treatments-xml/data/3E/10/E1/3E10E143A014D173FF16FEB0FDA9F8C6.xml

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<mods:title id="93059A746F7B8A0A9D23D3DC60BEB40D">Redescription of Alpheus vladivostokiensis (Vinogradov, 1950), a large and conspicuous snapping shrimp from the northern Sea of Japan (Decapoda: Caridea: Alpheidae)</mods:title>
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<mods:namePart id="43A4653D0AEA5E34DEE372D8CDA4849C">Anker, Arthur</mods:namePart>
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<mods:namePart id="821ACAFA4894CFC780D6205308D0D924">Marin, Ivan N.</mods:namePart>
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<paragraph id="B6065055A014D17AFF16FEB0FCCBFE04" blockId="3.[151,848,359,419]" box="[151,848,359,386]" pageId="3" pageNumber="174">
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<emphasis id="84CD8C47A014D17AFF16FEB0FCCBFE04" bold="true" box="[151,848,359,386]" pageId="3" pageNumber="174">
<taxonomicName id="71B92BD6A014D17AFF16FEB0FD59FE04" ID-CoL="85LQX" authority="Vinogradov, 1950" authorityName="Vinogradov" authorityYear="1950" box="[151,706,359,386]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="174" phylum="Arthropoda" rank="species" species="vladivostokiensis" status="comb. nov.">
<emphasis id="84CD8C47A014D17AFF16FEB0FE56FE07" bold="true" box="[151,461,359,385]" italics="true" pageId="3" pageNumber="174">Alpheus vladivostokiensis</emphasis>
(
<bibRefCitation id="D2282DA4A014D17AFE5DFEBFFD21FE04" author="Vinogradov" box="[476,698,360,386]" pageId="3" pageNumber="174" refString="Vinogradov, L. G. (1950) Keys to the shrimps, crayfishes and crabs of the Far East [Opredelitel' krevetok, rakov i krabov Dal'nego Vostoka]. Izvestia TINRO, 33 C, 179 - 350 [in Russian]" type="journal article" year="1950">Vinogradov, 1950</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="9FFE313CA014D17AFD4BFEBFFCCBFE04" box="[714,848,360,386]" pageId="3" pageNumber="174">comb. nov.</taxonomicNameLabel>
</emphasis>
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<paragraph id="B6065055A014D17AFF16FE5DFE90FE25" blockId="3.[151,848,359,419]" box="[151,267,394,419]" pageId="3" pageNumber="174">
(
<figureCitation id="2E824CD0A014D17AFF1EFE5DFE99FE25" box="[159,258,394,419]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStartId-0="5.[151,250,1858,1880]" captionStartId-1="6.[151,250,1709,1731]" captionStartId-2="7.[151,250,1413,1435]" captionStartId-3="8.[151,250,1581,1603]" captionTargetBox-0="[153,1429,200,1831]" captionTargetBox-1="[157,1419,203,1681]" captionTargetBox-2="[151,1435,176,1380]" captionTargetBox-3="[154,1434,176,1579]" captionTargetId-0="figure@5.[153,1438,196,1840]" captionTargetId-1="figure@6.[151,1436,193,1687]" captionTargetId-2="figure@7.[151,1436,154,1397]" captionTargetId-3="figure@8.[154,1434,164,1584]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionTargetPageId-2="7" captionTargetPageId-3="8" captionText-0="FIGURE 1. Alpheus vladivostokiensis (Vinogradov, 1950): male (cl 18.0 mm) from Troitza Bay (= Bukhta Troitzy), Russian Far East (OUMNH. ZC. 2016 - 01 - 013): A, frontal region, dorsal; B, same, lateral; C, telson, dorsal; D, ventromesial carina of first article of antennular peduncle, lateral; E, third maxilliped, lateral; F, second pereiopod, lateral; G, third pereiopod, lateral; H, same, dactylus, dorsal; I, fifth pereiopod, lateral; J, same, propodus and dactylus, ventral; K, second pleopod, detail of appendices masculina and interna, dorsal; L uropod, dorsal." captionText-1="FIGURE 2. Alpheus vladivostokiensis (Vinogradov, 1950): male (cl 18.0 mm) from Troitza Bay (= Bukhta Troitzy), Russian Far East (OUMNH. ZC. 2016 - 01 - 013) [A – G]; ovigerous female (cl 17.4 mm) from the same locality (MNHN-IU- 2014 - 5694) [H, I]: A, major cheliped, lateral; B, major chela, mesial; C, same, fingers open, lateral; D, same, merus, mesial; E, minor cheliped, lateral; F, minor chela, mesial; G, same, merus, mesial; H, minor chela, mesial; I, minor cheliped, merus, mesial." captionText-2="FIGURE 3. Alpheus vladivostokiensis (Vinogradov, 1950), colour pattern: neotype, male (cl 19.7 mm) from Troitza Bay (= Bukhta Troitzy), Russian Far East (MNHN-IU- 2014 - 5693), in dorsal (A) and lateral (B) views. Photographs by Ivan N. Marin." captionText-3="FIGURE 4. Alpheus vladivostokiensis (Vinogradov, 1950), colour pattern: ovigerous female (cl 16.3 mm) from Troitza Bay (= Bukhta Troitzy), Russian Far East (OUMNH. ZC. 2016 - 01 - 014), in dorsal (A) and lateral (B) views. Photographs by Ivan N. Marin." httpUri-0="https://zenodo.org/record/271759/files/figure.png" httpUri-1="https://zenodo.org/record/271760/files/figure.png" httpUri-2="https://zenodo.org/record/271761/files/figure.png" httpUri-3="https://zenodo.org/record/271762/files/figure.png" pageId="3" pageNumber="174">Figs 1–4</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="FEA303DEA014D17AFF16FE06FCCCFDC8" pageId="3" pageNumber="174" type="reference_group">
<paragraph id="B6065055A014D17AFF16FE06FC89FD8E" blockId="3.[151,1437,464,590]" pageId="3" pageNumber="174">
<treatmentCitationGroup id="96A9777BA014D17AFF16FE06FC89FD8E" pageId="3" pageNumber="174">
<taxonomicName id="71B92BD6A014D17AFF16FE06FD1DFE60" ID-CoL="85MYY" authority="Vinogradov 1950: 198" authorityName="Vinogradov" authorityPageNumber="198" authorityYear="1950" box="[151,646,464,486]" class="Malacostraca" family="Alpheidae" genus="Betaeus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="174" phylum="Arthropoda" rank="species" species="vladivostokiensis">
<emphasis id="84CD8C47A014D17AFF16FE06FE01FE60" box="[151,410,464,486]" italics="true" pageId="3" pageNumber="174">Betaeus vladivostokiensis</emphasis>
<treatmentCitation id="37187644A014D17AFE20FE07FD1DFE60" author="Vinogradov" box="[417,646,464,486]" page="198" pageId="3" pageNumber="174" year="1950">
<bibRefCitation id="D2282DA4A014D17AFE20FE07FDCEFE60" author="Vinogradov" box="[417,597,464,486]" pageId="3" pageNumber="174" refString="Vinogradov, L. G. (1950) Keys to the shrimps, crayfishes and crabs of the Far East [Opredelitel' krevetok, rakov i krabov Dal'nego Vostoka]. Izvestia TINRO, 33 C, 179 - 350 [in Russian]" type="journal article" year="1950">Vinogradov 1950</bibRefCitation>
: 198
</treatmentCitation>
</taxonomicName>
, pl. 10, fig. 31; 
<treatmentCitation id="37187644A014D17AFCA6FE06FBFFFE60" author="De" box="[807,1124,464,486]" page="475" pageId="3" pageNumber="174" year="2011">
<bibRefCitation id="D2282DA4A014D17AFCA6FE06FBAFFE60" author="De" box="[807,1076,464,486]" pageId="3" pageNumber="174" refString="De Grave, S. &amp; Fransen, C. H. J. M. (2011) Carideorum catalogus: the recent species of the dendrobranchiate, stenopodidean, procarididean and caridean shrimps (Crustacea: Decapoda). Zoologische Mededelingen, Leiden, 85, 195 - 589." type="journal article" year="2011">De Grave &amp; Fransen 2011</bibRefCitation>
: 475
</treatmentCitation>
(listed as 
<emphasis id="84CD8C47A014D17AFB4DFE06FAFAFE60" box="[1228,1377,464,486]" italics="true" pageId="3" pageNumber="174">nomen dubium</emphasis>
). 
<taxonomicName id="71B92BD6A014D17AFF16FE25FE07FD8E" ID-CoL="85LQX" box="[151,412,498,520]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="174" phylum="Arthropoda" rank="species" species="vladivostokiensis">
<emphasis id="84CD8C47A014D17AFF16FE25FE07FD8E" box="[151,412,498,520]" italics="true" pageId="3" pageNumber="174">Alpheus vladivostokiensis</emphasis>
</taxonomicName>
— 
<treatmentCitation id="37187644A014D17AFE34FE25FD1DFD8E" author="Pasternak" box="[437,646,498,520]" page="304" pageId="3" pageNumber="174" year="1988">
<bibRefCitation id="D2282DA4A014D17AFE34FE25FDCFFD8E" author="Pasternak" box="[437,596,498,520]" pageId="3" pageNumber="174" refString="Pasternak, R. K. (1988) Life of animals [Zhizn' zhivotnykh], Vol. 2. Second edition. Prosvezscheniye, Moscow, 447 pp. [in Russian]" type="book" year="1988">Pasternak 1988</bibRefCitation>
: 304
</treatmentCitation>
, pl. 55, fig. 1.
</treatmentCitationGroup>
</paragraph>
<paragraph id="B6065055A014D17AFF16FDC2FCCCFDC8" blockId="3.[151,1437,464,590]" pageId="3" pageNumber="174">
<treatmentCitationGroup id="96A9777BA014D17AFF16FDC2FCCCFDC8" pageId="3" pageNumber="174">
<taxonomicName id="71B92BD6A014D17AFF16FDC2FEC9FDAD" ID-CoL="84W83" box="[151,338,533,555]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="174" phylum="Arthropoda" rank="species" species="japonicus">
<emphasis id="84CD8C47A014D17AFF16FDC2FEC9FDAD" box="[151,338,533,555]" italics="true" pageId="3" pageNumber="174">Alpheus japonicus</emphasis>
</taxonomicName>
— 
<treatmentCitation id="37187644A014D17AFEECFDC2FD9EFDAD" author="Balss" box="[365,517,533,555]" page="40" pageId="3" pageNumber="174" year="1914">
<bibRefCitation id="D2282DA4A014D17AFEECFDC2FE7AFDAD" author="Balss" box="[365,481,533,555]" pageId="3" pageNumber="174" refString="Balss, H. (1914) Ostasiatischen Decapoden II. Die Natantia und Reptantia. Beitrage zur Naturgeschichte Ostasiens. Abhandlungen der Mathematisch-Physischen Klasse der Kayserlichen Bayerischen Akademie der Wissenschaften, II, Suppl. - Bd. 10, Abh., 1 - 101, 1 pl." type="book chapter" year="1914">Balss 1914</bibRefCitation>
: 40
</treatmentCitation>
(
<emphasis id="84CD8C47A014D17AFD94FDC1FDC3FDAD" box="[533,600,534,555]" italics="true" pageId="3" pageNumber="174">partim</emphasis>
); 
<treatmentCitation id="37187644A014D17AFDF1FDC2FC4AFDAD" author="Derjugin" box="[624,977,533,555]" page="142" pageId="3" pageNumber="174" year="1935">
<bibRefCitation id="D2282DA4A014D17AFDF1FDC2FC3BFDAD" author="Derjugin" box="[624,928,533,555]" pageId="3" pageNumber="174" refString="Derjugin, K. M. &amp; Kobjakowa, S. (1935) Zur Dekapodenfauna des Japanischen Meeres. Zoologischer Anzeiger, 112, 141 - 147." type="journal article" year="1935">Derjugin &amp; Kobjakowa 1935</bibRefCitation>
: 142
</treatmentCitation>
(
<emphasis id="84CD8C47A014D17AFC63FDC1FBBDFDAD" box="[994,1062,534,555]" italics="true" pageId="3" pageNumber="174">partim</emphasis>
?); 
<treatmentCitation id="37187644A014D17AFBC9FDC2FAAAFDAD" author="Vinogradov" box="[1096,1329,533,555]" page="197" pageId="3" pageNumber="174" year="1950">
<bibRefCitation id="D2282DA4A014D17AFBC9FDC2FB65FDAD" author="Vinogradov" box="[1096,1278,533,555]" pageId="3" pageNumber="174" refString="Vinogradov, L. G. (1950) Keys to the shrimps, crayfishes and crabs of the Far East [Opredelitel' krevetok, rakov i krabov Dal'nego Vostoka]. Izvestia TINRO, 33 C, 179 - 350 [in Russian]" type="journal article" year="1950">Vinogradov 1950</bibRefCitation>
: 197
</treatmentCitation>
, pl. 9, fig. 28; 
<treatmentCitation id="37187644A014D17AFF6CFDEFFE15FDC8" author="Marin" box="[237,398,568,590]" page="59" pageId="3" pageNumber="174" year="2013">
<bibRefCitation id="D2282DA4A014D17AFF6CFDEFFEF3FDC8" author="Marin" box="[237,360,568,590]" pageId="3" pageNumber="174" refString="Marin, I. N. (2013) Small atlas of decapod crustaceans of Russia [Malyi atlas desiatinogikh rakoobraznykh Rossii]. KMK Scientific Press, Moscow, 146 pp. [in Russian]" type="book" year="2013">Marin 2013</bibRefCitation>
: 59
</treatmentCitation>
, pl. 23, fig. 1 [not 
<taxonomicName id="71B92BD6A014D17AFDC8FDEEFCD6FDC8" ID-CoL="84W83" authority="Miers, 1879" authorityName="Miers" authorityYear="1879" box="[585,845,568,590]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="174" phylum="Arthropoda" rank="species" species="japonicus">
<emphasis id="84CD8C47A014D17AFDC8FDEEFD53FDC8" box="[585,712,569,590]" italics="true" pageId="3" pageNumber="174">A. japonicus</emphasis>
<bibRefCitation id="D2282DA4A014D17AFD4FFDEFFCD6FDC8" author="Miers" box="[718,845,568,590]" pageId="3" pageNumber="174" refString="Miers, E. J. (1879) On a collection of Crustacea made by Capt. H. C. St. John R. N. in the Corean and Japanese Seas. Part I." type="book" year="1879">Miers, 1879</bibRefCitation>
</taxonomicName>
].
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</paragraph>
</subSubSection>
<subSubSection id="FEA303DEA014D17AFF16FDAAFAF3F920" pageId="3" pageNumber="174" type="materials_examined">
<paragraph id="B6065055A014D17AFF16FDAAFB4EFC48" blockId="3.[151,1437,637,2014]" pageId="3" pageNumber="174">
<emphasis id="84CD8C47A014D17AFF16FDAAFEE7FD10" bold="true" box="[151,380,637,662]" pageId="3" pageNumber="174">Material examined</emphasis>
. 
<collectingCountry id="CEAE10C5A014D17AFE0CFDAAFE43FD10" box="[397,472,637,662]" name="Russia" pageId="3" pageNumber="174">Russia</collectingCountry>
: 
<typeStatus id="6902EEF7A014D17AFE6BFDA9FDDDFD10" box="[490,582,638,662]" pageId="3" pageNumber="174" type="neotype">neotype</typeStatus>
, male (cl 
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), MNHN-IU-2014-5693, Russian Far East, Primorskiy Krai, Troitza Bay (= Bukhta Troitzy), Andreevka, 
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, off jetty, depth: 
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, under rocks on mud-sand, coll. I.N. Marin, 
<date id="C2077695A014D17AFE78FD14FDF5FD5D" box="[505,622,707,731]" pageId="3" pageNumber="174" value="2011-06">June 2011</date>
; 
<specimenCount id="A0BF9BDCA014D17AFDFAFD14FD5CFD5D" box="[635,711,706,731]" pageId="3" pageNumber="174" type="male">1 male</specimenCount>
(cl 18.0 mm), 
<collectionCode id="D0A8C890A014D17AFCF3FD14FC79FD5D" box="[882,994,707,731]" pageId="3" pageNumber="174">OUMNH</collectionCode>
.ZC. 2016-01-013, same collection data as for 
<typeStatus id="6902EEF7A014D17AFF63FD31FEA5FD78" box="[226,318,742,766]" pageId="3" pageNumber="174" type="neotype">neotype</typeStatus>
; 1 ovigerous female (cl 
<quantity id="7141FDB0A014D17AFDD6FD31FD59FD78" box="[599,706,742,766]" metricMagnitude="-2" metricUnit="m" metricValue="1.7399999999999998" pageId="3" pageNumber="174" unit="mm" value="17.4">17.4 mm</quantity>
), MNHN-IU-2014-5694, same collection data as for 
<typeStatus id="6902EEF7A014D17AFAA5FD31FA1BFD78" box="[1316,1408,742,766]" pageId="3" pageNumber="174" type="neotype">neotype</typeStatus>
; 1 ovigerous female (cl 
<quantity id="7141FDB0A014D17AFE06FCD0FE6BFCA6" box="[391,496,775,800]" metricMagnitude="-2" metricUnit="m" metricValue="1.63" pageId="3" pageNumber="174" unit="mm" value="16.3">16.3 mm</quantity>
), 
<collectionCode id="D0A8C890A014D17AFD85FCDFFDEFFCA6" box="[516,628,776,800]" pageId="3" pageNumber="174">OUMNH</collectionCode>
.ZC. 2016-01-014, same collection data as for 
<typeStatus id="6902EEF7A014D17AFBFFFCDFFB40FCA6" box="[1150,1243,776,800]" pageId="3" pageNumber="174" type="neotype">neotype</typeStatus>
; 
<specimenCount id="A0BF9BDCA014D17AFB67FCDFFAA6FCA6" box="[1254,1341,775,800]" pageId="3" pageNumber="174" type="male">3 males</specimenCount>
(cl 9.5– 
<quantity id="7141FDB0A014D17AFF16FCFDFF62FCC5" box="[151,249,810,835]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" pageId="3" pageNumber="174" unit="mm" value="7.5">7.5 mm</quantity>
), 
<collectionCode id="D0A8C890A014D17AFE92FCFCFEEBFCC5" box="[275,368,811,835]" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="3" pageNumber="174">MZUSP</collectionCode>
34289, same collection data as for 
<typeStatus id="6902EEF7A014D17AFCB5FCFCFC0BFCC5" box="[820,912,811,835]" pageId="3" pageNumber="174" type="neotype">neotype</typeStatus>
; 
<specimenCount id="A0BF9BDCA014D17AFC22FCFCFB97FCC5" box="[931,1036,810,835]" pageId="3" pageNumber="174" type="female">1 female</specimenCount>
(cl 
<quantity id="7141FDB0A014D17AFBC6FCFCFB2CFCC5" box="[1095,1207,811,835]" metricMagnitude="-2" metricUnit="m" metricValue="1.21" pageId="3" pageNumber="174" unit="mm" value="12.1">12.1 mm</quantity>
), 
<collectionCode id="D0A8C890A014D17AFB50FCFCFAB3FCC4" box="[1233,1320,811,834]" httpUri="http://grbio.org/cool/k3qp-615s" name="Zoological Museum, Moscow Lomonosov State University" pageId="3" pageNumber="174">ZMMU</collectionCode>
Ma 608, Vladivostok, no other data, det. H. Balss as 
<taxonomicName id="71B92BD6A014D17AFD0EFC99FC86FCE3" box="[655,797,846,869]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="174" phylum="Arthropoda" rank="species" species="japonicus">
<emphasis id="84CD8C47A014D17AFD0EFC99FC86FCE3" box="[655,797,846,869]" italics="true" pageId="3" pageNumber="174">A. japonicus</emphasis>
</taxonomicName>
; 
<specimenCount id="A0BF9BDCA014D17AFCAFFC99FCE6FCE0" box="[814,893,845,870]" pageId="3" pageNumber="174" type="male">1 male</specimenCount>
, 
<specimenCount id="A0BF9BDCA014D17AFC0AFC99FC63FCE0" box="[907,1016,845,870]" pageId="3" pageNumber="174" type="female">2 females</specimenCount>
(cl not measured, specimens in very bad condition), 
<collectionCode id="D0A8C890A014D17AFECCFCA7FE3FFC01" box="[333,420,880,903]" httpUri="http://grbio.org/cool/k3qp-615s" name="Zoological Museum, Moscow Lomonosov State University" pageId="3" pageNumber="174">ZMMU</collectionCode>
Ma 1837, Krupenin, no other data, det. as 
<taxonomicName id="71B92BD6A014D17AFC1FFCA7FBB6FC01" box="[926,1069,880,903]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="174" phylum="Arthropoda" rank="species" species="japonicus">
<emphasis id="84CD8C47A014D17AFC1FFCA7FBB6FC01" box="[926,1069,880,903]" italics="true" pageId="3" pageNumber="174">A. japonicus</emphasis>
</taxonomicName>
. 
<collectingCountry id="CEAE10C5A014D17AFBBFFCA7FBE5FC01" box="[1086,1150,880,903]" name="Japan" pageId="3" pageNumber="174">Japan</collectingCountry>
: 
<specimenCount id="A0BF9BDCA014D17AFB11FCA7FB6FFC0E" box="[1168,1268,879,904]" pageId="3" pageNumber="174" type="female">1 female</specimenCount>
(cl 
<quantity id="7141FDB0A014D17AFAA4FCA7FA0BFC0E" box="[1317,1424,880,904]" metricMagnitude="-2" metricUnit="m" metricValue="1.31" pageId="3" pageNumber="174" unit="mm" value="13.1">13.1 mm</quantity>
), CBM-ZC 13263, southern Hokkaido, Hakodate Bay, Hokuto City, Kattoshi, 
<geoCoordinate id="D38D3692A014D17AFBC1FC45FB2EFC2D" box="[1088,1205,914,939]" direction="north" orientation="latitude" pageId="3" pageNumber="174" precision="92" value="41.743332">41°44.6’N</geoCoordinate>
<geoCoordinate id="D38D3692A014D17AFB44FC45FAD3FC2D" box="[1221,1352,914,939]" direction="east" orientation="longitude" pageId="3" pageNumber="174" precision="92" value="140.60333">140°36.2’E</geoCoordinate>
, lower intertidal (exposed at low tide), under rocks on sand and gravel, coll. S. Goshima, 
<date id="C2077695A014D17AFBB0FC61FB4AFC48" box="[1073,1233,949,974]" pageId="3" pageNumber="174" value="2015-04-22">22 April 2015</date>
.
</paragraph>
<paragraph id="B6065055A014D17AFF46FC00FAF3F920" blockId="3.[151,1437,637,2014]" pageId="3" pageNumber="174">
<emphasis id="84CD8C47A014D17AFF46FC00FE49FC76" bold="true" box="[199,466,983,1008]" pageId="3" pageNumber="174">Comparative material</emphasis>
. 
<taxonomicName id="71B92BD6A014D17AFE62FC0FFCD2FC76" authority="Miers, 1879" authorityName="Miers" authorityYear="1879" box="[483,841,983,1008]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="174" phylum="Arthropoda" rank="species" species="japonicus">
<emphasis id="84CD8C47A014D17AFE62FC0FFD28FC69" box="[483,691,984,1008]" italics="true" pageId="3" pageNumber="174">Alpheus japonicus</emphasis>
<bibRefCitation id="D2282DA4A014D17AFD3DFC00FCD2FC76" author="Miers" box="[700,841,983,1008]" pageId="3" pageNumber="174" refString="Miers, E. J. (1879) On a collection of Crustacea made by Capt. H. C. St. John R. N. in the Corean and Japanese Seas. Part I." type="book" year="1879">Miers, 1879</bibRefCitation>
</taxonomicName>
. 
<collectingCountry id="CEAE10C5A014D17AFCD7FC0FFC0DFC69" box="[854,918,984,1007]" name="Japan" pageId="3" pageNumber="174">Japan</collectingCountry>
: 
<specimenCount id="A0BF9BDCA014D17AFC26FC0FFB91FC76" box="[935,1034,983,1008]" pageId="3" pageNumber="174" type="female">1 female</specimenCount>
(cl 
<quantity id="7141FDB0A014D17AFBBBFC0FFB3EFC76" box="[1082,1189,984,1008]" metricMagnitude="-2" metricUnit="m" metricValue="1.08" pageId="3" pageNumber="174" unit="mm" value="10.8">10.8 mm</quantity>
), HUMZ-C 24, Iwate Prefecture, Miyako Bay, depth: 
<quantity id="7141FDB0A014D17AFE7CFC2DFDF9FB94" box="[509,610,1018,1043]" metricMagnitude="1" metricUnit="m" metricValue="1.25" metricValueMax="1.5" metricValueMin="1.0" pageId="3" pageNumber="174" unit="m" value="12.5" valueMax="15.0" valueMin="10.0">10–15 m</quantity>
, commercial round haul net, coll. T. Komai, 
<date id="C2077695A014D17AFBDBFC2DFB6DFB95" box="[1114,1270,1018,1043]" pageId="3" pageNumber="174" value="1982-10">October 1982</date>
; 
<specimenCount id="A0BF9BDCA014D17AFA83FC2CFAD6FB95" box="[1282,1357,1018,1043]" pageId="3" pageNumber="174" type="male">1 male</specimenCount>
(cl 
<quantity id="7141FDB0A014D17AFAF8FC2DFF5CFBB0" metricMagnitude="-3" metricUnit="m" metricValue="6.6" pageId="3" pageNumber="174" unit="mm" value="6.6">6.6 mm</quantity>
), 
<specimenCount id="A0BF9BDCA014D17AFF5AFBC9FEA6FBB0" box="[219,317,1053,1078]" pageId="3" pageNumber="174" type="female">1 female</specimenCount>
(cl 
<quantity id="7141FDB0A014D17AFEEAFBCAFE5DFBB0" box="[363,454,1053,1078]" metricMagnitude="-3" metricUnit="m" metricValue="5.1" pageId="3" pageNumber="174" unit="mm" value="5.1">5.1 mm</quantity>
), Funakoshi Bay, Iwate Prefecture, Yamada, 
<geoCoordinate id="D38D3692A014D17AFC45FBC9FBA2FBB0" box="[964,1081,1054,1078]" direction="north" orientation="latitude" pageId="3" pageNumber="174" precision="92" value="39.385">39°23.1’N</geoCoordinate>
<geoCoordinate id="D38D3692A014D17AFBC3FBCAFB5EFBB0" box="[1090,1221,1053,1078]" direction="east" orientation="longitude" pageId="3" pageNumber="174" precision="92" value="141.975">141°58.5’E</geoCoordinate>
, depth: 
<quantity id="7141FDB0A014D17AFAA1FBCAFAC7FBB3" box="[1312,1372,1053,1078]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="3" pageNumber="174" unit="m" value="50.0">50 m</quantity>
, sand with shell fragments, dredge, coll. T. Komai, 
<date id="C2077695A014D17AFD24FBE8FCD8FBDE" box="[677,835,1087,1112]" pageId="3" pageNumber="174" value="1995-05-25">25 May 1995</date>
; 
<specimenCount id="A0BF9BDCA014D17AFCD0FB97FC2EFBDE" box="[849,949,1087,1112]" pageId="3" pageNumber="174" type="female">1 female</specimenCount>
(cl 
<quantity id="7141FDB0A014D17AFC67FB97FBD9FBDE" box="[998,1090,1088,1112]" metricMagnitude="-3" metricUnit="m" metricValue="9.4" pageId="3" pageNumber="174" unit="mm" value="9.4">9.4 mm</quantity>
), 
<collectionCode id="D0A8C890A014D17AFBD9FB97FB09FBDE" box="[1112,1170,1088,1112]" httpUri="http://grbio.org/cool/6hz5-fzvb" name="Universite de Strasbourg, Musee de Zoologie" pageId="3" pageNumber="174">MZS</collectionCode>
70, Tanagawa, coll. L. Döderlein, 1881 [material “b” of 
<bibRefCitation id="D2282DA4A014D17AFD92FBB4FD57FBFD" author="Ortmann" box="[531,716,1123,1147]" pageId="3" pageNumber="174" refString="Ortmann, A. (1890) Die Unterordnung Natantia Boas. Theil I. Die Decapoden Krebse des Strassburger Museums, mit besonderer Berucksichtigung der von Herrn Dr. Doderlein bei Japan und bei den Liu-Kiu-Inseln gesammelten und z. Z. im Strassburger Museum aufbewahrten Formen. Zoologische Jahrbucher, Systematik, 5, 437 - 542, pls. 36, 37." type="journal article" year="1890">Ortmann (1890)</bibRefCitation>
, re-examined by 
<bibRefCitation id="D2282DA4A014D17AFC13FBB5FBACFBFD" author="Komai" box="[914,1079,1122,1147]" pageId="3" pageNumber="174" refString="Komai, T. (1999) Decapod Crustacea collected by L. Doderlein in Japan and reported by Ortmann (1890 - 1894) in the collection of the Musee Zoologique, Strasbourg. In: Nishikawa, T. (Ed.), Preliminary Taxonomic and Historical Studies on Prof. Ludwig Doderlein's collection of Japanese Animals made in 1880 - 81 and deposited at Several European Museum. Report on Activities in 1997 - 8 supported by Grant-in-Aid for International Scientific Research (Field Research) No. 0 9041155, pp. 53 - 101." type="journal article" year="1999">Komai (1999)</bibRefCitation>
]; 
<specimenCount id="A0BF9BDCA014D17AFBCAFBB5FB38FBFD" box="[1099,1187,1122,1147]" pageId="3" pageNumber="174" type="male">5 males</specimenCount>
(cl 16.0–19.0 mm), 
<specimenCount id="A0BF9BDCA014D17AFA0FFBB4FF75FB18" pageId="3" pageNumber="174" type="female">3 females</specimenCount>
(cl 16.5–18.0 mm), 
<collectionCode id="D0A8C890A014D17AFE60FB51FD80FB18" box="[481,539,1158,1182]" httpUri="http://grbio.org/cool/6hz5-fzvb" name="Universite de Strasbourg, Musee de Zoologie" pageId="3" pageNumber="174">MZS</collectionCode>
73, Tokyo Bay, 
<date id="C2077695A014D17AFD60FB52FCC7FB18" box="[737,860,1157,1182]" pageId="3" pageNumber="174" value="1880" valueMax="1881">1880–1881</date>
[material “a” of 
<bibRefCitation id="D2282DA4A014D17AFBA1FB51FB41FB18" author="Ortmann" box="[1056,1242,1158,1182]" pageId="3" pageNumber="174" refString="Ortmann, A. (1890) Die Unterordnung Natantia Boas. Theil I. Die Decapoden Krebse des Strassburger Museums, mit besonderer Berucksichtigung der von Herrn Dr. Doderlein bei Japan und bei den Liu-Kiu-Inseln gesammelten und z. Z. im Strassburger Museum aufbewahrten Formen. Zoologische Jahrbucher, Systematik, 5, 437 - 542, pls. 36, 37." type="journal article" year="1890">Ortmann (1890)</bibRefCitation>
, re-examined by 
<bibRefCitation id="D2282DA4A014D17AFF16FB70FEDAFB46" author="Komai" box="[151,321,1191,1216]" pageId="3" pageNumber="174" refString="Komai, T. (1999) Decapod Crustacea collected by L. Doderlein in Japan and reported by Ortmann (1890 - 1894) in the collection of the Musee Zoologique, Strasbourg. In: Nishikawa, T. (Ed.), Preliminary Taxonomic and Historical Studies on Prof. Ludwig Doderlein's collection of Japanese Animals made in 1880 - 81 and deposited at Several European Museum. Report on Activities in 1997 - 8 supported by Grant-in-Aid for International Scientific Research (Field Research) No. 0 9041155, pp. 53 - 101." type="journal article" year="1999">Komai (1999)</bibRefCitation>
]; 
<specimenCount id="A0BF9BDCA014D17AFEDBFB7FFE59FB46" box="[346,450,1191,1216]" pageId="3" pageNumber="174" type="female">1 female</specimenCount>
(cl 
<quantity id="7141FDB0A014D17AFE7DFB70FDC7FB46" box="[508,604,1191,1216]" metricMagnitude="-3" metricUnit="m" metricValue="9.6" pageId="3" pageNumber="174" unit="mm" value="9.6">9.6 mm</quantity>
), CBM-ZC 5044, Tokyo Bay, off Yokoyama, depth not recorded, 
<date id="C2077695A014D17AFA01FB7FFEC8FB65" pageId="3" pageNumber="174" value="1996-11-27">27 November 1996</date>
; 
<specimenCount id="A0BF9BDCA014D17AFEE0FB1CFE21FB65" box="[353,442,1226,1251]" pageId="3" pageNumber="174" type="male">2 males</specimenCount>
(cl 8.3, 13.0 mm), 1 ovigerous female (cl 
<quantity id="7141FDB0A014D17AFC30FB1CFB87FB65" box="[945,1052,1227,1251]" metricMagnitude="-2" metricUnit="m" metricValue="1.22" pageId="3" pageNumber="174" unit="mm" value="12.2">12.2 mm</quantity>
), CBM-ZC 10, Chiba Prefecture, Tokyo Bay, Banzu, Kisarazu, depth not recorded, scuba diving, coll. T. Furota, 
<date id="C2077695A014D17AFB9DFB39FB2CFA80" box="[1052,1207,1262,1286]" pageId="3" pageNumber="174" value="1984-05-10">10 May 1984</date>
; 1 ovigerous female (cl 
<quantity id="7141FDB0A014D17AFF43FAC7FEABFAAE" box="[194,304,1296,1320]" metricMagnitude="-2" metricUnit="m" metricValue="1.03" pageId="3" pageNumber="174" unit="mm" value="10.3">10.3 mm</quantity>
), CBM-ZC 16, Tokyo Bay, same locality, depth not recorded, scuba diving, coll. T. Furota, 
<date id="C2077695A014D17AFA0FFAC7FEC8FACD" pageId="3" pageNumber="174" value="1989-09-08">8 September 1989</date>
; 
<specimenCount id="A0BF9BDCA014D17AFEE0FAE4FE35FACD" box="[353,430,1330,1355]" pageId="3" pageNumber="174" type="male">1 male</specimenCount>
(cl 
<quantity id="7141FDB0A014D17AFE5FFAE4FDD3FACD" box="[478,584,1331,1355]" metricMagnitude="-2" metricUnit="m" metricValue="1.39" pageId="3" pageNumber="174" unit="mm" value="13.9">13.9 mm</quantity>
), 
<collectionCode id="D0A8C890A014D17AFDDCFAE4FD56FACD" box="[605,717,1331,1355]" pageId="3" pageNumber="174">OUMNH</collectionCode>
.ZC. 2016-01-015, Tokyo Bay, sta. 2, depth: 
<quantity id="7141FDB0A014D17AFB4AFAE4FA93FACC" box="[1227,1288,1331,1355]" metricMagnitude="1" metricUnit="m" metricValue="3.0" pageId="3" pageNumber="174" unit="m" value="30.0">30 m</quantity>
, beam trawl, coll. W. Doi, 
<date id="C2077695A014D17AFEB4FA81FE47FAE8" box="[309,476,1365,1390]" pageId="3" pageNumber="174" value="2006-04-18">18 April 2006</date>
; 
<specimenCount id="A0BF9BDCA014D17AFE6BFA81FDA3FAE8" box="[490,568,1365,1390]" pageId="3" pageNumber="174" type="male">1 male</specimenCount>
(cl 
<quantity id="7141FDB0A014D17AFDEBFA81FD4EFAE8" box="[618,725,1366,1390]" metricMagnitude="-2" metricUnit="m" metricValue="1.43" pageId="3" pageNumber="174" unit="mm" value="14.3">14.3 mm</quantity>
), MNHN-IU-2014-5695, same collection data as for previous specimen; 
<specimenCount id="A0BF9BDCA014D17AFE91FAAFFEE9FA16" box="[272,370,1399,1424]" pageId="3" pageNumber="174" type="female">1 female</specimenCount>
(cl 12.0 mm), MNHN-IU-2014-5696, same collection data as for previous specimen; 
<specimenCount id="A0BF9BDCA014D17AFAC4FAAFFA07FA16" box="[1349,1436,1399,1424]" pageId="3" pageNumber="174" type="male">3 males</specimenCount>
(cl 14.1– 
<quantity id="7141FDB0A014D17AFF7AFA4CFEF8FA35" box="[251,355,1435,1459]" metricMagnitude="-2" metricUnit="m" metricValue="1.13" pageId="3" pageNumber="174" unit="mm" value="11.3">11.3 mm</quantity>
), 
<collectionCode id="D0A8C890A014D17AFEF7FA4CFE48FA35" box="[374,467,1435,1459]" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="3" pageNumber="174">MZUSP</collectionCode>
34290, same collection data as for previous specimen; 
<specimenCount id="A0BF9BDCA014D17AFBC4FA4CFB0AFA35" box="[1093,1169,1434,1459]" pageId="3" pageNumber="174" type="male">1 male</specimenCount>
(cl 
<quantity id="7141FDB0A014D17AFB3FFA4DFABCFA35" box="[1214,1319,1434,1459]" metricMagnitude="-2" metricUnit="m" metricValue="1.06" pageId="3" pageNumber="174" unit="mm" value="10.6">10.6 mm</quantity>
), 
<specimenCount id="A0BF9BDCA014D17AFABAFA4CFA07FA35" box="[1339,1436,1434,1459]" pageId="3" pageNumber="174" type="female">1 female</specimenCount>
(cl 
<quantity id="7141FDB0A014D17AFF41FA6AFEB6FA50" box="[192,301,1469,1494]" metricMagnitude="-2" metricUnit="m" metricValue="1.26" pageId="3" pageNumber="174" unit="mm" value="12.6">12.6 mm</quantity>
), CBM-ZC 8109, Seto Inland Sea, off Kurahashi Island, Hiroshima Prefecture, depth: 
<quantity id="7141FDB0A014D17AFAAFFA69FA0CFA53" box="[1326,1431,1470,1494]" metricMagnitude="1" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="3" pageNumber="174" unit="m" value="15.0" valueMax="20.0" valueMin="10.0">10–20 m</quantity>
, commercial trawler, coll. M. Fujiwara, 
<date id="C2077695A014D17AFDD9FA37FC96FA7E" box="[600,781,1503,1528]" pageId="3" pageNumber="174" value="2004-10-09">9 October 2004</date>
; 1 ovigerous female (cl 
<quantity id="7141FDB0A014D17AFBA4FA37FB14FA7E" box="[1061,1167,1504,1528]" metricMagnitude="-2" metricUnit="m" metricValue="1.73" pageId="3" pageNumber="174" unit="mm" value="17.3">17.3 mm</quantity>
), Seto Inland Sea, Suonada, 
<geoCoordinate id="D38D3692A014D17AFF5FF9D4FECAF99D" box="[222,337,1539,1563]" direction="north" orientation="latitude" pageId="3" pageNumber="174" precision="92" value="33.716667">33°43.0’N</geoCoordinate>
<geoCoordinate id="D38D3692A014D17AFEDFF9D5FE45F99D" box="[350,478,1538,1563]" direction="east" orientation="longitude" pageId="3" pageNumber="174" precision="92" value="131.25">131°15.0’E</geoCoordinate>
, depth: 
<quantity id="7141FDB0A014D17AFDC1F9D4FD1BF99C" box="[576,640,1539,1563]" metricMagnitude="1" metricUnit="m" metricValue="1.8" pageId="3" pageNumber="174" unit="m" value="18.0">18 m</quantity>
, TRV “Toyoshio-maru”, 
<date id="C2077695A014D17AFC24F9D4FB9AF99D" box="[933,1025,1539,1563]" pageId="3" pageNumber="174" value="2000" valueMax="2010">2000-10</date>
cruise, sta. 15, sledge net, coll. S. Ohtsuka, 
<date id="C2077695A014D17AFF7EF9F1FE14F9B8" box="[255,399,1573,1598]" pageId="3" pageNumber="174" value="2000-07-14">14 July 2000</date>
; 
<specimenCount id="A0BF9BDCA014D17AFE18F9F1FE6CF9B8" box="[409,503,1573,1598]" pageId="3" pageNumber="174" type="female">1 female</specimenCount>
(cl 
<quantity id="7141FDB0A014D17AFDA0F9F2FDE3F9B8" box="[545,632,1573,1598]" metricMagnitude="-3" metricUnit="m" metricValue="7.6" pageId="3" pageNumber="174" unit="mm" value="7.6">7.6 mm</quantity>
), Kyushu, Ohita Prefecture, Beppu Bay, 
<geoCoordinate id="D38D3692A014D17AFBB3F9F2FB3EF9B8" box="[1074,1189,1573,1598]" direction="north" orientation="latitude" pageId="3" pageNumber="174" precision="92" value="33.28">33°16.8’N</geoCoordinate>
<geoCoordinate id="D38D3692A014D17AFB2AF9F2FAB7F9B8" box="[1195,1324,1573,1598]" direction="east" orientation="longitude" pageId="3" pageNumber="174" precision="92" value="131.77667">131°46.6’E</geoCoordinate>
, depth: 
<quantity id="7141FDB0A014D17AFA00F9F1FF34F9D9" metricMagnitude="1" metricUnit="m" metricValue="4.8" pageId="3" pageNumber="174" unit="m" value="48.0">48 m</quantity>
, dredge, TRV “Toyoshio-maru”, 1996-5 cruise, sta. 18, coll. T. Komai, 
<date id="C2077695A014D17AFC3FF990FBDEF9E6" box="[958,1093,1607,1632]" pageId="3" pageNumber="174" value="1996-06-06">6 June 1996</date>
. 
<collectingCountry id="CEAE10C5A014D17AFBD1F99FFB08F9D9" box="[1104,1171,1608,1631]" name="South Korea" pageId="3" pageNumber="174">Korea</collectingCountry>
: 
<specimenCount id="A0BF9BDCA014D17AFB20F99FFA9BF9E6" box="[1185,1280,1607,1632]" pageId="3" pageNumber="174" type="female">1 female</specimenCount>
(cl 
<quantity id="7141FDB0A014D17AFAAAF990FA0BF9E6" box="[1323,1424,1607,1632]" metricMagnitude="-2" metricUnit="m" metricValue="1.05" pageId="3" pageNumber="174" unit="mm" value="10.5">10.5 mm</quantity>
), CBM-ZC 5515; Kwangyang, depth: ca. 
<quantity id="7141FDB0A014D17AFDD8F9BCFD0EF904" box="[601,661,1643,1667]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="3" pageNumber="174" unit="m" value="10.0">10 m</quantity>
, commercial trawler, coll. F. Muto, 
<date id="C2077695A014D17AFBA9F9BDFB58F905" box="[1064,1219,1642,1667]" pageId="3" pageNumber="174" value="1999-10">October 1999</date>
. 
<collectingCountry id="CEAE10C5A014D17AFB4EF9BDFA8BF905" box="[1231,1296,1642,1667]" name="China" pageId="3" pageNumber="174">China</collectingCountry>
: 
<specimenCount id="A0BF9BDCA014D17AFAA1F9BCFAECF905" box="[1312,1399,1642,1667]" pageId="3" pageNumber="174" type="male">3 males</specimenCount>
(cl 13.9– 
<quantity id="7141FDB0A014D17AFF56F959FEBEF920" box="[215,293,1678,1702]" metricMagnitude="-2" metricUnit="m" metricValue="1.3" pageId="3" pageNumber="174" unit="mm" value="13.0">13 mm</quantity>
), 
<specimenCount id="A0BF9BDCA014D17AFEB9F959FE03F920" box="[312,408,1677,1702]" pageId="3" pageNumber="174" type="female">1 female</specimenCount>
(cl 
<quantity id="7141FDB0A014D17AFE40F95AFDBCF920" box="[449,551,1677,1702]" metricMagnitude="-2" metricUnit="m" metricValue="1.46" pageId="3" pageNumber="174" unit="mm" value="14.6">14.6 mm</quantity>
), 
<collectionCode id="D0A8C890A014D17AFDB8F959FDEAF920" box="[569,625,1678,1702]" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="3" pageNumber="174">ZRC</collectionCode>
.1998.1080, 
<collectingCountry id="CEAE10C5A014D17AFD76F95AFC16F920" box="[759,909,1677,1702]" name="China" pageId="3" pageNumber="174">Jiaozhou Bay</collectingCountry>
, coll. Zhong, 
<date id="C2077695A014D17AFBA2F95AFB43F920" box="[1059,1240,1677,1702]" pageId="3" pageNumber="174" value="1982-09">September 1982</date>
, det. J.Y.Liu.
</paragraph>
</subSubSection>
<subSubSection id="FEA303DEA014D17DFF46F978FC7CF96D" lastPageId="4" lastPageNumber="175" pageId="3" pageNumber="174" type="description">
<paragraph id="B6065055A014D17AFF46F978FA07F83D" blockId="3.[151,1437,637,2014]" pageId="3" pageNumber="174">
<emphasis id="84CD8C47A014D17AFF46F978FECAF94E" bold="true" box="[199,337,1711,1736]" pageId="3" pageNumber="174">Description</emphasis>
. Large-sized species of 
<taxonomicName id="71B92BD6A014D17AFDE2F967FCA9F94E" box="[611,818,1712,1736]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="174" phylum="Arthropoda" rank="species" species="edwardsii">
<emphasis id="84CD8C47A014D17AFDE2F967FCA9F94E" box="[611,818,1712,1736]" italics="true" pageId="3" pageNumber="174">Alpheus edwardsii</emphasis>
</taxonomicName>
species group, cl and tl of largest specimen (
<typeStatus id="6902EEF7A014D17AFAB0F967FA0AF94E" box="[1329,1425,1712,1736]" pageId="3" pageNumber="174" type="neotype">neotype</typeStatus>
): 
<quantity id="7141FDB0A014D17AFF16F904FE9BF96C" box="[151,256,1747,1771]" metricMagnitude="-2" metricUnit="m" metricValue="1.97" pageId="3" pageNumber="174" unit="mm" value="19.7">19.7 mm</quantity>
and 
<quantity id="7141FDB0A014D17AFEC0F905FE37F96C" box="[321,428,1746,1771]" metricMagnitude="-2" metricUnit="m" metricValue="6.45" pageId="3" pageNumber="174" unit="mm" value="64.5">64.5 mm</quantity>
, respectively. Rostrum conical, acute distally; rostral carina broad, not conspicuously demarked, smoothly flattening between orbital hoods; orbital hoods moderately swollen, rounded, unarmed; adrostral furrows shallow (
<figureCitation id="2E824CD0A014D17AFE42F8C0FD9BF8B6" box="[451,512,1815,1840]" captionStart="FIGURE 1" captionStartId="5.[151,250,1858,1880]" captionTargetBox="[153,1429,200,1831]" captionTargetId="figure@5.[153,1438,196,1840]" captionTargetPageId="5" captionText="FIGURE 1. Alpheus vladivostokiensis (Vinogradov, 1950): male (cl 18.0 mm) from Troitza Bay (= Bukhta Troitzy), Russian Far East (OUMNH. ZC. 2016 - 01 - 013): A, frontal region, dorsal; B, same, lateral; C, telson, dorsal; D, ventromesial carina of first article of antennular peduncle, lateral; E, third maxilliped, lateral; F, second pereiopod, lateral; G, third pereiopod, lateral; H, same, dactylus, dorsal; I, fifth pereiopod, lateral; J, same, propodus and dactylus, ventral; K, second pleopod, detail of appendices masculina and interna, dorsal; L uropod, dorsal." httpUri="https://zenodo.org/record/271759/files/figure.png" pageId="3" pageNumber="174">Fig. 1</figureCitation>
A, B). Carapace with pterygostomial angle rounded (
<figureCitation id="2E824CD0A014D17AFBD3F8C0FB0AF8B6" box="[1106,1169,1815,1840]" captionStart="FIGURE 1" captionStartId="5.[151,250,1858,1880]" captionTargetBox="[153,1429,200,1831]" captionTargetId="figure@5.[153,1438,196,1840]" captionTargetPageId="5" captionText="FIGURE 1. Alpheus vladivostokiensis (Vinogradov, 1950): male (cl 18.0 mm) from Troitza Bay (= Bukhta Troitzy), Russian Far East (OUMNH. ZC. 2016 - 01 - 013): A, frontal region, dorsal; B, same, lateral; C, telson, dorsal; D, ventromesial carina of first article of antennular peduncle, lateral; E, third maxilliped, lateral; F, second pereiopod, lateral; G, third pereiopod, lateral; H, same, dactylus, dorsal; I, fifth pereiopod, lateral; J, same, propodus and dactylus, ventral; K, second pleopod, detail of appendices masculina and interna, dorsal; L uropod, dorsal." httpUri="https://zenodo.org/record/271759/files/figure.png" pageId="3" pageNumber="174">Fig. 1</figureCitation>
B); cardiac notch deep. All pleonal pleura rounded marginally; sixth pleonite without articulated flap. Telson moderately broad, subrectangular, tapering posteriorly, dorsal spiniform setae well developed, situated at some distance from lateral margins, first pair anterior to telson mid-length, second pair at about 0.7–0.8 of telson length; posterior margin broadly rounded, each posterolateral angle with two small spiniform setae, lateral much shorter than mesial (
<figureCitation id="2E824CD0A014D17AFABBF875FAE3F83D" box="[1338,1400,1954,1979]" captionStart="FIGURE 1" captionStartId="5.[151,250,1858,1880]" captionTargetBox="[153,1429,200,1831]" captionTargetId="figure@5.[153,1438,196,1840]" captionTargetPageId="5" captionText="FIGURE 1. Alpheus vladivostokiensis (Vinogradov, 1950): male (cl 18.0 mm) from Troitza Bay (= Bukhta Troitzy), Russian Far East (OUMNH. ZC. 2016 - 01 - 013): A, frontal region, dorsal; B, same, lateral; C, telson, dorsal; D, ventromesial carina of first article of antennular peduncle, lateral; E, third maxilliped, lateral; F, second pereiopod, lateral; G, third pereiopod, lateral; H, same, dactylus, dorsal; I, fifth pereiopod, lateral; J, same, propodus and dactylus, ventral; K, second pleopod, detail of appendices masculina and interna, dorsal; L uropod, dorsal." httpUri="https://zenodo.org/record/271759/files/figure.png" pageId="3" pageNumber="174">Fig. 1</figureCitation>
C).
</paragraph>
<paragraph id="B6065055A014D17AFF46F812FA94F858" blockId="3.[151,1437,637,2014]" box="[199,1295,1989,2014]" pageId="3" pageNumber="174">Eyestalk with moderately developed, anterolaterally situated cornea; ocellar beak well developed.</paragraph>
<paragraph id="B6065055A013D17DFF46FF40FCBBFED8" blockId="4.[151,1437,151,2014]" pageId="4" pageNumber="175">
Antennular peduncle with second article slightly more than twice as long as wide; stylocerite reaching or almost reaching distal margin of first article; ventromesial carina of first article with triangular and usually subacutely pointed tooth (
<figureCitation id="2E824CD0A013D17DFE46FF0AFD91FF70" box="[455,522,221,246]" captionStart="FIGURE 1" captionStartId="5.[151,250,1858,1880]" captionTargetBox="[153,1429,200,1831]" captionTargetId="figure@5.[153,1438,196,1840]" captionTargetPageId="5" captionText="FIGURE 1. Alpheus vladivostokiensis (Vinogradov, 1950): male (cl 18.0 mm) from Troitza Bay (= Bukhta Troitzy), Russian Far East (OUMNH. ZC. 2016 - 01 - 013): A, frontal region, dorsal; B, same, lateral; C, telson, dorsal; D, ventromesial carina of first article of antennular peduncle, lateral; E, third maxilliped, lateral; F, second pereiopod, lateral; G, third pereiopod, lateral; H, same, dactylus, dorsal; I, fifth pereiopod, lateral; J, same, propodus and dactylus, ventral; K, second pleopod, detail of appendices masculina and interna, dorsal; L uropod, dorsal." httpUri="https://zenodo.org/record/271759/files/figure.png" pageId="4" pageNumber="175">Fig. 1</figureCitation>
A, B, D). Antenna with basicerite distally ending in large ventrolateral tooth; scaphocerite relatively broad, with slightly concave lateral margin and narrow blade, latter not over-reaching very strong distolateral tooth, separated from distal portion of blade by deep cleft; carpocerite reaching slightly beyond scaphocerite and end of antennular peduncle (
<figureCitation id="2E824CD0A013D17DFD18FE92FD4CFED8" box="[665,727,325,350]" captionStart="FIGURE 1" captionStartId="5.[151,250,1858,1880]" captionTargetBox="[153,1429,200,1831]" captionTargetId="figure@5.[153,1438,196,1840]" captionTargetPageId="5" captionText="FIGURE 1. Alpheus vladivostokiensis (Vinogradov, 1950): male (cl 18.0 mm) from Troitza Bay (= Bukhta Troitzy), Russian Far East (OUMNH. ZC. 2016 - 01 - 013): A, frontal region, dorsal; B, same, lateral; C, telson, dorsal; D, ventromesial carina of first article of antennular peduncle, lateral; E, third maxilliped, lateral; F, second pereiopod, lateral; G, third pereiopod, lateral; H, same, dactylus, dorsal; I, fifth pereiopod, lateral; J, same, propodus and dactylus, ventral; K, second pleopod, detail of appendices masculina and interna, dorsal; L uropod, dorsal." httpUri="https://zenodo.org/record/271759/files/figure.png" pageId="4" pageNumber="175">Fig. 1</figureCitation>
A, B).
</paragraph>
<paragraph id="B6065055A013D17DFF46FEB0FC12FE6E" blockId="4.[151,1437,151,2014]" pageId="4" pageNumber="175">
Mouthparts typical for 
<taxonomicName id="71B92BD6A013D17DFE55FEBFFDB5FE06" box="[468,558,360,384]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="175" phylum="Arthropoda" rank="genus">
<emphasis id="84CD8C47A013D17DFE55FEBFFDB5FE06" box="[468,558,360,384]" italics="true" pageId="4" pageNumber="175">Alpheus</emphasis>
</taxonomicName>
. Third maxilliped slender, pediform, not operculate; antepenultimate article flattened ventrolaterally; penultimate article about 1.5 times as long as high, widening distally; ultimate article about as long as antepenultimate article, gradually tapering distally, very setose; lateral plate of coxa dorsally protruding, subacute; single arthrobranch well developed (
<figureCitation id="2E824CD0A013D17DFCA4FE18FCFEFE6E" box="[805,869,463,488]" captionStart="FIGURE 1" captionStartId="5.[151,250,1858,1880]" captionTargetBox="[153,1429,200,1831]" captionTargetId="figure@5.[153,1438,196,1840]" captionTargetPageId="5" captionText="FIGURE 1. Alpheus vladivostokiensis (Vinogradov, 1950): male (cl 18.0 mm) from Troitza Bay (= Bukhta Troitzy), Russian Far East (OUMNH. ZC. 2016 - 01 - 013): A, frontal region, dorsal; B, same, lateral; C, telson, dorsal; D, ventromesial carina of first article of antennular peduncle, lateral; E, third maxilliped, lateral; F, second pereiopod, lateral; G, third pereiopod, lateral; H, same, dactylus, dorsal; I, fifth pereiopod, lateral; J, same, propodus and dactylus, ventral; K, second pleopod, detail of appendices masculina and interna, dorsal; L uropod, dorsal." httpUri="https://zenodo.org/record/271759/files/figure.png" pageId="4" pageNumber="175">Fig. 1</figureCitation>
E).
</paragraph>
<paragraph id="B6065055A013D17DFF46FE25FB35FCA6" blockId="4.[151,1437,151,2014]" pageId="4" pageNumber="175">
Major cheliped generally similar in both sexes, larger and with sometimes slightly different proportions in males. Ischium very short, unarmed. Merus very stout, blunt distodorsally; ventromesial margin rugose, with short spiniform setae, distally with large sharp tooth. Carpus very short, cup-shaped. Chela with palm about 1.8 times as long as fingers, distodorsal margin with broad notch extending onto lateral and mesial surfaces as shallow longitudinal grooves; dorsal shoulder distinctly overhanging distodorsal notch, rounded; ventral margin with very deep, broad notch, adjacent areas depressed on both mesial and lateral surface, ventral shoulder markedly protruding anteriorly, rounded; mesial surface of palm with small granules near ventral margin; distal margin of palm with sharp tooth laterally and acute or subacute (when not broken) tooth mesially; dactylus slightly overreaching pollex, with moderately developed, not greatly protruding plunger (
<figureCitation id="2E824CD0A013D17DFBA5FCD0FBFCFCA6" box="[1060,1127,775,800]" captionStart="FIGURE 2" captionStartId="6.[151,250,1709,1731]" captionTargetBox="[157,1419,203,1681]" captionTargetId="figure@6.[151,1436,193,1687]" captionTargetPageId="6" captionText="FIGURE 2. Alpheus vladivostokiensis (Vinogradov, 1950): male (cl 18.0 mm) from Troitza Bay (= Bukhta Troitzy), Russian Far East (OUMNH. ZC. 2016 - 01 - 013) [A – G]; ovigerous female (cl 17.4 mm) from the same locality (MNHN-IU- 2014 - 5694) [H, I]: A, major cheliped, lateral; B, major chela, mesial; C, same, fingers open, lateral; D, same, merus, mesial; E, minor cheliped, lateral; F, minor chela, mesial; G, same, merus, mesial; H, minor chela, mesial; I, minor cheliped, merus, mesial." httpUri="https://zenodo.org/record/271760/files/figure.png" pageId="4" pageNumber="175">Fig. 2</figureCitation>
A–D).
</paragraph>
<paragraph id="B6065055A013D17DFF46FCFDFEFEFBB0" blockId="4.[151,1437,151,2014]" pageId="4" pageNumber="175">
Male minor cheliped about as long as major cheliped, however, much more slender. Ischium very short, unarmed. Merus and carpus similar to those of major cheliped, noticeably more slender. Chela with palm approximately 1.4 times as long as fingers; distodorsal margin with moderately deep notch extending onto lateral surface as shallow longitudinal groove and onto mesial surface as short triangular groove; ventral margin with shallow, sometimes barely noticeable sinus; distal margin of palm with sharp tooth on both lateral and mesial sides; ventromesial surface of palm usually with some granules; dactylus somewhat expanded, especially distally, lateral and mesial ridges of distal half furnished with balaeniceps setae; pollex also with row of setae on lateral and mesial sides (
<figureCitation id="2E824CD0A013D17DFF5FFBCAFEBAFBB0" box="[222,289,1053,1078]" captionStart="FIGURE 2" captionStartId="6.[151,250,1709,1731]" captionTargetBox="[157,1419,203,1681]" captionTargetId="figure@6.[151,1436,193,1687]" captionTargetPageId="6" captionText="FIGURE 2. Alpheus vladivostokiensis (Vinogradov, 1950): male (cl 18.0 mm) from Troitza Bay (= Bukhta Troitzy), Russian Far East (OUMNH. ZC. 2016 - 01 - 013) [A – G]; ovigerous female (cl 17.4 mm) from the same locality (MNHN-IU- 2014 - 5694) [H, I]: A, major cheliped, lateral; B, major chela, mesial; C, same, fingers open, lateral; D, same, merus, mesial; E, minor cheliped, lateral; F, minor chela, mesial; G, same, merus, mesial; H, minor chela, mesial; I, minor cheliped, merus, mesial." httpUri="https://zenodo.org/record/271760/files/figure.png" pageId="4" pageNumber="175">Fig. 2</figureCitation>
E–G).
</paragraph>
<paragraph id="B6065055A013D17DFF46FBE8FDECFB46" blockId="4.[151,1437,151,2014]" pageId="4" pageNumber="175">
Female minor cheliped with ischium, merus and carpus similar to those of male minor cheliped. Chela with fingers almost as long as palm; palm with shallow distodorsal depression, not extending onto lateral and mesial surfaces, ventral margin without sinus; dactylus not expanded, without ridge and without balaeniceps setae; pollex also without balaeniceps setae (
<figureCitation id="2E824CD0A013D17DFE79FB70FDA0FB46" box="[504,571,1191,1216]" captionStart="FIGURE 2" captionStartId="6.[151,250,1709,1731]" captionTargetBox="[157,1419,203,1681]" captionTargetId="figure@6.[151,1436,193,1687]" captionTargetPageId="6" captionText="FIGURE 2. Alpheus vladivostokiensis (Vinogradov, 1950): male (cl 18.0 mm) from Troitza Bay (= Bukhta Troitzy), Russian Far East (OUMNH. ZC. 2016 - 01 - 013) [A – G]; ovigerous female (cl 17.4 mm) from the same locality (MNHN-IU- 2014 - 5694) [H, I]: A, major cheliped, lateral; B, major chela, mesial; C, same, fingers open, lateral; D, same, merus, mesial; E, minor cheliped, lateral; F, minor chela, mesial; G, same, merus, mesial; H, minor chela, mesial; I, minor cheliped, merus, mesial." httpUri="https://zenodo.org/record/271760/files/figure.png" pageId="4" pageNumber="175">Fig. 2</figureCitation>
H, I).
</paragraph>
<paragraph id="B6065055A013D17DFF46FB1DFA07FA7E" blockId="4.[151,1437,151,2014]" pageId="4" pageNumber="175">
Second pereiopod moderately slender; merus subequal to ischium; carpus five-jointed, ratio of joints approximately equal to 5:4:1:1:2; chela as long as second carpal joint (
<figureCitation id="2E824CD0A013D17DFC34FB3AFC6EFA80" box="[949,1013,1261,1286]" captionStart="FIGURE 1" captionStartId="5.[151,250,1858,1880]" captionTargetBox="[153,1429,200,1831]" captionTargetId="figure@5.[153,1438,196,1840]" captionTargetPageId="5" captionText="FIGURE 1. Alpheus vladivostokiensis (Vinogradov, 1950): male (cl 18.0 mm) from Troitza Bay (= Bukhta Troitzy), Russian Far East (OUMNH. ZC. 2016 - 01 - 013): A, frontal region, dorsal; B, same, lateral; C, telson, dorsal; D, ventromesial carina of first article of antennular peduncle, lateral; E, third maxilliped, lateral; F, second pereiopod, lateral; G, third pereiopod, lateral; H, same, dactylus, dorsal; I, fifth pereiopod, lateral; J, same, propodus and dactylus, ventral; K, second pleopod, detail of appendices masculina and interna, dorsal; L uropod, dorsal." httpUri="https://zenodo.org/record/271759/files/figure.png" pageId="4" pageNumber="175">Fig. 1</figureCitation>
F). Third pereiopod relatively robust; ischium with stout ventrolateral spiniform seta; merus about five times as long as wide, unarmed; carpus about half-length of merus, with long setae on dorsal surface; propodus with numerous long setae arising especially from dorsal and ventral surfaces and row of stout spiniform setae along ventral margin, including one pair of longer distoventral spiniform setae adjacent to dactylus; dactylus about one-third of propodus length, subspatulate (
<figureCitation id="2E824CD0A013D17DFAEEFAA0FF3AFA35" captionStart="FIGURE 1" captionStartId="5.[151,250,1858,1880]" captionTargetBox="[153,1429,200,1831]" captionTargetId="figure@5.[153,1438,196,1840]" captionTargetPageId="5" captionText="FIGURE 1. Alpheus vladivostokiensis (Vinogradov, 1950): male (cl 18.0 mm) from Troitza Bay (= Bukhta Troitzy), Russian Far East (OUMNH. ZC. 2016 - 01 - 013): A, frontal region, dorsal; B, same, lateral; C, telson, dorsal; D, ventromesial carina of first article of antennular peduncle, lateral; E, third maxilliped, lateral; F, second pereiopod, lateral; G, third pereiopod, lateral; H, same, dactylus, dorsal; I, fifth pereiopod, lateral; J, same, propodus and dactylus, ventral; K, second pleopod, detail of appendices masculina and interna, dorsal; L uropod, dorsal." httpUri="https://zenodo.org/record/271759/files/figure.png" pageId="4" pageNumber="175">Fig. 1</figureCitation>
G, H). Fourth pereiopod similar to third. Fifth pereiopod shorter and more slender than third or fourth pereiopods; ischium unarmed; carpus 0.75 length of merus; propodus with well-developed setal brush distally and several stout spiniform setae along ventromesial margin; dactylus less expanded compared to that of third pereiopod (
<figureCitation id="2E824CD0A013D17DFAA6FA08FAF3FA7E" box="[1319,1384,1503,1528]" captionStart="FIGURE 1" captionStartId="5.[151,250,1858,1880]" captionTargetBox="[153,1429,200,1831]" captionTargetId="figure@5.[153,1438,196,1840]" captionTargetPageId="5" captionText="FIGURE 1. Alpheus vladivostokiensis (Vinogradov, 1950): male (cl 18.0 mm) from Troitza Bay (= Bukhta Troitzy), Russian Far East (OUMNH. ZC. 2016 - 01 - 013): A, frontal region, dorsal; B, same, lateral; C, telson, dorsal; D, ventromesial carina of first article of antennular peduncle, lateral; E, third maxilliped, lateral; F, second pereiopod, lateral; G, third pereiopod, lateral; H, same, dactylus, dorsal; I, fifth pereiopod, lateral; J, same, propodus and dactylus, ventral; K, second pleopod, detail of appendices masculina and interna, dorsal; L uropod, dorsal." httpUri="https://zenodo.org/record/271759/files/figure.png" pageId="4" pageNumber="175">
Fig. 
<date id="C2077695A013D17DFADDFA37FAF3FA7E" box="[1372,1384,1504,1528]" pageId="4" pageNumber="175">1</date>
</figureCitation>
I, J).
</paragraph>
<paragraph id="B6065055A013D17DFF46F9D5FAE1F905" blockId="4.[151,1437,151,2014]" pageId="4" pageNumber="175">
Male second pleopod with appendix masculina slightly shorter than appendix interna, furnished with slender spiniform setae, as illustrated (
<figureCitation id="2E824CD0A013D17DFE77F9F2FDACF9B8" box="[502,567,1573,1598]" captionStart="FIGURE 1" captionStartId="5.[151,250,1858,1880]" captionTargetBox="[153,1429,200,1831]" captionTargetId="figure@5.[153,1438,196,1840]" captionTargetPageId="5" captionText="FIGURE 1. Alpheus vladivostokiensis (Vinogradov, 1950): male (cl 18.0 mm) from Troitza Bay (= Bukhta Troitzy), Russian Far East (OUMNH. ZC. 2016 - 01 - 013): A, frontal region, dorsal; B, same, lateral; C, telson, dorsal; D, ventromesial carina of first article of antennular peduncle, lateral; E, third maxilliped, lateral; F, second pereiopod, lateral; G, third pereiopod, lateral; H, same, dactylus, dorsal; I, fifth pereiopod, lateral; J, same, propodus and dactylus, ventral; K, second pleopod, detail of appendices masculina and interna, dorsal; L uropod, dorsal." httpUri="https://zenodo.org/record/271759/files/figure.png" pageId="4" pageNumber="175">Fig. 1</figureCitation>
K). Uropod with both lobes of protopod ending in acute or subacute points; exopod and endopod very broad, rounded; diaeresis of exopod slightly sinuous, lateral portion with broad rounded lobe adjacent to stout distolateral spiniform seta; distal margin of endopod with row of small spinules (
<figureCitation id="2E824CD0A013D17DFA97F9BDFACEF905" box="[1302,1365,1642,1667]" captionStart="FIGURE 1" captionStartId="5.[151,250,1858,1880]" captionTargetBox="[153,1429,200,1831]" captionTargetId="figure@5.[153,1438,196,1840]" captionTargetPageId="5" captionText="FIGURE 1. Alpheus vladivostokiensis (Vinogradov, 1950): male (cl 18.0 mm) from Troitza Bay (= Bukhta Troitzy), Russian Far East (OUMNH. ZC. 2016 - 01 - 013): A, frontal region, dorsal; B, same, lateral; C, telson, dorsal; D, ventromesial carina of first article of antennular peduncle, lateral; E, third maxilliped, lateral; F, second pereiopod, lateral; G, third pereiopod, lateral; H, same, dactylus, dorsal; I, fifth pereiopod, lateral; J, same, propodus and dactylus, ventral; K, second pleopod, detail of appendices masculina and interna, dorsal; L uropod, dorsal." httpUri="https://zenodo.org/record/271759/files/figure.png" pageId="4" pageNumber="175">Fig. 1</figureCitation>
L).
</paragraph>
<paragraph id="B6065055A013D17DFF46F95AFC7CF96D" blockId="4.[151,1437,151,2014]" pageId="4" pageNumber="175">
<emphasis id="84CD8C47A013D17DFF46F95AFEE7F920" bold="true" box="[199,380,1677,1702]" pageId="4" pageNumber="175">Colour pattern</emphasis>
. Overall dull olive-green or brown with some whitish areas on carapace; chelipeds greenishbrown or brown; walking legs reddish with white articulations; uropod greenish-olive proximally, bluish-grey distally; eggs olive-green (
<figureCitation id="2E824CD0A013D17DFE40F905FD8BF96D" box="[449,528,1746,1771]" captionStart="FIGURE 3" captionStartId="7.[151,250,1413,1435]" captionTargetBox="[151,1435,176,1380]" captionTargetId="figure@7.[151,1436,154,1397]" captionTargetPageId="7" captionText="FIGURE 3. Alpheus vladivostokiensis (Vinogradov, 1950), colour pattern: neotype, male (cl 19.7 mm) from Troitza Bay (= Bukhta Troitzy), Russian Far East (MNHN-IU- 2014 - 5693), in dorsal (A) and lateral (B) views. Photographs by Ivan N. Marin." httpUri="https://zenodo.org/record/271761/files/figure.png" pageId="4" pageNumber="175">Figs. 3</figureCitation>
, 
<figureCitation id="2E824CD0A013D17DFD9CF904FDB5F96D" box="[541,558,1747,1771]" captionStart="FIGURE 4" captionStartId="8.[151,250,1581,1603]" captionTargetBox="[154,1434,176,1579]" captionTargetId="figure@8.[154,1434,164,1584]" captionTargetPageId="8" captionText="FIGURE 4. Alpheus vladivostokiensis (Vinogradov, 1950), colour pattern: ovigerous female (cl 16.3 mm) from Troitza Bay (= Bukhta Troitzy), Russian Far East (OUMNH. ZC. 2016 - 01 - 014), in dorsal (A) and lateral (B) views. Photographs by Ivan N. Marin." httpUri="https://zenodo.org/record/271762/files/figure.png" pageId="4" pageNumber="175">4</figureCitation>
; see also 
<bibRefCitation id="D2282DA4A013D17DFD19F905FCDCF96D" author="Pasternak" box="[664,839,1746,1771]" pageId="4" pageNumber="175" refString="Pasternak, R. K. (1988) Life of animals [Zhizn' zhivotnykh], Vol. 2. Second edition. Prosvezscheniye, Moscow, 447 pp. [in Russian]" type="book" year="1988">Pasternak 1988</bibRefCitation>
; 
<bibRefCitation id="D2282DA4A013D17DFCD2F905FC40F96D" author="Marin" box="[851,987,1746,1771]" pageId="4" pageNumber="175" refString="Marin, I. N. (2013) Small atlas of decapod crustaceans of Russia [Malyi atlas desiatinogikh rakoobraznykh Rossii]. KMK Scientific Press, Moscow, 146 pp. [in Russian]" type="book" year="2013">Marin 2013</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="FEA303DEA013D17DFF46F922FC62F8B6" pageId="4" pageNumber="175" type="materials_examined">
<paragraph id="B6065055A013D17DFF46F922FC62F8B6" blockId="4.[151,1437,151,2014]" pageId="4" pageNumber="175">
<emphasis id="84CD8C47A013D17DFF46F922FEC6F888" bold="true" box="[199,349,1781,1806]" pageId="4" pageNumber="175">
<typeStatus id="6902EEF7A013D17DFF46F922FE99F888" box="[199,258,1781,1806]" pageId="4" pageNumber="175">Type</typeStatus>
locality
</emphasis>
. Zolotoi Rog Bay (= Bukhta Zolotoi Rog) near Vladivostok, 
<collectingCountry id="CEAE10C5A013D17DFB8CF922FBC3F888" box="[1037,1112,1781,1806]" name="Russia" pageId="4" pageNumber="175">Russia</collectingCountry>
(
<bibRefCitation id="D2282DA4A013D17DFBE7F922FAB4F888" author="Vinogradov" box="[1126,1327,1781,1806]" pageId="4" pageNumber="175" refString="Vinogradov, L. G. (1950) Keys to the shrimps, crayfishes and crabs of the Far East [Opredelitel' krevetok, rakov i krabov Dal'nego Vostoka]. Izvestia TINRO, 33 C, 179 - 350 [in Russian]" type="journal article" year="1950">Vinogradov 1950</bibRefCitation>
); 
<typeStatus id="6902EEF7A013D17DFAC2F921FA07F888" box="[1347,1436,1782,1806]" pageId="4" pageNumber="175" type="neotype">neotype</typeStatus>
from Troitza Bay (= Bukhta Troitzy), about 
<quantity id="7141FDB0A013D17DFD04F8C0FD57F8B6" box="[645,716,1815,1840]" metricMagnitude="4" metricUnit="m" metricValue="6.0" pageId="4" pageNumber="175" unit="km" value="60.0">60 km</quantity>
southwest of Vladivostok.
</paragraph>
</subSubSection>
<subSubSection id="FEA303DEA013D17DFF46F8EDFBA0F858" pageId="4" pageNumber="175" type="distribution">
<paragraph id="B6065055A013D17DFF46F8EDFB3DF81E" blockId="4.[151,1437,151,2014]" pageId="4" pageNumber="175">
<emphasis id="84CD8C47A013D17DFF46F8EDFEC2F8D5" bold="true" box="[199,345,1850,1875]" pageId="4" pageNumber="175">Distribution</emphasis>
. Northwestern Pacific: northern Sea of 
<collectingCountry id="CEAE10C5A013D17DFC9CF8ECFCFBF8D4" box="[797,864,1851,1874]" name="Japan" pageId="4" pageNumber="175">Japan</collectingCountry>
. Presently known only from a few localities in the Russian Far East (Primorskiy Krai), e.g., Vladivostok area, Peter the Great Bay (= Zaliv Petra Velikogo), Troitza Bay (= Bukhta Troitzy), as well as from Hakodate Bay in southern Hokkaido, 
<collectingCountry id="CEAE10C5A013D17DFB85F856FBDFF81E" box="[1028,1092,1921,1944]" name="Japan" pageId="4" pageNumber="175">Japan</collectingCountry>
(
<figureCitation id="2E824CD0A013D17DFBD2F8A8FB03F81E" box="[1107,1176,1919,1944]" captionStart="FIGURE 7" captionStartId="11.[151,250,1178,1200]" captionTargetBox="[151,1435,195,1156]" captionTargetId="figure@11.[151,1435,195,1156]" captionTargetPageId="11" captionText="FIGURE 7. Map of the eastern Asian seas showing the presently known distributions of Alpheus vladivostokiensis Vinogradov, 1950 (red circles) and A. japonicus Miers, 1879 (blue circles), compiled based on the material examined in the present study and previously published, confirmable records (e. g., Miers 1879; Spence Bate 1888; Ortmann 1890; Yokoya 1930; Kim 1977; Banner &amp; Banner 1984; Komai et al. 1992; Anker 2001; Cha et al. 2001; Xu et al. 2015)." httpUri="https://zenodo.org/record/271765/files/figure.png" pageId="4" pageNumber="175">Fig. 7</figureCitation>
).
</paragraph>
<paragraph id="B6065055A013D17DFF46F875FBA0F858" blockId="4.[151,1437,151,2014]" pageId="4" pageNumber="175">
<emphasis id="84CD8C47A013D17DFF46F875FEBFF83D" bold="true" box="[199,292,1954,1979]" pageId="4" pageNumber="175">Ecology</emphasis>
. Intertidal and shallow subtidal to at least 
<quantity id="7141FDB0A013D17DFC80F874FCA6F83C" box="[769,829,1955,1979]" metricMagnitude="1" metricUnit="m" metricValue="3.0" pageId="4" pageNumber="175" unit="m" value="30.0">30 m</quantity>
; on muddy or mixed muddy-rocky bottoms; in simple burrows dug in mud or mud-sand under large rocks; typically in male-female pairs.
</paragraph>
</subSubSection>
<caption id="E2C600DDA012D17CFF16F895FD68F855" httpUri="https://zenodo.org/record/271759/files/figure.png" pageId="5" pageNumber="176" targetBox="[153,1429,200,1831]" targetPageId="5">
<paragraph id="B6065055A012D17CFF16F895FD68F855" blockId="5.[151,1436,1858,2003]" pageId="5" pageNumber="176">
<emphasis id="84CD8C47A012D17CFF16F895FE95F8D1" bold="true" box="[151,270,1858,1880]" pageId="5" pageNumber="176">FIGURE 1</emphasis>
. 
<taxonomicName id="71B92BD6A012D17CFE9AF895FD6FF8DE" authority="Vinogradov, 1950" authorityName="Vinogradov" authorityYear="1950" box="[283,756,1858,1880]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="176" phylum="Arthropoda" rank="species" species="vladivostokiensis">
<emphasis id="84CD8C47A012D17CFE9AF895FDBAF8DE" box="[283,545,1858,1880]" italics="true" pageId="5" pageNumber="176">Alpheus vladivostokiensis</emphasis>
(Vinogradov, 1950)
</taxonomicName>
: male (cl 18.0 mm) from Troitza Bay (= Bukhta Troitzy), Russian Far East (OUMNH.ZC. 2016-01-013): A, frontal region, dorsal; B, same, lateral; C, telson, dorsal; D, ventromesial carina of first article of antennular peduncle, lateral; E, third maxilliped, lateral; F, second pereiopod, lateral; G, third pereiopod, lateral; H, same, dactylus, dorsal; I, fifth pereiopod, lateral; J, same, propodus and dactylus, ventral; K, second pleopod, detail of appendices masculina and interna, dorsal; L uropod, dorsal.
</paragraph>
</caption>
<caption id="E2C600DDA011D17FFF16F97AFAF1F899" httpUri="https://zenodo.org/record/271760/files/figure.png" pageId="6" pageNumber="177" targetBox="[157,1419,203,1681]" targetPageId="6">
<paragraph id="B6065055A011D17FFF16F97AFAF1F899" blockId="6.[151,1436,1708,1823]" pageId="6" pageNumber="177">
<emphasis id="84CD8C47A011D17FFF16F97AFE95F944" bold="true" box="[151,270,1709,1731]" pageId="6" pageNumber="177">FIGURE 2</emphasis>
. 
<taxonomicName id="71B92BD6A011D17FFE9AF97BFD6FF944" authority="Vinogradov, 1950" authorityName="Vinogradov" authorityYear="1950" box="[283,756,1708,1730]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="177" phylum="Arthropoda" rank="species" species="vladivostokiensis">
<emphasis id="84CD8C47A011D17FFE9AF97BFDBAF944" box="[283,545,1708,1730]" italics="true" pageId="6" pageNumber="177">Alpheus vladivostokiensis</emphasis>
(Vinogradov, 1950)
</taxonomicName>
: male (cl 18.0 mm) from Troitza Bay (= Bukhta Troitzy), Russian Far East (OUMNH.ZC. 2016-01-013) [A–G]; ovigerous female (cl 17.4 mm) from the same locality (MNHN-IU-2014-5694) [H, I]: A, major cheliped, lateral; B, major chela, mesial; C, same, fingers open, lateral; D, same, merus, mesial; E, minor cheliped, lateral; F, minor chela, mesial; G, same, merus, mesial; H, minor chela, mesial; I, minor cheliped, merus, mesial.
</paragraph>
</caption>
<caption id="E2C600DDA010D17EFF16FA52FA06FA3C" httpUri="https://zenodo.org/record/271761/files/figure.png" pageId="7" pageNumber="178" targetBox="[151,1435,176,1380]" targetPageId="7">
<paragraph id="B6065055A010D17EFF16FA52FA06FA3C" blockId="7.[151,1437,1413,1466]" pageId="7" pageNumber="178">
<emphasis id="84CD8C47A010D17EFF16FA52FE94FA1D" bold="true" box="[151,271,1413,1435]" pageId="7" pageNumber="178">FIGURE 3</emphasis>
. 
<taxonomicName id="71B92BD6A010D17EFE9CFA52FD6DFA1D" authority="Vinogradov, 1950" authorityName="Vinogradov" authorityYear="1950" box="[285,758,1413,1435]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="178" phylum="Arthropoda" rank="species" species="vladivostokiensis">
<emphasis id="84CD8C47A010D17EFE9CFA52FDBFFA1D" box="[285,548,1413,1435]" italics="true" pageId="7" pageNumber="178">Alpheus vladivostokiensis</emphasis>
(Vinogradov, 1950)
</taxonomicName>
, colour pattern: neotype, male (cl 19.7 mm) from Troitza Bay (= Bukhta Troitzy), Russian Far East (MNHN-IU-2014-5693), in dorsal (A) and lateral (B) views. Photographs by Ivan N. Marin.
</paragraph>
</caption>
<subSubSection id="FEA303DEA010D173FF46FA31FDA9F8C6" lastPageId="10" lastPageNumber="181" pageId="7" pageNumber="178" type="discussion">
<paragraph id="B6065055A010D17EFF46FA31FC21F872" blockId="7.[151,1436,1510,2036]" pageId="7" pageNumber="178">
<emphasis id="84CD8C47A010D17EFF46FA31FEAFFA79" bold="true" box="[199,308,1510,1535]" pageId="7" pageNumber="178">Remarks</emphasis>
. 
<taxonomicName id="71B92BD6A010D17EFEC7FA30FDF0FA79" box="[326,619,1511,1535]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="178" phylum="Arthropoda" rank="species" species="vladivostokiensis">
<emphasis id="84CD8C47A010D17EFEC7FA30FDF0FA79" box="[326,619,1511,1535]" italics="true" pageId="7" pageNumber="178">Alpheus vladivostokiensis</emphasis>
</taxonomicName>
is closely related to 
<taxonomicName id="71B92BD6A010D17EFCE8FA30FC62FA79" box="[873,1017,1511,1535]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="178" phylum="Arthropoda" rank="species" species="japonicus">
<emphasis id="84CD8C47A010D17EFCE8FA30FC62FA79" box="[873,1017,1511,1535]" italics="true" pageId="7" pageNumber="178">A. japonicus</emphasis>
</taxonomicName>
(
<figureCitation id="2E824CD0A010D17EFB8CFA31FBFAFA79" box="[1037,1121,1510,1535]" captionStart="FIGURE 5" captionStartId="9.[151,250,1805,1827]" captionTargetBox="[213,1387,381,1749]" captionTargetId="figure@9.[199,1399,375,1763]" captionTargetPageId="9" captionText="FIGURE 5. Alpheus japonicus Miers, 1879: male (13.9 mm) from Tokyo Bay, Japan (OUMNH. ZC. 2016 - 01 - 015): A, frontal region, dorsal; B, telson, dorsal; C, first article of antennular peduncle, lateral; D, major chela, mesial; E, minor chela, mesial; F, third pereiopod, dactylus, dorsal; G, second pleopod, detail of appendices masculina and interna, dorsal. For additional illustrations see Liu (1955), Kim (1977) and Banner &amp; Banner (1984)." httpUri="https://zenodo.org/record/271763/files/figure.png" pageId="7" pageNumber="178">Figs. 5</figureCitation>
, 
<figureCitation id="2E824CD0A010D17EFBF0FA31FB1AFA79" box="[1137,1153,1510,1535]" captionStart="FIGURE 6" captionStartId="10.[151,250,1097,1119]" captionTargetBox="[151,1435,193,1074]" captionTargetId="figure@10.[151,1435,193,1075]" captionTargetPageId="10" captionText="FIGURE 6. Alpheus japonicus Miers, 1879, colour pattern: A, B, young male dredged at 50 – 60 m off Awaji Island, Japan (aquarium photograph, specimen not deposited); C, four adult males from a seafood market in Tokyo, Japan (OUMNH. ZC. 2016 - 01 - 015, MNHN-IU- 2014 - 5695, and MZUSP 34290); D, adult female from unknown locality in southern Korea. Photographs courtesy of Tomoyasu Tamego (A, B), Wataru Doi (C) and NFRDI (D)." httpUri="https://zenodo.org/record/271764/files/figure.png" pageId="7" pageNumber="178">6</figureCitation>
), a species occurring in deeper water (below 
<quantity id="7141FDB0A010D17EFE00F9DEFE20F9A6" box="[385,443,1545,1569]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="7" pageNumber="178" unit="m" value="10.0">10 m</quantity>
, typically 
<quantity id="7141FDB0A010D17EFDAEF9DFFD08F9A7" box="[559,659,1544,1569]" metricMagnitude="1" metricUnit="m" metricValue="5.0" metricValueMax="6.0" metricValueMin="4.0" pageId="7" pageNumber="178" unit="m" value="50.0" valueMax="60.0" valueMin="40.0">40–60 m</quantity>
) off the coasts of 
<collectingCountry id="CEAE10C5A010D17EFCD8F9DEFC07F9A6" box="[857,924,1545,1568]" name="Japan" pageId="7" pageNumber="178">Japan</collectingCountry>
, 
<collectingCountry id="CEAE10C5A010D17EFC26F9DEFC75F9A6" box="[935,1006,1545,1568]" name="South Korea" pageId="7" pageNumber="178">Korea</collectingCountry>
, 
<collectingCountry id="CEAE10C5A010D17EFC79F9DFFBA0F9A7" box="[1016,1083,1544,1569]" name="China" pageId="7" pageNumber="178">China</collectingCountry>
and 
<collectingCountry id="CEAE10C5A010D17EFBF0F9DFFB58F9A7" box="[1137,1219,1544,1569]" name="Taiwan" pageId="7" pageNumber="178">Taiwan</collectingCountry>
(
<bibRefCitation id="D2282DA4A010D17EFB51F9DFFAF1F9A7" author="Yokoya" box="[1232,1386,1544,1569]" pageId="7" pageNumber="178" refString="Yokoya, Y. (1930) Report of the biological survey of Mutsu Bay. 16. Macrura of Mutsu Bay. Science Reports of the Tohoku Imperial University, 4 ser., Biology, 5, 525 - 548, pl. 1." type="journal article" year="1930">Yokoya 1930</bibRefCitation>
; 
<bibRefCitation id="D2282DA4A010D17EFAF4F9DFFF49F9C4" author="Liu" pageId="7" pageNumber="178" refString="Liu, J. Y. (1955) Economic shrimps and prawns of northern China. Marine Biological Laboratory, Academy of Sciences, Scientific Publishing House, Beijing, viii + 73 pp., 24 pls. [in Chinese]" type="book" year="1955">Liu 1955</bibRefCitation>
; 
<bibRefCitation id="D2282DA4A010D17EFF5FF9FEFEC8F9C4" author="Kim" box="[222,339,1577,1602]" pageId="7" pageNumber="178" refString="Kim, H. S. (1977) Macrura. Illustrated flora and fauna of Korea, vol. 19. Samwha, Seoul, 414 pp., 56 pls. [in Korean]" type="book" year="1977">Kim 1977</bibRefCitation>
; 
<bibRefCitation id="D2282DA4A010D17EFEDEF9FDFDF2F9C4" author="Banner" box="[351,617,1578,1602]" pageId="7" pageNumber="178" refString="Banner, D. M. &amp; Banner, A. H. (1984) Old and unreported collections of alpheid shrimp from the Zoologisches Museum Berlin, principally from Melanesia. Pacific Science, 38, 34 - 50." type="journal article" year="1984">Banner &amp; Banner 1984</bibRefCitation>
; 
<bibRefCitation id="D2282DA4A010D17EFDF4F9FEFCAAF9C4" author="Baba" box="[629,817,1577,1602]" pageId="7" pageNumber="178" refString="Baba, K., Hayashi, K. I. &amp; Toriyama, M. (1986) Decapod crustaceans from continental shelf and slope around Japan. Japan Fisheries Resource Conservation Association, Tokyo, 336 pp." type="book" year="1986">
Baba 
<emphasis id="84CD8C47A010D17EFD37F9FDFD74F9C4" box="[694,751,1578,1602]" italics="true" pageId="7" pageNumber="178">et al.</emphasis>
1986
</bibRefCitation>
; 
<bibRefCitation id="D2282DA4A010D17EFCBCF9FEFC47F9C4" author="Hayashi" box="[829,988,1577,1602]" pageId="7" pageNumber="178" refString="Hayashi, K. - I. (1998) Prawns, shrimps and lobsters from Japan (102). Family Alpheidae -- Genus Alpheus 11. Aquabiology, 118, 390 - 395. [in Japanese]" type="journal article" year="1998">Hayashi 1998</bibRefCitation>
; 
<bibRefCitation id="D2282DA4A010D17EFC69F9FEFBEAF9C4" author="Anker" box="[1000,1137,1577,1602]" pageId="7" pageNumber="178" refString="Anker, A. (2001) Taxonomie et evolution des Alpheidae (Crustacea, Decapoda). Vol. 1 &amp; 2. PhD thesis. Museum National d'Histoire Naturelle, Paris, 547 pp. &amp; 331 pp." type="book" year="2001">Anker 2001</bibRefCitation>
; 
<bibRefCitation id="D2282DA4A010D17EFBFCF9FEFAB5F9C4" author="Cha" box="[1149,1326,1577,1602]" pageId="7" pageNumber="178" refString="Cha, H. K., Lee, J. U., Park, C. S., Baik, C. I., Hong, S. Y., Park, J. H., Lee, D. W., Choi, Y. M., Hwang, K., Kim, Z. G., Choi, K. H., Sohn, H., Sohn, M. H., Kim, D. H. &amp; Choi, J. H. (2001) Shrimps of the Korean Waters. National Fisheries Research and Development Institute, Pusan, 188 pp." type="book" year="2001">
Cha 
<emphasis id="84CD8C47A010D17EFB33F9FDFB78F9C4" box="[1202,1251,1578,1602]" italics="true" pageId="7" pageNumber="178">et al</emphasis>
. 2001
</bibRefCitation>
; 
<bibRefCitation id="D2282DA4A010D17EFABBF9FDFF4AF9E2" author="Xu" pageId="7" pageNumber="178" refString="Xu, Z. L., Shen, X. M. &amp; Gao, Q. (2015) Marine biology of the Changjiang (Yangtze River) estuary and adjacent East China Sea shelf. In: Zhang, J. (Ed.), Ecological continuum from the Changjiang (Yangtze River) watersheds to the East China Sea continental margin, Springer, pp. 161 - 180." type="book chapter" year="2015">
Xu 
<emphasis id="84CD8C47A010D17EFAE2F9FDFA07F9C4" box="[1379,1436,1578,1602]" italics="true" pageId="7" pageNumber="178">et al.</emphasis>
2015
</bibRefCitation>
; see also 
<figureCitation id="2E824CD0A010D17EFEB6F99CFEE2F9E5" box="[311,377,1611,1636]" captionStart="FIGURE 7" captionStartId="11.[151,250,1178,1200]" captionTargetBox="[151,1435,195,1156]" captionTargetId="figure@11.[151,1435,195,1156]" captionTargetPageId="11" captionText="FIGURE 7. Map of the eastern Asian seas showing the presently known distributions of Alpheus vladivostokiensis Vinogradov, 1950 (red circles) and A. japonicus Miers, 1879 (blue circles), compiled based on the material examined in the present study and previously published, confirmable records (e. g., Miers 1879; Spence Bate 1888; Ortmann 1890; Yokoya 1930; Kim 1977; Banner &amp; Banner 1984; Komai et al. 1992; Anker 2001; Cha et al. 2001; Xu et al. 2015)." httpUri="https://zenodo.org/record/271765/files/figure.png" pageId="7" pageNumber="178">Fig. 7</figureCitation>
). The records from 
<collectingCountry id="CEAE10C5A010D17EFDCEF99CFD05F9E2" box="[591,670,1611,1636]" name="Taiwan" pageId="7" pageNumber="178">Taiwan</collectingCountry>
are based on material from Dasi and Nanfang-ao fishing ports trawled off northwestern 
<collectingCountry id="CEAE10C5A010D17EFED0F9BBFE3BF903" box="[337,416,1644,1669]" name="Taiwan" pageId="7" pageNumber="178">Taiwan</collectingCountry>
(
<bibRefCitation id="D2282DA4A010D17EFE2EF9BBFCDFF903" author="Anker" box="[431,836,1644,1669]" pageId="7" pageNumber="178" refString="Anker, A. (2001) Taxonomie et evolution des Alpheidae (Crustacea, Decapoda). Vol. 1 &amp; 2. PhD thesis. Museum National d'Histoire Naturelle, Paris, 547 pp. &amp; 331 pp." type="book" year="2001">Anker 2001; T.Y. Chan, pers. comm.</bibRefCitation>
). The two species can be distinguished from each other by a suite of morphological characters (cf. 
<figureCitation id="2E824CD0A010D17EFDEAF95AFD2DF920" box="[619,694,1677,1702]" captionStart="FIGURE 1" captionStartId="5.[151,250,1858,1880]" captionTargetBox="[153,1429,200,1831]" captionTargetId="figure@5.[153,1438,196,1840]" captionTargetPageId="5" captionText="FIGURE 1. Alpheus vladivostokiensis (Vinogradov, 1950): male (cl 18.0 mm) from Troitza Bay (= Bukhta Troitzy), Russian Far East (OUMNH. ZC. 2016 - 01 - 013): A, frontal region, dorsal; B, same, lateral; C, telson, dorsal; D, ventromesial carina of first article of antennular peduncle, lateral; E, third maxilliped, lateral; F, second pereiopod, lateral; G, third pereiopod, lateral; H, same, dactylus, dorsal; I, fifth pereiopod, lateral; J, same, propodus and dactylus, ventral; K, second pleopod, detail of appendices masculina and interna, dorsal; L uropod, dorsal." httpUri="https://zenodo.org/record/271759/files/figure.png" pageId="7" pageNumber="178">Figs. 1</figureCitation>
, 
<figureCitation id="2E824CD0A010D17EFD42F959FD48F920" box="[707,723,1678,1702]" captionStart="FIGURE 2" captionStartId="6.[151,250,1709,1731]" captionTargetBox="[157,1419,203,1681]" captionTargetId="figure@6.[151,1436,193,1687]" captionTargetPageId="6" captionText="FIGURE 2. Alpheus vladivostokiensis (Vinogradov, 1950): male (cl 18.0 mm) from Troitza Bay (= Bukhta Troitzy), Russian Far East (OUMNH. ZC. 2016 - 01 - 013) [A – G]; ovigerous female (cl 17.4 mm) from the same locality (MNHN-IU- 2014 - 5694) [H, I]: A, major cheliped, lateral; B, major chela, mesial; C, same, fingers open, lateral; D, same, merus, mesial; E, minor cheliped, lateral; F, minor chela, mesial; G, same, merus, mesial; H, minor chela, mesial; I, minor cheliped, merus, mesial." httpUri="https://zenodo.org/record/271760/files/figure.png" pageId="7" pageNumber="178">2</figureCitation>
, 
<figureCitation id="2E824CD0A010D17EFD5EF95AFD75F920" box="[735,750,1677,1702]" captionStart="FIGURE 5" captionStartId="9.[151,250,1805,1827]" captionTargetBox="[213,1387,381,1749]" captionTargetId="figure@9.[199,1399,375,1763]" captionTargetPageId="9" captionText="FIGURE 5. Alpheus japonicus Miers, 1879: male (13.9 mm) from Tokyo Bay, Japan (OUMNH. ZC. 2016 - 01 - 015): A, frontal region, dorsal; B, telson, dorsal; C, first article of antennular peduncle, lateral; D, major chela, mesial; E, minor chela, mesial; F, third pereiopod, dactylus, dorsal; G, second pleopod, detail of appendices masculina and interna, dorsal. For additional illustrations see Liu (1955), Kim (1977) and Banner &amp; Banner (1984)." httpUri="https://zenodo.org/record/271763/files/figure.png" pageId="7" pageNumber="178">5</figureCitation>
). In 
<taxonomicName id="71B92BD6A010D17EFCA0F959FC62F923" box="[801,1017,1677,1701]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="178" phylum="Arthropoda" rank="species" species="vladivostokiensis">
<emphasis id="84CD8C47A010D17EFCA0F959FC62F923" box="[801,1017,1677,1701]" italics="true" pageId="7" pageNumber="178">A. vladivostokiensis</emphasis>
</taxonomicName>
, the rostral carina is slightly widening posteriorly between the orbital hoods (
<figureCitation id="2E824CD0A010D17EFDC7F979FD1EF941" box="[582,645,1710,1735]" captionStart="FIGURE 1" captionStartId="5.[151,250,1858,1880]" captionTargetBox="[153,1429,200,1831]" captionTargetId="figure@5.[153,1438,196,1840]" captionTargetPageId="5" captionText="FIGURE 1. Alpheus vladivostokiensis (Vinogradov, 1950): male (cl 18.0 mm) from Troitza Bay (= Bukhta Troitzy), Russian Far East (OUMNH. ZC. 2016 - 01 - 013): A, frontal region, dorsal; B, same, lateral; C, telson, dorsal; D, ventromesial carina of first article of antennular peduncle, lateral; E, third maxilliped, lateral; F, second pereiopod, lateral; G, third pereiopod, lateral; H, same, dactylus, dorsal; I, fifth pereiopod, lateral; J, same, propodus and dactylus, ventral; K, second pleopod, detail of appendices masculina and interna, dorsal; L uropod, dorsal." httpUri="https://zenodo.org/record/271759/files/figure.png" pageId="7" pageNumber="178">Fig. 1</figureCitation>
A), which is not the case of 
<taxonomicName id="71B92BD6A010D17EFC4DF978FBCDF941" box="[972,1110,1711,1735]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="178" phylum="Arthropoda" rank="species" species="japonicus">
<emphasis id="84CD8C47A010D17EFC4DF978FBCDF941" box="[972,1110,1711,1735]" italics="true" pageId="7" pageNumber="178">A. japonicus</emphasis>
</taxonomicName>
, in which the carina remains relatively narrow (
<figureCitation id="2E824CD0A010D17EFEF3F907FE27F96F" box="[370,444,1744,1769]" captionStart="FIGURE 5" captionStartId="9.[151,250,1805,1827]" captionTargetBox="[213,1387,381,1749]" captionTargetId="figure@9.[199,1399,375,1763]" captionTargetPageId="9" captionText="FIGURE 5. Alpheus japonicus Miers, 1879: male (13.9 mm) from Tokyo Bay, Japan (OUMNH. ZC. 2016 - 01 - 015): A, frontal region, dorsal; B, telson, dorsal; C, first article of antennular peduncle, lateral; D, major chela, mesial; E, minor chela, mesial; F, third pereiopod, dactylus, dorsal; G, second pleopod, detail of appendices masculina and interna, dorsal. For additional illustrations see Liu (1955), Kim (1977) and Banner &amp; Banner (1984)." httpUri="https://zenodo.org/record/271763/files/figure.png" pageId="7" pageNumber="178">Fig. 5</figureCitation>
A). The dactylus of the third and fourth pereiopods is much less expanded in 
<taxonomicName id="71B92BD6A010D17EFA05F906FECAF88C" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="178" phylum="Arthropoda" rank="species" species="vladivostokiensis">
<emphasis id="84CD8C47A010D17EFA05F906FECAF88C" italics="true" pageId="7" pageNumber="178">A. vladivostokiensis</emphasis>
</taxonomicName>
(
<figureCitation id="2E824CD0A010D17EFEDEF926FE00F88C" box="[351,411,1777,1802]" captionStart="FIGURE 1" captionStartId="5.[151,250,1858,1880]" captionTargetBox="[153,1429,200,1831]" captionTargetId="figure@5.[153,1438,196,1840]" captionTargetPageId="5" captionText="FIGURE 1. Alpheus vladivostokiensis (Vinogradov, 1950): male (cl 18.0 mm) from Troitza Bay (= Bukhta Troitzy), Russian Far East (OUMNH. ZC. 2016 - 01 - 013): A, frontal region, dorsal; B, same, lateral; C, telson, dorsal; D, ventromesial carina of first article of antennular peduncle, lateral; E, third maxilliped, lateral; F, second pereiopod, lateral; G, third pereiopod, lateral; H, same, dactylus, dorsal; I, fifth pereiopod, lateral; J, same, propodus and dactylus, ventral; K, second pleopod, detail of appendices masculina and interna, dorsal; L uropod, dorsal." httpUri="https://zenodo.org/record/271759/files/figure.png" pageId="7" pageNumber="178">Fig. 1</figureCitation>
H) compared to that of 
<taxonomicName id="71B92BD6A010D17EFD1AF925FCB8F88F" box="[667,803,1778,1801]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="178" phylum="Arthropoda" rank="species" species="japonicus">
<emphasis id="84CD8C47A010D17EFD1AF925FCB8F88F" box="[667,803,1778,1801]" italics="true" pageId="7" pageNumber="178">A. japonicus</emphasis>
</taxonomicName>
(
<figureCitation id="2E824CD0A010D17EFCB0F926FCEAF88C" box="[817,881,1777,1802]" captionStart="FIGURE 5" captionStartId="9.[151,250,1805,1827]" captionTargetBox="[213,1387,381,1749]" captionTargetId="figure@9.[199,1399,375,1763]" captionTargetPageId="9" captionText="FIGURE 5. Alpheus japonicus Miers, 1879: male (13.9 mm) from Tokyo Bay, Japan (OUMNH. ZC. 2016 - 01 - 015): A, frontal region, dorsal; B, telson, dorsal; C, first article of antennular peduncle, lateral; D, major chela, mesial; E, minor chela, mesial; F, third pereiopod, dactylus, dorsal; G, second pleopod, detail of appendices masculina and interna, dorsal. For additional illustrations see Liu (1955), Kim (1977) and Banner &amp; Banner (1984)." httpUri="https://zenodo.org/record/271763/files/figure.png" pageId="7" pageNumber="178">Fig. 5</figureCitation>
F). The male chelipeds in 
<taxonomicName id="71B92BD6A010D17EFB0BF925FAFAF88C" box="[1162,1377,1778,1802]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="178" phylum="Arthropoda" rank="species" species="vladivostokiensis">
<emphasis id="84CD8C47A010D17EFB0BF925FAFAF88C" box="[1162,1377,1778,1802]" italics="true" pageId="7" pageNumber="178">A. vladivostokiensis</emphasis>
</taxonomicName>
(
<figureCitation id="2E824CD0A010D17EFAF1F926FF3EF8AD" captionStart="FIGURE 2" captionStartId="6.[151,250,1709,1731]" captionTargetBox="[157,1419,203,1681]" captionTargetId="figure@6.[151,1436,193,1687]" captionTargetPageId="6" captionText="FIGURE 2. Alpheus vladivostokiensis (Vinogradov, 1950): male (cl 18.0 mm) from Troitza Bay (= Bukhta Troitzy), Russian Far East (OUMNH. ZC. 2016 - 01 - 013) [A – G]; ovigerous female (cl 17.4 mm) from the same locality (MNHN-IU- 2014 - 5694) [H, I]: A, major cheliped, lateral; B, major chela, mesial; C, same, fingers open, lateral; D, same, merus, mesial; E, minor cheliped, lateral; F, minor chela, mesial; G, same, merus, mesial; H, minor chela, mesial; I, minor cheliped, merus, mesial." httpUri="https://zenodo.org/record/271760/files/figure.png" pageId="7" pageNumber="178">Fig. 2</figureCitation>
A, B, E, F) are noticeably stouter than in 
<taxonomicName id="71B92BD6A010D17EFDFCF8C4FC93F8AD" box="[637,776,1811,1835]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="178" phylum="Arthropoda" rank="species" species="japonicus">
<emphasis id="84CD8C47A010D17EFDFCF8C4FC93F8AD" box="[637,776,1811,1835]" italics="true" pageId="7" pageNumber="178">A. japonicus</emphasis>
</taxonomicName>
(
<figureCitation id="2E824CD0A010D17EFC98F8C4FCC0F8AA" box="[793,859,1811,1836]" captionStart="FIGURE 5" captionStartId="9.[151,250,1805,1827]" captionTargetBox="[213,1387,381,1749]" captionTargetId="figure@9.[199,1399,375,1763]" captionTargetPageId="9" captionText="FIGURE 5. Alpheus japonicus Miers, 1879: male (13.9 mm) from Tokyo Bay, Japan (OUMNH. ZC. 2016 - 01 - 015): A, frontal region, dorsal; B, telson, dorsal; C, first article of antennular peduncle, lateral; D, major chela, mesial; E, minor chela, mesial; F, third pereiopod, dactylus, dorsal; G, second pleopod, detail of appendices masculina and interna, dorsal. For additional illustrations see Liu (1955), Kim (1977) and Banner &amp; Banner (1984)." httpUri="https://zenodo.org/record/271763/files/figure.png" pageId="7" pageNumber="178">Fig. 5</figureCitation>
D, E) and also differ by the stouter, less protruding dorsal and ventral shoulders on the palm of the major cheliped, and relatively shorter, distally more expanded dactylus of the minor cheliped. In addition, 
<taxonomicName id="71B92BD6A010D17EFD95F881FD70F8EB" box="[532,747,1877,1901]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="178" phylum="Arthropoda" rank="species" species="vladivostokiensis">
<emphasis id="84CD8C47A010D17EFD95F881FD70F8EB" box="[532,747,1877,1901]" italics="true" pageId="7" pageNumber="178">A. vladivostokiensis</emphasis>
</taxonomicName>
differs from 
<taxonomicName id="71B92BD6A010D17EFCFDF881FB9FF8EB" box="[892,1028,1878,1901]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="178" phylum="Arthropoda" rank="species" species="japonicus">
<emphasis id="84CD8C47A010D17EFCFDF881FB9FF8EB" box="[892,1028,1878,1901]" italics="true" pageId="7" pageNumber="178">A. japonicus</emphasis>
</taxonomicName>
by the olive-green colour (
<figureCitation id="2E824CD0A010D17EFAACF882FAE1F8E8" box="[1325,1402,1877,1902]" captionStart="FIGURE 3" captionStartId="7.[151,250,1413,1435]" captionTargetBox="[151,1435,176,1380]" captionTargetId="figure@7.[151,1436,154,1397]" captionTargetPageId="7" captionText="FIGURE 3. Alpheus vladivostokiensis (Vinogradov, 1950), colour pattern: neotype, male (cl 19.7 mm) from Troitza Bay (= Bukhta Troitzy), Russian Far East (MNHN-IU- 2014 - 5693), in dorsal (A) and lateral (B) views. Photographs by Ivan N. Marin." httpUri="https://zenodo.org/record/271761/files/figure.png" pageId="7" pageNumber="178">Figs. 3</figureCitation>
, 
<figureCitation id="2E824CD0A010D17EFA06F881FA0CF8E8" box="[1415,1431,1878,1902]" captionStart="FIGURE 4" captionStartId="8.[151,250,1581,1603]" captionTargetBox="[154,1434,176,1579]" captionTargetId="figure@8.[154,1434,164,1584]" captionTargetPageId="8" captionText="FIGURE 4. Alpheus vladivostokiensis (Vinogradov, 1950), colour pattern: ovigerous female (cl 16.3 mm) from Troitza Bay (= Bukhta Troitzy), Russian Far East (OUMNH. ZC. 2016 - 01 - 014), in dorsal (A) and lateral (B) views. Photographs by Ivan N. Marin." httpUri="https://zenodo.org/record/271762/files/figure.png" pageId="7" pageNumber="178">4</figureCitation>
; see also 
<bibRefCitation id="D2282DA4A010D17EFF78F8A1FE3EF809" author="Pasternak" box="[249,421,1910,1935]" pageId="7" pageNumber="178" refString="Pasternak, R. K. (1988) Life of animals [Zhizn' zhivotnykh], Vol. 2. Second edition. Prosvezscheniye, Moscow, 447 pp. [in Russian]" type="book" year="1988">Pasternak 1988</bibRefCitation>
; 
<bibRefCitation id="D2282DA4A010D17EFE35F8A1FDA1F809" author="Marin" box="[436,570,1910,1935]" pageId="7" pageNumber="178" refString="Marin, I. N. (2013) Small atlas of decapod crustaceans of Russia [Malyi atlas desiatinogikh rakoobraznykh Rossii]. KMK Scientific Press, Moscow, 146 pp. [in Russian]" type="book" year="2013">Marin 2013</bibRefCitation>
), which contrasts to the reddish colour (with a more conspicuous banding in younger individuals) of 
<taxonomicName id="71B92BD6A010D17EFE1FF84EFDBCF836" box="[414,551,1945,1968]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="178" phylum="Arthropoda" rank="species" species="japonicus">
<emphasis id="84CD8C47A010D17EFE1FF84EFDBCF836" box="[414,551,1945,1968]" italics="true" pageId="7" pageNumber="178">A. japonicus</emphasis>
</taxonomicName>
(
<figureCitation id="2E824CD0A010D17EFDB7F84FFDE2F837" box="[566,633,1944,1969]" captionStart="FIGURE 6" captionStartId="10.[151,250,1097,1119]" captionTargetBox="[151,1435,193,1074]" captionTargetId="figure@10.[151,1435,193,1075]" captionTargetPageId="10" captionText="FIGURE 6. Alpheus japonicus Miers, 1879, colour pattern: A, B, young male dredged at 50 – 60 m off Awaji Island, Japan (aquarium photograph, specimen not deposited); C, four adult males from a seafood market in Tokyo, Japan (OUMNH. ZC. 2016 - 01 - 015, MNHN-IU- 2014 - 5695, and MZUSP 34290); D, adult female from unknown locality in southern Korea. Photographs courtesy of Tomoyasu Tamego (A, B), Wataru Doi (C) and NFRDI (D)." httpUri="https://zenodo.org/record/271764/files/figure.png" pageId="7" pageNumber="178">Fig. 6</figureCitation>
; see also 
<bibRefCitation id="D2282DA4A010D17EFD62F84FFCEDF837" author="Miyake" box="[739,886,1944,1969]" pageId="7" pageNumber="178" refString="Miyake, S. (1982) Japanese crustacean decapods and stomatopods in colour. Vol. 1. Hoikusha, Osaka, 261 pp. [in Japanese]" type="book" year="1982">Miyake 1982</bibRefCitation>
; 
<bibRefCitation id="D2282DA4A010D17EFC02F84FFBA0F837" author="Baba" box="[899,1083,1944,1969]" pageId="7" pageNumber="178" refString="Baba, K., Hayashi, K. I. &amp; Toriyama, M. (1986) Decapod crustaceans from continental shelf and slope around Japan. Japan Fisheries Resource Conservation Association, Tokyo, 336 pp." type="book" year="1986">
Baba 
<emphasis id="84CD8C47A010D17EFC43F84EFC62F836" box="[962,1017,1944,1968]" italics="true" pageId="7" pageNumber="178">et al.</emphasis>
1986
</bibRefCitation>
). The ventromesial carina of the first article of the antennular peduncle exhibits some variation in both 
<taxonomicName id="71B92BD6A010D17EFC0CF86DFBFBF854" box="[909,1120,1978,2002]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="178" phylum="Arthropoda" rank="species" species="vladivostokiensis">
<emphasis id="84CD8C47A010D17EFC0CF86DFBFBF854" box="[909,1120,1978,2002]" italics="true" pageId="7" pageNumber="178">A. vladivostokiensis</emphasis>
</taxonomicName>
and 
<taxonomicName id="71B92BD6A010D17EFB13F86DFA8DF857" box="[1170,1302,1978,2001]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="178" phylum="Arthropoda" rank="species" species="japonicus">
<emphasis id="84CD8C47A010D17EFB13F86DFA8DF857" box="[1170,1302,1978,2001]" italics="true" pageId="7" pageNumber="178">A. japonicus</emphasis>
</taxonomicName>
, and may be either bluntly triangular (as in 
<figureCitation id="2E824CD0A010D17EFE58F80CFD88F875" box="[473,531,2011,2036]" captionStart="FIGURE 1" captionStartId="5.[151,250,1858,1880]" captionTargetBox="[153,1429,200,1831]" captionTargetId="figure@5.[153,1438,196,1840]" captionTargetPageId="5" captionText="FIGURE 1. Alpheus vladivostokiensis (Vinogradov, 1950): male (cl 18.0 mm) from Troitza Bay (= Bukhta Troitzy), Russian Far East (OUMNH. ZC. 2016 - 01 - 013): A, frontal region, dorsal; B, same, lateral; C, telson, dorsal; D, ventromesial carina of first article of antennular peduncle, lateral; E, third maxilliped, lateral; F, second pereiopod, lateral; G, third pereiopod, lateral; H, same, dactylus, dorsal; I, fifth pereiopod, lateral; J, same, propodus and dactylus, ventral; K, second pleopod, detail of appendices masculina and interna, dorsal; L uropod, dorsal." httpUri="https://zenodo.org/record/271759/files/figure.png" pageId="7" pageNumber="178">Fig. 1</figureCitation>
D) or with a sharp tooth (as in 
<figureCitation id="2E824CD0A010D17EFCDBF80CFC02F872" box="[858,921,2011,2036]" captionStart="FIGURE 5" captionStartId="9.[151,250,1805,1827]" captionTargetBox="[213,1387,381,1749]" captionTargetId="figure@9.[199,1399,375,1763]" captionTargetPageId="9" captionText="FIGURE 5. Alpheus japonicus Miers, 1879: male (13.9 mm) from Tokyo Bay, Japan (OUMNH. ZC. 2016 - 01 - 015): A, frontal region, dorsal; B, telson, dorsal; C, first article of antennular peduncle, lateral; D, major chela, mesial; E, minor chela, mesial; F, third pereiopod, dactylus, dorsal; G, second pleopod, detail of appendices masculina and interna, dorsal. For additional illustrations see Liu (1955), Kim (1977) and Banner &amp; Banner (1984)." httpUri="https://zenodo.org/record/271763/files/figure.png" pageId="7" pageNumber="178">Fig. 5</figureCitation>
C).
</paragraph>
<caption id="E2C600DDA01FD171FF16F9FAFF40F906" httpUri="https://zenodo.org/record/271762/files/figure.png" pageId="8" pageNumber="179" targetBox="[154,1434,176,1579]" targetPageId="8">
<paragraph id="B6065055A01FD171FF16F9FAFF40F906" blockId="8.[151,1436,1581,1664]" pageId="8" pageNumber="179">
<emphasis id="84CD8C47A01FD171FF16F9FAFE95F9C4" bold="true" box="[151,270,1581,1603]" pageId="8" pageNumber="179">FIGURE 4</emphasis>
. 
<taxonomicName id="71B92BD6A01FD171FE9DF9FAFD72F9C5" authority="Vinogradov, 1950" authorityName="Vinogradov" authorityYear="1950" box="[284,745,1581,1603]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="179" phylum="Arthropoda" rank="species" species="vladivostokiensis">
<emphasis id="84CD8C47A01FD171FE9DF9FAFD86F9C5" box="[284,541,1581,1603]" italics="true" pageId="8" pageNumber="179">Alpheus vladivostokiensis</emphasis>
(Vinogradov, 1950)
</taxonomicName>
, colour pattern: ovigerous female (cl 16.3 mm) from Troitza Bay (= Bukhta Troitzy), Russian Far East (OUMNH.ZC. 2016-01-014), in dorsal (A) and lateral (B) views. Photographs by Ivan N. Marin.
</paragraph>
</caption>
<paragraph id="B6065055A01FD171FF46F970FA07F859" blockId="8.[151,1436,1703,2016]" pageId="8" pageNumber="179">
The specimen from Vladivostok identified and reported by 
<bibRefCitation id="D2282DA4A01FD171FCDFF970FC77F946" author="Balss" box="[862,1004,1703,1728]" pageId="8" pageNumber="179" refString="Balss, H. (1914) Ostasiatischen Decapoden II. Die Natantia und Reptantia. Beitrage zur Naturgeschichte Ostasiens. Abhandlungen der Mathematisch-Physischen Klasse der Kayserlichen Bayerischen Akademie der Wissenschaften, II, Suppl. - Bd. 10, Abh., 1 - 101, 1 pl." type="book chapter" year="1914">Balss (1914)</bibRefCitation>
as 
<taxonomicName id="71B92BD6A01FD171FB90F97FFB06F939" box="[1041,1181,1704,1727]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="179" phylum="Arthropoda" rank="species" species="japonicus">
<emphasis id="84CD8C47A01FD171FB90F97FFB06F939" box="[1041,1181,1704,1727]" italics="true" pageId="8" pageNumber="179">A. japonicus</emphasis>
</taxonomicName>
(ZMMU Ma 608) was re-examined and confirmed as 
<taxonomicName id="71B92BD6A01FD171FE6EF91BFD57F962" box="[495,716,1740,1764]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="179" phylum="Arthropoda" rank="species" species="vladivostokiensis">
<emphasis id="84CD8C47A01FD171FE6EF91BFD57F962" box="[495,716,1740,1764]" italics="true" pageId="8" pageNumber="179">A. vladivostokiensis</emphasis>
</taxonomicName>
. The listing of 
<taxonomicName id="71B92BD6A01FD171FCF7F91BFB99F965" box="[886,1026,1740,1763]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="179" phylum="Arthropoda" rank="species" species="japonicus">
<emphasis id="84CD8C47A01FD171FCF7F91BFB99F965" box="[886,1026,1740,1763]" italics="true" pageId="8" pageNumber="179">A. japonicus</emphasis>
</taxonomicName>
from the Sea of 
<collectingCountry id="CEAE10C5A01FD171FB3BF91BFB61F965" box="[1210,1274,1740,1763]" name="Japan" pageId="8" pageNumber="179">Japan</collectingCountry>
in 
<bibRefCitation id="D2282DA4A01FD171FA9DF91CFEF1F88E" author="Derjugin" pageId="8" pageNumber="179" refString="Derjugin, K. M. &amp; Kobjakowa, S. (1935) Zur Dekapodenfauna des Japanischen Meeres. Zoologischer Anzeiger, 112, 141 - 147." type="journal article" year="1935">Derjugin &amp; Kobjakowa (1935)</bibRefCitation>
is problematic as it may refer to both 
<taxonomicName id="71B92BD6A01FD171FC99F927FC3FF881" box="[792,932,1776,1799]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="179" phylum="Arthropoda" rank="species" species="japonicus">
<emphasis id="84CD8C47A01FD171FC99F927FC3FF881" box="[792,932,1776,1799]" italics="true" pageId="8" pageNumber="179">A. japonicus</emphasis>
</taxonomicName>
and/or 
<taxonomicName id="71B92BD6A01FD171FC7AF927FB7FF88E" box="[1019,1252,1776,1800]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="179" phylum="Arthropoda" rank="species" species="vladisvostokiensis">
<emphasis id="84CD8C47A01FD171FC7AF927FB7FF88E" box="[1019,1252,1776,1800]" italics="true" pageId="8" pageNumber="179">A. vladisvostokiensis</emphasis>
</taxonomicName>
. As far as it can presently be ascertained, 
<taxonomicName id="71B92BD6A01FD171FE3EF8C3FDCBF8AD" box="[447,592,1812,1835]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="179" phylum="Arthropoda" rank="species" species="japonicus">
<emphasis id="84CD8C47A01FD171FE3EF8C3FDCBF8AD" box="[447,592,1812,1835]" italics="true" pageId="8" pageNumber="179">A. japonicus</emphasis>
</taxonomicName>
appears to be absent (or at least has yet not been confirmed) from the northwestern Sea of 
<collectingCountry id="CEAE10C5A01FD171FE0DF8EFFE57F8C9" box="[396,460,1848,1871]" name="Japan" pageId="8" pageNumber="179">Japan</collectingCountry>
(including the Russian Far East), occurring in the deeper near-shore waters of the southern Sea of 
<collectingCountry id="CEAE10C5A01FD171FEDDF88BFE07F8F5" box="[348,412,1884,1907]" name="Japan" pageId="8" pageNumber="179">Japan</collectingCountry>
off the coasts of 
<collectingCountry id="CEAE10C5A01FD171FDF6F88CFC93F8F5" box="[631,776,1883,1908]" name="South Korea" pageId="8" pageNumber="179">South Korea</collectingCountry>
and central-southern 
<collectingCountry id="CEAE10C5A01FD171FB8DF88BFBD7F8F5" box="[1036,1100,1884,1907]" name="Japan" pageId="8" pageNumber="179">Japan</collectingCountry>
(from southern and western Hokkaido to Kyushu), as well as in the Yellow and East 
<collectingCountry id="CEAE10C5A01FD171FC8DF8A8FCD4F81E" box="[780,847,1919,1944]" name="China" pageId="8" pageNumber="179">China</collectingCountry>
Seas and the Pacific coast of 
<collectingCountry id="CEAE10C5A01FD171FB1AF857FB40F811" box="[1179,1243,1920,1943]" name="Japan" pageId="8" pageNumber="179">Japan</collectingCountry>
(from Honshu to Kyushu) (
<figureCitation id="2E824CD0A01FD171FE89F874FED6F83A" box="[264,333,1955,1980]" captionStart="FIGURE 7" captionStartId="11.[151,250,1178,1200]" captionTargetBox="[151,1435,195,1156]" captionTargetId="figure@11.[151,1435,195,1156]" captionTargetPageId="11" captionText="FIGURE 7. Map of the eastern Asian seas showing the presently known distributions of Alpheus vladivostokiensis Vinogradov, 1950 (red circles) and A. japonicus Miers, 1879 (blue circles), compiled based on the material examined in the present study and previously published, confirmable records (e. g., Miers 1879; Spence Bate 1888; Ortmann 1890; Yokoya 1930; Kim 1977; Banner &amp; Banner 1984; Komai et al. 1992; Anker 2001; Cha et al. 2001; Xu et al. 2015)." httpUri="https://zenodo.org/record/271765/files/figure.png" pageId="8" pageNumber="179">Fig. 7</figureCitation>
). 
<taxonomicName id="71B92BD6A01FD171FEE5F873FD1EF83A" box="[356,645,1956,1980]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="179" phylum="Arthropoda" rank="species" species="vladivostokiensis">
<emphasis id="84CD8C47A01FD171FEE5F873FD1EF83A" box="[356,645,1956,1980]" italics="true" pageId="8" pageNumber="179">Alpheus vladivostokiensis</emphasis>
</taxonomicName>
, on the other hand, seems to be confined to the shallow waters of the northern Sea of 
<collectingCountry id="CEAE10C5A01FD171FEC9F81FFE10F859" box="[328,395,1992,2015]" name="Japan" pageId="8" pageNumber="179">Japan</collectingCountry>
, i.e. the Russian Far East and northern 
<collectingCountry id="CEAE10C5A01FD171FCBDF81FFCE7F859" box="[828,892,1992,2015]" name="Japan" pageId="8" pageNumber="179">Japan</collectingCountry>
(Hokkaido) (
<figureCitation id="2E824CD0A01FD171FB93F810FBCDF866" box="[1042,1110,1991,2016]" captionStart="FIGURE 7" captionStartId="11.[151,250,1178,1200]" captionTargetBox="[151,1435,195,1156]" captionTargetId="figure@11.[151,1435,195,1156]" captionTargetPageId="11" captionText="FIGURE 7. Map of the eastern Asian seas showing the presently known distributions of Alpheus vladivostokiensis Vinogradov, 1950 (red circles) and A. japonicus Miers, 1879 (blue circles), compiled based on the material examined in the present study and previously published, confirmable records (e. g., Miers 1879; Spence Bate 1888; Ortmann 1890; Yokoya 1930; Kim 1977; Banner &amp; Banner 1984; Komai et al. 1992; Anker 2001; Cha et al. 2001; Xu et al. 2015)." httpUri="https://zenodo.org/record/271765/files/figure.png" pageId="8" pageNumber="179">Fig. 7</figureCitation>
). 
<bibRefCitation id="D2282DA4A01FD171FBEBF810FA8EF866" author="Igarashi" box="[1130,1301,1991,2016]" pageId="8" pageNumber="179" refString="Igarashi, T. (1969) A list of marine decapod crustaceans from Hokkaido, deposited at the Fisheries Museum, Faculty of Fisheries, Hokkaido University I. Macrura. Contribution from the Fisheries Museum, Faculty of Fisheries, Hokkaido University, 11, 1 - 15, pls. 1 - 20." type="journal article" year="1969">Igarashi (1969)</bibRefCitation>
recorded 
<emphasis id="84CD8C47A01FD171FA05F81FFA07F859" box="[1412,1436,1992,2015]" italics="true" pageId="8" pageNumber="179">A.</emphasis>
</paragraph>
<paragraph id="B6065055A01ED170FF16FF4FFC89FEC6" blockId="9.[151,1436,151,320]" pageId="9" pageNumber="180">
<taxonomicName id="71B92BD6A01ED170FF16FF4FFE9FFF29" box="[151,260,152,175]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="180" phylum="Arthropoda" rank="species" species="japonicus">
<emphasis id="84CD8C47A01ED170FF16FF4FFE9FFF29" box="[151,260,152,175]" italics="true" pageId="9" pageNumber="180">japonicus</emphasis>
</taxonomicName>
from Hokkaido on the basis of material from Oshoro Bay and Ishikari Bay, both on the Sea of 
<collectingCountry id="CEAE10C5A01ED170FAA9FF4FFAF3FF29" box="[1320,1384,152,175]" name="Japan" pageId="9" pageNumber="180">Japan</collectingCountry>
side of Hokkaido. The photograph of a specimen of 
<taxonomicName id="71B92BD6A01ED170FD35FF6BFCDAFF52" box="[692,833,188,212]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="180" phylum="Arthropoda" rank="species" species="japonicus">
<emphasis id="84CD8C47A01ED170FD35FF6BFCDAFF52" box="[692,833,188,212]" italics="true" pageId="9" pageNumber="180">A. japonicus</emphasis>
</taxonomicName>
(with slender chelipeds) provided by 
<bibRefCitation id="D2282DA4A01ED170FB6EFF6BFA07FF53" author="Igarashi" box="[1263,1436,188,213]" pageId="9" pageNumber="180" refString="Igarashi, T. (1969) A list of marine decapod crustaceans from Hokkaido, deposited at the Fisheries Museum, Faculty of Fisheries, Hokkaido University I. Macrura. Contribution from the Fisheries Museum, Faculty of Fisheries, Hokkaido University, 11, 1 - 15, pls. 1 - 20." type="journal article" year="1969">Igarashi (1969)</bibRefCitation>
clearly shows that his identification was correct. The listing of 
<taxonomicName id="71B92BD6A01ED170FCDAFF37FC7CFF71" box="[859,999,224,247]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="180" phylum="Arthropoda" rank="species" species="japonicus">
<emphasis id="84CD8C47A01ED170FCDAFF37FC7CFF71" box="[859,999,224,247]" italics="true" pageId="9" pageNumber="180">A. japonicus</emphasis>
</taxonomicName>
from Hokkaido in Komai 
<emphasis id="84CD8C47A01ED170FA93FF36FAD1FF7E" box="[1298,1354,224,248]" italics="true" pageId="9" pageNumber="180">et al.</emphasis>
(1992) was based on Igarashi’s (1969) record. Thus, both 
<taxonomicName id="71B92BD6A01ED170FD60FED3FCEBFE9A" box="[737,880,260,284]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="180" phylum="Arthropoda" rank="species" species="japonicus">
<emphasis id="84CD8C47A01ED170FD60FED3FCEBFE9A" box="[737,880,260,284]" italics="true" pageId="9" pageNumber="180">A. japonicus</emphasis>
</taxonomicName>
and 
<taxonomicName id="71B92BD6A01ED170FC2DFED3FB17FE9A" box="[940,1164,260,284]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="180" phylum="Arthropoda" rank="species" species="vladivostokiensis">
<emphasis id="84CD8C47A01ED170FC2DFED3FB17FE9A" box="[940,1164,260,284]" italics="true" pageId="9" pageNumber="180">A. vladivostokiensis</emphasis>
</taxonomicName>
are present in southern Hokkaido, however, appear to be ecologically separated.
</paragraph>
<caption id="E2C600DDA01ED170FF16F8DAFCFBF8F9" httpUri="https://zenodo.org/record/271763/files/figure.png" pageId="9" pageNumber="180" targetBox="[213,1387,381,1749]" targetPageId="9">
<paragraph id="B6065055A01ED170FF16F8DAFCFBF8F9" blockId="9.[151,1436,1805,1919]" pageId="9" pageNumber="180">
<emphasis id="84CD8C47A01ED170FF16F8DAFE96F8A5" bold="true" box="[151,269,1805,1827]" pageId="9" pageNumber="180">FIGURE 5</emphasis>
. 
<taxonomicName id="71B92BD6A01ED170FE9BF8DAFDC1F8A5" authority="Miers, 1879" authorityName="Miers" authorityYear="1879" box="[282,602,1805,1827]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="180" phylum="Arthropoda" rank="species" species="japonicus">
<emphasis id="84CD8C47A01ED170FE9BF8DAFE4DF8A5" box="[282,470,1805,1827]" italics="true" pageId="9" pageNumber="180">Alpheus japonicus</emphasis>
Miers, 1879
</taxonomicName>
: male (13.9 mm) from Tokyo Bay, Japan (OUMNH.ZC. 2016-01-015): A, frontal region, dorsal; B, telson, dorsal; C, first article of antennular peduncle, lateral; D, major chela, mesial; E, minor chela, mesial; F, third pereiopod, dactylus, dorsal; G, second pleopod, detail of appendices masculina and interna, dorsal. For additional illustrations see Liu (1955), Kim (1977) and Banner &amp; Banner (1984).
</paragraph>
</caption>
<caption id="E2C600DDA01DD173FF16FB9EFC6AFB3C" httpUri="https://zenodo.org/record/271764/files/figure.png" pageId="10" pageNumber="181" targetBox="[151,1435,193,1074]" targetPageId="10">
<paragraph id="B6065055A01DD173FF16FB9EFC6AFB3C" blockId="10.[151,1437,1097,1210]" pageId="10" pageNumber="181">
<emphasis id="84CD8C47A01DD173FF16FB9EFE94FBD8" bold="true" box="[151,271,1097,1119]" pageId="10" pageNumber="181">FIGURE 6</emphasis>
. 
<taxonomicName id="71B92BD6A01DD173FE9FFB9EFDF9FBD9" authority="Miers, 1879" authorityName="Miers" authorityYear="1879" box="[286,610,1097,1119]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="181" phylum="Arthropoda" rank="species" species="japonicus">
<emphasis id="84CD8C47A01DD173FE9FFB9EFE40FBD9" box="[286,475,1097,1119]" italics="true" pageId="10" pageNumber="181">Alpheus japonicus</emphasis>
Miers, 1879
</taxonomicName>
, colour pattern: A, B, young male dredged at 50–60 m off Awaji Island, Japan (aquarium photograph, specimen not deposited); C, four adult males from a seafood market in Tokyo, Japan (OUMNH.ZC. 2016-01-015, MNHN-IU-2014-5695, and MZUSP 34290); D, adult female from unknown locality in southern Korea. Photographs courtesy of Tomoyasu Tamego (A, B), Wataru Doi (C) and NFRDI (D).
</paragraph>
</caption>
<paragraph id="B6065055A01DD173FF46FB30FDA9F8C6" blockId="10.[151,1436,1255,1856]" pageId="10" pageNumber="181">
The somewhat confusing synonymy of 
<taxonomicName id="71B92BD6A01DD173FD04FB3FFC89FB79" box="[645,786,1256,1279]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="181" phylum="Arthropoda" rank="species" species="japonicus">
<emphasis id="84CD8C47A01DD173FD04FB3FFC89FB79" box="[645,786,1256,1279]" italics="true" pageId="10" pageNumber="181">A. japonicus</emphasis>
</taxonomicName>
was briefly reviewed by 
<bibRefCitation id="D2282DA4A01DD173FBB0FB3FFAD6FA86" author="Banner" box="[1073,1357,1256,1280]" pageId="10" pageNumber="181" refString="Banner, D. M. &amp; Banner, A. H. (1984) Old and unreported collections of alpheid shrimp from the Zoologisches Museum Berlin, principally from Melanesia. Pacific Science, 38, 34 - 50." type="journal article" year="1984">Banner &amp; Banner (1984)</bibRefCitation>
. These authors found some, possibly maturity-related variation, in the development of the balaeniceps setae on the minor cheliped. The earlier suggested synonymy of 
<taxonomicName id="71B92BD6A01DD173FD16FAE7FB88FACE" authority="Spence Bate, 1888" authorityName="Spence Bate" authorityYear="1888" box="[663,1043,1327,1352]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="181" phylum="Arthropoda" rank="species" species="longimanus">
<emphasis id="84CD8C47A01DD173FD16FAE7FCA1FAC1" box="[663,826,1327,1351]" italics="true" pageId="10" pageNumber="181">A. longimanus</emphasis>
<bibRefCitation id="D2282DA4A01DD173FCC0FAE7FB88FACE" author="Spence" box="[833,1043,1328,1352]" pageId="10" pageNumber="181" refString="Spence Bate, C. (1888) Report on the Crustacea Macrura dredged by H. M. S. Challenger during the years 1873 - 76. The voyage of H. M. S. Challenger, Zoology, 24, xi + 942 pp., 157 pls. (separate volume), Eyre &amp; Spottiswoode, London." type="book" year="1888">Spence Bate, 1888</bibRefCitation>
</taxonomicName>
with 
<taxonomicName id="71B92BD6A01DD173FBD5FAE7FB7AFAC1" box="[1108,1249,1328,1351]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="181" phylum="Arthropoda" rank="species" species="japonicus">
<emphasis id="84CD8C47A01DD173FBD5FAE7FB7AFAC1" box="[1108,1249,1328,1351]" italics="true" pageId="10" pageNumber="181">A. japonicus</emphasis>
</taxonomicName>
(
<bibRefCitation id="D2282DA4A01DD173FB6EFAE7FA0CFACE" author="Ortmann" box="[1263,1431,1328,1352]" pageId="10" pageNumber="181" refString="Ortmann, A. (1890) Die Unterordnung Natantia Boas. Theil I. Die Decapoden Krebse des Strassburger Museums, mit besonderer Berucksichtigung der von Herrn Dr. Doderlein bei Japan und bei den Liu-Kiu-Inseln gesammelten und z. Z. im Strassburger Museum aufbewahrten Formen. Zoologische Jahrbucher, Systematik, 5, 437 - 542, pls. 36, 37." type="journal article" year="1890">Ortmann 1890</bibRefCitation>
; 
<bibRefCitation id="D2282DA4A01DD173FF16FA83FEADFAEA" author="De" box="[151,310,1364,1388]" pageId="10" pageNumber="181" refString="De Man, J. G. (1907) On a collection of Crustacea, Decapoda and Stomatopoda, chiefly from the Inland Sea of Japan; with descriptions of new species. Transactions of the Linnean Society of London, II, 9, 387 - 454, pls. 19 - 27." type="journal article" year="1907">De Man 1907</bibRefCitation>
) was confirmed, although one of Spence Bate’s specimens from 
<collectingCountry id="CEAE10C5A01DD173FBA4FA84FBC4FAEA" box="[1061,1119,1363,1388]" name="India" pageId="10" pageNumber="181">India</collectingCountry>
was considered to possibly belong to a different species, more closely related to 
<taxonomicName id="71B92BD6A01DD173FD72FAAFFBD3FA16" authority="Audouin, 1826" authorityName="Audouin" authorityYear="1826" box="[755,1096,1399,1424]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="181" phylum="Arthropoda" rank="species" species="edwardsii">
<emphasis id="84CD8C47A01DD173FD72FAAFFC1AFA09" box="[755,897,1399,1423]" italics="true" pageId="10" pageNumber="181">A. edwardsii</emphasis>
(
<bibRefCitation id="D2282DA4A01DD173FC10FAA0FBDBFA16" author="Audouin" box="[913,1088,1399,1424]" pageId="10" pageNumber="181" refString="Audouin, J. V. (1826) Explication sommaire des planches de crustaces de l'Egypte et de la Syrie, publiees par Jules-Cesar Savigny, Membre de l'Institut; offrant un expose des caracteres naturels des genres, avec la distinction des especes. In: Savigny, J. - C. (Ed.), Description de l'Egypte, Histoire Naturelle, 1 (4), 77 - 98." type="journal article" year="1826">Audouin, 1826</bibRefCitation>
)
</taxonomicName>
. Part of the Japanese material identified as 
<taxonomicName id="71B92BD6A01DD173FEAAFA4BFDD3FA32" authority="Ortmann, 1890" authorityName="Ortmann" authorityYear="1890" box="[299,584,1436,1460]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="181" phylum="Arthropoda" rank="species" species="haanii">
<emphasis id="84CD8C47A01DD173FEAAFA4BFE0FFA32" box="[299,404,1436,1460]" italics="true" pageId="10" pageNumber="181">A. haanii</emphasis>
<bibRefCitation id="D2282DA4A01DD173FE1DFA4BFDD3FA32" author="Ortmann" box="[412,584,1436,1460]" pageId="10" pageNumber="181" refString="Ortmann, A. (1890) Die Unterordnung Natantia Boas. Theil I. Die Decapoden Krebse des Strassburger Museums, mit besonderer Berucksichtigung der von Herrn Dr. Doderlein bei Japan und bei den Liu-Kiu-Inseln gesammelten und z. Z. im Strassburger Museum aufbewahrten Formen. Zoologische Jahrbucher, Systematik, 5, 437 - 542, pls. 36, 37." type="journal article" year="1890">Ortmann, 1890</bibRefCitation>
</taxonomicName>
by 
<bibRefCitation id="D2282DA4A01DD173FDF4FA4CFC84FA32" author="Yokoya" box="[629,799,1435,1460]" pageId="10" pageNumber="181" refString="Yokoya, Y. (1939) Macrura and Anomura of decapod Crustacea found in the neighborhood of Onogawa, Miyagi-ken. Scientific Reports of the Tohoku Imperial University, series IV, 4, 261 - 289." type="journal article" year="1939">Yokoya (1939)</bibRefCitation>
was tentatively re-assigned to 
<taxonomicName id="71B92BD6A01DD173FB03FA4BFA8BFA35" box="[1154,1296,1436,1459]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="181" phylum="Arthropoda" rank="species" species="japonicus">
<emphasis id="84CD8C47A01DD173FB03FA4BFA8BFA35" box="[1154,1296,1436,1459]" italics="true" pageId="10" pageNumber="181">A. japonicus</emphasis>
</taxonomicName>
, whereas 
<taxonomicName id="71B92BD6A01DD173FA05FA4BFF44FA51" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="181" phylum="Arthropoda" rank="species" species="haanii">
<emphasis id="84CD8C47A01DD173FA05FA4BFF44FA51" italics="true" pageId="10" pageNumber="181">A. haanii</emphasis>
</taxonomicName>
was considered as a possible synonym of 
<taxonomicName id="71B92BD6A01DD173FD44FA17FCC9FA51" box="[709,850,1471,1495]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="181" phylum="Arthropoda" rank="species" species="edwardsii">
<emphasis id="84CD8C47A01DD173FD44FA17FCC9FA51" box="[709,850,1471,1495]" italics="true" pageId="10" pageNumber="181">A. edwardsii</emphasis>
</taxonomicName>
. However, since the latter species is clearly a large species complex (Anker &amp; De Grave, in press), the taxonomic status of 
<taxonomicName id="71B92BD6A01DD173FC68FA33FBC8FA7A" box="[1001,1107,1508,1532]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="181" phylum="Arthropoda" rank="species" species="haanii">
<emphasis id="84CD8C47A01DD173FC68FA33FBC8FA7A" box="[1001,1107,1508,1532]" italics="true" pageId="10" pageNumber="181">A. haanii</emphasis>
</taxonomicName>
remains unsettled. 
<bibRefCitation id="D2282DA4A01DD173FAB9FA33FF7AF9A6" author="Ortmann" pageId="10" pageNumber="181" refString="Ortmann, A. (1890) Die Unterordnung Natantia Boas. Theil I. Die Decapoden Krebse des Strassburger Museums, mit besonderer Berucksichtigung der von Herrn Dr. Doderlein bei Japan und bei den Liu-Kiu-Inseln gesammelten und z. Z. im Strassburger Museum aufbewahrten Formen. Zoologische Jahrbucher, Systematik, 5, 437 - 542, pls. 36, 37." type="journal article" year="1890">Ortmann (1890)</bibRefCitation>
based the description of 
<taxonomicName id="71B92BD6A01DD173FD8DF9DFFDEDF999" box="[524,630,1543,1567]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="181" phylum="Arthropoda" rank="species" species="haanii">
<emphasis id="84CD8C47A01DD173FD8DF9DFFDEDF999" box="[524,630,1543,1567]" italics="true" pageId="10" pageNumber="181">A. haanii</emphasis>
</taxonomicName>
on two lots, each with three specimens, from two different Japanese localities, one from Tokyo Bay and one from Kagoshima. 
<bibRefCitation id="D2282DA4A01DD173FCD3F9FCFC6EF9C2" author="Komai" box="[850,1013,1579,1604]" pageId="10" pageNumber="181" refString="Komai, T. (1999) Decapod Crustacea collected by L. Doderlein in Japan and reported by Ortmann (1890 - 1894) in the collection of the Musee Zoologique, Strasbourg. In: Nishikawa, T. (Ed.), Preliminary Taxonomic and Historical Studies on Prof. Ludwig Doderlein's collection of Japanese Animals made in 1880 - 81 and deposited at Several European Museum. Report on Activities in 1997 - 8 supported by Grant-in-Aid for International Scientific Research (Field Research) No. 0 9041155, pp. 53 - 101." type="journal article" year="1999">Komai (1999)</bibRefCitation>
re-examined Ortmann’s Kagoshima material (MZS 69) and found that one of the three specimens in this lot represented 
<taxonomicName id="71B92BD6A01DD173FBD2F987FA02F9EE" authority="De Haan, 1844" authorityName="De Haan" authorityYear="1844" box="[1107,1433,1615,1640]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="181" phylum="Arthropoda" rank="species" species="bisincisus">
<emphasis id="84CD8C47A01DD173FBD2F987FB79F9E1" box="[1107,1250,1615,1639]" italics="true" pageId="10" pageNumber="181">A. bisincisus</emphasis>
<bibRefCitation id="D2282DA4A01DD173FB6BF987FA02F9EE" author="De" box="[1258,1433,1616,1640]" pageId="10" pageNumber="181" refString="De Haan, W. (1844) [1833 - 1850] Crustacea. In: von Siebold, P. F. (Ed.), Fauna Japonica sive Descriptio Animalium, quae in Itinere per Japoniam, Jussu et Auspiciis Superiorum, qui Summum in India Batava Imperium Tenent, Suspecto, Annis 1823 - 1830 Collegit, Notis, Observationibus et Adumbrationibus Illustravit: i - xxxi, ix - xvi, 1 - 243, pls. A - J, L - Q, 1 - 55. Lugduni-Batavorum." type="book chapter" year="1844">De Haan, 1844</bibRefCitation>
</taxonomicName>
. The other two specimens could be identified only as an unidentifiable species of the 
<taxonomicName id="71B92BD6A01DD173FBE2F9A3FB6AF90A" box="[1123,1265,1652,1676]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="181" phylum="Arthropoda" rank="species" species="edwardsii">
<emphasis id="84CD8C47A01DD173FBE2F9A3FB6AF90A" box="[1123,1265,1652,1676]" italics="true" pageId="10" pageNumber="181">A. edwardsii</emphasis>
</taxonomicName>
species group, morphologically similar to 
<taxonomicName id="71B92BD6A01DD173FE56F94FFC87F936" authority="De Haan, 1844" authorityName="De Haan" authorityYear="1844" box="[471,796,1687,1712]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="181" phylum="Arthropoda" rank="species" species="lobidens">
<emphasis id="84CD8C47A01DD173FE56F94FFDC0F929" box="[471,603,1687,1711]" italics="true" pageId="10" pageNumber="181">A. lobidens</emphasis>
<bibRefCitation id="D2282DA4A01DD173FDE7F94FFC87F936" author="De" box="[614,796,1688,1712]" pageId="10" pageNumber="181" refString="De Haan, W. (1844) [1833 - 1850] Crustacea. In: von Siebold, P. F. (Ed.), Fauna Japonica sive Descriptio Animalium, quae in Itinere per Japoniam, Jussu et Auspiciis Superiorum, qui Summum in India Batava Imperium Tenent, Suspecto, Annis 1823 - 1830 Collegit, Notis, Observationibus et Adumbrationibus Illustravit: i - xxxi, ix - xvi, 1 - 243, pls. A - J, L - Q, 1 - 55. Lugduni-Batavorum." type="book chapter" year="1844">De Haan, 1844</bibRefCitation>
</taxonomicName>
. On the other side, we concur with the conclusion of 
<bibRefCitation id="D2282DA4A01DD173FF16F96BFED5F952" author="Ortmann" box="[151,334,1724,1748]" pageId="10" pageNumber="181" refString="Ortmann, A. (1890) Die Unterordnung Natantia Boas. Theil I. Die Decapoden Krebse des Strassburger Museums, mit besonderer Berucksichtigung der von Herrn Dr. Doderlein bei Japan und bei den Liu-Kiu-Inseln gesammelten und z. Z. im Strassburger Museum aufbewahrten Formen. Zoologische Jahrbucher, Systematik, 5, 437 - 542, pls. 36, 37." type="journal article" year="1890">Ortmann (1890)</bibRefCitation>
, 
<bibRefCitation id="D2282DA4A01DD173FEDAF96BFD9CF952" box="[347,519,1724,1748]" pageId="10" pageNumber="181" refString="De Man, J. G. (1907) On a collection of Crustacea, Decapoda and Stomatopoda, chiefly from the Inland Sea of Japan; with descriptions of new species. Transactions of the Linnean Society of London, II, 9, 387 - 454, pls. 19 - 27." type="journal article">De Man (1907)</bibRefCitation>
and 
<bibRefCitation id="D2282DA4A01DD173FDBFF96BFCCDF952" author="Banner" box="[574,854,1724,1748]" pageId="10" pageNumber="181" refString="Banner, D. M. &amp; Banner, A. H. (1984) Old and unreported collections of alpheid shrimp from the Zoologisches Museum Berlin, principally from Melanesia. Pacific Science, 38, 34 - 50." type="journal article" year="1984">Banner &amp; Banner (1984)</bibRefCitation>
on the status of 
<taxonomicName id="71B92BD6A01DD173FB91F96BFB29F952" box="[1040,1202,1724,1748]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="181" phylum="Arthropoda" rank="species" species="longimanus">
<emphasis id="84CD8C47A01DD173FB91F96BFB29F952" box="[1040,1202,1724,1748]" italics="true" pageId="10" pageNumber="181">A. longimanus</emphasis>
</taxonomicName>
as a junior synonym of 
<emphasis id="84CD8C47A01DD173FF35F937FEEFF971" box="[180,372,1760,1783]" italics="true" pageId="10" pageNumber="181">
<taxonomicName id="71B92BD6A01DD173FF35F937FEDBF971" box="[180,320,1760,1783]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="181" phylum="Arthropoda" rank="species" sensu="stricto" species="japonicus">A. japonicus</taxonomicName>
<taxonomicNameLabel id="9FFE313CA01DD173FEC7F937FEEFF971" box="[326,372,1760,1783]" pageId="10" pageNumber="181" sensu="stricto">s.str</taxonomicNameLabel>
</emphasis>
. With 
<taxonomicName id="71B92BD6A01DD173FE41F937FDBEF971" box="[448,549,1759,1783]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="181" phylum="Arthropoda" rank="species" species="haanii">
<emphasis id="84CD8C47A01DD173FE41F937FDBEF971" box="[448,549,1759,1783]" italics="true" pageId="10" pageNumber="181">A. haanii</emphasis>
</taxonomicName>
ruled out and with no other available name for a taxon morphologically similar to 
<taxonomicName id="71B92BD6A01DD173FF37F8D3FEDEF89D" box="[182,325,1796,1819]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="181" phylum="Arthropoda" rank="species" species="japonicus">
<emphasis id="84CD8C47A01DD173FF37F8D3FEDEF89D" box="[182,325,1796,1819]" italics="true" pageId="10" pageNumber="181">A. japonicus</emphasis>
</taxonomicName>
, the application of the name 
<taxonomicName id="71B92BD6A01DD173FD1DF8D3FCE7F89A" box="[668,892,1796,1820]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="181" phylum="Arthropoda" rank="species" species="vladivostokiensis">
<emphasis id="84CD8C47A01DD173FD1DF8D3FCE7F89A" box="[668,892,1796,1820]" italics="true" pageId="10" pageNumber="181">A. vladivostokiensis</emphasis>
</taxonomicName>
for 
<taxonomicName id="71B92BD6A01DD173FC30F8D3FAF1F89A" authority="Vinogradov (1950)" authorityName="Vinogradov" authorityYear="1950" box="[945,1386,1795,1820]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="181" phylum="Arthropoda" rank="species" species="japonicus">
<emphasis id="84CD8C47A01DD173FC30F8D3FB1CF89D" box="[945,1159,1796,1819]" italics="true" pageId="10" pageNumber="181">A. japonicus sensu</emphasis>
<bibRefCitation id="D2282DA4A01DD173FB10F8D4FAF1F89A" author="Vinogradov" box="[1169,1386,1795,1820]" pageId="10" pageNumber="181" refString="Vinogradov, L. G. (1950) Keys to the shrimps, crayfishes and crabs of the Far East [Opredelitel' krevetok, rakov i krabov Dal'nego Vostoka]. Izvestia TINRO, 33 C, 179 - 350 [in Russian]" type="journal article" year="1950">Vinogradov (1950)</bibRefCitation>
</taxonomicName>
and 
<bibRefCitation id="D2282DA4A01DD173FF16F8F0FEB6F8C6" author="Marin" box="[151,301,1831,1856]" pageId="10" pageNumber="181" refString="Marin, I. N. (2013) Small atlas of decapod crustaceans of Russia [Malyi atlas desiatinogikh rakoobraznykh Rossii]. KMK Scientific Press, Moscow, 146 pp. [in Russian]" type="book" year="2013">Marin (2013)</bibRefCitation>
is now well supported.
</paragraph>
</subSubSection>
</treatment>
</document>