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<mods:title id="E88DCEA83D96C1EDDD55D9E97E8F83D7">A taxonomic revision of the genus Thalia Blumenbach, 1798; Weelia Yount, 1954; Brooksia Metcalf, 1918 (Salpida: Salpidae) from East Coast of Peninsular Malaysia</mods:title>
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<mods:namePart id="CB49E1A65025B52333749185E62BD98F">Ishak, Nurul Huda Binti Ahmad</mods:namePart>
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<mods:namePart id="4C098A02FFA43FDBBA6C71EA439B4933"> Aliah, Nur</mods:namePart>
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<treatment id="CF1C87B4FFF2452CFF77F904FDD7FEB9" ID-DOI="http://doi.org/10.5281/zenodo.5951778" ID-GBIF-Taxon="144128880" ID-Zenodo-Dep="5951778" LSID="urn:lsid:plazi:treatment:CF1C87B4FFF2452CFF77F904FDD7FEB9" httpUri="http://treatment.plazi.org/id/CF1C87B4FFF2452CFF77F904FDD7FEB9" lastPageId="14" lastPageNumber="465" pageId="8" pageNumber="459">
<subSubSection id="0FAF6529FFF2452AFF77F904FDB3FFBC" box="[151,553,1715,1741]" pageId="8" pageNumber="459" type="nomenclature">
<paragraph id="470A36A2FFF2452AFF77F904FDB3FFBC" blockId="8.[151,553,1715,1741]" box="[151,553,1715,1741]" pageId="8" pageNumber="459">
<heading id="1C4281CEFFF2452AFF77F904FDB3FFBC" bold="true" box="[151,553,1715,1741]" fontSize="11" level="1" pageId="8" pageNumber="459" reason="1">
<emphasis id="75C1EAB0FFF2452AFF77F904FDB3FFBC" bold="true" box="[151,553,1715,1741]" pageId="8" pageNumber="459">
3.
<taxonomicName id="80B54D21FFF2452AFF54F904FDB3FFBC" ID-CoL="5BW74" authority="Cuvier, 1804" authorityName="Cuvier" authorityYear="1804" box="[180,553,1715,1741]" class="Thaliacea" family="Salpidae" genus="Weelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Salpida" pageId="8" pageNumber="459" phylum="Chordata" rank="species" species="cylindrica">
<emphasis id="75C1EAB0FFF2452AFF54F904FE1FFFBC" bold="true" box="[180,389,1715,1741]" italics="true" pageId="8" pageNumber="459">Weelia cylindrica</emphasis>
Cuvier, 1804
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="0FAF6529FFF2452AFF77F94DFD4BFE1D" pageId="8" pageNumber="459" type="reference_group">
<paragraph id="470A36A2FFF2452AFF77F94DFD94FE5F" blockId="8.[151,1436,1786,1930]" pageId="8" pageNumber="459">
<emphasis id="75C1EAB0FFF2452AFF77F94DFF4BFE61" box="[151,209,1786,1808]" italics="true" pageId="8" pageNumber="459">Salpa</emphasis>
<taxonomicName id="80B54D21FFF2452AFF3DF94DFE40FE61" ID-CoL="5BW74" authority="CUvIER, 1804" authorityName="Cuvier" authorityYear="1804" box="[221,474,1786,1808]" class="Thaliacea" family="Salpidae" genus="Weelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Salpida" pageId="8" pageNumber="459" phylum="Chordata" rank="species" species="cylindrica">
<emphasis id="75C1EAB0FFF2452AFF3DF94DFEDEFE61" box="[221,324,1786,1808]" italics="true" pageId="8" pageNumber="459">cylindrica</emphasis>
Cuvier, 1804
</taxonomicName>
(cited from van
<bibRefCitation id="23244B53FFF2452AFD7CF94DFC8CFE61" box="[668,790,1786,1808]" pageId="8" pageNumber="459" refId="ref12791" refString="VAN SOEST, R. W. M. (1973 A) THE GENUS Thalia BlUmENbACH, 1798 (TUNICATA, THAlIACEA) WITH DESCRIPTIONS OF TWO NEW SPECIES. Beaufortia, 20 (271), 193 - 212." type="journal article">Soest 1973</bibRefCitation>
); (
<bibRefCitation id="23244B53FFF2452AFCD3F94DFBB4FE61" box="[819,1070,1786,1808]" pageId="8" pageNumber="459" refId="ref11848" refString="RITTER, W. E. &amp; BYXbEE, E. S. (1905) REPORTS ON THE SCIENTIFIC RESUlTS OF THE EXPEDITION TO THE TROPICAl PACIFIC IN THE AlbATROSS VIII. THE PElAGIC TUNICATA. Memoirs of the Museum for Comparative Zoology Harvard, 26 (5), 195 - 214." type="journal article">Ritter &amp; Byxbee 1905</bibRefCitation>
); (Survey 1918); (
<bibRefCitation id="23244B53FFF2452AFB17F94DFA0AFE61" box="[1271,1424,1786,1808]" pageId="8" pageNumber="459" refId="ref12476" refString="TOkIOkA, T. (1937) NOTES ON SAlPAS AND DOlIOlUmS OCCURRING IN THE PACIFIC COAST OF MIDDlE JAPAN. The Zoological Society of Japan, 16 (3), 219 - 235." type="journal article">Tokioka 1937</bibRefCitation>
);
<bibRefCitation id="23244B53FFF2452AFF27F8AFFEF5FE5F" box="[199,367,1816,1838]" pageId="8" pageNumber="459" refId="ref12451" refString="THOmPSON, H. (1948) Pelagic tunicates of Australia. COmmONWEAlTH COUNCIl FOR SCIENTIFIC AND INDUSTRIAl RESEARCH, MElbOURNE, 196 PP." type="book">Thompson 1948</bibRefCitation>
; Godeaux 1998
</paragraph>
<paragraph id="470A36A2FFF2452AFF77F88FFB1BFE3C" blockId="8.[151,1436,1786,1930]" box="[151,1153,1847,1869]" pageId="8" pageNumber="459">
<taxonomicName id="80B54D21FFF2452AFF77F88FFE51FE3C" ID-CoL="3P727" authority="Savigny, 1816" authorityName="Savigny" authorityYear="1816" baseAuthorityName="Cuvier" baseAuthorityYear="1804" box="[151,459,1847,1869]" class="Thaliacea" family="Salpidae" genus="Iasis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Salpida" pageId="8" pageNumber="459" phylum="Chordata" rank="species" species="cylindrica">
<emphasis id="75C1EAB0FFF2452AFF77F88FFEAEFE3C" box="[151,308,1847,1869]" italics="true" pageId="8" pageNumber="459">Iasis cylindrica</emphasis>
Savigny, 1816
</taxonomicName>
(cited by
<bibRefCitation id="23244B53FFF2452AFDD0F880FD01FE3C" box="[560,667,1847,1869]" pageId="8" pageNumber="459" refId="ref10785" refString="KOTT, P. (2005) Catalogue of Tunicata in Australian waters. AUSTRAlIAN BIOlOGICAl RESOURCES STUDY, DEPARTmENT OF THE ENvIRONmENT AND HERITAGE, CANbERRA, 301 PP." type="book">Kott 2005</bibRefCitation>
);
<bibRefCitation id="23244B53FFF2452AFD4CF880FCFDFE3C" box="[684,871,1847,1869]" pageId="8" pageNumber="459" refId="ref9367" refString="FRANCO, P., DAHmS, H. U., LO, W. T. &amp; HWANG, J. S. (2017): PElAGIC TUNICATES IN THE CHINA SEAS, Journal of Natural History, 51, 917 - 936." type="journal article">
Franco
<emphasis id="75C1EAB0FFF2452AFD19F88FFCB6FE3C" box="[761,812,1847,1869]" italics="true" pageId="8" pageNumber="459">et al.</emphasis>
2017
</bibRefCitation>
;
<bibRefCitation id="23244B53FFF2452AFC91F880FB1BFE3C" box="[881,1153,1847,1869]" pageId="8" pageNumber="459" refId="ref11574" refString="PURUSHOTHAmAN, J., HANSDA, S., DEY, J., MOHAN, S., BASI, A. &amp; VENkATARAmAN, K. (2017) AN ANNOTATED CHECklIST OF THAlIACEANS (CHORDATA: TUNICATES). Marine Biodiversity, 2017, 1 - 28. [PUblISHED ONlINE]" type="journal article">
Purushothaman
<emphasis id="75C1EAB0FFF2452AFBF5F88FFBD3FE3C" box="[1045,1097,1847,1869]" italics="true" pageId="8" pageNumber="459">et al.</emphasis>
2017
</bibRefCitation>
</paragraph>
<paragraph id="470A36A2FFF2452AFF77F8E1FD4BFE1D" blockId="8.[151,1436,1786,1930]" box="[151,721,1878,1900]" pageId="8" pageNumber="459">
<emphasis id="75C1EAB0FFF2452AFF77F8E1FEC0FE1D" box="[151,346,1878,1900]" italics="true" pageId="8" pageNumber="459">Salpa coerulescens</emphasis>
Chamisso 1819 (cited by
<bibRefCitation id="23244B53FFF2452AFD83F8E1FD50FE1D" box="[611,714,1878,1900]" pageId="8" pageNumber="459" refId="ref10785" refString="KOTT, P. (2005) Catalogue of Tunicata in Australian waters. AUSTRAlIAN BIOlOGICAl RESOURCES STUDY, DEPARTmENT OF THE ENvIRONmENT AND HERITAGE, CANbERRA, 301 PP." type="book">Kott 2005</bibRefCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="0FAF6529FFF2452FFF77F8C3FE6AFF9B" lastPageId="13" lastPageNumber="464" pageId="8" pageNumber="459" type="description">
<paragraph id="470A36A2FFF2452AFF77F8C3FB62FEFB" blockId="8.[151,1436,1786,1930]" box="[151,1272,1908,1930]" pageId="8" pageNumber="459">
<taxonomicName id="80B54D21FFF2452AFF77F8C3FED2FEFB" authorityName="Cuvier" authorityYear="1804" box="[151,328,1908,1930]" class="Thaliacea" family="Salpidae" genus="Weelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Salpida" pageId="8" pageNumber="459" phylum="Chordata" rank="species" species="cylindrica">
<emphasis id="75C1EAB0FFF2452AFF77F8C3FED2FEFB" box="[151,328,1908,1930]" italics="true" pageId="8" pageNumber="459">Weelia cylindrica</emphasis>
</taxonomicName>
; (Yount 1954); Van Soest 1975, 1998; Al-Yamani
<emphasis id="75C1EAB0FFF2452AFCA9F8C2FCE6FEFB" box="[841,892,1908,1930]" italics="true" pageId="8" pageNumber="459">et al.</emphasis>
2011; Hereu &amp; Suárez-Morales 2012
</paragraph>
<paragraph id="470A36A2FFF2452AFF77F800FD26FE85" blockId="8.[151,1438,1975,2036]" pageId="8" pageNumber="459">
<materialsCitation id="F7DD3CFFFFF2452AFF77F800FA00FEBE" ID-GBIF-Occurrence="1845321952" box="[151,1434,1975,2000]" collectingDate="2016-06" collectionCode="SOLITARY" country="Malaysia" latitude="5.4260917" location="MALAYSIA" longLatPrecision="1" longitude="103.29242" pageId="8" pageNumber="459" specimenCount="1">
<emphasis id="75C1EAB0FFF2452AFF77F800FE0CFEA1" bold="true" box="[151,406,1975,2000]" pageId="8" pageNumber="459">Specimens examined.</emphasis>
1
<collectionCode id="21A4AE67FFF2452AFE52F800FD9CFEBE" box="[434,518,1975,1999]" pageId="8" pageNumber="459">solitary</collectionCode>
zooids, 17 aggregate zooids,
<geoCoordinate id="22815065FFF2452AFCB3F800FC74FEA1" box="[851,1006,1975,2000]" direction="north" orientation="latitude" pageId="8" pageNumber="459" precision="1" value="5.4260917">5°25'33.93&quot;N</geoCoordinate>
,
<geoCoordinate id="22815065FFF2452AFC1AF800FB34FEBE" box="[1018,1198,1975,1999]" direction="east" orientation="longitude" pageId="8" pageNumber="459" precision="1" value="103.29242">103°17'32.72&quot;E</geoCoordinate>
,
<date id="330B1062FFF2452AFB59F80FFABEFEA1" box="[1209,1316,1975,2000]" pageId="8" pageNumber="459" value="2016-06">
<collectingDate id="234FE98AFFF2452AFB59F80FFABEFEA1" box="[1209,1316,1975,2000]" pageId="8" pageNumber="459" value="2016-06">Jun 2016</collectingDate>
</date>
.
<collectingCountry id="3FA27632FFF2452AFACEF800FA00FEBE" box="[1326,1434,1975,1999]" name="Malaysia" pageId="8" pageNumber="459">Malaysia</collectingCountry>
</materialsCitation>
:
<materialsCitation id="F7DD3CFFFFF2452AFF77F86BFD8AFE85" ID-GBIF-Occurrence="1845321954" box="[151,528,2011,2036]" collectionCode="EAST, COST" country="Malaysia" location="Of" pageId="8" pageNumber="459" specimenCount="1">
<collectionCode id="21A4AE67FFF2452AFF77F86BFE9BFE85" box="[151,257,2012,2036]" pageId="8" pageNumber="459">East Cost</collectionCode>
<location id="426A6079FFF2452AFEE8F86CFE85FE85" LSID="urn:lsid:plazi:treatment:CF1C87B4FFF2452CFF77F904FDD7FEB9:426A6079FFF2452AFEE8F86CFE85FE85" box="[264,287,2011,2036]" country="Malaysia" name="Of" pageId="8" pageNumber="459">of</location>
Peninsular
<collectingCountry id="3FA27632FFF2452AFE44F86BFD8AFE85" box="[420,528,2012,2036]" name="Malaysia" pageId="8" pageNumber="459">Malaysia</collectingCountry>
</materialsCitation>
. UMTTn 0 0 0 6.
</paragraph>
<caption id="13CA662AFFF3452BFF77F94FFD64FE3A" httpUri="https://zenodo.org/record/1253193/files/figure.png" pageId="9" pageNumber="460" startId="9.[151,250,1784,1806]" targetBox="[151,1423,193,1762]" targetPageId="9">
<paragraph id="470A36A2FFF3452BFF77F94FFD64FE3A" blockId="9.[151,1437,1784,1867]" pageId="9" pageNumber="460">
<emphasis id="75C1EAB0FFF3452BFF77F94FFEBBFE7F" bold="true" box="[151,289,1784,1806]" pageId="9" pageNumber="460">FIGURE 4a.</emphasis>
<taxonomicName id="80B54D21FFF3452BFEC9F94FFE59FE7F" authorityName="van Soest" authorityYear="1973" box="[297,451,1784,1806]" class="Thaliacea" family="Salpidae" genus="Thalia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Salpida" pageId="9" pageNumber="460" phylum="Chordata" rank="species" species="sibogae">
<emphasis id="75C1EAB0FFF3452BFEC9F94FFE59FE7F" box="[297,451,1784,1806]" italics="true" pageId="9" pageNumber="460">Thalia sibogae</emphasis>
</taxonomicName>
(van Soest, 1973). Top, Solitary, dorsal view; Bottom, ventral view. Br, branchial septum; Dt., dorsal tubercle; End, endostyle; G, ganglion, i.m., intermediate muscle; Lp. Lateral projection; Mp., medioventral projection; N, nucleus; o.m., oral musculature; Pp., posterior projection.
</paragraph>
</caption>
<caption id="13CA662AFFF04528FF77F8EFFB9EFEDA" httpUri="https://zenodo.org/record/1253195/files/figure.png" pageId="10" pageNumber="461" startId="10.[151,250,1880,1902]" targetBox="[151,1436,193,1858]" targetPageId="10">
<paragraph id="470A36A2FFF04528FF77F8EFFB9EFEDA" blockId="10.[151,1437,1880,1963]" pageId="10" pageNumber="461">
<emphasis id="75C1EAB0FFF04528FF77F8EFFEBCFE1F" bold="true" box="[151,294,1880,1902]" pageId="10" pageNumber="461">FIGURE 4b.</emphasis>
<taxonomicName id="80B54D21FFF04528FED1F8EFFE57FE1F" authorityName="van Soest" authorityYear="1973" box="[305,461,1880,1902]" class="Thaliacea" family="Salpidae" genus="Thalia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Salpida" pageId="10" pageNumber="461" phylum="Chordata" rank="species" species="sibogae">
<emphasis id="75C1EAB0FFF04528FED1F8EFFE57FE1F" box="[305,461,1880,1902]" italics="true" pageId="10" pageNumber="461">Thalia sibogae</emphasis>
</taxonomicName>
(van Soest, 1973). Top, Solitary, lateral view; Bottom, posterior part. Ap., atrial palp; Br, branchial septum; Dt., dorsal tubercle; End, endostyle; G, ganglion, i.m., intermediate muscle; Lp. Lateral projection, Mp., medioventral projection; N, nucleus; o.m., oral musculature; Pp., posterior projection.
</paragraph>
</caption>
<caption id="13CA662AFFF14529FF77F96FFEFCFE5A" httpUri="https://zenodo.org/record/1253197/files/figure.png" pageId="11" pageNumber="462" startId="11.[151,250,1752,1774]" targetBox="[154,1433,196,1728]" targetPageId="11">
<paragraph id="470A36A2FFF14529FF77F96FFEFCFE5A" blockId="11.[151,1435,1752,1835]" pageId="11" pageNumber="462">
<emphasis id="75C1EAB0FFF14529FF77F96FFE87FF9F" bold="true" box="[151,285,1752,1774]" pageId="11" pageNumber="462">FIGURE 4c.</emphasis>
<taxonomicName id="80B54D21FFF14529FEC3F96FFE20FF9F" authorityName="van Soest" authorityYear="1973" box="[291,442,1752,1774]" class="Thaliacea" family="Salpidae" genus="Thalia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Salpida" pageId="11" pageNumber="462" phylum="Chordata" rank="species" species="sibogae">
<emphasis id="75C1EAB0FFF14529FEC3F96FFE20FF9F" box="[291,442,1752,1774]" italics="true" pageId="11" pageNumber="462">Thalia sibogae</emphasis>
</taxonomicName>
(van Soest, 1973). Top, Aggregate, ventral view; Bottom, Chain of aggregate zooids. Br, branchial septum; Dt., dorsal tubercle; End, endostyle; G, ganglion, i.m., intermediate muscle; N, nucleus; o.m., oral musculature; Pp., posterior projection.
</paragraph>
</caption>
<caption id="13CA662AFFF6452EFF77FA43FAE8FF58" httpUri="https://zenodo.org/record/1253199/files/figure.png" pageId="12" pageNumber="463" startId="12.[151,250,1524,1546]" targetBox="[340,1247,213,1473]" targetPageId="12">
<paragraph id="470A36A2FFF6452EFF77FA43FAE8FF58" blockId="12.[151,1437,1524,1577]" pageId="12" pageNumber="463">
<emphasis id="75C1EAB0FFF6452EFF77FA43FEBBFF7B" bold="true" box="[151,289,1524,1546]" pageId="12" pageNumber="463">FIGURE 4d.</emphasis>
Comparison between A:
<taxonomicName id="80B54D21FFF6452EFDC7FA43FD25FF7B" authorityName="van Soest" authorityYear="1973" box="[551,703,1524,1546]" class="Thaliacea" family="Salpidae" genus="Thalia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Salpida" pageId="12" pageNumber="463" phylum="Chordata" rank="species" species="sibogae">
<emphasis id="75C1EAB0FFF6452EFDC7FA43FD25FF7B" box="[551,703,1524,1546]" italics="true" pageId="12" pageNumber="463">Thalia sibogae</emphasis>
</taxonomicName>
(van Soest, 1973) and B:
<taxonomicName id="80B54D21FFF6452EFC2BFA43FB11FF7B" authorityName="Quoy &amp; Gaimard" authorityYear="1824" box="[971,1163,1524,1546]" class="Thaliacea" family="Salpidae" genus="Thalia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Salpida" pageId="12" pageNumber="463" phylum="Chordata" rank="species" species="rhomboides">
<emphasis id="75C1EAB0FFF6452EFC2BFA43FB11FF7B" box="[971,1163,1524,1546]" italics="true" pageId="12" pageNumber="463">Thalia rhomboides</emphasis>
</taxonomicName>
(Quoy &amp; Gaimard, 1824). Top, Solitary, dorsal view, showing length of posterior projections; Bottom, body test part showing presence of echination.
</paragraph>
</caption>
<paragraph id="470A36A2FFF6452EFF27F9E2FA07FEAA" blockId="12.[151,1438,1621,2012]" pageId="12" pageNumber="463">
<emphasis id="75C1EAB0FFF6452EFF27F9E2FD3FFF1F" bold="true" box="[199,677,1621,1646]" pageId="12" pageNumber="463">
Description. Solitary (
<figureCitation id="DF8E2A27FFF6452EFE31F9E2FDBCFF1F" box="[465,550,1621,1646]" captionStart="FIGURE 6" captionStartId="15.[151,250,1804,1826]" captionTargetBox="[151,1433,195,1783]" captionTargetId="figure@15.[151,1436,195,1783]" captionTargetPageId="15" captionText="FIGURE 6a. Weelia cylindrica (Cuvier, 1804). Top, Solitary; Bottom, Aggregate. Br, branchial septum; Dt., dorsal tubercle; End, endostyle; G, ganglion, i.m., intermediate muscle; N, nucleus; o.m., oral musculature." httpUri="https://zenodo.org/record/1253205/files/figure.png" pageId="12" pageNumber="463">Fig. 6a</figureCitation>
, Top; 6b):
</emphasis>
Body length reaches 20.7 mm. Body is elongated and cylindrical; rounded anteriorly, and squarely cut off posteriorly. The test is thin, soft and smooth with no projection or echination. There are 9 body muscles that extend rather far ventrally and are usually separated. The first four body muscles; MI
<emphasis id="75C1EAB0FFF6452EFEC2F976FEAAFFAB" bold="true" box="[290,304,1729,1754]" pageId="12" pageNumber="463"></emphasis>
MIV are seen closely approaching in the mid-dorsal line. The muscle fibers for MIMIX is a total of 181. There are 3 sphincters present in the upper lip. The broad oral retractor, after crossing the intermediate muscle and the 3rd sphincter of the upper lip, gives 2nd sphincter where it appears to be continuous across the middle line. The retractor finally divided into the 1st short sphincter of the upper lip and the 1st and 2nd sphincters of the lower lip, which both the lower lips are continuous across the middle line. The transverse ribs of the gill bar meet dorsally, narrowing sharply toward the ventral surface, so that they are far apart from one another ventrally. The dorsal tubercle is a simple elongated arched stick. The alimentary canal is coiled into a tight nucleus. The gill is long and slender. Stolon runs forward in a straight course on the mid ventral line. There is no outgrowths from the ganglion.
</paragraph>
<caption id="13CA662AFFF7452FFF77FA81FB9EFCFB" httpUri="https://zenodo.org/record/1253201/files/figure.png" pageId="13" pageNumber="464" startId="13.[151,250,1334,1356]" targetBox="[170,1395,221,1312]" targetPageId="13">
<paragraph id="470A36A2FFF7452FFF77FA81FB9EFCFB" blockId="13.[151,1437,1334,1418]" pageId="13" pageNumber="464">
<emphasis id="75C1EAB0FFF7452FFF77FA81FEB9FC3D" bold="true" box="[151,291,1334,1356]" pageId="13" pageNumber="464">FIGURE 4e.</emphasis>
Drawing of
<taxonomicName id="80B54D21FFF7452FFE53FA81FDD5FC3D" authorityName="van Soest" authorityYear="1973" box="[435,591,1334,1356]" class="Thaliacea" family="Salpidae" genus="Thalia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Salpida" pageId="13" pageNumber="464" phylum="Chordata" rank="species" species="sibogae">
<emphasis id="75C1EAB0FFF7452FFE53FA81FDD5FC3D" box="[435,591,1334,1356]" italics="true" pageId="13" pageNumber="464">Thalia sibogae</emphasis>
</taxonomicName>
(van Soest, 1973). Left, Solitary; Right, aggregate zooid. Ap., atrial palp; Br, branchial septum; Dt., dorsal tubercle; End, endostyle; G, ganglion, i.m., intermediate muscle; Lp. Lateral projection; Mp., medioventral projection; N, nucleus; o.m., oral musculature; Pp., posterior projection.
</paragraph>
</caption>
<paragraph id="470A36A2FFF7452FFF27FA06FE6AFF9B" blockId="13.[151,1438,1457,2022]" pageId="13" pageNumber="464">
<emphasis id="75C1EAB0FFF7452FFF27FA06FDCDFCBB" bold="true" box="[199,599,1457,1482]" pageId="13" pageNumber="464">
Aggregate (
<figureCitation id="DF8E2A27FFF7452FFEB4FA06FE31FCBB" box="[340,427,1457,1482]" captionStart="FIGURE 6" captionStartId="15.[151,250,1804,1826]" captionTargetBox="[151,1433,195,1783]" captionTargetId="figure@15.[151,1436,195,1783]" captionTargetPageId="15" captionText="FIGURE 6a. Weelia cylindrica (Cuvier, 1804). Top, Solitary; Bottom, Aggregate. Br, branchial septum; Dt., dorsal tubercle; End, endostyle; G, ganglion, i.m., intermediate muscle; N, nucleus; o.m., oral musculature." httpUri="https://zenodo.org/record/1253205/files/figure.png" pageId="13" pageNumber="464">Fig. 6a</figureCitation>
, Bottom, 6b):
</emphasis>
Body length reaches 3.1 mm. Body is oval in form, with short hollow projections anteriorly and posteriorly present, which are slightly asymmetricaly arranged. The test is voluminous, thickly fusiform, soft and smooth with no echination. Five body muscles, all approaching in the mid-dorsal line in two separate groups; MI
<materialsCitation id="F7DD3CFFFFF7452FFE5AF9A9FB82FF44" ID-GBIF-Occurrence="1845321956" box="[442,1048,1565,1590]" collectionCode="MIII, ARE, MIV" country="American Samoa" location="AS" pageId="13" pageNumber="464" specimenCount="1">
<collectionCode id="21A4AE67FFF7452FFE5AF9A9FD83FF47" box="[442,537,1566,1590]" pageId="13" pageNumber="464">MIII are</collectionCode>
merged in the mid-dorsal region
<collectingCountry id="3FA27632FFF7452FFC77F9A9FC34FF47" box="[919,942,1566,1590]" name="American Samoa" pageId="13" pageNumber="464">as</collectingCountry>
<collectionCode id="21A4AE67FFF7452FFC57F9A9FC43FF47" box="[951,985,1566,1590]" pageId="13" pageNumber="464">are</collectionCode>
<collectionCode id="21A4AE67FFF7452FFC01F9A9FB82FF44" box="[993,1048,1566,1589]" country="Italy" httpUri="http://grbio.org/cool/mev6-t33n" name="Universita' degli Studi di Milano, Medicina Veterinaria, Sez. Parassitologia" pageId="13" pageNumber="464">MIV</collectionCode>
</materialsCitation>
MV; the two groups for the most part touch in the mid-dorsal line. The MV is branched, where its posterior branches on each side passing the base of the atrial siphon; and it unites behind the siphon. The transverse ribs of the gill bar meet dorsally, tapering severely toward the ventral surface, so that they are far-off from one another ventrally. The dorsal tubercle is a simple elongated arched stick. The alimentary canal is coiled into a tight nucleus. The gill is thick and short. Ovary and embryo are situated at MIV ¼.
</paragraph>
</subSubSection>
<subSubSection id="0FAF6529FFF7452CFF27F942FF6BFB1F" lastPageId="14" lastPageNumber="465" pageId="13" pageNumber="464" type="discussion">
<paragraph id="470A36A2FFF7452CFF27F942FF6BFB1F" blockId="13.[151,1438,1457,2022]" lastBlockId="14.[151,1437,151,622]" lastPageId="14" lastPageNumber="465" pageId="13" pageNumber="464">
<emphasis id="75C1EAB0FFF7452FFF27F942FEA1FE7F" bold="true" box="[199,315,1781,1806]" pageId="13" pageNumber="464">Remarks.</emphasis>
Both solitary and aggregate zooids of this species were found during the sampling of this study. According to Sewell in 1926 (p. 81, fig. 11), in the solitary stage some specimens have ten body muscles instead of nine while van Soest in 1975a (p. 111) stated that specimens from Atlantic Ocean were found to have 8 body muscles instead of nine. This specimen has nine body muscles. Sewell (p. 81, fig. 10, A
<emphasis id="75C1EAB0FFF7452FFB6CF8D6FB00FE0B" bold="true" box="[1164,1178,1889,1914]" pageId="13" pageNumber="464"></emphasis>
D) have also notes the fours variations in the connections of the MIMIV. The four anterior muscles of this specimen were observed to be closely matching with variation A, where the first MIMIII muscles are connected together and MIV is approaching MIII. Metcalf in 1918 (p. 95) describes the oral musculature
<collectingCountry id="3FA27632FFF7452FFC34F879FC71FE97" box="[980,1003,1998,2022]" name="American Samoa" pageId="13" pageNumber="464">as</collectingCountry>
“consisting of an oral retractor, which give rise to two sphincters of the lower lip and is connected with the 1st sphincter of the upper lip, a broad independent 2nd sphincter of the upper lip, and the intermediate muscle, both of these latter lying internal to the retractor”. From near the dorsal end of the intermediate muscle, a short horizontal muscle runs forwards. Sewell states that in two large Indian Ocean specimens examined by him, there was an additional sphincter in the upper lip. There are 3 sphincters present in the upper lip (p. 80, fig. 9). This specimen resembles of Sewells but with a slight different on the upper lip length. The broad oral retractor, after crossing the intermediate muscle and the 3rd sphincter of the upper lip, gives of a 2nd sphincter where it appeared to be continuous across the middle line. The retractor divided into the 1st short sphincter of the upper lip and the 1st and 2nd sphincters of the lower lip, which both the lower lips are continuous across the middle line. A distinct variation is also found in the number of muscular fiber. Van Soest (p. 112) stated the number of fibers could varies from 91261 fibers. In this specimen, for MIMIX there is a total of 181 muscle fibers. For aggregate zooids, this specimens closely resemble of Thompsons rather than the elongated variation collected from
<collectingCountry id="3FA27632FFF4452CFC86FD85FC0FFB3B" box="[870,917,562,586]" name="India" pageId="14" pageNumber="465">
<collectingRegion id="8571F840FFF4452CFC86FD85FC0FFB3B" box="[870,917,562,586]" country="India" name="Goa" pageId="14" pageNumber="465">Goa</collectingRegion>
</collectingCountry>
by Dr. S.W. Kemp
<collectingCountry id="3FA27632FFF4452CFB98FD85FB15FB3B" box="[1144,1167,562,586]" name="American Samoa" pageId="14" pageNumber="465">as</collectingCountry>
noted in Sewell (p. 82, fig. 12).
</paragraph>
</subSubSection>
<caption id="13CA662AFFF4452CFF77FAA3FD19FC39" httpUri="https://zenodo.org/record/1253203/files/figure.png" pageId="14" pageNumber="465" startId="14.[151,250,1300,1322]" targetBox="[345,1246,683,1278]" targetPageId="14">
<paragraph id="470A36A2FFF4452CFF77FAA3FD19FC39" blockId="14.[151,1436,1300,1353]" pageId="14" pageNumber="465">
<emphasis id="75C1EAB0FFF4452CFF77FAA3FE89FC5B" bold="true" box="[151,275,1300,1322]" pageId="14" pageNumber="465">FIGURE 5.</emphasis>
Distribution of
<taxonomicName id="80B54D21FFF4452CFE56FAA3FDD7FC5B" authorityName="van Soest" authorityYear="1973" box="[438,589,1300,1322]" class="Thaliacea" family="Salpidae" genus="Thalia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Salpida" pageId="14" pageNumber="465" phylum="Chordata" rank="species" species="sibogae">
<emphasis id="75C1EAB0FFF4452CFE56FAA3FDD7FC5B" box="[438,589,1300,1322]" italics="true" pageId="14" pageNumber="465">Thalia sibogae</emphasis>
</taxonomicName>
(van Soest, 1973). Red dots: specimen examined by the author; black dots: records based on compilation of previous literature data.
</paragraph>
</caption>
<subSubSection id="0FAF6529FFF4452CFF27FAC7FDD7FEB9" pageId="14" pageNumber="465" type="distribution">
<paragraph id="470A36A2FFF4452CFF27FAC7FDD7FEB9" blockId="14.[151,1436,1392,1993]" pageId="14" pageNumber="465">
<emphasis id="75C1EAB0FFF4452CFF27FAC7FEFAFCF6" bold="true" box="[199,352,1392,1417]" pageId="14" pageNumber="465">Distribution.</emphasis>
<taxonomicName id="80B54D21FFF4452CFE88FAC7FDB7FCF8" authorityName="Cuvier" authorityYear="1804" box="[360,557,1392,1417]" class="Thaliacea" family="Salpidae" genus="Weelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Salpida" pageId="14" pageNumber="465" phylum="Chordata" rank="species" species="cylindrica">
<emphasis id="75C1EAB0FFF4452CFE88FAC7FDB7FCF8" box="[360,557,1392,1417]" italics="true" pageId="14" pageNumber="465">Weelia cylindrica</emphasis>
</taxonomicName>
is the most common species of
<taxonomicName id="80B54D21FFF4452CFC79FAC7FC63FCF8" authorityName="Lahille" authorityYear="1888" box="[921,1017,1392,1417]" class="Thaliacea" family="Salpidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Salpida" pageId="14" pageNumber="465" phylum="Chordata" rank="family">Salpidae</taxonomicName>
(van Soest 1975), widely distributed in the Atlantic, Indian and Pacific Ocean. Records from the Atlantic Ocean includes of Dalls (1871), occurrence on the coast of Rhode
<collectingCountry id="3FA27632FFF4452CFE47FA0FFE71FCA0" box="[423,491,1464,1489]" name="Iceland" pageId="14" pageNumber="465">Island</collectingCountry>
and Block
<collectingCountry id="3FA27632FFF4452CFD99FA0FFD27FCA0" box="[633,701,1464,1489]" name="Iceland" pageId="14" pageNumber="465">Island</collectingCountry>
(Metcalf 1918), Sardinera Lagoon of
<collectingCountry id="3FA27632FFF4452CFB62FA0FFA91FCA1" box="[1154,1291,1464,1488]" name="Puerto Rico" pageId="14" pageNumber="465">Puerto Rico</collectingCountry>
(
<bibRefCitation id="23244B53FFF4452CFAFCFA0FFE1DFC84" pageId="14" pageNumber="465" refId="ref12607" refString="TOkIOkA, T. &amp; BHAvANARAYANA, P. B. (1979) NOTES ON THE OCCURRENCE OF A SWARm OF Salpa cylindrica CUvIER (TUNICATA: SAlPIDAE) IN SARDINERA LAGOON, PUERTO RICO. Proceedings of the Biological Society of Washington, 92 (3), 572 - 576." type="journal article">Tokioka &amp; Bhavanarayana 1979</bibRefCitation>
),
<bibRefCitation id="23244B53FFF4452CFE7BFA6BFD0FFC84" box="[411,661,1500,1525]" pageId="14" pageNumber="465" refId="ref9780" refString="HARbISON, G. R., MADIN, L. P. &amp; SWANbERG, N. R. (1978) ON THE NATURAl HISTORY AND DISTRIbUTION OF OCEANIC CTENOPHORES. Deep Sea Research, 25, 233 - 256." type="journal article">
Harbison
<emphasis id="75C1EAB0FFF4452CFDE9FA6AFDD8FC84" box="[521,578,1500,1525]" italics="true" pageId="14" pageNumber="465">et al.</emphasis>
(1978)
</bibRefCitation>
,
<bibRefCitation id="23244B53FFF4452CFD42FA6BFC6EFC84" box="[674,1012,1500,1525]" pageId="14" pageNumber="465" refId="ref9720" refString="HARbISON, G. R. &amp; CAmPENOT, R. B. (1979) EFFECTS OF TEmPERATURE ON THE SWImmING OF SAlPS (TUNICATA, THAlIACEA): ImPlICATIONS FOR VERTICAl MIGRATION. Limnology and Oceanography, 24 (6), 1081 - 1091." type="journal article">Harbison &amp; Campenot (1979)</bibRefCitation>
,
<bibRefCitation id="23244B53FFF4452CFBE1FA6AFAB3FC84" box="[1025,1321,1500,1525]" pageId="14" pageNumber="465" refId="ref11701" refString="POmEROY, L. R. &amp; DEIbEl, D. (1980) AGGREGATION OF ORGANIC MATTER bY PElAGIC TUNICATES. Limnology and Oceanography, 25 (4), 643 - 652." type="journal article">Pomeroy &amp; Deibel (1980)</bibRefCitation>
,
<bibRefCitation id="23244B53FFF4452CFAD6FA6BFEBCFF69" pageId="14" pageNumber="465" refId="ref11146" refString="MADIN, L. P. &amp; CETTA, C. M. (1984) THE USE OF GUT FlUORESCENCE TO ESTImATE GRAzING bY OCEANIC SAlPS. Journal of Plankton Research, 6 (3), 475 - 492." type="journal article">Madin &amp; Cetta (1984)</bibRefCitation>
,
<bibRefCitation id="23244B53FFF4452CFED2F9B7FD90FF69" box="[306,522,1536,1561]" pageId="14" pageNumber="465" refId="ref8533" refString="CARON, D. A., MADIN, L. P. &amp; COlE, J. J. (1989) COmPOSITION AND DEGRADATION OF SAlP FECAl PEllETS: ImPlICATIONS FOR vERTICAl FlUX IN OCEANIC ENvIRONmENTS. Journal of Marine Research, 47, 829 - 850." type="journal article">
Caron
<emphasis id="75C1EAB0FFF4452CFE9EF9B6FE2CFF68" box="[382,438,1536,1561]" italics="true" pageId="14" pageNumber="465">et al.</emphasis>
(1989)
</bibRefCitation>
, southeast of
<collectingCountry id="3FA27632FFF4452CFD44F9B7FC90FF68" box="[676,778,1536,1561]" name="Bermuda" pageId="14" pageNumber="465">Bermuda</collectingCountry>
(Madin
<emphasis id="75C1EAB0FFF4452CFC8CF9B6FC3EFF68" box="[876,932,1536,1561]" italics="true" pageId="14" pageNumber="465">et al.</emphasis>
1996), Mid-Atlantic Bight (
<bibRefCitation id="23244B53FFF4452CFB3FF9B7FF4BFF4C" pageId="14" pageNumber="465" refId="ref13050" refString="VARGAS, C. A. &amp; MADIN, L. P. (2004) ZOOPlANkTON FEEDING ECOlOGY: ClEARANCE AND INGESTION RATES OF THE SAlPS Thalia democratica, Cyclosalpa affinis AND Salpa cylindrica ON NATURAllY OCCURRING PARTIClES IN THE MID-ATlANTIC BIGHT. Journal of Plankton Research, 26 (7), 827 - 833." type="journal article">Vargas &amp; Madin 2004</bibRefCitation>
) and Caribbean Sea and the Gulf of
<collectingCountry id="3FA27632FFF4452CFD97F992FD56FF4C" box="[631,716,1573,1597]" name="Mexico" pageId="14" pageNumber="465">
<collectingRegion id="8571F840FFF4452CFD97F992FD56FF4C" box="[631,716,1573,1597]" country="Mexico" name="Mexico" pageId="14" pageNumber="465">Mexico</collectingRegion>
</collectingCountry>
(Hereu &amp; Suárez-Morales 2012). It has also been found in the Arabian Sea (including the
<collectingCountry id="3FA27632FFF4452CFE2FF9FFFDD8FF10" box="[463,578,1608,1633]" name="India" pageId="14" pageNumber="465">
<collectingRegion id="8571F840FFF4452CFE2FF9FFFDD8FF10" box="[463,578,1608,1633]" country="India" name="Lakshadweep" pageId="14" pageNumber="465">Laccadive</collectingRegion>
</collectingCountry>
Sea), the Bay of Bengal and the Andaman Sea (
<bibRefCitation id="23244B53FFF4452CFB86F9FFFB60FF10" box="[1126,1274,1608,1633]" pageId="14" pageNumber="465" refId="ref12143" refString="SEWEll, R. B. S. (1926) THE SAlPS OF THE INDIAN SEAS. Records of the Indian Museum, 28 (2), 65 - 126." type="journal article">Sewell 1926</bibRefCitation>
), Red Sea and Gulf of Suez (
<bibRefCitation id="23244B53FFF4452CFEDAF9DBFE7DFFF4" box="[314,487,1644,1669]" pageId="14" pageNumber="465" refId="ref9594" refString="GODEAUX, J. (1974) THAlIACeS ReCOlTeS AU lARGE DES CoTES eGYPTIENNES DE lA MeDITERRANeE ET DE lA mER ROUGE (TUNICATA, THAlIACEA). Beaufortia, 22 (291), 83 - 103." type="journal article">Godeaux 1974</bibRefCitation>
; Al-Yamani
<emphasis id="75C1EAB0FFF4452CFD9AF9DAFD2FFFF4" box="[634,693,1644,1669]" italics="true" pageId="14" pageNumber="465">et al.</emphasis>
2011 &amp; Kannathasan,
<emphasis id="75C1EAB0FFF4452CFC5FF9DAFC60FFF4" box="[959,1018,1644,1669]" italics="true" pageId="14" pageNumber="465">et al.</emphasis>
2014), Gulf of Aden &amp; Hanish Hill (
<bibRefCitation id="23244B53FFF4452CFF7CF927FED0FFD9" box="[156,330,1680,1705]" pageId="14" pageNumber="465" refId="ref9633" refString="GODEAUX, J. (1987) DISTRIbUTION OF THAlIACEA ON TRANSECT FROm THE GUlF OF ADEN DURING THE WINTER mONSOON (MARCH 1979). Oceanologica Acta, 10 (2), 197 - 204." type="journal article">Godeaux 1987</bibRefCitation>
). Records found from the Pacific Ocean includes (Yount 1954; 1958), Australian Seas (
<bibRefCitation id="23244B53FFF4452CFAD0F927FF55FFBC" pageId="14" pageNumber="465" refId="ref10117" refString="HERDmAN, W. A. (1891). A REvISED ClASSIFICATION OF THE TUNICATA, WITH DEFINITIONS OF THE ORDERS, SUb-ORDERS, FAmIlIES, SUb- FAmIlIES AND GENERA, AND ANAlYTICAl kEYS TO THE SPECIES. Journal of the Linnaean Society of London. Zoology, 23, 558 - 652." type="journal article">Herdman 1891</bibRefCitation>
&amp;
<bibRefCitation id="23244B53FFF4452CFF14F903FE35FFBC" box="[244,431,1716,1741]" pageId="14" pageNumber="465" refId="ref12451" refString="THOmPSON, H. (1948) Pelagic tunicates of Australia. COmmONWEAlTH COUNCIl FOR SCIENTIFIC AND INDUSTRIAl RESEARCH, MElbOURNE, 196 PP." type="book">Thompson 1948</bibRefCitation>
),
<bibRefCitation id="23244B53FFF4452CFE24F902FD49FFBC" box="[452,723,1716,1741]" pageId="14" pageNumber="465" refId="ref11848" refString="RITTER, W. E. &amp; BYXbEE, E. S. (1905) REPORTS ON THE SCIENTIFIC RESUlTS OF THE EXPEDITION TO THE TROPICAl PACIFIC IN THE AlbATROSS VIII. THE PElAGIC TUNICATA. Memoirs of the Museum for Comparative Zoology Harvard, 26 (5), 195 - 214." type="journal article">Ritter &amp; Byxbee (1905)</bibRefCitation>
, Great Barrier Reef lagoon (
<bibRefCitation id="23244B53FFF4452CFBF3F902FABDFFBC" box="[1043,1319,1716,1741]" pageId="14" pageNumber="465" refId="ref11903" refString="RUSSEll, F. S. &amp; COlmAN, J. S. (1935) THE ZOOPlANkTON IV. THE OCCURRENCE AND SEASONAl DISTRIbUTION OF THE TUNICATA, MOllUSCA AND COElENTERATA (SIPHONOPHORA). Scientific Rep. Great Barrier Reef Expedition, 2 (7), 205 - 224." type="journal article">Russell &amp; Colman1935</bibRefCitation>
), Province of Izu (
<bibRefCitation id="23244B53FFF4452CFF0FF96FFE0FFF81" box="[239,405,1752,1777]" pageId="14" pageNumber="465" refId="ref12476" refString="TOkIOkA, T. (1937) NOTES ON SAlPAS AND DOlIOlUmS OCCURRING IN THE PACIFIC COAST OF MIDDlE JAPAN. The Zoological Society of Japan, 16 (3), 219 - 235." type="journal article">Tokioka 1937</bibRefCitation>
;
<bibRefCitation id="23244B53FFF4452CFE44F96FFE47FF80" box="[420,477,1752,1777]" pageId="14" pageNumber="465" refId="ref12512" refString="TOkIOkA, T. (1960) STUDIES ON THE DISTRIbUTION OF APPENDICUlARIANS AND SOmE THAlIACEANS OF THE NORTH PACIFIC, WITH SOmE mORPHOlOGICAl NOTES. Publications of the Seto Marine Biological Laboratory, 8 (2), 351 - 443." type="journal article">1960</bibRefCitation>
) California Current region (
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),
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Current and Gulf of California
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(
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;
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),
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(Madin
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1996), European waters (van der Land &amp; van Soest 2001), Mid-Atlantic Bight (
<bibRefCitation id="23244B53FFF4452CFDEDF897FC97FE49" box="[525,781,1824,1849]" pageId="14" pageNumber="465" refId="ref13050" refString="VARGAS, C. A. &amp; MADIN, L. P. (2004) ZOOPlANkTON FEEDING ECOlOGY: ClEARANCE AND INGESTION RATES OF THE SAlPS Thalia democratica, Cyclosalpa affinis AND Salpa cylindrica ON NATURAllY OCCURRING PARTIClES IN THE MID-ATlANTIC BIGHT. Journal of Plankton Research, 26 (7), 827 - 833." type="journal article">Vargas &amp; Madin 2004</bibRefCitation>
), Kaoping Trench, southwestern waters of
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(
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), coast of
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(
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),
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tropical of
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(
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Hereu
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2010
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),
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waters (
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Liao
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2013
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&amp; Franco
<emphasis id="75C1EAB0FFF4452CFE66F8DEFE24FEF0" box="[390,446,1896,1921]" italics="true" pageId="14" pageNumber="465">et al.</emphasis>
2017),
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Straits (
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), northwest continental shelf of South
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Sea (
<bibRefCitation id="23244B53FFF4452CFA61F8DFFE8CFED4" pageId="14" pageNumber="465" refId="ref10885" refString="LI, K., YIN, J., HUANG, L., ZHANG, J., LIAN, S. &amp; LIU, C. (2011) DISTRIbUTION AND AbUNDANCE OF THAlIACEANS IN THE NORTHWEST CONTINENTAl SHElF OF SOUTH CHINA SEA, WITH RESPONSE TO ENvIRONmENTAl FACTORS DRIvEN bY mONSOON. Continental Shelf Research, 31, 979 - 989." type="journal article">
Li
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2011
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),
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, Borneo of
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. The compilation of previous literature and present data distribution of
<taxonomicName id="80B54D21FFF4452CFF77F807FEC1FEB8" authorityName="Cuvier" authorityYear="1804" box="[151,347,1968,1993]" class="Thaliacea" family="Salpidae" genus="Weelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Salpida" pageId="14" pageNumber="465" phylum="Chordata" rank="species" species="cylindrica">
<emphasis id="75C1EAB0FFF4452CFF77F807FEC1FEB8" box="[151,347,1968,1993]" italics="true" pageId="14" pageNumber="465">Weelia cylindrica</emphasis>
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is shown in
<figureCitation id="DF8E2A27FFF4452CFE08F806FDD7FEB9" box="[488,589,1969,1993]" captionStart="FIGURE 7" captionStartId="17.[151,250,1164,1186]" captionTargetBox="[345,1246,525,1136]" captionTargetId="figure@17.[340,1247,517,1142]" captionTargetPageId="17" captionText="FIGURE 7. Distribution of Weelia cylindrica (Cuvier, 1804) Red dots: specimen examined by the author; black dots: records based on compilation of previous literature data." httpUri="https://zenodo.org/record/1253209/files/figure.png" pageId="14" pageNumber="465">Figure 7.</figureCitation>
</paragraph>
</subSubSection>
</treatment>
</document>