287 lines
40 KiB
XML
287 lines
40 KiB
XML
<document id="989DDE0D19A21E265FF23B85490DB31A" ID-CLB-Dataset="30462" ID-DOI="10.11646/zootaxa.4399.1.12" ID-GBIF-Dataset="98a2a3b6-d2b0-4ab8-ae02-af89677449ca" ID-ISSN="1175-5326" ID-Zenodo-Dep="1206426" ID-ZooBank="13F6805A-830B-404F-BC9A-E13262AD4834" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1521898234142" checkinUser="plazi" docAuthor="Jałoszyński, Paweł" docDate="2018" docId="BD0B8784FFBB6903BCFEF8B5793DFE71" docLanguage="en" docName="zootaxa.4399.1.12.pdf" docOrigin="Zootaxa 4399 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Clidicus asymmetricus Jałoszyński 2018, sp. n." docType="treatment" docVersion="7" lastPageNumber="145" masterDocId="4132FFFCFFBB6907BC69FFB37D02FFB5" masterDocTitle="A new species of Clidicus with asymmetrical parameres (Coleoptera, Staphylinidae, Scydmaeninae)" masterLastPageNumber="145" masterPageNumber="141" pageNumber="141" updateTime="1698532579982" updateUser="ExternalLinkService">
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<mods:title id="B4A5E8EE2B42D71EAD0882F660DA7D13">A new species of Clidicus with asymmetrical parameres (Coleoptera, Staphylinidae, Scydmaeninae)</mods:title>
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<mods:name id="3EF4697CE3506E34B9185A9A5748443A" type="personal">
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<mods:namePart id="1F294C1F75089E84CFC74B2FB49323C0">Jałoszyński, Paweł</mods:namePart>
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<mods:title id="C7EAA6A55896A36412D06BB1C76DDE7A">Zootaxa</mods:title>
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<mods:part id="EB606F522FFE136F8A09F275A086C366">
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<mods:date id="E8AA0E020BA634B1A494217FD0BD7367">2018</mods:date>
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<mods:number id="19F2CAC1C8B53330B3E3D1BA158911CD">2018-03-20</mods:number>
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<treatment id="BD0B8784FFBB6903BCFEF8B5793DFE71" ID-DOI="http://doi.org/10.5281/zenodo.5958466" ID-GBIF-Taxon="142524159" ID-Zenodo-Dep="5958466" LSID="urn:lsid:plazi:treatment:BD0B8784FFBB6903BCFEF8B5793DFE71" httpUri="http://treatment.plazi.org/id/BD0B8784FFBB6903BCFEF8B5793DFE71" lastPageId="4" lastPageNumber="145" pageId="0" pageNumber="141">
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<subSubSection id="7DB86519FFBB6907BCFEF8B57F82F895" box="[151,640,1798,1824]" pageId="0" pageNumber="141" type="nomenclature">
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<paragraph id="351D3692FFBB6907BCFEF8B57F82F895" blockId="0.[151,1436,1764,2029]" box="[151,640,1798,1824]" pageId="0" pageNumber="141">
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<heading id="6E5581FEFFBB6907BCFEF8B57F82F895" bold="true" box="[151,640,1798,1824]" fontSize="11" level="1" pageId="0" pageNumber="141" reason="1">
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<emphasis id="07D6EA80FFBB6907BCFEF8B57F82F895" bold="true" box="[151,640,1798,1824]" pageId="0" pageNumber="141">
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<taxonomicName id="F2A24D11FFBB6907BCFEF8B57CA5F895" ID-CoL="W5G4" authority="Jałoszyński, 2018" authorityName="Jałoszyński" authorityYear="2018" box="[151,423,1798,1824]" class="Insecta" family="Staphylinidae" genus="Clidicus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="0" pageNumber="141" phylum="Arthropoda" rank="species" species="asymmetricus" status="sp. nov.">
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<emphasis id="07D6EA80FFBB6907BCFEF8B57CA5F895" bold="true" box="[151,423,1798,1824]" italics="true" pageId="0" pageNumber="141">Clidicus asymmetricus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="1CE557FBFFBB6907BDC6F8B57CF3F895" box="[431,497,1798,1824]" pageId="0" pageNumber="141" rank="species">sp. n.</taxonomicNameLabel>
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(
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<figureCitation id="AD992A17FFBB6907BE69F8B57F75F895" box="[512,631,1798,1824]" captionStart-0="FIGURES1–2" captionStart-1="FIGURES3–6" captionStart-2="FIGURES7–11" captionStartId-0="1.[151,258,1773,1795]" captionStartId-1="2.[151,258,1973,1995]" captionStartId-2="3.[151,258,1242,1264]" captionTargetBox-0="[171,1418,414,1735]" captionTargetBox-1="[151,1436,426,1960]" captionTargetBox-2="[155,1426,210,1220]" captionTargetId-0="figure@1.[151,1436,399,1752]" captionTargetId-1="figure@2.[151,1436,426,1960]" captionTargetId-2="figure@3.[151,1436,191,1220]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionTargetPageId-2="3" captionText-0="FIGURES1–2.Clidicus of the Philippines,dorsal habitus of males.Clidicus asymmetricus sp.n. (1); C.crocodylus Jałoszyński (2)." captionText-1="FIGURES3–6. Clidicus ofthe Philippines,pronotum and head of males (pronotumin dorsalview,head slightly tilted ventrad) (3, 5), andhead indorsal view(4, 6).Clidicus asymmetricus sp.n.(3–4); C. crocodylus Jałoszyński (5–6)." captionText-2="FIGURES7–11. Clidicus asymmetricus sp.n.Aedeagus inventral (7–8) andlateral (9–10) views, and'spermpump' (11)." httpUri-0="https://zenodo.org/record/1206428/files/figure.png" httpUri-1="https://zenodo.org/record/1206430/files/figure.png" httpUri-2="https://zenodo.org/record/1206432/files/figure.png" pageId="0" pageNumber="141">
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Figs
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<docIssue id="71C1497DFFBB6907BE54F8B57F4BF895" box="[573,585,1798,1824]" pageId="0" pageNumber="141">1</docIssue>
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–11
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</figureCitation>
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)
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="7DB86519FFBB6907BCFEF89A7F8BF81D" pageId="0" pageNumber="141" type="materials_examined">
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<paragraph id="351D3692FFBB6907BCFEF89A7F8BF81D" blockId="0.[151,1436,1764,2029]" pageId="0" pageNumber="141">
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<emphasis id="07D6EA80FFBB6907BCFEF89A7C56F8F5" bold="true" box="[151,340,1833,1856]" pageId="0" pageNumber="141">Material studied</emphasis>
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.
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<emphasis id="07D6EA80FFBB6907BD08F89A7E7AF8F5" bold="true" box="[353,888,1833,1856]" pageId="0" pageNumber="141">Holotype: male (PHILIPPINES, MINDANAO)</emphasis>
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: three labels: "PHILIPPINES / 07°39'N 124°42'E / Mindanao, prov. Lanao de Sul / Wao /
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<date id="411C1052FFBB6907BE5BF8FF7F7CF8D6" box="[562,638,1867,1891]" pageId="0" pageNumber="141" value="2017-05">
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<emphasis id="07D6EA80FFBB6907BE5BF8FF7F7CF8D6" box="[562,638,1867,1891]" italics="true" pageId="0" pageNumber="141">V 2017</emphasis>
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</date>
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/ leg. Ismael Lumawig(local coll.)" [white, printed], "5960 / 156526 / coll. Upper Silesian Museum / (USMB) Bytom, Poland" [white, printed] "
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<taxonomicName id="F2A24D11FFBB6907BFE0F8DE7EE0F830" authorityName="Laporte" authorityYear="1832" box="[905,994,1901,1925]" class="Insecta" family="Staphylinidae" genus="Clidicus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="0" pageNumber="141" phylum="Arthropoda" rank="genus">
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<emphasis id="07D6EA80FFBB6907BFE0F8DE7EE0F830" box="[905,994,1901,1925]" italics="true" pageId="0" pageNumber="141">Clidicus</emphasis>
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</taxonomicName>
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/
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<taxonomicName id="F2A24D11FFBB6907BF93F8DC798EF833" authorityName="Jałoszyński" authorityYear="2018" box="[1018,1164,1903,1926]" class="Insecta" family="Staphylinidae" genus="Clidicus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="0" pageNumber="141" phylum="Arthropoda" rank="species" species="asymmetricus">
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<emphasis id="07D6EA80FFBB6907BF93F8DC798EF833" box="[1018,1164,1903,1926]" italics="true" pageId="0" pageNumber="141">asymmetricus</emphasis>
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</taxonomicName>
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m. / P. JAŁOSZYŃSKI,
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<docYear id="2DC4DA69FFBB6907BCFEF8227DCEF81D" box="[151,204,1937,1960]" pageId="0" pageNumber="141">2018</docYear>
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/ HOLOTYPUS" [red, printed] (USMB).
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</paragraph>
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</subSubSection>
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<subSubSection id="7DB86519FFBB6906BCAEF8077CCCFF41" lastPageId="1" lastPageNumber="142" pageId="0" pageNumber="141" type="diagnosis">
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<paragraph id="351D3692FFBB6906BCAEF8077CCCFF41" blockId="0.[151,1436,1764,2029]" lastBlockId="1.[151,1436,152,348]" lastPageId="1" lastPageNumber="142" pageId="0" pageNumber="141">
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<emphasis id="07D6EA80FFBB6907BCAEF8077C3EF87E" bold="true" box="[199,316,1972,1995]" pageId="0" pageNumber="141">Diagnosis.</emphasis>
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Each of antennomeres II–XI distinctly elongate; anterior margin of labrum with three shallow and broad subtriangular emarginations; head slightly narrower than pronotum; tempora evenly rounded; aedeagus very slender and in lateral view strongly curved, one paramere much longer than the other and with its apical region strongly flattened laterally, in lateral view parameral apex broadened and curved ventrad; 'funnels' of 'sperm pump' equal in diameter, each very shallow and very broad.
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</paragraph>
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</subSubSection>
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<subSubSection id="7DB86519FFBA6904BCAEFEB37C9AF9DC" lastPageId="3" lastPageNumber="144" pageId="1" pageNumber="142" type="description">
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<paragraph id="351D3692FFBA6906BCAEFEB37C64FEE9" blockId="1.[151,1436,152,348]" pageId="1" pageNumber="142">
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<emphasis id="07D6EA80FFBA6906BCAEFEB37C50FEA2" bold="true" box="[199,338,256,279]" pageId="1" pageNumber="142">Description.</emphasis>
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Body of male (
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<figureCitation id="AD992A17FFBA6906BD93FEB37F3AFEA2" box="[506,568,256,279]" captionStart="FIGURES1–2" captionStartId="1.[151,258,1773,1795]" captionTargetBox="[171,1418,414,1735]" captionTargetId="figure@1.[151,1436,399,1752]" captionTargetPageId="1" captionText="FIGURES1–2.Clidicus of the Philippines,dorsal habitus of males.Clidicus asymmetricus sp.n. (1); C.crocodylus Jałoszyński (2)." httpUri="https://zenodo.org/record/1206428/files/figure.png" pageId="1" pageNumber="142">Fig. 1</figureCitation>
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) elongate and slender, head flattened, prothorax and elytra strongly convex; legs and antennae long and slender, BL
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<quantity id="F25A9B77FFBA6906BE75FE917F83FE8C" box="[540,641,290,313]" metricMagnitude="-3" metricUnit="m" metricValue="6.05" pageId="1" pageNumber="142" unit="mm" value="6.05">6.05 mm</quantity>
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; cuticle moderately glossy, pigmentation uniformly brown, setae slightly lighter than cuticle.
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</paragraph>
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<caption id="61DD661AFFBA6906BCFEF95E7890F8B6" box="[151,1426,1773,1795]" httpUri="https://zenodo.org/record/1206428/files/figure.png" pageId="1" pageNumber="142" startId="1.[151,258,1773,1795]" targetBox="[171,1418,414,1735]" targetPageId="1">
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<paragraph id="351D3692FFBA6906BCFEF95E7890F8B6" blockId="1.[151,1426,1773,1795]" box="[151,1426,1773,1795]" pageId="1" pageNumber="142">
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<emphasis id="07D6EA80FFBA6906BCFEF95E7C30F8B6" bold="true" box="[151,306,1773,1795]" pageId="1" pageNumber="142">FIGURES 1–2.</emphasis>
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<taxonomicName id="F2A24D11FFBA6906BD5EF95E7C85F8B7" authorityName="Laporte" authorityYear="1832" box="[311,391,1773,1794]" class="Insecta" family="Staphylinidae" genus="Clidicus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="142" phylum="Arthropoda" rank="genus">
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<emphasis id="07D6EA80FFBA6906BD5EF95E7C85F8B7" box="[311,391,1773,1794]" italics="true" pageId="1" pageNumber="142">Clidicus</emphasis>
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</taxonomicName>
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of the Philippines, dorsal habitus of males.
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<taxonomicName id="F2A24D11FFBA6906BF45F95E7907F8B7" authority="Jałoszyński, 2018" authorityName="Jałoszyński" authorityYear="2018" box="[812,1029,1773,1794]" class="Insecta" family="Staphylinidae" genus="Clidicus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="142" phylum="Arthropoda" rank="species" species="asymmetricus" status="sp. nov.">
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<emphasis id="07D6EA80FFBA6906BF45F95E7907F8B7" box="[812,1029,1773,1794]" italics="true" pageId="1" pageNumber="142">Clidicus asymmetricus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="1CE557FBFFBA6906B865F95E7940F8B6" box="[1036,1090,1773,1795]" pageId="1" pageNumber="142" rank="species">
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<emphasis id="07D6EA80FFBA6906B865F95E7940F8B6" bold="true" box="[1036,1090,1773,1795]" pageId="1" pageNumber="142">sp. n.</emphasis>
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</taxonomicNameLabel>
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(1);
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<taxonomicName id="F2A24D11FFBA6906B819F95E786BF8B6" authority="Jaloszynski" authorityName="Jaloszynski" box="[1136,1385,1773,1795]" class="Insecta" family="Staphylinidae" genus="Clidicus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="142" phylum="Arthropoda" rank="species" species="crocodylus">
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<emphasis id="07D6EA80FFBA6906B819F95E79F6F8B7" box="[1136,1268,1773,1794]" italics="true" pageId="1" pageNumber="142">C. crocodylus</emphasis>
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Jałoszyński
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</taxonomicName>
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(2).
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</paragraph>
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</caption>
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<paragraph id="351D3692FFBA6905BCAEF89E7E8CFF41" blockId="1.[151,1436,1837,2034]" lastBlockId="2.[151,1436,152,383]" lastPageId="2" lastPageNumber="143" pageId="1" pageNumber="142">
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Head (
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<figureCitation id="AD992A17FFBA6906BD66F89E7C72F8F1" box="[271,368,1837,1860]" captionStart="FIGURES3–6" captionStartId="2.[151,258,1973,1995]" captionTargetBox="[151,1436,426,1960]" captionTargetId="figure@2.[151,1436,426,1960]" captionTargetPageId="2" captionText="FIGURES3–6. Clidicus ofthe Philippines,pronotum and head of males (pronotumin dorsalview,head slightly tilted ventrad) (3, 5), andhead indorsal view(4, 6).Clidicus asymmetricus sp.n.(3–4); C. crocodylus Jałoszyński (5–6)." httpUri="https://zenodo.org/record/1206430/files/figure.png" pageId="1" pageNumber="142">Figs 3–4</figureCitation>
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) broadest shortly behind eyes, HL
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<quantity id="F25A9B77FFBA6906BE98F89E7E57F8F1" box="[753,853,1837,1860]" metricMagnitude="-4" metricUnit="m" metricValue="9.8" pageId="1" pageNumber="142" unit="mm" value="0.98">0.98 mm</quantity>
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, HW
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<quantity id="F25A9B77FFBA6906BFFFF89E7EF8F8F1" box="[918,1018,1837,1860]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="1" pageNumber="142" unit="mm" value="1.4">1.40 mm</quantity>
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; tempora in dorsal view (
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<figureCitation id="AD992A17FFBA6906B97BF89E7856F8F1" box="[1298,1364,1837,1860]" captionStart="FIGURES3–6" captionStartId="2.[151,258,1973,1995]" captionTargetBox="[151,1436,426,1960]" captionTargetId="figure@2.[151,1436,426,1960]" captionTargetPageId="2" captionText="FIGURES3–6. Clidicus ofthe Philippines,pronotum and head of males (pronotumin dorsalview,head slightly tilted ventrad) (3, 5), andhead indorsal view(4, 6).Clidicus asymmetricus sp.n.(3–4); C. crocodylus Jałoszyński (5–6)." httpUri="https://zenodo.org/record/1206430/files/figure.png" pageId="1" pageNumber="142">Fig. 4</figureCitation>
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) more than 4 times as long as eyes and evenly, strongly rounded; vertex with deep posteromedian impression and well-visible pits marking the sites where dorsal tentorial arms are fused with the head wall, sides of vertex convex; frons between antennal insertions slightly impressed, both frons and vertex with narrow median longitudinal groove, which is more pronounced posteriorly; supraantennal tubercles small but distinctly elevated; composite eyes small, nearly round, not projecting laterally from the head silhouette and relatively finely faceted. Punctures on vertex and frons distinct but shallow, except for supraantennal tubercles which are virtually impunctate, also median impression on vertex and median line on frons and vertex bear indistinct punctures; setae moderately long, sparse, suberect. Antennae (
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<figureCitation id="AD992A17FFB96905B8BEFF097814FF64" box="[1239,1302,186,209]" captionStart="FIGURES1–2" captionStartId="1.[151,258,1773,1795]" captionTargetBox="[171,1418,414,1735]" captionTargetId="figure@1.[151,1436,399,1752]" captionTargetPageId="1" captionText="FIGURES1–2.Clidicus of the Philippines,dorsal habitus of males.Clidicus asymmetricus sp.n. (1); C.crocodylus Jałoszyński (2)." httpUri="https://zenodo.org/record/1206428/files/figure.png" pageId="2" pageNumber="143">Fig. 1</figureCitation>
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) slender, all antennomeres elongate, scape slightly longer than head, AnL
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<quantity id="F25A9B77FFB96905BF41FF6E7E88FF41" box="[808,906,221,244]" metricMagnitude="-3" metricUnit="m" metricValue="4.18" pageId="2" pageNumber="143" unit="mm" value="4.18">4.18 mm</quantity>
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.
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</paragraph>
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<paragraph id="351D3692FFB96905BCAEFEB37890FECA" blockId="2.[151,1436,152,383]" pageId="2" pageNumber="143">
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Pronotum (
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<figureCitation id="AD992A17FFB96905BD56FEB37C7DFEA2" box="[319,383,256,279]" captionStart="FIGURES3–6" captionStartId="2.[151,258,1973,1995]" captionTargetBox="[151,1436,426,1960]" captionTargetId="figure@2.[151,1436,426,1960]" captionTargetPageId="2" captionText="FIGURES3–6. Clidicus ofthe Philippines,pronotum and head of males (pronotumin dorsalview,head slightly tilted ventrad) (3, 5), andhead indorsal view(4, 6).Clidicus asymmetricus sp.n.(3–4); C. crocodylus Jałoszyński (5–6)." httpUri="https://zenodo.org/record/1206430/files/figure.png" pageId="2" pageNumber="143">Fig. 3</figureCitation>
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) with nearly round disc and short posterior 'collar', broadest near anterior third; PL
|
||
<quantity id="F25A9B77FFB96905B96FFEB3786AFEA2" box="[1286,1384,256,279]" metricMagnitude="-3" metricUnit="m" metricValue="1.58" pageId="2" pageNumber="143" unit="mm" value="1.58">1.58 mm</quantity>
|
||
, PW
|
||
<quantity id="F25A9B77FFB96905BCFEFE917DFBFE8C" box="[151,249,290,313]" metricMagnitude="-3" metricUnit="m" metricValue="1.48" pageId="2" pageNumber="143" unit="mm" value="1.48">1.48 mm</quantity>
|
||
; 'collar' demarcated by narrow transverse groove with ten irregular and diffuse pits; additionally two larger pits are present on each side of pronotum in its constricted portion. Punctures on disc distinct but relatively small, shallow and diffuse, sides of pronotum (above each procoxal cavity) nearly impunctate; setae moderately long, sparse, suberect.
|
||
</paragraph>
|
||
<caption id="61DD661AFFB96905BCFEF8067951F85C" httpUri="https://zenodo.org/record/1206430/files/figure.png" pageId="2" pageNumber="143" startId="2.[151,258,1973,1995]" targetBox="[151,1436,426,1960]" targetPageId="2">
|
||
<paragraph id="351D3692FFB96905BCFEF8067951F85C" blockId="2.[151,1435,1973,2025]" pageId="2" pageNumber="143">
|
||
<emphasis id="07D6EA80FFB96905BCFEF8067C32F87E" bold="true" box="[151,304,1973,1995]" pageId="2" pageNumber="143">FIGURES 3–6.</emphasis>
|
||
<taxonomicName id="F2A24D11FFB96905BD5EF8057C85F87E" authorityName="Laporte" authorityYear="1832" box="[311,391,1974,1995]" class="Insecta" family="Staphylinidae" genus="Clidicus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="143" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="07D6EA80FFB96905BD5EF8057C85F87E" box="[311,391,1974,1995]" italics="true" pageId="2" pageNumber="143">Clidicus</emphasis>
|
||
</taxonomicName>
|
||
of the Philippines, pronotum and head of males (pronotum in dorsal view, head slightly tilted ventrad) (3, 5), and head in dorsal view (4, 6).
|
||
<taxonomicName id="F2A24D11FFB96905BDA9F8607F98F85D" authority="Jałoszyński, 2018" authorityName="Jałoszyński" authorityYear="2018" box="[448,666,2003,2024]" class="Insecta" family="Staphylinidae" genus="Clidicus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="143" phylum="Arthropoda" rank="species" species="asymmetricus" status="sp. nov.">
|
||
<emphasis id="07D6EA80FFB96905BDA9F8607F98F85D" box="[448,666,2003,2024]" italics="true" pageId="2" pageNumber="143">Clidicus asymmetricus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="1CE557FBFFB96905BEC9F8607FD6F85D" box="[672,724,2003,2024]" pageId="2" pageNumber="143" rank="species">sp. n.</taxonomicNameLabel>
|
||
(3–4);
|
||
<taxonomicName id="F2A24D11FFB96905BF73F8607910F85C" authority="Jaloszynski" authorityName="Jaloszynski" box="[794,1042,2003,2025]" class="Insecta" family="Staphylinidae" genus="Clidicus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="143" phylum="Arthropoda" rank="species" species="crocodylus">
|
||
<emphasis id="07D6EA80FFB96905BF73F8607E9CF85D" box="[794,926,2003,2024]" italics="true" pageId="2" pageNumber="143">C. crocodylus</emphasis>
|
||
Jałoszyński
|
||
</taxonomicName>
|
||
(5–6).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="61DD661AFFB86904BCFEFB69782FFB5A" box="[151,1325,1241,1264]" httpUri="https://zenodo.org/record/1206432/files/figure.png" pageId="3" pageNumber="144" startId="3.[151,258,1242,1264]" targetBox="[155,1426,210,1220]" targetPageId="3">
|
||
<paragraph id="351D3692FFB86904BCFEFB69782FFB5A" blockId="3.[151,1325,1241,1264]" box="[151,1325,1241,1264]" pageId="3" pageNumber="144">
|
||
<emphasis id="07D6EA80FFB86904BCFEFB697C39FB45" bold="true" box="[151,315,1242,1264]" pageId="3" pageNumber="144">FIGURES 7–11.</emphasis>
|
||
<taxonomicName id="F2A24D11FFB86904BD2BFB697F1EFB5A" authority="Jałoszyński, 2018" authorityName="Jałoszyński" authorityYear="2018" box="[322,540,1242,1263]" class="Insecta" family="Staphylinidae" genus="Clidicus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="144" phylum="Arthropoda" rank="species" species="asymmetricus" status="sp. nov.">
|
||
<emphasis id="07D6EA80FFB86904BD2BFB697F1EFB5A" box="[322,540,1242,1263]" italics="true" pageId="3" pageNumber="144">Clidicus asymmetricus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="1CE557FBFFB86904BE4BFB697F5AFB45" box="[546,600,1242,1264]" pageId="3" pageNumber="144" rank="species">
|
||
<emphasis id="07D6EA80FFB86904BE4BFB697F5AFB45" bold="true" box="[546,600,1242,1264]" pageId="3" pageNumber="144">sp. n.</emphasis>
|
||
</taxonomicNameLabel>
|
||
Aedeagus in ventral (7–8) and lateral (9–10) views, and 'sperm pump' (11).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="351D3692FFB86904BCAEFAA97E6CFAC3" blockId="3.[151,1436,1306,2022]" pageId="3" pageNumber="144">
|
||
Elytra oval, broadest clearly in front of middle, EL
|
||
<quantity id="F25A9B77FFB86904BE9EFAA97E58FA84" box="[759,858,1306,1329]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="3" pageNumber="144" unit="mm" value="3.5">3.50 mm</quantity>
|
||
, EW
|
||
<quantity id="F25A9B77FFB86904BFFEFAA97EF8FA84" box="[919,1018,1306,1329]" metricMagnitude="-3" metricUnit="m" metricValue="1.98" pageId="3" pageNumber="144" unit="mm" value="1.98">1.98 mm</quantity>
|
||
, EI 1.77; each elytron with five dorsal and one lateral rows of large and deep pits, cuticle between rows with very fine and sparse setiferous punctures; setae moderately long, denser than those on head and pronotum, suberect.
|
||
</paragraph>
|
||
<paragraph id="351D3692FFB86904BCAEFA317C97FA2C" blockId="3.[151,1436,1306,2022]" box="[199,405,1410,1433]" pageId="3" pageNumber="144">Hind wings absent.</paragraph>
|
||
<paragraph id="351D3692FFB86904BCAEFA167F42FA09" blockId="3.[151,1436,1306,2022]" box="[199,576,1445,1468]" pageId="3" pageNumber="144">Legs long and slender, unmodified.</paragraph>
|
||
<paragraph id="351D3692FFB86904BCAEFA747FECF9F3" blockId="3.[151,1436,1306,2022]" pageId="3" pageNumber="144">
|
||
Aedeagus (
|
||
<figureCitation id="AD992A17FFB86904BD2DFA747CB4FA6B" box="[324,438,1479,1502]" captionStart="FIGURES7–11" captionStartId="3.[151,258,1242,1264]" captionTargetBox="[155,1426,210,1220]" captionTargetId="figure@3.[151,1436,191,1220]" captionTargetPageId="3" captionText="FIGURES7–11. Clidicus asymmetricus sp.n.Aedeagus inventral (7–8) andlateral (9–10) views, and'spermpump' (11)." httpUri="https://zenodo.org/record/1206432/files/figure.png" pageId="3" pageNumber="144">Figs 7–10</figureCitation>
|
||
) strongly elongate, slender, AeL
|
||
<quantity id="F25A9B77FFB86904BF51FA747EA3FA6B" box="[824,929,1479,1502]" metricMagnitude="-3" metricUnit="m" metricValue="2.33" pageId="3" pageNumber="144" unit="mm" value="2.33">2.33 mm</quantity>
|
||
; parameres of unequal length, the short one truncated at apex, the long one with broadened apical region, which is curved ventrad; flagellum not differentiated from ductus ejaculatorius, not broadened and straight. 'Sperm pump' (
|
||
<figureCitation id="AD992A17FFB86904BF38F9BE7E9CF991" box="[849,926,1549,1572]" captionStart="FIGURES7–11" captionStartId="3.[151,258,1242,1264]" captionTargetBox="[155,1426,210,1220]" captionTargetId="figure@3.[151,1436,191,1220]" captionTargetPageId="3" captionText="FIGURES7–11. Clidicus asymmetricus sp.n.Aedeagus inventral (7–8) andlateral (9–10) views, and'spermpump' (11)." httpUri="https://zenodo.org/record/1206432/files/figure.png" pageId="3" pageNumber="144">Fig. 11</figureCitation>
|
||
)
|
||
<quantity id="F25A9B77FFB86904BFC6F9BE7912F991" box="[943,1040,1549,1572]" metricMagnitude="-4" metricUnit="m" metricValue="6.5" pageId="3" pageNumber="144" unit="mm" value="0.65">0.65 mm</quantity>
|
||
in length (measured between lateral edges of 'funnels') with large 'funnels' equal in diameter.
|
||
</paragraph>
|
||
<paragraph id="351D3692FFB86904BCAEF9E17C9AF9DC" blockId="3.[151,1436,1306,2022]" box="[199,408,1618,1641]" pageId="3" pageNumber="144">
|
||
<emphasis id="07D6EA80FFB86904BCAEF9E17C18F9DC" bold="true" box="[199,282,1618,1641]" pageId="3" pageNumber="144">Female</emphasis>
|
||
. Unknown.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="7DB86519FFB86904BCAEF9C67F2AF939" box="[199,552,1653,1676]" pageId="3" pageNumber="144" type="distribution">
|
||
<paragraph id="351D3692FFB86904BCAEF9C67F2AF939" blockId="3.[151,1436,1306,2022]" box="[199,552,1653,1676]" pageId="3" pageNumber="144">
|
||
<emphasis id="07D6EA80FFB86904BCAEF9C67C50F939" bold="true" box="[199,338,1653,1676]" pageId="3" pageNumber="144">Distribution</emphasis>
|
||
.
|
||
<collectingRegion id="F766F870FFB86904BD36F9C67CAFF939" box="[351,429,1653,1676]" country="Sri Lanka" name="Central" pageId="3" pageNumber="144">Central</collectingRegion>
|
||
Mindanao.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="7DB86519FFB86904BCAEF9247E41F91B" box="[199,835,1687,1710]" pageId="3" pageNumber="144" type="etymology">
|
||
<paragraph id="351D3692FFB86904BCAEF9247E41F91B" blockId="3.[151,1436,1306,2022]" box="[199,835,1687,1710]" pageId="3" pageNumber="144">
|
||
<emphasis id="07D6EA80FFB86904BCAEF9247C45F91B" bold="true" box="[199,327,1687,1710]" pageId="3" pageNumber="144">Etymology.</emphasis>
|
||
The name refers to the asymmetrical aedeagus.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="7DB86519FFB86903BCAEF909793DFE71" lastPageId="4" lastPageNumber="145" pageId="3" pageNumber="144" type="discussion">
|
||
<paragraph id="351D3692FFB86904BCAEF909789EF853" blockId="3.[151,1436,1306,2022]" pageId="3" pageNumber="144">
|
||
<emphasis id="07D6EA80FFB86904BCAEF9097C37F964" bold="true" box="[199,309,1722,1745]" pageId="3" pageNumber="144">Remarks.</emphasis>
|
||
<taxonomicName id="F2A24D11FFB86904BD29F90A7F35F964" authorityName="Jałoszyński" authorityYear="2018" box="[320,567,1721,1745]" class="Insecta" family="Staphylinidae" genus="Clidicus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="144" phylum="Arthropoda" rank="species" species="asymmetricus">
|
||
<emphasis id="07D6EA80FFB86904BD29F90A7F35F964" box="[320,567,1721,1745]" italics="true" pageId="3" pageNumber="144">Clidicus asymmetricus</emphasis>
|
||
</taxonomicName>
|
||
has a strikingly slender aedeagus with strongly asymmetrical parameres; this structure alone can be used to identify the species. Strongly asymmetrical parameres can also be found in
|
||
<taxonomicName id="F2A24D11FFB86904B94FF96E7C3AF8A3" authority="Orousset, 2014" authorityName="Orousset" authorityYear="2014" class="Insecta" family="Staphylinidae" genus="Clidicus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="144" phylum="Arthropoda" rank="species" species="minutus">
|
||
<emphasis id="07D6EA80FFB86904B94FF96E7899F941" box="[1318,1435,1757,1780]" italics="true" pageId="3" pageNumber="144">C. minutus</emphasis>
|
||
<bibRefCitation id="51334B63FFB86904BCFEF94C7C3AF8A3" box="[151,312,1791,1814]" pageId="3" pageNumber="144" refId="ref1753" refString="Orousset, J. (2014) Contribution a la connaissance du genre Clidicus Laporte de Castelnau, 1832 (Coleoptera, Staphylinidae, Scydmaeninae). Le Coleopteriste, 17, 116 - 135." type="journal article">Orousset, 2014</bibRefCitation>
|
||
</taxonomicName>
|
||
(aedeagus very stout, 'sperm pump' extremely elongate and lacking 'funnels'),
|
||
<taxonomicName id="F2A24D11FFB86904B816F94C789EF8A3" authority="Orousset, 2014" authorityName="Orousset" authorityYear="2014" box="[1151,1436,1791,1815]" class="Insecta" family="Staphylinidae" genus="Clidicus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="144" phylum="Arthropoda" rank="species" species="armipes">
|
||
<emphasis id="07D6EA80FFB86904B816F94C79F6F8A2" box="[1151,1268,1791,1815]" italics="true" pageId="3" pageNumber="144">C. armipes</emphasis>
|
||
<bibRefCitation id="51334B63FFB86904B892F94C789EF8A3" box="[1275,1436,1791,1814]" pageId="3" pageNumber="144" refId="ref1753" refString="Orousset, J. (2014) Contribution a la connaissance du genre Clidicus Laporte de Castelnau, 1832 (Coleoptera, Staphylinidae, Scydmaeninae). Le Coleopteriste, 17, 116 - 135." type="journal article">Orousset, 2014</bibRefCitation>
|
||
</taxonomicName>
|
||
(aedeagus moderately stout, protibiae with two large ventral preapical teeth), and
|
||
<taxonomicName id="F2A24D11FFB86904B86EF891782BF88C" authority="Orousset, 2014" authorityName="Orousset" authorityYear="2014" box="[1031,1321,1825,1849]" class="Insecta" family="Staphylinidae" genus="Clidicus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="144" phylum="Arthropoda" rank="species" species="dohertyi">
|
||
<emphasis id="07D6EA80FFB86904B86EF8917982F88C" box="[1031,1152,1825,1849]" italics="true" pageId="3" pageNumber="144">C. dohertyi</emphasis>
|
||
<bibRefCitation id="51334B63FFB86904B8EEF891782BF88C" box="[1159,1321,1826,1849]" pageId="3" pageNumber="144" refId="ref1753" refString="Orousset, J. (2014) Contribution a la connaissance du genre Clidicus Laporte de Castelnau, 1832 (Coleoptera, Staphylinidae, Scydmaeninae). Le Coleopteriste, 17, 116 - 135." type="journal article">Orousset, 2014</bibRefCitation>
|
||
</taxonomicName>
|
||
(aedeagus very stout with conspicuous subtriangular lateral projections, 'sperm pump' with 'funnels' of unequal length). Externally, it is most similar to the sympatric
|
||
<taxonomicName id="F2A24D11FFB86904BE6BF8D47E5FF8CB" authority="Jaloszynski, 2009" authorityName="Jaloszynski" authorityYear="2009" box="[514,861,1894,1918]" class="Insecta" family="Staphylinidae" genus="Clidicus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="144" phylum="Arthropoda" rank="species" species="crocodylus">
|
||
<emphasis id="07D6EA80FFB86904BE6BF8D47F97F8CB" box="[514,661,1894,1918]" italics="true" pageId="3" pageNumber="144">C. crocodylus</emphasis>
|
||
<bibRefCitation id="51334B63FFB86904BEF5F8D47E5FF8CB" box="[668,861,1895,1918]" pageId="3" pageNumber="144" refId="ref1615" refString="Jaloszynski, P. (2009) Clidicus crocodylus sp. nov. from Mindanao, the first representatives of the Mastiginae from the Philippines (Coleoptera, Scydmaenidae). Genus, 20 (2), 203 - 207." type="journal article">Jałoszyński, 2009</bibRefCitation>
|
||
</taxonomicName>
|
||
. The latter species has adults showing a variable form of elytra; in males the elytra can be nearly parallel-sided (as illustrated for the
|
||
<typeStatus id="EA198830FFB86904BF8CF8397940F814" box="[997,1090,1930,1953]" pageId="3" pageNumber="144" type="holotype">holotype</typeStatus>
|
||
in
|
||
<bibRefCitation id="51334B63FFB86904B80FF8397830F814" box="[1126,1330,1930,1953]" pageId="3" pageNumber="144" refId="ref1615" refString="Jaloszynski, P. (2009) Clidicus crocodylus sp. nov. from Mindanao, the first representatives of the Mastiginae from the Philippines (Coleoptera, Scydmaenidae). Genus, 20 (2), 203 - 207." type="journal article">Jałoszyński (2009)</bibRefCitation>
|
||
, fig. 7) to more rounded at sides, resembling those of females (an example of such a stout male is shown in
|
||
<figureCitation id="AD992A17FFB86904B8C7F81E79EDF871" box="[1198,1263,1965,1988]" captionStart="FIGURES1–2" captionStartId="1.[151,258,1773,1795]" captionTargetBox="[171,1418,414,1735]" captionTargetId="figure@1.[151,1436,399,1752]" captionTargetPageId="1" captionText="FIGURES1–2.Clidicus of the Philippines,dorsal habitus of males.Clidicus asymmetricus sp.n. (1); C.crocodylus Jałoszyński (2)." httpUri="https://zenodo.org/record/1206428/files/figure.png" pageId="3" pageNumber="144">Fig. 2</figureCitation>
|
||
). Independently of the variation in the elytral shape,
|
||
<taxonomicName id="F2A24D11FFB86904BE71F87C7FA9F853" box="[536,683,1998,2022]" class="Insecta" family="Staphylinidae" genus="Clidicus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="144" phylum="Arthropoda" rank="species" species="crocodylus">
|
||
<emphasis id="07D6EA80FFB86904BE71F87C7FA9F853" box="[536,683,1998,2022]" italics="true" pageId="3" pageNumber="144">C. crocodylus</emphasis>
|
||
</taxonomicName>
|
||
has the antennae shorter in relation to the body length than those in
|
||
<emphasis id="07D6EA80FFB86904B9EDF87C789EF853" box="[1412,1436,1999,2022]" italics="true" pageId="3" pageNumber="144">C.</emphasis>
|
||
</paragraph>
|
||
<paragraph id="351D3692FFBF6903BCFEFF2B793DFE71" blockId="4.[151,1436,152,452]" pageId="4" pageNumber="145">
|
||
<taxonomicName id="F2A24D11FFBF6903BCFEFF2B7C28FF1A" authorityName="Jałoszyński" authorityYear="2018" box="[151,298,152,175]" class="Insecta" family="Staphylinidae" genus="Clidicus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="145" phylum="Arthropoda" rank="species" species="asymmetricus">
|
||
<emphasis id="07D6EA80FFBF6903BCFEFF2B7C28FF1A" box="[151,298,152,175]" italics="true" pageId="4" pageNumber="145">asymmetricus</emphasis>
|
||
</taxonomicName>
|
||
, the antennomeres being clearly more elongate in the latter species (
|
||
<figureCitation id="AD992A17FFBF6903B82FFF2B7989FF1A" box="[1094,1163,152,175]" captionStart="FIGURES1–2" captionStartId="1.[151,258,1773,1795]" captionTargetBox="[171,1418,414,1735]" captionTargetId="figure@1.[151,1436,399,1752]" captionTargetPageId="1" captionText="FIGURES1–2.Clidicus of the Philippines,dorsal habitus of males.Clidicus asymmetricus sp.n. (1); C.crocodylus Jałoszyński (2)." httpUri="https://zenodo.org/record/1206428/files/figure.png" pageId="4" pageNumber="145">Fig. 1</figureCitation>
|
||
vs. 2). The head in
