168 lines
15 KiB
XML
168 lines
15 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.205.3491" ID-GBIF-Dataset="16f08073-854b-40a2-aef9-72a97c85c8bd" ID-PMC="PMC3391731" ID-Pensoft-Pub="1313-2970-205-59" ID-PubMed="22792035" ModsDocAuthor="" ModsDocDate="2012" ModsDocID="1313-2970-205-59" ModsDocOrigin="ZooKeys 205" ModsDocTitle="A review of the spider genus Haplodrassus Chamberlin, 1922 in Crimea (Ukraine) and adjacent areas (Araneae, Gnaphosidae)" checkinTime="1451248915768" checkinUser="pensoft" docAuthor="Kovblyuk, Mykola M., Kastrygina, Zoya A. & Omelko, Mikhail M." docDate="2012" docId="2FA3B5621D53E9C128783A5E24357EEF" docLanguage="en" docName="ZooKeys 205: 59-89" docOrigin="ZooKeys 205" docSource="http://dx.doi.org/10.3897/zookeys.205.3491" docTitle="Haplodrassus isaevi Ponomarev & Tsvetkov 2006" docType="treatment" docVersion="3" lastPageNumber="65" masterDocId="FFC2BD6FFFD62C69FFC3EC55350AFFA3" masterDocTitle="A review of the spider genus Haplodrassus Chamberlin, 1922 in Crimea (Ukraine) and adjacent areas (Araneae, Gnaphosidae)" masterLastPageNumber="89" masterPageNumber="59" pageNumber="65" updateTime="1668154130483" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>A review of the spider genus Haplodrassus Chamberlin, 1922 in Crimea (Ukraine) and adjacent areas (Araneae, Gnaphosidae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Kovblyuk, Mykola M.</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Kastrygina, Zoya A.</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Omelko, Mikhail M.</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2012</mods:date>
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<mods:detail type="volume">
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<mods:number>205</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>59</mods:start>
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<mods:end>89</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.205.3491</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.205.3491</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-205-59</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="152036251" LSID="urn:lsid:plazi:treatment:2FA3B5621D53E9C128783A5E24357EEF" httpUri="http://treatment.plazi.org/id/2FA3B5621D53E9C128783A5E24357EEF" lastPageId="7" lastPageNumber="65" pageId="6" pageNumber="65">
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<subSubSection pageId="6" pageNumber="65" type="nomenclature">
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<paragraph pageId="6" pageNumber="65">
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<taxonomicName LSID="http://species-id.net/wiki/Haplodrassus_isaevi" authority="Ponomarev & Tsvetkov, 2006" authorityName="Ponomarev & Tsvetkov" authorityYear="2006" class="Arachnida" family="Gnaphosidae" genus="Haplodrassus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Haplodrassus isaevi" order="Araneae" pageId="6" pageNumber="65" phylum="Arthropoda" rank="species" species="isaevi">
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<pageBreakToken pageId="6" pageNumber="65" start="start">Haplodrassus</pageBreakToken>
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isaevi Ponomarev & Tsvetkov, 2006
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</taxonomicName>
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Figs 25
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<normalizedToken originalValue="–2730–">-2730-</normalizedToken>
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31
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</paragraph>
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</subSubSection>
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<subSubSection pageId="6" pageNumber="65" type="reference_group">
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<paragraph pageId="6" pageNumber="65">
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<taxonomicName class="Arachnida" family="Gnaphosidae" genus="Haplodrassus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Haplodrassus isaevi" order="Araneae" pageId="6" pageNumber="65" phylum="Arthropoda" rank="species" species="isaevi">Haplodrassus isaevi</taxonomicName>
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Ponomarev & Tsvetkov, 2006: 9, f. 12-14 (♂♀).
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</paragraph>
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<paragraph pageId="6" pageNumber="65">
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<taxonomicName class="Arachnida" family="Gnaphosidae" genus="Haplodrassus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Haplodrassus isaevi" order="Araneae" pageId="6" pageNumber="65" phylum="Arthropoda" rank="species" species="isaevi">Haplodrassus isaevi</taxonomicName>
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:
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<bibRefCitation author="Piterkina, TV" journalOrPublisher="Zoologicheskyi Zhurnal" pageId="14" pageNumber="73" pagination="1425 - 1432" title="Interesting findings of spiders of the families Gnaphosidae and Liocranidae (Aranei) from semi-desert of the northern Caspian lowland." volume="86" year="2007">Piterkina and Ovtsharenko 2007</bibRefCitation>
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: 1426, f. 1.1-6 (♂♀).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="6" pageNumber="65" type="records from crimea">
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<paragraph pageId="6" pageNumber="65">Records from Crimea.</paragraph>
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<paragraph pageId="6" pageNumber="65">
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<bibRefCitation author="Kovblyuk, MM" journalOrPublisher="N. Orianda, Simpheropol" pageId="13" pageNumber="72" title="Brief atlas of spiders (Arachnida, Aranei) of Karadag Nature Reserve." year="2008">Kovblyuk et al. (2008)</bibRefCitation>
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;
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<bibRefCitation author="Kovblyuk, MM" journalOrPublisher="Karadag Nature Reserve of Ukrainian Academy of Sciences," pageId="13" pageNumber="72" pagination="162 - 166" title="Arachnida. New records of spiders (Arachnida: Aranei) from Karadag." volume="24" year="2009">Kovblyuk et al. (2009)</bibRefCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="6" pageNumber="65" type="type material">
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<paragraph pageId="6" pageNumber="65">Type material.</paragraph>
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<paragraph pageId="6" pageNumber="65">RUSSIA, ROSTOV AREA: 3 ♂♂ paratypes (TNU from CP 18.24.8), Orlovskyi Distr., Rostov Reserve, 6.10.2002, A.V. Ponomarev.</paragraph>
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</subSubSection>
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<subSubSection pageId="6" pageNumber="65" type="material">
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<paragraph pageId="6" pageNumber="65">Material.</paragraph>
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<paragraph pageId="6" pageNumber="65">UKRAINE, CRIMEA: Sudak Distr.: 2 ♂♂, 3 ♀♀ (TNU), 10 km W Sudak, Mezhdurechie Vill., 3.10.-7.11.2010, A.K. Yusufova. Feodosiya Distr.: 21 ♂♂, 10 ♀♀ (TNU), Karadag Nature Reserve, 28.05.2003-19.12.2008, M.M. Kovblyuk, O.V. Kukushkin.</paragraph>
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</subSubSection>
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<subSubSection pageId="6" pageNumber="65" type="diagnosis">
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<paragraph pageId="6" pageNumber="65">Diagnosis.</paragraph>
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<paragraph pageId="6" pageNumber="65">
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<taxonomicName class="Arachnida" family="Gnaphosidae" genus="Haplodrassus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Haplodrassus isaevi" order="Araneae" pageId="6" pageNumber="65" phylum="Arthropoda" rank="species" species="isaevi">Haplodrassus isaevi</taxonomicName>
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is most similar to
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<taxonomicName class="Arachnida" family="Gnaphosidae" genus="Haplodrassus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Haplodrassus dalmatensis" order="Araneae" pageId="6" pageNumber="65" phylum="Arthropoda" rank="species" species="dalmatensis">Haplodrassus dalmatensis</taxonomicName>
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but differs by the shape of the terminal apophysis, which has only one tooth-like process (two tooth-like apical processes in
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<taxonomicName class="Arachnida" family="Gnaphosidae" genus="Haplodrassus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Haplodrassus dalmatensis" order="Araneae" pageId="6" pageNumber="65" phylum="Arthropoda" rank="species" species="dalmatensis">Haplodrassus dalmatensis</taxonomicName>
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), in lacking a tooth on the embolus (embolic tooth present in
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<taxonomicName class="Arachnida" family="Gnaphosidae" genus="Haplodrassus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Haplodrassus dalmatensis" order="Araneae" pageId="6" pageNumber="65" phylum="Arthropoda" rank="species" species="dalmatensis">Haplodrassus dalmatensis</taxonomicName>
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), and also by the proportions of the epigyne. Differences also occur in the spination of certain leg segments: male metatarsus I with two ventral spines in
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<taxonomicName class="Arachnida" family="Gnaphosidae" genus="Haplodrassus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Haplodrassus isaevi" order="Araneae" pageId="6" pageNumber="65" phylum="Arthropoda" rank="species" species="isaevi">Haplodrassus isaevi</taxonomicName>
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, but without spines in
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<taxonomicName class="Arachnida" family="Gnaphosidae" genus="Haplodrassus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Haplodrassus dalmatensis" order="Araneae" pageId="6" pageNumber="65" phylum="Arthropoda" rank="species" species="dalmatensis">Haplodrassus dalmatensis</taxonomicName>
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; female metatarsus IV with 4-5 retrolateral spines in
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<taxonomicName class="Arachnida" family="Gnaphosidae" genus="Haplodrassus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Haplodrassus isaevi" order="Araneae" pageId="6" pageNumber="65" phylum="Arthropoda" rank="species" species="isaevi">Haplodrassus isaevi</taxonomicName>
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, but with 3 spines in
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<taxonomicName class="Arachnida" family="Gnaphosidae" genus="Haplodrassus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Haplodrassus dalmatensis" order="Araneae" pageId="6" pageNumber="65" phylum="Arthropoda" rank="species" species="dalmatensis">Haplodrassus dalmatensis</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="6" pageNumber="65" type="description">
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<paragraph pageId="6" pageNumber="65">Description.</paragraph>
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<paragraph pageId="6" pageNumber="65">Males (n =5) and females (n = 5). Measurements (♂ / ♀): total length 5.4-7.2 (6.4) / 5.5-7.4 (6.3); carapace 2.3-2.9 (2.7) / 2.4-3.0 (2.7) long, 1.9-2.4 (2.2) / 1.9-2.2 (2.0) wide; abdomen 3.1-4.3 (3.7) / 3.0-4.4 (3.7) long, 1.7-2.3 (2.0) / 1.8-2.8 (2.3) wide.</paragraph>
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<paragraph pageId="6" pageNumber="65">Length of leg segments:</paragraph>
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<paragraph pageId="6" pageNumber="65">
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<table pageId="6" pageNumber="65">
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<tr pageId="6" pageNumber="65">
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<th colspan="1" pageId="6" pageNumber="65" rowspan="1">Leg</th>
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<th colspan="1" pageId="6" pageNumber="65" rowspan="1">Femur</th>
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<th colspan="1" pageId="6" pageNumber="65" rowspan="1">Patella</th>
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<th colspan="1" pageId="6" pageNumber="65" rowspan="1">Tibia</th>
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<th colspan="1" pageId="6" pageNumber="65" rowspan="1">Metatarsus</th>
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<th colspan="1" pageId="6" pageNumber="65" rowspan="1">Tarsus</th>
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<th colspan="1" pageId="6" pageNumber="65" rowspan="1">Total</th>
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</tr>
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</table>
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</paragraph>
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<paragraph pageId="6" pageNumber="65">Length of palp segments (male / female): femur 1.0-1.2 (1.1) / 0.8-1.1 (1.0), patella 0.4-0.5 (0.4) / 0.4-0.5 (0.5), tibia 0.3-0.4 (0.3) / 0.3-0.4 (0.4), tarsus 1.0-1.2 (1.0) / 0.6-0.7 (0.7).</paragraph>
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<paragraph pageId="6" pageNumber="65">Chelicerae with 2-3 promarginal and 2 retromarginal teeth in males and females. Number of promarginal teeth varies from 3 (most common) to 2 (seldom). One female studied had one chelicera with 3 and the other chelicera with 2 promarginal teeth. Coloration grey.</paragraph>
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<paragraph pageId="6" pageNumber="65">Male palp as in Figs 25-27. Terminal apophysis sharply turned, without ridge, but with tooth (Tt) in subterminal part, embolus without tooth.</paragraph>
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<paragraph pageId="6" pageNumber="65">Epigyne as in Figs 30-31. Fovea long, lateral pockets slightly converging, foveal width less that spermathecal span.</paragraph>
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</subSubSection>
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<subSubSection pageId="6" pageNumber="65" type="distribution">
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<paragraph pageId="6" pageNumber="65">Distribution.</paragraph>
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<paragraph pageId="6" pageNumber="65">
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Greece, Ukraine (Crimea), Russia (Rostov Area), Kazakhstan (West-Kazakhstan Area) (
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<bibRefCitation author="Ponomarev, AV" journalOrPublisher="Caucasian Entomological Bulletin" pageId="14" pageNumber="73" pagination="5 - 13" title="New and rare spiders of family Gnaphosidae (Aranei) from a southeast of Europe." volume="2" year="2006">Ponomarev and Tsvetkov 2006</bibRefCitation>
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;
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<bibRefCitation author="Piterkina, TV" journalOrPublisher="Zoologicheskyi Zhurnal" pageId="14" pageNumber="73" pagination="1425 - 1432" title="Interesting findings of spiders of the families Gnaphosidae and Liocranidae (Aranei) from semi-desert of the northern Caspian lowland." volume="86" year="2007">Piterkina and Ovtsharenko 2007</bibRefCitation>
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;
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<bibRefCitation pageId="6" pageNumber="65">Platnick 2012</bibRefCitation>
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; present data).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="6" pageNumber="65" type="habitats">
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<paragraph pageId="6" pageNumber="65">Habitats.</paragraph>
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<paragraph pageId="6" pageNumber="65">Steppes.</paragraph>
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</subSubSection>
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<subSubSection pageId="6" pageNumber="65" type="phenology">
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<paragraph pageId="6" pageNumber="65">Phenology.</paragraph>
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<paragraph pageId="6" pageNumber="65">In Crimea ♂♀ - X-XII, ♀♀ - II-III, V, the peak activity of adults occurs in December.</paragraph>
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</subSubSection>
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<subSubSection lastPageId="7" lastPageNumber="66" pageId="6" pageNumber="65" type="comments">
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<paragraph pageId="6" pageNumber="65">Comments.</paragraph>
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<paragraph lastPageId="7" lastPageNumber="66" pageId="6" pageNumber="65">
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In Crimea we found both closely related species,
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<taxonomicName class="Arachnida" family="Gnaphosidae" genus="Haplodrassus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Haplodrassus dalmatensis" order="Araneae" pageId="6" pageNumber="65" phylum="Arthropoda" rank="species" species="dalmatensis">Haplodrassus dalmatensis</taxonomicName>
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and
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<taxonomicName class="Arachnida" family="Gnaphosidae" genus="Haplodrassus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Haplodrassus isaevi" order="Araneae" pageId="6" pageNumber="65" phylum="Arthropoda" rank="species" species="isaevi">Haplodrassus isaevi</taxonomicName>
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, to be syntopical in two localities (Sudak Distr., 10 km W Sudak, Mezhdurechie Vill. and Feodosiya Distr., Karadag Nature Reserve). However, these species have quite different phenologies and adults of the two different sp
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<pageBreakToken pageId="7" pageNumber="66" start="start">ecies</pageBreakToken>
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do not co-occur. The reproductive period of
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<taxonomicName class="Arachnida" family="Gnaphosidae" genus="Haplodrassus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Haplodrassus dalmatensis" order="Araneae" pageId="7" pageNumber="66" phylum="Arthropoda" rank="species" species="dalmatensis">Haplodrassus dalmatensis</taxonomicName>
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is in May-July with the peak in June, and in
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<taxonomicName class="Arachnida" family="Gnaphosidae" genus="Haplodrassus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Haplodrassus isaevi" order="Araneae" pageId="7" pageNumber="66" phylum="Arthropoda" rank="species" species="isaevi">Haplodrassus isaevi</taxonomicName>
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adults can be found in October-December, with their peak of activity in December.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |