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<document id="4FEA68E1759D799246FBF0C7FBE2D1D0" ID-CLB-Dataset="31846" ID-DOI="10.11646/zootaxa.4330.1.1" ID-GBIF-Dataset="4603132a-4e96-4119-bf2f-21ffe40dce69" ID-ISSN="1175-5326" ID-Zenodo-Dep="1010167" ID-ZooBank="Bf4D6091-5346-42Fd-8F64-D8E5Ca407415" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1507885096493" checkinUser="plazi" docAuthor="Häckel, Martin" docDate="2017" docId="BD7387D9FFDA5118FF07FF45FE5AFBD3" docLanguage="en" docName="zootaxa.4330.1.1.pdf.imf" docOrigin="Zootaxa 4330 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Craspedophorus ruficroides subsp. ruficroides Häckel 2017, n. sp." docType="treatment" docVersion="10" lastPageNumber="22" masterDocId="414AFFA1FFCE510DFF90FFDDFC0BFFE7" masterDocTitle="A contribution to the knowledge of the subfamily Panagaeinae Hope, 1838 from Africa. Part 3. Revision of the Craspedophorus strachani and C. brevicollis groups (Coleoptera: Carabidae)" masterLastPageNumber="67" masterPageNumber="1" pageNumber="21" updateTime="1698469524835" updateUser="ExternalLinkService">
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<mods:title id="CF6E3CC82E3F1EFC6DC2BAC846C1606E">A contribution to the knowledge of the subfamily Panagaeinae Hope, 1838 from Africa. Part 3. Revision of the Craspedophorus strachani and C. brevicollis groups (Coleoptera: Carabidae)</mods:title>
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<mods:date id="2401A7AAA53657842B48386767091DE1">2017</mods:date>
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<treatment id="BD7387D9FFDA5118FF07FF45FE5AFBD3" ID-DOI="http://doi.org/10.5281/zenodo.6019391" ID-GBIF-Taxon="135435415" ID-Zenodo-Dep="6019391" LSID="urn:lsid:plazi:treatment:BD7387D9FFDA5118FF07FF45FE5AFBD3" httpUri="http://treatment.plazi.org/id/BD7387D9FFDA5118FF07FF45FE5AFBD3" lastPageId="21" lastPageNumber="22" pageId="20" pageNumber="21">
<subSubSection id="7DC06544FFDA5119FF07FF45FEF3FF55" box="[151,760,151,178]" pageId="20" pageNumber="21" type="nomenclature">
<paragraph id="356536CFFFDA5119FF07FF45FEF3FF55" blockId="20.[151,782,151,211]" box="[151,760,151,178]" pageId="20" pageNumber="21">
<heading id="6E2D81A3FFDA5119FF07FF45FEF3FF55" bold="true" box="[151,760,151,178]" fontSize="11" level="1" pageId="20" pageNumber="21" reason="1">
<emphasis id="07AEEADDFFDA5119FF07FF45FEF3FF55" bold="true" box="[151,760,151,178]" pageId="20" pageNumber="21">
14a.
<taxonomicName id="F2DA4D4CFFDA5119FF41FF4AFEA4FF56" ID-CoL="8MLV7" authority="Häckel, 2017" authorityName="Häckel" authorityYear="2017" box="[209,687,151,177]" class="Insecta" family="Carabidae" genus="Craspedophorus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="subSpecies" species="ruficroides" status="sp. nov." subSpecies="ruficroides">
<emphasis id="07AEEADDFFDA5119FF41FF4AFEA4FF56" bold="true" box="[209,687,151,177]" italics="true" pageId="20" pageNumber="21">Craspedophorus ruficroides ruficroides</emphasis>
</taxonomicName>
<taxonomicNameLabel id="1C9D57A6FFDA5119FD26FF45FEF3FF55" box="[694,760,152,178]" pageId="20" pageNumber="21" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="7DC06544FFDA5119FF07FF67FF05FF34" box="[151,782,186,211]" pageId="20" pageNumber="21" type="description">
<paragraph id="356536CFFFDA5119FF07FF67FF05FF34" blockId="20.[151,782,151,211]" box="[151,782,186,211]" pageId="20" pageNumber="21">
(
<figureCitation id="ADE12A4AFFDA5119FF0EFF67FCE4FF34" box="[158,239,186,211]" captionStart="PLATE 6" captionStartId="55.[151,234,1836,1858]" captionTargetBox="[354,1236,1149,1798]" captionTargetId="figure@55.[340,1247,1146,1826]" captionTargetPageId="55" captionText="PLATE 6. Habitus of Craspedophorus strachani group V (Scale bar: 10 mm). 44. C. ruficroides ruficroides n. sp., HT (male); 45. C. ruficroides ruficroides n. sp., PT female, Guinea; 46. C. ruficroides thomsoni n. ssp., PT male, Central Africa; 47 C. ruficroides thomsoni n. ssp., PT, female, Cameroon, Centre; 48. C. ruficroides thomsoni n. ssp., HT (male); 49. C. congoanus Kolbe, 1883, female, DR Congo: Équateur (comparative type Basilewsky 1954); 50. C. congoanus Kolbe, 1883, male, DR Congo: Équateur; 51. C. congoanus Kolbe, 1883, female, DR Congo: Orientale; 52. C. phenax Basilewsky, 1987, HT (male); 53. C. phenax Basilewsky, 1987, female, DR Congo: Kasaï-Oriental; 54. C. phenax Basilewsky, 1987, male, Camerooon, West; 55. C. phenax Basilewsky, 1987, male, Camerooon, West." httpUri="https://zenodo.org/record/1010179/files/figure.png" pageId="20" pageNumber="21">Plate 6</figureCitation>
, Figs 4445,
<figureCitation id="ADE12A4AFFDA5119FE16FF67FDDFFF34" box="[390,468,186,211]" captionStart="PLATE 8" captionStartId="56.[151,234,1807,1829]" captionTargetBox="[341,1246,1107,1760]" captionTargetId="figure@56.[340,1247,1106,1786]" captionTargetPageId="56" captionText="PLATE 8. Craspedophorus strachani group, C. ruficrus complex, head and pronotum. 67 a. C. ruficrus (Chaudoir, 1861), female Guinea Equatorial. 68 a. C. lemariei n. sp., HT (male). 69 a. C. dicranulothorax, HT (female). 70. C. phenacoides n. sp., HT (male). 71. C. r. ruficroides n. sp., HT (male)." httpUri="https://zenodo.org/record/1010183/files/figure.png" pageId="20" pageNumber="21">Plate 8</figureCitation>
, Fig. 71,
<figureCitation id="ADE12A4AFFDA5119FDAFFF67FE97FF34" box="[575,668,186,211]" captionStart="PLATE 13" captionStartId="59.[151,234,815,837]" captionTargetBox="[348,1233,199,779]" captionTargetId="figure@59.[340,1247,193,794]" captionTargetPageId="59" captionText="PLATE 13 Craspedophorus strachani group II, aedeagus in frontal and right lateral views: 101. C. chevalieri (Alluaud, 1915), Central Africa; 102. C. cuneatus cuneatus (Alluaud, 1915), Central Africa; 103. C. cuneatus paromius Basilewsky, 1987, Orientale, DR Congo; 104. C. cuneatus rotundatus new subspecies, HT; 105. C. glaber Bates, 1886, Cameroon, Southwestern; 106. C. ruficroides ruficroides new species, HT; 107. C. ruficroides thomsoni new subspecies, PT RCA; 108. C. phenax Basilewsky, 1987, PT Fontem, Cameroon, West; 109. C. congoanus Kolbe, 1889, Eala Bokatola-Bikoro, Équateur, DR Congo; 110. C. phenacoides new species, HT; 111. C. lemariei new species, HT." httpUri="https://zenodo.org/record/1010195/files/figure.png" pageId="20" pageNumber="21">Plate 13</figureCitation>
, Fig. 106)
</paragraph>
</subSubSection>
<subSubSection id="7DC06544FFDA5119FF07FEDEFD2CFEB1" pageId="20" pageNumber="21" type="reference_group">
<paragraph id="356536CFFFDA5119FF07FEDEFD2CFEB1" blockId="20.[151,1437,258,342]" pageId="20" pageNumber="21">
<treatmentCitationGroup id="15CA11E1FFDA5119FF07FEDEFD2CFEB1" pageId="20" pageNumber="21">
<taxonomicName id="F2DA4D4CFFDA5119FF07FEDEFE73FEFF" ID-CoL="3QC65" authority="Laferte-Senectere, 1851: 221" authorityName="Laferte-Senectere" authorityPageNumber="221" authorityYear="1851" box="[151,632,258,280]" class="Insecta" family="Carabidae" genus="Isotarsus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="ruficrus">
<emphasis id="07AEEADDFFDA5119FF07FEDEFD41FEFF" box="[151,330,258,280]" italics="true" pageId="20" pageNumber="21">Isotarsus ruficrus</emphasis>
Laferté-Sénectere, 1851: 221
</taxonomicName>
(
<typeStatus id="EA61886DFFDA5119FD15FEDFFEB9FEFF" box="[645,690,258,280]" pageId="20" pageNumber="21">type</typeStatus>
locality “
<collectingCountry id="4DCD765FFFDA5119FC85FEDFFF6EFEFF" box="[789,869,258,280]" name="Guinea-Bissau" pageId="20" pageNumber="21">Guinea</collectingCountry>
”).
<taxonomicName id="F2DA4D4CFFDA5119FC10FEDFF992FEFF" authority="Gemminger and Harold 1868: 209" authorityName="Gemminger and Harold" authorityPageNumber="209" authorityYear="1868" box="[896,1433,258,280]" class="Magnoliopsida" family="Brassicaceae" genus="Eudema" kingdom="Plantae" order="Brassicales" pageId="20" pageNumber="21" phylum="Tracheophyta" rank="species" species="ruficrum">
<emphasis id="07AEEADDFFDA5119FC10FEDFF83AFEFF" box="[896,1073,258,280]" italics="true" pageId="20" pageNumber="21">Eudema ruficrum</emphasis>
<treatmentCitation id="B47B10DEFFDA5119FBA8FEDFF992FEFF" author="Gemminger" box="[1080,1433,258,280]" page="209" pageId="20" pageNumber="21" year="1868">
<bibRefCitation id="514B4B3EFFDA5119FBA8FEDFF96EFEFF" author="Gemminger" box="[1080,1381,258,280]" pageId="20" pageNumber="21" refString="Gemminger, M. &amp; Harold von, E. (1868) Catalogus coleopterorum hucusque descriptorum synonymicus et systematicus. Tom. I. Cicindelidae-Carabidae. Monachii, E. H. Gummi, xxxvi + 424 + 8 unn. pp." type="journal article" year="1868">Gemminger and Harold 1868</bibRefCitation>
: 209
</treatmentCitation>
</taxonomicName>
.
<taxonomicName id="F2DA4D4CFFDA5119FF57FEFCFE7EFED0" ID-CoL="Z5YW" authority="Alluaud 1930: 2" authorityName="Alluaud" authorityPageNumber="2" authorityYear="1930" box="[199,629,289,311]" class="Insecta" family="Carabidae" genus="Craspedophorus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="ruficrus">
<emphasis id="07AEEADDFFDA5119FF57FEFCFDCCFED0" box="[199,455,289,311]" italics="true" pageId="20" pageNumber="21">Craspedophorus ruficrus</emphasis>
<treatmentCitation id="B47B10DEFFDA5119FE5EFEFCFE7EFED0" author="Alluaud" box="[462,629,289,311]" page="2" pageId="20" pageNumber="21" year="1930">
<bibRefCitation id="514B4B3EFFDA5119FE5EFEFCFE55FED0" author="Alluaud" box="[462,606,289,311]" pageId="20" pageNumber="21" refString="Alluaud, C. (1930) Note sur les Panageides dAfrique tropicale. Afra, Cahiers d'Entomologie, 1, 1 - 9." type="journal article" year="1930">Alluaud 1930</bibRefCitation>
: 2
</treatmentCitation>
</taxonomicName>
(nec. Chaudoir).
<treatmentCitation id="B47B10DEFFDA5119FCB9FEFCF805FED0" author="Basilewsky" box="[809,1038,289,311]" page="383" pageId="20" pageNumber="21" year="1963">
<bibRefCitation id="514B4B3EFFDA5119FCB9FEFCFFD0FED0" author="Basilewsky" box="[809,987,289,311]" pageId="20" pageNumber="21" refString="Basilewsky, P. (1963) La reserve naturelle integrale du Mont Nimba. XV. Coleoptera Carabidae (troisieme note). Memoires de l'Institut Francais de l'Afrique Noire, 66, 367 - 391." type="journal article" year="1963">Basilewsky 1963</bibRefCitation>
: 383
</treatmentCitation>
,
<treatmentCitation id="B47B10DEFFDA5119FB89FEFFF8DAFED0" author="Lorenz" box="[1049,1233,289,311]" page="321" pageId="20" pageNumber="21" year="2005">
<bibRefCitation id="514B4B3EFFDA5119FB89FEFFF895FED0" author="Lorenz" box="[1049,1182,289,311]" pageId="20" pageNumber="21" refString="Lorenz, W. (2005) A systematic list of extant ground beetles of the World (Insecta, Coleoptera, Adephaga: Trachypachidae &amp; Carabidae incl. Paussinae, Cicindelinae, Rhysodinae). 2 nd Edition. Published by the author, Tutzing, 530 pp." type="book" year="2005">Lorenz 2005</bibRefCitation>
: 321
</treatmentCitation>
,
<treatmentCitation id="B47B10DEFFDA5119FB4CFEFCFD28FEB1" author="Hackel" page="83" pageId="20" pageNumber="21" year="2012">
<bibRefCitation id="514B4B3EFFDA5119FB4CFEFCFCF5FEB1" author="Hackel" pageId="20" pageNumber="21" refString="Hackel, M. &amp; Farkac, J. (2012) A check-list of the subfamily Panagaeinae Hope, 1838 of the World (Coleoptera: Carabidae). Studies and Reports, Taxonomical Series, 8 (1 - 2), 67 - 116." type="journal article" year="2012">Häckel and Farkač 2012</bibRefCitation>
: 83
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="7DC06544FFDA5119FF07FE5FFE4AFB54" pageId="20" pageNumber="21" type="materials_examined">
<paragraph id="356536CFFFDA5119FF07FE5FFE18FE7C" blockId="20.[151,1437,386,2031]" box="[151,531,386,411]" pageId="20" pageNumber="21">
<materialsCitation id="85B23C92FFDA5119FF07FE5FFE18FE7C" ID-GBIF-Occurrence="1671737065" box="[151,531,386,411]" country="Guinea-Bissau" location="Nimba" pageId="20" pageNumber="21" specimenCount="1" stateProvince="Nimba" typeStatus="holotype">
<emphasis id="07AEEADDFFDA5119FF07FE5FFD3EFE7C" bold="true" box="[151,309,386,411]" pageId="20" pageNumber="21">
<typeStatus id="EA61886DFFDA5119FF07FE5FFCD9FE7C" box="[151,210,386,411]" pageId="20" pageNumber="21">Type</typeStatus>
locality.
</emphasis>
<collectingRegion id="F71EF82DFFDA5119FED6FE5FFD9EFE7C" box="[326,405,386,411]" country="Liberia" name="Nimba" pageId="20" pageNumber="21">Nimba</collectingRegion>
(
<collectingCountry id="4DCD765FFFDA5119FE34FE5FFDF6FE7C" box="[420,509,386,411]" name="Guinea-Bissau" pageId="20" pageNumber="21">Guinée</collectingCountry>
)”.
</materialsCitation>
</paragraph>
<paragraph id="356536CFFFDA5119FF57FE7BF996FD70" blockId="20.[151,1437,386,2031]" pageId="20" pageNumber="21">
<emphasis id="07AEEADDFFDA5119FF57FE7BFD7EFE58" bold="true" box="[199,373,422,447]" pageId="20" pageNumber="21">Type material.</emphasis>
Holotype (♂): “Comp. typ. / Basilewsky [printed in black on red circumscribed label]// Camp des /
<taxonomicName id="F2DA4D4CFFDA5119FF42FE17FD24FE04" box="[210,303,458,483]" class="Polypodiopsida" family="Cyatheaceae" genus="Cyathea" kingdom="Plantae" order="Cyatheales" pageId="20" pageNumber="21" phylum="Tracheophyta" rank="genus">
<emphasis id="07AEEADDFFDA5119FF42FE17FD24FE04" box="[210,303,458,483]" italics="true" pageId="20" pageNumber="21">Cyathea</emphasis>
</taxonomicName>
[handwritten in black on white label]// Nimba (Guinée) / Lamotte /
<date id="4164100FFFDA5119FBBFFE17F8D7FE04" box="[1071,1244,458,483]" pageId="20" pageNumber="21" value="1957-02-23" valueMax="1957-02-28" valueMin="1957-02-23">2328II1957</date>
[printed in black on blue label]// Mus. Roy. Afr. Centr. / (don M. Lamotte) / Coll. P. Basilewsky [printed in black on white label]//
<emphasis id="07AEEADDFFDA5119FF07FDCEFCE8FDCD" box="[151,227,531,554]" italics="true" pageId="20" pageNumber="21">Crasp.</emphasis>
/
<taxonomicName id="F2DA4D4CFFDA5119FF6AFDCEFDA7FDCC" authority="Chaud." authorityName="Chaudoir" authorityYear="1879" box="[250,428,530,555]" class="Insecta" family="Carabidae" genus="Craspedophorus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="ruficrus">
<emphasis id="07AEEADDFFDA5119FF6AFDCEFD58FDCC" box="[250,339,531,555]" italics="true" pageId="20" pageNumber="21">ruficrus</emphasis>
Chaud.
</taxonomicName>
[handwritten in black]/ P. Basilewsky det., 19 [printed on white label]” (
<figureCitation id="ADE12A4AFFDA5119FB76FDCFF933FDCC" box="[1254,1336,530,555]" captionStart="PLATE 6" captionStartId="55.[151,234,1836,1858]" captionTargetBox="[354,1236,1149,1798]" captionTargetId="figure@55.[340,1247,1146,1826]" captionTargetPageId="55" captionText="PLATE 6. Habitus of Craspedophorus strachani group V (Scale bar: 10 mm). 44. C. ruficroides ruficroides n. sp., HT (male); 45. C. ruficroides ruficroides n. sp., PT female, Guinea; 46. C. ruficroides thomsoni n. ssp., PT male, Central Africa; 47 C. ruficroides thomsoni n. ssp., PT, female, Cameroon, Centre; 48. C. ruficroides thomsoni n. ssp., HT (male); 49. C. congoanus Kolbe, 1883, female, DR Congo: Équateur (comparative type Basilewsky 1954); 50. C. congoanus Kolbe, 1883, male, DR Congo: Équateur; 51. C. congoanus Kolbe, 1883, female, DR Congo: Orientale; 52. C. phenax Basilewsky, 1987, HT (male); 53. C. phenax Basilewsky, 1987, female, DR Congo: Kasaï-Oriental; 54. C. phenax Basilewsky, 1987, male, Camerooon, West; 55. C. phenax Basilewsky, 1987, male, Camerooon, West." httpUri="https://zenodo.org/record/1010179/files/figure.png" pageId="20" pageNumber="21">Plate 6</figureCitation>
, Fig. 44,
<figureCitation id="ADE12A4AFFDA5119FF07FDEBFCEEFDA8" box="[151,229,566,591]" captionStart="PLATE 8" captionStartId="56.[151,234,1807,1829]" captionTargetBox="[341,1246,1107,1760]" captionTargetId="figure@56.[340,1247,1106,1786]" captionTargetPageId="56" captionText="PLATE 8. Craspedophorus strachani group, C. ruficrus complex, head and pronotum. 67 a. C. ruficrus (Chaudoir, 1861), female Guinea Equatorial. 68 a. C. lemariei n. sp., HT (male). 69 a. C. dicranulothorax, HT (female). 70. C. phenacoides n. sp., HT (male). 71. C. r. ruficroides n. sp., HT (male)." httpUri="https://zenodo.org/record/1010183/files/figure.png" pageId="20" pageNumber="21">Plate 8</figureCitation>
, Fig. 71,
<figureCitation id="ADE12A4AFFDA5119FEC0FDEBFDA6FDA8" box="[336,429,566,591]" captionStart="PLATE 13" captionStartId="59.[151,234,815,837]" captionTargetBox="[348,1233,199,779]" captionTargetId="figure@59.[340,1247,193,794]" captionTargetPageId="59" captionText="PLATE 13 Craspedophorus strachani group II, aedeagus in frontal and right lateral views: 101. C. chevalieri (Alluaud, 1915), Central Africa; 102. C. cuneatus cuneatus (Alluaud, 1915), Central Africa; 103. C. cuneatus paromius Basilewsky, 1987, Orientale, DR Congo; 104. C. cuneatus rotundatus new subspecies, HT; 105. C. glaber Bates, 1886, Cameroon, Southwestern; 106. C. ruficroides ruficroides new species, HT; 107. C. ruficroides thomsoni new subspecies, PT RCA; 108. C. phenax Basilewsky, 1987, PT Fontem, Cameroon, West; 109. C. congoanus Kolbe, 1889, Eala Bokatola-Bikoro, Équateur, DR Congo; 110. C. phenacoides new species, HT; 111. C. lemariei new species, HT." httpUri="https://zenodo.org/record/1010195/files/figure.png" pageId="20" pageNumber="21">Plate 13</figureCitation>
, Fig. 106, NMPC). Paratypes. 1♂,2♀ same labeled as holotype except: “Nimba,
<quantity id="F2229B2AFFDA5119FAABFDEAF986FDA8" box="[1339,1421,567,591]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="20" pageNumber="21" unit="m" value="1000.0">1000m</quantity>
”, 1♀ same labeled except: “Nimba, Camp des
<taxonomicName id="F2DA4D4CFFDA5119FD0BFD87FF6BFD95" box="[667,864,602,627]" class="Polypodiopsida" family="Cyatheaceae" genus="Cyathea" kingdom="Plantae" order="Cyatheales" pageId="20" pageNumber="21" phylum="Tracheophyta" rank="genus">
Cyathea,
<quantity id="F2229B2AFFDA5119FC97FD86FF6BFD95" box="[775,864,603,627]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="20" pageNumber="21" unit="m" value="1000.0">1000 m</quantity>
</taxonomicName>
, 1♂,1♀ same labeled except: “Nimba”, 1 ♀ same labeled except: “Nimba, Yalanzou”, 1♀: same labeled except “Nimba, Kéoulenta” (
<figureCitation id="ADE12A4AFFDA5119FBABFDA3F880FD70" box="[1083,1163,638,663]" captionStart="PLATE 6" captionStartId="55.[151,234,1836,1858]" captionTargetBox="[354,1236,1149,1798]" captionTargetId="figure@55.[340,1247,1146,1826]" captionTargetPageId="55" captionText="PLATE 6. Habitus of Craspedophorus strachani group V (Scale bar: 10 mm). 44. C. ruficroides ruficroides n. sp., HT (male); 45. C. ruficroides ruficroides n. sp., PT female, Guinea; 46. C. ruficroides thomsoni n. ssp., PT male, Central Africa; 47 C. ruficroides thomsoni n. ssp., PT, female, Cameroon, Centre; 48. C. ruficroides thomsoni n. ssp., HT (male); 49. C. congoanus Kolbe, 1883, female, DR Congo: Équateur (comparative type Basilewsky 1954); 50. C. congoanus Kolbe, 1883, male, DR Congo: Équateur; 51. C. congoanus Kolbe, 1883, female, DR Congo: Orientale; 52. C. phenax Basilewsky, 1987, HT (male); 53. C. phenax Basilewsky, 1987, female, DR Congo: Kasaï-Oriental; 54. C. phenax Basilewsky, 1987, male, Camerooon, West; 55. C. phenax Basilewsky, 1987, male, Camerooon, West." httpUri="https://zenodo.org/record/1010179/files/figure.png" pageId="20" pageNumber="21">Plate 6</figureCitation>
, Fig. 45, MRAC, cMH).
</paragraph>
<paragraph id="356536CFFFDA5119FF57FD7FFE4AFB54" blockId="20.[151,1437,386,2031]" pageId="20" pageNumber="21">
<emphasis id="07AEEADDFFDA5119FF57FD7FFD0DFD5C" bold="true" box="[199,262,674,699]" pageId="20" pageNumber="21">Note.</emphasis>
Laferté-Sénectere (1851) described
<taxonomicName id="F2DA4D4CFFDA5119FD31FD7EFF17FD5C" authorityName="Chaudoir" authorityYear="1879" box="[673,796,675,699]" class="Insecta" family="Carabidae" genus="Craspedophorus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="ruficrus">
<emphasis id="07AEEADDFFDA5119FD31FD7EFF17FD5C" box="[673,796,675,699]" italics="true" pageId="20" pageNumber="21">C. ruficrus</emphasis>
</taxonomicName>
very briefly, in a manner applicable to the entire species group (“Première division”). Description (in part, see key to species in LaFerté-Sénectere 1851: 220): “Grande taille (de 10 à 28 millimètres). Antennes plus longues que la moitié du corps, atténuées vers ľextrémité; le troisième article plus que du double du deuxième.
<collectingRegion id="F71EF82DFFDA5119FDF0FCD2FE83FCC1" box="[608,648,783,806]" country="Cameroon" name="East" pageId="20" pageNumber="21">Les</collectingRegion>
pattes entièrement noires”. Its sections (1851: 220 a-bbb) are devoted to differential diagnosis and the shape of the pronotum, and characterize
<taxonomicName id="F2DA4D4CFFDA5119FC55FCEEF835FCAC" authorityName="Chaudoir" authorityYear="1879" box="[965,1086,819,843]" class="Insecta" family="Carabidae" genus="Craspedophorus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="ruficrus">
<emphasis id="07AEEADDFFDA5119FC55FCEEF835FCAC" box="[965,1086,819,843]" italics="true" pageId="20" pageNumber="21">C. ruficrus</emphasis>
</taxonomicName>
as follows: “(aa) pronotum not convex, margins more-or-less elevated, (bb) transverse, not too annular, with sinuate margins, (cc) without constriction at base (peduncle)” [from French]. As the country of origin is given “
<collectingCountry id="4DCD765FFFDA5119FBA5FCA7F881FC74" box="[1077,1162,890,915]" name="Guinea-Bissau" pageId="20" pageNumber="21">Guinea</collectingCountry>
” (1851: 221). However, the
<collectionCode id="53CBAE0AFFDA5119FF54FC42FD12FC51" box="[196,281,927,950]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="20" pageNumber="21">MNHN</collectionCode>
<typeStatus id="EA61886DFFDA5119FEB4FC43FD80FC50" box="[292,395,926,951]" pageId="20" pageNumber="21" type="lectotype">lectotype</typeStatus>
selected by
<bibRefCitation id="514B4B3EFFDA5119FDB0FC43FF11FC50" author="Chaudoir" box="[544,794,926,951]" pageId="20" pageNumber="21" refString="Chaudoir, M. de (1861) Revision des especes qui rentrent dans l`ancien genre Panagaeus. Bulletin da la Societe Imperiale des Naturalistes de Moscou, 34 (4), 335 - 360." type="journal article" year="1861">Chaudoir (1861: 340)</bibRefCitation>
in my opinion belongs to a species different from that inhabiting
<collectingCountry id="4DCD765FFFDA5119FE81FC1FFD69FC3C" box="[273,354,962,987]" name="Guinea-Bissau" pageId="20" pageNumber="21">Guinea</collectingCountry>
(see discussion). The valid name therefore becomes
<taxonomicName id="F2DA4D4CFFDA5119FC2CFC1EF8F4FC3C" authority="(Chaudoir, 1861)" authorityName="Häckel" authorityYear="2017" baseAuthorityName="Chaudoir" baseAuthorityYear="1861" box="[956,1279,962,987]" class="Insecta" family="Carabidae" genus="Craspedophorus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="ruficrus">
<emphasis id="07AEEADDFFDA5119FC2CFC1EF83DFC3C" box="[956,1078,963,987]" italics="true" pageId="20" pageNumber="21">C. ruficrus</emphasis>
(
<bibRefCitation id="514B4B3EFFDA5119FBD6FC1FF8FCFC3C" author="Chaudoir" box="[1094,1271,962,987]" pageId="20" pageNumber="21" refString="Chaudoir, M. de (1861) Revision des especes qui rentrent dans l`ancien genre Panagaeus. Bulletin da la Societe Imperiale des Naturalistes de Moscou, 34 (4), 335 - 360." type="journal article" year="1861">Chaudoir, 1861</bibRefCitation>
)
</taxonomicName>
with the
<typeStatus id="EA61886DFFDA5119FAFCFC1EF997FC3C" box="[1388,1436,963,987]" pageId="20" pageNumber="21">type</typeStatus>
locality “
<collectingCountry id="4DCD765FFFDA5119FF6EFC3BFD5BFC18" box="[254,336,998,1023]" name="Gabon" pageId="20" pageNumber="21">Gabon</collectingCountry>
“. Lafertés name and description cannot be applied to any taxon and must be treated as a
<emphasis id="07AEEADDFFDA5119FAC1FC3AFCE0FBC4" italics="true" pageId="20" pageNumber="21">nomen dubium</emphasis>
(Article 11.5.2 of the Code)]. The
<collectingCountry id="4DCD765FFFDA5119FD3AFBD7FEF0FBC4" box="[682,763,1034,1059]" name="Guinea-Bissau" pageId="20" pageNumber="21">Guinea</collectingCountry>
species (although it may comply with Lafertés vague description) must be redescribed under a new name and
<typeStatus id="EA61886DFFDA5119FCAAFBF3FF97FBA0" box="[826,924,1070,1095]" pageId="20" pageNumber="21" type="holotype">holotype</typeStatus>
designation. I have selected a specimen in Basilewskys collection that agrees with both the original Lafertés description and Basilewskys opinion.
<collectingRegion id="F71EF82DFFDA5119FA18FB8EF997FB8C" box="[1416,1436,1107,1131]" country="Nigeria" name="Rivers" pageId="20" pageNumber="21">A</collectingRegion>
suitable candidate for
<typeStatus id="EA61886DFFDA5119FE05FBABFDFCFB68" box="[405,503,1142,1167]" pageId="20" pageNumber="21" type="holotype">holotype</typeStatus>
of the new species is Basilewskys male specimen from Mt.
<collectingRegion id="F71EF82DFFDA5119FB2BFBABF90CFB68" box="[1211,1287,1142,1167]" country="Liberia" name="Nimba" pageId="20" pageNumber="21">Nimba</collectingRegion>
(comparated with the
<typeStatus id="EA61886DFFDA5119FF69FB46FD27FB54" box="[249,300,1179,1203]" pageId="20" pageNumber="21">type</typeStatus>
,
<bibRefCitation id="514B4B3EFFDA5119FEA7FB47FDF0FB54" author="Basilewsky" box="[311,507,1178,1203]" pageId="20" pageNumber="21" refString="Basilewsky, P. (1963) La reserve naturelle integrale du Mont Nimba. XV. Coleoptera Carabidae (troisieme note). Memoires de l'Institut Francais de l'Afrique Noire, 66, 367 - 391." type="journal article" year="1963">Basilewsky 1963</bibRefCitation>
: 383).
</paragraph>
</subSubSection>
<subSubSection id="7DC06544FFDA5118FF57FB63FDB8FD17" lastPageId="21" lastPageNumber="22" pageId="20" pageNumber="21" type="description">
<paragraph id="356536CFFFDA5119FF57FB63F894FB1C" blockId="20.[151,1437,386,2031]" pageId="20" pageNumber="21">
<emphasis id="07AEEADDFFDA5119FF57FB63FDE6FB30" bold="true" box="[199,493,1214,1239]" pageId="20" pageNumber="21">
Description of
<typeStatus id="EA61886DFFDA5119FEEFFB63FDE3FB30" box="[383,488,1214,1239]" pageId="20" pageNumber="21" type="holotype">holotype</typeStatus>
.
</emphasis>
Length
<quantity id="F2229B2AFFDA5119FDC2FB62FEB7FB31" box="[594,700,1215,1239]" metricMagnitude="-2" metricUnit="m" metricValue="1.94" pageId="20" pageNumber="21" unit="mm" value="19.4">19.4 mm</quantity>
, width
<quantity id="F2229B2AFFDA5119FC87FB62FF7EFB31" box="[791,885,1215,1239]" metricMagnitude="-3" metricUnit="m" metricValue="8.1" pageId="20" pageNumber="21" unit="mm" value="8.1">8.1 mm</quantity>
. Proportions: Pronotum 1.44x wider than long, 2.01x wider than head with eyes, elytra 1.21x wider than pronotum, 1.50x longer than wide.
</paragraph>
<paragraph id="356536CFFFDA5119FF57FADBFE7CFA6C" blockId="20.[151,1437,386,2031]" pageId="20" pageNumber="21">Colouration. Body throughout piceous black, opaque, densely covered with black setae. Femora black, genae, tarsi, antennae and mandibles brownish. Each elytron with two yellow-red maculae, humeral macula slightly transverse, semicircular, reaching from V to IX interval, overlapping to IV., elytral margin black. Preapical macula quadrate, reaching from V to VIII interval.</paragraph>
<paragraph id="356536CFFFDA5119FF57FA4BFFB6F915" blockId="20.[151,1437,386,2031]" pageId="20" pageNumber="21">Head subquadrate, labrum and mandibles smooth, glossy, terminal labial palpomere prolonged, kidney-shaped (male). Eyes weakly convex. Frons flat, only a small glossy protuberance behind labium weakly elevated. Surface softly, densely punctured without grooves, in front of eyes with two longitudinal depressions, lateral in the form of a deep and wide ridge running from the clypeus to eyes mid-length (supra-antennary ridge), wider than in other species, medial wider, shallow, almost indistinct, running from clypeus base to the anterior margin of eye, softly and densely punctured as the frontal center. Vertex punctured somewhat more sparsely without furrows, occiput and neck punctured as well, without strangulation. Antennae whip-shaped, distally not widening nor flattening, reaching almost up to the half of elytral length. The basal antennomere (scapus) twice longer than the second, the third twice longer than the first, the fourth one only weakly shorter than the first, surface densely covered with short setae, longer setae cumulated at the distal end of the antennomere.</paragraph>
<paragraph id="356536CFFFDA5118FF57F922FF11FED8" blockId="20.[151,1437,386,2031]" lastBlockId="21.[151,1437,151,1077]" lastPageId="21" lastPageNumber="22" pageId="20" pageNumber="21">Pronotum not too tranverse 1.44x wider than long, cordiform; anterior margin straight, as long as head with eyes is wide, parallel to base in its middle, straight part as long as neck is wide, laterally changing to anterior angles, weakly rounded and distinctly protruded forward, lateral margins widely curved, narrowing towards base with a distinct sinuosity, maximum width behind mid-length, in front of posterior angles mild incisure, posterior angles rectangular, posteriorly weakly protruded to elytral base. Base straight at the center, at the margins up to one fourth indistinctly angled towards elytral humeri without any peduncle. Pronotal disc convex, separated by a shallow depression from wide lateral rims which are distinct already from the anterior pronotal margin, the rim posteriorly distinctly widening, elevated upwards along the entire length, the greatest elevation in its basal fourth, then the rim flattening, thus posterior angles jutting out weakly upwards, neither backwards nor to the sides, without being keel-like. Disc and basal impressions grossly, densely and regularly punctured, as the frons, elevated lateral pronotal margins distinctly more glabrous than disc, sparsely and softly punctured, almost smooth posteriorly. Dorsal surface covered with long black setae.</paragraph>
<paragraph id="356536CFFFDB5118FF57FE91FD82FD87" blockId="21.[151,1437,151,1077]" pageId="21" pageNumber="22">
Elytra fused, moderately convex, less ovoid, almost parallel-sided, weakly widening to midlength, then narrowing, maximum width in midlength. Disc moderately symmetrically convex with indistinct collar-like basal depression. Humeri distinct, weakly rounded without bevelling, elytral base with distinct rim separating elytral base from striae between scutellum and VI interval, laterally disappearing. Elytra margined with a narrow rim, regularly and softly punctured longwise, without widening. Striae deep, strial punctuation very soft and regular. Intervals convex, densely punctured in four to five lines. Intervals with soft distinct isodiametric microsculpure among punctuation. Elytra sparsely covered with dark setae which are shorter than those covering the pronotum (
<figureCitation id="ADE12A4AFFDB5118FF0EFD9AFCE4FD87" box="[158,239,583,608]" captionStart="PLATE 6" captionStartId="55.[151,234,1836,1858]" captionTargetBox="[354,1236,1149,1798]" captionTargetId="figure@55.[340,1247,1146,1826]" captionTargetPageId="55" captionText="PLATE 6. Habitus of Craspedophorus strachani group V (Scale bar: 10 mm). 44. C. ruficroides ruficroides n. sp., HT (male); 45. C. ruficroides ruficroides n. sp., PT female, Guinea; 46. C. ruficroides thomsoni n. ssp., PT male, Central Africa; 47 C. ruficroides thomsoni n. ssp., PT, female, Cameroon, Centre; 48. C. ruficroides thomsoni n. ssp., HT (male); 49. C. congoanus Kolbe, 1883, female, DR Congo: Équateur (comparative type Basilewsky 1954); 50. C. congoanus Kolbe, 1883, male, DR Congo: Équateur; 51. C. congoanus Kolbe, 1883, female, DR Congo: Orientale; 52. C. phenax Basilewsky, 1987, HT (male); 53. C. phenax Basilewsky, 1987, female, DR Congo: Kasaï-Oriental; 54. C. phenax Basilewsky, 1987, male, Camerooon, West; 55. C. phenax Basilewsky, 1987, male, Camerooon, West." httpUri="https://zenodo.org/record/1010179/files/figure.png" pageId="21" pageNumber="22">Plate 6</figureCitation>
, Figs 44, 45).
</paragraph>
<paragraph id="356536CFFFDB5118FF57FDB1FF91FD40" blockId="21.[151,1437,151,1077]" pageId="21" pageNumber="22">Underside covered with short setae, finely punctured medially, coarsely punctured near margins; metepisterna rhombic, as long as wide. Legs moderately covered with black setae.</paragraph>
<paragraph id="356536CFFFDB5118FF57FD69FDB8FD17" blockId="21.[151,1437,151,1077]" pageId="21" pageNumber="22">
Aedeagus (
<figureCitation id="ADE12A4AFFDB5118FED4FD69FDA8FD2B" box="[324,419,692,717]" captionStart="PLATE 13" captionStartId="59.[151,234,815,837]" captionTargetBox="[348,1233,199,779]" captionTargetId="figure@59.[340,1247,193,794]" captionTargetPageId="59" captionText="PLATE 13 Craspedophorus strachani group II, aedeagus in frontal and right lateral views: 101. C. chevalieri (Alluaud, 1915), Central Africa; 102. C. cuneatus cuneatus (Alluaud, 1915), Central Africa; 103. C. cuneatus paromius Basilewsky, 1987, Orientale, DR Congo; 104. C. cuneatus rotundatus new subspecies, HT; 105. C. glaber Bates, 1886, Cameroon, Southwestern; 106. C. ruficroides ruficroides new species, HT; 107. C. ruficroides thomsoni new subspecies, PT RCA; 108. C. phenax Basilewsky, 1987, PT Fontem, Cameroon, West; 109. C. congoanus Kolbe, 1889, Eala Bokatola-Bikoro, Équateur, DR Congo; 110. C. phenacoides new species, HT; 111. C. lemariei new species, HT." httpUri="https://zenodo.org/record/1010195/files/figure.png" pageId="21" pageNumber="22">Plate 13</figureCitation>
, Fig. 106) is similar to that in
<taxonomicName id="F2DA4D4CFFDB5118FD69FD68F833FD2B" authority="Basilewsky, 1987" authorityName="Basilewsky" authorityYear="1987" box="[761,1080,692,717]" class="Insecta" family="Carabidae" genus="Craspdophorus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="phenax">
<emphasis id="07AEEADDFFDB5118FD69FD68FF60FD2A" box="[761,875,692,717]" italics="true" pageId="21" pageNumber="22">C. phenax</emphasis>
<bibRefCitation id="514B4B3EFFDB5118FCE2FD69F833FD2B" author="Basilewsky" box="[882,1080,692,717]" pageId="21" pageNumber="22" refString="Basilewsky, P. (1987) Descriptions de quelques especes africaines nouvelles du genre Craspedophorus Hope et notes synonymiques (Coleoptera Carabidae Panagaeini). Bulletin et Annales de La Societe Royale Belge d'entomologie, 123 (7 - 9), 185 - 201." type="journal article" year="1987">Basilewsky, 1987</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="F2DA4D4CFFDB5118FBFFFD68F997FD2B" authority="Kolbe, 1883" authorityName="Kolbe" authorityYear="1883" box="[1135,1436,692,717]" class="Insecta" family="Carabidae" genus="Craspedophorus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="congoanus">
<emphasis id="07AEEADDFFDB5118FBFFFD68F902FD2B" box="[1135,1289,693,716]" italics="true" pageId="21" pageNumber="22">C. congoanus</emphasis>
Kolbe, 1883
</taxonomicName>
(
<figureCitation id="ADE12A4AFFDB5118FF0EFD0AFCF6FD17" box="[158,253,727,752]" captionStart="PLATE 13" captionStartId="59.[151,234,815,837]" captionTargetBox="[348,1233,199,779]" captionTargetId="figure@59.[340,1247,193,794]" captionTargetPageId="59" captionText="PLATE 13 Craspedophorus strachani group II, aedeagus in frontal and right lateral views: 101. C. chevalieri (Alluaud, 1915), Central Africa; 102. C. cuneatus cuneatus (Alluaud, 1915), Central Africa; 103. C. cuneatus paromius Basilewsky, 1987, Orientale, DR Congo; 104. C. cuneatus rotundatus new subspecies, HT; 105. C. glaber Bates, 1886, Cameroon, Southwestern; 106. C. ruficroides ruficroides new species, HT; 107. C. ruficroides thomsoni new subspecies, PT RCA; 108. C. phenax Basilewsky, 1987, PT Fontem, Cameroon, West; 109. C. congoanus Kolbe, 1889, Eala Bokatola-Bikoro, Équateur, DR Congo; 110. C. phenacoides new species, HT; 111. C. lemariei new species, HT." httpUri="https://zenodo.org/record/1010195/files/figure.png" pageId="21" pageNumber="22">Plate 13</figureCitation>
, Figs 108, 109).
</paragraph>
</subSubSection>
<subSubSection id="7DC06544FFDB5118FF57FD21FFF8FC2F" pageId="21" pageNumber="22" type="diagnosis">
<paragraph id="356536CFFFDB5118FF57FD21FFF8FC2F" blockId="21.[151,1437,151,1077]" pageId="21" pageNumber="22">
<emphasis id="07AEEADDFFDB5118FF57FD21FDC5FCF2" bold="true" box="[199,462,764,789]" pageId="21" pageNumber="22">Differential diagnosis.</emphasis>
The species mostly resembles
<taxonomicName id="F2DA4D4CFFDB5118FCBAFD20F85DFCF3" authority="Kolbe, 1889" authorityName="Kolbe" authorityYear="1889" box="[810,1110,764,789]" class="Insecta" family="Carabidae" genus="Craspedophorus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="congoanus">
<emphasis id="07AEEADDFFDB5118FCBAFD20FFCFFCF3" box="[810,964,765,788]" italics="true" pageId="21" pageNumber="22">C. congoanus</emphasis>
<bibRefCitation id="514B4B3EFFDB5118FC5BFD21F85DFCF3" author="Kolbe" box="[971,1110,764,789]" pageId="21" pageNumber="22" refString="Kolbe, H. J. (1889) Bericht uber die von Herrn Dr. R. Buttner im Gebiete des unteren Quango und Kongo gesammelten Coleopteren. Entomologische Zeitung, 1889 (4 - 66), 113 - 133." type="journal article" year="1889">
Kolbe,
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</bibRefCitation>
</taxonomicName>
in its size, in the shape of its body and its pronotum, differing from it in its elytra, especially in elytral shape and sculpure.
<taxonomicName id="F2DA4D4CFFDB5118FB52FCFDF976FCDF" authority="Häckel, 2017" authorityName="Häckel" authorityYear="2017" box="[1218,1405,799,824]" class="Insecta" family="Carabidae" genus="Craspedophorus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="subSpecies" species="ruficroides" status="sp. nov." subSpecies="ruficroides">
In
<emphasis id="07AEEADDFFDB5118FB71FCFDF976FCDF" box="[1249,1405,799,824]" italics="true" pageId="21" pageNumber="22">C. ruficroides</emphasis>
</taxonomicName>
<emphasis id="07AEEADDFFDB5118FA15FCC2FCB3FCBA" bold="true" pageId="21" pageNumber="22">
<taxonomicNameLabel id="1C9D57A6FFDB5118FA15FCC2FCB3FCBA" pageId="21" pageNumber="22" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
elytra wider and less widening posteriorly, head punctuated more densely, neck distinctly punctuated (in
<taxonomicName id="F2DA4D4CFFDB5118FA12FC98FD1BFC67" authorityName="Kolbe" authorityYear="1889" class="Insecta" family="Carabidae" genus="Craspedophorus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="congoanus">
<emphasis id="07AEEADDFFDB5118FA12FC98FD1BFC67" italics="true" pageId="21" pageNumber="22">C. congoanus</emphasis>
</taxonomicName>
neck is smooth), elytral intervals more densely punctuated as well (
<taxonomicName id="F2DA4D4CFFDB5118FB97FCBAF8B4FC67" authority="Häckel, 2017" authorityName="Häckel" authorityYear="2017" box="[1031,1215,871,896]" class="Insecta" family="Carabidae" genus="Craspedophorus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="subSpecies" species="ruficroides" status="sp. nov." subSpecies="ruficroides">
in
<emphasis id="07AEEADDFFDB5118FBB4FCB5F8B4FC67" box="[1060,1215,871,896]" italics="true" pageId="21" pageNumber="22">C. ruficroides</emphasis>
</taxonomicName>
punctuation in four to six lines, in
<taxonomicName id="F2DA4D4CFFDB5118FEDCFC50FDE1FC43" authorityName="Kolbe" authorityYear="1889" box="[332,490,909,932]" class="Insecta" family="Carabidae" genus="Craspedophorus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="congoanus">
<emphasis id="07AEEADDFFDB5118FEDCFC50FDE1FC43" box="[332,490,909,932]" italics="true" pageId="21" pageNumber="22">C. congoanus</emphasis>
</taxonomicName>
in three to four lines). The species also resembles
<taxonomicName id="F2DA4D4CFFDB5118FBDBFC50F993FC43" authority="Basilewsky, 1987" authorityName="Basilewsky" authorityYear="1987" box="[1099,1432,908,933]" class="Insecta" family="Carabidae" genus="Craspdophorus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="phenax">
<emphasis id="07AEEADDFFDB5118FBDBFC50F8CBFC42" box="[1099,1216,908,933]" italics="true" pageId="21" pageNumber="22">C. phenax</emphasis>
<bibRefCitation id="514B4B3EFFDB5118FB5CFC51F993FC43" author="Basilewsky" box="[1228,1432,908,933]" pageId="21" pageNumber="22" refString="Basilewsky, P. (1987) Descriptions de quelques especes africaines nouvelles du genre Craspedophorus Hope et notes synonymiques (Coleoptera Carabidae Panagaeini). Bulletin et Annales de La Societe Royale Belge d'entomologie, 123 (7 - 9), 185 - 201." type="journal article" year="1987">Basilewsky, 1987</bibRefCitation>
</taxonomicName>
, differing from it in the same characters and in the elytral colouration as well.
</paragraph>
</subSubSection>
<subSubSection id="7DC06544FFDB5118FF57FC09FDC8FBE8" pageId="21" pageNumber="22" type="etymology">
<paragraph id="356536CFFFDB5118FF57FC09FDC8FBE8" blockId="21.[151,1437,151,1077]" pageId="21" pageNumber="22">
<emphasis id="07AEEADDFFDB5118FF57FC09FD45FC0A" bold="true" box="[199,334,980,1005]" pageId="21" pageNumber="22">Etymology.</emphasis>
The name refers to Laferte's original taxon with insufficient description (
<taxonomicName id="F2DA4D4CFFDB5118FB44FC08F95EFC0B" authorityName="Häckel" authorityYear="2017" baseAuthorityName="Chaudoir" baseAuthorityYear="1861" box="[1236,1365,980,1004]" class="Insecta" family="Carabidae" genus="Craspedophorus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="ruficrus">
<emphasis id="07AEEADDFFDB5118FB44FC08F95EFC0B" box="[1236,1365,980,1004]" italics="true" pageId="21" pageNumber="22">C. ruficrus</emphasis>
</taxonomicName>
) here considered
<emphasis id="07AEEADDFFDB5118FE87FC24FDB6FBF7" box="[279,445,1015,1040]" italics="true" pageId="21" pageNumber="22">nomen dubium</emphasis>
.
</paragraph>
</subSubSection>
<subSubSection id="7DC06544FFDB5118FF57FBC1FE5AFBD3" box="[199,593,1052,1077]" pageId="21" pageNumber="22" type="distribution">
<paragraph id="356536CFFFDB5118FF57FBC1FE5AFBD3" blockId="21.[151,1437,151,1077]" box="[199,593,1052,1077]" pageId="21" pageNumber="22">
<emphasis id="07AEEADDFFDB5118FF57FBC1FD6BFBD2" bold="true" box="[199,352,1052,1077]" pageId="21" pageNumber="22">Distribution.</emphasis>
<collectingCountry id="4DCD765FFFDB5118FEF7FBC1FDB0FBD2" box="[359,443,1052,1077]" name="Guinea-Bissau" pageId="21" pageNumber="22">Guinea</collectingCountry>
,
<collectingCountry id="4DCD765FFFDB5118FE56FBC0FE46FBD3" box="[454,589,1052,1077]" name="Ivory Coast" pageId="21" pageNumber="22">
Ivory
<collectingRegion id="F71EF82DFFDB5118FD9AFBC1FE46FBD3" box="[522,589,1052,1076]" country="Tanzania" name="Pwani" pageId="21" pageNumber="22">Coast</collectingRegion>
</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>