" species="jaegeri">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">S. jaegeri</emphasis>
|
||
</taxonomicName>
|
||
species group. The previously described
|
||
<taxonomicName lsidName="S. barabani" pageId="0" pageNumber="1" rank="species" species="barabani">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">S. barabani</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName lsidName="S. hageni" pageId="0" pageNumber="1" rank="species" species="hageni">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">S. hageni</emphasis>
|
||
</taxonomicName>
|
||
, also only known from female specimens, do not provide any clear distinguishing characters that would allow their placement in one of the other fossil species groups defined for
|
||
<taxonomicName class="Insecta" family="Metretopodidae" genus="Siphloplecton" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Siphloplecton" order="Ephemeroptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Siphloplecton</emphasis>
|
||
</taxonomicName>
|
||
, which can only be grouped based on similarities in the male genitalia.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="1" type="etymology">
|
||
<paragraph pageId="0" pageNumber="1">Etymology.</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
Following our tradition of naming new fossil species of
|
||
<taxonomicName class="Insecta" family="Metretopodidae" genus="Siphloplecton" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Siphloplecton" order="Ephemeroptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Siphloplecton</emphasis>
|
||
</taxonomicName>
|
||
after Swiss ephemeropterists, this species is named after Denise Studemann, Lausanne, to honour her contributions to the knowledge of Swiss mayflies.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |