treatments-xml/data/03/FC/1A/03FC1A09FFC9F770FF024B410DA9F9A0.xml
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<document id="9F9EDDCE8E9A1392E6BFF2DB434D6495" ID-DOI="10.5281/zenodo.5352440" ID-ISSN="1942-1354" ID-Zenodo-Dep="5352440" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630208260396" checkinUser="felipe" docAuthor="Alvarez, Héctor Jaime Gasca, da, Claudio Ruy Vasconcelos, Fonseca &amp; Ratcliffe, Brett C." docDate="2008" docId="03FC1A09FFC9F770FF024B410DA9F9A0" docLanguage="en" docName="0061.pdf" docOrigin="Insecta Mundi 2008 (61)" docStyle="DocumentStyle:A87B5EC635B7D5A430FC1D805C706BD1.3:InsectaMundi.2007-2008.journal_article" docStyleId="A87B5EC635B7D5A430FC1D805C706BD1" docStyleName="InsectaMundi.2007-2008.journal_article" docStyleVersion="3" docTitle="Heterogomphus eteocles Burmeister 1847" docType="treatment" docVersion="3" lastPageNumber="22" masterDocId="FFC56271FFDFF767FFC24F620949FFF3" masterDocTitle="Synopsis of the Oryctini (Coleoptera: Scarabaeidae: Dynastinae) from the Brazilian Amazon" masterLastPageNumber="62" masterPageNumber="1" pageNumber="21" updateTime="1699185735107" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="F067AEEBAECB49F000A89D780A464C7B">Synopsis of the Oryctini (Coleoptera: Scarabaeidae: Dynastinae) from the Brazilian Amazon</mods:title>
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<mods:namePart id="B6B04A17042AB7002EFCDDDD25462A99">Alvarez, Héctor Jaime Gasca</mods:namePart>
<mods:affiliation id="DF50CC56040F1AC48182895A2247BA66">Instituto Nacional de Pesquisas da Amazônia-INPA Coordenação de Pesquisas em Entomologia Av. André Araújo, 2936 - Petrópolis CEP 69011 - 970 Manaus, Amazonas, Brazil</mods:affiliation>
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<mods:namePart id="3FD59B8C2E09D25BCF9F6E151419067F">da, Claudio Ruy Vasconcelos</mods:namePart>
<mods:affiliation id="170651CCC72120C2C95E98199A647F05">Instituto Nacional de Pesquisas da Coordenação de Pesquisas em Av. André Araújo, 2936 - Petrópolis CEP 69011 - 970 Manaus, Amazonas</mods:affiliation>
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<mods:namePart id="AAA376389413EFFC7693237FFFE88F67">Fonseca</mods:namePart>
<mods:affiliation id="EB76B33CFFB060165FA8752CEAF51424">Amazônia-INPA Entomologia Petrópolis, Brazil</mods:affiliation>
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<mods:namePart id="B633098827031C8854DDCDEB6F6BAED5">Ratcliffe, Brett C.</mods:namePart>
<mods:affiliation id="7AFFBFB3CBEB84B27FA3A3FD1F1FB209">University of Nebraska State Museum W 436 Nebraska Hall Lincoln, NE 68588 - 0514, USA</mods:affiliation>
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<mods:title id="1809FE5B4B3A7DCFBF091C13F5814853">Insecta Mundi</mods:title>
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<mods:date id="77377AEB0754CD1088AFC02F288FB534">2008</mods:date>
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<mods:number id="5B50C8701B1ECAB7390E6164F424AFC4">2008</mods:number>
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<mods:number id="BA7622CE6B8C2391352749D31B450C98">61</mods:number>
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<mods:identifier id="EBAA766A7D1834A02BC2B4162A934729" type="ISSN">1942-1354</mods:identifier>
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<subSubSection id="C34FF894FFC9F771FF024B410BA9FBC8" box="[192,736,1059,1083]" pageId="22" pageNumber="21" type="nomenclature">