|
||
<taxonomicName id="F2A24D11FFBF6903B9EDFF2B7C28FF64" authorityName="Jałoszyński" authorityYear="2018" class="Insecta" family="Staphylinidae" genus="Clidicus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="145" phylum="Arthropoda" rank="species" species="asymmetricus">
|
||
<emphasis id="07D6EA80FFBF6903B9EDFF2B7C28FF64" italics="true" pageId="4" pageNumber="145">C. asymmetricus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="AD992A17FFBF6903BD51FF097C9BFF64" box="[312,409,186,209]" captionStart="FIGURES3–6" captionStartId="2.[151,258,1973,1995]" captionTargetBox="[151,1436,426,1960]" captionTargetId="figure@2.[151,1436,426,1960]" captionTargetPageId="2" captionText="FIGURES3–6. Clidicus ofthe Philippines,pronotum and head of males (pronotumin dorsalview,head slightly tilted ventrad) (3, 5), andhead indorsal view(4, 6).Clidicus asymmetricus sp.n.(3–4); C. crocodylus Jałoszyński (5–6)." httpUri="https://zenodo.org/record/1206430/files/figure.png" pageId="4" pageNumber="145">Figs 3–4</figureCitation>
|
||
) is broadest shortly behind the eyes, slightly narrower than pronotum and with evenly rounded tempora, whereas the head in
|
||
<taxonomicName id="F2A24D11FFBF6903BDB1FF6E7F6EFF41" box="[472,620,220,244]" class="Insecta" family="Staphylinidae" genus="Clidicus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="145" phylum="Arthropoda" rank="species" species="crocodylus">
|
||
<emphasis id="07D6EA80FFBF6903BDB1FF6E7F6EFF41" box="[472,620,220,244]" italics="true" pageId="4" pageNumber="145">C. crocodylus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="AD992A17FFBF6903BE13FF6E7FDBFF41" box="[634,729,221,244]" captionStart="FIGURES3–6" captionStartId="2.[151,258,1973,1995]" captionTargetBox="[151,1436,426,1960]" captionTargetId="figure@2.[151,1436,426,1960]" captionTargetPageId="2" captionText="FIGURES3–6. Clidicus ofthe Philippines,pronotum and head of males (pronotumin dorsalview,head slightly tilted ventrad) (3, 5), andhead indorsal view(4, 6).Clidicus asymmetricus sp.n.(3–4); C. crocodylus Jałoszyński (5–6)." httpUri="https://zenodo.org/record/1206430/files/figure.png" pageId="4" pageNumber="145">Figs 5–6</figureCitation>
|
||
) is broadest more posteriorly, slightly broader than pronotum and more triangular, with tempora behind eyes nearly straight and then rapidly curved mesad. Also the shape of pronotum slightly differs in these species; that in
|
||
<taxonomicName id="F2A24D11FFBF6903BE25FE917E00FE8C" authorityName="Jałoszyński" authorityYear="2018" box="[588,770,290,313]" class="Insecta" family="Staphylinidae" genus="Clidicus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="145" phylum="Arthropoda" rank="species" species="asymmetricus">
|
||
<emphasis id="07D6EA80FFBF6903BE25FE917E00FE8C" box="[588,770,290,313]" italics="true" pageId="4" pageNumber="145">C. asymmetricus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="AD992A17FFBF6903BF7AFE917E57FE8C" box="[787,853,290,313]" captionStart="FIGURES3–6" captionStartId="2.[151,258,1973,1995]" captionTargetBox="[151,1436,426,1960]" captionTargetId="figure@2.[151,1436,426,1960]" captionTargetPageId="2" captionText="FIGURES3–6. Clidicus ofthe Philippines,pronotum and head of males (pronotumin dorsalview,head slightly tilted ventrad) (3, 5), andhead indorsal view(4, 6).Clidicus asymmetricus sp.n.(3–4); C. crocodylus Jałoszyński (5–6)." httpUri="https://zenodo.org/record/1206430/files/figure.png" pageId="4" pageNumber="145">Fig. 3</figureCitation>
|
||
) has a longer posterior 'collar' with its sides slightly divergent caudad, whereas in
|
||
<taxonomicName id="F2A24D11FFBF6903BDBDFEF67F65FEE9" box="[468,615,324,348]" class="Insecta" family="Staphylinidae" genus="Clidicus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="145" phylum="Arthropoda" rank="species" species="crocodylus">
|
||
<emphasis id="07D6EA80FFBF6903BDBDFEF67F65FEE9" box="[468,615,324,348]" italics="true" pageId="4" pageNumber="145">C. crocodylus</emphasis>
|
||
</taxonomicName>
|
||
the short collar has its lateral margins parallel or slightly convergent caudad. The latter species has distinctly narrower 'funnels' of the 'sperm pump', a broadening of the ductus ejaculatorius that presumably can contract by means of longitudinal muscles fibers connecting sclerotized funnel-like ends of the 'pump'; its function remains unknown, the term 'sperm pump' was in use since
|
||
<bibRefCitation id="51334B63FFBF6903BFE1FE1E7938FE71" box="[904,1082,429,452]" pageId="4" pageNumber="145" refId="ref1586" refString="Besuchet, C. (1971) Les Clidicus de Ceylon (Col. Scydmaenidae). Mitteilungen der Schweizerischen Entomologischen Gesellschaft, 43, 249 - 257." type="journal article">Besuchet (1971)</bibRefCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |