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<document id="83D06714573DD2469897F67C6CC23380" ID-DOI="10.5281/zenodo.212417" ID-GBIF-Dataset="4a20fc78-d49d-440e-9829-00290cc0e482" ID-ISSN="1175-5326" ID-Zenodo-Dep="212417" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460479156746" checkinUser="plazi" docAuthor="Bayartogtokh, Badamdorj" docDate="2012" docId="3E6087FFFF93194DFF53FAC523A0F9A0" docLanguage="en" docName="zt03302p060.pdf" docOrigin="Zootaxa 3302" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Cultroribula rarisetosa Bayartogtokh, 2012, sp. nov." docType="treatment" docVersion="8" lastPageNumber="51" masterDocId="C259FF87FF96194AFFC4FFFB2177FFEB" masterDocTitle="The genus Cultroribula (Acari: Oribatida: Astegistidae) in Mongolia, with new findings from Altai Mountains and remarks on known species of the world" masterLastPageNumber="60" masterPageNumber="44" pageNumber="49" updateTime="1698307244462" updateUser="plazi">
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<mods:title id="944F02706B3F6A97686E030883BB2C5D">The genus Cultroribula (Acari: Oribatida: Astegistidae) in Mongolia, with new findings from Altai Mountains and remarks on known species of the world</mods:title>
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<mods:namePart id="EC725697D873291E9BD5F4C34C8DBC51">Bayartogtokh, Badamdorj</mods:namePart>
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<mods:date id="56DA5FE247551B020582D3D0A011CBEA">2012</mods:date>
<mods:detail id="362FABEBF9DBD203BE267A98B00D8818" type="volume">
<mods:number id="E8FEBA1A45F1D9C66568DD23D89AC1E3">3302</mods:number>
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<mods:classification id="2E829DEF93B43CCB43479855244B3FDC">journal article</mods:classification>
<mods:identifier id="DE4C9E6F12B43AADDF053D94A4FB14DF" type="DOI">10.5281/zenodo.212417</mods:identifier>
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<mods:identifier id="B53417375E69CEC4353D3FA6C1F187A4" type="Zenodo-Dep">212417</mods:identifier>
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<treatment id="3E6087FFFF93194DFF53FAC523A0F9A0" ID-DOI="http://doi.org/10.5281/zenodo.6176743" ID-GBIF-Taxon="119516810" ID-Zenodo-Dep="6176743" LSID="urn:lsid:plazi:treatment:3E6087FFFF93194DFF53FAC523A0F9A0" httpUri="http://treatment.plazi.org/id/3E6087FFFF93194DFF53FAC523A0F9A0" lastPageId="7" lastPageNumber="51" pageId="5" pageNumber="49">
<subSubSection id="FED36562FF93194FFF53FAC5219CFA92" pageId="5" pageNumber="49" type="nomenclature">
<paragraph id="B67636E9FF93194FFF53FAC52366FAB3" blockId="5.[151,529,1342,1401]" box="[151,529,1342,1368]" pageId="5" pageNumber="49">
<heading id="ED3E8185FF93194FFF53FAC52366FAB3" bold="true" box="[151,529,1342,1368]" fontSize="11" level="1" pageId="5" pageNumber="49" reason="1">
<emphasis id="84BDEAFBFF93194FFF53FAC52366FAB3" bold="true" box="[151,529,1342,1368]" pageId="5" pageNumber="49">
<taxonomicName id="71C94D6AFF93194FFF53FAC520DBFAB3" box="[151,428,1342,1368]" class="Arachnida" family="Astegistidae" genus="Cultroribula" kingdom="Animalia" order="Sarcoptiformes" pageId="5" pageNumber="49" phylum="Arthropoda" rank="species" species="rarisetosa" status="sp. nov.">
<emphasis id="84BDEAFBFF93194FFF53FAC520DBFAB3" bold="true" box="[151,428,1342,1368]" italics="true" pageId="5" pageNumber="49">Cultroribula rarisetosa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9F8E5780FF93194FFE70FAC52366FAB3" box="[436,529,1342,1368]" pageId="5" pageNumber="49" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="B67636E9FF93194FFF53FA9A219CFA92" blockId="5.[151,529,1342,1401]" box="[151,235,1377,1401]" pageId="5" pageNumber="49">
(
<figureCitation id="2EF22A6CFF93194FFF5BFA9A2194FA92" box="[159,227,1377,1401]" captionStart="FIGURE 4" captionStartId="6.[151,249,1948,1971]" captionTargetBox="[152,1431,181,1947]" captionTargetId="figure@6.[151,1436,169,1965]" captionTargetPageId="6" captionText="FIGURE 4. Cultroribula rarisetosa sp. nov. (A) Dorsal view of idiosoma; (B) Variation of rostrum, dorso-frontal view; (C) Rostrum, ventral view; (D) Cusp of lamella; (E) Lateral view of prodorsum and anterior part of notogaster; (F) Ventral view of idiosoma. B – E to same scale." httpUri="https://zenodo.org/record/212421/files/figure.png" pageId="5" pageNumber="49">Fig. 4</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="FED36562FF93194FFF53FA512578F99E" pageId="5" pageNumber="49" type="diagnosis">
<paragraph id="B67636E9FF93194FFF53FA512578F99E" blockId="5.[151,1436,1449,2014]" pageId="5" pageNumber="49">
<emphasis id="84BDEAFBFF93194FFF53FA512067FA2A" box="[151,272,1450,1473]" italics="true" pageId="5" pageNumber="49">Diagnosis.</emphasis>
Medium-sized species, covered with thin cerotegument; rostrum with 13–15 teeth, among them four central teeth much larger than lateral teeth; lamellae wide, fused medially; lamellar cusp long and wide, with large outer tooth, but without inner tooth; rostral and lamellar setae moderately long, with conspicuous barbs, interlamellar seta short, smooth; sensillus with short stalk and club-shaped head with minute barbs; tutorium wide at base, with long cusp sharply pointed distally; humeral process of notogaster not sharply projected, but rounded distally; ten pairs of notogastral setae short; four pairs of genital setae, legs tridactylous.
</paragraph>
</subSubSection>
<subSubSection id="FED36562FF93194DFF0DF97924E2FC56" lastPageId="7" lastPageNumber="51" pageId="5" pageNumber="49" type="description">
<paragraph id="B67636E9FF93194FFF0DF97920FEF956" blockId="5.[151,1436,1449,2014]" pageId="5" pageNumber="49">
<emphasis id="84BDEAFBFF93194FFF0DF9792004F972" box="[201,371,1666,1689]" italics="true" pageId="5" pageNumber="49">Measurements.</emphasis>
Body length: 338–346 (342) μm; width of notogaster 187–194 (191) μm. In total two specimens were measured.
</paragraph>
<paragraph id="B67636E9FF93194FFF0DF93124F8F8EE" blockId="5.[151,1436,1449,2014]" pageId="5" pageNumber="49">
<emphasis id="84BDEAFBFF93194FFF0DF9312030F90A" box="[201,327,1738,1761]" italics="true" pageId="5" pageNumber="49">Integument</emphasis>
. Body yellowish to yellowish-brown in color. Surface of body and leg segments with very thin, nearly smooth cerotegument. Integument microtuberculate on lateral part of prodorsum and around leg acetabula.
</paragraph>
<paragraph id="B67636E9FF93194DFF0DF8EA21CCFF1C" blockId="5.[151,1436,1449,2014]" lastBlockId="7.[151,1436,151,1611]" lastPageId="7" lastPageNumber="51" pageId="5" pageNumber="49">
<emphasis id="84BDEAFBFF93194FFF0DF8EA2031F8C2" box="[201,326,1809,1833]" italics="true" pageId="5" pageNumber="49">Prodorsum</emphasis>
. Rostrum with 13–15 teeth, among them four central teeth much larger than lateral teeth; two most central teeth slightly shorter than next lateral teeth; size of rostral teeth becoming smaller toward posterior direction (
<figureCitation id="2EF22A6CFF93194FFF5BF8A22195F89A" box="[159,226,1881,1905]" captionStart="FIGURE 4" captionStartId="6.[151,249,1948,1971]" captionTargetBox="[152,1431,181,1947]" captionTargetId="figure@6.[151,1436,169,1965]" captionTargetPageId="6" captionText="FIGURE 4. Cultroribula rarisetosa sp. nov. (A) Dorsal view of idiosoma; (B) Variation of rostrum, dorso-frontal view; (C) Rostrum, ventral view; (D) Cusp of lamella; (E) Lateral view of prodorsum and anterior part of notogaster; (F) Ventral view of idiosoma. B – E to same scale." httpUri="https://zenodo.org/record/212421/files/figure.png" pageId="5" pageNumber="49">Fig. 4</figureCitation>
B &amp; C). Rostral seta moderately long, with long barbs. Lamella wide, fused medially; lamellar cusp slightly longer, but narrower than basal part of lamella, with large outer tooth, but without inner tooth (
<figureCitation id="2EF22A6CFF93194FFB03F886247AF87E" box="[1223,1293,1917,1941]" captionStart="FIGURE 4" captionStartId="6.[151,249,1948,1971]" captionTargetBox="[152,1431,181,1947]" captionTargetId="figure@6.[151,1436,169,1965]" captionTargetPageId="6" captionText="FIGURE 4. Cultroribula rarisetosa sp. nov. (A) Dorsal view of idiosoma; (B) Variation of rostrum, dorso-frontal view; (C) Rostrum, ventral view; (D) Cusp of lamella; (E) Lateral view of prodorsum and anterior part of notogaster; (F) Ventral view of idiosoma. B – E to same scale." httpUri="https://zenodo.org/record/212421/files/figure.png" pageId="5" pageNumber="49">Fig. 4</figureCitation>
D). Lamellar seta long, about 1.6 times as long as
<emphasis id="84BDEAFBFF93194FFDF8F8592324F852" box="[572,595,1954,1977]" italics="true" pageId="5" pageNumber="49">ro</emphasis>
, sparsely barbed; interlamellar seta short, thin, smooth. Bothridium large, with wide opening directed anterolaterad, partly concealed under anterior margin of notogaster. Exobothridial seta short, as long as
<emphasis id="84BDEAFBFF91194DFE92FF62201BFF5B" box="[342,364,153,176]" italics="true" pageId="7" pageNumber="51">in</emphasis>
, smooth. Sensillus with short stalk and long fusiform head, distal end of which setiform, long. Tutorium wide at base, with long cusp sharply pointed distally, almost reaching alveolus of rostral seta (
<figureCitation id="2EF22A6CFF91194DFAECFF47241DFF3F" box="[1320,1386,188,212]" captionStart="FIGURE 4" captionStartId="6.[151,249,1948,1971]" captionTargetBox="[152,1431,181,1947]" captionTargetId="figure@6.[151,1436,169,1965]" captionTargetPageId="6" captionText="FIGURE 4. Cultroribula rarisetosa sp. nov. (A) Dorsal view of idiosoma; (B) Variation of rostrum, dorso-frontal view; (C) Rostrum, ventral view; (D) Cusp of lamella; (E) Lateral view of prodorsum and anterior part of notogaster; (F) Ventral view of idiosoma. B – E to same scale." httpUri="https://zenodo.org/record/212421/files/figure.png" pageId="7" pageNumber="51">Fig. 4</figureCitation>
A &amp; G).
</paragraph>
<caption id="E2B66661FF90194CFF53F86720B5F81B" httpUri="https://zenodo.org/record/212421/files/figure.png" pageId="6" pageNumber="50" targetBox="[152,1431,181,1947]" targetPageId="6">
<paragraph id="B67636E9FF90194CFF53F86720B5F81B" blockId="6.[151,1436,1948,2032]" pageId="6" pageNumber="50">
<emphasis id="84BDEAFBFF90194CFF53F8672063F85A" bold="true" box="[151,276,1948,1971]" pageId="6" pageNumber="50">FIGURE 4.</emphasis>
<taxonomicName id="71C94D6AFF90194CFED8F867237DF859" box="[284,522,1948,1970]" class="Arachnida" family="Astegistidae" genus="Cultroribula" kingdom="Animalia" order="Sarcoptiformes" pageId="6" pageNumber="50" phylum="Arthropoda" rank="species" species="rarisetosa" status="sp. nov.">
<emphasis id="84BDEAFBFF90194CFED8F867237DF859" box="[284,522,1948,1970]" italics="true" pageId="6" pageNumber="50">Cultroribula rarisetosa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9F8E5780FF90194CFDD6F8662313F859" box="[530,612,1949,1970]" pageId="6" pageNumber="50" rank="species">
<emphasis id="84BDEAFBFF90194CFDD6F8662313F859" bold="true" box="[530,612,1949,1970]" pageId="6" pageNumber="50">sp. nov.</emphasis>
</taxonomicNameLabel>
(A) Dorsal view of idiosoma; (B) Variation of rostrum, dorso-frontal view; (C) Rostrum, ventral view; (D) Cusp of lamella; (E) Lateral view of prodorsum and anterior part of notogaster; (F) Ventral view of idiosoma. B–E to same scale.
</paragraph>
</caption>
<paragraph id="B67636E9FF91194DFF03FEFE2081FE94" blockId="7.[151,1436,151,1611]" pageId="7" pageNumber="51">
<emphasis id="84BDEAFBFF91194DFF03FEFE2035FEF7" box="[199,322,261,284]" italics="true" pageId="7" pageNumber="51">Notogaster</emphasis>
. Oval, longer than wide, anterior margin almost straight, slightly rounded; humeral process not sharply projected, but rounded distally in both dorsal and lateral views (
<figureCitation id="2EF22A6CFF91194DFC70FEDE2282FED6" box="[948,1013,293,317]" captionStart="FIGURE 4" captionStartId="6.[151,249,1948,1971]" captionTargetBox="[152,1431,181,1947]" captionTargetId="figure@6.[151,1436,169,1965]" captionTargetPageId="6" captionText="FIGURE 4. Cultroribula rarisetosa sp. nov. (A) Dorsal view of idiosoma; (B) Variation of rostrum, dorso-frontal view; (C) Rostrum, ventral view; (D) Cusp of lamella; (E) Lateral view of prodorsum and anterior part of notogaster; (F) Ventral view of idiosoma. B – E to same scale." httpUri="https://zenodo.org/record/212421/files/figure.png" pageId="7" pageNumber="51">Fig. 4</figureCitation>
A &amp; G). Ten pairs of notogastral setae short, thin, smooth, all setae equal in length. Lyrifissures
<emphasis id="84BDEAFBFF91194DFCDFFEBC2240FEB5" box="[795,823,327,350]" italics="true" pageId="7" pageNumber="51">im</emphasis>
,
<emphasis id="84BDEAFBFF91194DFC81FEBD222CFEB5" box="[837,859,326,350]" italics="true" pageId="7" pageNumber="51">ih</emphasis>
,
<emphasis id="84BDEAFBFF91194DFCAEFEBC22FCFEB5" box="[874,907,327,350]" italics="true" pageId="7" pageNumber="51">ips</emphasis>
,
<emphasis id="84BDEAFBFF91194DFC5DFEBC22D8FEB5" box="[921,943,327,350]" italics="true" pageId="7" pageNumber="51">ip</emphasis>
and opisthonotal gland opening well developed; lyrifissure
<emphasis id="84BDEAFBFF91194DFE90FE92201DFE6B" box="[340,362,361,384]" italics="true" pageId="7" pageNumber="51">ia</emphasis>
not evident.
</paragraph>
<paragraph id="B67636E9FF91194DFF0DFE722060FDDA" blockId="7.[151,1436,151,1611]" pageId="7" pageNumber="51">
<emphasis id="84BDEAFBFF91194DFF0DFE722028FE4A" box="[201,351,393,417]" italics="true" pageId="7" pageNumber="51">
<taxonomicName id="71C94D6AFF91194DFF0DFE72202BFE4A" box="[201,348,393,417]" class="Secernentea" family="Gnathostomatidae" genus="Gnathosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirurida" pageId="7" pageNumber="51" phylum="Nematoda" rank="genus">Gnathosoma</taxonomicName>
.
</emphasis>
Subcapitular mentum nearly as long as wide, with minute microtubercles. Hypostomal setae
<emphasis id="84BDEAFBFF91194DFAA8FE71240DFE4A" box="[1388,1402,394,417]" italics="true" pageId="7" pageNumber="51">a</emphasis>
,
<emphasis id="84BDEAFBFF91194DFA4CFE7124EBFE4A" box="[1416,1436,394,417]" italics="true" pageId="7" pageNumber="51">m</emphasis>
and
<emphasis id="84BDEAFBFF91194DFF02FE5621A3FE2E" box="[198,212,429,453]" italics="true" pageId="7" pageNumber="51">h</emphasis>
medium long, thin, smooth (
<figureCitation id="2EF22A6CFF91194DFDDFFE562328FE2E" box="[539,607,429,453]" captionStart="FIGURE 4" captionStartId="6.[151,249,1948,1971]" captionTargetBox="[152,1431,181,1947]" captionTargetId="figure@6.[151,1436,169,1965]" captionTargetPageId="6" captionText="FIGURE 4. Cultroribula rarisetosa sp. nov. (A) Dorsal view of idiosoma; (B) Variation of rostrum, dorso-frontal view; (C) Rostrum, ventral view; (D) Cusp of lamella; (E) Lateral view of prodorsum and anterior part of notogaster; (F) Ventral view of idiosoma. B – E to same scale." httpUri="https://zenodo.org/record/212421/files/figure.png" pageId="7" pageNumber="51">Fig. 4</figureCitation>
F). Chelicera typical for genus as shown in the previous species, with few slightly sclerotized blunt teeth; seta
<emphasis id="84BDEAFBFF91194DFDFFFE2A2313FE02" box="[571,612,465,489]" italics="true" pageId="7" pageNumber="51">cha</emphasis>
barbed, about 1.5 times as long as smooth
<emphasis id="84BDEAFBFF91194DFBA1FE2A25FAFE02" box="[1125,1165,465,489]" italics="true" pageId="7" pageNumber="51">chb</emphasis>
; Trägårdh’s organ well developed, large. Palp typical for genus as in the previous species, palpal setation: 0-2-1-3-8, including solenidion ω of tarsus.
</paragraph>
<paragraph id="B67636E9FF91194DFF0DFDC625C5FD76" blockId="7.[151,1436,151,1611]" pageId="7" pageNumber="51">
<emphasis id="84BDEAFBFF91194DFF0DFDC620F6FDBE" box="[201,385,573,597]" italics="true" pageId="7" pageNumber="51">Epimeral region</emphasis>
. Apodemes
<emphasis id="84BDEAFBFF91194DFDCBFDC62339FDBE" box="[527,590,573,597]" italics="true" pageId="7" pageNumber="51">apo.2</emphasis>
,
<emphasis id="84BDEAFBFF91194DFD99FDC523F0FDBE" box="[605,647,574,597]" italics="true" pageId="7" pageNumber="51">apo</emphasis>
.
<emphasis id="84BDEAFBFF91194DFD4AFDC523D6FDBE" box="[654,673,574,597]" italics="true" pageId="7" pageNumber="51">sj</emphasis>
and
<emphasis id="84BDEAFBFF91194DFD1DFDC62260FDBE" box="[729,791,573,597]" italics="true" pageId="7" pageNumber="51">apo.3</emphasis>
well developed, slightly obliquely or nearly transversely oriented. Epimeral setae short, smooth; setal formula: 3-1-3-3. Discidium well developed, projecting distally; circumpedal carina well developed, its anterior tectum reaching level of pedotectum II (
<figureCitation id="2EF22A6CFF91194DFB8BFD7E25E4FD76" box="[1103,1171,645,669]" captionStart="FIGURE 4" captionStartId="6.[151,249,1948,1971]" captionTargetBox="[152,1431,181,1947]" captionTargetId="figure@6.[151,1436,169,1965]" captionTargetPageId="6" captionText="FIGURE 4. Cultroribula rarisetosa sp. nov. (A) Dorsal view of idiosoma; (B) Variation of rostrum, dorso-frontal view; (C) Rostrum, ventral view; (D) Cusp of lamella; (E) Lateral view of prodorsum and anterior part of notogaster; (F) Ventral view of idiosoma. B – E to same scale." httpUri="https://zenodo.org/record/212421/files/figure.png" pageId="7" pageNumber="51">Fig. 4</figureCitation>
F).
</paragraph>
<paragraph id="B67636E9FF91194DFF0DFD522444FCC6" blockId="7.[151,1436,151,1611]" pageId="7" pageNumber="51">
<emphasis id="84BDEAFBFF91194DFF0DFD5220EAFD2A" box="[201,413,681,705]" italics="true" pageId="7" pageNumber="51">Ano-genital region</emphasis>
. Anal and genital apertures medium in size, situated far from each other. Genital aperture nearly rounded, genital plates with four pairs of short, smooth setae (
<figureCitation id="2EF22A6CFF91194DFC72FD36228BFD0E" box="[950,1020,717,741]" captionStart="FIGURE 4" captionStartId="6.[151,249,1948,1971]" captionTargetBox="[152,1431,181,1947]" captionTargetId="figure@6.[151,1436,169,1965]" captionTargetPageId="6" captionText="FIGURE 4. Cultroribula rarisetosa sp. nov. (A) Dorsal view of idiosoma; (B) Variation of rostrum, dorso-frontal view; (C) Rostrum, ventral view; (D) Cusp of lamella; (E) Lateral view of prodorsum and anterior part of notogaster; (F) Ventral view of idiosoma. B – E to same scale." httpUri="https://zenodo.org/record/212421/files/figure.png" pageId="7" pageNumber="51">Fig. 4</figureCitation>
E). Aggenital seta as long as genital setae, thin, smooth. Anal aperture widened posteriorly, slightly longer than wide; anal and adanal setae short, equal in size. Adanal lyrifissure well developed, situated adjacent to anterolateral corner of anal plate (
<figureCitation id="2EF22A6CFF91194DFB14FCEE2463FCC6" box="[1232,1300,789,813]" captionStart="FIGURE 4" captionStartId="6.[151,249,1948,1971]" captionTargetBox="[152,1431,181,1947]" captionTargetId="figure@6.[151,1436,169,1965]" captionTargetPageId="6" captionText="FIGURE 4. Cultroribula rarisetosa sp. nov. (A) Dorsal view of idiosoma; (B) Variation of rostrum, dorso-frontal view; (C) Rostrum, ventral view; (D) Cusp of lamella; (E) Lateral view of prodorsum and anterior part of notogaster; (F) Ventral view of idiosoma. B – E to same scale." httpUri="https://zenodo.org/record/212421/files/figure.png" pageId="7" pageNumber="51">Fig. 4</figureCitation>
F).
</paragraph>
<paragraph id="B67636E9FF91194DFF0DFCC124E2FC56" blockId="7.[151,1436,151,1611]" pageId="7" pageNumber="51">
<emphasis id="84BDEAFBFF91194DFF0DFCC12189FCBA" box="[201,254,826,849]" italics="true" pageId="7" pageNumber="51">Legs</emphasis>
. Tarsi heterotridactylous, median claw conspicuously thicker than lateral claws. Trochanter and femora of all legs with large porose areas. Setation of legs as in the former species. Most of tarsal and tibial setae barbed, but those of genua, femora and trochanters mostly smooth. Formula of leg setation (including famulus): I (
<date id="C2771029FF91194DFAD6FC7A24F9FC72" box="[1298,1422,897,921]" pageId="7" pageNumber="51" value="1918-03-01" valueMax="1918-04-05" valueMin="1918-03-01">1-5-3-4-18</date>
), II (
<date id="C2771029FF91194DFF7DFC5E2042FC56" box="[185,309,933,957]" pageId="7" pageNumber="51" value="1915-02-01" valueMax="1915-04-05" valueMin="1915-02-01">1-5-2-4-15</date>
), III (
<date id="C2771029FF91194DFEB2FC5E2087FC56" box="[374,496,933,957]" pageId="7" pageNumber="51" value="1915-01-01" valueMax="1915-03-03" valueMin="1915-01-01">1-3-1-3-15</date>
); IV (
<date id="C2771029FF91194DFDF0FC5E23C7FC56" box="[564,688,933,957]" pageId="7" pageNumber="51" value="1912-02-01" valueMax="1912-03-03" valueMin="1912-02-01">1-3-2-3-12</date>
); formula of solenidia: I (1-2-2); II (1-1-2); III (1-1-0); IV (0-1-0).
</paragraph>
</subSubSection>
<subSubSection id="FED36562FF91194DFF0DFC322029FB73" pageId="7" pageNumber="51" type="materials_examined">
<paragraph id="B67636E9FF91194DFF0DFC322029FB73" blockId="7.[151,1436,151,1611]" pageId="7" pageNumber="51">
<emphasis id="84BDEAFBFF91194DFF0DFC3220D7FC0A" box="[201,416,969,993]" italics="true" pageId="7" pageNumber="51">Material examined</emphasis>
.
<typeStatus id="6972884BFF91194DFE76FC32236DFC0A" box="[434,538,969,993]" pageId="7" pageNumber="51" type="holotype">Holotype</typeStatus>
(female): Mts. Mongol Altai, close to Lake Dayan, District Sagsai, Province Bayan-Ulgii, foliose or subfruticose lichens growing on bare rocks (
<taxonomicName id="71C94D6AFF91194DFC66FC1525F0FBEE" box="[930,1159,1005,1029]" class="Arachnida" family="Astegistidae" genus="Xantoria" kingdom="Animalia" order="Sarcoptiformes" pageId="7" pageNumber="51" phylum="Arthropoda" rank="species" species="candelaria">
<emphasis id="84BDEAFBFF91194DFC66FC1525F0FBEE" box="[930,1159,1005,1029]" italics="true" pageId="7" pageNumber="51">Xantoria candelaria</emphasis>
</taxonomicName>
,
<taxonomicName id="71C94D6AFF91194DFB53FC1621CCFBC7" class="Lecanoromycetes" family="Parmeliaceae" genus="Melanelixia" kingdom="Fungi" order="Lecanorales" pageId="7" pageNumber="51" phylum="Ascomycota" rank="species" species="exasperatula">
<emphasis id="84BDEAFBFF91194DFB53FC1621CCFBC7" italics="true" pageId="7" pageNumber="51">Melanelixia exasperatula</emphasis>
</taxonomicName>
,
<taxonomicName id="71C94D6AFF91194DFF0EFBEF20FCFBC7" box="[202,395,1044,1068]" class="Lecanoromycetes" family="Parmeliaceae" genus="Parmelia" kingdom="Fungi" order="Lecanorales" pageId="7" pageNumber="51" phylum="Ascomycota" rank="species" species="sulcata">
<emphasis id="84BDEAFBFF91194DFF0EFBEF20FCFBC7" box="[202,395,1044,1068]" italics="true" pageId="7" pageNumber="51">Parmelia sulcata</emphasis>
</taxonomicName>
),
<geoCoordinate id="D3FD502EFF91194DFE67FBEF2375FBC7" box="[419,514,1041,1068]" direction="north" orientation="latitude" pageId="7" pageNumber="51" precision="925" value="48.25">N48o15’</geoCoordinate>
,
<geoCoordinate id="D3FD502EFF91194DFDCBFBEF231CFBC7" box="[527,619,1041,1068]" direction="east" orientation="longitude" pageId="7" pageNumber="51" precision="925" value="88.9">E88o54’</geoCoordinate>
, elevation
<quantity id="71319B0CFF91194DFD23FBEF224AFBC7" box="[743,829,1044,1068]" metricMagnitude="3" metricUnit="m" metricValue="2.3369999999999997" pageId="7" pageNumber="51" unit="m" value="2337.0">2337 m</quantity>
a.s.l.,
<date id="C2771029FF91194DFC43FBEF256AFBC7" box="[903,1053,1044,1068]" pageId="7" pageNumber="51" value="2010-07-16">16 July 2010</date>
, Coll. B. Bayartogtokh; one
<typeStatus id="6972884BFF91194DFAA6FBEE21B0FBBA" pageId="7" pageNumber="51" type="paratype">paratype</typeStatus>
(female): same data as
<typeStatus id="6972884BFF91194DFE3BFBC32312FBBB" box="[511,613,1080,1104]" pageId="7" pageNumber="51" type="holotype">holotype</typeStatus>
. The
<typeStatus id="6972884BFF91194DFD6BFBC32266FBBB" box="[687,785,1080,1104]" pageId="7" pageNumber="51" type="holotype">holotype</typeStatus>
and a
<typeStatus id="6972884BFF91194DFCABFBC222A7FBBA" box="[879,976,1081,1105]" pageId="7" pageNumber="51" type="paratype">paratype</typeStatus>
are deposited in the collection of the Department of Zoology, National University of
<collectingCountry id="CEDE7679FF91194DFD01FBA72242FB9F" box="[709,821,1116,1140]" name="Mongolia" pageId="7" pageNumber="51">Mongolia</collectingCountry>
, Ulaanbaatar,
<collectingCountry id="CEDE7679FF91194DFC1FFBA7253CFB9F" box="[987,1099,1116,1140]" name="Mongolia" pageId="7" pageNumber="51">Mongolia</collectingCountry>
. All
<typeStatus id="6972884BFF91194DFB4CFBA625CFFB9E" box="[1160,1208,1117,1141]" pageId="7" pageNumber="51">type</typeStatus>
specimens are preserved in alcohol.
</paragraph>
</subSubSection>
<subSubSection id="FED36562FF91194DFF0DFB5F23A0F9A0" pageId="7" pageNumber="51" type="discussion">
<paragraph id="B67636E9FF91194DFF0DFB5F239AFA9B" blockId="7.[151,1436,151,1611]" pageId="7" pageNumber="51">
<emphasis id="84BDEAFBFF91194DFF0DFB5F205CFB57" box="[201,299,1188,1212]" italics="true" pageId="7" pageNumber="51">Remarks</emphasis>
.
<taxonomicName id="71C94D6AFF91194DFEFEFB5F2348FB57" box="[314,575,1188,1212]" class="Arachnida" family="Astegistidae" genus="Cultroribula" kingdom="Animalia" order="Sarcoptiformes" pageId="7" pageNumber="51" phylum="Arthropoda" rank="species" species="rarisetosa" status="sp. nov.">
<emphasis id="84BDEAFBFF91194DFEFEFB5F2348FB57" box="[314,575,1188,1212]" italics="true" pageId="7" pageNumber="51">Cultroribula rarisetosa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9F8E5780FF91194DFD83FB5F23D5FB57" box="[583,674,1188,1212]" pageId="7" pageNumber="51" rank="species">
<emphasis id="84BDEAFBFF91194DFD83FB5F23D5FB57" bold="true" box="[583,674,1188,1212]" pageId="7" pageNumber="51">sp. nov.</emphasis>
</taxonomicNameLabel>
is clearly distinguishable from other species of
<taxonomicName id="71C94D6AFF91194DFB05FB5F2439FB57" box="[1217,1358,1188,1212]" class="Arachnida" family="Astegistidae" genus="Cultroribula" kingdom="Animalia" order="Sarcoptiformes" pageId="7" pageNumber="51" phylum="Arthropoda" rank="genus">
<emphasis id="84BDEAFBFF91194DFB05FB5F2439FB57" box="[1217,1358,1188,1212]" italics="true" pageId="7" pageNumber="51">Cultroribula</emphasis>
</taxonomicName>
by the combination of following characters: absence of inner tooth of lamellar cusp, presence of only four pairs of genital setae and dentation of rostrum. Among the known species of
<taxonomicName id="71C94D6AFF91194DFC9EFB172291FAEF" box="[858,998,1260,1284]" class="Arachnida" family="Astegistidae" genus="Cultroribula" kingdom="Animalia" order="Sarcoptiformes" pageId="7" pageNumber="51" phylum="Arthropoda" rank="genus">
<emphasis id="84BDEAFBFF91194DFC9EFB172291FAEF" box="[858,998,1260,1284]" italics="true" pageId="7" pageNumber="51">Cultroribula</emphasis>
</taxonomicName>
, only four Palaearctic species,
<taxonomicName id="71C94D6AFF91194DFA89FB162060FAC3" authority="Aoki, 1961" authorityName="Aoki" authorityYear="1961" class="Arachnida" family="Astegistidae" genus="Cultroribula" kingdom="Animalia" order="Sarcoptiformes" pageId="7" pageNumber="51" phylum="Arthropoda" rank="species" species="lata">
<emphasis id="84BDEAFBFF91194DFA89FB1624ECFAEF" box="[1357,1435,1260,1284]" italics="true" pageId="7" pageNumber="51">C. lata</emphasis>
<bibRefCitation id="D2584B18FF91194DFF53FAEB2060FAC3" author="Aoki" box="[151,279,1296,1320]" pageId="7" pageNumber="51" refString="Aoki, J. (1961) Beschreibungen von neuen Oribatiden Japans. Japanese Journal of Applied Entomology and Zoology, 5, 64 - 69." type="journal article" year="1961">Aoki, 1961</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="71C94D6AFF91194DFEE7FAEA2347FAC3" authority="Aoki, 1984" authorityName="Aoki" authorityYear="1984" box="[291,560,1296,1320]" class="Arachnida" family="Astegistidae" genus="Cultroribula" kingdom="Animalia" order="Sarcoptiformes" pageId="7" pageNumber="51" phylum="Arthropoda" rank="species" species="angulata">
<emphasis id="84BDEAFBFF91194DFEE7FAEA20D0FAC3" box="[291,423,1296,1320]" italics="true" pageId="7" pageNumber="51">C. angulata</emphasis>
<bibRefCitation id="D2584B18FF91194DFE6AFAEB2347FAC3" author="Aoki" box="[430,560,1296,1320]" pageId="7" pageNumber="51" refString="Aoki, J. (1984) New and unrecorded Oribatid mites from Kanagawa, Central Japan (I). Bulletin of the Institute of Environmental Science and Technology, Yokohama National University, 11, 107 - 118." type="journal article" year="1984">Aoki, 1984</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="71C94D6AFF91194DFDFFFAEA2204FAC3" authority="Fujikawa, 1972" authorityName="Fujikawa" authorityYear="1972" box="[571,883,1296,1320]" class="Arachnida" family="Astegistidae" genus="Cultroribula" kingdom="Animalia" order="Sarcoptiformes" pageId="7" pageNumber="51" phylum="Arthropoda" rank="species" species="elongata">
<emphasis id="84BDEAFBFF91194DFDFFFAEA23C9FAC3" box="[571,702,1296,1320]" italics="true" pageId="7" pageNumber="51">C. elongata</emphasis>
<bibRefCitation id="D2584B18FF91194DFD00FAEB2204FAC3" author="Fujikawa" box="[708,883,1296,1320]" pageId="7" pageNumber="51" refString="Fujikawa, T. (1972) A contribution to the knowledge of the oribatid fauna of Hokkaido (Acari: Oribatei). Insecta Matsumurana, 35, 127 - 183." type="journal article" year="1972">Fujikawa, 1972</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="71C94D6AFF91194DFC6EFAEA2583FAC3" authority="Subias, 2004" authorityName="Subias" authorityYear="2004" box="[938,1268,1296,1320]" class="Arachnida" family="Astegistidae" genus="Cultroribula" kingdom="Animalia" order="Sarcoptiformes" pageId="7" pageNumber="51" phylum="Arthropoda" rank="species" species="neonominata">
<emphasis id="84BDEAFBFF91194DFC6EFAEA252BFAC3" box="[938,1116,1297,1320]" italics="true" pageId="7" pageNumber="51">C. neonominata</emphasis>
<bibRefCitation id="D2584B18FF91194DFBA0FAEB2583FAC3" author="Subias" box="[1124,1268,1296,1320]" pageId="7" pageNumber="51" refString="Subias, L. S. (2004) Listado sistematico, sinonimico y biogeografico de los Acaros Oribatidos (Acariformes, Oribatida) del mundo (1748 - 2002). Graellsia, 60, 3 - 305." type="journal article" year="2004">Subías, 2004</bibRefCitation>
</taxonomicName>
are being with four pairs of genital setae. However, none of these species show outer teeth of lamellar cusps, and moreover they differ in the structure of rostral dentation and sensilli.
</paragraph>
<paragraph id="B67636E9FF91194DFF0DFA8623A0F9A0" blockId="7.[151,1436,151,1611]" pageId="7" pageNumber="51">
In the description of a Siberian species,
<taxonomicName id="71C94D6AFF91194DFD4BFA8622BEFA7F" authority="Grishina (1980)" authorityName="Grishina" authorityYear="1980" box="[655,969,1404,1428]" class="Arachnida" family="Astegistidae" genus="Cultroribula" kingdom="Animalia" order="Sarcoptiformes" pageId="7" pageNumber="51" phylum="Arthropoda" rank="species" species="ligulata">
<emphasis id="84BDEAFBFF91194DFD4BFA862270FA7F" box="[655,775,1404,1428]" italics="true" pageId="7" pageNumber="51">C. ligulata</emphasis>
,
<bibRefCitation id="D2584B18FF91194DFCD2FA8722BEFA7F" author="Grishina" box="[790,969,1404,1428]" pageId="7" pageNumber="51" refString="Grishina, L. G. (1980) New oribatid mites (Sarcoptiformes, Oribatei) from the Krasnoyarsk region. Systematics and Ecology of Animals (New and little known species of the fauna of Siberia). Nauka Press, Novosibirsk, pp. 40 - 48. (in Russian)" type="book chapter" year="1980">Grishina (1980)</bibRefCitation>
</taxonomicName>
illustrated four pairs of genital setae, but she did not mention about this structure in the description. Judging the figure 4 of her work, it is obvious that she overlooked the genital seta
<emphasis id="84BDEAFBFF91194DFE0FFA3E2096FA0B" box="[459,481,1477,1504]" italics="true" pageId="7" pageNumber="51">g1</emphasis>
, which is always at the anterior edge of genital plate. Moreover,
<taxonomicName id="71C94D6AFF91194DFB7DFA3E2447FA37" box="[1209,1328,1476,1500]" class="Arachnida" family="Astegistidae" genus="Cultroribula" kingdom="Animalia" order="Sarcoptiformes" pageId="7" pageNumber="51" phylum="Arthropoda" rank="species" species="ligulata">
<emphasis id="84BDEAFBFF91194DFB7DFA3E2447FA37" box="[1209,1328,1476,1500]" italics="true" pageId="7" pageNumber="51">C. ligulata</emphasis>
</taxonomicName>
have different structures of rostrum (with only three large teeth deeply incised in-between), lamella with smoothly rounded narrow cusp, sensillus with rough barbs throughout its head, and very short interlamellar and notogastral setae, in which its clearly differs from
<taxonomicName id="71C94D6AFF91194DFE20F9CF2301F9A0" box="[484,630,1588,1611]" class="Arachnida" family="Astegistidae" genus="Cultroribula" kingdom="Animalia" order="Sarcoptiformes" pageId="7" pageNumber="51" phylum="Arthropoda" rank="species" species="rarisetosa" status="sp. nov.">
<emphasis id="84BDEAFBFF91194DFE20F9CF2301F9A0" box="[484,630,1588,1611]" italics="true" pageId="7" pageNumber="51">C. rarisetosa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9F8E5780FF91194DFDB9F9C823A0F9A0" box="[637,727,1587,1611]" pageId="7" pageNumber="51" rank="species">
<emphasis id="84BDEAFBFF91194DFDB9F9C823A0F9A0" bold="true" box="[637,727,1587,1611]" pageId="7" pageNumber="51">sp. nov.</emphasis>
</taxonomicNameLabel>
</paragraph>
</subSubSection>
</treatment>
</document>
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