treatments-xml/data/BD/73/87/BD7387D9FFE95125FF07FF45F8E3FE60.xml
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<document id="CECB5D821C0872727A6868FB98840562" ID-CLB-Dataset="31846" ID-DOI="10.11646/zootaxa.4330.1.1" ID-GBIF-Dataset="4603132a-4e96-4119-bf2f-21ffe40dce69" ID-ISSN="1175-5326" ID-Zenodo-Dep="1010167" ID-ZooBank="Bf4D6091-5346-42Fd-8F64-D8E5Ca407415" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1507885096493" checkinUser="plazi" docAuthor="Häckel, Martin" docDate="2017" docId="BD7387D9FFE95125FF07FF45F8E3FE60" docLanguage="en" docName="zootaxa.4330.1.1.pdf.imf" docOrigin="Zootaxa 4330 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Craspedophorus latifrons" docType="treatment" docVersion="11" lastPageNumber="41" masterDocId="414AFFA1FFCE510DFF90FFDDFC0BFFE7" masterDocTitle="A contribution to the knowledge of the subfamily Panagaeinae Hope, 1838 from Africa. Part 3. Revision of the Craspedophorus strachani and C. brevicollis groups (Coleoptera: Carabidae)" masterLastPageNumber="67" masterPageNumber="1" pageNumber="40" updateTime="1698469524835" updateUser="ExternalLinkService">
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<mods:title id="CB28FD69CA7D4F1A1479A5938FCC0F89">A contribution to the knowledge of the subfamily Panagaeinae Hope, 1838 from Africa. Part 3. Revision of the Craspedophorus strachani and C. brevicollis groups (Coleoptera: Carabidae)</mods:title>
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<mods:date id="8D2EE853F08190F535080BABA552EABE">2017</mods:date>
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<treatment id="BD7387D9FFE95125FF07FF45F8E3FE60" ID-DOI="http://doi.org/10.5281/zenodo.6019501" ID-GBIF-Taxon="135435405" ID-Zenodo-Dep="6019501" LSID="urn:lsid:plazi:treatment:BD7387D9FFE95125FF07FF45F8E3FE60" httpUri="http://treatment.plazi.org/id/BD7387D9FFE95125FF07FF45F8E3FE60" lastPageId="40" lastPageNumber="41" pageId="39" pageNumber="40">
<subSubSection id="7DC06544FFE9512AFF07FF45FEC9FF55" box="[151,706,151,178]" pageId="39" pageNumber="40" type="nomenclature">
<paragraph id="356536CFFFE9512AFF07FF45FEC9FF55" blockId="39.[151,722,151,211]" box="[151,706,151,178]" pageId="39" pageNumber="40">
<heading id="6E2D81A3FFE9512AFF07FF45FEC9FF55" bold="true" box="[151,706,151,178]" fontSize="11" level="1" pageId="39" pageNumber="40" reason="1">
<emphasis id="07AEEADDFFE9512AFF07FF45FEC9FF55" bold="true" box="[151,706,151,178]" pageId="39" pageNumber="40">
3.
<taxonomicName id="F2DA4D4CFFE9512AFF24FF4AFEC9FF55" ID-CoL="Z5WT" authority="(Chaudoir, 1876)" baseAuthorityName="Chaudoir" baseAuthorityYear="1876" box="[180,706,151,178]" class="Insecta" family="Carabidae" genus="Craspedophorus" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="latifrons">
<emphasis id="07AEEADDFFE9512AFF24FF4AFDE2FF56" bold="true" box="[180,489,151,177]" italics="true" pageId="39" pageNumber="40">Craspedophorus latifrons</emphasis>
(
<bibRefCitation id="514B4B3EFFE9512AFE68FF45FEB1FF55" author="Chaudoir" box="[504,698,152,178]" pageId="39" pageNumber="40" refString="Chaudoir, M. de (1876) Catalogue des Cicindeletes et des Carabiques recueillis par Mr. Achille Raffray en Abyssinie, avec la description des especes nouvelles. Revue et Magasin de Zoologie Pure et Appliquee, Series 3, 4, 329 - 388." type="journal article" year="1876">Chaudoir, 1876</bibRefCitation>
)
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="7DC06544FFE9512AFF07FF67FED9FF34" box="[151,722,186,211]" pageId="39" pageNumber="40" type="description">
<paragraph id="356536CFFFE9512AFF07FF67FED9FF34" blockId="39.[151,722,151,211]" box="[151,722,186,211]" pageId="39" pageNumber="40">
(
<figureCitation id="ADE12A4AFFE9512AFF0EFF67FCF6FF34" box="[158,253,186,211]" captionStart="PLATE 15" captionStartId="60.[151,234,871,893]" captionTargetBox="[340,1244,199,838]" captionTargetId="figure@60.[340,1247,193,850]" captionTargetPageId="60" captionText="PLATE 15. Habitus of Craspedophorus brevicollis group. (Scale bar: 10 mm). 119 a. C. brevicollis brevicollis (Dejean, 1831): a) male, Senegal; b) male, Senegal (lectotype established by Chaudoir, 1861); 120. C. brevicollis brevicollis (Dejean, 1831), female, Kankan, Guinea; 121. C. brevicollis brevicollis (Dejean, 1831) trans ad C. brevicollis beninensis new subspecies: a) male, northern Benin, b) female, Burkina Faso; 122. C. brevicollis beninensis new subspecies, male, southern Benin; 123. C. latifrons (Chaudoir, 1876): a) male, “ Abyssinie, A. Raffray, Voy. 1881 ” (= Etiopia, MNHN), b) male, Boran Negele, Oromiya, Ethiopia; 124. C. latifrons (Chaudoir, 1876): a) female (Collectio Plason, MNHN), b) female, Moyale, Oromiya, Ethiopia; 125. C. abnormis Bates, 1886, HT (male); 126. C. abnormis Bates, 1886, female, Shurugwi District, Midlands, Zimbabwe (labeled by Péringuey as Eudema sexmaculatum Péringuey, 1898); 127. C. abnormis Bates, 1886, male, Iringa, Tanzania; 128. C. abnormis Bates, 1886, female, Mbeya, Tanzania." httpUri="https://zenodo.org/record/1010199/files/figure.png" pageId="39" pageNumber="40">Plate 15</figureCitation>
, Figs 123a, b; 124a, b,
<figureCitation id="ADE12A4AFFE9512AFD92FF67FE54FF34" box="[514,607,186,211]" captionStart="PLATE 16" captionStartId="60.[151,234,1580,1602]" captionTargetBox="[359,1235,1198,1551]" captionTargetId="figure@60.[340,1247,1188,1559]" captionTargetPageId="60" captionText="PLATE 16. Craspedophorus brevicollis group, aedeagus in frontal and right lateral views: 129. C. brevicollis brevicollis (Dejean, 1831), Senegal; 130. C. brevicollis brevicollis (Dejean, 1831) trans ad C. b. beninensis new subspecies, Sina-Issire, northern Benin; 131. C. abnormis Bates, 1886, Iringa, Tanzania; 132. C. latifrons (Chaudoir, 1876), Boran Negele, Oromiya, Ethiopia." httpUri="https://zenodo.org/record/1010201/files/figure.png" pageId="39" pageNumber="40">Plate 16</figureCitation>
, Fig. 132)
</paragraph>
</subSubSection>
<subSubSection id="7DC06544FFE9512AFF07FEDFFFD1FEB1" pageId="39" pageNumber="40" type="reference_group">
<paragraph id="356536CFFFE9512AFF07FEDFF892FED0" blockId="39.[151,1436,258,342]" pageId="39" pageNumber="40">
<treatmentCitationGroup id="15CA11E1FFE9512AFF07FEDFF892FED0" pageId="39" pageNumber="40">
<taxonomicName id="F2DA4D4CFFE9512AFF07FEDFFE20FEFF" authority="Chaudoir, 1876: 354" authorityName="Chaudoir" authorityPageNumber="354" authorityYear="1876" box="[151,555,258,280]" class="Magnoliopsida" family="Brassicaceae" genus="Eudema" kingdom="Plantae" order="Brassicales" pageId="39" pageNumber="40" phylum="Tracheophyta" rank="species" species="latifrons">
<emphasis id="07AEEADDFFE9512AFF07FEDFFD42FEFF" box="[151,329,258,280]" italics="true" pageId="39" pageNumber="40">Eudema latifrons</emphasis>
<treatmentCitation id="B47B10DEFFE9512AFEC3FEDFFE20FEFF" author="Chaudoir" box="[339,555,258,280]" page="354" pageId="39" pageNumber="40" year="1876">
<bibRefCitation id="514B4B3EFFE9512AFEC3FEDFFDFCFEFF" author="Chaudoir" box="[339,503,258,280]" pageId="39" pageNumber="40" refString="Chaudoir, M. de (1876) Catalogue des Cicindeletes et des Carabiques recueillis par Mr. Achille Raffray en Abyssinie, avec la description des especes nouvelles. Revue et Magasin de Zoologie Pure et Appliquee, Series 3, 4, 329 - 388." type="journal article" year="1876">Chaudoir, 1876</bibRefCitation>
: 354
</treatmentCitation>
</taxonomicName>
(type locality “Adowa en Abyssinie [=Adwa, Tigray, Ethiopia]”).
<taxonomicName id="F2DA4D4CFFE9512AFB63FEDFFDC3FED0" ID-CoL="Z5WT" authority="Alluaud 1930: 2" authorityName="Alluaud" authorityPageNumber="2" authorityYear="1930" class="Insecta" family="Carabidae" genus="Craspedophorus" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="latifrons">
<emphasis id="07AEEADDFFE9512AFB63FEDFFD17FED0" italics="true" pageId="39" pageNumber="40">Craspedophorus latifrons</emphasis>
<treatmentCitation id="B47B10DEFFE9512AFEB2FEFCFDC3FED0" author="Alluaud" box="[290,456,289,311]" page="2" pageId="39" pageNumber="40" year="1930">
<bibRefCitation id="514B4B3EFFE9512AFEB2FEFCFDBBFED0" author="Alluaud" box="[290,432,289,311]" pageId="39" pageNumber="40" refString="Alluaud, C. (1930) Note sur les Panageides dAfrique tropicale. Afra, Cahiers d'Entomologie, 1, 1 - 9." type="journal article" year="1930">Alluaud 1930</bibRefCitation>
: 2
</treatmentCitation>
</taxonomicName>
,
<treatmentCitation id="B47B10DEFFE9512AFE43FEFCFEA1FED0" author="Basilewsky" box="[467,682,289,311]" page="84" pageId="39" pageNumber="40" year="1978">
<bibRefCitation id="514B4B3EFFE9512AFE43FEFCFE8EFED0" author="Basilewsky" box="[467,645,289,311]" pageId="39" pageNumber="40" refString="Basilewsky, P. (1978) Nouvelle contribution r l'etude des coleopteres Carabidae du Nord-Yemen. Bulletin et Annales de la Societe Royale Belge d'Entomologie, 114, 77 - 86." type="journal article" year="1978">Basilewsky 1978</bibRefCitation>
: 84
</treatmentCitation>
,
<treatmentCitation id="B47B10DEFFE9512AFD24FEFFFF60FED0" author="Lorenz" box="[692,875,289,311]" page="320" pageId="39" pageNumber="40" year="2005">
<bibRefCitation id="514B4B3EFFE9512AFD24FEFFFF33FED0" author="Lorenz" box="[692,824,289,311]" pageId="39" pageNumber="40" refString="Lorenz, W. (2005) A systematic list of extant ground beetles of the World (Insecta, Coleoptera, Adephaga: Trachypachidae &amp; Carabidae incl. Paussinae, Cicindelinae, Rhysodinae). 2 nd Edition. Published by the author, Tutzing, 530 pp." type="book" year="2005">Lorenz 2005</bibRefCitation>
: 320
</treatmentCitation>
.