<paragraph id="8BEAAB1FFFC9F771FF024B410BA9FBC8" blockId="22.[192,736,1059,1083]" box="[192,736,1059,1083]" pageId="22" pageNumber="21">
<heading id="D0A21C73FFC9F771FF024B410BA9FBC8" bold="true" box="[192,736,1059,1083]" fontSize="10" level="2" pageId="22" pageNumber="21" reason="4">
<taxonomicName id="4C55D09CFFC9F771FF024B410BA9FBC8" authority="Burmeister 1847" authorityName="Burmeister" authorityYear="1847" box="[192,736,1059,1083]" class="Insecta" family="Dynastidae" genus="Heterogomphus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="21" phylum="Arthropoda" rank="species" species="eteocles">
<emphasis id="B921770DFFC9F771FF024B410BA9FBC8" bold="true" box="[192,736,1059,1083]" pageId="22" pageNumber="21">
<emphasis id="B921770DFFC9F771FF024B410B49FBC8" bold="true" box="[192,512,1059,1083]" italics="true" pageId="22" pageNumber="21">Heterogomphus eteocles</emphasis>
<bibRefCitation id="EFC4D6EEFFC9F771FDC64B410BA9FBC8" author="Burmeister, H. C. C." box="[516,736,1059,1083]" pageId="22" pageNumber="21" refId="ref23787" refString="Burmeister, H. C. C. 1847. Handbuch der Entomologie. (Coleoptera, Lamellicornia Xylophila et Pectinicornia) Vol. 5. T. C. F. Enslin; Berlin, Germany. 584 p." type="book" year="1847">Burmeister 1847</bibRefCitation>
</emphasis>
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</heading>
</paragraph>
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<subSubSection id="C34FF894FFC9F771FF024B210801FBA8" box="[192,328,1091,1115]" pageId="22" pageNumber="21" type="description">
<paragraph id="8BEAAB1FFFC9F771FF024B210801FBA8" blockId="22.[192,328,1091,1115]" box="[192,328,1091,1115]" pageId="22" pageNumber="21">
(
<figureCitation id="136EB79AFFC9F771FF0B4B210809FBA8" box="[201,320,1091,1115]" captionStart-0="Figure 29-31" captionStart-1="Figure 32" captionStartId-0="22.[192,276,782,804]" captionStartId-1="23.[195,279,1105,1127]" captionTargetBox-0="[201,1393,204,749]" captionTargetBox-1="[208,1432,199,1069]" captionTargetId-0="figure-372@22.[192,1413,191,762]" captionTargetId-1="figure-155@23.[195,1444,189,1075]" captionTargetPageId-0="22" captionTargetPageId-1="23" captionText-0="Figure 29-31. Heterogomphus eteocles. 29) Metatarsus. 30) Parameres frontal view. 31) Parameres lateral view (scale line: 2 mm)." captionText-1="Figure 32. Distribution of Heterogomphus eteocles in Brazilian Amazonia." figureDoi-0="http://doi.org/10.5281/zenodo.5352464" figureDoi-1="http://doi.org/10.5281/zenodo.5352466" httpUri-0="https://zenodo.org/record/5352464/files/figure.png" httpUri-1="https://zenodo.org/record/5352466/files/figure.png" pageId="22" pageNumber="21">Fig. 29-32</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C34FF894FFC9F771FF024BE10A7FFB48" pageId="22" pageNumber="21" type="reference_group">
<paragraph id="8BEAAB1FFFC9F771FF024BE10BA7FB68" blockId="22.[192,822,1155,1211]" box="[192,750,1155,1179]" pageId="22" pageNumber="21">
<treatmentCitationGroup id="AB458C31FFC9F771FF024BE10BA7FB68" box="[192,750,1155,1179]" pageId="22" pageNumber="21">
<treatmentCitation id="0AF48D0EFFC9F771FF024BE10BA7FB68" author="Burmeister, H. C. C." box="[192,750,1155,1179]" page="228" pageId="22" pageNumber="21" year="1847">
<taxonomicName id="4C55D09CFFC9F771FF024BE10BA7FB68" authority="Burmeister 1847: 228" authorityName="Burmeister" authorityPageNumber="228" authorityYear="1847" box="[192,750,1155,1179]" class="Insecta" family="Dynastidae" genus="Heterogomphus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="21" phylum="Arthropoda" rank="species" species="eteocles">
<emphasis id="B921770DFFC9F771FF024BE108A8FB68" box="[192,481,1155,1179]" italics="true" pageId="22" pageNumber="21">Heterogomphus eteocles</emphasis>
<bibRefCitation id="EFC4D6EEFFC9F771FE244BE10BA7FB68" author="Burmeister, H. C. C." box="[486,750,1155,1179]" pageId="22" pageNumber="21" refId="ref23787" refString="Burmeister, H. C. C. 1847. Handbuch der Entomologie. (Coleoptera, Lamellicornia Xylophila et Pectinicornia) Vol. 5. T. C. F. Enslin; Berlin, Germany. 584 p." type="book" year="1847">Burmeister 1847: 228</bibRefCitation>
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</paragraph>
<paragraph id="8BEAAB1FFFC9F771FF024BC10A7FFB48" blockId="22.[192,822,1155,1211]" box="[192,822,1187,1211]" pageId="22" pageNumber="21">
<treatmentCitationGroup id="AB458C31FFC9F771FF024BC10A7FFB48" box="[192,822,1187,1211]" pageId="22" pageNumber="21">
<treatmentCitation id="0AF48D0EFFC9F771FF024BC10BF9FB48" author="Prell, H." box="[192,688,1187,1211]" page="109" pageId="22" pageNumber="21" year="1912">
<taxonomicName id="4C55D09CFFC9F771FF024BC10BF9FB48" authority="Prell 1912: 109" authorityName="Prell" authorityPageNumber="109" authorityYear="1912" box="[192,688,1187,1211]" class="Insecta" family="Dynastidae" genus="Heterogomphus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="21" phylum="Arthropoda" rank="species" species="polynices">
<emphasis id="B921770DFFC9F771FF024BC108BCFB48" box="[192,501,1187,1211]" italics="true" pageId="22" pageNumber="21">Heterogomphus polynices</emphasis>
<bibRefCitation id="EFC4D6EEFFC9F771FE384BC10BF9FB48" author="Prell, H." box="[506,688,1187,1211]" pageId="22" pageNumber="21" pagination="93 - 176" refId="ref27079" refString="Prell, H. 1912. Revision des Dynastinen-Gattung Heterogomphus Burm. Memoirs de la Societe Entomologique de Belgique 20: 93 - 176." type="journal article" year="1912">Prell 1912: 109</bibRefCitation>
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(synonym)
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</paragraph>
</subSubSection>
<subSubSection id="C34FF894FFC9F771FF024B810B39F928" pageId="22" pageNumber="21" type="description">
<paragraph id="8BEAAB1FFFC9F771FF024B8108A7FAE8" blockId="22.[192,1441,1251,1755]" pageId="22" pageNumber="21">
<emphasis id="B921770DFFC9F771FF024B8108D2FB08" bold="true" box="[192,411,1251,1275]" pageId="22" pageNumber="21">DESCRIPTION.</emphasis>
Length:
<quantity id="4CAD06FAFFC9F771FDCF4B810BF3FB08" box="[525,698,1251,1275]" metricMagnitude="-2" metricUnit="m" metricValue="4.85" metricValueMax="5.55" metricValueMin="4.15" pageId="22" pageNumber="21" unit="mm" value="48.5" valueMax="55.5" valueMin="41.5">41.5-55.5 mm</quantity>
. Width: 24.5-26.0 mm. Color: Dark reddish brown to black; pronotum brown to black
</paragraph>
<paragraph id="8BEAAB1FFFC9F771FF324A410BEBF948" blockId="22.[192,1441,1251,1755]" pageId="22" pageNumber="21">
<emphasis id="B921770DFFC9F771FF324A410800FAC8" bold="true" box="[240,329,1315,1339]" pageId="22" pageNumber="21">Males.</emphasis>
<emphasis id="B921770DFFC9F771FE924A4108DBFAC8" box="[336,402,1315,1339]" italics="true" pageId="22" pageNumber="21">Head</emphasis>
: Frons with long, recurved horn, posterior margin of horn with a small prominence, apex narrowly truncate, slightly bilobed, surface densely rugose. Eye canthus with apex rounded, surface punctate and rugose. Antenna with 10 segments, club subequal in length to segments 2-7. Mandibles with 2 long, subequal teeth, apices of teeth rounded.
<emphasis id="B921770DFFC9F771FCFB4AE60AFAFA68" box="[825,947,1412,1435]" italics="true" pageId="22" pageNumber="21">Pronotum</emphasis>
: Prominence long, high, projecting at 90
<superScript id="7C200657FFC9F771FA5A4AE30CE9FA63" attach="none" box="[1432,1440,1409,1424]" fontSize="6" pageId="22" pageNumber="21">o</superScript>
from the body; apex bifurcate forming 2 small quadrangular horns, horns projecting forward. Sides with broad rugose area in basal half. Fovea broad, surface smooth with small punctures.
<emphasis id="B921770DFFC9F771FB004AA10C58FA28" box="[1218,1297,1475,1499]" italics="true" pageId="22" pageNumber="21">Elytra</emphasis>
: Impressed sutural stria present. Surface usually smooth with small, sparse punctures, lateral margin weakly rugose.
<taxonomicName id="4C55D09CFFC9F771FF314961089BF9E8" authority=": Surface" authorityName="Surface" box="[243,466,1539,1563]" class="Actinopterygii" family="Trichomycteridae" genus="Pygidium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="22" pageNumber="21" phylum="Chordata" rank="genus">
<emphasis id="B921770DFFC9F771FF3149610821F9E8" box="[243,360,1539,1563]" italics="true" pageId="22" pageNumber="21">Pygidium</emphasis>
: Surface
</taxonomicName>
finely punctate and rugose with a transverse band of long, reddish brown setae on posterior margin. Angles weakly rugose. Surface convex in lateral view.
<emphasis id="B921770DFFC9F771FBF449460D27F9C8" box="[1078,1134,1572,1595]" italics="true" pageId="22" pageNumber="21">Legs</emphasis>
: Protibia quadridentate. Apex of posterior tibia strongly crenulate. Apex of first tarsomere of posterior tarsus subtriangular, elongate, spur-like (
<figureCitation id="136EB79AFFC9F771FEBF4901089AF988" box="[381,467,1635,1659]" captionStart="Figure 29-31" captionStartId="22.[192,276,782,804]" captionTargetBox="[201,1393,204,749]" captionTargetId="figure-372@22.[192,1413,191,762]" captionTargetPageId="22" captionText="Figure 29-31. Heterogomphus eteocles. 29) Metatarsus. 30) Parameres frontal view. 31) Parameres lateral view (scale line: 2 mm)." figureDoi="http://doi.org/10.5281/zenodo.5352464" httpUri="https://zenodo.org/record/5352464/files/figure.png" pageId="22" pageNumber="21">Fig. 29</figureCitation>
).
<emphasis id="B921770DFFC9F771FE2649060B7AF988" box="[484,563,1636,1659]" italics="true" pageId="22" pageNumber="21">Venter</emphasis>
: Prosternal process short to moderate in length, conical, apex acuminate.
<emphasis id="B921770DFFC9F771FF0249E6080DF968" box="[192,324,1668,1691]" italics="true" pageId="22" pageNumber="21">Parameres</emphasis>
: Oval, contracted toward apices, sides curved. Posterior region with deep punctures. Apex setigerous on inner margin (
<figureCitation id="136EB79AFFC9F771FDD549C10BDCF948" box="[535,661,1699,1723]" captionStart="Figure 29-31" captionStartId="22.[192,276,782,804]" captionTargetBox="[201,1393,204,749]" captionTargetId="figure-372@22.[192,1413,191,762]" captionTargetPageId="22" captionText="Figure 29-31. Heterogomphus eteocles. 29) Metatarsus. 30) Parameres frontal view. 31) Parameres lateral view (scale line: 2 mm)." figureDoi="http://doi.org/10.5281/zenodo.5352464" httpUri="https://zenodo.org/record/5352464/files/figure.png" pageId="22" pageNumber="21">Fig. 30-31</figureCitation>
).
</paragraph>
<paragraph id="8BEAAB1FFFC9F771FF3249A10B39F928" blockId="22.[192,1441,1251,1755]" box="[240,624,1731,1755]" pageId="22" pageNumber="21">
<emphasis id="B921770DFFC9F771FF3249A10828F928" bold="true" box="[240,353,1731,1755]" pageId="22" pageNumber="21">Females</emphasis>
. Females unavailable.
</paragraph>
</subSubSection>
<subSubSection id="C34FF894FFC9F771FF0248610852F8A8" pageId="22" pageNumber="21" type="diagnosis">
<paragraph id="8BEAAB1FFFC9F771FF0248610852F8A8" blockId="22.[192,1440,1793,1883]" pageId="22" pageNumber="21">
<emphasis id="B921770DFFC9F771FF0248610838F8E8" bold="true" box="[192,369,1795,1819]" pageId="22" pageNumber="21">DIAGNOSIS.</emphasis>
Adults of
<taxonomicName id="4C55D09CFFC9F771FE3048660B3AF8E8" authorityName="Burmeister" authorityYear="1847" box="[498,627,1795,1819]" class="Insecta" family="Dynastidae" genus="Heterogomphus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="21" phylum="Arthropoda" rank="species" species="eteocles">
<emphasis id="B921770DFFC9F771FE3048660B3AF8E8" box="[498,627,1795,1819]" italics="true" pageId="22" pageNumber="21">H. eteocles</emphasis>
</taxonomicName>
can be recognized by the pronotal horn projecting at 90
<superScript id="7C200657FFC9F771FAE448630C67F8E3" attach="left" box="[1318,1326,1793,1808]" fontSize="6" pageId="22" pageNumber="21">o</superScript>
from the body, bifurcate at the apex with two quadrangular lobes. The frontal horn is long, recurved, and weakly bilobed.
</paragraph>
</subSubSection>
<caption id="DF2AFB97FFC8F770FF014B330D4BFB94" ID-DOI="http://doi.org/10.5281/zenodo.5352466" ID-Zenodo-Dep="5352466" box="[195,1026,1105,1127]" httpUri="https://zenodo.org/record/5352466/files/figure.png" pageId="23" pageNumber="22" startId="23.[195,279,1105,1127]" targetBox="[208,1432,199,1069]" targetPageId="23">
<paragraph id="8BEAAB1FFFC8F770FF014B330D4BFB94" blockId="23.[195,1026,1105,1127]" box="[195,1026,1105,1127]" pageId="23" pageNumber="22">
<emphasis id="B921770DFFC8F770FF014B330809FB95" bold="true" box="[195,320,1105,1127]" pageId="23" pageNumber="22">Figure 32.</emphasis>
Distribution of
<taxonomicName id="4C55D09CFFC8F770FE334B330BB1FB95" authorityName="Burmeister" authorityYear="1847" box="[497,760,1105,1126]" class="Insecta" family="Dynastidae" genus="Heterogomphus" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="22" phylum="Arthropoda" rank="species" species="eteocles">
<emphasis id="B921770DFFC8F770FE334B330BB1FB95" box="[497,760,1105,1126]" italics="true" pageId="23" pageNumber="22">Heterogomphus eteocles</emphasis>
</taxonomicName>
in Brazilian Amazonia.
</paragraph>
</caption>
<subSubSection id="C34FF894FFC8F770FF024BF90BF1FA20" pageId="23" pageNumber="22" type="distribution">
<paragraph id="8BEAAB1FFFC8F770FF024BF90BABFB00" blockId="23.[192,1440,1179,1267]" pageId="23" pageNumber="22">
<emphasis id="B921770DFFC8F770FF024BF908EEFB40" bold="true" box="[192,423,1179,1203]" pageId="23" pageNumber="22">DISTRIBUTION.</emphasis>
<taxonomicName id="4C55D09CFFC8F770FE6F4BF90B87FB40" authorityName="Burmeister" authorityYear="1847" box="[429,718,1179,1203]" class="Insecta" family="Dynastidae" genus="Heterogomphus" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="22" phylum="Arthropoda" rank="species" species="eteocles">
<emphasis id="B921770DFFC8F770FE6F4BF90B87FB40" box="[429,718,1179,1203]" italics="true" pageId="23" pageNumber="22">Heterogomphus eteocles</emphasis>
</taxonomicName>
is distributed only in South America, occurring from southern
<collectingCountry id="F342EB8FFFC8F770FF2D4BD90873FB20" box="[239,314,1211,1235]" name="Brazil" pageId="23" pageNumber="22">Brazil</collectingCountry>
to northern
<collectingCountry id="F342EB8FFFC8F770FE0D4BD90B06FB20" box="[463,591,1211,1235]" name="Argentina" pageId="23" pageNumber="22">Argentina</collectingCountry>
. The specimens listed below from Amazonas and Pará states represent NEW RECORDS from the Brazilian Amazon.
</paragraph>
<paragraph id="8BEAAB1FFFC8F770FF024A790888FAA0" blockId="23.[192,1440,1307,1363]" pageId="23" pageNumber="22">
<materialsCitation id="3B3DA142FFC8F770FF024A79082AFAA0" collectionCode="LOCALITY, RECORDS" pageId="23" pageNumber="22" specimenCount="4" specimenCount-male="2">
<emphasis id="B921770DFFC8F770FF024A7908BBFAC0" bold="true" box="[192,498,1307,1331]" pageId="23" pageNumber="22">
<collectionCode id="ED4433DAFFC8F770FF024A7908A4FAC0" box="[192,493,1307,1331]" pageId="23" pageNumber="22">LOCALITY RECORDS</collectionCode>
.
</emphasis>
(
<figureCitation id="136EB79AFFC8F770FDC24A790B1BFAC0" box="[512,594,1307,1331]" captionStart="Figure 32" captionStartId="23.[195,279,1105,1127]" captionTargetBox="[208,1432,199,1069]" captionTargetId="figure-155@23.[195,1444,189,1075]" captionTargetPageId="23" captionText="Figure 32. Distribution of Heterogomphus eteocles in Brazilian Amazonia." figureDoi="http://doi.org/10.5281/zenodo.5352466" httpUri="https://zenodo.org/record/5352466/files/figure.png" pageId="23" pageNumber="22">Fig. 32</figureCitation>
)
<specimenCount id="9D536096FFC8F770FD9D4A790BA7FAC0" box="[607,750,1307,1331]" pageId="23" pageNumber="22" type="generic">2 specimens</specimenCount>
examined (
<specimenCount id="9D536096FFC8F770FCB44A790A9CFAC0" box="[886,981,1307,1331]" pageId="23" pageNumber="22" type="male">2 males</specimenCount>
). Specimens were seen from the following collection
</materialsCitation>
: MZSP.
</paragraph>
<paragraph id="8BEAAB1FFFC8F770FF024A190ABEFA67" blockId="23.[192,1015,1402,1428]" box="[192,1015,1402,1428]" pageId="23" pageNumber="22">
<materialsCitation id="3B3DA142FFC8F770FF024A190A5DFA60" box="[192,788,1403,1427]" location="Rio Javari" pageId="23" pageNumber="22" specimenCount="1" stateProvince="Amazonas">
<collectingRegion id="499165FDFFC8F770FF024A190817FA60" box="[192,350,1403,1427]" country="Venezuela" name="Amazonas" pageId="23" pageNumber="22">AMAZONAS</collectingRegion>
(1): Benjamin Constant (
<location id="8E8AFDC4FFC8F770FD484A190A42FA60" LSID="urn:lsid:plazi:treatment:03FC1A09FFC9F770FF024B410DA9F9A0:8E8AFDC4FFC8F770FD484A190A42FA60" box="[650,779,1403,1427]" name="Rio Javari" pageId="23" pageNumber="22" stateProvince="Amazonas">Rio Javari</location>
)
</materialsCitation>
.
<collectingRegion id="499165FDFFC8F770FCDF4A180A21FA67" box="[797,872,1402,1428]" country="Suriname" name="Para" pageId="23" pageNumber="22">PARÁ</collectingRegion>
(1): Óbidos.
</paragraph>
<paragraph id="8BEAAB1FFFC8F770FF024AD90BF1FA20" blockId="23.[192,696,1467,1491]" box="[192,696,1467,1491]" pageId="23" pageNumber="22">
<emphasis id="B921770DFFC8F770FF024AD90B19FA20" bold="true" box="[192,592,1467,1491]" pageId="23" pageNumber="22">TEMPORAL DISTRIBUTION.</emphasis>
April (2)
</paragraph>
</subSubSection>
<subSubSection id="C34FF894FFC8F770FF024A990DA9F9A0" pageId="23" pageNumber="22" type="biology_ecology">
<paragraph id="8BEAAB1FFFC8F770FF024A990DA9F9A0" blockId="23.[192,1440,1531,1619]" pageId="23" pageNumber="22">
<emphasis id="B921770DFFC8F770FF024A99081BF9E0" bold="true" box="[192,338,1531,1555]" pageId="23" pageNumber="22">BIOLOGY.</emphasis>
Little is known about the biology of this species.
<taxonomicName id="4C55D09CFFC8F770FC624A990D88F9E0" authorityName="Burmeister" authorityYear="1847" box="[928,1217,1531,1555]" class="Insecta" family="Dynastidae" genus="Heterogomphus" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="22" phylum="Arthropoda" rank="species" species="eteocles">
<emphasis id="B921770DFFC8F770FC624A990D88F9E0" box="[928,1217,1531,1555]" italics="true" pageId="23" pageNumber="22">Heterogomphus eteocles</emphasis>
</taxonomicName>
has been reported as a pest of
<taxonomicName id="4C55D09CFFC8F770FE8A49790A70F9C0" authority="Lindm." authorityName="Lindm." box="[328,825,1563,1587]" class="Liliopsida" family="Poaceae" genus="Chusquea" kingdom="Plantae" order="Poales" pageId="23" pageNumber="22" phylum="Tracheophyta" rank="species" species="ramosissima">
<emphasis id="B921770DFFC8F770FE8A49790B2BF9C0" box="[328,610,1563,1587]" italics="true" pageId="23" pageNumber="22">Chusquea ramosissima</emphasis>
Lindm. (Poaceae)
</taxonomicName>
in
<collectingCountry id="F342EB8FFFC8F770FC9E49790A91F9C0" box="[860,984,1563,1587]" name="Argentina" pageId="23" pageNumber="22">Argentina</collectingCountry>
(
<bibRefCitation id="EFC4D6EEFFC8F770FC2749790D3FF9C0" author="Keller, H. A." box="[997,1142,1563,1587]" pageId="23" pageNumber="22" pagination="461 - 471" refId="ref25681" refString="Keller, H. A. 2003. Mythical origin of Chusquea ramosissima (Poaceae), the ancient knife of the Guaranis. Economic Botany 57 (4): 461 - 471." type="journal article" year="2003">Keller 2003</bibRefCitation>
). In Brazilian Amazonia, adults have been collected from areas of seasonal whitewater inundation forest (várzea).
</paragraph>
</subSubSection>
</treatment>
</document>