272 lines
32 KiB
XML
272 lines
32 KiB
XML
<document id="F9F84A535FFE4C687CB058C40FA69356" ID-DOI="10.5281/zenodo.7225400" IM.bibliography_approvedBy="valdenar" IM.illustrations_approvedBy="valdenar" IM.materialsCitations_approvedBy="valdenar" IM.metadata_approvedBy="valdenar" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="valdenar" IM.treatmentCitations_approvedBy="valdenar" IM.treatments_approvedBy="valdenar" checkinTime="1702532099912" checkinUser="valdenar" docAuthor="Grzelak, Katarzyna & Sørensen, Martin V." docDate="2022" docId="3E3587AEFFE3FFCF078F75FBFD4617DE" docLanguage="en" docName="ZoolStud.55.52.1-13.pdf" docOrigin="Zoological Studies 55 (52)" docSource="http://dx.doi.org/10.5281/zenodo.7225400" docStyle="DocumentStyle:E2601EF9E8BBDC4537EE724C10FF512F.1:ZoolStud.2015-.journal_article" docStyleId="E2601EF9E8BBDC4537EE724C10FF512F" docStyleName="ZoolStud.2015-.journal_article" docStyleVersion="1" docTitle="Amynthas konkakinh Nguyen, Lam & Nguyen 2022" docType="treatment" docVersion="2" lastPageNumber="12" masterDocId="C20CFFD6FFEAFFC4071A7066FFA7117E" masterDocTitle="Echinoderes blazeji Grzelak & amp; Sørensen 2022, sp. nov." masterLastPageNumber="13" masterPageNumber="1" pageNumber="10" updateTime="1704842016616" updateUser="ExternalLinkService">
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<mods:titleInfo id="C06CEA253DD9EB25344F613347B50D13">
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<mods:title id="B7CCB3B592C26F9589EEF67C44C0872C">Echinoderes blazeji Grzelak & amp; Sørensen 2022, sp. nov.</mods:title>
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<mods:name id="881745C38EA7E3EB9D0679787A2E093D" type="personal">
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<mods:namePart id="1C3610D9EE1B849FD2C98B8FFF830855">Grzelak, Katarzyna</mods:namePart>
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<mods:name id="DBB512D3CB2E3EE69D7775D2BF4B47D8" type="personal">
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<mods:roleTerm id="496EE6CA46EA4FD4BB571436FAD620A7">Author</mods:roleTerm>
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<mods:namePart id="BC68063AA2AD6F3D277A23C5C1C189AD">Sørensen, Martin V.</mods:namePart>
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<mods:typeOfResource id="C976F1B87E1E26AE1F98A8BB80D706AE">text</mods:typeOfResource>
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<mods:title id="C2CF22884488A90E49DE0C36694BA46C">Zoological Studies</mods:title>
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</mods:titleInfo>
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<mods:part id="FEC579C33A99CDFF852B5B685DDC4195">
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<mods:date id="A7A7CFF62C690BF92561FAC5A815BC9A">2022</mods:date>
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<mods:detail id="C9124BD6929DEDCA74DEED91A05802C8" type="volume">
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<mods:number id="0634B33B31A223D1436F8AE8A40BCC80">55</mods:number>
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<mods:detail id="1B64A88EEF35AD3ED0CF5D260F9A5E88" type="issue">
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<mods:number id="B1DE87C98D3F798A722B6EBCEC232EA9">52</mods:number>
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<mods:extent id="646B6E7AE23CD49DAC999C7009002BB0" unit="page">
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<mods:start id="1C0D3D76497A055F0AAFBC583FCC922A">1</mods:start>
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<mods:end id="FC00CE262FE103AFC7AE48232F864883">13</mods:end>
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<mods:url id="752442978B5043D6B2EB2E06EC3CEBE1">http://dx.doi.org/10.5281/zenodo.7225400</mods:url>
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<mods:classification id="FBD2B3FB1DB6E8A1C99CFE0482F8B89A">journal article</mods:classification>
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<mods:identifier id="B5C5D63418C67DAED438F33A264FFF5C" type="DOI">10.5281/zenodo.7225400</mods:identifier>
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</mods:mods>
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<treatment id="3E3587AEFFE3FFCF078F75FBFD4617DE" LSID="urn:lsid:plazi:treatment:3E3587AEFFE3FFCF078F75FBFD4617DE" httpUri="http://treatment.plazi.org/id/3E3587AEFFE3FFCF078F75FBFD4617DE" lastPageId="11" lastPageNumber="12" pageId="9" pageNumber="10">
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<subSubSection id="FE866533FFE3FFCD078F75FBFDB61753" pageId="9" pageNumber="10" type="nomenclature">
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<paragraph id="B62336B8FFE3FFCD078F75FBFD7314C8" blockId="9.[149,724,1437,1581]" box="[149,724,1437,1462]" pageId="9" pageNumber="10">
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<heading id="ED6B81D4FFE3FFCD078F75FBFD7314C8" bold="true" box="[149,724,1437,1462]" centered="true" fontSize="10" level="2" pageId="9" pageNumber="10" reason="2">
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<taxonomicName id="719C4D3BFFE3FFCD078F75FBFD7314C8" authority="Nguyen, Lam & Nguyen" authorityName="Nguyen, Lam & Nguyen" authorityYear="2022" box="[149,724,1437,1462]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Crassiclitellata" pageId="9" pageNumber="10" phylum="Annelida" rank="species" species="konkakinh" status="sp. nov.">
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<emphasis id="84E8EAAAFFE3FFCD078F75FBFD7314C8" bold="true" box="[149,724,1437,1462]" pageId="9" pageNumber="10">
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<emphasis id="84E8EAAAFFE3FFCD078F75FBFE0614C8" bold="true" box="[149,417,1437,1462]" italics="true" pageId="9" pageNumber="10">Amynthas konkakinh</emphasis>
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Nguyen, Lam & Nguyen
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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<paragraph id="B62336B8FFE3FFCD069975DBFDB61753" blockId="9.[149,724,1437,1581]" pageId="9" pageNumber="10">
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<emphasis id="84E8EAAAFFE3FFCD069975DBFE4114A8" bold="true" box="[387,486,1469,1494]" pageId="9" pageNumber="10">sp. nov.</emphasis>
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lsid:zoobank.org:act:
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<uuid id="C23A0C6DFFE3FFCD069B75BAFDAE1771" pageId="9" pageNumber="10">833AA0A9-E3EF-4AE4-9F87- 9C75E84696A7</uuid>
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(
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<figureCitation id="2EA72A3DFFE3FFCD067A7672FE001753" box="[352,423,1556,1581]" captionStart="Fig" captionStartId="10.[174,209,1897,1917]" captionTargetBox="[244,1377,252,1869]" captionTargetId="figure-0@10.[234,1391,235,1883]" captionTargetPageId="10" captionText="Fig. 6. Amynthas konkakinh sp. nov., holotype. A, F: Male pore region, ventral view (mp = male pore; gm = gential markings); H: Spermathecal region (sp = spermathecal pore; gm = gential markings); C, E: Spermathecae (amp = ampulla; dv = diverticula); D, G: Intestinal caeca; B: Prostatic gland (ag = accessory gland). Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10476489" httpUri="https://zenodo.org/record/10476489/files/figure.png" pageId="9" pageNumber="10">Fig. 6</figureCitation>
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,
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<tableCitation id="FB1E0303FFE3FFCD06A87672FDA11753" box="[434,518,1556,1581]" captionStart="Table 1" captionStartId="3.[136,204,658,683]" captionText="Table 1. Morphological comparison among giant earthworm species" pageId="9" pageNumber="10">Table 1</tableCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="FE866533FFE3FFCF07DF7636FD4515E8" lastPageId="11" lastPageNumber="12" pageId="9" pageNumber="10" type="materials_examined">
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<paragraph id="B62336B8FFE3FFCD07DF7636FD4517B7" blockId="9.[136,738,1616,1961]" pageId="9" pageNumber="10">
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<emphasis id="84E8EAAAFFE3FFCD07DF7636FE161717" box="[197,433,1616,1641]" italics="true" pageId="9" pageNumber="10">Material examined</emphasis>
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:
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<materialsCitation id="06F43CE5FFE3FFCD06DF7636FD4517B7" collectingDate="2010-10-20" collectorName="Pham Thi Mai." country="Vietnam" location="One" municipality="Kon Ka Kinh National Park" pageId="9" pageNumber="10" specimenCode="CTU-EW038" specimenCount="1" stateProvince="Gia Lai" typeStatus="holotype">
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<typeStatus id="6927881AFFE3FFCD06DF7636FD9D1717" box="[453,570,1616,1641]" pageId="9" pageNumber="10" type="holotype">Holotype</typeStatus>
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:
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<location id="B3436063FFE3FFCD05537636FDDA1714" LSID="urn:lsid:plazi:treatment:3E3587AEFFE3FFCF078F75FBFD4617DE:B3436063FFE3FFCD05537636FDDA1714" box="[585,637,1616,1642]" country="Vietnam" municipality="Kon Ka Kinh National Park" name="One" pageId="9" pageNumber="10" stateProvince="Gia Lai">One</location>
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mature (
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<specimenCode id="E63A9EC3FFE3FFCD07957616FE9F17F4" box="[143,312,1648,1674]" collectionCode="CTU-EW" pageId="9" pageNumber="10">CTU-EW038</specimenCode>
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-p01) primary forest, southwestern part of
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<collectingMunicipality id="5647ACC2FFE3FFCD07EA76F6FDCB17D7" box="[240,620,1680,1705]" pageId="9" pageNumber="10">Kon Ka Kinh National Park</collectingMunicipality>
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,
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<collectingRegion id="7458F85AFFE3FFCD056476F6FF5617B7" country="Vietnam" name="Gia Lai" pageId="9" pageNumber="10">Gia Lai province</collectingRegion>
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,
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<collectingCountry id="CE8B7628FFE3FFCD07E176D6FEC617B7" box="[251,353,1712,1737]" name="Vietnam" pageId="9" pageNumber="10">Vietnam</collectingCountry>
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,
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<date id="C2221078FFE3FFCD067176D6FE5317B7" box="[363,500,1712,1737]" pageId="9" pageNumber="10" value="2010-10-20">
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<collectingDate id="D266E990FFE3FFCD067176D6FE5317B7" box="[363,500,1712,1737]" pageId="9" pageNumber="10" value="2010-10-20">20/10/2010</collectingDate>
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</date>
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, coll.
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<collectorName id="1B69536EFFE3FFCD052C76D6FD4517B7" box="[566,738,1712,1737]" pageId="9" pageNumber="10">Pham Thi Mai.</collectorName>
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</materialsCitation>
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</paragraph>
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<paragraph id="B62336B8FFE3FFCD07DF76B6FB831008" blockId="9.[136,738,1616,1961]" lastBlockId="9.[813,1414,253,1975]" pageId="9" pageNumber="10">
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<emphasis id="84E8EAAAFFE3FFCD07DF76B6FEEE1797" box="[197,329,1744,1769]" italics="true" pageId="9" pageNumber="10">Diagnosis</emphasis>
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: Large-size worm, length ca.
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<quantity id="71649B5DFFE3FFCD07927696FF4B1677" box="[136,236,1776,1801]" metricMagnitude="-1" metricUnit="m" metricValue="6.7" pageId="9" pageNumber="10" unit="mm" value="670.0">670 mm</quantity>
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, diameter ca.
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<quantity id="71649B5DFFE3FFCD068A7696FE5C1677" box="[400,507,1776,1801]" metricMagnitude="-2" metricUnit="m" metricValue="1.65" pageId="9" pageNumber="10" unit="mm" value="16.5">16.5 mm</quantity>
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. First dorsal pore in 12/13. Prostomium epilobous. Spermathecal pores round and small, ventrally paired in intersegmental furrows 5/6/7/8/9. Male pores located in xviii, without copulatory pouches. Genital makings present on both spermathecal and male regions (on viii and xviii). Holandric. Spermathecal ampulla cylindrical-shaped, duct short; diverticula coiled. Testis sacs separated. Intestinal caeca simple. Septa 8/9/10 absent.
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</paragraph>
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<paragraph id="B62336B8FFE3FFCD0473711BFB7010C8" blockId="9.[813,1414,253,1975]" pageId="9" pageNumber="10">
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<emphasis id="84E8EAAAFFE3FFCD0473711BFC4C10E8" box="[873,1003,381,406]" italics="true" pageId="9" pageNumber="10">Etymology</emphasis>
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:
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<taxonomicName id="719C4D3BFFE3FFCD04E5711BFB2110E8" authorityName="Nguyen, Lam & Nguyen" authorityYear="2022" box="[1023,1158,381,406]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Crassiclitellata" pageId="9" pageNumber="10" phylum="Annelida" rank="species" species="konkakinh">
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“
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<emphasis id="84E8EAAAFFE3FFCD0312711BFB2110E8" box="[1032,1158,381,406]" italics="true" pageId="9" pageNumber="10">konkakinh</emphasis>
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</taxonomicName>
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” is a noun in apposition to emphasize the
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<typeStatus id="6927881AFFE3FFCD035F71FBFBD010C8" box="[1093,1143,413,438]" pageId="9" pageNumber="10">type</typeStatus>
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locality.
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</paragraph>
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<paragraph id="B62336B8FFE3FFCD047371DBFC711528" blockId="9.[813,1414,253,1975]" pageId="9" pageNumber="10">
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<emphasis id="84E8EAAAFFE3FFCD047371DBFBA310A8" box="[873,1028,445,470]" italics="true" pageId="9" pageNumber="10">Description</emphasis>
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:
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<emphasis id="84E8EAAAFFE3FFCD030671DBFA8C10A8" box="[1052,1323,445,470]" italics="true" pageId="9" pageNumber="10">External characters</emphasis>
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: Body cylindrial, length
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<quantity id="71649B5DFFE3FFCD030E71BBFB241088" box="[1044,1155,477,502]" metricMagnitude="-1" metricUnit="m" metricValue="6.7" pageId="9" pageNumber="10" unit="mm" value="670.0">670 mm</quantity>
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, diameter
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<quantity id="71649B5DFFE3FFCD020871BBFA231088" box="[1298,1412,477,502]" metricMagnitude="-2" metricUnit="m" metricValue="1.65" pageId="9" pageNumber="10" unit="mm" value="16.5">16.5 mm</quantity>
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(
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<quantity id="71649B5DFFE3FFCD0429719BFC2B1368" box="[819,908,509,534]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" pageId="9" pageNumber="10" unit="mm" value="20.0">20 mm</quantity>
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/viii,
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<quantity id="71649B5DFFE3FFCD04DE719BFBBD1368" box="[964,1050,509,534]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="9" pageNumber="10" unit="mm" value="15.0">15 mm</quantity>
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/xvi,
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<quantity id="71649B5DFFE3FFCD034E719BFB0D1368" box="[1108,1194,509,534]" metricMagnitude="-2" metricUnit="m" metricValue="1.6" pageId="9" pageNumber="10" unit="mm" value="16.0">16 mm</quantity>
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/xviii,
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<quantity id="71649B5DFFE3FFCD03F5719BFAE01368" box="[1263,1351,509,534]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="9" pageNumber="10" unit="mm" value="15.0">15 mm</quantity>
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/xxx), segments 204, weight
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<quantity id="71649B5DFFE3FFCD034B727BFB671348" box="[1105,1216,541,566]" metricMagnitude="-1" metricUnit="kg" metricValue="1.1968" pageId="9" pageNumber="10" unit="g" value="119.68">119.68 g</quantity>
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. Dorsa blackish grey, but ventra paler. Prostomium epilobous. First dorsal pore large, in 12/13. Setae perichaetine, short; pre-clitellar setae stouter and sparser than post-clitellar setae,
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<quantity id="71649B5DFFE3FFCD030772FBFBCE13C8" box="[1053,1129,669,694]" metricMagnitude="0" metricUnit="m" metricValue="3.3274" pageId="9" pageNumber="10" unit="in" value="131.0">131 in</quantity>
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v,
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<quantity id="71649B5DFFE3FFCD038B72FBFB7913C8" box="[1169,1246,669,694]" metricMagnitude="0" metricUnit="m" metricValue="3.6576" pageId="9" pageNumber="10" unit="in" value="144.0">144 in</quantity>
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viii,
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<quantity id="71649B5DFFE3FFCD020072FBFAC113C8" box="[1306,1382,669,694]" metricMagnitude="0" metricUnit="m" metricValue="4.7498" pageId="9" pageNumber="10" unit="in" value="187.0">187 in</quantity>
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x,
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<quantity id="71649B5DFFE3FFCD043772DBFCD113A8" box="[813,886,701,726]" metricMagnitude="0" metricUnit="m" metricValue="5.5626" pageId="9" pageNumber="10" unit="in" value="219.0">219 in</quantity>
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xx,
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<quantity id="71649B5DFFE3FFCD04B272DBFC5513A8" box="[936,1010,701,726]" metricMagnitude="0" metricUnit="m" metricValue="5.740399999999999" pageId="9" pageNumber="10" unit="in" value="226.0">226 in</quantity>
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xxv,
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<quantity id="71649B5DFFE3FFCD032A72DBFBDE13A8" box="[1072,1145,701,726]" metricMagnitude="0" metricUnit="m" metricValue="5.2832" pageId="9" pageNumber="10" unit="in" value="208.0">208 in</quantity>
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xxx, 50 between male porophores in xviii; setal distance aa = ab, zz = 1.0-2.0zy. Clitellum annular, xiv-1/2xvii, blackish brown, smooth and without setae and dorsal pores. Female pore single, mid-ventral in xiv. Spermathecal pores very small, ventrally paired in intersegmental furrows 5/6/7/8/9, hardly visible externally. A genital marking ventrally present on viii. Male pores directly opened in xviii; ventral distance between male porophores about 0.35x body circumference. A pair of genital markings present in front of setal ring of xviii, slightly medial to male pores.
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</paragraph>
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<paragraph id="B62336B8FFE3FFCD0473743BFA8E17A8" blockId="9.[813,1414,253,1975]" pageId="9" pageNumber="10">
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<emphasis id="84E8EAAAFFE3FFCD0473743BFBC51508" box="[873,1122,1117,1142]" italics="true" pageId="9" pageNumber="10">Internal characters</emphasis>
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: Septa 4/5/6/7/8 very thickened, 8/9/10 absent, 10/11/12/13 thin. Oesophageal gizzard within viii-x. Intestinal origin at xv; caeca simple, originating at xxvii and extending anteriorly to xxii. Last hearts in xiii. Pharyngeal micronephridia developed in 5/6/7. Lymph glands present from 32/33. Typhlosole simple, lamelliform, poorly developed. Spermathecae paired ventrally in vi- ix. Spermathecal ampulla cylindrical-shaped; duct short, about 1/3 ampulla length. Diverticula very shorter than ampulla, opalescent, coiled, directly attached to base of ampulla. Accessory glands absent. Holandric. Testis sacs separated, in x and xi. Seminal vesicles whitish yellow, poorly developed within xi-xii. Oviduct poorly developed on septum 12/13 posteriorly; ovaries developed in 13/14. Prostate glands racemose, paired within xviii; prostatic ducts strong, slightly getting smaller distally. A pair of accessory glands present.
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</paragraph>
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<paragraph id="B62336B8FFE3FFCD047376BBFC4C1648" blockId="9.[813,1414,253,1975]" pageId="9" pageNumber="10">
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<emphasis id="84E8EAAAFFE3FFCD047376BBFC181788" box="[873,959,1757,1782]" italics="true" pageId="9" pageNumber="10">Habitat</emphasis>
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: The species was collected from soil surface or litter layer in the rich forest. It can be epigeic species.
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</paragraph>
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<paragraph id="B62336B8FFE3FFCF0473775BFEFA1028" blockId="9.[813,1414,253,1975]" lastBlockId="11.[136,738,253,1174]" lastPageId="11" lastPageNumber="12" pageId="9" pageNumber="10">
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<emphasis id="84E8EAAAFFE3FFCD0473775BFC731628" box="[873,980,1853,1878]" italics="true" pageId="9" pageNumber="10">Remarks</emphasis>
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: The new species can be classified into the
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<emphasis id="84E8EAAAFFE3FFCD0494773BFBAF1608" box="[910,1032,1885,1910]" italics="true" pageId="9" pageNumber="10">diffringens</emphasis>
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species group characterised by holandric and four pairs of spermathecal pores in intersegments 5/6/7/8/9 (
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<bibRefCitation id="D20D4B49FFE3FFCD034177FBFADC16C8" author="Sims RW & Easton EG" box="[1115,1403,1949,1975]" pageId="9" pageNumber="10" pagination="169 - 268" refId="ref6856" refString="Sims RW, Easton EG. 1972. A numerical revision of the earthworm genus Pheretima auct. (Megascolecidae: Oligochaeta) with the recognition of new genera and an appendix on the earthworms collected by the Royal Society North Borneo Expedition. Biological Journal of the Linnean Society 4: 169 - 268. doi: http: // dx. doi. org / 10.1111 / j. 1095 - 8312.1972. tb 00694. x." type="journal article" year="1972">Sims and Easton 1972</bibRefCitation>
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). However, it can be distinguished by its large size and pattern of genital markings in spermathecal and male regions.
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</paragraph>
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<caption id="E2E36630FFE0FFCE07B4770FFC2916CB" ID-DOI="http://doi.org/10.5281/zenodo.10476489" ID-Zenodo-Dep="10476489" httpUri="https://zenodo.org/record/10476489/files/figure.png" pageId="10" pageNumber="11" startId="10.[174,209,1897,1917]" targetBox="[244,1377,252,1869]" targetPageId="10" targetType="figure">
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<paragraph id="B62336B8FFE0FFCE07B4770FFC2916CB" blockId="10.[174,1452,1897,1973]" pageId="10" pageNumber="11">
|
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<emphasis id="84E8EAAAFFE0FFCE07B4770FFF491603" bold="true" box="[174,238,1897,1917]" pageId="10" pageNumber="11">Fig. 6.</emphasis>
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<taxonomicName id="719C4D3BFFE0FFCE07E5770FFE6F1603" authority="Grzelak & Sørensen, 2022" authorityName="Grzelak & Sørensen" authorityYear="2022" box="[255,456,1897,1917]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Crassiclitellata" pageId="10" pageNumber="11" phylum="Annelida" rank="species" species="konkakinh" status="sp. nov.">
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<emphasis id="84E8EAAAFFE0FFCE07E5770FFE6F1603" box="[255,456,1897,1917]" italics="true" pageId="10" pageNumber="11">Amynthas konkakinh</emphasis>
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</taxonomicName>
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|
<taxonomicNameLabel id="9FDB57D1FFE0FFCE06CB770FFD861603" box="[465,545,1897,1917]" pageId="10" pageNumber="11" rank="species">sp. nov.</taxonomicNameLabel>
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, holotype. A, F: Male pore region, ventral view (mp = male pore; gm = gential markings); H: Spermathecal region (sp = spermathecal pore; gm = gential markings); C, E: Spermathecae (amp = ampulla; dv = diverticula); D, G: Intestinal caeca; B: Prostatic gland (ag = accessory gland). Scale bar = 1 mm.
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</paragraph>
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</caption>
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|
<paragraph id="B62336B8FFE1FFCF07DF713BFF1A13E8" blockId="11.[136,738,253,1174]" pageId="11" pageNumber="12">
|
|
<taxonomicName id="719C4D3BFFE1FFCF07DF713BFE6D1008" authority="Grzelak & Sørensen, 2022" authorityName="Grzelak & Sørensen" authorityYear="2022" box="[197,458,349,374]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Crassiclitellata" pageId="11" pageNumber="12" phylum="Annelida" rank="species" species="konkakinh" status="sp. nov.">
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<emphasis id="84E8EAAAFFE1FFCF07DF713BFE6D1008" box="[197,458,349,374]" italics="true" pageId="11" pageNumber="12">Amynthas konkakinh</emphasis>
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</taxonomicName>
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|
<taxonomicNameLabel id="9FDB57D1FFE1FFCF06CC713BFD9D1008" box="[470,570,349,374]" pageId="11" pageNumber="12" rank="species">sp. nov.</taxonomicNameLabel>
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is especially similar to
|
|
<taxonomicName id="719C4D3BFFE1FFCF0608711BFE7C10E8" authority="Grzelak & Sørensen, 2022" authorityName="Grzelak & Sørensen" authorityYear="2022" box="[274,475,381,406]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Crassiclitellata" pageId="11" pageNumber="12" phylum="Annelida" rank="species" species="antoanensis" status="sp. nov.">
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<emphasis id="84E8EAAAFFE1FFCF0608711BFE7C10E8" box="[274,475,381,406]" italics="true" pageId="11" pageNumber="12">A. antoanensis</emphasis>
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</taxonomicName>
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|
<taxonomicNameLabel id="9FDB57D1FFE1FFCF06F2711BFDF510E8" box="[488,594,381,406]" pageId="11" pageNumber="12" rank="species">sp. nov.</taxonomicNameLabel>
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by having large size, first dorsal pore in 12/13, four pairs of spermathecal pores in 5/6/7/8/9, intestinal caeca simple, septa 8/9/10 absent, and prostomium epilobous (
|
|
<tableCitation id="FB1E0303FFE1FFCF0616719BFECF1368" box="[268,360,509,534]" captionStart="Table 1" captionStartId="3.[136,204,658,683]" captionText="Table 1. Morphological comparison among giant earthworm species" pageId="11" pageNumber="12">Table 1</tableCitation>
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). However,
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|
<taxonomicName id="719C4D3BFFE1FFCF06E4719BFD141368" authority="Grzelak & Sørensen, 2022" authorityName="Grzelak & Sørensen" authorityYear="2022" box="[510,691,509,534]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Crassiclitellata" pageId="11" pageNumber="12" phylum="Annelida" rank="species" species="antoanensis" status="sp. nov.">
|
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<emphasis id="84E8EAAAFFE1FFCF06E4719BFD141368" box="[510,691,509,534]" italics="true" pageId="11" pageNumber="12">A. antoanensis</emphasis>
|
|
</taxonomicName>
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|
<taxonomicNameLabel id="9FDB57D1FFE1FFCF05A7719BFF1E1348" pageId="11" pageNumber="12" rank="species">sp. nov.</taxonomicNameLabel>
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|
can be recognized from
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|
<taxonomicName id="719C4D3BFFE1FFCF06F9727BFDDB1348" authority="Grzelak & Sørensen, 2022" authorityName="Grzelak & Sørensen" authorityYear="2022" box="[483,636,541,566]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Crassiclitellata" pageId="11" pageNumber="12" phylum="Annelida" rank="species" species="konkakinh" status="sp. nov.">
|
|
<emphasis id="84E8EAAAFFE1FFCF06F9727BFDDB1348" box="[483,636,541,566]" italics="true" pageId="11" pageNumber="12">A. konkakinh</emphasis>
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</taxonomicName>
|
|
<taxonomicNameLabel id="9FDB57D1FFE1FFCF059F727BFD451348" box="[645,738,541,566]" pageId="11" pageNumber="12" rank="species">sp. nov.</taxonomicNameLabel>
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|
by having four pairs of genital markings in viii and ix, and two pairs of genital markings in mid-ventral xviii.
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|
</paragraph>
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|
<paragraph id="B62336B8FFE1FFCF07DF72FBFD4515E8" blockId="11.[136,738,253,1174]" pageId="11" pageNumber="12">
|
|
The new species is also fairly similar to the other large
|
|
<taxonomicName id="719C4D3BFFE1FFCF067F72DBFE4D13A8" authorityName="Kinberg" authorityYear="1867" box="[357,490,701,726]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Crassiclitellata" pageId="11" pageNumber="12" phylum="Annelida" rank="genus">
|
|
<emphasis id="84E8EAAAFFE1FFCF067F72DBFE4D13A8" box="[357,490,701,726]" italics="true" pageId="11" pageNumber="12">Amynthas</emphasis>
|
|
</taxonomicName>
|
|
earthworms from
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|
<collectingCountry id="CE8B7628FFE1FFCF079272BBFF521388" box="[136,245,733,758]" name="Vietnam" pageId="11" pageNumber="12">Vietnam</collectingCountry>
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,
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|
<taxonomicName id="719C4D3BFFE1FFCF061F72BBFD271388" authority="(Thai, 1984)" baseAuthorityName="Thai" baseAuthorityYear="1984" box="[261,640,733,758]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Crassiclitellata" pageId="11" pageNumber="12" phylum="Annelida" rank="species" species="dangi">
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|
<emphasis id="84E8EAAAFFE1FFCF061F72BBFE721388" box="[261,469,733,758]" italics="true" pageId="11" pageNumber="12">Amynthas dangi</emphasis>
|
|
(
|
|
<bibRefCitation id="D20D4B49FFE1FFCF06F372BBFDDE1388" author="Thai TB" box="[489,633,733,758]" pageId="11" pageNumber="12" pagination="1317 - 1327" refId="ref7022" refString="Thai TB. 1984. New species of the genus Pheretima in Vietnam. Zoologicheskii Jurnal 63 (9): 1317 - 1327." type="journal article" year="1984">Thai, 1984</bibRefCitation>
|
|
)
|
|
</taxonomicName>
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|
and
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|
<taxonomicName id="719C4D3BFFE1FFCF05DD72BBFDEB1268" authority="(Thai & Tran, 1986)" authorityName="Grzelak & Sørensen" authorityYear="2022" baseAuthorityName="Thai & Tran" baseAuthorityYear="1986" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Crassiclitellata" pageId="11" pageNumber="12" phylum="Annelida" rank="species" species="munglonganus">
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|
<emphasis id="84E8EAAAFFE1FFCF05DD72BBFEE71268" italics="true" pageId="11" pageNumber="12">A. munglonganus</emphasis>
|
|
(
|
|
<bibRefCitation id="D20D4B49FFE1FFCF0649729BFDE21268" author="Thai TB & Tran BC" box="[339,581,765,790]" pageId="11" pageNumber="12" pagination="4 - 12" refId="ref6936" refString="Thai TB, Tran BC. 1986. The fauna of earthworms of mountain district Ki Son (Nghe Tinh province, Vietnam) and descriptions of new species of the genus Pheretima Kinberg. TAP CHI SINH HOC 8 (4): 4 - 12." type="journal article" year="1986">Thai & Tran, 1986</bibRefCitation>
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|
)
|
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</taxonomicName>
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|
, by its very large size, holandric, prostomium epilobous, septa 8/9/10 absent, last hearts in xiii, and copulatory pouches absent. However,
|
|
<taxonomicName id="719C4D3BFFE1FFCF06D2733BFD8E1208" baseAuthorityName="Thai" baseAuthorityYear="1984" box="[456,553,861,886]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Crassiclitellata" pageId="11" pageNumber="12" phylum="Annelida" rank="species" species="dangi">
|
|
<emphasis id="84E8EAAAFFE1FFCF06D2733BFD8E1208" box="[456,553,861,886]" italics="true" pageId="11" pageNumber="12">A. dangi</emphasis>
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|
</taxonomicName>
|
|
differs from the new species in having two pairs of spermathecal pores in 7/8/9, a genital marking closed to the male pore and an additional three pairs of genital markings on ventral xviii and xix.
|
|
<taxonomicName id="719C4D3BFFE1FFCF050B73BBFD461288" authorityName="Grzelak & Sørensen" authorityYear="2022" baseAuthorityName="Thai & Tran" baseAuthorityYear="1986" box="[529,737,989,1014]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Crassiclitellata" pageId="11" pageNumber="12" phylum="Annelida" rank="species" species="munglonganus">
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|
<emphasis id="84E8EAAAFFE1FFCF050B73BBFD461288" box="[529,737,989,1014]" italics="true" pageId="11" pageNumber="12">A. munglonganus</emphasis>
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|
</taxonomicName>
|
|
is distinguished from the new species by the first dorsal pore in 12/13, three pairs of small genital markings located closely to spermathecal pores in 5/6/7/8, a medium genital marking next to the male pore, and different shape of spermathecal ampulla.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="FE866533FFE1FFCF079274DBFD4617DE" pageId="11" pageNumber="12" type="key">
|
|
<paragraph id="B62336B8FFE1FFCF079274DBFDEA15A8" blockId="11.[136,589,1213,1238]" box="[136,589,1213,1238]" pageId="11" pageNumber="12">
|
|
<emphasis id="84E8EAAAFFE1FFCF079274DBFDEA15A8" bold="true" box="[136,589,1213,1238]" pageId="11" pageNumber="12">
|
|
Key to large earthworms in
|
|
<collectingCountry id="CE8B7628FFE1FFCF06FC74DBFDEA15A8" box="[486,589,1213,1238]" name="Vietnam" pageId="11" pageNumber="12">Vietnam</collectingCountry>
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|
</emphasis>
|
|
</paragraph>
|
|
<paragraph id="B62336B8FFE1FFCF0792749AFD46146E" blockId="11.[136,738,1276,1696]" box="[136,737,1276,1296]" pageId="11" pageNumber="12">1. Four pairs of spermathecal pores in 5/6/7/8/9 .................. 2</paragraph>
|
|
<paragraph id="B62336B8FFE1FFCF07927571FD461455" blockId="11.[136,738,1276,1696]" box="[136,737,1302,1323]" pageId="11" pageNumber="12">- Only two or three pairs of spermathecal pores ................. 3</paragraph>
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|
<paragraph id="B62336B8FFE1FFCF07927557FD46141E" blockId="11.[136,738,1276,1696]" pageId="11" pageNumber="12">
|
|
2. Genital markings paired in viii, ix, and mid-ventral xviii. Spermathecal ampulla oval-shaped ...........
|
|
<taxonomicName id="719C4D3BFFE1FFCF054E752AFD46141E" authorityName="Grzelak & Sørensen" authorityYear="2022" box="[596,737,1356,1376]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Crassiclitellata" pageId="11" pageNumber="12" phylum="Annelida" rank="species" species="antoanensis">
|
|
<emphasis id="84E8EAAAFFE1FFCF054E752AFD46141E" box="[596,737,1356,1376]" italics="true" pageId="11" pageNumber="12">A. antoanensis</emphasis>
|
|
</taxonomicName>
|
|
</paragraph>
|
|
<paragraph id="B62336B8FFE1FFCF07927501FD4614CE" blockId="11.[136,738,1276,1696]" pageId="11" pageNumber="12">
|
|
- Genital markings single in viii, and paired in front of male pores in xviii. Spermathecal ampulla cylindrical .................. ............................. ...........................................
|
|
<taxonomicName id="719C4D3BFFE1FFCF057D75FAFD4614CE" authorityName="Grzelak & Sørensen" authorityYear="2022" box="[615,737,1436,1456]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Crassiclitellata" pageId="11" pageNumber="12" phylum="Annelida" rank="species" species="konkakinh">
|
|
<emphasis id="84E8EAAAFFE1FFCF057D75FAFD4614CE" box="[615,737,1436,1456]" italics="true" pageId="11" pageNumber="12">A. konkakinh</emphasis>
|
|
</taxonomicName>
|
|
</paragraph>
|
|
<paragraph id="B62336B8FFE1FFCF079275D1FD4614B5" blockId="11.[136,738,1276,1696]" box="[136,737,1463,1483]" pageId="11" pageNumber="12">
|
|
3. Two pairs of spermathecal pores in 7/8/9 .............
|
|
<taxonomicName id="719C4D3BFFE1FFCF058975D1FD4614B5" baseAuthorityName="Thai" baseAuthorityYear="1984" box="[659,737,1463,1483]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Crassiclitellata" pageId="11" pageNumber="12" phylum="Annelida" rank="species" species="dangi">
|
|
<emphasis id="84E8EAAAFFE1FFCF058975D1FD4614B5" box="[659,737,1463,1483]" italics="true" pageId="11" pageNumber="12">A. dangi</emphasis>
|
|
</taxonomicName>
|
|
</paragraph>
|
|
<paragraph id="B62336B8FFE1FFCF079275B7FD46177E" blockId="11.[136,738,1276,1696]" pageId="11" pageNumber="12">- Three pairs of spermathecal pores in 5/6/7/8 or 6/7/8/9 ..... ........................................................................................... 4</paragraph>
|
|
<paragraph id="B62336B8FFE1FFCF07927661FD471765" blockId="11.[136,738,1276,1696]" box="[136,736,1542,1563]" pageId="11" pageNumber="12">
|
|
4. Spermathecal pores in 6/7/8/9 ..........................
|
|
<taxonomicName id="719C4D3BFFE1FFCF056D7661FD471765" baseAuthorityName="Thai" baseAuthorityYear="1982" box="[631,736,1543,1563]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Crassiclitellata" pageId="11" pageNumber="12" phylum="Annelida" rank="species" species="ghilarovi">
|
|
<emphasis id="84E8EAAAFFE1FFCF056D7661FD471765" box="[631,736,1543,1563]" italics="true" pageId="11" pageNumber="12">A. ghilarovi</emphasis>
|
|
</taxonomicName>
|
|
</paragraph>
|
|
<paragraph id="B62336B8FFE1FFCF07927647FD46174B" blockId="11.[136,738,1276,1696]" box="[136,737,1569,1589]" pageId="11" pageNumber="12">- Spermathecal pores in 5/6/7/8 .......................................... 5</paragraph>
|
|
<paragraph id="B62336B8FFE1FFCF0792765AFD461715" blockId="11.[136,738,1276,1696]" pageId="11" pageNumber="12">
|
|
5. Genital markings absent in both spermathecal and male pore regions ............................................
|
|
<taxonomicName id="719C4D3BFFE1FFCF05217631FD461715" authorityName="Grzelak & Sørensen" authorityYear="2022" baseAuthorityName="Thai & Tran" baseAuthorityYear="1986" box="[571,737,1623,1643]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Crassiclitellata" pageId="11" pageNumber="12" phylum="Annelida" rank="species" species="munglonganus">
|
|
<emphasis id="84E8EAAAFFE1FFCF05217631FD461715" box="[571,737,1623,1643]" italics="true" pageId="11" pageNumber="12">A. munglonganus</emphasis>
|
|
</taxonomicName>
|
|
</paragraph>
|
|
<paragraph id="B62336B8FFE1FFCF07927617FD4617DE" blockId="11.[136,738,1276,1696]" pageId="11" pageNumber="12">
|
|
- Genital markings present in both spermathecal and male pore regions ...........................................
|
|
<taxonomicName id="719C4D3BFFE1FFCF052C76EAFD4617DE" authorityName="Grzelak & Sørensen" authorityYear="2022" box="[566,737,1676,1696]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Crassiclitellata" pageId="11" pageNumber="12" phylum="Annelida" rank="species" species="munglongoides">
|
|
<emphasis id="84E8EAAAFFE1FFCF052C76EAFD4617DE" box="[566,737,1676,1696]" italics="true" pageId="11" pageNumber="12">A. munglongoides</emphasis>
|
|
</taxonomicName>
|
|
</paragraph>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |