treatments-xml/data/03/E5/87/03E587F9FFA0FF9CFF06F9B218A3418B.xml

<document id="28824F76A51F098A37363624A169AE56" ID-CLB-Dataset="8217" ID-DOI="10.11646/zootaxa.4859.1.3" ID-GBIF-Dataset="ff9ddcdc-1340-4533-ae1e-da83b895158c" ID-ISSN="1175-5326" ID-Zenodo-Dep="4412809" ID-ZooBank="C213B95B-6707-4F80-A1EA-58245141F24B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1609692846665" checkinUser="plazi" docAuthor="Dallevo-Gomes, Caio I. A., Mattox, George M. T. &amp; Toledo-Piza, Mônica" docDate="2020" docId="03E587F9FFA0FF9CFF06F9B218A3418B" docLanguage="en" docName="zootaxa.4859.1.3.pdf" docOrigin="Zootaxa 4859 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pseudophallus elcapitanensis" docType="treatment" docVersion="7" lastPageNumber="94" masterDocId="FFDCFF81FFAAFF91FF91FF861C704562" masterDocTitle="Taxonomic review of the pipefish genus Pseudophallus Herald, with the description of a new species (Syngnathiformes: Syngnathidae)" masterLastPageNumber="112" masterPageNumber="81" pageNumber="91" updateTime="1698851676651" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="4FDB2C25529AC4BAA028554914AB382D" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="FDE807A62163C75BC44F616C9622BDDC">
<mods:title id="DBA1004DB00AC4031F94ECBF3A7218D7">Taxonomic review of the pipefish genus Pseudophallus Herald, with the description of a new species (Syngnathiformes: Syngnathidae)</mods:title>
</mods:titleInfo>
<mods:name id="7C74706DA70472368299BFEE8F01554F" type="personal">
<mods:role id="E1E5B28FEB901E3777BBF1BEF47404F9">
<mods:roleTerm id="07BA32BEAADF20DCBE55F351D5F5E45F">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="92F67A7DE9D07DAE113A549E952C6450">Dallevo-Gomes, Caio I. A.</mods:namePart>
<mods:nameIdentifier id="FD32531EB6DF9DE7607E17084839A506" type="ORCID">0000-0001-7788-5861</mods:nameIdentifier>
<mods:affiliation id="7D1E1087751D9F22C20C7E361415B500">Departamento de Zoologia, Instituto de Biociências da Universidade de São Paulo, Rua do Matão, travessa 14, no. 101, 05508 - 090, São Paulo, SP Brazil. &amp; caioisola @ yahoo. com. br; https: // orcid. org / 0000 - 0001 - 7788 - 5861</mods:affiliation>
<mods:nameIdentifier id="7C2D948FF850F4CD9FA8718B032671C2" type="email">caioisola@yahoo.com.br</mods:nameIdentifier>
</mods:name>
<mods:name id="A0C9A19B53922BB01447AAE1B4EC3CE8" type="personal">
<mods:role id="537AA1154D08771CFDE7216E291AD447">
<mods:roleTerm id="2CED18DD628B4F95045959EA357B0000">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="E9C5A8E82C65D302B89B8764EB7075C5">Mattox, George M. T.</mods:namePart>
<mods:affiliation id="589B908AB46AC6BCFBB2B2604EFD44BA">Departamento de Biologia, Universidade Federal de São Carlos, Campus Sorocaba, São Paulo, Brazil.</mods:affiliation>
</mods:name>
<mods:name id="35A46944091D14110F366E22468F59A4" type="personal">
<mods:role id="0ECFFAF7EDAE58000CE3FE4A98182F60">
<mods:roleTerm id="1A62CBE008E5A75005F631231D0B8BCA">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="616854BF5E69F843B62D9D4014683D84">Toledo-Piza, Mônica</mods:namePart>
<mods:nameIdentifier id="BE6E90092EE6BB8A899463569EDC2B66" type="ORCID">0000-0001-7596-1410</mods:nameIdentifier>
<mods:affiliation id="F861F10B94FC0E3AAF920F67FE9FAA63">Departamento de Zoologia, Instituto de Biociências da Universidade de São Paulo, Rua do Matão, travessa 14, no. 101, 05508 - 090, São Paulo, SP Brazil. &amp; mtpiza @ usp. br; https: // orcid. org / 0000 - 0001 - 7596 - 1410</mods:affiliation>
<mods:nameIdentifier id="B091AE5C560B96E118E0B6AB29A5FB3E" type="email">mtpiza@usp.br</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="F231B104673290068947F9D49A37055B">text</mods:typeOfResource>
<mods:relatedItem id="F0C6682E9824BAA9595E25890F17E07F" type="host">
<mods:titleInfo id="EA6A3765AEA8F3921FA8454F19BE8359">
<mods:title id="31D4338DFEDAEE843BC69F21EE5DF913">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="1B856936358F2AAA299AB87EE6ACD457">
<mods:date id="E892F4ED9B758BC1958CB7AC8DA7327D">2020</mods:date>
<mods:detail id="D0E5CCA690BD23E46663AE71D7D785D2" type="pubDate">
<mods:number id="FDE93773627CA20154481D8A60BE8A31">2020-10-05</mods:number>
</mods:detail>
<mods:detail id="9E0D970CE41189462FF4CA9323763B4D" type="volume">
<mods:number id="387E54ED6826CE22B01FD44229B8607B">4859</mods:number>
</mods:detail>
<mods:detail id="100A7B0ACE35B5AB76AACB0119BA45D1" type="issue">
<mods:number id="B31ABE605A9DDC69021F766D60B586E2">1</mods:number>
</mods:detail>
<mods:extent id="CE1487D7F8A619F96B81DA3EF4F5637A" unit="page">
<mods:start id="F4C1D64A57158EF32387FF814FB9F313">81</mods:start>
<mods:end id="5E5CAAA809198B0E5DB901564C5FAFFB">112</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="7F7F0D7976BE216B7B0DDE7B0F913F4C">journal article</mods:classification>
<mods:identifier id="0414A343F132317A90F69B5CF6690F37" type="CLB-Dataset">8217</mods:identifier>
<mods:identifier id="EAE93394B08162FE1D105278D2D56151" type="DOI">10.11646/zootaxa.4859.1.3</mods:identifier>
<mods:identifier id="6BAC7CB2D99213F810CAEB4F67A29ACC" type="GBIF-Dataset">ff9ddcdc-1340-4533-ae1e-da83b895158c</mods:identifier>
<mods:identifier id="DF502C3AEC977FA0A3C252292BF840DE" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="1A3CDB1B252F75ADC6BA3565DB36DBB0" type="Zenodo-Dep">4412809</mods:identifier>
<mods:identifier id="86A979D166EED184A4497F2E3B6D16A3" type="ZooBank">C213B95B-6707-4F80-A1EA-58245141F24B</mods:identifier>
</mods:mods>
<treatment id="03E587F9FFA0FF9CFF06F9B218A3418B" ID-DOI="http://doi.org/10.5281/zenodo.4412821" ID-GBIF-Taxon="177326236" ID-Zenodo-Dep="4412821" LSID="urn:lsid:plazi:treatment:03E587F9FFA0FF9CFF06F9B218A3418B" httpUri="http://treatment.plazi.org/id/03E587F9FFA0FF9CFF06F9B218A3418B" lastPageId="13" lastPageNumber="94" pageId="10" pageNumber="91">
<subSubSection id="C3566564FFA0FF9BFF06F9B21F26432C" box="[151,854,1588,1615]" pageId="10" pageNumber="91" type="nomenclature">
<paragraph id="8BF336EFFFA0FF9BFF06F9B21F26432C" blockId="10.[151,854,1588,1651]" box="[151,854,1588,1615]" pageId="10" pageNumber="91">
<heading id="D0BB8183FFA0FF9BFF06F9B21F26432C" bold="true" box="[151,854,1588,1615]" fontSize="11" level="1" pageId="10" pageNumber="91" reason="1">
<taxonomicName id="4C4C4D6CFFA0FF9BFF06F9B21F26432C" ID-CoL="4P6RF" authority="(Meek &amp; Hildebrand 1914)" baseAuthorityName="Meek &amp; Hildebrand" baseAuthorityYear="1914" box="[151,854,1588,1615]" class="Actinopterygii" family="Syngnathidae" genus="Pseudophallus" kingdom="Animalia" order="Syngnathiformes" pageId="10" pageNumber="91" phylum="Chordata" rank="species" species="elcapitanensis">
<emphasis id="B938EAFDFFA0FF9BFF06F9B21F26432C" bold="true" box="[151,854,1588,1615]" pageId="10" pageNumber="91">
<emphasis id="B938EAFDFFA0FF9BFF06F9B21D8B432D" bold="true" box="[151,507,1588,1615]" italics="true" pageId="10" pageNumber="91">Pseudophallus elcapitanensis</emphasis>
(Meek &amp; Hildebrand 1914)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3566564FFA0FF9BFF06F9DE1DDC4310" box="[151,428,1624,1651]" pageId="10" pageNumber="91" type="description">
<paragraph id="8BF336EFFFA0FF9BFF06F9DE1DDC4310" blockId="10.[151,854,1588,1651]" box="[151,428,1624,1651]" pageId="10" pageNumber="91">
(
<figureCitation id="13772A6AFFA0FF9BFF0EF9DE1D7B4310" box="[159,267,1624,1650]" captionStart-0="FIGURE 2" captionStart-1="FIGURE 3" captionStartId-0="6.[151,250,1798,1822]" captionStartId-1="7.[151,250,1816,1840]" captionTargetBox-0="[164,1430,358,1762]" captionTargetBox-1="[375,1220,194,1786]" captionTargetId-0="figure-100@6.[151,1436,345,1775]" captionTargetId-1="figure-17@7.[361,1226,181,1798]" captionTargetPageId-0="6" captionTargetPageId-1="7" captionText-0="FIGURE 2: Species of the genus Pseudophallus: A—Pseudophallus brasiliensis (paratype) USNM 212058, 69.1 mm SL, Igarapé Inó, Furo de Panaquera, Rio Tocantins, Pará, Brazil. B—Pseudophallus elcapitanensis MCZ 88623, 110.6 mm SL, Río Picuro, Darien, Panama. C—Pseudophallus mindii (holotype) USNM 81770, 90.3 mm SL, Mindi Creek, Canal Zone, Panama. D—Pseudophallus starksii USNM 208365, 171.1 mm SL, upper tributary of Río Jaque, Província de Darien, Panama. E—Pseudophallus galadrielae. (holotype) FMNH 126156, 63.1mm SL, Lago Izabal, Guatemala. Red arrow indicates the anal papilla." captionText-1="FIGURE 3: Detail of anterior portion of body of species of the genus Pseudophallus A—Pseudophallus brasiliensis (paratype) USNM 212058, 69.1 mm SL, Igarapé Inó Furo de Panaquera, Rio Tocantins, Pará, Brazil. B—Pseudophallus elcapitanensis MCZ 88623, 110.6 mm SL, Río Picuro, Darien, Panama. C—Pseudophallus mindii (holotype) USNM 81770, 90.3 mm SL, Mindi Creek, Canal Zone, Panama. D—Pseudophallus starksii USNM 208365, 171.1 mm SL, upper tributary of Río Jaque, Província de Darien, Panama. E—Pseudophallus galadrielae. (holotype), FMNH 126156, 63.1 mm SL, Lago Izabal, Guatemala." figureDoi-0="http://doi.org/10.5281/zenodo.4412815" figureDoi-1="http://doi.org/10.5281/zenodo.4412817" httpUri-0="https://zenodo.org/record/4412815/files/figure.png" httpUri-1="https://zenodo.org/record/4412817/files/figure.png" pageId="10" pageNumber="91">Figs. 2–3</figureCitation>
, 
<figureCitation id="13772A6AFFA0FF9BFE86F9DE1D574311" box="[279,295,1624,1651]" captionStart="FIGURE 5" captionStartId="12.[151,250,1045,1069]" captionTargetBox="[349,1239,321,1005]" captionTargetId="figure-77@12.[340,1247,309,1022]" captionTargetPageId="12" captionText="FIGURE 5. Distribution map of species of Pseudophallus occurring in drainages along the Pacific coast based on specimens examined in this study." figureDoi="http://doi.org/10.5281/zenodo.4412823" httpUri="https://zenodo.org/record/4412823/files/figure.png" pageId="10" pageNumber="91">5</figureCitation>
, 
<figureCitation id="13772A6AFFA0FF9BFEA2F9DF1D334311" box="[307,323,1625,1651]" captionStart="FIGURE 7" captionStartId="25.[151,250,633,657]" captionTargetBox="[154,1431,185,607]" captionTargetId="figure-17@25.[151,1436,181,610]" captionTargetPageId="25" captionText="FIGURE 7. Maps of distribution of specimens of Pseudophallus available for study. Left side: distribution of specimens examined for the morphological analysis in this study. Right side map: distribution of voucher specimens with genetic sequences on GenBank (as of April 2020)." figureDoi="http://doi.org/10.5281/zenodo.4412829" httpUri="https://zenodo.org/record/4412829/files/figure.png" pageId="10" pageNumber="91">7</figureCitation>
; 
<tableCitation id="C6CE0354FFA0FF9BFEDEF9DE1DD34310" box="[335,419,1624,1650]" captionStart="TABLE 2" captionStartId="4.[151,239,151,175]" captionTargetPageId="4" captionText="TABLE 2. Morphometric and meristic data of Pseudophallus species of the Pacific drainages. Standard length (SL) in millimeters, 2–10 are percentages of SL, 11–17 are percentages of head length. N= number of specimens analyzed." pageId="10" pageNumber="91">Table 2</tableCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3566564FFA0FF9AFF06F91B1E14477D" lastPageId="11" lastPageNumber="92" pageId="10" pageNumber="91" type="reference_group">
<paragraph id="8BF336EFFFA0FF9BFF06F91B1D9A43B6" blockId="10.[151,1437,1693,2024]" pageId="10" pageNumber="91">
<treatmentCitationGroup id="AB5C11C1FFA0FF9BFF06F91B1D9A43B6" pageId="10" pageNumber="91">
<taxonomicName id="4C4C4D6CFFA0FF9BFF06F91B1D1843D7" authorityName="Swain" authorityYear="1882" box="[151,360,1693,1717]" class="Polychaeta" family="Flabelligeridae" genus="Siphostoma" kingdom="Animalia" order="Terebellida" pageId="10" pageNumber="91" phylum="Annelida" rank="species" species="auliscus">
<emphasis id="B938EAFDFFA0FF9BFF06F91B1D1843D7" box="[151,360,1693,1717]" italics="true" pageId="10" pageNumber="91">Siphostoma auliscus</emphasis>
</taxonomicName>
(non Swain): 
<treatmentCitation id="0AED10FEFFA0FF9BFE6DF91B1F7E43D7" author="Gilbert, C. H. &amp; Starks, E. C." box="[508,782,1693,1717]" page="57" pageId="10" pageNumber="91" year="1904">
<bibRefCitation id="EFDD4B1EFFA0FF9BFE6DF91B1F7E43D7" author="Gilbert, C. H. &amp; Starks, E. C." box="[508,782,1693,1717]" pageId="10" pageNumber="91" pagination="1 - 304" refId="ref26422" refString="Gilbert, C. H. &amp; Starks, E. C. (1904) The fishes of Panama Bay. Memoir of the California Academy of Sciences, 1, 1 - 304." type="journal article" year="1904">Gilbert &amp; Starks, 1904: 57</bibRefCitation>
</treatmentCitation>
(Rio Grande, Miraflores, near 
<collectingCountry id="F35B767FFFA0FF9BFBDEF91B18D343D6" box="[1103,1187,1693,1716]" name="Panama" pageId="10" pageNumber="91">Panama</collectingCountry>
; comparison with specimens from Magdalena Bay).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BF336EFFFA0FF9BFF06F95C1F124390" blockId="10.[151,1437,1693,2024]" box="[151,866,1754,1779]" pageId="10" pageNumber="91">
<treatmentCitationGroup id="AB5C11C1FFA0FF9BFF06F95C1F124390" box="[151,866,1754,1779]" pageId="10" pageNumber="91">
<taxonomicName id="4C4C4D6CFFA0FF9BFF06F95C1D154390" ID-CoL="53WKN" baseAuthorityName="Swain" baseAuthorityYear="1882" box="[151,357,1754,1778]" class="Actinopterygii" family="Syngnathidae" genus="Syngnathus" kingdom="Animalia" order="Syngnathiformes" pageId="10" pageNumber="91" phylum="Chordata" rank="species" species="auliscus">
<emphasis id="B938EAFDFFA0FF9BFF06F95C1D154390" box="[151,357,1754,1778]" italics="true" pageId="10" pageNumber="91">Syngnathus auliscus</emphasis>
</taxonomicName>
(non Swain): 
<treatmentCitation id="0AED10FEFFA0FF9BFE67F95D1EEC4390" author="Regan, C. T." box="[502,668,1754,1779]" page="53" pageId="10" pageNumber="91" year="1908">
<bibRefCitation id="EFDD4B1EFFA0FF9BFE67F95D1EEC4390" author="Regan, C. T." box="[502,668,1754,1779]" pageId="10" pageNumber="91" pagination="1 - 203" refId="ref29147" refString="Regan, C. T. (1908) Pisces. Part 193. In: Godman, F. D. &amp; Salvin, O. (Eds.), Biologia Centrali-Americana. Published for the editors by R. H. Porter, London, pp. 1 - 203, 15 pls." type="journal article" year="1908">Regan, 1908: 53</bibRefCitation>
</treatmentCitation>
(brief description).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BF336EFFFA0FF9BFF06F97F1E90422C" blockId="10.[151,1437,1693,2024]" pageId="10" pageNumber="91">
<treatmentCitationGroup id="AB5C11C1FFA0FF9BFF06F97F1E90422C" pageId="10" pageNumber="91">
<treatmentCitation id="0AED10FEFFA0FF9BFF06F97F1F4F4273" author="Meek, S. E." box="[151,831,1785,1809]" page="119" pageId="10" pageNumber="91" year="1914">
<taxonomicName id="4C4C4D6CFFA0FF9BFF06F97F1F4F4273" authority="Meek &amp; Hildebrand in Meek, 1914: 119" authorityName="Meek &amp; Hildebrand in Meek" authorityPageNumber="119" authorityYear="1914" box="[151,831,1785,1809]" class="Polychaeta" family="Flabelligeridae" genus="Siphostoma" kingdom="Animalia" order="Terebellida" pageId="10" pageNumber="91" phylum="Annelida" rank="species" species="elcapitanense">
<emphasis id="B938EAFDFFA0FF9BFF06F97F1DEF4273" box="[151,415,1785,1809]" italics="true" pageId="10" pageNumber="91">Siphostoma elcapitanense</emphasis>
Meek &amp; Hildebrand 
<emphasis id="B938EAFDFFA0FF9BFDE8F97F1EFD4272" box="[633,653,1785,1808]" italics="true" pageId="10" pageNumber="91">in</emphasis>
<bibRefCitation id="EFDD4B1EFFA0FF9BFD02F97F1F4F4273" author="Meek, S. E." box="[659,831,1785,1809]" pageId="10" pageNumber="91" pagination="101 - 134" refId="ref28392" refString="Meek, S. E. (1914) An annotated list of fishes known to occur in the fresh waters of Costa Rica. Field Museum of Natural History, Zoological Series, 10, 101 - 134. https: // doi. org / 10.5962 / bhl. title. 5576" type="journal article" year="1914">Meek, 1914: 119</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(original description; type locality: “El Capitan, 
<collectingCountry id="F35B767FFFA0FF9BFAA0F97F19F84273" box="[1329,1416,1785,1809]" name="Panama" pageId="10" pageNumber="91">Panama</collectingCountry>
”); 
<treatmentCitation id="0AED10FEFFA0FF9BFF56F89E1D1D4252" author="Grey, M." box="[199,365,1816,1840]" page="187" pageId="10" pageNumber="91" year="1947">
<bibRefCitation id="EFDD4B1EFFA0FF9BFF56F89E1D1D4252" author="Grey, M." box="[199,365,1816,1840]" pageId="10" pageNumber="91" pagination="109 - 205" refId="ref26660" refString="Grey, M. (1947) Catalogue of type specimens of fishes in Chicago Natural History Museum. Fieldiana, Zoology, 32, 109 - 205." type="journal article" year="1947">Grey, 1947: 187</bibRefCitation>
</treatmentCitation>
(type specimens, listed); 
<treatmentCitation id="0AED10FEFFA0FF9BFDE7F89E1F45424D" author="Dawson, C. E." box="[630,821,1815,1840]" page="46" pageId="10" pageNumber="91" year="1982">
<bibRefCitation id="EFDD4B1EFFA0FF9BFDE7F89E1F45424D" author="Dawson, C. E." box="[630,821,1815,1840]" pageId="10" pageNumber="91" pagination="1 - 172" refId="ref25847" refString="Dawson, C. E. (1982) Fishes of the Western North Atlantic. The pipefishes (subfamilies Doryrhamphinae and Syngnathinae). Memoirs of the Sears Foundation for Marine Research, 1 (8), 1 - 172." type="journal article" year="1982">Dawson, 1982: 46</bibRefCitation>
</treatmentCitation>
, 53 (as a synonymy of 
<taxonomicName id="4C4C4D6CFFA0FF9BFBB6F89E18A2424D" ID-CoL="4P6RF" baseAuthorityName="Meek &amp; Hildebrand" baseAuthorityYear="1914" box="[1063,1234,1815,1839]" class="Actinopterygii" family="Syngnathidae" genus="Pseudophallus" kingdom="Animalia" order="Syngnathiformes" pageId="10" pageNumber="91" phylum="Chordata" rank="species" species="elcapitanensis">
<emphasis id="B938EAFDFFA0FF9BFBB6F89E18A2424D" box="[1063,1234,1815,1839]" italics="true" pageId="10" pageNumber="91">P. elcapitanensis</emphasis>
</taxonomicName>
); 
<bibRefCitation id="EFDD4B1EFFA0FF9BFB78F8911D6B422C" author="Ibarra, M. &amp; Stewart, D. J." pageId="10" pageNumber="91" pagination="1 - 112" refId="ref27089" refString="Ibarra, M. &amp; Stewart, D. J. (1987) Catalogue of type specimens of recent fishes in Field Museum of Natural History. Fieldiana Zoology, New Series, 35, 1 - 112. https: // doi. org / 10.5962 / bhl. title. 3000" type="journal article" year="1987">Ibarra &amp; Stewart, 1987:80</bibRefCitation>
(listed as 
<typeStatus id="54F7884DFFA0FF9BFE13F8B11D93422D" box="[386,483,1847,1871]" pageId="10" pageNumber="91" type="paratype">paratypes</typeStatus>
at FMNH, see remarks).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BF336EFFFA0FF9BFF06F8D31FD6428A" blockId="10.[151,1437,1693,2024]" pageId="10" pageNumber="91">
<treatmentCitationGroup id="AB5C11C1FFA0FF9BFF06F8D31FD6428A" pageId="10" pageNumber="91">
<treatmentCitation id="0AED10FEFFA0FF9BFF06F8D31E83420F" author="Meek, S. &amp; Hildebrand, S. F." box="[151,755,1877,1901]" page="260" pageId="10" pageNumber="91" year="1923">
<taxonomicName id="4C4C4D6CFFA0FF9BFF06F8D31E83420F" authority=": Meek &amp; Hildebrand, 1923: 260" authorityName=": Meek &amp; Hildebrand" authorityPageNumber="260" authorityYear="1923" box="[151,755,1877,1901]" class="Actinopterygii" family="Syngnathidae" genus="Syngnathus" kingdom="Animalia" order="Syngnathiformes" pageId="10" pageNumber="91" phylum="Chordata" rank="species" species="elcapitanense">
<emphasis id="B938EAFDFFA0FF9BFF06F8D31DEF420F" box="[151,415,1877,1901]" italics="true" pageId="10" pageNumber="91">Syngnathus elcapitanense</emphasis>
: 
<bibRefCitation id="EFDD4B1EFFA0FF9BFE3CF8D31E83420F" author="Meek, S. &amp; Hildebrand, S. F." box="[429,755,1877,1901]" pageId="10" pageNumber="91" pagination="1 - 330" refId="ref28446" refString="Meek, S. &amp; Hildebrand, S. F. (1923) The marine fishes of Panama. Part 1. Field Museum of Natural History, Zoological Series, 15, 1 - 330." type="journal article" year="1923">Meek &amp; Hildebrand, 1923: 260</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, pl. 19 (new combination; description; distribution in 
<collectingCountry id="F35B767FFFA0FF9BFAB1F8D31900420E" box="[1312,1392,1877,1900]" name="Panama" pageId="10" pageNumber="91">Panama</collectingCountry>
and 
<collectingCountry id="F35B767FFFA0FF9BFF56F8F21D4842E9" box="[199,312,1907,1932]" name="Costa Rica" pageId="10" pageNumber="91">Costa Rica</collectingCountry>
, USNM 81735 indicated as 
<typeStatus id="54F7884DFFA0FF9BFDC7F8F21EF642EE" box="[598,646,1908,1932]" pageId="10" pageNumber="91">type</typeStatus>
); 
<treatmentCitation id="0AED10FEFFA0FF9BFD06F8F51F3F42E9" author="Breder, C. M." box="[663,847,1907,1932]" page="136" pageId="10" pageNumber="91" year="1927">
<bibRefCitation id="EFDD4B1EFFA0FF9BFD06F8F51F3F42E9" author="Breder, C. M." box="[663,847,1907,1932]" pageId="10" pageNumber="91" pagination="91 - 176" refId="ref25304" refString="Breder, C. M. (1927) The fishes of the Rio Chacunaque drainage, eastern Panama. Bulletin of the American Museum of Natural History, 57, 91 - 176." type="journal article" year="1927">Breder, 1927: 136</bibRefCitation>
</treatmentCitation>
, 144, 148, 159, 165 (in key, euryhaline, behavior, listed in Río Chico, 
<collectingCountry id="F35B767FFFA0FF9BFEACF8141DE242C8" box="[317,402,1938,1962]" name="Panama" pageId="10" pageNumber="91">Panama</collectingCountry>
); 
<treatmentCitation id="0AED10FEFFA0FF9BFE32F8141EE542C8" author="Jordan, D. S. &amp; Evermann B. W. &amp; Clark, H. W." box="[419,661,1938,1962]" page="242" pageId="10" pageNumber="91" year="1930">
<bibRefCitation id="EFDD4B1EFFA0FF9BFE32F8141EE542C8" author="Jordan, D. S. &amp; Evermann B. W. &amp; Clark, H. W." box="[419,661,1938,1962]" pageId="10" pageNumber="91" pagination="1 - 670" refId="ref27499" refString="Jordan, D. S., Evermann B. W. &amp; Clark, H. W. (1930) Check list of the fishes and fishlike vertebrates of North and Middle America north of the northern boundary of Venezuela and Colombia. Report of the United States Commissioner of Fisheries for 1928, Part II, 1 - 670." type="journal article" year="1930">
<collectingCountry id="F35B767FFFA0FF9BFE32F8141D9742C8" box="[419,487,1938,1962]" name="Jordan" pageId="10" pageNumber="91">Jordan</collectingCountry>
<emphasis id="B938EAFDFFA0FF9BFE7FF8151E6B42C8" box="[494,539,1938,1962]" italics="true" pageId="10" pageNumber="91">et al</emphasis>
., 1930: 242
</bibRefCitation>
</treatmentCitation>
(listed; West Coast of 
<collectingCountry id="F35B767FFFA0FF9BFCEEF8151FA242C8" box="[895,978,1939,1962]" name="Panama" pageId="10" pageNumber="91">Panama</collectingCountry>
, synonymic list); 
<bibRefCitation id="EFDD4B1EFFA0FF9BFB18F814196342C8" author="Fowler, H. W." box="[1161,1299,1938,1962]" pageId="10" pageNumber="91" pagination="57 - 529" refId="ref26255" refString="Fowler, H. W. (1944) The fishes. In: Results of the fifth George Vanderbilt Expedition 1941. The Academy of Natural Sciences of Philadelphia, Monographs 6, pp. 57 - 529." type="journal article" year="1944">Fowler, 1944</bibRefCitation>
a: 496 (listed; distribution in Central America); Bussing, 1967: 236 (listed, 
<collectingCountry id="F35B767FFFA0FF9BFCD0F8371FC642AB" box="[833,950,1969,1993]" name="Costa Rica" pageId="10" pageNumber="91">Costa Rica</collectingCountry>
); 
<treatmentCitation id="0AED10FEFFA0FF9BFC58F83718E442AB" author="Whatley, E. C." box="[969,1172,1969,1993]" page="448" pageId="10" pageNumber="91" year="1969">
<bibRefCitation id="EFDD4B1EFFA0FF9BFC58F83718E442AB" author="Whatley, E. C." box="[969,1172,1969,1993]" pageId="10" pageNumber="91" pagination="437 - 474" refId="ref30116" refString="Whatley, E. C. (1969) A study of Syngnathus scovelli in fresh waters of Louisiana and salt waters of Mississippi. Gulf Research Papers, 2, 437 - 474. https: // doi. org / 10.18785 / grr. 0204.02" type="journal article" year="1969">Whatley, 1969: 448</bibRefCitation>
</treatmentCitation>
(cited as euryhaline fish, misspelled); 
<treatmentCitation id="0AED10FEFFA0FF9BFED8F8561E754285" author="Dawson, C. E." box="[329,517,1999,2024]" page="46" pageId="10" pageNumber="91" year="1982">
<bibRefCitation id="EFDD4B1EFFA0FF9BFED8F8561E754285" author="Dawson, C. E." box="[329,517,1999,2024]" pageId="10" pageNumber="91" pagination="1 - 172" refId="ref25847" refString="Dawson, C. E. (1982) Fishes of the Western North Atlantic. The pipefishes (subfamilies Doryrhamphinae and Syngnathinae). Memoirs of the Sears Foundation for Marine Research, 1 (8), 1 - 172." type="journal article" year="1982">Dawson, 1982: 46</bibRefCitation>
</treatmentCitation>
, 53 (synonymic list of 
<taxonomicName id="4C4C4D6CFFA0FF9BFD7CF8561FE74285" ID-CoL="4P6RF" baseAuthorityName="Meek &amp; Hildebrand" baseAuthorityYear="1914" box="[749,919,1999,2023]" class="Actinopterygii" family="Syngnathidae" genus="Pseudophallus" kingdom="Animalia" order="Syngnathiformes" pageId="10" pageNumber="91" phylum="Chordata" rank="species" species="elcapitanensis">
<emphasis id="B938EAFDFFA0FF9BFD7CF8561FE74285" box="[749,919,1999,2023]" italics="true" pageId="10" pageNumber="91">P. elcapitanensis</emphasis>
</taxonomicName>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BF336EFFFA1FF9AFF06FF111F0945AF" blockId="11.[151,1437,150,543]" pageId="11" pageNumber="92">
<treatmentCitationGroup id="AB5C11C1FFA1FF9AFF06FF111F0945AF" pageId="11" pageNumber="92">
<treatmentCitation id="0AED10FEFFA1FF9AFF06FF111E1845CD" author="Herald, E. S." box="[151,616,151,175]" page="61" pageId="11" pageNumber="92" year="1940">
<taxonomicName id="4C4C4D6CFFA1FF9AFF06FF111E1845CD" authority=": Herald, 1940: 61" authorityName="Herald" authorityPageNumber="61" authorityYear="1940" box="[151,616,151,175]" class="Actinopterygii" family="Syngnathidae" genus="Syngnathus" kingdom="Animalia" order="Syngnathiformes" pageId="11" pageNumber="92" phylum="Chordata" rank="species" species="elcapitanensis">
<emphasis id="B938EAFDFFA1FF9AFF06FF111DD645CD" box="[151,422,151,175]" italics="true" pageId="11" pageNumber="92">Syngnathus elcapitanensis</emphasis>
: 
<bibRefCitation id="EFDD4B1EFFA1FF9AFE27FF111E1845CD" author="Herald, E. S." box="[438,616,151,175]" pageId="11" pageNumber="92" pagination="51 - 64" refId="ref26868" refString="Herald, E. S. (1940) A key to the pipefishes of the Pacific American coasts with descriptions of new genera and species. Allan Hancock Pacific Expedition, 9, 51 - 64." type="journal article" year="1940">Herald, 1940: 61</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, 63 (identification key; 
<collectingCountry id="F35B767FFFA1FF9AFCF4FF111FCA45CD" box="[869,954,151,175]" name="Panama" pageId="11" pageNumber="92">Panama</collectingCountry>
); 
<treatmentCitation id="0AED10FEFFA1FF9AFC5FFF1118FB45CD" author="Gunter, G." box="[974,1163,150,175]" page="349" pageId="11" pageNumber="92" year="1956">
<bibRefCitation id="EFDD4B1EFFA1FF9AFC5FFF1118FB45CD" author="Gunter, G." box="[974,1163,150,175]" pageId="11" pageNumber="92" pagination="345 - 354" refId="ref26737" refString="Gunter, G. (1956) A revised list of euryhaline fishes of North and Middle. America Midland Naturalist, 56, 345 - 354. https: // doi. org / 10.2307 / 2422425" type="journal article" year="1956">Gunter, 1956: 349</bibRefCitation>
</treatmentCitation>
(listed; Central America); 
<treatmentCitation id="0AED10FEFFA1FF9AFF56FF331D0845AF" author="Miller, R. R." box="[199,376,181,205]" page="796" pageId="11" pageNumber="92" year="1966">
<bibRefCitation id="EFDD4B1EFFA1FF9AFF56FF331D0845AF" author="Miller, R. R." box="[199,376,181,205]" pageId="11" pageNumber="92" pagination="773 - 802" refId="ref28598" refString="Miller, R. R. (1966) Geographic distribution of central american freshwater fishes. Copeia, 1966, 773 - 802. https: // doi. org / 10.2307 / 1441406" type="journal article" year="1966">Miller, 1966: 796</bibRefCitation>
</treatmentCitation>
(listed; Pacific slopes of 
<collectingCountry id="F35B767FFFA1FF9AFDE8FF331E9745AF" box="[633,743,181,205]" name="Costa Rica" pageId="11" pageNumber="92">Costa Rica</collectingCountry>
and 
<collectingCountry id="F35B767FFFA1FF9AFC88FF331F1E45AF" box="[793,878,181,205]" name="Panama" pageId="11" pageNumber="92">Panama</collectingCountry>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BF336EFFFA1FF9AFF06FF521803458E" blockId="11.[151,1437,150,543]" box="[151,1139,212,236]" pageId="11" pageNumber="92">
<treatmentCitationGroup id="AB5C11C1FFA1FF9AFF06FF521803458E" box="[151,1139,212,236]" pageId="11" pageNumber="92">
<treatmentCitation id="0AED10FEFFA1FF9AFF06FF521E85458E" author="Dawson, C. E." box="[151,757,212,236]" page="53" pageId="11" pageNumber="92" year="1982">
<taxonomicName id="4C4C4D6CFFA1FF9AFF06FF521E85458E" authority=": Dawson, 1982: 53" authorityName=": Dawson" authorityPageNumber="53" authorityYear="1982" box="[151,757,212,236]" class="Actinopterygii" family="Syngnathidae" genus="Syngnathus" kingdom="Animalia" order="Syngnathiformes" pageId="11" pageNumber="92" phylum="Chordata" rank="species" species="elcapitanensis" subGenus="Syngnathus">
<emphasis id="B938EAFDFFA1FF9AFF06FF521E5F458E" box="[151,559,212,236]" italics="true" pageId="11" pageNumber="92">Syngnathus (Syngnathus) elcapitanensis</emphasis>
: 
<bibRefCitation id="EFDD4B1EFFA1FF9AFDACFF521E85458E" author="Dawson, C. E." box="[573,757,212,236]" pageId="11" pageNumber="92" pagination="1 - 172" refId="ref25847" refString="Dawson, C. E. (1982) Fishes of the Western North Atlantic. The pipefishes (subfamilies Doryrhamphinae and Syngnathinae). Memoirs of the Sears Foundation for Marine Research, 1 (8), 1 - 172." type="journal article" year="1982">Dawson, 1982: 53</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(synonymic list of 
<taxonomicName id="4C4C4D6CFFA1FF9AFC2AFF531815458E" authorityName=": Dawson" authorityYear="1982" box="[955,1125,212,236]" class="Actinopterygii" family="Syngnathidae" genus="Syngnathus" kingdom="Animalia" order="Syngnathiformes" pageId="11" pageNumber="92" phylum="Chordata" rank="species" species="elcapitanensis">
<emphasis id="B938EAFDFFA1FF9AFC2AFF531815458E" box="[955,1125,212,236]" italics="true" pageId="11" pageNumber="92">P. elcapitanensis</emphasis>
</taxonomicName>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BF336EFFFA1FF9AFF06FF751E14477D" blockId="11.[151,1437,150,543]" pageId="11" pageNumber="92">
<treatmentCitationGroup id="AB5C11C1FFA1FF9AFF06FF751E14477D" pageId="11" pageNumber="92">
<treatmentCitation id="0AED10FEFFA1FF9AFF06FF751EFA4469" author="Herald, E. S." box="[151,650,242,267]" page="473" pageId="11" pageNumber="92" year="1959">
<taxonomicName id="4C4C4D6CFFA1FF9AFF06FF751EFA4469" ID-CoL="4P6RF" authority=": Herald, 1959: 473" authorityName=": Herald" authorityPageNumber="473" authorityYear="1959" box="[151,650,242,267]" class="Actinopterygii" family="Syngnathidae" genus="Pseudophallus" kingdom="Animalia" order="Syngnathiformes" pageId="11" pageNumber="92" phylum="Chordata" rank="species" species="elcapitanensis">
<emphasis id="B938EAFDFFA1FF9AFF06FF751DB44469" box="[151,452,243,267]" italics="true" pageId="11" pageNumber="92">Pseudophallus elcapitanensis</emphasis>
: 
<bibRefCitation id="EFDD4B1EFFA1FF9AFE43FF751EFA4469" author="Herald, E. S." box="[466,650,242,267]" pageId="11" pageNumber="92" pagination="465 - 473" refId="ref26947" refString="Herald, E. S. (1959) From pipefish to seahorse - a study of phylogenetic relationships. Proceedings of the California Academy of Sciences, 29, 465 - 473." type="journal article" year="1959">Herald, 1959: 473</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(new combination; brief mention to brood-pouch morphology; occurrence in the Pacific coast); 
<treatmentCitation id="0AED10FEFFA1FF9AFE15FE971E49444B" author="Follett, W. I." box="[388,569,273,297]" page="224" pageId="11" pageNumber="92" year="1960">
<bibRefCitation id="EFDD4B1EFFA1FF9AFE15FE971E49444B" author="Follett, W. I." box="[388,569,273,297]" pageId="11" pageNumber="92" pagination="212 - 232" refId="ref26218" refString="Follett, W. I. (1960) The fresh-water fishes - their origins and affinities. Systematic Zoology, 9, 212 - 232. https: // doi. org / 10.2307 / 2411968" type="journal article" year="1960">Follett, 1960: 224</bibRefCitation>
</treatmentCitation>
(affinities with congeners); 
<treatmentCitation id="0AED10FEFFA1FF9AFCC9FE941850444B" author="Dawson, C. E." box="[856,1056,273,297]" page="409" pageId="11" pageNumber="92" year="1974">
<bibRefCitation id="EFDD4B1EFFA1FF9AFCC9FE941850444B" author="Dawson, C. E." box="[856,1056,273,297]" pageId="11" pageNumber="92" pagination="405 - 410" refId="ref25774" refString="Dawson, C. E. (1974) Pseudophallus brasiliensis (Pisces: Syngnathidae), a new freshwater pipefish from Brazil. Proceedings of the Biological Society of Washington, 87, 405 - 410." type="journal article" year="1974">Dawson, 1974: 409</bibRefCitation>
</treatmentCitation>
, 410 (meristic characters in table and identification key); 
<bibRefCitation id="EFDD4B1EFFA1FF9AFE1EFEB61E4E442A" author="Luling, K. H." box="[399,574,304,328]" pageId="11" pageNumber="92" pagination="193 - 205" refId="ref27912" refString="Luling, K. H. (1974) Wissenchaftliche Ergebnisse des Argentinien-Sudbrasilien-Expedition Dr. K. H. Luling 1970 / 71. Der Rio Mage-Roncador und seine charakteristischen Fische von der Einmundung in der Bucht von Rio de Janeiro nebst einer kurzen Notiz uber einen Bach im Orgelgebirge (Sudost-brasilien). Zoologischer Anzeiger, 193, 193 - 205." type="journal article" year="1974">Lüling, 1974:201</bibRefCitation>
(reference); 
<treatmentCitation id="0AED10FEFFA1FF9AFD50FEB61F1E442A" author="Dawson, C. E." box="[705,878,304,328]" page="5" pageId="11" pageNumber="92" year="1982">
<bibRefCitation id="EFDD4B1EFFA1FF9AFD50FEB61F1E442A" author="Dawson, C. E." box="[705,878,304,328]" pageId="11" pageNumber="92" pagination="1 - 172" refId="ref25847" refString="Dawson, C. E. (1982) Fishes of the Western North Atlantic. The pipefishes (subfamilies Doryrhamphinae and Syngnathinae). Memoirs of the Sears Foundation for Marine Research, 1 (8), 1 - 172." type="journal article" year="1982">Dawson, 1982: 5</bibRefCitation>
</treatmentCitation>
, 23, 45, 47–55, figs. 19, 20 (redescription, distribution, comparisons with congeners, identification key); 
<treatmentCitation id="0AED10FEFFA1FF9AFD54FEC91FE04405" author="Dawson, C. E." box="[709,912,334,359]" page="161" pageId="11" pageNumber="92" year="1985">
<bibRefCitation id="EFDD4B1EFFA1FF9AFD54FEC91FE04405" author="Dawson, C. E." box="[709,912,334,359]" pageId="11" pageNumber="92" refId="ref25922" refString="Dawson, C. E. (1985) Indo-Pacific Pipefishes (Red Sea to the Americas). Gulf Coast Research Laboratory, Ocean Springs, Mississippi, 230 pp." type="book" year="1985">Dawson, 1985: 161</bibRefCitation>
</treatmentCitation>
, 209, figs. 239, 241 (characters, distribution, comparisons with congeners, identification key); 
<treatmentCitation id="0AED10FEFFA1FF9AFD08FEE81FCE44E7" author="Lyons, J. &amp; Schneider, D. W." box="[665,958,365,390]" page="4" pageId="11" pageNumber="92" year="1990">
<bibRefCitation id="EFDD4B1EFFA1FF9AFD08FEE81FCE44E7" author="Lyons, J. &amp; Schneider, D. W." box="[665,958,365,390]" pageId="11" pageNumber="92" pagination="1 - 14" refId="ref27975" refString="Lyons, J. &amp; Schneider, D. W. (1990) Factors influencing fish distribution and community structure in a small coastal river in southwestern Costa Rica. Hydrobiologia, 203, 1 - 14. https: // doi. org / 10.1007 / BF 00005608" type="journal article" year="1990">Lyons &amp; Schneider, 1990: 4</bibRefCitation>
</treatmentCitation>
, 6, 7, 8 (listed; 
<collectingCountry id="F35B767FFFA1FF9AFBF5FEEB18A744E7" box="[1124,1239,365,389]" name="Costa Rica" pageId="11" pageNumber="92">Costa Rica</collectingCountry>
, biology, ecology); 
<treatmentCitation id="0AED10FEFFA1FF9AFF56FE0A1DCF44C6" author="Bussing, W. A." box="[199,447,396,420]" page="252" pageId="11" pageNumber="92" year="1998">
<bibRefCitation id="EFDD4B1EFFA1FF9AFF56FE0A1DCF44C6" author="Bussing, W. A." box="[199,447,396,420]" pageId="11" pageNumber="92" pagination="1 - 468" refId="ref25408" refString="Bussing, W. A. (1998) Peces de las aguas continentales de costa rica. Freshwater fishes of Costa Rica. Revista de Biologia Tropical, 46, 1 - 468." type="journal article" year="1998">Bussing, 1998: 252–254</bibRefCitation>
</treatmentCitation>
, figures and map (identification key, species account, distribution in 
<collectingCountry id="F35B767FFFA1FF9AFBFFFE0A189044C6" box="[1134,1248,396,420]" name="Costa Rica" pageId="11" pageNumber="92">Costa Rica</collectingCountry>
); 
<treatmentCitation id="0AED10FEFFA1FF9AFB61FE0A1C9D44A0" author="Ferraris Jr., C." page="591" pageId="11" pageNumber="92" year="2003">
<bibRefCitation id="EFDD4B1EFFA1FF9AFB61FE0A1C9D44A0" author="Ferraris Jr., C." pageId="11" pageNumber="92" pagination="591 - 593" refId="ref26106" refString="Ferraris Jr., C. (2003) Syngnathidae. In: Reis, R. E., Kullander, S. O. &amp; Ferraris Jr., C. J. (Eds.), Check list of the freshwater fishes of South and Central America (CLOFFSCA). Edipucrs, Porto Alegre, pp. 591 - 593." type="book chapter" year="2003">Ferraris Jr, 2003: 591</bibRefCitation>
</treatmentCitation>
(listed, distribution; USNM 81735 indicated as 
<typeStatus id="54F7884DFFA1FF9AFD71FE2D1F4D44A1" box="[736,829,427,451]" pageId="11" pageNumber="92" type="lectotype">lectotype</typeStatus>
of species); Vega 
<emphasis id="B938EAFDFFA1FF9AFC6DFE2D184644A1" box="[1020,1078,427,451]" italics="true" pageId="11" pageNumber="92">et al.,</emphasis>
2006: 92 (listed; 
<collectingCountry id="F35B767FFFA1FF9AFB7EFE2D193444A1" box="[1263,1348,427,451]" name="Panama" pageId="11" pageNumber="92">Panama</collectingCountry>
); 
<treatmentCitation id="0AED10FEFFA1FF9AFAC7FE2D1D5E4483" author="Kuiter, R. H." page="241" pageId="11" pageNumber="92" year="2009">
<bibRefCitation id="EFDD4B1EFFA1FF9AFAC7FE2D1D5E4483" author="Kuiter, R. H." pageId="11" pageNumber="92" refId="ref27592" refString="Kuiter, R. H. (2009) Seahorses and their relatives. Aquatic Photographics, Seaford, 334 pp." type="book" year="2009">Kuiter, 2009: 241</bibRefCitation>
</treatmentCitation>
(distribution, figure from 
<bibRefCitation id="EFDD4B1EFFA1FF9AFDAAFE4C1EA24483" author="Dawson, C. E." box="[571,722,457,481]" pageId="11" pageNumber="92" pagination="1 - 172" refId="ref25847" refString="Dawson, C. E. (1982) Fishes of the Western North Atlantic. The pipefishes (subfamilies Doryrhamphinae and Syngnathinae). Memoirs of the Sears Foundation for Marine Research, 1 (8), 1 - 172." type="journal article" year="1982">Dawson, 1982</bibRefCitation>
); 
<treatmentCitation id="0AED10FEFFA1FF9AFD72FE4F1FAD4483" author="Angulo, A. &amp; Garita-Alvarado, C. A. &amp; Bussing, W. A. &amp; Lopez, M. I." box="[739,989,457,481]" page="998" pageId="11" pageNumber="92" year="2013">
<bibRefCitation id="EFDD4B1EFFA1FF9AFD72FE4F1FAD4483" author="Angulo, A. &amp; Garita-Alvarado, C. A. &amp; Bussing, W. A. &amp; Lopez, M. I." box="[739,989,457,481]" pageId="11" pageNumber="92" pagination="987 - 1019" refId="ref24843" refString="Angulo, A., Garita-Alvarado, C. A., Bussing, W. A. &amp; Lopez, M. I. (2013) Annotated checklist of the freshwater fishes of continental and insular Costa Rica: additions and nomenclatural revisions. Check List, 9, 987 - 1019. https: // doi. org / 10.15560 / 9.5.987" type="journal article" year="2013">
Angulo 
<emphasis id="B938EAFDFFA1FF9AFCA7FE4C1F1F4483" box="[822,879,457,481]" italics="true" pageId="11" pageNumber="92">et al.,</emphasis>
2013: 998
</bibRefCitation>
</treatmentCitation>
(listed, distribution; 
<collectingCountry id="F35B767FFFA1FF9AFB22FE4F19574483" box="[1203,1319,457,481]" name="Costa Rica" pageId="11" pageNumber="92">Costa Rica</collectingCountry>
); 
<treatmentCitation id="0AED10FEFFA1FF9AFAA6FE4F1D4C449D" author="Angulo, A. &amp; Molina-Arias, A. &amp; Murase, A. &amp; Miyazaki, Y. &amp; Bussing, W. &amp; Lopez, M." page="7" pageId="11" pageNumber="92" year="2015">
<bibRefCitation id="EFDD4B1EFFA1FF9AFAA6FE4F1D4C449D" author="Angulo, A. &amp; Molina-Arias, A. &amp; Murase, A. &amp; Miyazaki, Y. &amp; Bussing, W. &amp; Lopez, M." pageId="11" pageNumber="92" pagination="1 - 15" refId="ref24908" refString="Angulo, A., Molina-Arias, A., Murase, A., Miyazaki, Y., Bussing, W. &amp; Lopez, M. (2015) Fishes from the Tusubres River basin, Pacific Coast, Costa Rica: checklist, identification key and photographic album. Check List, 11, 1 - 15. https: // doi. org / 10.15560 / 11.3.1666" type="journal article" year="2015">
Angulo 
<emphasis id="B938EAFDFFA1FF9AFA1BFE4C1C974762" italics="true" pageId="11" pageNumber="92">et al.,</emphasis>
2015: 7
</bibRefCitation>
</treatmentCitation>
, 9, 10; fig. 5J (listed, distribution in 
<collectingCountry id="F35B767FFFA1FF9AFD3EFE6E1F514762" box="[687,801,488,512]" name="Costa Rica" pageId="11" pageNumber="92">Costa Rica</collectingCountry>
, identification key); 
<treatmentCitation id="0AED10FEFFA1FF9AFC60FE6E18A24762" author="Angulo, A. &amp; Gil-Leon, J. S. &amp; Oviedo-Soto, A. &amp; Abarca-Odio, J. I. &amp; Umana-Villalobos, G." box="[1009,1234,488,512]" page="9" pageId="11" pageNumber="92" year="2020">
<bibRefCitation id="EFDD4B1EFFA1FF9AFC60FE6E18A24762" author="Angulo, A. &amp; Gil-Leon, J. S. &amp; Oviedo-Soto, A. &amp; Abarca-Odio, J. I. &amp; Umana-Villalobos, G." box="[1009,1234,488,512]" pageId="11" pageNumber="92" pagination="1 - 28" refId="ref24980" refString="Angulo, A., Gil-Leon, J. S., Oviedo-Soto, A., Abarca-Odio, J. I. &amp; Umana-Villalobos, G. (2020) The fishes from the Coto River basin, Pacific Coast, Costa Rica. Zootaxa, 4751 (1), 1 - 28. https: // doi. org / 10.11646 / zootaxa. 4751.1.1" type="journal article" year="2020">
Angulo 
<emphasis id="B938EAFDFFA1FF9AFBD5FE6F18014762" box="[1092,1137,488,512]" italics="true" pageId="11" pageNumber="92">et al</emphasis>
., 2020: 9
</bibRefCitation>
</treatmentCitation>
, 18, 19, 25; fig. 8M (listed in 
<collectingCountry id="F35B767FFFA1FF9AFEB6FD811DE8477D" box="[295,408,519,543]" name="Costa Rica" pageId="11" pageNumber="92">Costa Rica</collectingCountry>
, identification key).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3566564FFA1FF9AFF06FDC118A6463C" pageId="11" pageNumber="92" type="diagnosis">
<paragraph id="8BF336EFFFA1FF9AFF06FDC11D3547C8" blockId="11.[151,1437,583,2041]" pageId="11" pageNumber="92">
<emphasis id="B938EAFDFFA1FF9AFF06FDC11D624703" bold="true" box="[151,274,583,609]" pageId="11" pageNumber="92">Diagnosis.</emphasis>
<taxonomicName id="4C4C4D6CFFA1FF9AFE86FDC11E124703" authorityName=": Herald" authorityYear="1959" box="[279,610,583,609]" class="Actinopterygii" family="Syngnathidae" genus="Pseudophallus" kingdom="Animalia" order="Syngnathiformes" pageId="11" pageNumber="92" phylum="Chordata" rank="species" species="elcapitanensis">
<emphasis id="B938EAFDFFA1FF9AFE86FDC11E124703" box="[279,610,583,609]" italics="true" pageId="11" pageNumber="92">Pseudophallus elcapitanensis</emphasis>
</taxonomicName>
differs from species of 
<taxonomicName id="4C4C4D6CFFA1FF9AFCF9FDC1187D4703" authorityName="Herald" authorityYear="1940" box="[872,1037,583,609]" class="Actinopterygii" family="Syngnathidae" genus="Pseudophallus" kingdom="Animalia" order="Syngnathiformes" pageId="11" pageNumber="92" phylum="Chordata" rank="genus">
<emphasis id="B938EAFDFFA1FF9AFCF9FDC1187D4703" box="[872,1037,583,609]" italics="true" pageId="11" pageNumber="92">Pseudophallus</emphasis>
</taxonomicName>
from the Atlantic coastal drainages, 
<taxonomicName id="4C4C4D6CFFA1FF9AFF06FDEA1D4347E7" box="[151,307,619,645]" class="Actinopterygii" family="Syngnathidae" genus="Pseudophallus" kingdom="Animalia" order="Syngnathiformes" pageId="11" pageNumber="92" phylum="Chordata" rank="species" species="brasiliensis">
<emphasis id="B938EAFDFFA1FF9AFF06FDEA1D4347E7" box="[151,307,619,645]" italics="true" pageId="11" pageNumber="92">P. brasiliensis</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C4C4D6CFFA1FF9AFED0FDEA1DD447E7" baseAuthorityName="Meek &amp; Hildebrand" baseAuthorityYear="1923" box="[321,420,619,645]" class="Actinopterygii" family="Syngnathidae" genus="Pseudophallus" kingdom="Animalia" order="Syngnathiformes" pageId="11" pageNumber="92" phylum="Chordata" rank="species" species="mindii">
<emphasis id="B938EAFDFFA1FF9AFED0FDEA1DD447E7" box="[321,420,619,645]" italics="true" pageId="11" pageNumber="92">P. mindii</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C4C4D6CFFA1FF9AFE49FDEA1E0447E7" authorityName="Dallevo-Gomes &amp; Mattox &amp; Toledo-Piza" authorityYear="2020" box="[472,628,619,645]" class="Actinopterygii" family="Syngnathidae" genus="Pseudophallus" kingdom="Animalia" order="Syngnathiformes" pageId="11" pageNumber="92" phylum="Chordata" rank="species" species="galadrielae">
<emphasis id="B938EAFDFFA1FF9AFE49FDEA1E0447E7" box="[472,628,619,645]" italics="true" pageId="11" pageNumber="92">P. galadrielae</emphasis>
</taxonomicName>
in having all of the specimens with 14 trunk rings (
<emphasis id="B938EAFDFFA1FF9AFB20FDEA18B847E7" box="[1201,1224,620,645]" italics="true" pageId="11" pageNumber="92">vs</emphasis>
. most usually with 13 trunk rings).
</paragraph>
<paragraph id="8BF336EFFFA1FF9AFF56FD3518A6463C" blockId="11.[151,1437,583,2041]" pageId="11" pageNumber="92">
<taxonomicName id="4C4C4D6CFFA1FF9AFF56FD351E6347AF" authorityName=": Herald" authorityYear="1959" box="[199,531,691,717]" class="Actinopterygii" family="Syngnathidae" genus="Pseudophallus" kingdom="Animalia" order="Syngnathiformes" pageId="11" pageNumber="92" phylum="Chordata" rank="species" species="elcapitanensis">
<emphasis id="B938EAFDFFA1FF9AFF56FD351E6347AF" box="[199,531,691,717]" italics="true" pageId="11" pageNumber="92">Pseudophallus elcapitanensis</emphasis>
</taxonomicName>
differs from 
<taxonomicName id="4C4C4D6CFFA1FF9AFD34FD321F6247AF" box="[677,786,691,717]" class="Actinopterygii" family="Syngnathidae" genus="Pseudophallus" kingdom="Animalia" order="Syngnathiformes" pageId="11" pageNumber="92" phylum="Chordata" rank="species" species="starksii">
<emphasis id="B938EAFDFFA1FF9AFD34FD321F6247AF" box="[677,786,691,717]" italics="true" pageId="11" pageNumber="92">P. starksii</emphasis>
</taxonomicName>
in the following combination of characters: 30–35 dorsalfin rays (87.2% of specimens with 31–33), 47-51 body rings (89.7% of specimens with 48–49), 17–19 brood-pouch rings (90.9% of specimens with 17–18) and 6.75–7.5 subdorsal tail rings 
<emphasis id="B938EAFDFFA1FF9AFC58FD7A1F904677" box="[969,992,764,789]" italics="true" pageId="11" pageNumber="92">vs</emphasis>
. 36–44 dorsal-fin rays (89.5% of specimens with 38–41), 49–53 body rings (91.9% of specimens with 50–51), 19–21 brood-pouch rings (64.9% of adult male specimens with 21) and 8.5–10.5 subdorsal tail rings (86.7% of specimens with 9.25–10.0).
</paragraph>
</subSubSection>
<subSubSection id="C3566564FFA1FF9DFF56FCE11F7345B7" lastPageId="12" lastPageNumber="93" pageId="11" pageNumber="92" type="description">
<paragraph id="8BF336EFFFA1FF9AFF56FCE119E2411F" blockId="11.[151,1437,583,2041]" pageId="11" pageNumber="92">
<emphasis id="B938EAFDFFA1FF9AFF56FCE11D2946E3" bold="true" box="[199,345,871,897]" pageId="11" pageNumber="92">Description.</emphasis>
Meristic and morphometric characters detailed in 
<tableCitation id="C6CE0354FFA1FF9AFC1FFCE11F9446E0" box="[910,996,871,898]" captionStart="TABLE 2" captionStartId="4.[151,239,151,175]" captionTargetPageId="4" captionText="TABLE 2. Morphometric and meristic data of Pseudophallus species of the Pacific drainages. Standard length (SL) in millimeters, 2–10 are percentages of SL, 11–17 are percentages of head length. N= number of specimens analyzed." pageId="11" pageNumber="92">Table 2</tableCitation>
. General appearance in 
<figureCitation id="13772A6AFFA1FF9AFB60FCE1191C46E0" box="[1265,1388,871,898]" captionStart="FIGURE 2" captionStartId="6.[151,250,1798,1822]" captionTargetBox="[164,1430,358,1762]" captionTargetId="figure-100@6.[151,1436,345,1775]" captionTargetPageId="6" captionText="FIGURE 2: Species of the genus Pseudophallus: A—Pseudophallus brasiliensis (paratype) USNM 212058, 69.1 mm SL, Igarapé Inó, Furo de Panaquera, Rio Tocantins, Pará, Brazil. B—Pseudophallus elcapitanensis MCZ 88623, 110.6 mm SL, Río Picuro, Darien, Panama. C—Pseudophallus mindii (holotype) USNM 81770, 90.3 mm SL, Mindi Creek, Canal Zone, Panama. D—Pseudophallus starksii USNM 208365, 171.1 mm SL, upper tributary of Río Jaque, Província de Darien, Panama. E—Pseudophallus galadrielae. (holotype) FMNH 126156, 63.1mm SL, Lago Izabal, Guatemala. Red arrow indicates the anal papilla." figureDoi="http://doi.org/10.5281/zenodo.4412815" httpUri="https://zenodo.org/record/4412815/files/figure.png" pageId="11" pageNumber="92">Figures 2B</figureCitation>
and 
<figureCitation id="13772A6AFFA1FF9AFF06FC0A1CCB46C4" box="[151,187,908,934]" captionStart="FIGURE 3" captionStartId="7.[151,250,1816,1840]" captionTargetBox="[375,1220,194,1786]" captionTargetId="figure-17@7.[361,1226,181,1798]" captionTargetPageId="7" captionText="FIGURE 3: Detail of anterior portion of body of species of the genus Pseudophallus A—Pseudophallus brasiliensis (paratype) USNM 212058, 69.1 mm SL, Igarapé Inó Furo de Panaquera, Rio Tocantins, Pará, Brazil. B—Pseudophallus elcapitanensis MCZ 88623, 110.6 mm SL, Río Picuro, Darien, Panama. C—Pseudophallus mindii (holotype) USNM 81770, 90.3 mm SL, Mindi Creek, Canal Zone, Panama. D—Pseudophallus starksii USNM 208365, 171.1 mm SL, upper tributary of Río Jaque, Província de Darien, Panama. E—Pseudophallus galadrielae. (holotype), FMNH 126156, 63.1 mm SL, Lago Izabal, Guatemala." figureDoi="http://doi.org/10.5281/zenodo.4412817" httpUri="https://zenodo.org/record/4412817/files/figure.png" pageId="11" pageNumber="92">3B</figureCitation>
. Body without scales, covered with bony plates forming rings along its axis, bony plates horizontally oval, with an elevation in midline forming bony ridges along body. Trunk rings 14. Tail rings 33–37 (89.7% of specimens with 34–35). Body rings 47–51 (89.7% of specimens with 48–49). Subdorsal rings 6.75–7.5 (97.4% of specimens with 6.75–7.25). Subdorsal trunk 0–0.25 rings. Subdorsal tail rings 6.75–7.5 (97.4% of specimens with 6.75–7.25). Dorsal-fin origin between 0.25 of posteriormost trunk ring and anteriormost caudal ring. Posterior tip of superior trunk ridge at vertical through superior tail ridge, between 6.0 and 7.5 anteriormost tail rings. Lateral trunk ridge continuous with superior tail ridge with origin on first caudal ring. Inferior trunk ridge continuous with inferior tail ridge.
</paragraph>
<paragraph id="8BF336EFFFA1FF9AFF56FB011D5F4036" blockId="11.[151,1437,583,2041]" pageId="11" pageNumber="92">Body slender and cylindrical, transversal section of trunk heptagonal, with transition to quadrangular in tail at end of superior trunk ridge. Maximum depth of body of females and juveniles at middle of trunk; in males at region of the brood pouch. Body depth decreasing posteriorly along superior trunk ridge to minimum depth on caudal peduncle. Dorsal profile of head straight or slightly inclined on snout region, more inclined on orbital region and straight on posterior region, with two nuchal plates between occipital region and first trunk ring. Ventral profile of head straight.</paragraph>
<paragraph id="8BF336EFFFA1FF9AFF56FAD81F07434B" blockId="11.[151,1437,583,2041]" pageId="11" pageNumber="92">Dorsal fin not elevated, curved on first rays, straight along median portion with pointed end. Dorsal-fin rays of similar length, except the first two rays. Dorsal-fin rays 30–35 (82.1% of specimens with 31–33), posteriormost two rays closer to each other, difficult to distinguish in some specimens. Pectoral fin small, middle rays longer than outer and inner rays. Pectoral-fin origin on anteriormost trunk ring, distal margin of pectoral-fin rays rarely extending beyond third trunk ring. Pectoral-fin rays 13–16 (66.7% of specimens with 14). Caudal fin small and fan like, with 10 rays (5 rays in each hypural plate). Pelvic and anal fins absent.</paragraph>
<paragraph id="8BF336EFFFA1FF9AFF56F9B51E3B439C" blockId="11.[151,1437,583,2041]" pageId="11" pageNumber="92">Mouth terminal, slightly inclined dorsally, jaws edentulous. Lower jaw slightly prognathous. Snout short, tubular, without dorsal or lateral ridges. Dorsal profile of snout with slight inclination following dorsal margin of eyes. Eyes located longitudinally at middle of head. Opercle occupying most of lateral surface of head posterior to eye, opercular opening reduced to pore on posterodorsal margin of opercle. Opercular ridge generally indistinct, except for some specimens with small protuberance on anterior region of opercle, following superior margin of bone. Lateral line and lateral line canals absent.</paragraph>
<paragraph id="8BF336EFFFA1FF9AFF56F8811D3B42B7" blockId="11.[151,1437,583,2041]" pageId="11" pageNumber="92">
<emphasis id="B938EAFDFFA1FF9AFF56F8811DCA4243" bold="true" box="[199,442,1799,1825]" pageId="11" pageNumber="92">Sexual dimorphism.</emphasis>
Males with brood pouch. Origin of brood pouch on posteriormost trunk rings, brood pouch rings extending along 17–19 rings (90.9% of specimens with 17–18 brood-pouch rings). Bony plates of the brood pouch well developed and T-shaped encasing entire lateral portion of pouch, decreasing in size posteriorly and parallel with the inferior tail ridge. Ventral region of pouch with contralateral dermal folds extending towards the midline, dividing brood pouch longitudinally. Adult females with anal papilla on anal ring with small, barely visible spicules.
</paragraph>
<paragraph id="8BF336EFFFA1FF9DFF56F8591F7345B7" blockId="11.[151,1437,583,2041]" lastBlockId="12.[151,1436,151,285]" lastPageId="12" lastPageNumber="93" pageId="11" pageNumber="92">
<emphasis id="B938EAFDFFA1FF9AFF56F8591DFE429B" bold="true" box="[199,398,2015,2041]" pageId="11" pageNumber="92">Color in alcohol.</emphasis>
Head and body ranging from a uniform light to dark brown, with small white punctuations. Dorsal-fin rays with scattered brown chromatophores, pectoral fin hyaline. Caudal fin with the same coloration of the body, sometimes varying in tone (
<figureCitation id="13772A6AFFA6FF9DFDADFF3D1EEC45B7" box="[572,668,187,213]" captionStart="FIGURE 2" captionStartId="6.[151,250,1798,1822]" captionTargetBox="[164,1430,358,1762]" captionTargetId="figure-100@6.[151,1436,345,1775]" captionTargetPageId="6" captionText="FIGURE 2: Species of the genus Pseudophallus: A—Pseudophallus brasiliensis (paratype) USNM 212058, 69.1 mm SL, Igarapé Inó, Furo de Panaquera, Rio Tocantins, Pará, Brazil. B—Pseudophallus elcapitanensis MCZ 88623, 110.6 mm SL, Río Picuro, Darien, Panama. C—Pseudophallus mindii (holotype) USNM 81770, 90.3 mm SL, Mindi Creek, Canal Zone, Panama. D—Pseudophallus starksii USNM 208365, 171.1 mm SL, upper tributary of Río Jaque, Província de Darien, Panama. E—Pseudophallus galadrielae. (holotype) FMNH 126156, 63.1mm SL, Lago Izabal, Guatemala. Red arrow indicates the anal papilla." figureDoi="http://doi.org/10.5281/zenodo.4412815" httpUri="https://zenodo.org/record/4412815/files/figure.png" pageId="12" pageNumber="93">Figs. 2B</figureCitation>
and 
<figureCitation id="13772A6AFFA6FF9DFD42FF3D1E8645B7" box="[723,758,187,213]" captionStart="FIGURE 3" captionStartId="7.[151,250,1816,1840]" captionTargetBox="[375,1220,194,1786]" captionTargetId="figure-17@7.[361,1226,181,1798]" captionTargetPageId="7" captionText="FIGURE 3: Detail of anterior portion of body of species of the genus Pseudophallus A—Pseudophallus brasiliensis (paratype) USNM 212058, 69.1 mm SL, Igarapé Inó Furo de Panaquera, Rio Tocantins, Pará, Brazil. B—Pseudophallus elcapitanensis MCZ 88623, 110.6 mm SL, Río Picuro, Darien, Panama. C—Pseudophallus mindii (holotype) USNM 81770, 90.3 mm SL, Mindi Creek, Canal Zone, Panama. D—Pseudophallus starksii USNM 208365, 171.1 mm SL, upper tributary of Río Jaque, Província de Darien, Panama. E—Pseudophallus galadrielae. (holotype), FMNH 126156, 63.1 mm SL, Lago Izabal, Guatemala." figureDoi="http://doi.org/10.5281/zenodo.4412817" httpUri="https://zenodo.org/record/4412817/files/figure.png" pageId="12" pageNumber="93">3B</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3566564FFA6FF9DFF56FF581F41447F" pageId="12" pageNumber="93" type="distribution">
<paragraph id="8BF336EFFFA6FF9DFF56FF581F41447F" blockId="12.[151,1436,151,285]" pageId="12" pageNumber="93">
<emphasis id="B938EAFDFFA6FF9DFF56FF581D11459A" bold="true" box="[199,353,222,248]" pageId="12" pageNumber="93">Distribution.</emphasis>
Freshwater coastal drainages of the Pacific, from Río Mamoni, Province of El Capitain, 
<collectingCountry id="F35B767FFFA6FF9DFAACFF5919E9459B" box="[1341,1433,223,249]" name="Panama" pageId="12" pageNumber="93">Panama</collectingCountry>
, to Río Turrubales, Province of Orotina, 
<collectingCountry id="F35B767FFFA6FF9DFDC7FE851EBF447F" box="[598,719,259,285]" name="Costa Rica" pageId="12" pageNumber="93">Costa Rica</collectingCountry>
(
<figureCitation id="13772A6AFFA6FF9DFD4EFE851F53447F" box="[735,803,258,285]" captionStart="FIGURE 5" captionStartId="12.[151,250,1045,1069]" captionTargetBox="[349,1239,321,1005]" captionTargetId="figure-77@12.[340,1247,309,1022]" captionTargetPageId="12" captionText="FIGURE 5. Distribution map of species of Pseudophallus occurring in drainages along the Pacific coast based on specimens examined in this study." figureDoi="http://doi.org/10.5281/zenodo.4412823" httpUri="https://zenodo.org/record/4412823/files/figure.png" pageId="12" pageNumber="93">Fig. 5</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="DF336667FFA6FF9DFF06FB931DF24133" ID-DOI="http://doi.org/10.5281/zenodo.4412823" ID-Zenodo-Dep="4412823" httpUri="https://zenodo.org/record/4412823/files/figure.png" pageId="12" pageNumber="93" startId="12.[151,250,1045,1069]" targetBox="[349,1239,321,1005]" targetPageId="12">
<paragraph id="8BF336EFFFA6FF9DFF06FB931DF24133" blockId="12.[151,1437,1045,1105]" pageId="12" pageNumber="93">
<emphasis id="B938EAFDFFA6FF9DFF06FB931D64414C" bold="true" box="[151,276,1045,1070]" pageId="12" pageNumber="93">FIGURE 5.</emphasis>
Distribution map of species of 
<taxonomicName id="4C4C4D6CFFA6FF9DFDCBFB931E9F414F" authorityName="Herald" authorityYear="1940" box="[602,751,1045,1069]" class="Actinopterygii" family="Syngnathidae" genus="Pseudophallus" kingdom="Animalia" order="Syngnathiformes" pageId="12" pageNumber="93" phylum="Chordata" rank="genus">
<emphasis id="B938EAFDFFA6FF9DFDCBFB931E9F414F" box="[602,751,1045,1069]" italics="true" pageId="12" pageNumber="93">Pseudophallus</emphasis>
</taxonomicName>
occurring in drainages along the Pacific coast based on specimens examined in this study.
</paragraph>
</caption>
<subSubSection id="C3566564FFA6FF9CFF56FB061EFE475F" lastPageId="13" lastPageNumber="94" pageId="12" pageNumber="93" type="discussion">
<paragraph id="8BF336EFFFA6FF9DFF56FB061FCB4048" blockId="12.[151,1437,1152,2042]" pageId="12" pageNumber="93">
<emphasis id="B938EAFDFFA6FF9DFF56FB061D4B41F8" bold="true" box="[199,315,1152,1178]" pageId="12" pageNumber="93">Remarks.</emphasis>
Meek &amp; Hildebrand (in 
<bibRefCitation id="EFDD4B1EFFA6FF9DFDC1FB061EA641F8" author="Meek, S. E." box="[592,726,1152,1178]" pageId="12" pageNumber="93" pagination="101 - 134" refId="ref28392" refString="Meek, S. E. (1914) An annotated list of fishes known to occur in the fresh waters of Costa Rica. Field Museum of Natural History, Zoological Series, 10, 101 - 134. https: // doi. org / 10.5962 / bhl. title. 5576" type="journal article" year="1914">Meek, 1914</bibRefCitation>
) provided a brief description of 
<taxonomicName id="4C4C4D6CFFA6FF9DFBD1FB06191441F8" authorityName="Meek &amp; Hildebrand in Meek" authorityYear="1914" box="[1088,1380,1152,1178]" class="Polychaeta" family="Flabelligeridae" genus="Siphostoma" kingdom="Animalia" order="Terebellida" pageId="12" pageNumber="93" phylum="Annelida" rank="species" species="elcapitanense">
<emphasis id="B938EAFDFFA6FF9DFBD1FB06191441F8" box="[1088,1380,1152,1178]" italics="true" pageId="12" pageNumber="93">Siphostoma elcapitanense</emphasis>
</taxonomicName>
(sic) stating that the species would be further detailed in an upcoming account of fishes of 
<collectingCountry id="F35B767FFFA6FF9DFBFDFB2218B841DC" box="[1132,1224,1188,1214]" name="Panama" pageId="12" pageNumber="93">Panama</collectingCountry>
, a study that came out years later (
<bibRefCitation id="EFDD4B1EFFA6FF9DFED9FB4E1E084180" author="Meek, S. &amp; Hildebrand, S. F." box="[328,632,1224,1250]" pageId="12" pageNumber="93" pagination="1 - 330" refId="ref28446" refString="Meek, S. &amp; Hildebrand, S. F. (1923) The marine fishes of Panama. Part 1. Field Museum of Natural History, Zoological Series, 15, 1 - 330." type="journal article" year="1923">Meek &amp; Hildebrand, 1923</bibRefCitation>
). In the original description, the 
<typeStatus id="54F7884DFFA6FF9DFC7DFB4E186C4180" box="[1004,1052,1224,1250]" pageId="12" pageNumber="93">type</typeStatus>
locality was stated as El Capitan, 
<collectingCountry id="F35B767FFFA6FF9DFF06FB6A1C834064" box="[151,243,1260,1286]" name="Panama" pageId="12" pageNumber="93">Panama</collectingCountry>
. No catalogue number of lots was mentioned, but two specimens from 
<collectingCountry id="F35B767FFFA6FF9DFB9EFB6A18F94064" box="[1039,1161,1260,1286]" name="Costa Rica" pageId="12" pageNumber="93">Costa Rica</collectingCountry>
were listed at the end of the description: one from Jesus Maria and another from Río Turrubales.
</paragraph>
<paragraph id="8BF336EFFFA6FF9DFF56FAB21D6440BC" blockId="12.[151,1437,1152,2042]" pageId="12" pageNumber="93">
<materialsCitation id="3B243CB2FFA6FF9DFF56FAB21D7F40BC" ID-GBIF-Occurrence="3034656351" collectionCode="USNM" collectorName="The &amp; El Capitan" country="Panama" location="Rio Mamoni" pageId="12" pageNumber="93" specimenCode="USNM 81735" specimenCount="11">
<collectorName id="26B95339FFA6FF9DFF56FAB21C83402C" box="[199,243,1332,1358]" pageId="12" pageNumber="93">The</collectorName>
species was described in more detail under a different genus, as 
<taxonomicName id="4C4C4D6CFFA6FF9DFC75FAB2197E402C" authorityName=": Dawson" authorityYear="1982" box="[996,1294,1332,1358]" class="Actinopterygii" family="Syngnathidae" genus="Syngnathus" kingdom="Animalia" order="Syngnathiformes" pageId="12" pageNumber="93" phylum="Chordata" rank="species" species="elcapitanensis">
<emphasis id="B938EAFDFFA6FF9DFC75FAB2197E402C" box="[996,1294,1332,1358]" italics="true" pageId="12" pageNumber="93">Syngnathus elcapitanensis</emphasis>
</taxonomicName>
by 
<bibRefCitation id="EFDD4B1EFFA6FF9DFAACFAB21DD44010" author="Meek, S. &amp; Hildebrand, S. F." pageId="12" pageNumber="93" pagination="1 - 330" refId="ref28446" refString="Meek, S. &amp; Hildebrand, S. F. (1923) The marine fishes of Panama. Part 1. Field Museum of Natural History, Zoological Series, 15, 1 - 330." type="journal article" year="1923">Meek &amp; Hildebrand (1923: 260)</bibRefCitation>
, who also listed and illustrated a male specimen, from 
<location id="8E936034FFA6FF9DFB86FADE18D94010" LSID="urn:lsid:plazi:treatment:03E587F9FFA0FF9CFF06F9B218A3418B:8E936034FFA6FF9DFB86FADE18D94010" box="[1047,1193,1368,1394]" country="Panama" name="Rio Mamoni" pageId="12" pageNumber="93">Río Mamoni</location>
, 
<collectorName id="26B95339FFA6FF9DFB24FADE19404010" box="[1205,1328,1368,1394]" pageId="12" pageNumber="93">El Capitan</collectorName>
, 
<collectingCountry id="F35B767FFFA6FF9DFAACFADE19E94010" box="[1341,1433,1368,1394]" name="Panama" pageId="12" pageNumber="93">Panama</collectingCountry>
, catalogued as 
<specimenCode id="DBEA9E94FFA6FF9DFEA6FAFA1DA840F4" box="[311,472,1403,1430]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="12" pageNumber="93" type="Museum">USNM 81735</specimenCode>
, as the 
<typeStatus id="54F7884DFFA6FF9DFDBDFAFA1E2C40F4" box="[556,604,1404,1430]" pageId="12" pageNumber="93">type</typeStatus>
specimen. Those authors also mentioned that they had a total of 
<specimenCount id="9D4AFD66FFA6FF9DFAA6FAFA1CA540D8" pageId="12" pageNumber="93" type="generic">11 specimens</specimenCount>
, two males and nine females, from five different localities in 
<collectingCountry id="F35B767FFFA6FF9DFC05FA261F9D40D8" box="[916,1005,1440,1466]" name="Panama" pageId="12" pageNumber="93">Panama</collectingCountry>
and that all, except one, originated in freshwater
</materialsCitation>
.
</paragraph>
<paragraph id="8BF336EFFFA6FF9DFF56FA6E1EB243D4" blockId="12.[151,1437,1152,2042]" pageId="12" pageNumber="93">
Ten of those specimens are currently deposited in the fish collections of the National Museum of Natural History, Smithsonian Institution (
<specimenCount id="9D4AFD66FFA6FF9DFE74F98D1E024344" box="[485,626,1547,1574]" pageId="12" pageNumber="93" type="generic">5 specimens</specimenCount>
, one in each lot of USNM 79687, 79689–90; 81375 and 108645) and the Field Museum of Natural History (
<specimenCount id="9D4AFD66FFA6FF9DFD8CF9A91EDA4328" box="[541,682,1583,1610]" pageId="12" pageNumber="93" type="generic">5 specimens</specimenCount>
, one in each lot of FMNH 8290–8294). The last specimen of the 11 mentioned by 
<bibRefCitation id="EFDD4B1EFFA6FF9DFEA4F9D21E14430C" author="Meek, S. &amp; Hildebrand, S. F." box="[309,612,1620,1646]" pageId="12" pageNumber="93" pagination="1 - 330" refId="ref28446" refString="Meek, S. &amp; Hildebrand, S. F. (1923) The marine fishes of Panama. Part 1. Field Museum of Natural History, Zoological Series, 15, 1 - 330." type="journal article" year="1923">Meek &amp; Hildebrand (1923)</bibRefCitation>
originated in Chame Point, 
<collectingCountry id="F35B767FFFA6FF9DFC08F9D21F85430C" box="[921,1013,1620,1646]" name="Panama" pageId="12" pageNumber="93">Panama</collectingCountry>
, which according to those authors was not a locality from freshwater. According to 
<bibRefCitation id="EFDD4B1EFFA6FF9DFD02F9FE1F1E43F0" author="Dawson, C. E." box="[659,878,1655,1682]" pageId="12" pageNumber="93" pagination="1 - 172" refId="ref25847" refString="Dawson, C. E. (1982) Fishes of the Western North Atlantic. The pipefishes (subfamilies Doryrhamphinae and Syngnathinae). Memoirs of the Sears Foundation for Marine Research, 1 (8), 1 - 172." type="journal article" year="1982">Dawson (1982: 54)</bibRefCitation>
that is a specimen of 
<taxonomicName id="4C4C4D6CFFA6FF9DFBFBF9FE191943F0" box="[1130,1385,1656,1682]" class="Actinopterygii" family="Syngnathidae" genus="Pseudophallus" kingdom="Animalia" order="Syngnathiformes" pageId="12" pageNumber="93" phylum="Chordata" rank="species" species="starksii">
<emphasis id="B938EAFDFFA6FF9DFBFBF9FE191943F0" box="[1130,1385,1656,1682]" italics="true" pageId="12" pageNumber="93">Pseudophallus starksii</emphasis>
</taxonomicName>
that was misidentified by 
<bibRefCitation id="EFDD4B1EFFA6FF9DFE19F91A1ECE43D4" author="Meek, S. &amp; Hildebrand, S. F." box="[392,702,1692,1718]" pageId="12" pageNumber="93" pagination="1 - 330" refId="ref28446" refString="Meek, S. &amp; Hildebrand, S. F. (1923) The marine fishes of Panama. Part 1. Field Museum of Natural History, Zoological Series, 15, 1 - 330." type="journal article" year="1923">Meek &amp; Hildebrand (1923)</bibRefCitation>
.
</paragraph>
<paragraph id="8BF336EFFFA6FF9DFF56F94618414224" blockId="12.[151,1437,1152,2042]" pageId="12" pageNumber="93">Associated with each lot from the Field Museum of Natural History are handwritten labels with numbers that correspond to the catalogue numbers of the lots from the National Museum of Natural History as follows: FMNH 8290 (number in label: 79688), FMNH 8291–8292 (number in label: 79690, also marked as cotypes in a label), FMNH 8293 (number in label: 79689) and FMNH 8294 (number in label: 79687).</paragraph>
<paragraph id="8BF336EFFFA6FF9CFF56F8D61EFE475F" blockId="12.[151,1437,1152,2042]" lastBlockId="13.[151,1437,151,1257]" lastPageId="13" lastPageNumber="94" pageId="12" pageNumber="93">
All the 
<specimenCount id="9D4AFD66FFA6FF9DFE88F8D61DC04208" box="[281,432,1872,1898]" pageId="12" pageNumber="93" type="generic">10 specimens</specimenCount>
listed above were collected by S. E. Meek and S. F. Hildebrand in 
<date id="FFF2102FFFA6FF9DFB1FF8D619644208" box="[1166,1300,1872,1898]" pageId="12" pageNumber="93" value="1911-03">March 1911</date>
or 
<date id="FFF2102FFFA6FF9DFAA6F8D61CA242EC" pageId="12" pageNumber="93" value="1912-02">February 1912</date>
, as part of the expedition Smithsonian Biological Survey of the 
<collectingCountry id="F35B767FFFA6FF9DFC07F8F21F9F42EC" box="[918,1007,1908,1934]" name="Panama" pageId="12" pageNumber="93">Panama</collectingCountry>
Canal Zone. Of those, only five specimens: USNM 79690, USNM 81735, USNM 108645, FMNH 8291 and FMNH 8292 were collected at Río Mamoni, El Capitan, the type locality of 
<taxonomicName id="4C4C4D6CFFA6FF9DFE69F83A1EC442B4" authorityName=": Dawson" authorityYear="1982" box="[504,692,1980,2006]" class="Actinopterygii" family="Syngnathidae" genus="Syngnathus" kingdom="Animalia" order="Syngnathiformes" pageId="12" pageNumber="93" phylum="Chordata" rank="species" species="elcapitanensis">
<emphasis id="B938EAFDFFA6FF9DFE69F83A1EC442B4" box="[504,692,1980,2006]" italics="true" pageId="12" pageNumber="93">P. elcapitanensis</emphasis>
</taxonomicName>
. In the original description of the species, Meek &amp; Hildebrand (in 
<bibRefCitation id="EFDD4B1EFFA6FF9DFF06F8661D6E4298" author="Meek, S. E." box="[151,286,2016,2042]" pageId="12" pageNumber="93" pagination="101 - 134" refId="ref28392" refString="Meek, S. E. (1914) An annotated list of fishes known to occur in the fresh waters of Costa Rica. Field Museum of Natural History, Zoological Series, 10, 101 - 134. https: // doi. org / 10.5962 / bhl. title. 5576" type="journal article" year="1914">Meek, 1914</bibRefCitation>
) did not expressly indicate a 
<typeStatus id="54F7884DFFA6FF9DFDF6F8661EBD4298" box="[615,717,2016,2042]" pageId="12" pageNumber="93" type="holotype">holotype</typeStatus>
. Later, in addition to citing USNM 81735 as the type of the spe-cies, another specimen was indicated as a 
<typeStatus id="54F7884DFFA7FF9CFDEDFF111EAC45D3" box="[636,732,151,177]" pageId="13" pageNumber="94" type="paratype">paratype</typeStatus>
(
<bibRefCitation id="EFDD4B1EFFA7FF9CFD7CFF11185045D3" author="Meek, S. &amp; Hildebrand, S. F." box="[749,1056,151,177]" pageId="13" pageNumber="94" pagination="1 - 330" refId="ref28446" refString="Meek, S. &amp; Hildebrand, S. F. (1923) The marine fishes of Panama. Part 1. Field Museum of Natural History, Zoological Series, 15, 1 - 330." type="journal article" year="1923">Meek &amp; Hildebrand, 1923</bibRefCitation>
: plate XIX b), which is currently catalogued as USNM 108645 and, according to the institution catalogue records, was removed from the lot USNM 81735. The specimens FMNH 8291 and 8292 seem to have originated from USNM 79690 and were also available to the authors when they established the nominal species (based on information on the labels inside the jars, mentioned above). Therefore, those are the five specimens that constitute the type series of the species (ICZN, 1999: Art. 72.4.1.1 and 73.2). 
<bibRefCitation id="EFDD4B1EFFA7FF9CFE36FECD1E9F4407" author="Meek, S. &amp; Hildebrand, S. F." box="[423,751,331,357]" pageId="13" pageNumber="94" pagination="1 - 330" refId="ref28446" refString="Meek, S. &amp; Hildebrand, S. F. (1923) The marine fishes of Panama. Part 1. Field Museum of Natural History, Zoological Series, 15, 1 - 330." type="journal article" year="1923">Meek &amp; Hildebrand´s (1923)</bibRefCitation>
citation of USNM 81735 as type, would therefore constitute a 
<typeStatus id="54F7884DFFA7FF9CFF3AFEE91D6244EB" box="[171,274,367,393]" pageId="13" pageNumber="94" type="lectotype">lectotype</typeStatus>
designation, an interpretation in line with that of Ferraris Jr. (2003: 591). The additional five specimens from 
<collectingCountry id="F35B767FFFA7FF9CFF45FE151D4044CF" box="[212,304,403,429]" name="Panama" pageId="13" pageNumber="94">Panama</collectingCountry>
, (USNM 79687, 79689 and FMNH 8290, FMNH 8293 and FMNH 8294) and the two specimens from 
<collectingCountry id="F35B767FFFA7FF9CFF06FE311D6444B3" box="[151,276,439,465]" name="Costa Rica" pageId="13" pageNumber="94">Costa Rica</collectingCountry>
, listed at the end of the description of 
<taxonomicName id="4C4C4D6CFFA7FF9CFD2DFE311F0744B3" authorityName=": Dawson" authorityYear="1982" box="[700,887,439,465]" class="Actinopterygii" family="Syngnathidae" genus="Syngnathus" kingdom="Animalia" order="Syngnathiformes" pageId="13" pageNumber="94" phylum="Chordata" rank="species" species="elcapitanensis">
<emphasis id="B938EAFDFFA7FF9CFD2DFE311F0744B3" box="[700,887,439,465]" italics="true" pageId="13" pageNumber="94">P. elcapitanensis</emphasis>
</taxonomicName>
by Meek &amp; Hildebrand (in 
<bibRefCitation id="EFDD4B1EFFA7FF9CFB3CFE31194344B3" author="Meek, S. E." box="[1197,1331,439,465]" pageId="13" pageNumber="94" pagination="101 - 134" refId="ref28392" refString="Meek, S. E. (1914) An annotated list of fishes known to occur in the fresh waters of Costa Rica. Field Museum of Natural History, Zoological Series, 10, 101 - 134. https: // doi. org / 10.5962 / bhl. title. 5576" type="journal article" year="1914">Meek, 1914</bibRefCitation>
) (FMNH 7901 and FMNH 7916) have been sometimes erroneously considered as 
<typeStatus id="54F7884DFFA7FF9CFC50FE5D185C4497" box="[961,1068,475,501]" pageId="13" pageNumber="94" type="paratype">paratypes</typeStatus>
(e.g. 
<bibRefCitation id="EFDD4B1EFFA7FF9CFBFAFE5D189A4497" author="Grey, M." box="[1131,1258,475,501]" pageId="13" pageNumber="94" pagination="109 - 205" refId="ref26660" refString="Grey, M. (1947) Catalogue of type specimens of fishes in Chicago Natural History Museum. Fieldiana, Zoology, 32, 109 - 205." type="journal article" year="1947">Grey, 1947</bibRefCitation>
; 
<bibRefCitation id="EFDD4B1EFFA7FF9CFB64FE5D1D58477B" author="Ibarra, M. &amp; Stewart, D. J." pageId="13" pageNumber="94" pagination="1 - 112" refId="ref27089" refString="Ibarra, M. &amp; Stewart, D. J. (1987) Catalogue of type specimens of recent fishes in Field Museum of Natural History. Fieldiana Zoology, New Series, 35, 1 - 112. https: // doi. org / 10.5962 / bhl. title. 3000" type="journal article" year="1987">Ibarra &amp; Stewart, 1987: 80</bibRefCitation>
). The two specimens from 
<collectingCountry id="F35B767FFFA7FF9CFDC4FE791EBE477B" box="[597,718,511,537]" name="Costa Rica" pageId="13" pageNumber="94">Costa Rica</collectingCountry>
were identified by 
<bibRefCitation id="EFDD4B1EFFA7FF9CFC37FE791822477B" author="Dawson, C. E." box="[934,1106,511,537]" pageId="13" pageNumber="94" pagination="1 - 172" refId="ref25847" refString="Dawson, C. E. (1982) Fishes of the Western North Atlantic. The pipefishes (subfamilies Doryrhamphinae and Syngnathinae). Memoirs of the Sears Foundation for Marine Research, 1 (8), 1 - 172." type="journal article" year="1982">Dawson (1982)</bibRefCitation>
as 
<taxonomicName id="4C4C4D6CFFA7FF9CFBE6FE791904477B" box="[1143,1396,511,537]" class="Actinopterygii" family="Syngnathidae" genus="Pseudophallus" kingdom="Animalia" order="Syngnathiformes" pageId="13" pageNumber="94" phylum="Chordata" rank="species" species="starksii">
<emphasis id="B938EAFDFFA7FF9CFBE6FE791904477B" box="[1143,1396,511,537]" italics="true" pageId="13" pageNumber="94">Pseudophallus starksii</emphasis>
</taxonomicName>
, an identification confirmed in the present study.
</paragraph>
</subSubSection>
<subSubSection id="C3566564FFA7FF9CFF56FDC118A3418B" pageId="13" pageNumber="94" type="materials_examined">
<paragraph id="8BF336EFFFA7FF9CFF56FDC11CA847AC" blockId="13.[151,1437,151,1257]" pageId="13" pageNumber="94">
<emphasis id="B938EAFDFFA7FF9CFF56FDC11D074702" bold="true" box="[199,375,582,609]" pageId="13" pageNumber="94">Type material:</emphasis>
<materialsCitation id="3B243CB2FFA7FF9CFEECFDC11D9247E4" ID-GBIF-Occurrence="3034656432" collectingDate="1912-03-20" collectionCode="USNM" collectorName="S. E. Meek &amp; S. F. Hildebrand" country="Panama" location="Rio Mamoni" pageId="13" pageNumber="94" specimenCode="USNM 81735, 1" specimenCount="1" typeStatus="lectotype">
<emphasis id="B938EAFDFFA7FF9CFEECFDC11D9F4703" bold="true" box="[381,495,583,609]" pageId="13" pageNumber="94">
<typeStatus id="54F7884DFFA7FF9CFEECFDC11D9A4703" box="[381,490,583,609]" pageId="13" pageNumber="94" type="lectotype">lectotype</typeStatus>
.
</emphasis>
<specimenCode id="DBEA9E94FFA7FF9CFE67FDC11EE64703" box="[502,662,582,609]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="13" pageNumber="94" type="Museum">USNM 81735</specimenCode>
, 
<specimenCode id="DBEA9E94FFA7FF9CFD31FDC11EDE4703" box="[672,686,583,609]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="13" pageNumber="94" type="Museum">1</specimenCode>
, 
<collectorName id="26B95339FFA7FF9CFD2DFDC11F4D4703" box="[700,829,583,609]" pageId="13" pageNumber="94">El Capitain</collectorName>
, 
<location id="8E936034FFA7FF9CFCD6FDC11FA84703" LSID="urn:lsid:plazi:treatment:03E587F9FFA0FF9CFF06F9B218A3418B:8E936034FFA7FF9CFCD6FDC11FA84703" box="[839,984,583,609]" country="Panama" name="Rio Mamoni" pageId="13" pageNumber="94">Río Mamoni</location>
, 
<collectingCountry id="F35B767FFFA7FF9CFC72FDC1184F4703" box="[995,1087,583,609]" name="Panama" pageId="13" pageNumber="94">Panamá</collectingCountry>
, 
<collectorName id="26B95339FFA7FF9CFBD8FDC118B44703" box="[1097,1220,583,609]" pageId="13" pageNumber="94">S. E. Meek</collectorName>
&amp; 
<collectorName id="26B95339FFA7FF9CFB76FDC119EC4703" box="[1255,1436,583,609]" pageId="13" pageNumber="94">S. F. Hildebrand</collectorName>
<date id="FFF2102FFFA7FF9CFF06FDED1D5A47E7" box="[151,298,619,645]" pageId="13" pageNumber="94" value="1912-03-20">
<collectingDate id="EFB6E9C7FFA7FF9CFF06FDED1D5A47E7" box="[151,298,619,645]" pageId="13" pageNumber="94" value="1912-03-20">20 Mar 1912</collectingDate>
</date>
(not examined)
</materialsCitation>
. 
<materialsCitation id="3B243CB2FFA7FF9CFE7CFDED1CA447AC" ID-GBIF-Occurrence="3034656398" collectingDate="1912-03-20" collectionCode="USNM" country="Panama" location="Rio Mamoni" pageId="13" pageNumber="94" specimenCode="USNM 79690, 1, USNM 108645" specimenCount="3" typeStatus="paralectotype">
<typeStatus id="54F7884DFFA7FF9CFE7CFDED1EE447E7" box="[493,660,619,645]" pageId="13" pageNumber="94" type="paralectotype">Paralectotypes</typeStatus>
. FMNH 8291/8292, 2, 108.5– 
<quantity id="4CB49B0AFFA7FF9CFC7AFDEC182E47E7" box="[1003,1118,618,645]" metricMagnitude="-1" metricUnit="m" metricValue="1.308" pageId="13" pageNumber="94" unit="mm" value="130.8">130.8 mm</quantity>
SL, same data as lectotype; 
<specimenCode id="DBEA9E94FFA7FF9CFF06FD091D4B47CB" box="[151,315,655,681]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="13" pageNumber="94" type="Museum">USNM 79690</specimenCode>
, 
<specimenCode id="DBEA9E94FFA7FF9CFED8FD091D2747CB" box="[329,343,655,681]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="13" pageNumber="94" type="Museum">1</specimenCode>
, same data as 
<typeStatus id="54F7884DFFA7FF9CFD95FD091E1B47CB" box="[516,619,655,681]" pageId="13" pageNumber="94" type="lectotype">lectotype</typeStatus>
(not examined); 
<specimenCode id="DBEA9E94FFA7FF9CFCA5FD091F9547CB" box="[820,997,654,681]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="13" pageNumber="94" type="Museum">USNM 108645</specimenCode>
, 
<specimenCode id="DBEA9E94FFA7FF9CFC62FD09187147CB" box="[1011,1025,655,681]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="13" pageNumber="94" type="Museum">1</specimenCode>
, same data as 
<typeStatus id="54F7884DFFA7FF9CFB3EFD09196647CB" box="[1199,1302,655,681]" pageId="13" pageNumber="94" type="lectotype">lectotype</typeStatus>
(not examined)
</materialsCitation>
.
</paragraph>
<paragraph id="8BF336EFFFA7FF9CFF56FD51190E41C3" blockId="13.[151,1437,151,1257]" pageId="13" pageNumber="94">
<emphasis id="B938EAFDFFA7FF9CFF56FD511D384793" bold="true" box="[199,328,727,753]" pageId="13" pageNumber="94">Non-types.</emphasis>
<materialsCitation id="3B243CB2FFA7FF9CFEC1FD5119E74793" ID-GBIF-Occurrence="3034656337" box="[336,1431,726,753]" collectingDate="1924-02-24" collectorName="Marsh Expedition" country="Panama" location="Rio Chico" pageId="13" pageNumber="94" specimenCount="1" stateProvince="Darien">
<emphasis id="B938EAFDFFA7FF9CFEC1FD511DC24793" bold="true" box="[336,434,727,753]" pageId="13" pageNumber="94">
<collectingCountry id="F35B767FFFA7FF9CFEC1FD511DC24793" box="[336,434,727,753]" name="Panama" pageId="13" pageNumber="94">Panama</collectingCountry>
</emphasis>
: 
<emphasis id="B938EAFDFFA7FF9CFE52FD501E6F4792" bold="true" box="[451,543,726,752]" pageId="13" pageNumber="94">
<collectingRegion id="4988F80DFFA7FF9CFE52FD501E6A4792" box="[451,538,726,752]" country="Panama" name="Darien" pageId="13" pageNumber="94">Darien</collectingRegion>
:
</emphasis>
AMNH 11376, 1, 125.0 mm SL, 
<location id="8E936034FFA7FF9CFC34FD51186C4793" LSID="urn:lsid:plazi:treatment:03E587F9FFA0FF9CFF06F9B218A3418B:8E936034FFA7FF9CFC34FD51186C4793" box="[933,1052,727,753]" country="Panama" name="Rio Chico" pageId="13" pageNumber="94" stateProvince="Darien">Río Chico</location>
, 
<collectorName id="26B95339FFA7FF9CFBB8FD5118884793" box="[1065,1272,727,753]" pageId="13" pageNumber="94">Marsh Expedition</collectorName>
, 
<date id="FFF2102FFFA7FF9CFA94FD5119E74793" box="[1285,1431,727,753]" pageId="13" pageNumber="94" value="1924-02-24">
<collectingDate id="EFB6E9C7FFA7FF9CFA94FD5119E74793" box="[1285,1431,727,753]" pageId="13" pageNumber="94" value="1924-02-24">24 Feb 1924</collectingDate>
</date>
</materialsCitation>
; 
<materialsCitation id="3B243CB2FFA7FF9CFF06FD7D19494677" ID-GBIF-Occurrence="3034656402" box="[151,1337,763,789]" collectingDate="1912-02-28" collectorName="S. E. Meek &amp; S. F. Hildebrand" country="Panama" location="Boca de Cupe" municipality="Rio Tuyra" pageId="13" pageNumber="94" specimenCount="1" stateProvince="Darien">
FMNH 8290, 
<quantity id="4CB49B0AFFA7FF9CFEABFD7D1DBC4677" box="[314,460,763,789]" metricMagnitude="0" metricUnit="m" metricValue="1.1229" pageId="13" pageNumber="94" unit="mm" value="1122.9">1, 122.9 mm</quantity>
SL, 
<collectingMunicipality id="6B97AC95FFA7FF9CFD97FD7D1E0C4677" box="[518,636,763,789]" pageId="13" pageNumber="94">Río Tuyra</collectingMunicipality>
, 
<location id="8E936034FFA7FF9CFD18FD7D1F5D4677" LSID="urn:lsid:plazi:treatment:03E587F9FFA0FF9CFF06F9B218A3418B:8E936034FFA7FF9CFD18FD7D1F5D4677" box="[649,813,763,789]" country="Panama" municipality="Rio Tuyra" name="Boca de Cupe" pageId="13" pageNumber="94" stateProvince="Darien">Boca de Cupe</location>
, 
<collectorName id="26B95339FFA7FF9CFCAAFD7D1FCC4677" box="[827,956,763,789]" pageId="13" pageNumber="94">S. E. Meek</collectorName>
&amp; 
<collectorName id="26B95339FFA7FF9CFC75FD7D18EF4677" box="[996,1183,763,789]" pageId="13" pageNumber="94">S. F. Hildebrand</collectorName>
<date id="FFF2102FFFA7FF9CFB39FD7D19494677" box="[1192,1337,763,789]" pageId="13" pageNumber="94" value="1912-02-28">
<collectingDate id="EFB6E9C7FFA7FF9CFB39FD7D19494677" box="[1192,1337,763,789]" pageId="13" pageNumber="94" value="1912-02-28">28 Feb 1912</collectingDate>
</date>
</materialsCitation>
; 
<materialsCitation id="3B243CB2FFA7FF9CFADAFD7D18DB465B" ID-GBIF-Occurrence="3034656319" accessionNumber="MCZ 88623" collectingDate="1911-03-18" collectorName="S. E. Meek &amp; S. F. Hildebrand" country="Panama" location="Rio Calobra" pageId="13" pageNumber="94" specimenCount="1" stateProvince="Darien">
FMNH 8294, 1, 
<quantity id="4CB49B0AFFA7FF9CFF69FC991D2B465B" box="[248,347,799,825]" metricMagnitude="-2" metricUnit="m" metricValue="7.39" pageId="13" pageNumber="94" unit="mm" value="73.9">73.9 mm</quantity>
SL, 
<location id="8E936034FFA7FF9CFE1EFC991E69465B" LSID="urn:lsid:plazi:treatment:03E587F9FFA0FF9CFF06F9B218A3418B:8E936034FFA7FF9CFE1EFC991E69465B" box="[399,537,799,825]" country="Panama" name="Rio Calobra" pageId="13" pageNumber="94" stateProvince="Darien">Río Calobra</location>
, 
<collectorName id="26B95339FFA7FF9CFDB2FC991EEE465B" box="[547,670,799,825]" pageId="13" pageNumber="94">S. E. Meek</collectorName>
&amp; 
<collectorName id="26B95339FFA7FF9CFD51FC991F08465B" box="[704,888,799,825]" pageId="13" pageNumber="94">S. F. Hildebrand</collectorName>
, 
<date id="FFF2102FFFA7FF9CFC12FC991864465B" box="[899,1044,799,825]" pageId="13" pageNumber="94" value="1911-03-18">
<collectingDate id="EFB6E9C7FFA7FF9CFC12FC991864465B" box="[899,1044,799,825]" pageId="13" pageNumber="94" value="1911-03-18">18 Mar 1911</collectingDate>
</date>
; MCZ 88623
</materialsCitation>
, 
<materialsCitation id="3B243CB2FFA7FF9CFB24FC9918F6463F" ID-GBIF-Occurrence="3034656333" collectingDate="1985-02-16" collectorName="K. E. Hartel" country="Panama" latitude="8.0" location="Rio Tuira" longLatPrecision="1302" longitude="-77.53333" municipality="Rio Pucuro" pageId="13" pageNumber="94" specimenCount="1" stateProvince="Darien">
<quantity id="4CB49B0AFFA7FF9CFB24FC991930465B" box="[1205,1344,799,825]" metricMagnitude="0" metricUnit="m" metricValue="1.1105999999999998" pageId="13" pageNumber="94" unit="mm" value="1110.6">1, 110.6 mm</quantity>
SL, 
<collectingMunicipality id="6B97AC95FFA7FF9CFAE5FC991C96463F" pageId="13" pageNumber="94">Río Pucuro</collectingMunicipality>
just above confluence with 
<location id="8E936034FFA7FF9CFDB0FCC51EFF463F" LSID="urn:lsid:plazi:treatment:03E587F9FFA0FF9CFF06F9B218A3418B:8E936034FFA7FF9CFDB0FCC51EFF463F" box="[545,655,835,861]" country="Panama" latitude="8.0" longLatPrecision="1302" longitude="-77.53333" municipality="Rio Pucuro" name="Rio Tuira" pageId="13" pageNumber="94" stateProvince="Darien">Río Tuira</location>
, 
<geoCoordinate id="EE785028FFA7FF9CFD0BFCC51E9A463F" box="[666,746,835,861]" degrees="08" direction="north" minutes="0" orientation="latitude" pageId="13" pageNumber="94" precision="925" value="8.0">08º0’N</geoCoordinate>
<geoCoordinate id="EE785028FFA7FF9CFD60FCC51F26463F" box="[753,854,835,861]" degrees="77" direction="west" minutes="32" orientation="longitude" pageId="13" pageNumber="94" precision="925" value="-77.53333">77º32’W</geoCoordinate>
, 
<collectorName id="26B95339FFA7FF9CFCF0FCC51F9C463F" box="[865,1004,835,861]" pageId="13" pageNumber="94">K. E. Hartel</collectorName>
, 
<date id="FFF2102FFFA7FF9CFC66FCC518F6463F" box="[1015,1158,834,861]" pageId="13" pageNumber="94" value="1985-02-16">
<collectingDate id="EFB6E9C7FFA7FF9CFC66FCC518F6463F" box="[1015,1158,834,861]" pageId="13" pageNumber="94" value="1985-02-16">16 Feb 1985</collectingDate>
</date>
</materialsCitation>
; 
<materialsCitation id="3B243CB2FFA7FF9CFB00FCC51F5146C7" ID-GBIF-Occurrence="3034656437" collectingDate="1911-03-21" collectionCode="CAS" collectorName=" S. E. Meek &amp; S. F. Hildebrand" country="Panama" county="Rio Grande" location="Rio Mamoni" municipality="Hilbert" pageId="13" pageNumber="94" specimenCode="CAS 106919, 2" specimenCount="1">
<emphasis id="B938EAFDFFA7FF9CFB00FCC5188D463F" bold="true" box="[1169,1277,835,861]" pageId="13" pageNumber="94">
<collectingCountry id="F35B767FFFA7FF9CFB00FCC51887463F" box="[1169,1271,835,861]" name="Panama" pageId="13" pageNumber="94">Panama</collectingCountry>
:
</emphasis>
<specimenCode id="DBEA9E94FFA7FF9CFA95FCC519E8463F" box="[1284,1432,835,861]" collectionCode="CAS" country="USA" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="13" pageNumber="94" type="Museum">CAS 106919</specimenCode>
, 
<specimenCode id="DBEA9E94FFA7FF9CFF06FCE11CD746E3" box="[151,167,871,897]" collectionCode="CAS" country="USA" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="13" pageNumber="94" type="Museum">2</specimenCode>
, 
<quantity id="4CB49B0AFFA7FF9CFF25FCE11D1646E3" box="[180,358,871,897]" metricMagnitude="-1" metricUnit="m" metricValue="1.0294999999999999" metricValueMax="1.193" metricValueMin="0.866" pageId="13" pageNumber="94" unit="mm" value="102.94999999999999" valueMax="119.3" valueMin="86.6">86.6–119.3 mm</quantity>
SL, 
<collectingCounty id="62924E63FFA7FF9CFE0CFCE11E5346E3" box="[413,547,871,897]" pageId="13" pageNumber="94">Río Grande</collectingCounty>
, 
<collectorName id="26B95339FFA7FF9CFDA1FCE11E9E46E3" box="[560,750,871,897]" pageId="13" pageNumber="94">Miraflores, C. H</collectorName>
. 
<collectingMunicipality id="6B97AC95FFA7FF9CFD6BFCE11F3946E3" box="[762,841,871,897]" pageId="13" pageNumber="94">Hilbert</collectingMunicipality>
<emphasis id="B938EAFDFFA7FF9CFCC3FCE11FE246E3" box="[850,914,871,897]" italics="true" pageId="13" pageNumber="94">et al.,</emphasis>
1900; FMNH 8293, 1, 
<quantity id="4CB49B0AFFA7FF9CFB31FCE0197546E3" box="[1184,1285,870,897]" metricMagnitude="-2" metricUnit="m" metricValue="8.53" pageId="13" pageNumber="94" unit="mm" value="85.3">85.3 mm</quantity>
SL, 
<location id="8E936034FFA7FF9CFAACFCE11CA346C7" LSID="urn:lsid:plazi:treatment:03E587F9FFA0FF9CFF06F9B218A3418B:8E936034FFA7FF9CFAACFCE11CA346C7" country="Panama" county="Rio Grande" municipality="Hilbert" name="Rio Mamoni" pageId="13" pageNumber="94">Río Mamoni</location>
, 
<location id="8E936034FFA7FF9CFF4FFC0D1D5A46C7" LSID="urn:lsid:plazi:treatment:03E587F9FFA0FF9CFF06F9B218A3418B:8E936034FFA7FF9CFF4FFC0D1D5A46C7" box="[222,298,907,933]" country="Panama" county="Rio Grande" municipality="Hilbert" name="Chepo" pageId="13" pageNumber="94">Chepo</location>
, 
<collectorName id="26B95339FFA7FF9CFEA4FC0D1DC146C7" box="[309,433,907,933]" pageId="13" pageNumber="94">S. E. Meek</collectorName>
&amp; 
<collectorName id="26B95339FFA7FF9CFE42FC0D1EF946C7" box="[467,649,907,933]" pageId="13" pageNumber="94">S. F. Hildebrand</collectorName>
<date id="FFF2102FFFA7FF9CFD1EFC0D1F5146C7" box="[655,801,907,933]" pageId="13" pageNumber="94" value="1911-03-21">
<collectingDate id="EFB6E9C7FFA7FF9CFD1EFC0D1F5146C7" box="[655,801,907,933]" pageId="13" pageNumber="94" value="1911-03-21">21 Mar 1911</collectingDate>
</date>
</materialsCitation>
; 
<materialsCitation id="3B243CB2FFA7FF9CFCBDFC0C183546AB" ID-GBIF-Occurrence="3034656335" collectingDate="1961-11-18" collectorName=" Loftin &amp; E. L. Tyson" country="Panama" location="Boca del Monte" pageId="13" pageNumber="94" specimenCount="1" stateProvince="Chiriqui">
<emphasis id="B938EAFDFFA7FF9CFCBDFC0C1FEC46C6" bold="true" box="[812,924,906,932]" pageId="13" pageNumber="94">
<collectingRegion id="4988F80DFFA7FF9CFCBDFC0C1FE746C6" box="[812,919,906,932]" country="Panama" name="Chiriqui" pageId="13" pageNumber="94">Chiriqui</collectingRegion>
:
</emphasis>
ANSP 99933, 
<quantity id="4CB49B0AFFA7FF9CFBD3FC0D18BE46C7" box="[1090,1230,906,933]" metricMagnitude="0" metricUnit="m" metricValue="1.1159" pageId="13" pageNumber="94" unit="mm" value="1115.9">1, 115.9 mm</quantity>
SL, creek 
<quantity id="4CB49B0AFFA7FF9CFAD4FC0C190846C7" box="[1349,1400,906,933]" metricMagnitude="3" metricUnit="m" metricValue="8.04672" pageId="13" pageNumber="94" unit="mi" value="5.0">5 mi</quantity>
W. of 
<location id="8E936034FFA7FF9CFF27FC291D1A46AB" LSID="urn:lsid:plazi:treatment:03E587F9FFA0FF9CFF06F9B218A3418B:8E936034FFA7FF9CFF27FC291D1A46AB" box="[182,362,943,969]" country="Panama" name="Boca del Monte" pageId="13" pageNumber="94" stateProvince="Chiriqui">Boca del Monte</location>
on old road to 
<collectorName id="26B95339FFA7FF9CFD88FC291ED646AB" box="[537,678,943,969]" pageId="13" pageNumber="94">David, H. G</collectorName>
. 
<collectorName id="26B95339FFA7FF9CFD23FC291E8846AB" box="[690,760,943,969]" pageId="13" pageNumber="94">Loftin</collectorName>
&amp; 
<collectorName id="26B95339FFA7FF9CFC8CFC291FD346AB" box="[797,931,943,969]" pageId="13" pageNumber="94">E. L. Tyson</collectorName>
, 
<date id="FFF2102FFFA7FF9CFC3EFC29183546AB" box="[943,1093,943,969]" pageId="13" pageNumber="94" value="1961-11-18">
<collectingDate id="EFB6E9C7FFA7FF9CFC3EFC29183546AB" box="[943,1093,943,969]" pageId="13" pageNumber="94" value="1961-11-18">18 Nov 1961</collectingDate>
</date>
</materialsCitation>
; 
<materialsCitation id="3B243CB2FFA7FF9CFBC1FC291D0D4173" ID-GBIF-Occurrence="3034656411" collectingDate="1961-11-18" collectorName="H. G. Loftin &amp; E. L. Tyson" country="Panama" location="Creek" pageId="13" pageNumber="94" specimenCount="1" stateProvince="Chiriqui">
ANSP 104124, 4, 64.4–130.0 mm SL, 
<location id="8E936034FFA7FF9CFF69FC551D4B468F" LSID="urn:lsid:plazi:treatment:03E587F9FFA0FF9CFF06F9B218A3418B:8E936034FFA7FF9CFF69FC551D4B468F" box="[248,315,979,1005]" country="Panama" name="Creek" pageId="13" pageNumber="94" stateProvince="Chiriqui">Creek</location>
at bridge 
<quantity id="4CB49B0AFFA7FF9CFE3DFC541D96468F" box="[428,486,978,1005]" metricMagnitude="3" metricUnit="m" metricValue="8.04672" pageId="13" pageNumber="94" unit="mi" value="5.0">5 mi.</quantity>
W. of San Felix bridge on Interamerican Highway new section, 
<collectorName id="26B95339FFA7FF9CFB26FC551931468F" box="[1207,1345,979,1005]" pageId="13" pageNumber="94">H. G. Loftin</collectorName>
&amp; 
<collectorName id="26B95339FFA7FF9CFAF4FC551CAE4173" pageId="13" pageNumber="94">E. L. Tyson</collectorName>
, 
<date id="FFF2102FFFA7FF9CFF78FC711D0D4173" box="[233,381,1015,1041]" pageId="13" pageNumber="94" value="1961-11-18">
<collectingDate id="EFB6E9C7FFA7FF9CFF78FC711D0D4173" box="[233,381,1015,1041]" pageId="13" pageNumber="94" value="1961-11-18">18 Nov 1961</collectingDate>
</date>
</materialsCitation>
; 
<materialsCitation id="3B243CB2FFA7FF9CFE16FC711F994157" ID-GBIF-Occurrence="3034656386" collectingDate="1961-11-11" collectorName="H. G. Loftin &amp; E. L. Tyson &amp; Canipe" country="Panama" location="Rio Tabasara on Sona-Remedios" municipality="Rio Tinta" pageId="13" pageNumber="94" specimenCount="1" stateProvince="Chiriqui">
ANSP 104212, 2, 105.2– 
<quantity id="4CB49B0AFFA7FF9CFD0EFC701F7F4173" box="[671,783,1014,1041]" metricMagnitude="-1" metricUnit="m" metricValue="1.063" pageId="13" pageNumber="94" unit="mm" value="106.3">106.3 mm</quantity>
SL, 
<collectingMunicipality id="6B97AC95FFA7FF9CFCD5FC711FDF4173" box="[836,943,1015,1041]" pageId="13" pageNumber="94">Río Tinta</collectingMunicipality>
, creek at bridge 
<quantity id="4CB49B0AFFA7FF9CFBF7FC7118EF4173" box="[1126,1183,1015,1041]" metricMagnitude="3" metricUnit="m" metricValue="4.828032" pageId="13" pageNumber="94" unit="mi" value="3.0">3 mi.</quantity>
W. of 
<location id="8E936034FFA7FF9CFB79FC711D384157" LSID="urn:lsid:plazi:treatment:03E587F9FFA0FF9CFF06F9B218A3418B:8E936034FFA7FF9CFB79FC711D384157" country="Panama" municipality="Rio Tinta" name="Rio Tabasara on Sona-Remedios" pageId="13" pageNumber="94" stateProvince="Chiriqui">Río Tabasara on Sona-Remedios</location>
road, 
<collectorName id="26B95339FFA7FF9CFE01FB9D1E6E4157" box="[400,542,1051,1077]" pageId="13" pageNumber="94">H. G. Loftin</collectorName>
, 
<collectorName id="26B95339FFA7FF9CFDB8FB9D1EDB4157" box="[553,683,1051,1077]" pageId="13" pageNumber="94">E. L. Tyson</collectorName>
&amp; M. 
<collectorName id="26B95339FFA7FF9CFD67FB9D1F384157" box="[758,840,1051,1077]" pageId="13" pageNumber="94">Canipe</collectorName>
, 
<date id="FFF2102FFFA7FF9CFCC5FB9D1F994157" box="[852,1001,1051,1077]" pageId="13" pageNumber="94" value="1961-11-11">
<collectingDate id="EFB6E9C7FFA7FF9CFCC5FB9D1F994157" box="[852,1001,1051,1077]" pageId="13" pageNumber="94" value="1961-11-11">11 Nov 1961</collectingDate>
</date>
</materialsCitation>
; 
<materialsCitation id="3B243CB2FFA7FF9CFC62FB9D19E7413B" ID-GBIF-Occurrence="3034656368" collectingDate="1961-11-12" collectorName="Remedios, H. G &amp; Loftin, E. L &amp; Tyson &amp; Canipe" country="Panama" location="Creek Chacarrera on Sona-Remedios Rd." pageId="13" pageNumber="94" specimenCount="1" stateProvince="Chiriqui">
ANSP 104342, 1, 
<quantity id="4CB49B0AFFA7FF9CFB2EFB9D19544157" box="[1215,1316,1051,1077]" metricMagnitude="-2" metricUnit="m" metricValue="8.059999999999999" pageId="13" pageNumber="94" unit="mm" value="80.6">80.6 mm</quantity>
SL, 
<location id="8E936034FFA7FF9CFAC8FB9D1E6F413B" LSID="urn:lsid:plazi:treatment:03E587F9FFA0FF9CFF06F9B218A3418B:8E936034FFA7FF9CFAC8FB9D1E6F413B" country="Panama" name="Creek Chacarrera on Sona-Remedios Rd." pageId="13" pageNumber="94" stateProvince="Chiriqui">Creek Chacarrera on Sona-Remedios Rd.</location>
<quantity id="4CB49B0AFFA7FF9CFDB6FBB91E00413B" box="[551,624,1087,1113]" metricMagnitude="4" metricUnit="m" metricValue="2.2530816" pageId="13" pageNumber="94" unit="mi" value="14.0">14 mi.</quantity>
E. of 
<collectorName id="26B95339FFA7FF9CFD26FBB91F00413B" box="[695,880,1087,1113]" pageId="13" pageNumber="94">Remedios, H. G</collectorName>
. 
<collectorName id="26B95339FFA7FF9CFCEDFBB91875413B" box="[892,1029,1087,1113]" pageId="13" pageNumber="94">Loftin, E. L</collectorName>
. 
<collectorName id="26B95339FFA7FF9CFB81FBB91824413B" box="[1040,1108,1087,1113]" pageId="13" pageNumber="94">Tyson</collectorName>
&amp; M. 
<collectorName id="26B95339FFA7FF9CFB33FBB91884413B" box="[1186,1268,1087,1113]" pageId="13" pageNumber="94">Canipe</collectorName>
, 
<date id="FFF2102FFFA7FF9CFA91FBB919E7413B" box="[1280,1431,1087,1113]" pageId="13" pageNumber="94" value="1961-11-12">
<collectingDate id="EFB6E9C7FFA7FF9CFA91FBB919E7413B" box="[1280,1431,1087,1113]" pageId="13" pageNumber="94" value="1961-11-12">12 Nov 1961</collectingDate>
</date>
</materialsCitation>
; 
<materialsCitation id="3B243CB2FFA7FF9CFF06FBE51D5C41C3" ID-GBIF-Occurrence="3034656378" collectingDate="1961-11-18" collectionCode="USNM" collectorName="Am. Highway &amp; H. G. Loftin &amp; E. L. Tyson" country="Panama" elevation="14" location="San Juan" municipality="Creek" pageId="13" pageNumber="94" specimenCode="USNM 208369, 8" specimenCount="14" stateProvince="Chiriqui">
<specimenCode id="DBEA9E94FFA7FF9CFF06FBE51D35411F" box="[151,325,1123,1149]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="13" pageNumber="94" type="Museum">USNM 208369</specimenCode>
, 
<specimenCode id="DBEA9E94FFA7FF9CFEC1FBE51D2F411F" box="[336,351,1123,1149]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="13" pageNumber="94" type="Museum">8</specimenCode>
, 
<quantity id="4CB49B0AFFA7FF9CFEFAFBE51E6B411F" box="[363,539,1123,1149]" metricMagnitude="-2" metricUnit="m" metricValue="8.62" metricValueMax="10.01" metricValueMin="7.2299999999999995" pageId="13" pageNumber="94" unit="mm" value="86.19999999999999" valueMax="100.1" valueMin="72.3">72.3–100.1 mm</quantity>
SL, 
<collectingMunicipality id="6B97AC95FFA7FF9CFDDEFBE51EE2411F" box="[591,658,1123,1149]" pageId="13" pageNumber="94">Creek</collectingMunicipality>
<specimenCount id="9D4AFD66FFA7FF9CFD09FBE51EA7411F" box="[664,727,1123,1149]" pageId="13" pageNumber="94" type="generic">
<quantity id="4CB49B0AFFA7FF9CFD09FBE51EA3411F" box="[664,723,1123,1149]" metricMagnitude="1" metricUnit="m" metricValue="1.4" pageId="13" pageNumber="94" unit="m" value="14.0">
<elevation id="0061D1DCFFA7FF9CFD09FBE51EA3411F" box="[664,723,1123,1149]" metricMagnitude="1" metricUnit="m" metricValue="1.4" pageId="13" pageNumber="94" unit="m" value="14.0">14 m</elevation>
</quantity>
.
</specimenCount>
W. of 
<location id="8E936034FFA7FF9CFC8EFBE51FF3411F" LSID="urn:lsid:plazi:treatment:03E587F9FFA0FF9CFF06F9B218A3418B:8E936034FFA7FF9CFC8EFBE51FF3411F" box="[799,899,1123,1149]" country="Panama" municipality="Creek" name="San Juan" pageId="13" pageNumber="94" stateProvince="Chiriqui">San Juan</location>
on I. 
<collectorName id="26B95339FFA7FF9CFC51FBE51810411F" box="[960,1120,1123,1149]" pageId="13" pageNumber="94">Am. Highway</collectorName>
, 
<collectorName id="26B95339FFA7FF9CFBFBFBE51883411F" box="[1130,1267,1123,1149]" pageId="13" pageNumber="94">H. G. Loftin</collectorName>
&amp; 
<collectorName id="26B95339FFA7FF9CFA84FBE519E8411F" box="[1301,1432,1123,1149]" pageId="13" pageNumber="94">E. L. Tyson</collectorName>
, 
<date id="FFF2102FFFA7FF9CFF06FB011D5C41C3" box="[151,300,1159,1185]" pageId="13" pageNumber="94" value="1961-11-18">
<collectingDate id="EFB6E9C7FFA7FF9CFF06FB011D5C41C3" box="[151,300,1159,1185]" pageId="13" pageNumber="94" value="1961-11-18">18 Nov 1961</collectingDate>
</date>
</materialsCitation>
; 
<materialsCitation id="3B243CB2FFA7FF9CFEA9FB01190A41C3" ID-GBIF-Occurrence="3034656322" box="[312,1402,1158,1185]" collectingDate="1961-12-01" collectionCode="USNM" collectorName="David, H. G &amp; Loftin" country="Panama" location="Rio Estero Salado" pageId="13" pageNumber="94" specimenCode="USNM 208371, 12" specimenCount="1" stateProvince="Chiriqui">
<specimenCode id="DBEA9E94FFA7FF9CFEA9FB011D9741C3" box="[312,487,1159,1185]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="13" pageNumber="94" type="Museum">USNM 208371</specimenCode>
, 
<specimenCode id="DBEA9E94FFA7FF9CFE63FB011E6041C3" box="[498,528,1159,1185]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="13" pageNumber="94" type="Museum">12</specimenCode>
, 
<quantity id="4CB49B0AFFA7FF9CFD8DFB001EBD41C3" box="[540,717,1158,1185]" metricMagnitude="-1" metricUnit="m" metricValue="1.1139999999999999" metricValueMax="1.3519999999999999" metricValueMin="0.876" pageId="13" pageNumber="94" unit="mm" value="111.39999999999999" valueMax="135.2" valueMin="87.6">87.6–135.2 mm</quantity>
SL, 
<location id="8E936034FFA7FF9CFC93FB011FBC41C3" LSID="urn:lsid:plazi:treatment:03E587F9FFA0FF9CFF06F9B218A3418B:8E936034FFA7FF9CFC93FB011FBC41C3" box="[770,972,1159,1185]" country="Panama" name="Rio Estero Salado" pageId="13" pageNumber="94" stateProvince="Chiriqui">Río Estero Salado</location>
near 
<collectorName id="26B95339FFA7FF9CFB9AFB0118E641C3" box="[1035,1174,1159,1185]" pageId="13" pageNumber="94">David, H. G</collectorName>
. 
<collectorName id="26B95339FFA7FF9CFB33FB01189B41C3" box="[1186,1259,1159,1185]" pageId="13" pageNumber="94">Loftin</collectorName>
, 
<date id="FFF2102FFFA7FF9CFB67FB01190A41C3" box="[1270,1402,1159,1185]" pageId="13" pageNumber="94" value="1961-12-01">
<collectingDate id="EFB6E9C7FFA7FF9CFB67FB01190A41C3" box="[1270,1402,1159,1185]" pageId="13" pageNumber="94" value="1961-12-01">1 Dec 1961</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
<paragraph id="8BF336EFFFA7FF9CFF56FB2C18A3418B" blockId="13.[151,1437,151,1257]" pageId="13" pageNumber="94">
<materialsCitation id="3B243CB2FFA7FF9CFF56FB2C188041A7" ID-GBIF-Occurrence="3034656357" box="[199,1264,1194,1221]" collectingDate="1912-04-21" collectorName="Orotina, S. E" country="Costa Rica" location="Rio Turrubales" pageId="13" pageNumber="94" specimenCount="1">
<emphasis id="B938EAFDFFA7FF9CFF56FB2C1D3941A6" bold="true" box="[199,329,1194,1220]" pageId="13" pageNumber="94">
<collectingCountry id="F35B767FFFA7FF9CFF56FB2C1D3941A6" box="[199,329,1194,1220]" name="Costa Rica" pageId="13" pageNumber="94">Costa Rica</collectingCountry>
</emphasis>
: FMNH 7916, 
<quantity id="4CB49B0AFFA7FF9CFE66FB2D1EF641A7" box="[503,646,1195,1221]" metricMagnitude="0" metricUnit="m" metricValue="1.1047999999999998" pageId="13" pageNumber="94" unit="mm" value="1104.8">1, 104.8 mm</quantity>
SL, 
<location id="8E936034FFA7FF9CFD2AFB2D1F1641A7" LSID="urn:lsid:plazi:treatment:03E587F9FFA0FF9CFF06F9B218A3418B:8E936034FFA7FF9CFD2AFB2D1F1641A7" box="[699,870,1195,1221]" country="Costa Rica" name="Rio Turrubales" pageId="13" pageNumber="94">Río Turrubales</location>
, 
<collectorName id="26B95339FFA7FF9CFCE0FB2D187741A7" box="[881,1031,1195,1221]" pageId="13" pageNumber="94">Orotina, S. E</collectorName>
. Meek, 
<date id="FFF2102FFFA7FF9CFBF0FB2D188041A7" box="[1121,1264,1195,1221]" pageId="13" pageNumber="94" value="1912-04-21">
<collectingDate id="EFB6E9C7FFA7FF9CFBF0FB2D188041A7" box="[1121,1264,1195,1221]" pageId="13" pageNumber="94" value="1912-04-21">21 Apr 1912</collectingDate>
</date>
</materialsCitation>
; 
<materialsCitation id="3B243CB2FFA7FF9CFB6DFB2D18BE418B" ID-GBIF-Occurrence="3034656428" collectingDate="1986-08" collectorName="Rio Pavo, P. N &amp; K. Winemiller" country="Costa Rica" latitude="8.85" location="Rio Pavo" longLatPrecision="1300" longitude="-83.96667" pageId="13" pageNumber="94" specimenCount="1">
TNHC 14736, 2, 129.7– 
<quantity id="4CB49B0AFFA7FF9CFE91FB491D02418B" box="[256,370,1231,1257]" metricMagnitude="-1" metricUnit="m" metricValue="1.4819999999999998" pageId="13" pageNumber="94" unit="mm" value="148.2">148.2 mm</quantity>
SL, 
<collectorName id="26B95339FFA7FF9CFE36FB491E3E418B" box="[423,590,1231,1257]" pageId="13" pageNumber="94">
<location id="8E936034FFA7FF9CFE36FB491E62418B" LSID="urn:lsid:plazi:treatment:03E587F9FFA0FF9CFF06F9B218A3418B:8E936034FFA7FF9CFE36FB491E62418B" box="[423,530,1231,1257]" country="Costa Rica" latitude="8.85" longLatPrecision="1300" longitude="-83.96667" name="Rio Pavo" pageId="13" pageNumber="94">Río Pavo</location>
, P. N
</collectorName>
. Corcovado 
<geoCoordinate id="EE785028FFA7FF9CFD4AFB481F49418B" box="[731,825,1230,1257]" degrees="08" direction="north" minutes="51" orientation="latitude" pageId="13" pageNumber="94" precision="925" value="8.85">08º51’N</geoCoordinate>
<geoCoordinate id="EE785028FFA7FF9CFCD0FB481FD6418B" box="[833,934,1230,1257]" degrees="83" direction="west" minutes="58" orientation="longitude" pageId="13" pageNumber="94" precision="925" value="-83.96667">83º58’W</geoCoordinate>
, 
<collectorName id="26B95339FFA7FF9CFC20FB491822418B" box="[945,1106,1231,1257]" pageId="13" pageNumber="94">K. Winemiller</collectorName>
, 
<date id="FFF2102FFFA7FF9CFBCDFB4918BE418B" box="[1116,1230,1231,1257]" pageId="13" pageNumber="94" value="1986-08">
<collectingDate id="EFB6E9C7FFA7FF9CFBCDFB4918BE418B" box="[1116,1230,1231,1257]" pageId="13" pageNumber="94" value="1986-08">Aug 1986</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>