<treatmentCitation id="B47B10DEFFE9512AFCE5FEFCF89EFED0" author="Hackel" box="[885,1173,289,311]" page="81" pageId="39" pageNumber="40" year="2012">
<bibRefCitation id="514B4B3EFFE9512AFCE5FEFCF87BFED0" author="Hackel" box="[885,1136,289,311]" pageId="39" pageNumber="40" refString="Hackel, M. &amp; Farkac, J. (2012) A check-list of the subfamily Panagaeinae Hope, 1838 of the World (Coleoptera: Carabidae). Studies and Reports, Taxonomical Series, 8 (1 - 2), 67 - 116." type="journal article" year="2012">Häckel and Farkač 2012</bibRefCitation>
: 81
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="356536CFFFE9512AFF07FE9DFFD1FEB1" blockId="39.[151,1436,258,342]" box="[151,986,320,342]" pageId="39" pageNumber="40">
<treatmentCitationGroup id="15CA11E1FFE9512AFF07FE9DFFD1FEB1" box="[151,986,320,342]" pageId="39" pageNumber="40">
<taxonomicName id="F2DA4D4CFFE9512AFF07FE9DFE60FEB1" ID-CoL="Z5WN" authority="Chaudoir, 1879: 103" authorityName="Chaudoir" authorityPageNumber="103" authorityYear="1879" box="[151,619,320,342]" class="Insecta" family="Carabidae" genus="Craspedophorus" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="laticeps">
<emphasis id="07AEEADDFFE9512AFF07FE9DFD9EFEB1" box="[151,405,320,342]" italics="true" pageId="39" pageNumber="40">Craspedophorus laticeps</emphasis>
<treatmentCitation id="B47B10DEFFE9512AFE0BFE9DFE60FEB1" author="Chaudoir" box="[411,619,320,342]" page="103" pageId="39" pageNumber="40" year="1879">
<bibRefCitation id="514B4B3EFFE9512AFE0BFE9DFE30FEB1" author="Chaudoir" box="[411,571,320,342]" pageId="39" pageNumber="40" refString="Chaudoir, M. de (1879) Essai monographie sur les Panageides (part II). Annales de la Societe Entomologique des Belgique, 21, 85 - 186." type="journal article" year="1879">Chaudoir, 1879</bibRefCitation>
: 103
</treatmentCitation>
</taxonomicName>
by error.
<treatmentCitation id="B47B10DEFFE9512AFD5DFE9DFF78FEB1" author="Alluaud" box="[717,883,320,342]" page="2" pageId="39" pageNumber="40" year="1930">
<bibRefCitation id="514B4B3EFFE9512AFD5DFE9DFF50FEB1" author="Alluaud" box="[717,859,320,342]" pageId="39" pageNumber="40" refString="Alluaud, C. (1930) Note sur les Panageides dAfrique tropicale. Afra, Cahiers d'Entomologie, 1, 1 - 9." type="journal article" year="1930">Alluaud 1930</bibRefCitation>
: 2
</treatmentCitation>
, syn. nov.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="7DC06544FFE95125FF07FE5FF826FE82" lastPageId="40" lastPageNumber="41" pageId="39" pageNumber="40" type="materials_examined">
<paragraph id="356536CFFFE9512AFF07FE5FFD46FDA8" blockId="39.[151,1437,386,2037]" pageId="39" pageNumber="40">
<emphasis id="07AEEADDFFE9512AFF07FE5FFD80FE7C" bold="true" box="[151,395,386,411]" pageId="39" pageNumber="40">Compared material.</emphasis>
1♂: “Abyssinie / A. Raffray / Voy. 1881. [printed in black on white label]” (
<figureCitation id="ADE12A4AFFE9512AFA91FE5FF969FE7C" box="[1281,1378,386,411]" captionStart="PLATE 15" captionStartId="60.[151,234,871,893]" captionTargetBox="[340,1244,199,838]" captionTargetId="figure@60.[340,1247,193,850]" captionTargetPageId="60" captionText="PLATE 15. Habitus of Craspedophorus brevicollis group. (Scale bar: 10 mm). 119 a. C. brevicollis brevicollis (Dejean, 1831): a) male, Senegal; b) male, Senegal (lectotype established by Chaudoir, 1861); 120. C. brevicollis brevicollis (Dejean, 1831), female, Kankan, Guinea; 121. C. brevicollis brevicollis (Dejean, 1831) trans ad C. brevicollis beninensis new subspecies: a) male, northern Benin, b) female, Burkina Faso; 122. C. brevicollis beninensis new subspecies, male, southern Benin; 123. C. latifrons (Chaudoir, 1876): a) male, “ Abyssinie, A. Raffray, Voy. 1881 ” (= Etiopia, MNHN), b) male, Boran Negele, Oromiya, Ethiopia; 124. C. latifrons (Chaudoir, 1876): a) female (Collectio Plason, MNHN), b) female, Moyale, Oromiya, Ethiopia; 125. C. abnormis Bates, 1886, HT (male); 126. C. abnormis Bates, 1886, female, Shurugwi District, Midlands, Zimbabwe (labeled by Péringuey as Eudema sexmaculatum Péringuey, 1898); 127. C. abnormis Bates, 1886, male, Iringa, Tanzania; 128. C. abnormis Bates, 1886, female, Mbeya, Tanzania." httpUri="https://zenodo.org/record/1010199/files/figure.png" pageId="39" pageNumber="40">Plate 15</figureCitation>
, Fig. 123a, MNHN, Oberthür/Bates Collection). 3♂, 2♀ same labeled as previous (MNHN, Oberthür/Bates Collection). 1♂: “Abyssinie / Raffray [handwritten in black on white label]//
<taxonomicName id="F2DA4D4CFFE9512AFCE7FE17FFD8FE04" authorityName="Laporte de Castelnau" authorityYear="1840" box="[887,979,458,483]" class="Magnoliopsida" family="Brassicaceae" genus="Eudema" kingdom="Plantae" order="Brassicales" pageId="39" pageNumber="40" phylum="Tracheophyta" rank="genus">
<emphasis id="07AEEADDFFE9512AFCE7FE17FFD8FE04" box="[887,979,458,483]" italics="true" pageId="39" pageNumber="40">Eudema</emphasis>
</taxonomicName>
/
<taxonomicName id="F2DA4D4CFFE9512AFC7BFE17F88FFE04" authority="Chd." authorityName="Chd." box="[1003,1156,458,483]" class="Insecta" family="Carabidae" genus="Craspedophorus" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="latifrons">
<emphasis id="07AEEADDFFE9512AFC7BFE17F842FE04" box="[1003,1097,458,483]" italics="true" pageId="39" pageNumber="40">latifrons</emphasis>
Chd.
</taxonomicName>
[handwritten in black on white label]// Museum Paris / 1935 / Coll. M. Sédillot [printed in black on blue label]//
<taxonomicName id="F2DA4D4CFFE9512AFB5CFE32F997FDE0" authority="Chd." authorityName="Chd." box="[1228,1436,494,519]" class="Insecta" family="Carabidae" genus="Craspedophorus" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="latifrons">
<emphasis id="07AEEADDFFE9512AFB5CFE32F952FDE0" box="[1228,1369,494,519]" italics="true" pageId="39" pageNumber="40">Cr. latifrons</emphasis>
Chd.
</taxonomicName>
[handwritten in black on white label]/ P. Basilewsky det., 19[printed in black]54[handwritten in black on white label] (MNHN).
</paragraph>
<paragraph id="356536CFFFE9512AFF57FD87FE5DFC88" blockId="39.[151,1437,386,2037]" pageId="39" pageNumber="40">
<emphasis id="07AEEADDFFE9512AFF57FD87FE32FD94" bold="true" box="[199,569,602,627]" pageId="39" pageNumber="40">Additional material examined.</emphasis>
1♀ erroneously labeled as
<taxonomicName id="F2DA4D4CFFE9512AFCE9FD87F837FD94" box="[889,1084,602,627]" class="Magnoliopsida" family="Brassicaceae" genus="Eudema" kingdom="Plantae" order="Brassicales" pageId="39" pageNumber="40" phylum="Tracheophyta" rank="species" species="festivum">
<emphasis id="07AEEADDFFE9512AFCE9FD87F837FD94" box="[889,1084,602,627]" italics="true" pageId="39" pageNumber="40">Eudema festivum</emphasis>
</taxonomicName>
by unknown author and as
<taxonomicName id="F2DA4D4CFFE9512AFA12FD86FD3CFD70" authority="Klug." authorityName="Klug." class="Insecta" family="Carabidae" genus="Panagaeus" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="festivus">
<emphasis id="07AEEADDFFE9512AFA12FD86FCE0FD70" italics="true" pageId="39" pageNumber="40">C. festivus</emphasis>
Klug.
</taxonomicName>
by Kirschenhofer 1999: “Collect. Plason” (
<figureCitation id="ADE12A4AFFE9512AFCD2FDA3FFADFD70" box="[834,934,638,663]" captionStart="PLATE 15" captionStartId="60.[151,234,871,893]" captionTargetBox="[340,1244,199,838]" captionTargetId="figure@60.[340,1247,193,850]" captionTargetPageId="60" captionText="PLATE 15. Habitus of Craspedophorus brevicollis group. (Scale bar: 10 mm). 119 a. C. brevicollis brevicollis (Dejean, 1831): a) male, Senegal; b) male, Senegal (lectotype established by Chaudoir, 1861); 120. C. brevicollis brevicollis (Dejean, 1831), female, Kankan, Guinea; 121. C. brevicollis brevicollis (Dejean, 1831) trans ad C. brevicollis beninensis new subspecies: a) male, northern Benin, b) female, Burkina Faso; 122. C. brevicollis beninensis new subspecies, male, southern Benin; 123. C. latifrons (Chaudoir, 1876): a) male, “ Abyssinie, A. Raffray, Voy. 1881 ” (= Etiopia, MNHN), b) male, Boran Negele, Oromiya, Ethiopia; 124. C. latifrons (Chaudoir, 1876): a) female (Collectio Plason, MNHN), b) female, Moyale, Oromiya, Ethiopia; 125. C. abnormis Bates, 1886, HT (male); 126. C. abnormis Bates, 1886, female, Shurugwi District, Midlands, Zimbabwe (labeled by Péringuey as Eudema sexmaculatum Péringuey, 1898); 127. C. abnormis Bates, 1886, male, Iringa, Tanzania; 128. C. abnormis Bates, 1886, female, Mbeya, Tanzania." httpUri="https://zenodo.org/record/1010199/files/figure.png" pageId="39" pageNumber="40">Plate 15</figureCitation>
, Fig. 124a, MNHN).
