350 lines
55 KiB
XML
350 lines
55 KiB
XML
<document id="36B07CB7A4D9D30283E77E75B3B5FE57" ID-DOI="10.5281/zenodo.5740573" ID-ISSN="0374-1036" ID-Zenodo-Dep="5740573" ID-ZooBank="A9032795-05E1-441F-9305-85D76070B7CD" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1638256467574" checkinUser="marcus" docAuthor="Moravec, Jiří & Duran, Daniel P." docDate="2013" docId="655487FAE158FFB8B9A448FAFE43FE68" docLanguage="en" docName="ActaEntMusNatPra.2013.53.2.585-599.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae 53 (2)" docStyle="DocumentStyle:E9BE312A6EDF9B84FC5211AAA80D80BF.10:ActaEntMusNatPra.2005-2016.journal_article" docStyleId="E9BE312A6EDF9B84FC5211AAA80D80BF" docStyleName="ActaEntMusNatPra.2005-2016.journal_article" docStyleVersion="10" docTitle="Odontocheila gilli Johnson 2000" docType="treatment" docVersion="4" lastPageNumber="594" masterDocId="996DFF82E15AFFB1B86C493BFFC0FF9C" masterDocTitle="Taxonomic and nomenclatorial revision within the Neotropical genera of the subtribe Odontochilina. 6. Odontocheila fraternum sp. nov., a new species sister to O. gilli (Coleoptera: Cicindelidae)" masterLastPageNumber="599" masterPageNumber="585" pageNumber="587" updateTime="1699435954514" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="1F2D1A91780B837B87B1938BF0023727">Taxonomic and nomenclatorial revision within the Neotropical genera of the subtribe Odontochilina. 6. Odontocheila fraternum sp. nov., a new species sister to O. gilli (Coleoptera: Cicindelidae)</mods:title>
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<mods:name id="03A214B68EDA3278F52A5565E3FDDCB3" type="personal">
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<mods:roleTerm id="73A4ED1E5DD9FC120DC95D6D73DD9F11">Author</mods:roleTerm>
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<mods:namePart id="0A21CD4115BD6D4D19E850631CDAB8DB">Moravec, Jiří</mods:namePart>
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<mods:affiliation id="BBFAE52F8E210CB4D76EF26089CB55DE">Sadová 336 / 21, 679 04 Adamov 1, Czech Republic; e-mail: jirmor @ quick. cz</mods:affiliation>
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<mods:namePart id="143B895A15767D751B2FC61CA80F8BE7">Duran, Daniel P.</mods:namePart>
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<mods:affiliation id="6BC6145A716EDA5C328150830DE03012">Department of Biology, Drexel University, 3141 Chestnut St., Philadelphia, PA 19104, U. S. A; e-mail: dpd 56 @ drexel. edu</mods:affiliation>
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<mods:title id="3E9F9C87D023652969B9C880D5137F34">Acta Entomologica Musei Nationalis Pragae</mods:title>
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<mods:date id="D256506246D9ED6989ACB2CB04003735">2013</mods:date>
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<mods:number id="A2C74EDD8EAE9926D647B638052552DD">2013-11-15</mods:number>
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<mods:number id="0E76707DA010E794AFE6D699A82E5420">53</mods:number>
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<treatment id="655487FAE158FFB8B9A448FAFE43FE68" ID-DOI="http://doi.org/10.5281/zenodo.5740869" ID-Zenodo-Dep="5740869" LSID="urn:lsid:plazi:treatment:655487FAE158FFB8B9A448FAFE43FE68" httpUri="http://treatment.plazi.org/id/655487FAE158FFB8B9A448FAFE43FE68" lastPageId="9" lastPageNumber="594" pageId="2" pageNumber="587">
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<subSubSection id="A5E76567E158FFB3B9A448FAFCF5FE45" box="[456,821,449,474]" pageId="2" pageNumber="587" type="nomenclature">
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<paragraph id="ED4236ECE158FFB3B9A448FAFCF5FE45" blockId="2.[456,821,449,474]" box="[456,821,449,474]" pageId="2" pageNumber="587">
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<heading id="B60A8180E158FFB3B9A448FAFCF5FE45" bold="true" box="[456,821,449,474]" centered="true" fontSize="8" level="2" pageId="2" pageNumber="587" reason="3">
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<taxonomicName id="2AFD4D6FE158FFB3B9A448FAFCF5FE45" ID-CoL="48PKF" authority="Johnson, 2000" authorityName="Johnson" authorityYear="2000" box="[456,821,449,474]" class="Insecta" family="Carabidae" genus="Odontocheila" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="587" phylum="Arthropoda" rank="species" species="gilli">
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<emphasis id="DF89EAFEE158FFB3B9A448FAFCF5FE45" bold="true" box="[456,821,449,474]" pageId="2" pageNumber="587">
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<emphasis id="DF89EAFEE158FFB3B9A448FAFD4EFE46" bold="true" box="[456,654,449,474]" italics="true" pageId="2" pageNumber="587">Odontocheila gilli</emphasis>
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<bibRefCitation id="896C4B1DE158FFB3BAF848F9FCF5FE45" author="JOHNSON W. N." box="[660,821,449,474]" pageId="2" pageNumber="587" pagination="13 - 18" refId="ref7088" refString="JOHNSON W. N. 2000: A new species of Odontocheila from Panama (Coleoptera: Cicindelidae). Cicindela 32: 13 - 18." type="journal article" year="2000">Johnson, 2000</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="A5E76567E158FFB3BA5448DBFD05FE6F" box="[568,709,480,499]" pageId="2" pageNumber="587" type="description">
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<paragraph id="ED4236ECE158FFB3BA5448DBFD05FE6F" blockId="2.[151,709,480,533]" box="[568,709,480,499]" pageId="2" pageNumber="587">
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(
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<figureCitation id="75C62A69E158FFB3BA5248DBFD4BFE6F" box="[574,651,480,499]" captionStart="Figs 1–6" captionStartId="4.[151,188,1486,1505]" captionTargetBox="[185,1089,221,1439]" captionTargetId="figure-10@4.[183,1094,219,1444]" captionTargetPageId="4" captionText="Figs 1–6. Habitus of two species of Odontocheila Laporte de Castelnau, 1834. 1–2 – O. gilli Johnson, 2000:1 – male, 8 mm, Chepo-Carti Road, HT (CMNC); 2 – male, 8.1 mm, Nusagandi (DBCN).3–6 – O. fraternum sp. nov.: 3 – male, 8.9 mm, Darien, HT (USNM); 4 – male, 8.6 mm, Altos de Campana, PT (CCJM); 5 – female, 9.1 mm, Darien, AT (DBCN); 6 – female, 10 mm, Altos de Campana (CCJM)." figureDoi="http://doi.org/10.5281/zenodo.5740577" httpUri="https://zenodo.org/record/5740577/files/figure.png" pageId="2" pageNumber="587">Figs 1–2</figureCitation>
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,
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<figureCitation id="75C62A69E158FFB3BAF848DBFD7FFE6F" box="[660,703,480,499]" captionStart="Figs 7–21" captionStartId="5.[113,150,1468,1487]" captionTargetBox="[121,1083,222,1444]" captionTargetId="figure-17@5.[117,1084,219,1447]" captionTargetPageId="5" captionText="Figs 7–21. Odontocheila gilli Johnson, 2000. 7 – head, male, Nusagandi (DBCN); 8–10 – labrum: 8 – male, Chepo- Carti Road, HT (CMNC); 9 – male, Nusagandi; 10 – female, Chepo-Carti Road, AT (WJCM). 11–13 – pronotum: 11 – male, HT; 12 – male, Nusagandi; 13 – female, AT. 14–16 – elytron: 14 – male, HT; 15 – male, Nusagandi; 16 – female, AT. 17 – elytral apex, male, HT; 18–19 – elytral apex, back illumination: 18 – male, Nusagandi; 19 – female, AT. 20–21 – aedeagus: 20 – HT; 21 – Nusagandi. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5740579" httpUri="https://zenodo.org/record/5740579/files/figure.png" pageId="2" pageNumber="587">7–21</figureCitation>
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)
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||
</paragraph>
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||
</subSubSection>
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<subSubSection id="A5E76567E158FFB3B8FB4B38FDAFFD89" box="[151,623,514,533]" pageId="2" pageNumber="587" type="reference_group">
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<paragraph id="ED4236ECE158FFB3B8FB4B38FDAFFD89" blockId="2.[151,709,480,533]" box="[151,623,514,533]" pageId="2" pageNumber="587">
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<treatmentCitationGroup id="CDED11C2E158FFB3B8FB4B38FDAFFD89" box="[151,623,514,533]" pageId="2" pageNumber="587">
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<treatmentCitation id="6C5C10FDE158FFB3B8FB4B38FE16FD89" author="JOHNSON W. N." box="[151,470,514,533]" page="14" pageId="2" pageNumber="587" year="2000">
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<taxonomicName id="2AFD4D6FE158FFB3B8FB4B38FE16FD89" ID-CoL="48PKF" authority="Johnson, 2000: 14" authorityName="Johnson" authorityPageNumber="14" authorityYear="2000" box="[151,470,514,533]" class="Insecta" family="Carabidae" genus="Odontocheila" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="587" phylum="Arthropoda" rank="species" species="gilli">
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<emphasis id="DF89EAFEE158FFB3B8FB4B38FEF3FD89" box="[151,307,515,533]" italics="true" pageId="2" pageNumber="587">Odontocheila gilli</emphasis>
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<bibRefCitation id="896C4B1DE158FFB3B9544B39FE16FD89" author="JOHNSON W. N." box="[312,470,514,533]" pageId="2" pageNumber="587" pagination="13 - 18" refId="ref7088" refString="JOHNSON W. N. 2000: A new species of Odontocheila from Panama (Coleoptera: Cicindelidae). Cicindela 32: 13 - 18." type="journal article" year="2000">Johnson, 2000: 14</bibRefCitation>
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</taxonomicName>
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||
</treatmentCitation>
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, 15–17,
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<figureCitation id="75C62A69E158FFB3BA734B39FDABFD89" box="[543,619,514,533]" captionStart="Figs 1–6" captionStartId="4.[151,188,1486,1505]" captionTargetBox="[185,1089,221,1439]" captionTargetId="figure-10@4.[183,1094,219,1444]" captionTargetPageId="4" captionText="Figs 1–6. Habitus of two species of Odontocheila Laporte de Castelnau, 1834. 1–2 – O. gilli Johnson, 2000:1 – male, 8 mm, Chepo-Carti Road, HT (CMNC); 2 – male, 8.1 mm, Nusagandi (DBCN).3–6 – O. fraternum sp. nov.: 3 – male, 8.9 mm, Darien, HT (USNM); 4 – male, 8.6 mm, Altos de Campana, PT (CCJM); 5 – female, 9.1 mm, Darien, AT (DBCN); 6 – female, 10 mm, Altos de Campana (CCJM)." figureDoi="http://doi.org/10.5281/zenodo.5740577" httpUri="https://zenodo.org/record/5740577/files/figure.png" pageId="2" pageNumber="587">Figs 1–2</figureCitation>
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.
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||
</treatmentCitationGroup>
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</paragraph>
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||
</subSubSection>
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<subSubSection id="A5E76567E158FFB3B8FB4B17FDA0FCB3" pageId="2" pageNumber="587" type="materials_examined">
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||
<paragraph id="ED4236ECE158FFB3B8FB4B17FDE6FD66" blockId="2.[151,1128,555,815]" pageId="2" pageNumber="587">
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||
<emphasis id="DF89EAFEE158FFB3B8FB4B17FECAFDA3" bold="true" box="[151,266,555,575]" pageId="2" pageNumber="587">Type locality</emphasis>
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.
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<collectingCountry id="95EA767CE158FFB3B9794B17FE9AFDA3" box="[277,346,556,575]" name="Panama" pageId="2" pageNumber="587">Panama</collectingCountry>
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, San Blas province, Chepo-Carti Road,
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<quantity id="2A059B09E158FFB3BAC04B17FD22FDA3" box="[684,738,556,575]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="2" pageNumber="587" unit="m" value="400.0">400 m</quantity>
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a.s.l., El Llano-Carti area, San Blas Foothills.
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<emphasis id="DF89EAFEE158FFB3B8FB4B7CFEDBFDC6" bold="true" box="[151,283,582,602]" pageId="2" pageNumber="587">Type material.</emphasis>
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||
<typeStatus id="3246884EE158FFB3B9724B7CFEB2FDC4" box="[286,370,583,602]" pageId="2" pageNumber="587" type="holotype">HOLOTYPE</typeStatus>
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||
: J, “
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<collectingCountry id="95EA767CE158FFB3B9F24B7CFE26FDC6" box="[414,486,583,602]" name="Panama" pageId="2" pageNumber="587">Panama</collectingCountry>
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:
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<collectingCountry id="95EA767CE158FFB3B9804B7CFDF0FDC6" box="[492,560,583,602]" name="Panama" pageId="2" pageNumber="587">Panama</collectingCountry>
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/ Chepo-Carti Rd. /
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<date id="9943102CE158FFB3BAB44B7CFCF3FDC6" box="[728,819,583,602]" pageId="2" pageNumber="587" value="1982-06-06">6.VI. 1982</date>
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/ B. Gill
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<quantity id="2A059B09E158FFB3BB124B7CFC73FDC6" box="[894,947,583,602]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="2" pageNumber="587" unit="m" value="400.0">400 m</quantity>
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” [printed, date partly handwritten] // “
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<taxonomicName id="2AFD4D6FE158FFB3B94C4B5AFE66FDE9" authority="II" authorityName="II" box="[288,422,609,629]" class="Insecta" family="Carabidae" genus="Odontocheila" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="587" phylum="Arthropoda" rank="genus">Odontocheila II</taxonomicName>
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/ or III cf mexicana / cf quadrina” [handwritten] // “
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<typeStatus id="3246884EE158FFB3BB3F4B5AFC05FDE8" box="[851,965,609,628]" pageId="2" pageNumber="587" type="holotype">HOLOTYPE</typeStatus>
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” [red with doubled black frame, printed] // “
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<typeStatus id="3246884EE158FFB3B9044B47FE1AFD13" box="[360,474,636,655]" pageId="2" pageNumber="587" type="holotype">HOLOTYPE</typeStatus>
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/
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<taxonomicName id="2AFD4D6FE158FFB3B9864B47FD44FD13" authorityName="Johnson" authorityYear="2000" box="[490,644,636,655]" class="Insecta" family="Carabidae" genus="Odontocheila" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="587" phylum="Arthropoda" rank="species" species="gilli">Odontocheila gilli</taxonomicName>
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/ W. Johnson” [yellow-green, printed] // “CANADIAN / MUSEUM / OF NATURE / LP2012-0171” [printed, with coloured circular logo] // “Canadian Museum of / Musée canadien de la / NATURE / CMNEN 00011499” [printed and with QR code] (
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<collectionCode id="8BECAE29E158FFB3BB2B4B8AFC4BFD58" box="[839,907,689,708]" pageId="2" pageNumber="587">CMNC</collectionCode>
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).
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<typeStatus id="3246884EE158FFB3BBF54B8AFC2AFD5F" box="[921,1002,689,708]" pageId="2" pageNumber="587" type="allotype">ALLOTYPE</typeStatus>
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:
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<specimenCount id="FBFBFD65E158FFB3BB9B4B90FBC7FD58" box="[1015,1031,683,708]" pageId="2" pageNumber="587" type="female">♀</specimenCount>
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with same locality label and: “
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<typeStatus id="3246884EE158FFB3B9574BF7FE6AFD43" box="[315,426,716,735]" pageId="2" pageNumber="587" type="allotype">ALLOTYPE</typeStatus>
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||
/
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<taxonomicName id="2AFD4D6FE158FFB3B9D54BF7FD92FD43" authorityName="Johnson" authorityYear="2000" box="[441,594,716,735]" class="Insecta" family="Carabidae" genus="Odontocheila" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="587" phylum="Arthropoda" rank="species" species="gilli">Odontocheila gilli</taxonomicName>
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/ W. Johnson” [yellow, printed] (
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<collectionCode id="8BECAE29E158FFB3BB0B4BF7FC6AFD43" box="[871,938,716,735]" pageId="2" pageNumber="587">WJCM</collectionCode>
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||
). Both type specimens are pined, the
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<typeStatus id="3246884EE158FFB3B97C4BDCFE95FD66" box="[272,341,743,762]" pageId="2" pageNumber="587" type="allotype">allotype</typeStatus>
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is of a rather bad shape.
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</paragraph>
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<paragraph id="ED4236ECE158FFB3B8FB4A3BFDA0FCB3" blockId="2.[151,1128,555,815]" pageId="2" pageNumber="587">
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<emphasis id="DF89EAFEE158FFB3B8FB4A3BFE55FC88" bold="true" box="[151,405,768,788]" pageId="2" pageNumber="587">Other specimens examined.</emphasis>
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1 J, “
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<collectingCountry id="95EA767CE158FFB3B9BD4A3AFDD5FC88" box="[465,533,769,788]" name="Panama" pageId="2" pageNumber="587">Panama</collectingCountry>
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– San Blas / Nusagandi Reserve
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<quantity id="2A059B09E158FFB3BB514A3AFCADFC88" box="[829,877,769,788]" metricMagnitude="2" metricUnit="m" metricValue="3.5" pageId="2" pageNumber="587" unit="m" value="350.0">350m</quantity>
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||
/ D. Brzoska,
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<date id="9943102CE158FFB3BB834A3AFB8AFC88" box="[1007,1098,769,788]" pageId="2" pageNumber="587" value="1999-05-17">17-V-1999</date>
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” // “
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<taxonomicName id="2AFD4D6FE158FFB3B8CC4A27FEFBFCB3" authorityName="Johnson" authorityYear="2000" box="[160,315,796,815]" class="Insecta" family="Carabidae" genus="Odontocheila" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="587" phylum="Arthropoda" rank="species" species="gilli">Odontocheila gilli</taxonomicName>
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/ det. D. Brzoska 2001” (
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<collectionCode id="8BECAE29E158FFB3BA794A27FD95FCB3" box="[533,597,796,815]" pageId="2" pageNumber="587">DBCN</collectionCode>
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||
).
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="A5E76567E158FFB9B8FB4A7CFC5FFAC8" lastPageId="8" lastPageNumber="593" pageId="2" pageNumber="587" type="description">
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<paragraph id="ED4236ECE158FFB3B8FB4A7CFD2EFC23" blockId="2.[151,1127,839,1600]" pageId="2" pageNumber="587">
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||
<emphasis id="DF89EAFEE158FFB3B8FB4A7CFEE2FCC3" bold="true" box="[151,290,839,863]" pageId="2" pageNumber="587">Description.</emphasis>
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Body (
|
||
<figureCitation id="75C62A69E158FFB3B9184A7CFE14FCC3" box="[372,468,839,863]" captionStart="Figs 1–6" captionStartId="4.[151,188,1486,1505]" captionTargetBox="[185,1089,221,1439]" captionTargetId="figure-10@4.[183,1094,219,1444]" captionTargetPageId="4" captionText="Figs 1–6. Habitus of two species of Odontocheila Laporte de Castelnau, 1834. 1–2 – O. gilli Johnson, 2000:1 – male, 8 mm, Chepo-Carti Road, HT (CMNC); 2 – male, 8.1 mm, Nusagandi (DBCN).3–6 – O. fraternum sp. nov.: 3 – male, 8.9 mm, Darien, HT (USNM); 4 – male, 8.6 mm, Altos de Campana, PT (CCJM); 5 – female, 9.1 mm, Darien, AT (DBCN); 6 – female, 10 mm, Altos de Campana (CCJM)." figureDoi="http://doi.org/10.5281/zenodo.5740577" httpUri="https://zenodo.org/record/5740577/files/figure.png" pageId="2" pageNumber="587">Figs 1–2</figureCitation>
|
||
) small, length 8.10–8.60 (
|
||
<typeStatus id="3246884EE158FFB3BA9F4A7CFC92FCC3" box="[755,850,839,863]" pageId="2" pageNumber="587" type="holotype">holotype</typeStatus>
|
||
8.20,
|
||
<typeStatus id="3246884EE158FFB3BBF44A7CFC2EFCC3" box="[920,1006,839,863]" pageId="2" pageNumber="587" type="allotype">allotype</typeStatus>
|
||
8.60) mm, width 2.40–2.80 (
|
||
<typeStatus id="3246884EE158FFB3B9224A5CFE6CFCE3" box="[334,428,871,895]" pageId="2" pageNumber="587" type="holotype">holotype</typeStatus>
|
||
2.40,
|
||
<typeStatus id="3246884EE158FFB3B9874A5CFD80FCE3" box="[491,576,871,895]" pageId="2" pageNumber="587" type="allotype">allotype</typeStatus>
|
||
2.80) mm, median area of pronotum and elytra shiny metallic yellow-green with bronze-cupreous lustre or golden-cupreous, sublateral areas bright green to green-blue, and lateral areas purple-violaceous.
|
||
</paragraph>
|
||
<paragraph id="ED4236ECE158FFB3B8D94AFCFE34FC63" blockId="2.[151,1127,839,1600]" pageId="2" pageNumber="587">
|
||
Head (
|
||
<figureCitation id="75C62A69E158FFB3B8924AFCFE80FC43" box="[254,320,967,991]" captionStart="Figs 7–21" captionStartId="5.[113,150,1468,1487]" captionTargetBox="[121,1083,222,1444]" captionTargetId="figure-17@5.[117,1084,219,1447]" captionTargetPageId="5" captionText="Figs 7–21. Odontocheila gilli Johnson, 2000. 7 – head, male, Nusagandi (DBCN); 8–10 – labrum: 8 – male, Chepo- Carti Road, HT (CMNC); 9 – male, Nusagandi; 10 – female, Chepo-Carti Road, AT (WJCM). 11–13 – pronotum: 11 – male, HT; 12 – male, Nusagandi; 13 – female, AT. 14–16 – elytron: 14 – male, HT; 15 – male, Nusagandi; 16 – female, AT. 17 – elytral apex, male, HT; 18–19 – elytral apex, back illumination: 18 – male, Nusagandi; 19 – female, AT. 20–21 – aedeagus: 20 – HT; 21 – Nusagandi. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5740579" httpUri="https://zenodo.org/record/5740579/files/figure.png" pageId="2" pageNumber="587">Fig. 7</figureCitation>
|
||
) notably large with pronounced eyes, as wide as the body,
|
||
<quantity id="2A059B09E158FFB3BBA64AFCFBA6FC43" box="[970,1126,967,991]" metricMagnitude="-3" metricUnit="m" metricValue="2.625" metricValueMax="2.8" metricValueMin="2.45" pageId="2" pageNumber="587" unit="mm" value="2.625" valueMax="2.8" valueMin="2.45">2.45–2.80 mm</quantity>
|
||
wide, all head portions glabrous.
|
||
</paragraph>
|
||
<paragraph id="ED4236ECE158FFB3B8D94D33FC11FB03" blockId="2.[151,1127,839,1600]" pageId="2" pageNumber="587">Frons rather steeply sloped towards clypeus, with convex median area, clearly delimited from clypeus and separated from vertex by distinct, transverse, rather sharp edge; frons surface dark copper on lateral areas which are almost smooth or only indistinctly finely longitudinally wrinkled, median area ornamented by short, irregularly transverse to arcuate-arranged wavy rugae; supraantennal plates smooth, metallic green-blue with cupreous lustre.</paragraph>
|
||
<paragraph id="ED4236ECE158FFB3B8D94D93FDA7FAE3" blockId="2.[151,1127,839,1600]" pageId="2" pageNumber="587">Vertex with usual juxtaorbital sensory seta (on each side), almost flat, reddish cupreous in middle with well delimited bright greenish-blue sublateral areas and green-blue to violaceous postero-lateral areas passing onto temples; anteromedian area with irregularly transverse-wavy and arcuate rugae passing from frons and forming a rather distinct ornament, large juxtaorbital areas cupreous, distinctly longitudinally parallel-striate, striae on sublateral areas more irregular and wavy, divergent posteriad passing onto temples and postgenae; occipital area very finely irregularly vermicular-rugulose.</paragraph>
|
||
<paragraph id="ED4236ECE158FFB3B8D94CBCFC6DFA03" blockId="2.[151,1127,839,1600]" box="[181,941,1415,1440]" pageId="2" pageNumber="587">Clypeus iridescent green with bluish lateral areas, irregularly wrinkled.</paragraph>
|
||
<paragraph id="ED4236ECE158FFB3B8D94C9CFF18FA43" blockId="2.[151,1127,839,1600]" pageId="2" pageNumber="587">Genae metallic green-blue, almost smooth with barely recognizable, shallow, parallel striae.</paragraph>
|
||
<paragraph id="ED4236ECE158FFB2B8D94CDCFEC5FE0F" blockId="2.[151,1127,839,1600]" lastBlockId="3.[113,1089,219,1588]" lastPageId="3" lastPageNumber="588" pageId="2" pageNumber="587">
|
||
Labrum 4-setose, male labrum (
|
||
<figureCitation id="75C62A69E158FFB3BA4D4CDCFD46FA63" box="[545,646,1511,1535]" captionStart="Figs 7–21" captionStartId="5.[113,150,1468,1487]" captionTargetBox="[121,1083,222,1444]" captionTargetId="figure-17@5.[117,1084,219,1447]" captionTargetPageId="5" captionText="Figs 7–21. Odontocheila gilli Johnson, 2000. 7 – head, male, Nusagandi (DBCN); 8–10 – labrum: 8 – male, Chepo- Carti Road, HT (CMNC); 9 – male, Nusagandi; 10 – female, Chepo-Carti Road, AT (WJCM). 11–13 – pronotum: 11 – male, HT; 12 – male, Nusagandi; 13 – female, AT. 14–16 – elytron: 14 – male, HT; 15 – male, Nusagandi; 16 – female, AT. 17 – elytral apex, male, HT; 18–19 – elytral apex, back illumination: 18 – male, Nusagandi; 19 – female, AT. 20–21 – aedeagus: 20 – HT; 21 – Nusagandi. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5740579" httpUri="https://zenodo.org/record/5740579/files/figure.png" pageId="2" pageNumber="587">Figs 8–9</figureCitation>
|
||
) rather long, length
|
||
<quantity id="2A059B09E158FFB3BB194CDCFBDAFA63" box="[885,1050,1511,1535]" metricMagnitude="-4" metricUnit="m" metricValue="6.75" metricValueMax="7.0" metricValueMin="6.5" pageId="2" pageNumber="587" unit="mm" value="0.675" valueMax="0.7" valueMin="0.65">0.65–0.70 mm</quantity>
|
||
, width
|
||
<quantity id="2A059B09E158FFB3B8FB4F3CFEF5F983" box="[151,309,1543,1567]" metricMagnitude="-3" metricUnit="m" metricValue="1.075" metricValueMax="1.1" metricValueMin="1.05" pageId="2" pageNumber="587" unit="mm" value="1.075" valueMax="1.1" valueMin="1.05">1.05–1.10 mm</quantity>
|
||
, ochre-testaceous, with brown to black-brown darkened basal and sublateral areas (more obvious so in
|
||
<typeStatus id="3246884EE158FFB3B9DA4F1CFDD8F9A3" box="[438,536,1575,1599]" pageId="2" pageNumber="587" type="holotype">holotype</typeStatus>
|
||
), labral shape remarkable within the genus, possessing rounded lateral teeth, larger right-angled or rounded anterolateral teeth and prominent, acute anterior teeth of a horn-like shape; anterior margin between these teeth shallowly emarginate; female labrum (
|
||
<figureCitation id="75C62A69E159FFB2B9754820FEA4FEA8" box="[281,356,283,308]" captionStart="Figs 7–21" captionStartId="5.[113,150,1468,1487]" captionTargetBox="[121,1083,222,1444]" captionTargetId="figure-17@5.[117,1084,219,1447]" captionTargetPageId="5" captionText="Figs 7–21. Odontocheila gilli Johnson, 2000. 7 – head, male, Nusagandi (DBCN); 8–10 – labrum: 8 – male, Chepo- Carti Road, HT (CMNC); 9 – male, Nusagandi; 10 – female, Chepo-Carti Road, AT (WJCM). 11–13 – pronotum: 11 – male, HT; 12 – male, Nusagandi; 13 – female, AT. 14–16 – elytron: 14 – male, HT; 15 – male, Nusagandi; 16 – female, AT. 17 – elytral apex, male, HT; 18–19 – elytral apex, back illumination: 18 – male, Nusagandi; 19 – female, AT. 20–21 – aedeagus: 20 – HT; 21 – Nusagandi. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5740579" httpUri="https://zenodo.org/record/5740579/files/figure.png" pageId="3" pageNumber="588">Fig. 10</figureCitation>
|
||
) much longer, length
|
||
<quantity id="2A059B09E159FFB2BA274827FD6CFEAF" box="[587,684,284,308]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="3" pageNumber="588" unit="mm" value="1.1">1.10 mm</quantity>
|
||
, width
|
||
<quantity id="2A059B09E159FFB2BA944820FC98FEAF" box="[760,856,283,307]" metricMagnitude="-3" metricUnit="m" metricValue="1.15" pageId="3" pageNumber="588" unit="mm" value="1.15">1.15 mm</quantity>
|
||
, much darker, reddish brown with larger black basal and anterior marginal areas, with rounded lateral and anterolateral teeth and prominent, acutely tridentate median lobe with wider and more protruding median tooth.
|
||
</paragraph>
|
||
<paragraph id="ED4236ECE159FFB2B8FC48A0FEF1FDAF" blockId="3.[113,1089,219,1588]" pageId="3" pageNumber="588">
|
||
Mandibles (
|
||
<figureCitation id="75C62A69E159FFB2B96248A0FE8CFE28" box="[270,332,411,436]" captionStart="Figs 7–21" captionStartId="5.[113,150,1468,1487]" captionTargetBox="[121,1083,222,1444]" captionTargetId="figure-17@5.[117,1084,219,1447]" captionTargetPageId="5" captionText="Figs 7–21. Odontocheila gilli Johnson, 2000. 7 – head, male, Nusagandi (DBCN); 8–10 – labrum: 8 – male, Chepo- Carti Road, HT (CMNC); 9 – male, Nusagandi; 10 – female, Chepo-Carti Road, AT (WJCM). 11–13 – pronotum: 11 – male, HT; 12 – male, Nusagandi; 13 – female, AT. 14–16 – elytron: 14 – male, HT; 15 – male, Nusagandi; 16 – female, AT. 17 – elytral apex, male, HT; 18–19 – elytral apex, back illumination: 18 – male, Nusagandi; 19 – female, AT. 20–21 – aedeagus: 20 – HT; 21 – Nusagandi. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5740579" httpUri="https://zenodo.org/record/5740579/files/figure.png" pageId="3" pageNumber="588">Fig. 7</figureCitation>
|
||
) normally shaped with arcuate lateral margins, ochre to brownish-testaceous in male, dark brownish in female, subsymmetrical, each mandible with four teeth (and basal molar), left mandible with second and third tooth of approximately same size, fourth tooth smaller; right mandible with third tooth somewhat smaller than the second, fourth tooth markedly smaller.
|
||
</paragraph>
|
||
<paragraph id="ED4236ECE159FFB2B8FC4B00FD99FD68" blockId="3.[113,1089,219,1588]" pageId="3" pageNumber="588">
|
||
Palpi (
|
||
<figureCitation id="75C62A69E159FFB2B8BA4B00FED9FDC8" box="[214,281,571,596]" captionStart="Figs 7–21" captionStartId="5.[113,150,1468,1487]" captionTargetBox="[121,1083,222,1444]" captionTargetId="figure-17@5.[117,1084,219,1447]" captionTargetPageId="5" captionText="Figs 7–21. Odontocheila gilli Johnson, 2000. 7 – head, male, Nusagandi (DBCN); 8–10 – labrum: 8 – male, Chepo- Carti Road, HT (CMNC); 9 – male, Nusagandi; 10 – female, Chepo-Carti Road, AT (WJCM). 11–13 – pronotum: 11 – male, HT; 12 – male, Nusagandi; 13 – female, AT. 14–16 – elytron: 14 – male, HT; 15 – male, Nusagandi; 16 – female, AT. 17 – elytral apex, male, HT; 18–19 – elytral apex, back illumination: 18 – male, Nusagandi; 19 – female, AT. 20–21 – aedeagus: 20 – HT; 21 – Nusagandi. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5740579" httpUri="https://zenodo.org/record/5740579/files/figure.png" pageId="3" pageNumber="588">Fig. 7</figureCitation>
|
||
). Both maxillary and labial palpi with normal (elongate) shape of terminal palpomeres, in male yellow-ochre with gradually brownish-testaceous darkened terminal palpomeres, in female much darker and with apices of palpomeres blackish darkened, in female also whole labial palpi blackish darkened ventrally; penultimate (longest) palpomere of labial palpi rather narrow and elongate with only slightly and gradually dilated lateral margins towards apex (width
|
||
<quantity id="2A059B09E159FFB2B9C24BE0FD8DFD68" box="[430,589,731,756]" metricMagnitude="-4" metricUnit="m" metricValue="1.55" metricValueMax="1.6" metricValueMin="1.5" pageId="3" pageNumber="588" unit="mm" value="0.155" valueMax="0.16" valueMin="0.15">0.15–0.16 mm</quantity>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="ED4236ECE159FFB2B8FC4BC7FD3BFC4F" blockId="3.[113,1089,219,1588]" pageId="3" pageNumber="588">
|
||
Antennae very long, in male reaching elytral anteapical angle (the
|
||
<typeStatus id="3246884EE159FFB2BB314BC0FC7AFC8F" box="[861,954,763,787]" pageId="3" pageNumber="588" type="holotype">holotype</typeStatus>
|
||
missing last three antennomeres), in female somewhat shorter (incomplete in the
|
||
<typeStatus id="3246884EE159FFB2BB334A20FC7AFCAF" box="[863,954,795,819]" pageId="3" pageNumber="588" type="allotype">allotype</typeStatus>
|
||
); scape with only subapical seta, in male ochre-testaceous to brownish-testaceous with faint mahogany lustre, in female almost black with feeble metallic-green lustre; antennomeres
|
||
<quantity id="2A059B09E159FFB2BBAD4A67FBC5FCEF" box="[961,1029,859,884]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" metricValueMax="10.16" metricValueMin="5.08" pageId="3" pageNumber="588" unit="in" value="3.0" valueMax="4.0" valueMin="2.0">2–4 in</quantity>
|
||
male brownish-testaceous with mahogany lustre, in female much darker with almost black pedicel; antennomeres 5–11 dark brownish, gradually smoky-black darkened with normal micropubescence; all antennomeres with indistinct short apical setae.
|
||
</paragraph>
|
||
<paragraph id="ED4236ECE159FFB2B8FC4AE0FD21FA2F" blockId="3.[113,1089,219,1588]" pageId="3" pageNumber="588">
|
||
Thorax. Pronotum (
|
||
<figureCitation id="75C62A69E159FFB2B9094AE0FE1FFC68" box="[357,479,987,1012]" captionStart="Figs 7–21" captionStartId="5.[113,150,1468,1487]" captionTargetBox="[121,1083,222,1444]" captionTargetId="figure-17@5.[117,1084,219,1447]" captionTargetPageId="5" captionText="Figs 7–21. Odontocheila gilli Johnson, 2000. 7 – head, male, Nusagandi (DBCN); 8–10 – labrum: 8 – male, Chepo- Carti Road, HT (CMNC); 9 – male, Nusagandi; 10 – female, Chepo-Carti Road, AT (WJCM). 11–13 – pronotum: 11 – male, HT; 12 – male, Nusagandi; 13 – female, AT. 14–16 – elytron: 14 – male, HT; 15 – male, Nusagandi; 16 – female, AT. 17 – elytral apex, male, HT; 18–19 – elytral apex, back illumination: 18 – male, Nusagandi; 19 – female, AT. 20–21 – aedeagus: 20 – HT; 21 – Nusagandi. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5740579" httpUri="https://zenodo.org/record/5740579/files/figure.png" pageId="3" pageNumber="588">Figs 11–13</figureCitation>
|
||
) as long as wide, length and width
|
||
<quantity id="2A059B09E159FFB2BB0A4AE0FBC5FC6F" box="[870,1029,987,1011]" metricMagnitude="-3" metricUnit="m" metricValue="1.65" metricValueMax="1.7" metricValueMin="1.6" pageId="3" pageNumber="588" unit="mm" value="1.65" valueMax="1.7" valueMin="1.6">1.60–1.70 mm</quantity>
|
||
, both anterior and posterior sulci well pronounced; anterior lobe metallic-green with violaceous lateral areas, wider than the posterior, irregularly wavy-rugulose with a transversely vermicular rugulose ornament in middle; disc with lateral margins (including those of dorsally visible proepisterna) moderately convex and subparallel in middle (including inconspicuous but well obvious notopleural sutures); discal surface on median area bright bronze-cupreous with rather distinct transverse rugae, while iridescent blue-green sublateral areas are covered with much shallower rugae which become effaced towards smooth, shiny, purple-violaceous lateral areas; posterior lobe with rather distinct dorsolateral bulges and double-sutured posterior rim, iridescent green-blue with violaceous lustre, irregularly rugulose; all thoracic sterna glabrous; prosternum, mesosternum and metasternum smooth, metallic black-blue with green, bronze, cupreous and violaceous lustre; proepisterna smooth, shiny metallic-black with strong, cupreous lustre; mesepisterna and metepisterna metallic black-green, metepisterna indistinctly coriaceous-wrinkled; female mesepisternal coupling sulci unrecognizable, in form of longitudinal sulcus only somewhat deeper than in male.
|
||
</paragraph>
|
||
<paragraph id="ED4236ECE159FFB9B8FC4C80FE21FD4F" blockId="3.[113,1089,219,1588]" lastBlockId="8.[151,1127,219,1620]" lastPageId="8" lastPageNumber="593" pageId="3" pageNumber="588">
|
||
Elytra (
|
||
<figureCitation id="75C62A69E159FFB2B8B24C80FE97FA48" box="[222,343,1467,1492]" captionStart="Figs 7–21" captionStartId="5.[113,150,1468,1487]" captionTargetBox="[121,1083,222,1444]" captionTargetId="figure-17@5.[117,1084,219,1447]" captionTargetPageId="5" captionText="Figs 7–21. Odontocheila gilli Johnson, 2000. 7 – head, male, Nusagandi (DBCN); 8–10 – labrum: 8 – male, Chepo- Carti Road, HT (CMNC); 9 – male, Nusagandi; 10 – female, Chepo-Carti Road, AT (WJCM). 11–13 – pronotum: 11 – male, HT; 12 – male, Nusagandi; 13 – female, AT. 14–16 – elytron: 14 – male, HT; 15 – male, Nusagandi; 16 – female, AT. 17 – elytral apex, male, HT; 18–19 – elytral apex, back illumination: 18 – male, Nusagandi; 19 – female, AT. 20–21 – aedeagus: 20 – HT; 21 – Nusagandi. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5740579" httpUri="https://zenodo.org/record/5740579/files/figure.png" pageId="3" pageNumber="588">Figs 14–19</figureCitation>
|
||
) elongate, length
|
||
<quantity id="2A059B09E159FFB2BA784C80FD71FA4F" box="[532,689,1467,1491]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" metricValueMax="5.2" metricValueMin="4.8" pageId="3" pageNumber="588" unit="mm" value="5.0" valueMax="5.2" valueMin="4.8">4.80–5.20 mm</quantity>
|
||
, with rounded to subquadrate humeri, lateral margins subparallel, anteapical angles arcuate, then obliquely running towards apices which are in male subacute (only shortly rounded towards sutural spine) in female almost rounded; sutural spine short but distinct; microserrulation indistinct and very irregular; elytral dorsal surface regularly convex on posterior half of elytral disc, humeral impression moderate, discal impression rather distinct, clearly delimiting moderate basodiscal convexity; apical impression distinct; elytral surface punctate, punctures mostly isolated and notably larger within humeral impressions and on basodiscal convexity, indistinctly anastomosing into chains within the discal impression, becoming much smaller, isolated and sparser posteriad and very shallow on posterior declivity towards apices, nearly effaced on anteapical angles; appearance of the punctation varies depending on angle of illumination: the isolated punctures on apical elytral third are much more obvious in back illumination (
|
||
<figureCitation id="75C62A69E152FFB9BBB14880FB9AFE48" box="[989,1114,443,468]" captionStart="Figs 7–21" captionStartId="5.[113,150,1468,1487]" captionTargetBox="[121,1083,222,1444]" captionTargetId="figure-17@5.[117,1084,219,1447]" captionTargetPageId="5" captionText="Figs 7–21. Odontocheila gilli Johnson, 2000. 7 – head, male, Nusagandi (DBCN); 8–10 – labrum: 8 – male, Chepo- Carti Road, HT (CMNC); 9 – male, Nusagandi; 10 – female, Chepo-Carti Road, AT (WJCM). 11–13 – pronotum: 11 – male, HT; 12 – male, Nusagandi; 13 – female, AT. 14–16 – elytron: 14 – male, HT; 15 – male, Nusagandi; 16 – female, AT. 17 – elytral apex, male, HT; 18–19 – elytral apex, back illumination: 18 – male, Nusagandi; 19 – female, AT. 20–21 – aedeagus: 20 – HT; 21 – Nusagandi. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5740579" httpUri="https://zenodo.org/record/5740579/files/figure.png" pageId="8" pageNumber="593">Figs 18–19</figureCitation>
|
||
); elytral surface glabrous except for a few usual hairlike sensory setae indistinctly scattered mostly on basal area, and a few others adjacent to epipleura; elytral coloration on elytral disc shiny metallic yellow-green with bronze-cupreous lustre, sublateral areas bright green to green-blue, and lateral areas purple-violaceous (coloration varies depending on angle of illumination), white elytral maculation in male consisting of only sublateral-median macula and anteapical-apical lunule which in female appears as only anteapical macula because its lunule-like prolongation along the apical margin is in female barely recognizable; humeral macula is absent in both sexes.
|
||
</paragraph>
|
||
<caption id="B9826664E15EFFB5B8FB4CF5FD46F9AD" ID-DOI="http://doi.org/10.5281/zenodo.5740577" ID-Zenodo-Dep="5740577" httpUri="https://zenodo.org/record/5740577/files/figure.png" pageId="4" pageNumber="589" startId="4.[151,188,1486,1505]" targetBox="[185,1089,221,1439]" targetPageId="4">
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||
<paragraph id="ED4236ECE15EFFB5B8FB4CF5FD46F9AD" blockId="4.[151,1127,1486,1585]" pageId="4" pageNumber="589">
|
||
Figs 1–6. Habitus of two species of
|
||
<taxonomicName id="2AFD4D6FE15EFFB5B9D34CF4FCDFFA7D" authority="Laporte de Castelnau, 1834" authorityName="Laporte de Castelnau" authorityYear="1834" box="[447,799,1486,1505]" class="Insecta" family="Carabidae" genus="Odontocheila" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="589" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="DF89EAFEE15EFFB5B9D34CF4FDF3FA7D" box="[447,563,1487,1505]" italics="true" pageId="4" pageNumber="589">Odontocheila</emphasis>
|
||
Laporte de Castelnau, 1834
|
||
</taxonomicName>
|
||
. 1–2 –
|
||
<taxonomicName id="2AFD4D6FE15EFFB5BB344CF4FBE5FA7D" authority="Johnson, 2000: 1" authorityName="Johnson" authorityPageNumber="1" authorityYear="2000" box="[856,1061,1486,1505]" class="Insecta" family="Carabidae" genus="Odontocheila" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="589" phylum="Arthropoda" rank="species" species="gilli">
|
||
<emphasis id="DF89EAFEE15EFFB5BB344CF4FC53FA7D" box="[856,915,1487,1505]" italics="true" pageId="4" pageNumber="589">O. gilli</emphasis>
|
||
<bibRefCitation id="896C4B1DE15EFFB5BBFB4CF5FBE5FA7D" author="JOHNSON W. N." box="[919,1061,1486,1505]" pageId="4" pageNumber="589" pagination="13 - 18" refId="ref7088" refString="JOHNSON W. N. 2000: A new species of Odontocheila from Panama (Coleoptera: Cicindelidae). Cicindela 32: 13 - 18." type="journal article" year="2000">Johnson, 2000:1</bibRefCitation>
|
||
</taxonomicName>
|
||
– male, 8 mm, Chepo-Carti Road, HT (CMNC); 2 – male, 8.1 mm, Nusagandi (DBCN).3–6 –
|
||
<taxonomicName id="2AFD4D6FE15EFFB5BB0D4CD2FC0EFA67" authority="Moravec & Duran, 2013" authorityName="Moravec & Duran" authorityYear="2013" box="[865,974,1513,1531]" class="Insecta" family="Carabidae" genus="Odontocheila" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="589" phylum="Arthropoda" rank="species" species="fraternum" status="sp. nov.">
|
||
<emphasis id="DF89EAFEE15EFFB5BB0D4CD2FC0EFA67" box="[865,974,1513,1531]" italics="true" pageId="4" pageNumber="589">O. fraternum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="C4BA5785E15EFFB5BBBE4CD2FBD6FA60" box="[978,1046,1513,1532]" pageId="4" pageNumber="589" rank="species">sp. nov.</taxonomicNameLabel>
|
||
: 3 – male, 8.9 mm, Darien, HT (USNM); 4 – male, 8.6 mm, Altos de Campana, PT (CCJM); 5 – female, 9.1 mm, Darien, AT (DBCN); 6 – female, 10 mm, Altos de Campana (CCJM).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="B9826664E15FFFB4B81D4C87FCEEF9A6" ID-DOI="http://doi.org/10.5281/zenodo.5740579" ID-Zenodo-Dep="5740579" httpUri="https://zenodo.org/record/5740579/files/figure.png" pageId="5" pageNumber="590" startId="5.[113,150,1468,1487]" targetBox="[121,1083,222,1444]" targetPageId="5">
|
||
<paragraph id="ED4236ECE15FFFB4B81D4C87FCEEF9A6" blockId="5.[113,1090,1468,1594]" pageId="5" pageNumber="590">
|
||
Figs 7–21.
|
||
<taxonomicName id="2AFD4D6FE15FFFB4B8A34C86FE2CFA53" authority="Johnson, 2000" authorityName="Johnson" authorityYear="2000" box="[207,492,1468,1487]" class="Insecta" family="Carabidae" genus="Odontocheila" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="590" phylum="Arthropoda" rank="species" species="gilli">
|
||
<emphasis id="DF89EAFEE15FFFB4B8A34C86FEAAFA53" box="[207,362,1469,1487]" italics="true" pageId="5" pageNumber="590">Odontocheila gilli</emphasis>
|
||
<bibRefCitation id="896C4B1DE15FFFB4B9034C87FE2CFA53" author="JOHNSON W. N." box="[367,492,1468,1487]" pageId="5" pageNumber="590" pagination="13 - 18" refId="ref7088" refString="JOHNSON W. N. 2000: A new species of Odontocheila from Panama (Coleoptera: Cicindelidae). Cicindela 32: 13 - 18." type="journal article" year="2000">Johnson, 2000</bibRefCitation>
|
||
</taxonomicName>
|
||
. 7 – head, male, Nusagandi (DBCN); 8–10 – labrum: 8 – male, Chepo- Carti Road, HT (CMNC); 9 – male, Nusagandi; 10 – female, Chepo-Carti Road, AT (WJCM). 11–13 – pronotum: 11 – male, HT; 12 – male, Nusagandi; 13 – female, AT. 14–16 – elytron: 14 – male, HT; 15 – male, Nusagandi; 16 – female, AT. 17 – elytral apex, male, HT; 18–19 – elytral apex, back illumination: 18 – male, Nusagandi; 19 – female, AT. 20–21 – aedeagus: 20 – HT; 21 – Nusagandi. Scale bars = 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="B9826664E15CFFB7B8FB4C82FBCEF9AB" ID-DOI="http://doi.org/10.5281/zenodo.5740583" ID-Zenodo-Dep="5740583" httpUri="https://zenodo.org/record/5740583/files/figure.png" pageId="6" pageNumber="591" startId="6.[151,188,1465,1484]" targetBox="[175,1104,225,1433]" targetPageId="6">
|
||
<paragraph id="ED4236ECE15CFFB7B8FB4C82FBCEF9AB" blockId="6.[151,1127,1465,1591]" pageId="6" pageNumber="591">
|
||
Figs 22–38.
|
||
<taxonomicName id="2AFD4D6FE15CFFB7B96F4C81FE12FA50" authority="Moravec & Duran, 2013" authorityName="Moravec & Duran" authorityYear="2013" box="[259,466,1466,1484]" class="Insecta" family="Carabidae" genus="Odontocheila" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="591" phylum="Arthropoda" rank="species" species="fraternum" status="sp. nov.">
|
||
<emphasis id="DF89EAFEE15CFFB7B96F4C81FE12FA50" box="[259,466,1466,1484]" italics="true" pageId="6" pageNumber="591">Odontocheila fraternum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="C4BA5785E15CFFB7B9B54C81FDDDFA51" box="[473,541,1466,1485]" pageId="6" pageNumber="591" rank="species">sp. nov.</taxonomicNameLabel>
|
||
22–25 – male labrum: 22 – Darien, HT (USNM); 23–25 – Altos de Campana, PT (23 – DPDC; 24–25 – CCJM). 26 – head, HT. 27–29 – female labrum: 27 – Darien, AT (DBCN); 28 – Darien, PT (CCJM); 29 – Altos de Campana, PT (DBCN). 30–33 – male pronotum: 30 – HT; 31 – Darien, PT (DBCN); 32–33 – Altos de Campana, PT (32 – DBCN; 33 – CCJM). 34 – female pronotum, AT. 35–38 – aedeagus (apex): 35 – HT; 36 – Darien, PT (CCJM); 37–38 – Altos de Campana, PT (CCJM). Scale bars = 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="B9826664E15DFFB6B81D4CEFFEA9F9CE" ID-DOI="http://doi.org/10.5281/zenodo.5740585" ID-Zenodo-Dep="5740585" httpUri="https://zenodo.org/record/5740585/files/figure.png" pageId="7" pageNumber="592" startId="7.[113,150,1492,1511]" targetBox="[128,1082,223,1474]" targetPageId="7">
|
||
<paragraph id="ED4236ECE15DFFB6B81D4CEFFEA9F9CE" blockId="7.[113,1089,1492,1618]" pageId="7" pageNumber="592">
|
||
Figs 39–49.
|
||
<taxonomicName id="2AFD4D6FE15DFFB6B8B34CEEFE6FFA7B" authority="Moravec & Duran, 2013" authorityName="Moravec & Duran" authorityYear="2013" box="[223,431,1493,1511]" class="Insecta" family="Carabidae" genus="Odontocheila" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="592" phylum="Arthropoda" rank="species" species="fraternum" status="sp. nov.">
|
||
<emphasis id="DF89EAFEE15DFFB6B8B34CEEFE6FFA7B" box="[223,431,1493,1511]" italics="true" pageId="7" pageNumber="592">Odontocheila fraternum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="C4BA5785E15DFFB6B9DB4CEFFE3CFA7B" box="[439,508,1492,1511]" pageId="7" pageNumber="592" rank="species">sp. nov.</taxonomicNameLabel>
|
||
39–42 – male elytron: 39 – HT; 40–41 – Darien, PT (DBCN); 42 – Altos de Campana, PT (DBCN). 43–44 – elytral apex, back illumination: 43 – HT; 44 – Altos de Campana, PT (DBCN). 45–46 – female elytron: 45 – Darien, AT; 46 – Altos de Campana, PT (CCJM). 47–49 – cleared aedeagus showing internal sac: 47 – Altos de Campana, PT (CCJM); 48 – Darien, PT (CCJM); 49 – Altos de Campana, PT (DPDC). Scale bars = 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="ED4236ECE152FFB9B8D94BE7FEA2FCC8" blockId="8.[151,1127,219,1620]" pageId="8" pageNumber="593">Abdomen. Ventrites metallic-black with greenish, blue and violaceous lustre, in male with indistinctly testaceous areas on last three ventrites and apical pleurite, surface of the ventrites glabrous (except for usual, sparse and easily abraded hairlike sensory setae at their posterior margins).</paragraph>
|
||
<paragraph id="ED4236ECE152FFB9B8D94A67FC8BFB0F" blockId="8.[151,1127,219,1620]" pageId="8" pageNumber="593">Legs. Coxae in male with pro- and mesocoxae yellow to testaceous, in female dark testaceous, with only a few setae, metacoxae black with metallic-blue lustre and testaceous apices with only one central seta (lateral setae entirely absent); trochanters yellow-testaceous in male, dark testaceous in female, glabrous (except for usual easily abraded apical seta); femora in male yellow to testaceous, dorsally with feeble mahogany lustre, in female much darker, brownish-testaceous; femoral surface with only very sparse, short and indistinct, white to brownish semierect setae; tibiae concolorous with femora and with similar sparse setae and dense pad of greyish setae on basal third to half of pro- and mesotibiae; metatibiae with only sparse, semierect, short and stiffer setae; tarsi concolorous with tibiae except for darkened apices, first three tarsomeres in male with moderate dilatation and usual pad of dense, white setae.</paragraph>
|
||
<paragraph id="ED4236ECE152FFB9B8D94DA7FC5FFAC8" blockId="8.[151,1127,219,1620]" pageId="8" pageNumber="593">
|
||
Aedeagus (
|
||
<figureCitation id="75C62A69E152FFB9B9414DA7FE66FB28" box="[301,422,1180,1204]" captionStart="Figs 7–21" captionStartId="5.[113,150,1468,1487]" captionTargetBox="[121,1083,222,1444]" captionTargetId="figure-17@5.[117,1084,219,1447]" captionTargetPageId="5" captionText="Figs 7–21. Odontocheila gilli Johnson, 2000. 7 – head, male, Nusagandi (DBCN); 8–10 – labrum: 8 – male, Chepo- Carti Road, HT (CMNC); 9 – male, Nusagandi; 10 – female, Chepo-Carti Road, AT (WJCM). 11–13 – pronotum: 11 – male, HT; 12 – male, Nusagandi; 13 – female, AT. 14–16 – elytron: 14 – male, HT; 15 – male, Nusagandi; 16 – female, AT. 17 – elytral apex, male, HT; 18–19 – elytral apex, back illumination: 18 – male, Nusagandi; 19 – female, AT. 20–21 – aedeagus: 20 – HT; 21 – Nusagandi. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5740579" httpUri="https://zenodo.org/record/5740579/files/figure.png" pageId="8" pageNumber="593">Figs 20–21</figureCitation>
|
||
) moderately voluminous in middle,
|
||
<quantity id="2A059B09E152FFB9BB464DA7FC48FB2F" box="[810,904,1180,1204]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" pageId="8" pageNumber="593" unit="mm" value="2.8">2.80 mm</quantity>
|
||
long,
|
||
<quantity id="2A059B09E152FFB9BBA04DA0FBA6FB2F" box="[972,1126,1179,1203]" metricMagnitude="-4" metricUnit="m" metricValue="6.75" metricValueMax="7.0" metricValueMin="6.5" pageId="8" pageNumber="593" unit="mm" value="0.675" valueMax="0.7" valueMin="0.65">0.65–0.70 mm</quantity>
|
||
wide, apical portion with almost straight ventral margin, gradually attenuated towards conspicuously shaped apex which is rounded, dorsally obliquely sloped and sharpened due to small but deep dorsal excision forming small, crochet hook-like shape. Internal sac not examined from cleared aedeagus, but when the aedeagus was observed re-hydrated by water, it showed usual sclerites and long convoluted flagellum characteristic of the genus.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="A5E76567E152FFB8B8FB4C60FDEBFE8F" lastPageId="9" lastPageNumber="594" pageId="8" pageNumber="593" type="diagnosis">
|
||
<paragraph id="ED4236ECE152FFB9B8FB4C60FEDCFA68" blockId="8.[151,1127,219,1620]" pageId="8" pageNumber="593">
|
||
<emphasis id="DF89EAFEE152FFB9B8FB4C60FE58FAEF" bold="true" box="[151,408,1371,1395]" pageId="8" pageNumber="593">Differential diagnosis.</emphasis>
|
||
Immediately recognizable from externally similar
|
||
<taxonomicName id="2AFD4D6FE152FFB9BBBF4C66FDE3FA08" authority="Laporte de Castelnau, 1834" authorityName="Laporte de Castelnau" authorityYear="1834" class="Insecta" family="Carabidae" genus="Odontocheila" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="593" phylum="Arthropoda" rank="species" species="mexicana">
|
||
<emphasis id="DF89EAFEE152FFB9BBBF4C66FF3DFA08" italics="true" pageId="8" pageNumber="593">Odontocheila mexicana</emphasis>
|
||
Laporte de Castelnau, 1834
|
||
</taxonomicName>
|
||
by the unusual shape of the male labrum and aedeagus, and absence of humeral lunule in both sexes. These characters and also its elytral sculpture differentiate
|
||
<taxonomicName id="2AFD4D6FE152FFB9B9704C86FEA5FA48" authorityName="Johnson" authorityYear="2000" box="[284,357,1469,1492]" class="Insecta" family="Carabidae" genus="Odontocheila" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="593" phylum="Arthropoda" rank="species" species="gilli">
|
||
<emphasis id="DF89EAFEE152FFB9B9704C86FEA5FA48" box="[284,357,1469,1492]" italics="true" pageId="8" pageNumber="593">O. gilli</emphasis>
|
||
</taxonomicName>
|
||
from the other similarly coloured Central American species
|
||
<taxonomicName id="2AFD4D6FE152FFB9BBB84C86FED9FA68" authority="Bates, 1872" authorityName="Bates" authorityYear="1872" class="Insecta" family="Carabidae" genus="Odontocheila" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="593" phylum="Arthropoda" rank="species" species="iodopleura">
|
||
<emphasis id="DF89EAFEE152FFB9BBB84C86FBA7FA48" box="[980,1127,1469,1492]" italics="true" pageId="8" pageNumber="593">O. iodopleura</emphasis>
|
||
Bates, 1872
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="ED4236ECE152FFB8B8D94CC6FDEBFE8F" blockId="8.[151,1127,219,1620]" lastBlockId="9.[113,1089,220,500]" lastPageId="9" lastPageNumber="594" pageId="8" pageNumber="593">
|
||
<taxonomicName id="2AFD4D6FE152FFB9B8D94CC6FEB7F988" authorityName="Johnson" authorityYear="2000" box="[181,375,1533,1556]" class="Insecta" family="Carabidae" genus="Odontocheila" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="593" phylum="Arthropoda" rank="species" species="gilli">
|
||
<emphasis id="DF89EAFEE152FFB9B8D94CC6FEB7F988" box="[181,375,1533,1556]" italics="true" pageId="8" pageNumber="593">Odontocheila gilli</emphasis>
|
||
</taxonomicName>
|
||
shares the characteristic shape of the labrum and aedeagus with
|
||
<taxonomicName id="2AFD4D6FE152FFB9BC734CC6FF22F9A8" authority="Moravec & Duran, 2013" authorityName="Moravec & Duran" authorityYear="2013" class="Insecta" family="Carabidae" genus="Odontocheila" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="593" phylum="Arthropoda" rank="species" species="fraternum" status="sp. nov.">
|
||
<emphasis id="DF89EAFEE152FFB9BC734CC6FF22F9A8" italics="true" pageId="8" pageNumber="593">O. fraternum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="C4BA5785E152FFB9B88B4F27FEF8F9AF" box="[231,312,1564,1587]" pageId="8" pageNumber="593" rank="species">sp. nov.</taxonomicNameLabel>
|
||
described here, but immediately differs from the new species in having elytral punctures mostly isolated, sparser and smaller towards elytral apices, and by the absence of humeral macula in both sexes. In addition, the labrum and palpi of the female
|
||
<typeStatus id="3246884EE153FFB8BBC249E7FBC4FF68" box="[942,1028,220,244]" pageId="9" pageNumber="594" type="allotype">allotype</typeStatus>
|
||
of
|
||
<taxonomicName id="2AFD4D6FE153FFB8BC4B49E6FF5CFE88" authorityName="Johnson" authorityYear="2000" class="Insecta" family="Carabidae" genus="Odontocheila" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="594" phylum="Arthropoda" rank="species" species="gilli">
|
||
<emphasis id="DF89EAFEE153FFB8BC4B49E6FF5CFE88" italics="true" pageId="9" pageNumber="594">O. gilli</emphasis>
|
||
</taxonomicName>
|
||
are much darker with blackish areas.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="A5E76567E153FFB8B81D4820FE43FE68" pageId="9" pageNumber="594" type="distribution">
|
||
<paragraph id="ED4236ECE153FFB8B81D4820FBD6FEE8" blockId="9.[113,1089,220,500]" pageId="9" pageNumber="594">
|
||
<emphasis id="DF89EAFEE153FFB8B81D4820FE52FEAF" bold="true" box="[113,402,283,308]" pageId="9" pageNumber="594">Distribution and habitat.</emphasis>
|
||
<taxonomicName id="2AFD4D6FE153FFB8B9F74826FDA0FEA8" authorityName="Johnson" authorityYear="2000" box="[411,608,285,308]" class="Insecta" family="Carabidae" genus="Odontocheila" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="594" phylum="Arthropoda" rank="species" species="gilli">
|
||
<emphasis id="DF89EAFEE153FFB8B9F74826FDA0FEA8" box="[411,608,285,308]" italics="true" pageId="9" pageNumber="594">Odontocheila gilli</emphasis>
|
||
</taxonomicName>
|
||
is a very rare species. Apart from
|
||
<specimenCount id="FBFBFD65E153FFB8BB8E4827FB81FEA8" box="[994,1089,284,308]" pageId="9" pageNumber="594" type="generic" typeStatus="type">two type</specimenCount>
|
||
specimens, only one other male was caught by David D. Brzoska (Naples,
|
||
<collectingRegion id="2F39F80EE153FFB8BB104807FC0BFEC8" box="[892,971,316,340]" country="United States of America" name="Florida" pageId="9" pageNumber="594">Florida</collectingRegion>
|
||
) within the Nusagandi Indian Reserve, San Blas (
|
||
<collectingRegion id="2F39F80EE153FFB8BA684867FDB7FEE8" box="[516,631,348,372]" country="Panama" name="Kuna Yala" pageId="9" pageNumber="594">Kuna Yala</collectingRegion>
|
||
) region, situated near the
|
||
<typeStatus id="3246884EE153FFB8BBE74867FC79FEE8" box="[907,953,348,372]" pageId="9" pageNumber="594">type</typeStatus>
|
||
locality.
|
||
</paragraph>
|
||
<paragraph id="ED4236ECE153FFB8B8FC4847FE43FE68" blockId="9.[113,1089,220,500]" pageId="9" pageNumber="594">
|
||
The habitat and collecting circumstances were mentioned by
|
||
<bibRefCitation id="896C4B1DE153FFB8BB444847FC0FFE08" author="JOHNSON W. N." box="[808,975,380,404]" pageId="9" pageNumber="594" pagination="13 - 18" refId="ref7088" refString="JOHNSON W. N. 2000: A new species of Odontocheila from Panama (Coleoptera: Cicindelidae). Cicindela 32: 13 - 18." type="journal article" year="2000">JOHNSON (2000)</bibRefCitation>
|
||
according to information given by the collector Bruce Gill (Ottawa,
|
||
<collectingRegion id="2F39F80EE153FFB8BABF48A7FCE7FE28" box="[723,807,412,436]" country="Canada" name="Ontario" pageId="9" pageNumber="594">Ontario</collectingRegion>
|
||
,
|
||
<collectingCountry id="95EA767CE153FFB8BB5D48A7FC43FE28" box="[817,899,412,436]" name="Canada" pageId="9" pageNumber="594">Canada</collectingCountry>
|
||
) that the
|
||
<specimenCount id="FBFBFD65E153FFB8BB8948A7FB81FE28" box="[997,1089,412,436]" pageId="9" pageNumber="594" type="generic" typeStatus="type">two type</specimenCount>
|
||
specimens were taken in a “flight intercept trap” placed along trails within tropical rain forest of the El Llano-Carti site.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |