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<mods:title id="41C9A102DAD7D93E9711A6884495C521">Revision and cladistic analysis of the Afrotropical endemic genus Smeringopus Simon, 1890 (Araneae: Pholcidae)</mods:title>
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<treatment id="03FBB532FFF41770FF6A0D6A3B2FF84B" ID-DOI="http://doi.org/10.5281/zenodo.5256566" ID-GBIF-Taxon="194469068" ID-Zenodo-Dep="5256566" LSID="urn:lsid:plazi:treatment:03FBB532FFF41770FF6A0D6A3B2FF84B" httpUri="http://treatment.plazi.org/id/03FBB532FFF41770FF6A0D6A3B2FF84B" lastPageId="115" lastPageNumber="116" pageId="113" pageNumber="114">
<subSubSection id="C34857AFFFF41772FF6A0D6A38DCF98C" pageId="113" pageNumber="114" type="nomenclature">
<paragraph id="8BED0424FFF41772FF6A0D6A3872F9A3" blockId="113.[151,758,1597,1656]" box="[151,600,1597,1623]" pageId="113" pageNumber="114">
<heading id="D0A5B348FFF41772FF6A0D6A3872F9A3" bold="true" box="[151,600,1597,1623]" fontSize="11" level="1" pageId="113" pageNumber="114" reason="1">
<taxonomicName id="4C527FA7FFF41772FF6A0D6A3872F9A3" ID-CoL="4XVJR" authority="Simon, 1907" authorityName="Simon" authorityYear="1907" box="[151,600,1597,1623]" class="Arachnida" family="Pholcidae" genus="Smeringopus" kingdom="Animalia" order="Araneae" pageId="113" pageNumber="114" phylum="Arthropoda" rank="species" species="thomensis">
<emphasis id="B926D836FFF41772FF6A0D6A3872F9A3" bold="true" box="[151,600,1597,1623]" pageId="113" pageNumber="114">
<emphasis id="B926D836FFF41772FF6A0D6A3B92F9A3" bold="true" box="[151,440,1597,1623]" italics="true" pageId="113" pageNumber="114">Smeringopus thomensis</emphasis>
<bibRefCitation id="EFC379D5FFF41772FE420D6A3872F9A3" author="Simon, E." box="[447,600,1597,1623]" firstAuthor="Simon" journalOrPublisher="Annales di Museo Civico, Genova" pageId="113" pageNumber="114" pagination="218 - 323" part="43" refId="ref81576" refString="Simon, E. (1907) Arachnides recueillis par L. Fea sur la cote occidentale d'Afrique. Annales di Museo Civico, Genova, 43: 218 - 323." title="Arachnides recueillis par L. Fea sur la cote occidentale d'Afrique" type="journal article" year="1907">Simon, 1907</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BED0424FFF41772FF6A0D3738DCF98C" blockId="113.[151,758,1597,1656]" box="[151,758,1632,1656]" pageId="113" pageNumber="114">
<figureCitation id="136918A1FFF41772FF6A0D373B80F98C" box="[151,426,1632,1656]" captionStart="FIGURES 699708" captionStartId="114.[151,263,1415,1438]" captionTargetBox="[156,1436,153,1388]" captionTargetId="figure-308@114.[151,1436,153,1401]" captionTargetPageId="114" captionText="FIGURES 699708. Smeringopus thomensis group. S. thomensis (699700, 705706), S. principe (702704, 707708), and S. mayombe (701). 699, 701, 703. Males, dorsal views. 700, 702. Male prosomata, oblique frontal views. 704. Female, ventral view. 705708. Left male palps, prolateral and retrolateral views." figureDoi="http://doi.org/10.5281/zenodo.6418204" httpUri="https://zenodo.org/record/6418204/files/figure.png" pageId="113" pageNumber="114">Figs. 699700, 705706</figureCitation>
, 709711, 719724, 732740
</paragraph>
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<paragraph id="8BED0424FFF41772FF6A0DF6385FF943" blockId="113.[151,629,1697,1719]" box="[151,629,1697,1719]" pageId="113" pageNumber="114">
<treatmentCitationGroup id="AB42230AFFF41772FF6A0DF6385FF943" box="[151,629,1697,1719]" pageId="113" pageNumber="114">
<treatmentCitation id="0AF32235FFF41772FF6A0DF6385BF943" author="Simon, E." box="[151,625,1697,1719]" page="252" pageId="113" pageNumber="114" year="1907">
<taxonomicName id="4C527FA7FFF41772FF6A0DF6385BF943" ID-CoL="4XVJR" authority="Simon 1907: 251 - 252" authorityName="Simon" authorityPageNumber="251 - 252" authorityYear="1907" box="[151,625,1697,1719]" class="Arachnida" family="Pholcidae" genus="Smeringopus" kingdom="Animalia" order="Araneae" pageId="113" pageNumber="114" phylum="Arthropoda" rank="species" species="thomensis">
<emphasis id="B926D836FFF41772FF6A0DF63BACF943" box="[151,390,1697,1719]" italics="true" pageId="113" pageNumber="114">Smeringopus thomensis</emphasis>
<bibRefCitation id="EFC379D5FFF41772FE700DF6385BF943" author="Simon, E." box="[397,625,1697,1719]" firstAuthor="Simon" journalOrPublisher="Annales di Museo Civico, Genova" page="251 - 252" pageId="113" pageNumber="114" pagination="218 - 323" part="43" refId="ref81576" refString="Simon, E. (1907) Arachnides recueillis par L. Fea sur la cote occidentale d'Afrique. Annales di Museo Civico, Genova, 43: 218 - 323." title="Arachnides recueillis par L. Fea sur la cote occidentale d'Afrique" type="journal article" year="1907">Simon 1907: 251252</bibRefCitation>
</taxonomicName>
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.
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</paragraph>
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<paragraph id="8BED0424FFF41772FF6A0D893E4AF8ED" blockId="113.[151,1436,1757,2033]" pageId="113" pageNumber="114">
Type.
<materialsCitation id="3B3A0E79FFF41772FF200D8A3E70F8ED" ID-GBIF-Occurrence="3747820825" collectorName="L. Fea" country="Sao Tome and Principe" latitude="0.35" location="Ribeira Palma" longLatPrecision="1308" longitude="6.5833335" municipality="Museo Civico di Storia Naturale" pageId="113" pageNumber="114" specimenCount="1" stateProvince="Sao Tome Island" typeStatus="holotype">
<specimenCount id="9D54CFADFFF41772FF200D8A3B3CF901" box="[221,278,1757,1781]" pageId="113" pageNumber="114" type="male">Male</specimenCount>
<typeStatus id="54E9BA86FFF41772FEE30D8A3BAAF901" box="[286,384,1757,1781]" pageId="113" pageNumber="114" type="holotype">holotype</typeStatus>
from
<collectingCountry id="F34544B4FFF41772FE380D8A38E7F901" box="[453,717,1757,1781]" name="Sao Tome and Principe" pageId="113" pageNumber="114">São Tomé and Principe</collectingCountry>
,
<collectingRegion id="4996CAC6FFF41772FD250D8A39B2F901" box="[728,920,1757,1781]" pageId="113" pageNumber="114">São Tomé Island</collectingRegion>
,
<location id="8E8D52FFFFF41772FC5E0D8A3E69F901" LSID="urn:lsid:plazi:treatment:03FBB532FFF41770FF6A0D6A3B2FF84B:8E8D52FFFFF41772FC5E0D8A3E69F901" box="[931,1091,1757,1781]" country="Sao Tome and Principe" latitude="0.35" longLatPrecision="1308" longitude="6.5833335" municipality="Museo Civico di Storia Naturale" name="Ribeira Palma" pageId="113" pageNumber="114" stateProvince="Sao Tome Island">Ribeira Palma</location>
[
<geoCoordinate id="EE6662E3FFF41772FBAF0D8A3E83F901" box="[1106,1193,1757,1781]" degrees="0" direction="north" minutes="21" orientation="latitude" pageId="113" pageNumber="114" precision="925" value="0.35">0°21N</geoCoordinate>
,
<geoCoordinate id="EE6662E3FFF41772FB480D8A3F23F901" box="[1205,1289,1757,1781]" degrees="6" direction="east" minutes="35" orientation="longitude" pageId="113" pageNumber="114" precision="925" value="6.5833335">6°35E</geoCoordinate>
], leg.
<collectorName id="26A761F2FFF41772FAB00D893FB2F901" box="[1357,1432,1758,1781]" pageId="113" pageNumber="114">L. Fea</collectorName>
, possibly in Museo Civico di Storia Naturale &quot;Giacomo Doria&quot;, Genova, not examined
</materialsCitation>
.
</paragraph>
<paragraph id="8BED0424FFF41772FF3A0C723851F805" blockId="113.[151,1436,1757,2033]" pageId="113" pageNumber="114">
Diagnosis. Distinguished from similar congeners (other species of the
<taxonomicName id="4C527FA7FFF41772FC120C723E75F8C9" authorityName="Simon" authorityYear="1907" box="[1007,1119,1829,1853]" class="Arachnida" family="Pholcidae" genus="Smeringopus" kingdom="Animalia" order="Araneae" pageId="113" pageNumber="114" phylum="Arthropoda" rank="species" species="thomensis">
<emphasis id="B926D836FFF41772FC120C723E75F8C9" box="[1007,1119,1829,1853]" italics="true" pageId="113" pageNumber="114">thomensis</emphasis>
</taxonomicName>
species group) by shape of embolus (two large rounded apophyses arising from embolus dorsally;
<figureCitation id="136918A1FFF41772FC210C1E3EACF895" box="[988,1158,1865,1889]" captionStart="FIGURES 719724" captionStartId="116.[151,257,1954,1975]" captionTargetBox="[300,1299,910,1935]" captionTargetId="figure-85@116.[288,1303,900,1945]" captionTargetPageId="116" captionText="FIGURES 719724. Smeringopus thomensis. 719. Left cymbium and procursus, retrolateral view. 720. Left procursus, prolateral view. 721. Male chelicerae, frontal view. 722723. Left embolus, prolateral and dorsal (slightly distal) views. 724. Cleared female genitalia, dorsal view. Scale lines: 0.3 mm." figureDoi="http://doi.org/10.5281/zenodo.6415830" httpUri="https://zenodo.org/record/6415830/files/figure.png" pageId="113" pageNumber="114">Figs. 722, 723</figureCitation>
) and absence of pair of sclerotized areas on posterior rim of epigynum (
<figureCitation id="136918A1FFF41772FD3D0C3A390FF871" box="[704,805,1901,1925]" captionStart="FIGURES 709717" captionStartId="115.[151,263,1495,1518]" captionTargetBox="[151,1436,191,1473]" captionTargetId="figure-188@115.[151,1436,191,1473]" captionTargetPageId="115" captionText="FIGURES 709717. Smeringopus thomensis group. Epigyna, ventral views and cleared female genitalia, ventral and dorsal views. 709711. S. thomensis. 712714. S. principe. 715717. S. mayombe." figureDoi="http://doi.org/10.5281/zenodo.6415826" httpUri="https://zenodo.org/record/6415826/files/figure.png" pageId="113" pageNumber="114">Fig. 709</figureCitation>
); from
<taxonomicName id="4C527FA7FFF41772FC870C3939DCF871" authorityName="Huber" authorityYear="2012" box="[890,1014,1902,1926]" class="Arachnida" family="Pholcidae" genus="Smeringopus" kingdom="Animalia" order="Araneae" pageId="113" pageNumber="114" phylum="Arthropoda" rank="species" species="principe">
<emphasis id="B926D836FFF41772FC870C3939DCF871" box="[890,1014,1902,1926]" italics="true" pageId="113" pageNumber="114">S. principe</emphasis>
</taxonomicName>
also by absence of dorsal projection from procursus tip (compare
<figureCitation id="136918A1FFF41772FE210CC63869F85D" box="[476,579,1937,1961]" captionStart="FIGURES 719724" captionStartId="116.[151,257,1954,1975]" captionTargetBox="[300,1299,910,1935]" captionTargetId="figure-85@116.[288,1303,900,1945]" captionTargetPageId="116" captionText="FIGURES 719724. Smeringopus thomensis. 719. Left cymbium and procursus, retrolateral view. 720. Left procursus, prolateral view. 721. Male chelicerae, frontal view. 722723. Left embolus, prolateral and dorsal (slightly distal) views. 724. Cleared female genitalia, dorsal view. Scale lines: 0.3 mm." figureDoi="http://doi.org/10.5281/zenodo.6415830" httpUri="https://zenodo.org/record/6415830/files/figure.png" pageId="113" pageNumber="114">Figs. 720</figureCitation>
and
<figureCitation id="136918A1FFF41772FD840CC6388EF85D" box="[633,676,1937,1961]" captionStart="FIGURES 725731" captionStartId="117.[151,258,1442,1463]" captionTargetBox="[213,1382,153,1416]" captionTargetId="figure-208@117.[175,1419,143,1416]" captionTargetPageId="117" captionText="FIGURES 725731. Smeringopus principe (725728) and S. mayombe (729731). 725. Left procursus, prolateral view. 726727, 729730. Left emboli, prolateral and dorsal (slightly distal) views. 728, 731. Cleared female genitalia, dorsal views. Scale lines: 0.3 mm." figureDoi="http://doi.org/10.5281/zenodo.6415832" httpUri="https://zenodo.org/record/6415832/files/figure.png" pageId="113" pageNumber="114">725</figureCitation>
) and much narrower pore plate (compare
<figureCitation id="136918A1FFF41772FB850CC63ECAF85D" box="[1144,1248,1937,1961]" captionStart="FIGURES 719724" captionStartId="116.[151,257,1954,1975]" captionTargetBox="[300,1299,910,1935]" captionTargetId="figure-85@116.[288,1303,900,1945]" captionTargetPageId="116" captionText="FIGURES 719724. Smeringopus thomensis. 719. Left cymbium and procursus, retrolateral view. 720. Left procursus, prolateral view. 721. Male chelicerae, frontal view. 722723. Left embolus, prolateral and dorsal (slightly distal) views. 724. Cleared female genitalia, dorsal view. Scale lines: 0.3 mm." figureDoi="http://doi.org/10.5281/zenodo.6415830" httpUri="https://zenodo.org/record/6415830/files/figure.png" pageId="113" pageNumber="114">Figs. 724</figureCitation>
and
<figureCitation id="136918A1FFF41772FAE80CC63F69F85D" box="[1301,1347,1937,1961]" captionStart="FIGURES 725731" captionStartId="117.[151,258,1442,1463]" captionTargetBox="[213,1382,153,1416]" captionTargetId="figure-208@117.[175,1419,143,1416]" captionTargetPageId="117" captionText="FIGURES 725731. Smeringopus principe (725728) and S. mayombe (729731). 725. Left procursus, prolateral view. 726727, 729730. Left emboli, prolateral and dorsal (slightly distal) views. 728, 731. Cleared female genitalia, dorsal views. Scale lines: 0.3 mm." figureDoi="http://doi.org/10.5281/zenodo.6415832" httpUri="https://zenodo.org/record/6415832/files/figure.png" pageId="113" pageNumber="114">728</figureCitation>
). Easily distinguished from all other congeners by distinctive color pattern dorsally on abdomen (figure composed of three black elements on posterior half;
<figureCitation id="136918A1FFF41772FDF00C8E3844F805" box="[525,622,2009,2033]" captionStart="FIGURES 699708" captionStartId="114.[151,263,1415,1438]" captionTargetBox="[156,1436,153,1388]" captionTargetId="figure-308@114.[151,1436,153,1401]" captionTargetPageId="114" captionText="FIGURES 699708. Smeringopus thomensis group. S. thomensis (699700, 705706), S. principe (702704, 707708), and S. mayombe (701). 699, 701, 703. Males, dorsal views. 700, 702. Male prosomata, oblique frontal views. 704. Female, ventral view. 705708. Left male palps, prolateral and retrolateral views." figureDoi="http://doi.org/10.5281/zenodo.6418204" httpUri="https://zenodo.org/record/6418204/files/figure.png" pageId="113" pageNumber="114">Fig. 699</figureCitation>
).
</paragraph>
<caption id="DF2D54ACFFF71771FF6A0ED03906FA2E" ID-DOI="http://doi.org/10.5281/zenodo.6418204" ID-Zenodo-Dep="6418204" httpUri="https://zenodo.org/record/6418204/files/figure.png" pageId="114" pageNumber="115" startId="114.[151,263,1415,1438]" targetBox="[156,1436,153,1388]" targetPageId="114">
<paragraph id="8BED0424FFF71771FF6A0ED03906FA2E" blockId="114.[151,1437,1415,1498]" pageId="114" pageNumber="115">
<emphasis id="B926D836FFF71771FF6A0ED03B46FA6A" bold="true" box="[151,364,1415,1438]" pageId="114" pageNumber="115">FIGURES 699708.</emphasis>
<taxonomicName id="4C527FA7FFF71771FE8E0ED03848FA69" authorityName="Simon" authorityYear="1907" box="[371,610,1415,1437]" class="Arachnida" family="Pholcidae" genus="Smeringopus" kingdom="Animalia" order="Araneae" pageId="114" pageNumber="115" phylum="Arthropoda" rank="species" species="thomensis">
<emphasis id="B926D836FFF71771FE8E0ED03848FA69" box="[371,610,1415,1437]" italics="true" pageId="114" pageNumber="115">Smeringopus thomensis</emphasis>
</taxonomicName>
group.
<taxonomicName id="4C527FA7FFF71771FD4D0ED03904FA69" authorityName="Simon" authorityYear="1907" box="[688,814,1415,1437]" class="Arachnida" family="Pholcidae" genus="Smeringopus" kingdom="Animalia" order="Araneae" pageId="114" pageNumber="115" phylum="Arthropoda" rank="species" species="thomensis">
<emphasis id="B926D836FFF71771FD4D0ED03904FA69" box="[688,814,1415,1437]" italics="true" pageId="114" pageNumber="115">S. thomensis</emphasis>
</taxonomicName>
(699700, 705706),
<taxonomicName id="4C527FA7FFF71771FBF20ED03E57FA69" authorityName="Huber" authorityYear="2012" box="[1039,1149,1415,1437]" class="Arachnida" family="Pholcidae" genus="Smeringopus" kingdom="Animalia" order="Araneae" pageId="114" pageNumber="115" phylum="Arthropoda" rank="species" species="principe">
<emphasis id="B926D836FFF71771FBF20ED03E57FA69" box="[1039,1149,1415,1437]" italics="true" pageId="114" pageNumber="115">S. principe</emphasis>
</taxonomicName>
(702704, 707708), and
<taxonomicName id="4C527FA7FFF71771FA740ED03AD2FA48" authorityName="Huber" authorityYear="2012" class="Arachnida" family="Pholcidae" genus="Smeringopus" kingdom="Animalia" order="Araneae" pageId="114" pageNumber="115" phylum="Arthropoda" rank="species" species="mayombe">
<emphasis id="B926D836FFF71771FA740ED03AD2FA48" italics="true" pageId="114" pageNumber="115">S. mayombe</emphasis>
</taxonomicName>
(701). 699, 701, 703. Males, dorsal views. 700, 702. Male prosomata, oblique frontal views. 704. Female, ventral view. 705708. Left male palps, prolateral and retrolateral views.
</paragraph>
</caption>
<paragraph id="8BED0424FFF71771FF3A0D503F2FF807" blockId="114.[151,1436,1543,2035]" pageId="114" pageNumber="115">
Male (Palha Plantation). Total body length 5.0, carapace width 1.5. Leg 1: 45.3 (11.9 + 0.5 + 10.8 + 20.1 + 2.0), tibia 2: 7.2, tibia 3: 5.1, tibia 4: 7.2; tibia 1 L/d: 64. Habitus as in
<figureCitation id="136918A1FFF71771FC1A0D7C3E61F9B7" box="[999,1099,1579,1603]" captionStart="FIGURES 699708" captionStartId="114.[151,263,1415,1438]" captionTargetBox="[156,1436,153,1388]" captionTargetId="figure-308@114.[151,1436,153,1401]" captionTargetPageId="114" captionText="FIGURES 699708. Smeringopus thomensis group. S. thomensis (699700, 705706), S. principe (702704, 707708), and S. mayombe (701). 699, 701, 703. Males, dorsal views. 700, 702. Male prosomata, oblique frontal views. 704. Female, ventral view. 705708. Left male palps, prolateral and retrolateral views." figureDoi="http://doi.org/10.5281/zenodo.6418204" httpUri="https://zenodo.org/record/6418204/files/figure.png" pageId="114" pageNumber="115">Fig. 699</figureCitation>
. Carapace ochre-yellow with median and lateral dark pattern, clypeus distal half darker, without pair of lines, sternum dark brown with some small light spots, legs ochre-yellow, femora and tibiae with dark subdistal rings and light tips, abdomen grey with distinctive dorsal pattern, ventrally with dark anterior and posterior plates and three indistinct bands in median section. Distance PME-PME 125 µm, diameter PME 150 µm, distance PME-ALE 55 µm, distance AME-AME 30 µm, diameter AME 120 µm. Ocular area slightly elevated, secondary eyes with small dark pseudo-lenses; deep thoracic pit. Chelicerae with pair of small distal apophyses (
<figureCitation id="136918A1FFF71771FCC00C5439B4F8EF" box="[829,926,1795,1819]" captionStart="FIGURES 719724" captionStartId="116.[151,257,1954,1975]" captionTargetBox="[300,1299,910,1935]" captionTargetId="figure-85@116.[288,1303,900,1945]" captionTargetPageId="116" captionText="FIGURES 719724. Smeringopus thomensis. 719. Left cymbium and procursus, retrolateral view. 720. Left procursus, prolateral view. 721. Male chelicerae, frontal view. 722723. Left embolus, prolateral and dorsal (slightly distal) views. 724. Cleared female genitalia, dorsal view. Scale lines: 0.3 mm." figureDoi="http://doi.org/10.5281/zenodo.6415830" httpUri="https://zenodo.org/record/6415830/files/figure.png" pageId="114" pageNumber="115">Fig. 721</figureCitation>
), each provided with one modified hair at tip (
<figureCitation id="136918A1FFF71771FF620C703B2FF8CB" box="[159,261,1831,1855]" captionStart="FIGURES 732744" captionStartId="119.[151,263,1718,1741]" captionTargetBox="[151,1436,155,1695]" captionTargetId="figure-181@119.[151,1436,155,1695]" captionTargetPageId="119" captionText="FIGURES 732744. Smeringopus thomensis (732740) and S. principe (741744). 732. Left male palpal femur, retrolateral view. 733. Left procursus tip, prolateral view. 734. Male cheliceral apophysis. 735. Left embolus, distal view. 736. Male gonopore. 737. Female ALS. 738. Epigynum. 739. Cleared female genitalia, dorsal view. 740. Detail of pore plate. 741. Left embolus, distal view. 742. Left procursus tip, prolateral view. 743. Cleared female genitalia, dorsal view. 744. Detail of pore plate. Scale lines: 10 µm (734, 737), 20 µm (740), 30 µm (744), 80 µm (733, 735736), 100 µm (739, 741742), 200 µm (732, 738, 743)." figureDoi="http://doi.org/10.5281/zenodo.6415834" httpUri="https://zenodo.org/record/6415834/files/figure.png" pageId="114" pageNumber="115">Fig. 734</figureCitation>
). Palps as in
<figureCitation id="136918A1FFF71771FE5E0C70385DF8CB" box="[419,631,1831,1855]" captionStart="FIGURES 699708" captionStartId="114.[151,263,1415,1438]" captionTargetBox="[156,1436,153,1388]" captionTargetId="figure-308@114.[151,1436,153,1401]" captionTargetPageId="114" captionText="FIGURES 699708. Smeringopus thomensis group. S. thomensis (699700, 705706), S. principe (702704, 707708), and S. mayombe (701). 699, 701, 703. Males, dorsal views. 700, 702. Male prosomata, oblique frontal views. 704. Female, ventral view. 705708. Left male palps, prolateral and retrolateral views." figureDoi="http://doi.org/10.5281/zenodo.6418204" httpUri="https://zenodo.org/record/6418204/files/figure.png" pageId="114" pageNumber="115">Figs. 705 and 706</figureCitation>
, coxa with distinct retrolateral apophysis, trochanter barely modified, femur with deep retrolateral furrow, proximal rim distinct (
<figureCitation id="136918A1FFF71771FCBD0C1C398EF897" box="[832,932,1867,1891]" captionStart="FIGURES 732744" captionStartId="119.[151,263,1718,1741]" captionTargetBox="[151,1436,155,1695]" captionTargetId="figure-181@119.[151,1436,155,1695]" captionTargetPageId="119" captionText="FIGURES 732744. Smeringopus thomensis (732740) and S. principe (741744). 732. Left male palpal femur, retrolateral view. 733. Left procursus tip, prolateral view. 734. Male cheliceral apophysis. 735. Left embolus, distal view. 736. Male gonopore. 737. Female ALS. 738. Epigynum. 739. Cleared female genitalia, dorsal view. 740. Detail of pore plate. 741. Left embolus, distal view. 742. Left procursus tip, prolateral view. 743. Cleared female genitalia, dorsal view. 744. Detail of pore plate. Scale lines: 10 µm (734, 737), 20 µm (740), 30 µm (744), 80 µm (733, 735736), 100 µm (739, 741742), 200 µm (732, 738, 743)." figureDoi="http://doi.org/10.5281/zenodo.6415834" httpUri="https://zenodo.org/record/6415834/files/figure.png" pageId="114" pageNumber="115">Fig. 732</figureCitation>
), procursus with distinctive distal processes (
<figureCitation id="136918A1FFF71771FF620C383B66F873" box="[159,332,1903,1927]" captionStart="FIGURES 719724" captionStartId="116.[151,257,1954,1975]" captionTargetBox="[300,1299,910,1935]" captionTargetId="figure-85@116.[288,1303,900,1945]" captionTargetPageId="116" captionText="FIGURES 719724. Smeringopus thomensis. 719. Left cymbium and procursus, retrolateral view. 720. Left procursus, prolateral view. 721. Male chelicerae, frontal view. 722723. Left embolus, prolateral and dorsal (slightly distal) views. 724. Cleared female genitalia, dorsal view. Scale lines: 0.3 mm." figureDoi="http://doi.org/10.5281/zenodo.6415830" httpUri="https://zenodo.org/record/6415830/files/figure.png" pageId="114" pageNumber="115">Figs. 719, 720</figureCitation>
), bulb with complex embolus with distinctive rounded dorsal processes (
<figureCitation id="136918A1FFF71771FB5F0C383F65F873" box="[1186,1359,1903,1927]" captionStart="FIGURES 719724" captionStartId="116.[151,257,1954,1975]" captionTargetBox="[300,1299,910,1935]" captionTargetId="figure-85@116.[288,1303,900,1945]" captionTargetPageId="116" captionText="FIGURES 719724. Smeringopus thomensis. 719. Left cymbium and procursus, retrolateral view. 720. Left procursus, prolateral view. 721. Male chelicerae, frontal view. 722723. Left embolus, prolateral and dorsal (slightly distal) views. 724. Cleared female genitalia, dorsal view. Scale lines: 0.3 mm." figureDoi="http://doi.org/10.5281/zenodo.6415830" httpUri="https://zenodo.org/record/6415830/files/figure.png" pageId="114" pageNumber="115">Figs. 722, 723</figureCitation>
). Legs without spines, few vertical hairs, without curved hairs, retrolateral trichobothrium on tibia 1 at 2%; prolateral trichobothrium present on tibia 1. Gonopore with two epiandrous spigots (
<figureCitation id="136918A1FFF71771FC150CE03E66F83B" box="[1000,1100,1975,1999]" captionStart="FIGURES 732744" captionStartId="119.[151,263,1718,1741]" captionTargetBox="[151,1436,155,1695]" captionTargetId="figure-181@119.[151,1436,155,1695]" captionTargetPageId="119" captionText="FIGURES 732744. Smeringopus thomensis (732740) and S. principe (741744). 732. Left male palpal femur, retrolateral view. 733. Left procursus tip, prolateral view. 734. Male cheliceral apophysis. 735. Left embolus, distal view. 736. Male gonopore. 737. Female ALS. 738. Epigynum. 739. Cleared female genitalia, dorsal view. 740. Detail of pore plate. 741. Left embolus, distal view. 742. Left procursus tip, prolateral view. 743. Cleared female genitalia, dorsal view. 744. Detail of pore plate. Scale lines: 10 µm (734, 737), 20 µm (740), 30 µm (744), 80 µm (733, 735736), 100 µm (739, 741742), 200 µm (732, 738, 743)." figureDoi="http://doi.org/10.5281/zenodo.6415834" httpUri="https://zenodo.org/record/6415834/files/figure.png" pageId="114" pageNumber="115">Fig. 736</figureCitation>
); ALS with one widened and one pointed spigot each, other spigots apparently reduced to two tiny remnants (cf. female,
<figureCitation id="136918A1FFF71771FB6A0C8C3ED2F807" box="[1175,1272,2011,2035]" captionStart="FIGURES 732744" captionStartId="119.[151,263,1718,1741]" captionTargetBox="[151,1436,155,1695]" captionTargetId="figure-181@119.[151,1436,155,1695]" captionTargetPageId="119" captionText="FIGURES 732744. Smeringopus thomensis (732740) and S. principe (741744). 732. Left male palpal femur, retrolateral view. 733. Left procursus tip, prolateral view. 734. Male cheliceral apophysis. 735. Left embolus, distal view. 736. Male gonopore. 737. Female ALS. 738. Epigynum. 739. Cleared female genitalia, dorsal view. 740. Detail of pore plate. 741. Left embolus, distal view. 742. Left procursus tip, prolateral view. 743. Cleared female genitalia, dorsal view. 744. Detail of pore plate. Scale lines: 10 µm (734, 737), 20 µm (740), 30 µm (744), 80 µm (733, 735736), 100 µm (739, 741742), 200 µm (732, 738, 743)." figureDoi="http://doi.org/10.5281/zenodo.6415834" httpUri="https://zenodo.org/record/6415834/files/figure.png" pageId="114" pageNumber="115">Fig. 737</figureCitation>
).
</paragraph>
<caption id="DF2D54ACFFF61770FF6A0E8039B9F9FF" ID-DOI="http://doi.org/10.5281/zenodo.6415826" ID-Zenodo-Dep="6415826" httpUri="https://zenodo.org/record/6415826/files/figure.png" pageId="115" pageNumber="116" startId="115.[151,263,1495,1518]" targetBox="[151,1436,191,1473]" targetPageId="115">
<paragraph id="8BED0424FFF61770FF6A0E8039B9F9FF" blockId="115.[151,1436,1495,1548]" pageId="115" pageNumber="116">
<emphasis id="B926D836FFF61770FF6A0E803B44FA1A" bold="true" box="[151,366,1495,1518]" pageId="115" pageNumber="116">FIGURES 709717.</emphasis>
<taxonomicName id="4C527FA7FFF61770FE8A0E803843FA19" authorityName="Simon" authorityYear="1907" box="[375,617,1495,1517]" class="Arachnida" family="Pholcidae" genus="Smeringopus" kingdom="Animalia" order="Araneae" pageId="115" pageNumber="116" phylum="Arthropoda" rank="species" species="thomensis">
<emphasis id="B926D836FFF61770FE8A0E803843FA19" box="[375,617,1495,1517]" italics="true" pageId="115" pageNumber="116">Smeringopus thomensis</emphasis>
</taxonomicName>
group. Epigyna, ventral views and cleared female genitalia, ventral and dorsal views. 709711.
<taxonomicName id="4C527FA7FFF61770FEBE0EA13BEBF9F8" authorityName="Simon" authorityYear="1907" box="[323,449,1526,1548]" class="Arachnida" family="Pholcidae" genus="Smeringopus" kingdom="Animalia" order="Araneae" pageId="115" pageNumber="116" phylum="Arthropoda" rank="species" species="thomensis">
<emphasis id="B926D836FFF61770FEBE0EA13BEBF9F8" box="[323,449,1526,1548]" italics="true" pageId="115" pageNumber="116">S. thomensis</emphasis>
</taxonomicName>
. 712714.
<taxonomicName id="4C527FA7FFF61770FDCE0EA1388BF9FF" authorityName="Huber" authorityYear="2012" box="[563,673,1526,1548]" class="Arachnida" family="Pholcidae" genus="Smeringopus" kingdom="Animalia" order="Araneae" pageId="115" pageNumber="116" phylum="Arthropoda" rank="species" species="principe">
<emphasis id="B926D836FFF61770FDCE0EA1388BF9FF" box="[563,673,1526,1548]" italics="true" pageId="115" pageNumber="116">S. principe</emphasis>
</taxonomicName>
. 715717.
<taxonomicName id="4C527FA7FFF61770FCEE0EA139A7F9F8" authorityName="Huber" authorityYear="2012" box="[787,909,1526,1548]" class="Arachnida" family="Pholcidae" genus="Smeringopus" kingdom="Animalia" order="Araneae" pageId="115" pageNumber="116" phylum="Arthropoda" rank="species" species="mayombe">
<emphasis id="B926D836FFF61770FCEE0EA139A7F9F8" box="[787,909,1526,1548]" italics="true" pageId="115" pageNumber="116">S. mayombe</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="8BED0424FFF61770FF3A0D1838D6F993" blockId="115.[151,1436,1615,1983]" box="[199,764,1615,1639]" pageId="115" pageNumber="116">
Variation. Tibia
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3 other males: 8.9, 10.1, 10.7.
</paragraph>
<paragraph id="8BED0424FFF61770FF3A0D243AD2F927" blockId="115.[151,1436,1615,1983]" pageId="115" pageNumber="116">
Female. In general similar to male; tibia
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<specimenCount id="9D54CFADFFF61770FD1F0D24397DF97F" box="[738,855,1651,1675]" pageId="115" pageNumber="116" type="female">5 females</specimenCount>
: 8.49.7 (mean 9.1). Epigynum a simple, roughly triangular plate with pockets (
<figureCitation id="136918A1FFF61770FE160DC038BEF95B" box="[491,660,1687,1711]" captionStart="FIGURES 709717" captionStartId="115.[151,263,1495,1518]" captionTargetBox="[151,1436,191,1473]" captionTargetId="figure-188@115.[151,1436,191,1473]" captionTargetPageId="115" captionText="FIGURES 709717. Smeringopus thomensis group. Epigyna, ventral views and cleared female genitalia, ventral and dorsal views. 709711. S. thomensis. 712714. S. principe. 715717. S. mayombe." figureDoi="http://doi.org/10.5281/zenodo.6415826" httpUri="https://zenodo.org/record/6415826/files/figure.png" pageId="115" pageNumber="116">Figs. 709, 710</figureCitation>
), posterior plate semicircular; internal genitalia as in
<figureCitation id="136918A1FFF61770FB0E0DC03F74F95B" box="[1267,1374,1687,1711]" captionStart="FIGURES 709717" captionStartId="115.[151,263,1495,1518]" captionTargetBox="[151,1436,191,1473]" captionTargetId="figure-188@115.[151,1436,191,1473]" captionTargetPageId="115" captionText="FIGURES 709717. Smeringopus thomensis group. Epigyna, ventral views and cleared female genitalia, ventral and dorsal views. 709711. S. thomensis. 712714. S. principe. 715717. S. mayombe." figureDoi="http://doi.org/10.5281/zenodo.6415826" httpUri="https://zenodo.org/record/6415826/files/figure.png" pageId="115" pageNumber="116">Figs. 711</figureCitation>
,
<figureCitation id="136918A1FFF61770FA960DC03FB2F95B" box="[1387,1432,1687,1711]" captionStart="FIGURES 719724" captionStartId="116.[151,257,1954,1975]" captionTargetBox="[300,1299,910,1935]" captionTargetId="figure-85@116.[288,1303,900,1945]" captionTargetPageId="116" captionText="FIGURES 719724. Smeringopus thomensis. 719. Left cymbium and procursus, retrolateral view. 720. Left procursus, prolateral view. 721. Male chelicerae, frontal view. 722723. Left embolus, prolateral and dorsal (slightly distal) views. 724. Cleared female genitalia, dorsal view. Scale lines: 0.3 mm." figureDoi="http://doi.org/10.5281/zenodo.6415830" httpUri="https://zenodo.org/record/6415830/files/figure.png" pageId="115" pageNumber="116">724</figureCitation>
, and 739.
</paragraph>
<paragraph id="8BED0424FFF61770FF3A0D88394CF903" blockId="115.[151,1436,1615,1983]" box="[199,870,1759,1783]" pageId="115" pageNumber="116">
Distribution. Known from
<collectingCountry id="F34544B4FFF61770FE0E0D883884F903" box="[499,686,1759,1783]" name="Sao Tome Island" pageId="115" pageNumber="116">São Tomé Island</collectingCountry>
only (
<figureCitation id="136918A1FFF61770FD0A0D883973F903" box="[759,857,1759,1783]" captionStart="FIGURE 718" captionStartId="116.[151,244,819,840]" captionTargetBox="[151,1436,155,797]" captionTargetId="figure-81@116.[151,1436,155,797]" captionTargetPageId="116" captionText="FIGURE 718. Known distributions of the thomensis and roeweri groups. “?”: unknown type locality of S. roeweri in Rwanda. Squares: further undescribed species." figureDoi="http://doi.org/10.5281/zenodo.6415828" httpUri="https://zenodo.org/record/6415828/files/figure.png" pageId="115" pageNumber="116">Fig. 718</figureCitation>
).
</paragraph>
<paragraph id="8BED0424FFF61770FF3A0C573B2FF84B" blockId="115.[151,1436,1615,1983]" pageId="115" pageNumber="116">
<collectingMunicipality id="6B899E5EFFF61770FF3A0C573B0CF8EC" box="[199,294,1792,1816]" pageId="115" pageNumber="116">Material</collectingMunicipality>
examined.
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<collectingCountry id="F34544B4FFF61770FE440C573906F8EC" box="[441,812,1792,1816]" name="Sao Tome and Principe" pageId="115" pageNumber="116">SÃO TOMÉ AND PRÍNCIPE</collectingCountry>
:
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</emphasis>
:
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near
<collectingRegion id="4996CAC6FFF61770FB260C573F7EF8EC" box="[1243,1364,1792,1816]" country="Sao Tome and Principe" name="Sao Tome" pageId="115" pageNumber="116">São Tomé</collectingRegion>
town (
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,
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),
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<collectingDate id="EFA8DB0CFFF61770FE610C763838F8CD" box="[412,530,1825,1849]" pageId="115" pageNumber="116" value="2001-04-12">12.iv.2001</collectingDate>
</date>
(
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),
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<specimenCount id="9D54CFADFFF61770FCF10C763905F8CD" box="[780,815,1825,1849]" pageId="115" pageNumber="116" type="female">5♀</specimenCount>
in
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</materialsCitation>
.
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(
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,
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), in culvert along roadside,
<date id="FFEC22E4FFF61770FE570C14380AF8AF" box="[426,544,1859,1883]" pageId="115" pageNumber="116" value="2001-04-27">
<collectingDate id="EFA8DB0CFFF61770FE570C14380AF8AF" box="[426,544,1859,1883]" pageId="115" pageNumber="116" value="2001-04-27">27.iv.2001</collectingDate>
</date>
(
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),
<specimenCount id="9D54CFADFFF61770FD1A0C143920F8AF" box="[743,778,1859,1883]" pageId="115" pageNumber="116" type="female">1♀</specimenCount>
3 juvs in
<collectionCode id="ED439CE1FFF61770FC740C1439E9F8AF" box="[905,963,1859,1883]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="115" pageNumber="116" type="Museum">CAS</collectionCode>
</materialsCitation>
.
<materialsCitation id="3B3A0E79FFF61770FC2F0C1439D6F888" ID-GBIF-Occurrence="3747820649" collectingDate="2001-04-13" collectionCode="CAS" collectorName="C. E. Griswold" country="Sao Tome and Principe" elevation="220" latitude="0.36499998" location="Rio de Oro Plantation" longLatPrecision="130" longitude="6.645" municipality="Augustino Neto" pageId="115" pageNumber="116" specimenCount="1" specimenCount-female="1" stateProvince="Sao Tome">
<collectingMunicipality id="6B899E5EFFF61770FC2F0C143EA1F8AF" box="[978,1163,1859,1883]" pageId="115" pageNumber="116">Augustino Neto</collectingMunicipality>
,
<location id="8E8D52FFFFF61770FB670C143FB1F8AF" LSID="urn:lsid:plazi:treatment:03FBB532FFF41770FF6A0D6A3B2FF84B:8E8D52FFFFF61770FB670C143FB1F8AF" box="[1178,1435,1859,1883]" country="Sao Tome and Principe" latitude="0.36499998" longLatPrecision="130" longitude="6.645" municipality="Augustino Neto" name="Rio de Oro Plantation" pageId="115" pageNumber="116" stateProvince="Sao Tome">Rio de Oro Plantation</location>
(
<geoCoordinate id="EE6662E3FFF61770FF620C333B20F888" box="[159,266,1892,1916]" degrees="0" direction="north" minutes="21.9" orientation="latitude" pageId="115" pageNumber="116" precision="92" value="0.36499998">0°21.9N</geoCoordinate>
,
<geoCoordinate id="EE6662E3FFF61770FEE40C333BABF888" box="[281,385,1892,1916]" degrees="6" direction="east" minutes="38.7" orientation="longitude" pageId="115" pageNumber="116" precision="92" value="6.645">6°38.7E</geoCoordinate>
),
<elevation id="007FE317FFF61770FE6A0C33380AF888" box="[407,544,1892,1916]" metricMagnitude="2" metricUnit="m" metricValue="2.2" pageId="115" pageNumber="116" unit="m" value="220.0">
<quantity id="4CAAA9C1FFF61770FE6A0C333BCBF888" box="[407,481,1892,1916]" metricMagnitude="2" metricUnit="m" metricValue="2.2" pageId="115" pageNumber="116" unit="m" value="220.0">220 m</quantity>
a.s.l.
</elevation>
,
<date id="FFEC22E4FFF61770FDD20C33388FF888" box="[559,677,1892,1916]" pageId="115" pageNumber="116" value="2001-04-13">
<collectingDate id="EFA8DB0CFFF61770FDD20C33388FF888" box="[559,677,1892,1916]" pageId="115" pageNumber="116" value="2001-04-13">13.iv.2001</collectingDate>
</date>
(
<collectorName id="26A761F2FFF61770FD450C33394BF888" box="[696,865,1892,1916]" pageId="115" pageNumber="116">C.E. Griswold</collectorName>
),
<specimenCount id="9D54CFADFFF61770FC8B0C3339B3F888" box="[886,921,1892,1916]" pageId="115" pageNumber="116" type="female">1♀</specimenCount>
in
<collectionCode id="ED439CE1FFF61770FC3F0C3339D6F888" box="[962,1020,1892,1916]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="115" pageNumber="116" type="Museum">CAS</collectionCode>
</materialsCitation>
.
<materialsCitation id="3B3A0E79FFF61770FBF40C333930F86A" ID-GBIF-Occurrence="3747820850" collectingDate="1999-11-01" collectionCode="MRAC" collectorName="Tavernier" country="Sao Tome and Principe" location="Sao Tome" pageId="115" pageNumber="116" specimenCount="2" specimenCount-male="1" stateProvince="Sao Tome">
<collectingRegion id="4996CAC6FFF61770FBF40C333E55F888" box="[1033,1151,1892,1916]" country="Sao Tome and Principe" name="Sao Tome" pageId="115" pageNumber="116">São Tomé</collectingRegion>
, no further locality data,
<date id="FFEC22E4FFF61770FF6A0CD13B2AF86A" box="[151,256,1926,1950]" pageId="115" pageNumber="116" value="1999-11-01">
<collectingDate id="EFA8DB0CFFF61770FF6A0CD13B2AF86A" box="[151,256,1926,1950]" pageId="115" pageNumber="116" value="1999-11-01">1.xi.1999</collectingDate>
</date>
(W.
<collectorName id="26A761F2FFF61770FEC60CD13B81F86A" box="[315,427,1926,1950]" pageId="115" pageNumber="116">Tavernier</collectorName>
),
<specimenCount id="9D54CFADFFF61770FE3D0CD13BC9F869" box="[448,483,1926,1950]" pageId="115" pageNumber="116" type="male">1♂</specimenCount>
<specimenCount id="9D54CFADFFF61770FE110CD13807F86A" box="[492,557,1926,1950]" pageId="115" pageNumber="116" type="generic">1 juv.</specimenCount>
in
<collectionCode id="ED439CE1FFF61770FDA80CD1388DF86A" box="[597,679,1926,1950]" country="Belgium" httpUri="http://grbio.org/cool/z14b-bb31" name="Musée Royal de lAfrique Centrale" pageId="115" pageNumber="116" type="Museum">MRAC</collectionCode>
(209235)
</materialsCitation>
;
<materialsCitation id="3B3A0E79FFF61770FCD50CD13B2FF84B" ID-GBIF-Occurrence="3747820892" collectingDate="1999-11-17" collectionCode="MRAC" collectorName="Tavernier" country="Sao Tome and Principe" location="Sao Tome" pageId="115" pageNumber="116" specimenCount="2" specimenCount-male="1" stateProvince="Sao Tome">
same data but garden,
<date id="FFEC22E4FFF61770FBD10CD13E8CF86A" box="[1068,1190,1926,1950]" pageId="115" pageNumber="116" value="1999-11-17">
<collectingDate id="EFA8DB0CFFF61770FBD10CD13E8CF86A" box="[1068,1190,1926,1950]" pageId="115" pageNumber="116" value="1999-11-17">17.xi.1999</collectingDate>
</date>
,
<specimenCount id="9D54CFADFFF61770FB490CD13EF2F869" box="[1204,1240,1926,1950]" pageId="115" pageNumber="116" type="male">1♂</specimenCount>
<specimenCount id="9D54CFADFFF61770FB1C0CD13F08F86A" box="[1249,1314,1926,1950]" pageId="115" pageNumber="116" type="generic">1 juv.</specimenCount>
in
<collectionCode id="ED439CE1FFF61770FAB40CD13FB1F86A" box="[1353,1435,1926,1950]" country="Belgium" httpUri="http://grbio.org/cool/z14b-bb31" name="Musée Royal de lAfrique Centrale" pageId="115" pageNumber="116" type="Museum">MRAC</collectionCode>
(209256).
</materialsCitation>
</paragraph>
</subSubSection>
</treatment>
</document>