<emphasis id="07AEEADDFFE9512AFB3DFDA3F997FD70" bold="true" box="[1197,1436,638,663]" pageId="39" pageNumber="40">Ethiopia: Oromiya.</emphasis>
3♂,1♀: “Oromiya Reg.: Boran Negele, 5°13'20” N 39°38'10” E,
<quantity id="F2229B2AFFE9512AFCE6FD7FFFC6FD5C" box="[886,973,674,699]" metricMagnitude="3" metricUnit="m" metricValue="1.538" pageId="39" pageNumber="40" unit="m" value="1538.0">1538 m</quantity>
” (
<figureCitation id="ADE12A4AFFE9512AFC77FD7FF84DFD5C" box="[999,1094,674,699]" captionStart="PLATE 15" captionStartId="60.[151,234,871,893]" captionTargetBox="[340,1244,199,838]" captionTargetId="figure@60.[340,1247,193,850]" captionTargetPageId="60" captionText="PLATE 15. Habitus of Craspedophorus brevicollis group. (Scale bar: 10 mm). 119 a. C. brevicollis brevicollis (Dejean, 1831): a) male, Senegal; b) male, Senegal (lectotype established by Chaudoir, 1861); 120. C. brevicollis brevicollis (Dejean, 1831), female, Kankan, Guinea; 121. C. brevicollis brevicollis (Dejean, 1831) trans ad C. brevicollis beninensis new subspecies: a) male, northern Benin, b) female, Burkina Faso; 122. C. brevicollis beninensis new subspecies, male, southern Benin; 123. C. latifrons (Chaudoir, 1876): a) male, “ Abyssinie, A. Raffray, Voy. 1881 ” (= Etiopia, MNHN), b) male, Boran Negele, Oromiya, Ethiopia; 124. C. latifrons (Chaudoir, 1876): a) female (Collectio Plason, MNHN), b) female, Moyale, Oromiya, Ethiopia; 125. C. abnormis Bates, 1886, HT (male); 126. C. abnormis Bates, 1886, female, Shurugwi District, Midlands, Zimbabwe (labeled by Péringuey as Eudema sexmaculatum Péringuey, 1898); 127. C. abnormis Bates, 1886, male, Iringa, Tanzania; 128. C. abnormis Bates, 1886, female, Mbeya, Tanzania." httpUri="https://zenodo.org/record/1010199/files/figure.png" pageId="39" pageNumber="40">Plate 15</figureCitation>
, Fig. 123b,
<figureCitation id="ADE12A4AFFE9512AFB5DFD7FF920FD5C" box="[1229,1323,674,699]" captionStart="PLATE 16" captionStartId="60.[151,234,1580,1602]" captionTargetBox="[359,1235,1198,1551]" captionTargetId="figure@60.[340,1247,1188,1559]" captionTargetPageId="60" captionText="PLATE 16. Craspedophorus brevicollis group, aedeagus in frontal and right lateral views: 129. C. brevicollis brevicollis (Dejean, 1831), Senegal; 130. C. brevicollis brevicollis (Dejean, 1831) trans ad C. b. beninensis new subspecies, Sina-Issire, northern Benin; 131. C. abnormis Bates, 1886, Iringa, Tanzania; 132. C. latifrons (Chaudoir, 1876), Boran Negele, Oromiya, Ethiopia." httpUri="https://zenodo.org/record/1010201/files/figure.png" pageId="39" pageNumber="40">Plate 16</figureCitation>
, Fig. 132, cMH, cRS); 1♀: “
<quantity id="F2229B2AFFE9512AFEF7FD1AFDA4FD38" box="[359,431,710,735]" metricMagnitude="4" metricUnit="m" metricValue="1.1" pageId="39" pageNumber="40" unit="km" value="11.0">11 km</quantity>
n Moyale, 03°51'05”N 38°46'17”E,
<quantity id="F2229B2AFFE9512AFCC2FD1BFFA1FD38" box="[850,938,710,735]" metricMagnitude="3" metricUnit="m" metricValue="1.295" pageId="39" pageNumber="40" unit="m" value="1295.0">1295 m</quantity>
” (
<figureCitation id="ADE12A4AFFE9512AFC55FD1BF82EFD38" box="[965,1061,710,735]" captionStart="PLATE 15" captionStartId="60.[151,234,871,893]" captionTargetBox="[340,1244,199,838]" captionTargetId="figure@60.[340,1247,193,850]" captionTargetPageId="60" captionText="PLATE 15. Habitus of Craspedophorus brevicollis group. (Scale bar: 10 mm). 119 a. C. brevicollis brevicollis (Dejean, 1831): a) male, Senegal; b) male, Senegal (lectotype established by Chaudoir, 1861); 120. C. brevicollis brevicollis (Dejean, 1831), female, Kankan, Guinea; 121. C. brevicollis brevicollis (Dejean, 1831) trans ad C. brevicollis beninensis new subspecies: a) male, northern Benin, b) female, Burkina Faso; 122. C. brevicollis beninensis new subspecies, male, southern Benin; 123. C. latifrons (Chaudoir, 1876): a) male, “ Abyssinie, A. Raffray, Voy. 1881 ” (= Etiopia, MNHN), b) male, Boran Negele, Oromiya, Ethiopia; 124. C. latifrons (Chaudoir, 1876): a) female (Collectio Plason, MNHN), b) female, Moyale, Oromiya, Ethiopia; 125. C. abnormis Bates, 1886, HT (male); 126. C. abnormis Bates, 1886, female, Shurugwi District, Midlands, Zimbabwe (labeled by Péringuey as Eudema sexmaculatum Péringuey, 1898); 127. C. abnormis Bates, 1886, male, Iringa, Tanzania; 128. C. abnormis Bates, 1886, female, Mbeya, Tanzania." httpUri="https://zenodo.org/record/1010199/files/figure.png" pageId="39" pageNumber="40">Plate 15</figureCitation>
, Fig. 124b, cMH); 1♀: “Bedele” (cOH).
<emphasis id="07AEEADDFFE9512AFF61FD37F80BFCE4" bold="true" box="[241,1024,746,771]" pageId="39" pageNumber="40">Southern Nations, Nationalities, and People's Region [SNNPR].</emphasis>
1♀: “Gamo Gofa Zone,
<quantity id="F2229B2AFFE9512AFABEFD36F972FCE4" box="[1326,1401,746,771]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="39" pageNumber="40" unit="km" value="10.0">10 km</quantity>
se Konso,
<quantity id="F2229B2AFFE9512AFF7FFCD3FD4DFCC0" box="[239,326,782,807]" metricMagnitude="3" metricUnit="m" metricValue="1.15" pageId="39" pageNumber="40" unit="m" value="1150.0">1150 m</quantity>
” (cMH); 2♂,1♀: “Mago National Park, Jinka” (cIB); 1♂,1♀: “Woyto, near river” (cRK).
<emphasis id="07AEEADDFFE9512AFAD5FCD3FCE9FCAC" bold="true" pageId="39" pageNumber="40">Kenya: Coast.</emphasis>
1♀: “Coast Province, sw of Voi” (cMH); 1♀: “Tsavo NP, Sagala Hills, Voi env” (cRK); 3♂,4♀: “Gede Forest, 3°16'S 39°55'E” (NMWC, cPS).
</paragraph>
<paragraph id="356536CFFFE95125FF57FCA7F826FE82" blockId="39.[151,1437,386,2037]" lastBlockId="40.[151,1436,151,392]" lastPageId="40" lastPageNumber="41" pageId="39" pageNumber="40">
<emphasis id="07AEEADDFFE9512AFF57FCA7FD0DFC74" bold="true" box="[199,262,890,915]" pageId="39" pageNumber="40">Note.</emphasis>
This species was based on a single male specimen, labeled “Adowa en
<collectingCountry id="4DCD765FFFE9512AFBA8FCA7F8A5FC74" box="[1080,1198,890,915]" name="Ethiopia" pageId="39" pageNumber="40">Abyssinie</collectingCountry>
”. Adwa (also spelled Adowa, Aduwa) is a market town surrounded by Adwa woreda in northern
<collectingCountry id="4DCD765FFFE9512AFC7AFC43F846FC50" box="[1002,1101,926,951]" name="Ethiopia" pageId="39" pageNumber="40">Ethiopia</collectingCountry>
. Maákelay, or Mehakelegnaw (or “The central”) is a Zone in the Ethiopian Region of
<collectingRegion id="F71EF82DFFE9512AFCB6FC1FFF79FC3C" box="[806,882,962,987]" country="Ethiopia" name="Tigray" pageId="39" pageNumber="40">Tigray</collectingRegion>
. Description (in part, see
<bibRefCitation id="514B4B3EFFE9512AFB0FFC1FF95FFC3C" author="Chaudoir" box="[1183,1364,962,987]" pageId="39" pageNumber="40" refString="Chaudoir, M. de (1876) Catalogue des Cicindeletes et des Carabiques recueillis par Mr. Achille Raffray en Abyssinie, avec la description des especes nouvelles. Revue et Magasin de Zoologie Pure et Appliquee, Series 3, 4, 329 - 388." type="journal article" year="1876">Chaudoir, 1876</bibRefCitation>
: 354). “Length
<quantity id="F2229B2AFFE9512AFF6AFC3AFD46FC19" box="[250,333,999,1023]" metricMagnitude="-2" metricUnit="m" metricValue="1.8" pageId="39" pageNumber="40" unit="mm" value="18.0">18 mm</quantity>
. It resembles the species of
<taxonomicName id="F2DA4D4CFFE9512AFD1AFC3AFF0DFC19" baseAuthorityName="Hope" baseAuthorityYear="1842" box="[650,774,999,1022]" class="Insecta" family="Carabidae" genus="Craspedophorus" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="tropicus">
<emphasis id="07AEEADDFFE9512AFD1AFC3AFF0DFC19" box="[650,774,999,1022]" italics="true" pageId="39" pageNumber="40">C. tropicus</emphasis>
</taxonomicName>
… -
<taxonomicName id="F2DA4D4CFFE9512AFCABFC3AFFBAFC19" authorityName="Murray" authorityYear="1857" box="[827,945,999,1022]" class="Insecta" family="Carabidae" genus="Craspedophorus" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="conicus">
<emphasis id="07AEEADDFFE9512AFCABFC3AFFBAFC19" box="[827,945,999,1022]" italics="true" pageId="39" pageNumber="40">C. conicus</emphasis>
</taxonomicName>
… group [=
<taxonomicName id="F2DA4D4CFFE9512AFBA7FC3AF8A5FC18" box="[1079,1198,999,1023]" class="Insecta" family="Carabidae" genus="Panagaeus" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="festivus">
<emphasis id="07AEEADDFFE9512AFBA7FC3AF8A5FC18" box="[1079,1198,999,1023]" italics="true" pageId="39" pageNumber="40">C. festivus</emphasis>
</taxonomicName>
species group in this article], but wider. Mainly its head is less narrow. Head as wide as long, very abruptly strangulated behind eyes, with longitudinal depression widely and deeply impressed close to the margin between eyes, eyes more convex, hemispherical, neck not as gross as in
<taxonomicName id="F2DA4D4CFFE9512AFDCDFB8EFF89FB8C" authority="(Hope, 1842)" baseAuthorityName="Hope" baseAuthorityYear="1842" box="[605,898,1107,1131]" class="Insecta" family="Carabidae" genus="Craspedophorus" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="tropicus">
<emphasis id="07AEEADDFFE9512AFDCDFB8EFED7FB8C" box="[605,732,1107,1131]" italics="true" pageId="39" pageNumber="40">C. tropicus</emphasis>
(
<bibRefCitation id="514B4B3EFFE9512AFD7EFB8EFF71FB8C" author="Hope" box="[750,890,1107,1131]" pageId="39" pageNumber="40" refString="Hope, F. W. (1842) On some new Insects from Western Africa. The Annals and Magazine of Natural History including Zoology, Botany and Geology, 10, 91 - 93. [London]" type="journal article" year="1842">Hope, 1842</bibRefCitation>
)
</taxonomicName>
; frons and vertex punctured and covered with erected setae. Pronotum wider, anterior angles and lateral margins anteriorly distinctly more rounded, creating shell-shaped form; posteriorly margins each with an extruded dent in front of posterior angles, also more rounded, surface with equal sculpture and punctuation as that in the head. Elytra differ more distinctly in their width and interval and they are more grossly punctured. Body, antennae and legs black, as well as the setation cover; humeral elytral macula lemon yellow, consists of 6 macular spots, forming weakly oblique fascia, with margins quite serrate because of macular spots on IV and VI interval which are moved more posteriorly than the others; preapical macula as in [
<emphasis id="07AEEADDFFE9512AFEA2FA92FD4EFA81" box="[306,325,1359,1382]" italics="true" pageId="39" pageNumber="40">C</emphasis>
.]
<emphasis id="07AEEADDFFE9512AFECCFA93FDC0FA80" box="[348,459,1358,1383]" italics="true" pageId="39" pageNumber="40">laevifrons</emphasis>
(Schaum, 1863). One specimen collected in Adowa,
<collectingCountry id="4DCD765FFFE9512AFB8FFA93F899FA80" box="[1055,1170,1358,1383]" name="Ethiopia" pageId="39" pageNumber="40">Abyssinia</collectingCountry>
. The external margin of the terminal labial palpomere is very prolonged and pointed in males, distinctly longer than the internal one, in contrast to that in [
<emphasis id="07AEEADDFFE9512AFEFCFA4AFD74FA49" box="[364,383,1431,1454]" italics="true" pageId="39" pageNumber="40">C</emphasis>
.]
<emphasis id="07AEEADDFFE9512AFE07FA4BFE0DFA48" box="[407,518,1430,1455]" italics="true" pageId="39" pageNumber="40">laevifrons</emphasis>
, with terminal palpomere triangular-shaped and simply obliquely snipped” [from French]. Description of
<taxonomicName id="F2DA4D4CFFE9512AFE35FA67FEB5FA34" authorityName="Chaudoir" authorityYear="1879" box="[421,702,1466,1491]" class="Insecta" family="Carabidae" genus="Craspedophorus" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="laticeps">
<emphasis id="07AEEADDFFE9512AFE35FA67FEB5FA34" box="[421,702,1466,1491]" italics="true" pageId="39" pageNumber="40">Craspedophorus laticeps</emphasis>
</taxonomicName>
[=
<emphasis id="07AEEADDFFE9512AFD4FFA67FF45FA34" box="[735,846,1466,1491]" italics="true" pageId="39" pageNumber="40">laevifrons</emphasis>
] (in part, see
<bibRefCitation id="514B4B3EFFE9512AFC7AFA67F89DFA34" author="Chaudoir" box="[1002,1174,1466,1491]" pageId="39" pageNumber="40" refString="Chaudoir, M. de (1879) Essai monographie sur les Panageides (part II). Annales de la Societe Entomologique des Belgique, 21, 85 - 186." type="journal article" year="1879">Chaudoir 1879</bibRefCitation>
: 103). “Length
<quantity id="F2229B2AFFE9512AFAD6FA66F99CFA35" box="[1350,1431,1467,1491]" metricMagnitude="-2" metricUnit="m" metricValue="1.8" pageId="39" pageNumber="40" unit="mm" value="18.0">18 mm</quantity>
, width
<quantity id="F2229B2AFFE9512AFF4EFA02FD3CFA11" box="[222,311,1503,1527]" metricMagnitude="-3" metricUnit="m" metricValue="8.8" pageId="39" pageNumber="40" unit="mm" value="8.8">8.8 mm</quantity>
. It resembles some form of
<taxonomicName id="F2DA4D4CFFE9512AFDFFFA02FFB3FA10" authority="(Dejean, 1831)" baseAuthorityName="Dejean" baseAuthorityYear="1831" box="[623,952,1502,1527]" class="Insecta" family="Carabidae" genus="Craspedophorus" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="brevicollis">
<emphasis id="07AEEADDFFE9512AFDFFFA02FF0EFA10" box="[623,773,1502,1527]" italics="true" pageId="39" pageNumber="40">C. brevicollis</emphasis>
(
<bibRefCitation id="514B4B3EFFE9512AFC84FA03FFA4FA10" author="Dejean" box="[788,943,1502,1527]" pageId="39" pageNumber="40" refString="Dejean, P. F. M. A. (1831) Species general des coleopteres, de la collection de M. le Comte Dejean. Tome cinquieme. Mequignon-Marvis, Paris, viii + 883 pp." type="book" year="1831">Dejean, 1831</bibRefCitation>
)
</taxonomicName>
, but differs from it because alate, and in its differently shaped pronotum, which is less convex anteriorly, as well as in its elytral coloration, with fasciae less expanded. Pronotum weakly narrower, lateral margins less curved anteriorly from midlength, disc less convex, with lateral depressions less marked, and weakly protruded to basal excavations. Elytra narrower, seem more elongated, lateral margins more parallel, humeri less widely rounded; dorsum markedly less convex posteriorly, descending less rapidly to apex mainly posteriorly; striae and intervals not too similarly punctured; elytral fasciae somewhat darker, distinctly less expanded, mainly more transverse, and with more serrate margins; humeral macula reaching from II to VIII stria, consists of 6 macular spots markedly shorter, spots on VII and VIII stria [=next interval] closer towards the base than the others, and by a half more posterior than the spot on VI interval, spots on III and V interval more prolonged towards the base than the contiguous which is prolonged towards the apex; preapical macula consists of five spots, reaching from III to VIII stria, with spots on IV and VI interval twice longer than three others, and spots on VII and VIII interval, each posteriorly arranged with spot on VI interval, anteriorly prolonged by a half of its length. This species was collected in Adoua,
<collectingCountry id="4DCD765FFFE9512AFBA6F849F8A2F84A" box="[1078,1193,1940,1965]" name="Ethiopia" pageId="39" pageNumber="40">Abyssinia</collectingCountry>
, by Mr. Raffray, who gave me a single specimen,
<specimenCount id="23DCFD46FFE9512AFE47F865FDE6F837" box="[471,493,1976,2000]" pageId="39" pageNumber="40" type="male"></specimenCount>
” [from French].
<bibRefCitation id="514B4B3EFFE9512AFD3FF865FFA5F837" author="Basilewsky" box="[687,942,1976,2001]" pageId="39" pageNumber="40" refString="Basilewsky, P. (1978) Nouvelle contribution r l'etude des coleopteres Carabidae du Nord-Yemen. Bulletin et Annales de la Societe Royale Belge d'Entomologie, 114, 77 - 86." type="journal article" year="1978">Basilewsky (1978: 84)</bibRefCitation>
recorded one specimen from Wadi-Sohoul,
<collectingCountry id="4DCD765FFFE9512AFF07F800FCE3F813" box="[151,232,2013,2036]" name="Yemen" pageId="39" pageNumber="40">Yemen</collectingCountry>
,
<quantity id="F2229B2AFFE9512AFF65F800FD46F813" box="[245,333,2013,2037]" metricMagnitude="3" metricUnit="m" metricValue="2.1" pageId="39" pageNumber="40" unit="m" value="2100.0">2100 m</quantity>
a.s.l. According to Basilewsky's map (1978: 78) this wadi is located south of Yarim, a town in southwestern
<collectingCountry id="4DCD765FFFE65125FEAFFF45FD9AFF48" box="[319,401,152,175]" name="Yemen" pageId="40" pageNumber="41">Yemen</collectingCountry>
. Basilewsky's note represents the first and still the only record of Afrotropical
<taxonomicName id="F2DA4D4CFFE65125FF07FF61FD59FF32" authorityName="Häckel" authorityYear="2017" box="[151,338,188,213]" class="Insecta" family="Carabidae" genus="Craspedophorus" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="genus">
<emphasis id="07AEEADDFFE65125FF07FF61FD59FF32" box="[151,338,188,213]" italics="true" pageId="40" pageNumber="41">Craspedophorus</emphasis>
</taxonomicName>
species outside the African continent. Yet it is not the first species with such distribution, and this record only supports a different concept of Afrotropical region than that presented in Palearctic Catalogue (
<bibRefCitation id="514B4B3EFFE65125FAC7FF02FCD4FEFA" author="Baehr" pageId="40" pageNumber="41" refString="Baehr, M. (2003 b) Tribe Panagaeini. In: Lobl, I. &amp; Smetana, A. (Eds.), Catalogue of Palearctic Coleoptera. Fol. 1. Apollo Books, Stenstrup, pp. 446 - 448." type="journal article" year="2003" yearSuffix="b">Baehr 2003b</bibRefCitation>
,
<bibRefCitation id="514B4B3EFFE65125FF7AFED9FD76FEFB" author="Hackel" box="[234,381,260,285]" pageId="40" pageNumber="41" refString="Hackel, M. (2017) A contribution to the knowledge of the subfamily Panagaeinae Hope, 1838 from Africa. Part 2. Revision of the Craspedophorus leprieuri and C. regalis groups (Coleoptera: Carabidae). Zootaxa, 4236 (2), 201 - 243. https: // doi. org / 10.11646 / zootaxa. 4236.2.1" type="journal article" year="2017">Häckel 2017</bibRefCitation>
). The Palearctic catalogue includes the southern portion of the Arabian peninsula in the region, whereas other publications include at least
<collectingCountry id="4DCD765FFFE65125FD12FEF5FEDBFED8" box="[642,720,296,319]" name="Yemen" pageId="40" pageNumber="41">Yemen</collectingCountry>
and
<collectingCountry id="4DCD765FFFE65125FC9AFEF5FF44FEA7" box="[778,847,296,320]" name="Oman" pageId="40" pageNumber="41">Oman</collectingCountry>
either in the Afrotropical region or in a transitional zone between the Palearctic, Afrotropical and
<collectingRegion id="F71EF82DFFE65125FD0EFE91FEF1FE82" box="[670,762,332,357]" country="Oman" name="Shamal ash Sharqiyah" pageId="40" pageNumber="41">Oriental</collectingRegion>
regions (Häckel 2012: 58).
</paragraph>
</subSubSection>
<subSubSection id="7DC06544FFE65125FF57FEB2F8E3FE60" box="[199,1256,367,392]" pageId="40" pageNumber="41" type="distribution">
<paragraph id="356536CFFFE65125FF57FEB2F8E3FE60" blockId="40.[151,1436,151,392]" box="[199,1256,367,392]" pageId="40" pageNumber="41">
<emphasis id="07AEEADDFFE65125FF57FEB2FD6BFE6F" bold="true" box="[199,352,367,392]" pageId="40" pageNumber="41">Distribution.</emphasis>
<collectingCountry id="4DCD765FFFE65125FEF7FEB2FDBDFE6F" box="[359,438,367,392]" name="Eritrea" pageId="40" pageNumber="41">Eritrea</collectingCountry>
,
<collectingCountry id="4DCD765FFFE65125FE51FEB2FE2FFE6F" box="[449,548,367,392]" name="Ethiopia" pageId="40" pageNumber="41">Ethiopia</collectingCountry>
:
<collectingRegion id="F71EF82DFFE65125FDBFFEB2FE9EFE6F" box="[559,661,367,392]" country="Ethiopia" name="Oromiya" pageId="40" pageNumber="41">Oromiya</collectingRegion>
, S.N.N.P.R.,
<collectingRegion id="F71EF82DFFE65125FCB9FEB2FF7DFE6F" box="[809,886,367,392]" country="Ethiopia" name="Tigray" pageId="40" pageNumber="41">Tigray</collectingRegion>
;
<collectingCountry id="4DCD765FFFE65125FC11FEADFFC4FE6F" box="[897,975,368,392]" name="Kenya" pageId="40" pageNumber="41">Kenya</collectingCountry>
:
<collectingRegion id="F71EF82DFFE65125FC49FEADF812FE6F" box="[985,1049,368,392]" country="Tanzania" name="Pwani" pageId="40" pageNumber="41">Coast</collectingRegion>
Province;
<collectingCountry id="4DCD765FFFE65125FB02FEADF8EFFE60" box="[1170,1252,368,391]" name="Yemen" pageId="40" pageNumber="41">Yemen</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>