358 lines
57 KiB
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358 lines
57 KiB
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<document id="229D32511E141D9361BAACB22DCF664C" ID-CLB-Dataset="56065" ID-DOI="10.5852/ejt.2022.809.1719" ID-GBIF-Dataset="f76de735-01e3-454c-a7f1-bbafd26686f5" ID-ISSN="2118-9773" ID-Zenodo-Dep="6419425" ID-ZooBank="A754493E-5466-4479-B515-AABEDDE09D93" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1649273451180" checkinUser="felipe" docAuthor="Chigray, Svetlana N., Nabozhenko, Maxim V., Chigray, Ivan A. & Abakumov, Evgeny V." docDate="2022" docId="03FC86665574FFA6507CBF5EFC32FE06" docLanguage="en" docName="ejt-809_1-71.pdf" docOrigin="European Journal of Taxonomy 809" docSource="http://dx.doi.org/10.5852/ejt.2022.809.1719" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Mantichorula Semenovi Rtt. Coll. Reitter 1889" docType="treatment" docVersion="9" lastPageNumber="32" masterDocId="FFC5FE1E556AFF865230BB19FFBAFF8B" masterDocTitle="A revision of the Palaearctic Pimeliini (Coleoptera: Tenebrionidae): a comparative analysis and systematic position of Eastern European and Asian taxa with dorso-lateral eyes" masterLastPageNumber="71" masterPageNumber="1" pageNumber="31" updateTime="1699319505913" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="74BC278D24DC7632F07411D8FB1DDE11">A revision of the Palaearctic Pimeliini (Coleoptera: Tenebrionidae): a comparative analysis and systematic position of Eastern European and Asian taxa with dorso-lateral eyes</mods:title>
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<mods:namePart id="867CE8739BEACB7A61F326E9A5480C96">Chigray, Svetlana N.</mods:namePart>
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<mods:affiliation id="915D6138FF39E246B6B6C167E3F5285D">St. Petersburg State University, 16 line of Vasilevskiy Island, 29, St. Petersburg 199178 Russia.</mods:affiliation>
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<mods:namePart id="BB01A6B86737217F652355D859706355">Nabozhenko, Maxim V.</mods:namePart>
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<mods:affiliation id="4BDA2697C51F09EA678C83ACE42E5140">Precaspian Institute of Biological Resources of the Daghestan Federal Research Centre of the Russian Academy of Sciences, M. Gadzhiev str., 45, Makhachkala, Republic of Dagestan 367000 Russia. & Dagestan State University, M. Gadzhiev str., 43 a, Makhachkala, Republic of Dagestan 367000 Russia. & Severtsov Institute of ecology and evolution of the Russian Academy of sciences, Leninsky Prospect, 33, Moscow 119071 Russia.</mods:affiliation>
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<mods:affiliation id="40F16B9F387B3A049031C0BEA12218EB">Zoological Institute of the Russian Academy of Sciences, Universitetskaya Emb. 1, St. Petersburg 199034 Russia.</mods:affiliation>
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Genus
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<taxonomicName id="4C554CF35574FF9850AEBF5EFC52FBEA" authority="Reitter, 1889" authorityName="Semenovi Rtt. Coll. Reitter" authorityYear="1889" box="[670,1000,1095,1122]" class="Insecta" family="Tenebrionidae" genus="Mantichorula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="genus">
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<bibRefCitation id="EFC44A815574FF98517FBF5EFC52FBEA" author="Reitter E." box="[847,1000,1095,1121]" pageId="30" pageNumber="31" pagination="678 - 710" refId="ref32704" refString="Reitter E. 1889. Insecta, a cl. G. N. Potanin in China et in Mongolia novissime lecta. XIII. Tenebrionidae. Horae Societatis Entomologicae Rossicae 23: 678 - 710." type="journal article" year="1889">Reitter, 1889</bibRefCitation>
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<figureCitation id="136E2BF55574FF9853E6BF73FDE3FB0F" box="[470,601,1130,1156]" captionStart="Fig" captionStartId="9.[189,232,1892,1918]" captionTargetBox="[233,1353,649,1847]" captionTargetId="figure-90@9.[227,1361,643,1855]" captionTargetPageId="9" captionText="Fig. 4. Species of the genera Przewalskia Semenov, 1893, Mantichorula Reitter, 1889 and Platyope Fischer von Waldheim, 1820, habitus. A. Przewalskia dilatata (Reitter, 1887), ♀ (ZIN), dorsal view. B. Ditto, ventral view. C. M. semenowi Reitter, 1889, ♂ (ZIN), dorsal view. D. Ditto, ventral view. E. Platyope leucogramma (Pallas, 1773), ♂ (ZIN), dorsal view. F. Ditto, ventral view." figureDoi="http://doi.org/10.5281/zenodo.6419435" httpUri="https://zenodo.org/record/6419435/files/figure.png" pageId="30" pageNumber="31">Figs 4C–D</figureCitation>
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,
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<figureCitation id="136E2BF55574FF985054BF73FD08FB0F" box="[612,690,1130,1156]" captionStart="Fig" captionStartId="20.[189,232,1892,1918]" captionTargetBox="[189,1398,431,1851]" captionTargetId="figure-17@20.[189,1398,431,1853]" captionTargetPageId="20" captionText="Fig. 11. Przewalskia Semenov, 1893 and Mantichorula Reitter, 1889, details of structure. A. P. dilatata (Reitter, 1887), ♂ (ZIN), pronotum. B. Ditto, hemispherical large tubercles on pronotum. C. Ditto, base of left elytron. D. Ditto, pubescence of elytron. E. M. semenowi Reitter, 1889, ♂ (ZIN), pronotum. F. Ditto, tubercles and setation on sides of pronotum." figureDoi="http://doi.org/10.5281/zenodo.6419457" httpUri="https://zenodo.org/record/6419457/files/figure.png" pageId="30" pageNumber="31">11E–F</figureCitation>
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,
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<figureCitation id="136E2BF55574FF98508EBF73FCAFFB0F" box="[702,789,1130,1156]" captionStart="Fig" captionStartId="23.[189,232,1892,1918]" captionTargetBox="[189,1398,505,1853]" captionTargetId="figure-77@23.[189,1398,505,1853]" captionTargetPageId="23" captionText="Fig. 12. Mantichorula Reitter, 1889 and Platyope Fischer von Waldheim, 1820, details of structure. A. M. semenowi Reitter, 1889, ♂ (ZIN), base of left elytron. B. Ditto, pubescence of elytron. C. P. leucogramma (Pallas, 1773), ♂ (ZIN), pronotum. D. Ditto, hemispherical tubercles and pubescence of pronotum. E. Ditto, basal half of left elytron. F. Ditto, pubescence of elytron." figureDoi="http://doi.org/10.5281/zenodo.6419461" httpUri="https://zenodo.org/record/6419461/files/figure.png" pageId="30" pageNumber="31">12A–B</figureCitation>
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<figureCitation id="136E2BF55574FF985111BF73FCFAFB0F" box="[801,832,1130,1156]" captionStart="Fig" captionStartId="40.[191,234,1857,1883]" captionTargetBox="[189,1398,539,1819]" captionTargetId="figure-91@40.[189,1398,529,1819]" captionTargetPageId="40" captionText="Fig. 19. Mantichorula semenowi Reitter, 1889, genitalia (ZIN). A. Tegmen of aedeagus, ventrally. B. Ditto, dorsally. C. Ditto, laterally. D. Median lobe, ventrally. E. Ditto, dorsally. F. Ditto, laterally. G. Spiculum gastrale. H. ♂, inner sternite VIII. I. ♀, inner sternite VIII and spiculum ventrale. J. Ovipositor, ventrally. K. Ditto, dorsally. L. Ditto, laterally. Not to scale. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.6419481" httpUri="https://zenodo.org/record/6419481/files/figure.png" pageId="30" pageNumber="31">19</figureCitation>
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<figureCitation id="136E2BF55574FF98517DBF73FC3AFB0F" box="[845,896,1130,1156]" captionStart="Fig" captionStartId="46.[189,232,1822,1848]" captionTargetBox="[208,1379,488,1777]" captionTargetId="figure-18@46.[200,1387,478,1783]" captionTargetPageId="46" captionText="Fig. 23. Female genital ducts of Pimeliini. A. Apatopsis grombczewskii Semenov, 1891 (ZIN). B. Habrobates vernalis Semenov, 1903 (ZIN). C. Dietomorpha pardalis Kühnelt, 1957 (ZIN). D. Przewalskia dilatata (Reitter, 1887) (ZIN). E. Mantichorula semenowi Reitter, 1889 (ZIN). F. Platyope leucogramma (Pallas, 1773) (ZIN). G. Platyope grumi (Semenov, 1893), paralectotype (ZIN). H. Earophanta serrata (Semenov, 1893) (ZIN). Not to scale. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.6419493" httpUri="https://zenodo.org/record/6419493/files/figure.png" pageId="30" pageNumber="31">23E</figureCitation>
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<figureCitation id="136E2BF55574FF9851BBBF73FC05FB0F" box="[907,959,1130,1156]" captionStart="Fig" captionStartId="49.[188,231,1892,1918]" captionTargetBox="[191,1398,531,1851]" captionTargetId="figure-73@49.[189,1398,530,1855]" captionTargetPageId="49" captionText="Fig. 24. Apical antennomeres of different taxa of Pimeliini. A. Habrobates vernalis Semenov, 1903 ♂, paralectotype (ZIN). B. Przewalskia dilatata (Reitter, 1887), ♂. C. Mantichorula semenowi Reitter, 1889, ♂ (ZIN). D. Platyope leucogramma (Pallas, 1773), ♀. E. Earophanta serrata (Semenov, 1893), ♂ (ZIN)." figureDoi="http://doi.org/10.5281/zenodo.6419497" httpUri="https://zenodo.org/record/6419497/files/figure.png" pageId="30" pageNumber="31">24C</figureCitation>
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,
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<figureCitation id="136E2BF55574FF9851FBBF73FBBAFB0F" box="[971,1024,1130,1156]" captionStart="Fig" captionStartId="52.[189,232,1822,1848]" captionTargetBox="[290,1286,773,1776]" captionTargetId="figure-71@52.[281,1307,763,1783]" captionTargetPageId="52" captionText="Fig. 26. Protibiae of Pimeliini, optic microscopy. A. Habrochiton vernus Semenov-Tjan-Shansky, 1907, ♂, lectotype (ZIN). B. Habrobates gabrieli (Schuster, 1935) comb. nov., paralectotype (NMB). C. Dietomorpha pardalis Kühnelt, 1957, ♂ (ZIN). D. Mantichorula semenowi Reitter, 1889, ♂ (ZIN). E. Przewalskia dilatata (Reitter, 1887), ♂ (ZIN). F. Apatopsis grombczewskii Semenov, 1891, ♀, paralectotype (ZIN). G. Earophanta serrata (Semenov, 1893), ♂ (ZIN). H. Platyope leucogramma (Pallas, 1773), ♂ (ZIN). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.6419501" httpUri="https://zenodo.org/record/6419501/files/figure.png" pageId="30" pageNumber="31">26D</figureCitation>
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,
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<figureCitation id="136E2BF55574FF98563CBF73FBE4FB0F" box="[1036,1118,1130,1156]" captionStart="Fig" captionStartId="53.[189,232,1857,1883]" captionTargetBox="[189,1398,449,1817]" captionTargetId="figure-26@53.[189,1398,449,1818]" captionTargetPageId="53" captionText="Fig. 27. Protibiae of Pimeliini, SEM. A. Apatopsis grombczewskii Semenov, 1891, ♀ (ZIN), dorsal view. B. Ditto, apical part. C. Habrobates vernalis Semenov, 1903, ♂, paralectotype (ZIN), ventral view. D. Ditto, dorsal view. E. Habrochiton vernus Semenov-Tjan-Shansky, 1907, dorsal view (ZIN). F. Mantichorula semenowi Reitter, 1889, ♂ (ZIN), ventral view. G. Ditto, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.6419505" httpUri="https://zenodo.org/record/6419505/files/figure.png" pageId="30" pageNumber="31">27F–G</figureCitation>
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<treatmentCitation id="0AF411615574FF98528DBFB5FD85FB4C" author="Reitter E." box="[189,575,1196,1223]" page="695" pageId="30" pageNumber="31" year="1889">
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<emphasis id="B921EB625574FF98528DBFB5FEDAFB4D" box="[189,352,1196,1222]" italics="true" pageId="30" pageNumber="31">Mantichorula</emphasis>
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<bibRefCitation id="EFC44A815574FF985357BFB5FD85FB4C" author="Reitter E." box="[359,575,1196,1223]" pageId="30" pageNumber="31" pagination="678 - 710" refId="ref32704" refString="Reitter E. 1889. Insecta, a cl. G. N. Potanin in China et in Mongolia novissime lecta. XIII. Tenebrionidae. Horae Societatis Entomologicae Rossicae 23: 678 - 710." type="journal article" year="1889">Reitter, 1889: 695</bibRefCitation>
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.
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||
<taxonomicName id="4C554CF35574FF98528DBFF4FEDAFA8C" authorityName="Semenovi Rtt. Coll. Reitter" authorityYear="1889" box="[189,352,1261,1287]" class="Insecta" family="Tenebrionidae" genus="Mantichorula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B921EB625574FF98528DBFF4FEDAFA8C" box="[189,352,1261,1287]" italics="true" pageId="30" pageNumber="31">Mantichorula</emphasis>
|
||
</taxonomicName>
|
||
–
|
||
<treatmentCitation id="0AF411615574FF9853B7BFF4FDD9FA8C" author="Reitter E." box="[391,611,1261,1287]" page="246" pageId="30" pageNumber="31" year="1893">
|
||
<bibRefCitation id="EFC44A815574FF9853B7BFF4FDD9FA8C" author="Reitter E." box="[391,611,1261,1287]" pageId="30" pageNumber="31" pagination="201 - 250" refId="ref32741" refString="Reitter E. 1893. Bestimmungs-Tabelle der unechten Pimeliden aus der palaearctischen Fauna. Verhandlungen des Naturforschenden Vereines in Brunn 31 (1892): 201 - 250." type="journal article" year="1893">Reitter 1893: 246</bibRefCitation>
|
||
</treatmentCitation>
|
||
. —
|
||
<treatmentCitation id="0AF411615574FF9850ACBFF4FC2FFA8C" author="Semenov A. P." box="[668,917,1261,1287]" page="263" pageId="30" pageNumber="31" year="1893">
|
||
<bibRefCitation id="EFC44A815574FF9850ACBFF4FC2FFA8C" author="Semenov A. P." box="[668,917,1261,1287]" pageId="30" pageNumber="31" pagination="1892 - 1893" refId="ref33186" refString="Semenov A. P. 1893. Symbolae ad cognitionem pimeliidarum. I - III. Horae Societatis Entomologicae Rossicae 27 [1892 - 1893]: 249 - 264." type="journal article" year="1893">Semenov 1893: 263</bibRefCitation>
|
||
</treatmentCitation>
|
||
. —
|
||
<treatmentCitation id="0AF411615574FF9851FEBFF4FAC8FA8C" author="Semenov-Tjan-Shansky A. P." box="[974,1394,1261,1287]" page="179" pageId="30" pageNumber="31" year="1907">
|
||
<bibRefCitation id="EFC44A815574FF9851FEBFF4FAC8FA8C" author="Semenov-Tjan-Shansky A. P." box="[974,1394,1261,1287]" pageId="30" pageNumber="31" pagination="1907 - 1908" refId="ref33261" refString="Semenov-Tjan-Shansky A. P. 1907. Synopsis generum tribus Platyopinorum (Coleoptera, Tenebrionidae Pimeliini). Horae Societatis Entomologicae Rossicae 38 [1907 - 1908]: 175 - 184." type="journal article" year="1907">Semenov-Tjan-Shansky 1907: 179</bibRefCitation>
|
||
</treatmentCitation>
|
||
. —
|
||
<treatmentCitation id="0AF411615574FF985322BE09FE55FAA0" author="Schuster A." box="[274,495,1296,1323]" page="27" pageId="30" pageNumber="31" year="1935">
|
||
<bibRefCitation id="EFC44A815574FF985322BE09FE55FAA0" author="Schuster A." box="[274,495,1296,1323]" pageId="30" pageNumber="31" pagination="20 - 29" refId="ref33019" refString="Schuster A. 1935. Die Tenebrioniden-Ausbeute Dr. Gabriels in Persien. Koleopterologische Rundschau 21: 20 - 29." type="journal article" year="1935">Schuster 1935: 27</bibRefCitation>
|
||
</treatmentCitation>
|
||
. —
|
||
<treatmentCitation id="0AF411615574FF985015BE09FC9DFAA0" author="Medvedev G. S." box="[549,807,1296,1323]" page="125" pageId="30" pageNumber="31" year="1990">
|
||
<bibRefCitation id="EFC44A815574FF985015BE09FC9DFAA0" author="Medvedev G. S." box="[549,807,1296,1323]" pageId="30" pageNumber="31" refId="ref31976" refString="Medvedev G. S. 1990. Key to the darkling beetles of Mongolia. Proceedings of the Zoological Institute 220. Zoological Institute of the USSR, Leningrad. [In Russian.]" type="book" year="1990">Medvedev 1990: 125</bibRefCitation>
|
||
</treatmentCitation>
|
||
. —
|
||
<treatmentCitation id="0AF411615574FF98516DBE08FBF4FAA1" author="Ren G. - D. & Yu Y." box="[861,1102,1296,1323]" page="67" pageId="30" pageNumber="31" year="1999">
|
||
<bibRefCitation id="EFC44A815574FF98516DBE08FBF4FAA1" author="Ren G. - D. & Yu Y." box="[861,1102,1296,1323]" pageId="30" pageNumber="31" refId="ref32918" refString="Ren G. - D. & Yu Y. 1999. The Darkling Beetles from Deserts and Semideserts of China (Coleoptera: Tenebrionidae). Hebei University Publishing House, Baoding. [In Chinese.]" type="book" year="1999">Ren & Yu 1999: 67</bibRefCitation>
|
||
</treatmentCitation>
|
||
, 74, 76–78, figs 23, 30– 31. —
|
||
<treatmentCitation id="0AF411615574FF98530BBE2DFDE8FAC5" author="Ren G. - D. & Dong S." box="[315,594,1332,1358]" page="297" pageId="30" pageNumber="31" year="2001">
|
||
<bibRefCitation id="EFC44A815574FF98530BBE2DFDE8FAC5" author="Ren G. - D. & Dong S." box="[315,594,1332,1358]" pageId="30" pageNumber="31" pagination="296 - 302" refId="ref32868" refString="Ren G. - D. & Dong S. 2001. The beetles of the tribe Platyopini in China and description of two new species (Coleoptera: Tenebrionidae). Journal of Hebei University (Natural Science Edition) 21 (3): 296 - 302." type="journal article" year="2001">Ren & Dong 2001: 297</bibRefCitation>
|
||
</treatmentCitation>
|
||
.
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34F64FB5574FF98528DBE6CFC91FA35" pageId="30" pageNumber="31" type="type_taxon">
|
||
<paragraph id="8BEA37705574FF98528DBE6CFEE3FA04" blockId="30.[189,345,1397,1423]" box="[189,345,1397,1423]" pageId="30" pageNumber="31">
|
||
<heading id="D0A2801C5574FF98528DBE6CFEE3FA04" bold="true" box="[189,345,1397,1423]" fontSize="11" level="3" pageId="30" pageNumber="31" reason="3">
|
||
<emphasis id="B921EB625574FF98528DBE6CFEE3FA04" bold="true" box="[189,345,1397,1423]" pageId="30" pageNumber="31">
|
||
<typeStatus id="54EE89D25574FF98528DBE6CFF41FA04" box="[189,251,1397,1423]" pageId="30" pageNumber="31">Type</typeStatus>
|
||
species
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BEA37705574FF98528DBEBAFC91FA35" blockId="30.[189,811,1443,1470]" box="[189,811,1443,1470]" pageId="30" pageNumber="31">
|
||
<taxonomicName id="4C554CF35574FF98528DBEBAFDC6FA35" authority="Reitter, 1889" authorityName="Reitter" authorityYear="1889" box="[189,636,1443,1470]" class="Insecta" family="Tenebrionidae" genus="Mantichorula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="semenowi">
|
||
<emphasis id="B921EB625574FF98528DBEBAFE67FA36" box="[189,477,1443,1469]" italics="true" pageId="30" pageNumber="31">Mantichorula semenowi</emphasis>
|
||
<bibRefCitation id="EFC44A815574FF9853D4BEBAFDC6FA35" author="Reitter E." box="[484,636,1443,1470]" pageId="30" pageNumber="31" pagination="678 - 710" refId="ref32704" refString="Reitter E. 1889. Insecta, a cl. G. N. Potanin in China et in Mongolia novissime lecta. XIII. Tenebrionidae. Horae Societatis Entomologicae Rossicae 23: 678 - 710." type="journal article" year="1889">Reitter, 1889</bibRefCitation>
|
||
</taxonomicName>
|
||
, by monotypy.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34F64FB5574FF98528DBEFCFB0DF9A5" pageId="30" pageNumber="31" type="reference_group">
|
||
<paragraph id="8BEA37705574FF98528DBEFCFE34FA74" blockId="30.[189,398,1509,1535]" box="[189,398,1509,1535]" pageId="30" pageNumber="31">
|
||
<heading id="D0A2801C5574FF98528DBEFCFE34FA74" bold="true" box="[189,398,1509,1535]" fontSize="11" level="3" pageId="30" pageNumber="31" reason="3">
|
||
<emphasis id="B921EB625574FF98528DBEFCFE34FA74" bold="true" box="[189,398,1509,1535]" pageId="30" pageNumber="31">Species included</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BEA37705574FF98528DBD0DFB0DF9A5" blockId="30.[189,1207,1555,1582]" box="[189,1207,1555,1582]" pageId="30" pageNumber="31">
|
||
<taxonomicName id="4C554CF35574FF98528DBD0DFE46F9A5" authority="Reitter, 1889" authorityName="Reitter" authorityYear="1889" box="[189,508,1555,1582]" class="Insecta" family="Tenebrionidae" genus="Mantichorula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="semenowi">
|
||
<emphasis id="B921EB625574FF98528DBD0DFEE0F9A6" box="[189,346,1556,1581]" italics="true" pageId="30" pageNumber="31">M. semenowi</emphasis>
|
||
<bibRefCitation id="EFC44A815574FF985351BD0AFE46F9A5" author="Reitter E." box="[353,508,1555,1582]" pageId="30" pageNumber="31" pagination="678 - 710" refId="ref32704" refString="Reitter E. 1889. Insecta, a cl. G. N. Potanin in China et in Mongolia novissime lecta. XIII. Tenebrionidae. Horae Societatis Entomologicae Rossicae 23: 678 - 710." type="journal article" year="1889">Reitter, 1889</bibRefCitation>
|
||
</taxonomicName>
|
||
;
|
||
<emphasis id="B921EB625574FF985038BD0DFD93F9A6" box="[520,553,1556,1581]" italics="true" pageId="30" pageNumber="31">М.</emphasis>
|
||
<taxonomicName id="4C554CF35574FF985000BD0AFCF0F9A5" authority="Semenov, 1893" authorityName="Semenov" authorityYear="1893" box="[560,842,1555,1582]" class="Insecta" family="Tenebrionidae" genus="Mantichorula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="grandis">
|
||
<emphasis id="B921EB625574FF985000BD0AFD31F9A6" box="[560,651,1555,1581]" italics="true" pageId="30" pageNumber="31">grandis</emphasis>
|
||
<bibRefCitation id="EFC44A815574FF9850A2BD0DFCF0F9A5" author="Semenov A. P." box="[658,842,1556,1582]" pageId="30" pageNumber="31" pagination="1892 - 1893" refId="ref33186" refString="Semenov A. P. 1893. Symbolae ad cognitionem pimeliidarum. I - III. Horae Societatis Entomologicae Rossicae 27 [1892 - 1893]: 249 - 264." type="journal article" year="1893">Semenov, 1893</bibRefCitation>
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName id="4C554CF35574FF985166BD0DFB08F9A5" authority="Schuster, 1940" authorityName="Schuster" authorityYear="1940" box="[854,1202,1555,1582]" class="Insecta" family="Tenebrionidae" genus="Mantichorula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="mongolica">
|
||
<emphasis id="B921EB625574FF985166BD0DFC40F9A6" box="[854,1018,1555,1581]" italics="true" pageId="30" pageNumber="31">M. mongolica</emphasis>
|
||
Schuster, 1940
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34F64FB5574FF99528DBD4CFAE7F928" lastPageId="31" lastPageNumber="32" pageId="30" pageNumber="31" type="diagnosis">
|
||
<paragraph id="8BEA37705574FF98528DBD4CFE8DF9E4" blockId="30.[189,311,1621,1647]" box="[189,311,1621,1647]" pageId="30" pageNumber="31">
|
||
<heading id="D0A2801C5574FF98528DBD4CFE8DF9E4" bold="true" box="[189,311,1621,1647]" fontSize="11" level="3" pageId="30" pageNumber="31" reason="3">
|
||
<emphasis id="B921EB625574FF98528DBD4CFE8DF9E4" bold="true" box="[189,311,1621,1647]" pageId="30" pageNumber="31">Diagnosis</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BEA37705574FF98528DBD9AFCE0F88C" blockId="30.[189,1399,1667,1800]" pageId="30" pageNumber="31">
|
||
Body black, wide, dorsoventrally flattened, disc-shaped, without dense pubescence dorsally, only some parts of pronotum and head with setae, elytra dorsally and laterally with very short, fine, sparse spines (
|
||
<figureCitation id="136E2BF55574FF9852F6BDD3FEFCF96F" box="[198,326,1738,1764]" captionStart="Fig" captionStartId="9.[189,232,1892,1918]" captionTargetBox="[233,1353,649,1847]" captionTargetId="figure-90@9.[227,1361,643,1855]" captionTargetPageId="9" captionText="Fig. 4. Species of the genera Przewalskia Semenov, 1893, Mantichorula Reitter, 1889 and Platyope Fischer von Waldheim, 1820, habitus. A. Przewalskia dilatata (Reitter, 1887), ♀ (ZIN), dorsal view. B. Ditto, ventral view. C. M. semenowi Reitter, 1889, ♂ (ZIN), dorsal view. D. Ditto, ventral view. E. Platyope leucogramma (Pallas, 1773), ♂ (ZIN), dorsal view. F. Ditto, ventral view." figureDoi="http://doi.org/10.5281/zenodo.6419435" httpUri="https://zenodo.org/record/6419435/files/figure.png" pageId="30" pageNumber="31">Fig. 4C–D</figureCitation>
|
||
) and sometimes with small area of setation along apical part of suture. Body length
|
||
<quantity id="4CAD9A955574FF98577ABDD3FEA9F883" metricMagnitude="-2" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="30" pageNumber="31" unit="mm" value="15.0" valueMax="20.0" valueMin="10.0">10– 20 mm</quantity>
|
||
. Eyes large, circular, moderately convex dorsally.
|
||
</paragraph>
|
||
<paragraph id="8BEA37705574FF98528DBC36FACCF85E" blockId="30.[187,1399,1838,2006]" pageId="30" pageNumber="31">
|
||
Pronotum strongly transverse (more than 2 × as wide as long). Anterolateral angles weakly projecting. Base of pronotum strongly widely emarginated, and posterolateral angles deeply protruding and impressed into elytral base (
|
||
<figureCitation id="136E2BF55574FF9853CEBC6CFDE6F804" box="[510,604,1909,1935]" captionStart="Fig" captionStartId="9.[189,232,1892,1918]" captionTargetBox="[233,1353,649,1847]" captionTargetId="figure-90@9.[227,1361,643,1855]" captionTargetPageId="9" captionText="Fig. 4. Species of the genera Przewalskia Semenov, 1893, Mantichorula Reitter, 1889 and Platyope Fischer von Waldheim, 1820, habitus. A. Przewalskia dilatata (Reitter, 1887), ♀ (ZIN), dorsal view. B. Ditto, ventral view. C. M. semenowi Reitter, 1889, ♂ (ZIN), dorsal view. D. Ditto, ventral view. E. Platyope leucogramma (Pallas, 1773), ♂ (ZIN), dorsal view. F. Ditto, ventral view." figureDoi="http://doi.org/10.5281/zenodo.6419435" httpUri="https://zenodo.org/record/6419435/files/figure.png" pageId="30" pageNumber="31">Figs 4C</figureCitation>
|
||
,
|
||
<figureCitation id="136E2BF55574FF985056BC6CFD22F804" box="[614,664,1909,1935]" captionStart="Fig" captionStartId="20.[189,232,1892,1918]" captionTargetBox="[189,1398,431,1851]" captionTargetId="figure-17@20.[189,1398,431,1853]" captionTargetPageId="20" captionText="Fig. 11. Przewalskia Semenov, 1893 and Mantichorula Reitter, 1889, details of structure. A. P. dilatata (Reitter, 1887), ♂ (ZIN), pronotum. B. Ditto, hemispherical large tubercles on pronotum. C. Ditto, base of left elytron. D. Ditto, pubescence of elytron. E. M. semenowi Reitter, 1889, ♂ (ZIN), pronotum. F. Ditto, tubercles and setation on sides of pronotum." figureDoi="http://doi.org/10.5281/zenodo.6419457" httpUri="https://zenodo.org/record/6419457/files/figure.png" pageId="30" pageNumber="31">11E</figureCitation>
|
||
). Disc of pronotum with one medial and two lateral longitudinal stripes of recumbent, moderately dense, short setae and coarse, semispherical tubercles on lateral sides (
|
||
<figureCitation id="136E2BF55574FF9852F6BCA2FEF7F85E" box="[198,333,1979,2005]" captionStart="Fig" captionStartId="20.[189,232,1892,1918]" captionTargetBox="[189,1398,431,1851]" captionTargetId="figure-17@20.[189,1398,431,1853]" captionTargetPageId="20" captionText="Fig. 11. Przewalskia Semenov, 1893 and Mantichorula Reitter, 1889, details of structure. A. P. dilatata (Reitter, 1887), ♂ (ZIN), pronotum. B. Ditto, hemispherical large tubercles on pronotum. C. Ditto, base of left elytron. D. Ditto, pubescence of elytron. E. M. semenowi Reitter, 1889, ♂ (ZIN), pronotum. F. Ditto, tubercles and setation on sides of pronotum." figureDoi="http://doi.org/10.5281/zenodo.6419457" httpUri="https://zenodo.org/record/6419457/files/figure.png" pageId="30" pageNumber="31">Fig. 11E–F</figureCitation>
|
||
); base widely, sharply depressed and flattened. Prosternum before procoxae near 2× shorter
|
||
</paragraph>
|
||
<paragraph id="8BEA37705575FF99528DBA10FB1CFECD" blockId="31.[189,1399,265,327]" pageId="31" pageNumber="32">
|
||
than longitudinal diameter of one procoxa. Prosternal process very large and broad, raised between procoxae, strongly protruding beyond procoxae, extending to mesoventrite (
|
||
<figureCitation id="136E2BF55575FF99560ABA35FB20FECD" box="[1082,1178,300,326]" captionStart="Fig" captionStartId="9.[189,232,1892,1918]" captionTargetBox="[233,1353,649,1847]" captionTargetId="figure-90@9.[227,1361,643,1855]" captionTargetPageId="9" captionText="Fig. 4. Species of the genera Przewalskia Semenov, 1893, Mantichorula Reitter, 1889 and Platyope Fischer von Waldheim, 1820, habitus. A. Przewalskia dilatata (Reitter, 1887), ♀ (ZIN), dorsal view. B. Ditto, ventral view. C. M. semenowi Reitter, 1889, ♂ (ZIN), dorsal view. D. Ditto, ventral view. E. Platyope leucogramma (Pallas, 1773), ♂ (ZIN), dorsal view. F. Ditto, ventral view." figureDoi="http://doi.org/10.5281/zenodo.6419435" httpUri="https://zenodo.org/record/6419435/files/figure.png" pageId="31" pageNumber="32">Fig. 4D</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BEA37705575FF99528DBA77FC86FE7A" blockId="31.[187,1399,366,498]" pageId="31" pageNumber="32">
|
||
Scutellar shield not concealed by base of pronotum, triangular. Elytra short, oval, with (
|
||
<figureCitation id="136E2BF55575FF9956D3BA77FAF1FE03" box="[1251,1355,366,392]" captionStart="Fig" captionStartId="20.[189,232,1892,1918]" captionTargetBox="[189,1398,431,1851]" captionTargetId="figure-17@20.[189,1398,431,1853]" captionTargetPageId="20" captionText="Fig. 11. Przewalskia Semenov, 1893 and Mantichorula Reitter, 1889, details of structure. A. P. dilatata (Reitter, 1887), ♂ (ZIN), pronotum. B. Ditto, hemispherical large tubercles on pronotum. C. Ditto, base of left elytron. D. Ditto, pubescence of elytron. E. M. semenowi Reitter, 1889, ♂ (ZIN), pronotum. F. Ditto, tubercles and setation on sides of pronotum." figureDoi="http://doi.org/10.5281/zenodo.6419457" httpUri="https://zenodo.org/record/6419457/files/figure.png" pageId="31" pageNumber="32">Fig. 11E</figureCitation>
|
||
) or without depressions around scutellar shield, apical part abducted. Elytra with double humeral rib. Ventral side of body covered with short, dense setae. Metaventrite much shorter, than mesoventrite. Intercoxal process 1.2–1.24 × as wide as one metacoxa (
|
||
<figureCitation id="136E2BF55575FF9950E1BACEFC8AFE7A" box="[721,816,471,497]" captionStart="Fig" captionStartId="9.[189,232,1892,1918]" captionTargetBox="[233,1353,649,1847]" captionTargetId="figure-90@9.[227,1361,643,1855]" captionTargetPageId="9" captionText="Fig. 4. Species of the genera Przewalskia Semenov, 1893, Mantichorula Reitter, 1889 and Platyope Fischer von Waldheim, 1820, habitus. A. Przewalskia dilatata (Reitter, 1887), ♀ (ZIN), dorsal view. B. Ditto, ventral view. C. M. semenowi Reitter, 1889, ♂ (ZIN), dorsal view. D. Ditto, ventral view. E. Platyope leucogramma (Pallas, 1773), ♂ (ZIN), dorsal view. F. Ditto, ventral view." figureDoi="http://doi.org/10.5281/zenodo.6419435" httpUri="https://zenodo.org/record/6419435/files/figure.png" pageId="31" pageNumber="32">Fig. 4D</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BEA37705575FF99528DB900FB2DFC18" blockId="31.[187,1399,537,915]" pageId="31" pageNumber="32">
|
||
Protrochanters and inner side of profemora covered with very long, dense setae. Femora robust, weakly curved outward. All tibiae weakly curved outward. Protibiae strongly gradually widened to apex, triangular, flattened, without projecting process at apex of outer margin, with dense, short, strong spines, but with longer and finer spines and dense, long setae along lateral margins (more recumbent on inner side) (
|
||
<figureCitation id="136E2BF55575FF995334B9BCFEC8FD34" box="[260,370,677,703]" captionStart="Fig" captionStartId="52.[189,232,1822,1848]" captionTargetBox="[290,1286,773,1776]" captionTargetId="figure-71@52.[281,1307,763,1783]" captionTargetPageId="52" captionText="Fig. 26. Protibiae of Pimeliini, optic microscopy. A. Habrochiton vernus Semenov-Tjan-Shansky, 1907, ♂, lectotype (ZIN). B. Habrobates gabrieli (Schuster, 1935) comb. nov., paralectotype (NMB). C. Dietomorpha pardalis Kühnelt, 1957, ♂ (ZIN). D. Mantichorula semenowi Reitter, 1889, ♂ (ZIN). E. Przewalskia dilatata (Reitter, 1887), ♂ (ZIN). F. Apatopsis grombczewskii Semenov, 1891, ♀, paralectotype (ZIN). G. Earophanta serrata (Semenov, 1893), ♂ (ZIN). H. Platyope leucogramma (Pallas, 1773), ♂ (ZIN). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.6419501" httpUri="https://zenodo.org/record/6419501/files/figure.png" pageId="31" pageNumber="32">Figs 26D</figureCitation>
|
||
,
|
||
<figureCitation id="136E2BF55575FF99534DB9BFFE69FD4B" box="[381,467,678,704]" captionStart="Fig" captionStartId="53.[189,232,1857,1883]" captionTargetBox="[189,1398,449,1817]" captionTargetId="figure-26@53.[189,1398,449,1818]" captionTargetPageId="53" captionText="Fig. 27. Protibiae of Pimeliini, SEM. A. Apatopsis grombczewskii Semenov, 1891, ♀ (ZIN), dorsal view. B. Ditto, apical part. C. Habrobates vernalis Semenov, 1903, ♂, paralectotype (ZIN), ventral view. D. Ditto, dorsal view. E. Habrochiton vernus Semenov-Tjan-Shansky, 1907, dorsal view (ZIN). F. Mantichorula semenowi Reitter, 1889, ♂ (ZIN), ventral view. G. Ditto, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.6419505" httpUri="https://zenodo.org/record/6419505/files/figure.png" pageId="31" pageNumber="32">27F–G</figureCitation>
|
||
); strong spines denser at apex of outer margin. Meso- and metatibiae with similar structures, but not dorsoventrally flattened and widened, often additionally covered with cream coloured scales. Protibial terminal spurs large and widened, not lanceolate (
|
||
<figureCitation id="136E2BF55575FF9951D8B9F5FBC2FC8D" box="[1000,1144,748,774]" captionStart="Fig" captionStartId="53.[189,232,1857,1883]" captionTargetBox="[189,1398,449,1817]" captionTargetId="figure-26@53.[189,1398,449,1818]" captionTargetPageId="53" captionText="Fig. 27. Protibiae of Pimeliini, SEM. A. Apatopsis grombczewskii Semenov, 1891, ♀ (ZIN), dorsal view. B. Ditto, apical part. C. Habrobates vernalis Semenov, 1903, ♂, paralectotype (ZIN), ventral view. D. Ditto, dorsal view. E. Habrochiton vernus Semenov-Tjan-Shansky, 1907, dorsal view (ZIN). F. Mantichorula semenowi Reitter, 1889, ♂ (ZIN), ventral view. G. Ditto, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.6419505" httpUri="https://zenodo.org/record/6419505/files/figure.png" pageId="31" pageNumber="32">Fig. 27F–G</figureCitation>
|
||
), subequal in length, extending to base of protarsomere 5. Meso- and metatibial terminal spurs different in length, inner tibial terminal spurs longer than outer ones, extending to protarsomere 3 and half length of metatarsomere 1 subsequently. Protarsi with long, recumbent spines dorsally and ventrally and short setae laterally; meso- and metatarsi flattened from sides, with very dense, long setae on dorsal side.
|
||
</paragraph>
|
||
<paragraph id="8BEA37705575FF9952DAB8A3FACCFC7C" blockId="31.[189,1400,954,1121]" pageId="31" pageNumber="32">
|
||
<emphasis id="B921EB625575FF9952DAB8A3FE25FC5F" bold="true" box="[234,415,954,980]" pageId="31" pageNumber="32">Male genitalia</emphasis>
|
||
(
|
||
<figureCitation id="136E2BF55575FF99539FB8A3FD84FC5E" box="[431,574,954,981]" captionStart="Fig" captionStartId="40.[191,234,1857,1883]" captionTargetBox="[189,1398,539,1819]" captionTargetId="figure-91@40.[189,1398,529,1819]" captionTargetPageId="40" captionText="Fig. 19. Mantichorula semenowi Reitter, 1889, genitalia (ZIN). A. Tegmen of aedeagus, ventrally. B. Ditto, dorsally. C. Ditto, laterally. D. Median lobe, ventrally. E. Ditto, dorsally. F. Ditto, laterally. G. Spiculum gastrale. H. ♂, inner sternite VIII. I. ♀, inner sternite VIII and spiculum ventrale. J. Ovipositor, ventrally. K. Ditto, dorsally. L. Ditto, laterally. Not to scale. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.6419481" httpUri="https://zenodo.org/record/6419481/files/figure.png" pageId="31" pageNumber="32">Fig. 19A, H</figureCitation>
|
||
) Inner sternite VIII (
|
||
<figureCitation id="136E2BF55575FF995398B8C4FDAFFC7C" box="[424,533,989,1015]" captionStart="Fig" captionStartId="40.[191,234,1857,1883]" captionTargetBox="[189,1398,539,1819]" captionTargetId="figure-91@40.[189,1398,529,1819]" captionTargetPageId="40" captionText="Fig. 19. Mantichorula semenowi Reitter, 1889, genitalia (ZIN). A. Tegmen of aedeagus, ventrally. B. Ditto, dorsally. C. Ditto, laterally. D. Median lobe, ventrally. E. Ditto, dorsally. F. Ditto, laterally. G. Spiculum gastrale. H. ♂, inner sternite VIII. I. ♀, inner sternite VIII and spiculum ventrale. J. Ovipositor, ventrally. K. Ditto, dorsally. L. Ditto, laterally. Not to scale. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.6419481" httpUri="https://zenodo.org/record/6419481/files/figure.png" pageId="31" pageNumber="32">Fig. 19H</figureCitation>
|
||
) sclerotized on margins and membranous in middle, densely pubescent;
|
||
</paragraph>
|
||
<paragraph id="8BEA37705575FF99528FBF19FE70FBEA" blockId="31.[189,1400,954,1121]" pageId="31" pageNumber="32">
|
||
posterior margin deeply emarginated medially; gland absent (at least, not observed). Spiculum gastrale (
|
||
<figureCitation id="136E2BF55575FF9952F6BF3DFE82FBB5" box="[198,312,1060,1086]" captionStart="Fig" captionStartId="40.[191,234,1857,1883]" captionTargetBox="[189,1398,539,1819]" captionTargetId="figure-91@40.[189,1398,529,1819]" captionTargetPageId="40" captionText="Fig. 19. Mantichorula semenowi Reitter, 1889, genitalia (ZIN). A. Tegmen of aedeagus, ventrally. B. Ditto, dorsally. C. Ditto, laterally. D. Median lobe, ventrally. E. Ditto, dorsally. F. Ditto, laterally. G. Spiculum gastrale. H. ♂, inner sternite VIII. I. ♀, inner sternite VIII and spiculum ventrale. J. Ovipositor, ventrally. K. Ditto, dorsally. L. Ditto, laterally. Not to scale. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.6419481" httpUri="https://zenodo.org/record/6419481/files/figure.png" pageId="31" pageNumber="32">Fig. 19G</figureCitation>
|
||
) with roundly connected rods and rounded derivatives of inner sternite IX with short pubescence terminally.
|
||
</paragraph>
|
||
<paragraph id="8BEA37705575FF99528DBF90FD2FFAA4" blockId="31.[187,1399,1160,1328]" pageId="31" pageNumber="32">
|
||
Tegmen of aedeagus (
|
||
<figureCitation id="136E2BF55575FF99538DBF91FDF0FB29" box="[445,586,1160,1186]" captionStart="Fig" captionStartId="40.[191,234,1857,1883]" captionTargetBox="[189,1398,539,1819]" captionTargetId="figure-91@40.[189,1398,529,1819]" captionTargetPageId="40" captionText="Fig. 19. Mantichorula semenowi Reitter, 1889, genitalia (ZIN). A. Tegmen of aedeagus, ventrally. B. Ditto, dorsally. C. Ditto, laterally. D. Median lobe, ventrally. E. Ditto, dorsally. F. Ditto, laterally. G. Spiculum gastrale. H. ♂, inner sternite VIII. I. ♀, inner sternite VIII and spiculum ventrale. J. Ovipositor, ventrally. K. Ditto, dorsally. L. Ditto, laterally. Not to scale. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.6419481" httpUri="https://zenodo.org/record/6419481/files/figure.png" pageId="31" pageNumber="32">Fig. 19A–C</figureCitation>
|
||
) elongate, slender, with subequal basal and apical pieces. Basal piece weakly wider than apical piece, both fusiform.Apical piece glabrous, weakly curved, ventral aphophyses not expressed, dorsal apophyses short, merged in triangular plate. Basal piece with sub-acutely angulate apex. Median lobe acutely angulate apically, with narrow baculi and wide, membranous area at middle, slightly bifurcated at base (
|
||
<figureCitation id="136E2BF55575FF9953CBBE0CFD32FAA4" box="[507,648,1301,1327]" captionStart="Fig" captionStartId="40.[191,234,1857,1883]" captionTargetBox="[189,1398,539,1819]" captionTargetId="figure-91@40.[189,1398,529,1819]" captionTargetPageId="40" captionText="Fig. 19. Mantichorula semenowi Reitter, 1889, genitalia (ZIN). A. Tegmen of aedeagus, ventrally. B. Ditto, dorsally. C. Ditto, laterally. D. Median lobe, ventrally. E. Ditto, dorsally. F. Ditto, laterally. G. Spiculum gastrale. H. ♂, inner sternite VIII. I. ♀, inner sternite VIII and spiculum ventrale. J. Ovipositor, ventrally. K. Ditto, dorsally. L. Ditto, laterally. Not to scale. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.6419481" httpUri="https://zenodo.org/record/6419481/files/figure.png" pageId="31" pageNumber="32">Fig. 19D–F</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BEA37705575FF9952DABE4EFE01FAFB" blockId="31.[187,1398,1366,1463]" box="[234,443,1366,1393]" pageId="31" pageNumber="32">
|
||
<emphasis id="B921EB625575FF9952DABE4EFE01FAFB" bold="true" box="[234,443,1366,1393]" pageId="31" pageNumber="32">Female genitalia</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BEA37705575FF99528DBE63FD76FA3C" blockId="31.[187,1398,1366,1463]" pageId="31" pageNumber="32">
|
||
Spiculum ventrale with elongate common stem and narrow rods (
|
||
<figureCitation id="136E2BF55575FF99519DBE63FBB6FA1F" box="[941,1036,1402,1428]" captionStart="Fig" captionStartId="40.[191,234,1857,1883]" captionTargetBox="[189,1398,539,1819]" captionTargetId="figure-91@40.[189,1398,529,1819]" captionTargetPageId="40" captionText="Fig. 19. Mantichorula semenowi Reitter, 1889, genitalia (ZIN). A. Tegmen of aedeagus, ventrally. B. Ditto, dorsally. C. Ditto, laterally. D. Median lobe, ventrally. E. Ditto, dorsally. F. Ditto, laterally. G. Spiculum gastrale. H. ♂, inner sternite VIII. I. ♀, inner sternite VIII and spiculum ventrale. J. Ovipositor, ventrally. K. Ditto, dorsally. L. Ditto, laterally. Not to scale. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.6419481" httpUri="https://zenodo.org/record/6419481/files/figure.png" pageId="31" pageNumber="32">Fig. 19I</figureCitation>
|
||
). Sternite VIII narrow, strongly transverse, with moderately short apophyses.
|
||
</paragraph>
|
||
<paragraph id="8BEA37705575FF99528DBEC7FBCDF990" blockId="31.[189,1397,1502,1564]" pageId="31" pageNumber="32">
|
||
Ovipositor (
|
||
<figureCitation id="136E2BF55575FF99537ABEC7FE70FA73" box="[330,458,1502,1528]" captionStart="Fig" captionStartId="40.[191,234,1857,1883]" captionTargetBox="[189,1398,539,1819]" captionTargetId="figure-91@40.[189,1398,529,1819]" captionTargetPageId="40" captionText="Fig. 19. Mantichorula semenowi Reitter, 1889, genitalia (ZIN). A. Tegmen of aedeagus, ventrally. B. Ditto, dorsally. C. Ditto, laterally. D. Median lobe, ventrally. E. Ditto, dorsally. F. Ditto, laterally. G. Spiculum gastrale. H. ♂, inner sternite VIII. I. ♀, inner sternite VIII and spiculum ventrale. J. Ovipositor, ventrally. K. Ditto, dorsally. L. Ditto, laterally. Not to scale. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.6419481" httpUri="https://zenodo.org/record/6419481/files/figure.png" pageId="31" pageNumber="32">Fig. 19J–L</figureCitation>
|
||
) is very similar to that in
|
||
<taxonomicName id="4C554CF35575FF9950C1BEC7FC38FA73" authorityName="Semenov" authorityYear="1893" box="[753,898,1502,1528]" class="Magnoliopsida" family="Solanaceae" genus="Przewalskia" kingdom="Plantae" order="Solanales" pageId="31" pageNumber="32" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="B921EB625575FF9950C1BEC7FC38FA73" box="[753,898,1502,1528]" italics="true" pageId="31" pageNumber="32">Przewalskia</emphasis>
|
||
</taxonomicName>
|
||
, but differs in the absence of sclerotization on vulva, apical lobes turned to dorsal side and widely weakly rounded proctiger.
|
||
</paragraph>
|
||
<paragraph id="8BEA37705575FF9952DABD5AFDD4F9D6" blockId="31.[189,1399,1603,1700]" box="[234,622,1603,1629]" pageId="31" pageNumber="32">
|
||
<emphasis id="B921EB625575FF9952DABD5AFE54F9D6" bold="true" box="[234,494,1603,1629]" pageId="31" pageNumber="32">Female genital ducts</emphasis>
|
||
(
|
||
<figureCitation id="136E2BF55575FF9953CFBD5AFDDCF9D6" box="[511,614,1603,1629]" captionStart="Fig" captionStartId="46.[189,232,1822,1848]" captionTargetBox="[208,1379,488,1777]" captionTargetId="figure-18@46.[200,1387,478,1783]" captionTargetPageId="46" captionText="Fig. 23. Female genital ducts of Pimeliini. A. Apatopsis grombczewskii Semenov, 1891 (ZIN). B. Habrobates vernalis Semenov, 1903 (ZIN). C. Dietomorpha pardalis Kühnelt, 1957 (ZIN). D. Przewalskia dilatata (Reitter, 1887) (ZIN). E. Mantichorula semenowi Reitter, 1889 (ZIN). F. Platyope leucogramma (Pallas, 1773) (ZIN). G. Platyope grumi (Semenov, 1893), paralectotype (ZIN). H. Earophanta serrata (Semenov, 1893) (ZIN). Not to scale. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.6419493" httpUri="https://zenodo.org/record/6419493/files/figure.png" pageId="31" pageNumber="32">Fig. 23E</figureCitation>
|
||
)
|
||
</paragraph>
|
||
<paragraph id="8BEA37705575FF99528DBD7FFAE7F928" blockId="31.[189,1399,1603,1700]" pageId="31" pageNumber="32">Vagina elongated, tube-shaped, not widened, with short primary bursa copulatrix at apex. Spermatheca absent. Accessory gland of spermatheca very short, tube-shaped, not modified, without constrictions.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34F64FB5575FF99528DBDD2FC47F89F" box="[189,1021,1739,1812]" pageId="31" pageNumber="32" type="distribution">
|
||
<paragraph id="8BEA37705575FF99528DBDD2FEE2F96E" blockId="31.[189,344,1739,1765]" box="[189,344,1739,1765]" pageId="31" pageNumber="32">
|
||
<heading id="D0A2801C5575FF99528DBDD2FEE2F96E" bold="true" box="[189,344,1739,1765]" fontSize="11" level="3" pageId="31" pageNumber="32" reason="3">
|
||
<emphasis id="B921EB625575FF99528DBDD2FEE2F96E" bold="true" box="[189,344,1739,1765]" pageId="31" pageNumber="32">Distribution</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BEA37705575FF99528DBDE0FC47F89F" blockId="31.[189,1021,1785,1812]" box="[189,1021,1785,1812]" pageId="31" pageNumber="32">
|
||
<collectingCountry id="F34277E05575FF99528DBDE0FE88F898" box="[189,306,1785,1811]" name="Mongolia" pageId="31" pageNumber="32">Mongolia</collectingCountry>
|
||
,
|
||
<collectingCountry id="F34277E05575FF99530EBDE0FE3EF898" box="[318,388,1785,1811]" name="China" pageId="31" pageNumber="32">China</collectingCountry>
|
||
(
|
||
<collectingRegion id="4991F9925575FF9953A3BDE3FDF5F898" box="[403,591,1785,1812]" country="China" name="Nei Mongol" pageId="31" pageNumber="32">Inner Mongolia</collectingRegion>
|
||
,
|
||
<collectingRegion id="4991F9925575FF99506BBDE0FD07F898" box="[603,701,1785,1811]" country="China" name="Ningxia" pageId="31" pageNumber="32">Ningxia</collectingRegion>
|
||
,
|
||
<collectingRegion id="4991F9925575FF9950F8BDE3FCACF89F" box="[712,790,1786,1812]" country="China" name="Gansu" pageId="31" pageNumber="32">Gansu</collectingRegion>
|
||
,
|
||
<collectingRegion id="4991F9925575FF995112BDE0FCC9F898" box="[802,883,1785,1811]" country="China" name="Shanxi" pageId="31" pageNumber="32">Shanxi</collectingRegion>
|
||
provinces).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34F64FB5575FFA6528DBC22FC32FE06" lastPageId="32" lastPageNumber="33" pageId="31" pageNumber="32" type="discussion">
|
||
<paragraph id="8BEA37705575FF99528DBC22FEBEF8DE" blockId="31.[189,260,1851,1877]" box="[189,260,1851,1877]" pageId="31" pageNumber="32">
|
||
<heading id="D0A2801C5575FF99528DBC22FEBEF8DE" bold="true" box="[189,260,1851,1877]" fontSize="11" level="3" pageId="31" pageNumber="32" reason="3">
|
||
<emphasis id="B921EB625575FF99528DBC22FEBEF8DE" bold="true" box="[189,260,1851,1877]" pageId="31" pageNumber="32">Notes</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BEA37705575FFA6528DBC70FC32FE06" blockId="31.[189,1398,1897,1994]" lastBlockId="32.[187,1399,265,397]" lastPageId="32" lastPageNumber="33" pageId="31" pageNumber="32">
|
||
Schuster (1940) believed that all three species of
|
||
<taxonomicName id="4C554CF35575FF995122BC70FC0FF808" authorityName="Semenovi Rtt. Coll. Reitter" authorityYear="1889" box="[786,949,1897,1923]" class="Insecta" family="Tenebrionidae" genus="Mantichorula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B921EB625575FF995122BC70FC0FF808" box="[786,949,1897,1923]" italics="true" pageId="31" pageNumber="32">Mantichorula</emphasis>
|
||
</taxonomicName>
|
||
should be regarded as valid, and that
|
||
<taxonomicName id="4C554CF35575FF99528FBC94FEFFF82D" authorityName="Semenov" authorityYear="1893" box="[191,325,1932,1958]" class="Insecta" family="Tenebrionidae" genus="Mantichorula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="grandis">
|
||
<emphasis id="B921EB625575FF99528FBC94FEFFF82D" box="[191,325,1932,1958]" italics="true" pageId="31" pageNumber="32">M. grandis</emphasis>
|
||
</taxonomicName>
|
||
distinctly differs from
|
||
<taxonomicName id="4C554CF35575FF995051BC94FCBBF82D" authorityName="Reitter" authorityYear="1889" box="[609,769,1933,1958]" class="Insecta" family="Tenebrionidae" genus="Mantichorula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="semenowi">
|
||
<emphasis id="B921EB625575FF995051BC94FCBBF82D" box="[609,769,1933,1958]" italics="true" pageId="31" pageNumber="32">M. semenowi</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C554CF35575FF995172BC94FC50F82D" authorityName="Schuster" authorityYear="1940" box="[834,1002,1932,1958]" class="Insecta" family="Tenebrionidae" genus="Mantichorula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="mongolica">
|
||
<emphasis id="B921EB625575FF995172BC94FC50F82D" box="[834,1002,1932,1958]" italics="true" pageId="31" pageNumber="32">M. mongolica</emphasis>
|
||
</taxonomicName>
|
||
because of their short antennae, which do not extend far beyond the base of the pronotum.
|
||
<bibRefCitation id="EFC44A815575FF9951A7BCA9FBD6F841" author="Medvedev G. S." box="[919,1132,1968,1994]" pageId="31" pageNumber="32" refId="ref31976" refString="Medvedev G. S. 1990. Key to the darkling beetles of Mongolia. Proceedings of the Zoological Institute 220. Zoological Institute of the USSR, Leningrad. [In Russian.]" type="book" year="1990">Medvedev (1990)</bibRefCitation>
|
||
interpreted the genus
|
||
<taxonomicName id="4C554CF3554AFFA6528DBA10FEDAFEA8" authorityName="Semenovi Rtt. Coll. Reitter" authorityYear="1889" box="[189,352,265,291]" class="Insecta" family="Tenebrionidae" genus="Mantichorula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B921EB62554AFFA6528DBA10FEDAFEA8" box="[189,352,265,291]" italics="true" pageId="32" pageNumber="33">Mantichorula</emphasis>
|
||
</taxonomicName>
|
||
as monotypic. We studied the
|
||
<typeStatus id="54EE89D2554AFFA6508BBA13FD54FEAF" box="[699,750,266,292]" pageId="32" pageNumber="33">type</typeStatus>
|
||
series of
|
||
<taxonomicName id="4C554CF3554AFFA65166BA13FC63FEA8" authorityName="Semenov" authorityYear="1893" box="[854,985,265,291]" class="Insecta" family="Tenebrionidae" genus="Mantichorula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="grandis">
|
||
<emphasis id="B921EB62554AFFA65166BA13FC63FEA8" box="[854,985,265,291]" italics="true" pageId="32" pageNumber="33">M. grandis</emphasis>
|
||
</taxonomicName>
|
||
and discovered that this species and some specimens standing as
|
||
<taxonomicName id="4C554CF3554AFFA65023BA34FD0AFECD" authorityName="Reitter" authorityYear="1889" box="[531,688,301,326]" class="Insecta" family="Tenebrionidae" genus="Mantichorula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="semenowi">
|
||
<emphasis id="B921EB62554AFFA65023BA34FD0AFECD" box="[531,688,301,326]" italics="true" pageId="32" pageNumber="33">M. semenowi</emphasis>
|
||
</taxonomicName>
|
||
in the ZIN collection do indeed have shorter antennomeres. Studies of the
|
||
<typeStatus id="54EE89D2554AFFA65358BA49FE21FEE1" box="[360,411,336,362]" pageId="32" pageNumber="33">type</typeStatus>
|
||
specimens of
|
||
<taxonomicName id="4C554CF3554AFFA65075BA49FD58FEE2" authorityName="Reitter" authorityYear="1889" box="[581,738,336,361]" class="Insecta" family="Tenebrionidae" genus="Mantichorula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="semenowi">
|
||
<emphasis id="B921EB62554AFFA65075BA49FD58FEE2" box="[581,738,336,361]" italics="true" pageId="32" pageNumber="33">M. semenowi</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C554CF3554AFFA6512CBA49FC7BFEE2" authorityName="Schuster" authorityYear="1940" box="[796,961,335,361]" class="Insecta" family="Tenebrionidae" genus="Mantichorula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="mongolica">
|
||
<emphasis id="B921EB62554AFFA6512CBA49FC7BFEE2" box="[796,961,335,361]" italics="true" pageId="32" pageNumber="33">M. mongolica</emphasis>
|
||
</taxonomicName>
|
||
, as well as a detailed examination of male and female genitalia, are necessary to revise this genus.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |