treatments-xml/data/03/FB/A4/03FBA436FFCCFFA8B9C9F93EFDB9FE22.xml
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<document id="204E1F31A011382F19FF45F12565E1A2" ID-DOI="10.11646/zootaxa.4098.3.7" ID-GBIF-Dataset="11a55b63-d8bc-4f9e-a5bb-417152f7512e" ID-ISSN="1175-5326" ID-Zenodo-Dep="264774" ID-ZooBank="FB74AD01-13B0-4D39-A575-E871D68AB4C4" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460013227175" checkinUser="plazi" docAuthor="Miranda, Gustavo Silva De, Giupponi, Alessandro Ponce De Leão &amp; Wizen, Gil" docDate="2016" docId="03FBA436FFCCFFA8B9C9F93EFDB9FE22" docLanguage="en" docName="zootaxa.4098.3.7.pdf.imf" docOrigin="Zootaxa 4098 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Charinus reddelli Miranda, Giupponi &amp; Wizen, 2016, sp. nov." docType="treatment" docUuid="F47C0435-D972-4F18-8C06-B808670E6378" docUuidSource="ZooBank" docVersion="10" lastPageNumber="554" masterDocId="FFC2DC4EFFC9FFA1B95EFFD2FFC2FFAA" masterDocTitle="Two new species of whip spider (Amblypygi): an epigean and a cave dwelling Charinus Simon, 1892 from Belize" masterLastPageNumber="559" masterPageNumber="545" pageNumber="550" updateTime="1698233481388" updateUser="plazi">
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<mods:title id="ADB54C28D13DA59B99CFD20BB5A4E609">Two new species of whip spider (Amblypygi): an epigean and a cave dwelling Charinus Simon, 1892 from Belize</mods:title>
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<mods:namePart id="46C29E2ED6484E3129AC0652AC7C6F8D">Miranda, Gustavo Silva De</mods:namePart>
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<mods:namePart id="157322D5CB3D63C50F4CA4278D0ED2EC">Giupponi, Alessandro Ponce De Leão</mods:namePart>
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<mods:namePart id="4D15893F070161C06E08552BF402FF71">Wizen, Gil</mods:namePart>
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<mods:title id="AA48E663CDE58E217F6D1DF429F45216">Zootaxa</mods:title>
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<mods:date id="12021885988D3AB2B1E33DA499563881">2016</mods:date>
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<treatment id="03FBA436FFCCFFA8B9C9F93EFDB9FE22" ID-DOI="http://doi.org/10.5281/zenodo.6086690" ID-GBIF-Taxon="119381578" ID-Zenodo-Dep="6086690" LSID="urn:lsid:zoobank.org:act:F47C0435-D972-4F18-8C06-B808670E6378" httpUri="http://treatment.plazi.org/id/03FBA436FFCCFFA8B9C9F93EFDB9FE22" lastPageId="9" lastPageNumber="554" pageId="5" pageNumber="550">
<subSubSection id="C34846ABFFCCFFA4B9C9F93EFC14F882" pageId="5" pageNumber="550" type="nomenclature">
<paragraph id="8BED1520FFCCFFA4B9C9F93EFE12F8AC" blockId="5.[151,982,1772,1832]" box="[151,464,1772,1798]" pageId="5" pageNumber="550">
<heading id="D0A5A24CFFCCFFA4B9C9F93EFE12F8AC" bold="true" box="[151,464,1772,1798]" fontSize="11" level="1" pageId="5" pageNumber="550" reason="1">
<emphasis id="B926C932FFCCFFA4B9C9F93EFE12F8AC" bold="true" box="[151,464,1772,1798]" pageId="5" pageNumber="550">
<taxonomicName id="4C526EA3FFCCFFA4B9C9F93EFEA9F8AC" ID-CoL="TNW5" box="[151,363,1772,1798]" class="Arachnida" family="Charinidae" genus="Charinus" kingdom="Animalia" order="Amblypygi" pageId="5" pageNumber="550" phylum="Arthropoda" rank="species" species="reddelli" status="sp. nov.">
<emphasis id="B926C932FFCCFFA4B9C9F93EFEA9F8AC" bold="true" box="[151,363,1772,1798]" italics="true" pageId="5" pageNumber="550">Charinus reddelli</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2157449FFCCFFA4B82DF93EFE12F8AC" box="[371,464,1772,1798]" pageId="5" pageNumber="550" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BED1520FFCCFFA4B9C9F8DDFC14F882" blockId="5.[151,982,1772,1832]" box="[151,982,1807,1832]" pageId="5" pageNumber="550">
<figureCitation id="136909A5FFCCFFA4B9C9F8DDFF22F882" box="[151,224,1807,1832]" captionStart="FIGURE 1" captionStartId="3.[151,250,1623,1645]" captionTargetBox="[151,1436,531,1601]" captionTargetId="figure@3.[151,1436,517,1602]" captionTargetPageId="3" captionText="FIGURE 1. Habitus (A C) and sternum (D F) of Charinus belizensis sp. nov. (holotype: A, D) and C. reddelli sp. nov. (holotype: B and E; paratype from Footprint cave: C and F). The distinctly short tristosternum of the holotype (E) is an abnormality and is not found in any other specimen of the species. Scale bars: 1 mm." httpUri="https://zenodo.org/record/264775/files/figure.png" pageId="5" pageNumber="550">Figs. 1</figureCitation>
B, C, E, F; 2CH; 3B, D; 4B, D; 5CF, H; 6B, C, E, F, H; 7B, C; 8.
</paragraph>
</subSubSection>
<subSubSection id="C34846ABFFCCFFA4B9C9F885FE6AF876" pageId="5" pageNumber="550" type="diagnosis">
<paragraph id="8BED1520FFCCFFA4B9C9F885FE6AF876" blockId="5.[151,1437,1879,2012]" pageId="5" pageNumber="550">
<emphasis id="B926C932FFCCFFA4B9C9F885FEC8F8DA" bold="true" box="[151,266,1879,1904]" pageId="5" pageNumber="550">Diagnosis</emphasis>
: median eyes and tubercle absent, large size (carapace mean width: 3.77; pedipalp femur mean length: 2.61; femur I mean length: 8.63), tibia I with 23 articles, tarsus I with 41 articles, proximal article of tarsus I 1.4 to 1.9 times longer than the second, cheliceral claw with four teeth, and cusps of the bifid tooth of the basal segment of the chelicerae as p=d.
</paragraph>
</subSubSection>
<caption id="DF2D45A8FFCFFFA7B9C9FD94FCBDFD30" httpUri="https://zenodo.org/record/264778/files/figure.png" pageId="6" pageNumber="551" targetBox="[151,1436,193,560]" targetPageId="6">
<paragraph id="8BED1520FFCFFFA7B9C9FD94FCBDFD30" blockId="6.[151,1435,582,666]" pageId="6" pageNumber="551">
<emphasis id="B926C932FFCFFFA7B9C9FD94FECFFDF6" bold="true" box="[151,269,582,604]" pageId="6" pageNumber="551">FIGURE 4</emphasis>
. Part of tibia and tarsus I and last segments of tarsus IV, all lateral view. Tibia and tarsus I of
<taxonomicName id="4C526EA3FFCFFFA7BD9AFD95FA87FDF6" box="[1220,1349,583,604]" class="Arachnida" family="Charinidae" genus="Charinus" kingdom="Animalia" order="Amblypygi" pageId="6" pageNumber="551" phylum="Arthropoda" rank="species" species="belizensis" status="sp. nov.">
<emphasis id="B926C932FFCFFFA7BD9AFD95FA87FDF6" box="[1220,1349,583,604]" italics="true" pageId="6" pageNumber="551">C. belizensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2157449FFCFFFA7BC15FD95FA59FDF6" box="[1355,1435,583,604]" pageId="6" pageNumber="551" rank="species">
<emphasis id="B926C932FFCFFFA7BC15FD95FA59FDF6" bold="true" box="[1355,1435,583,604]" pageId="6" pageNumber="551">sp. nov.</emphasis>
</taxonomicNameLabel>
(A) and
<taxonomicName id="4C526EA3FFCFFFA7B9AEFDB7FE9CFDD0" box="[240,350,613,634]" class="Arachnida" family="Charinidae" genus="Charinus" kingdom="Animalia" order="Amblypygi" pageId="6" pageNumber="551" phylum="Arthropoda" rank="species" species="reddelli" status="sp. nov.">
<emphasis id="B926C932FFCFFFA7B9AEFDB7FE9CFDD0" box="[240,350,613,634]" italics="true" pageId="6" pageNumber="551">C. reddelli</emphasis>
</taxonomicName>
<emphasis id="B926C932FFCFFFA7B839FDB7FE78FDD0" bold="true" box="[359,442,613,634]" pageId="6" pageNumber="551">
<taxonomicNameLabel id="A2157449FFCFFFA7B839FDB7FE78FDD0" box="[359,442,613,634]" pageId="6" pageNumber="551" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(B). Tarsus IV of
<taxonomicName id="4C526EA3FFCFFFA7BBDEFDB7FCC1FDD0" box="[640,771,613,634]" class="Arachnida" family="Charinidae" genus="Charinus" kingdom="Animalia" order="Amblypygi" pageId="6" pageNumber="551" phylum="Arthropoda" rank="species" species="belizensis" status="sp. nov.">
<emphasis id="B926C932FFCFFFA7BBDEFDB7FCC1FDD0" box="[640,771,613,634]" italics="true" pageId="6" pageNumber="551">C. belizensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2157449FFCFFFA7BA55FDB7FC9CFDD0" box="[779,862,613,634]" pageId="6" pageNumber="551" rank="species">
<emphasis id="B926C932FFCFFFA7BA55FDB7FC9CFDD0" bold="true" box="[779,862,613,634]" pageId="6" pageNumber="551">sp. nov.</emphasis>
</taxonomicNameLabel>
(C) and
<taxonomicName id="4C526EA3FFCFFFA7BA9EFDB7FBECFDD0" box="[960,1070,613,634]" class="Arachnida" family="Charinidae" genus="Charinus" kingdom="Animalia" order="Amblypygi" pageId="6" pageNumber="551" phylum="Arthropoda" rank="species" species="reddelli" status="sp. nov.">
<emphasis id="B926C932FFCFFFA7BA9EFDB7FBECFDD0" box="[960,1070,613,634]" italics="true" pageId="6" pageNumber="551">C. reddelli</emphasis>
</taxonomicName>
<emphasis id="B926C932FFCFFFA7BD68FDB7FB4BFDD0" bold="true" box="[1078,1161,613,634]" pageId="6" pageNumber="551">
<taxonomicNameLabel id="A2157449FFCFFFA7BD68FDB7FB4BFDD0" box="[1078,1161,613,634]" pageId="6" pageNumber="551" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(D). Arrows points to the membranose ring in the second segment of the tarsus. Scale bars: 0.5mm.
</paragraph>
</caption>
<subSubSection id="C34846ABFFCFFFA7B999FD14FB6FFCA9" pageId="6" pageNumber="551" type="etymology">
<paragraph id="8BED1520FFCFFFA7B999FD14FB6FFCA9" blockId="6.[151,1437,710,2031]" pageId="6" pageNumber="551">
<emphasis id="B926C932FFCFFFA7B999FD14FE84FD75" bold="true" box="[199,326,710,735]" pageId="6" pageNumber="551">Etymology</emphasis>
. the epithet is a patronym to Dr. James R. Reddel for the first finding of
<taxonomicName id="4C526EA3FFCFFFA7BD23FD14FB27FD74" box="[1149,1253,710,734]" class="Arachnida" family="Charinidae" genus="Charinus" kingdom="Animalia" order="Amblypygi" pageId="6" pageNumber="551" phylum="Arthropoda" rank="genus">
<emphasis id="B926C932FFCFFFA7BD23FD14FB27FD74" box="[1149,1253,710,734]" italics="true" pageId="6" pageNumber="551">Charinus</emphasis>
</taxonomicName>
in the Footprint cave and in Central
<collectingCountry id="F34555B0FFCFFFA7B829FD38FE18FCA9" box="[375,474,746,771]" name="United States of America" pageId="6" pageNumber="551">America</collectingCountry>
, and for his enormous contribution to the study of the arachnids.
</paragraph>
</subSubSection>
<subSubSection id="C34846ABFFCFFFA7B999FCDCFC5EFBC1" pageId="6" pageNumber="551" type="materials_examined">
<paragraph id="8BED1520FFCFFFA7B999FCDCFC5EFBC1" blockId="6.[151,1437,710,2031]" pageId="6" pageNumber="551">
<emphasis id="B926C932FFCFFFA7B999FCDCFEACFC8D" bold="true" box="[199,366,782,807]" pageId="6" pageNumber="551">
<typeStatus id="54E9AB82FFCFFFA7B999FCDCFEC0FC8D" box="[199,258,782,807]" pageId="6" pageNumber="551">Type</typeStatus>
material
</emphasis>
:
<emphasis id="B926C932FFCFFFA7B823FCDCFE28FC8D" bold="true" box="[381,490,782,807]" pageId="6" pageNumber="551">
<typeStatus id="54E9AB82FFCFFFA7B823FCDCFE28FC8D" box="[381,490,782,807]" pageId="6" pageNumber="551" type="holotype">Holotype</typeStatus>
</emphasis>
.
<collectingCountry id="F34555B0FFCFFFA7B8A6FCDDFD99FC8C" box="[504,603,783,806]" name="Belize" pageId="6" pageNumber="551">BELIZE</collectingCountry>
,
<collectingCountry id="F34555B0FFCFFFA7BB39FCDDFD2FFC8D" box="[615,749,783,807]" name="Armenia" pageId="6" pageNumber="551">ARMENIA</collectingCountry>
,
<collectionCode id="ED438DE5FFCFFFA7BBA6FCDDFC87FC8D" box="[760,837,783,807]" pageId="6" pageNumber="551">CAYO</collectionCode>
DISTRICT: Waterfall Cave (Actun Lubul Ha), Karst cave, in cave,
<quantity id="4CAAB8C5FFCFFFA7B867FCE0FEBBFCE1" box="[313,377,818,843]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="6" pageNumber="551" unit="m" value="500.0">500m</quantity>
from entrance, on walls and ceiling,
<date id="FFEC33E0FFCFFFA7BA7EFCE0FC68FCE1" box="[800,938,818,843]" pageId="6" pageNumber="551" value="2014-08-30">30.viii.2014</date>
, Wizen G.
<emphasis id="B926C932FFCFFFA7BD78FCE1FB8AFCE1" box="[1062,1096,819,843]" italics="true" pageId="6" pageNumber="551">leg</emphasis>
., 17,108611'N, -88,730833'W (
<specimenCount id="9D54DEA9FFCFFFA7B9FCFC85FECBFCC5" box="[162,265,854,879]" pageId="6" pageNumber="551" type="female">1 female</specimenCount>
, HUJINVAMB115).
<emphasis id="B926C932FFCFFFA7B8A0FC84FDB4FCC5" bold="true" box="[510,630,854,879]" pageId="6" pageNumber="551">
<typeStatus id="54E9AB82FFCFFFA7B8A0FC84FDB4FCC5" box="[510,630,854,879]" pageId="6" pageNumber="551" type="paratype">Paratypes</typeStatus>
</emphasis>
: Waterfall Cave (Actun Lubul Ha), Karst cave, in cave,
<quantity id="4CAAB8C5FFCFFFA7BC44FC84FA98FCC5" box="[1306,1370,854,879]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="6" pageNumber="551" unit="m" value="500.0">500m</quantity>
from entrance, on walls and ceiling,
<date id="FFEC33E0FFCFFFA7BB4FFCA8FD59FC39" box="[529,667,890,915]" pageId="6" pageNumber="551" value="2014-08-30">30.viii.2014</date>
, Wizen G.
<emphasis id="B926C932FFCFFFA7BA78FCA9FC8AFC39" box="[806,840,891,915]" italics="true" pageId="6" pageNumber="551">leg</emphasis>
., 17,108611'N, -88,730833'W (
<specimenCount id="9D54DEA9FFCFFFA7BDE1FCA9FAD3FC39" box="[1215,1297,890,915]" pageId="6" pageNumber="551" type="male">1 male</specimenCount>
,
<specimenCount id="9D54DEA9FFCFFFA7BC7CFCA9FA5AFC39" box="[1314,1432,890,915]" pageId="6" pageNumber="551" type="female">2 females</specimenCount>
, HUJINVAMB116); Waterfall Cave,
<date id="FFEC33E0FFCFFFA7BB71FC4CFD59FC1D" box="[559,667,926,951]" pageId="6" pageNumber="551" value="2014-09-03">3.ix.2014</date>
, Wizen G.
<emphasis id="B926C932FFCFFFA7BA4DFC4CFCF7FC1C" box="[787,821,926,950]" italics="true" pageId="6" pageNumber="551">leg</emphasis>
. (
<specimenCount id="9D54DEA9FFCFFFA7BA12FC4DFC7EFC1D" box="[844,956,926,951]" pageId="6" pageNumber="551" type="female">2 females</specimenCount>
,
<collectionCode id="ED438DE5FFCFFFA7BA98FC4DFBD3FC1C" box="[966,1041,927,950]" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="6" pageNumber="551">MNRJ</collectionCode>
9305); Footprint Cave, Karst cave, in cave,
<quantity id="4CAAB8C5FFCFFFA7B9ABFC11FEF6FC71" box="[245,308,963,987]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="6" pageNumber="551" unit="m" value="200.0">200m</quantity>
from entrance, on sand next to cave walls,
<date id="FFEC33E0FFCFFFA7BA47FC10FC51FC71" box="[793,915,962,987]" pageId="6" pageNumber="551" value="2014-09-07">07.ix.2014</date>
, Naskrecki P. and Wizen G.
<emphasis id="B926C932FFCFFFA7BD93FC11FB2DFC71" box="[1229,1263,963,987]" italics="true" pageId="6" pageNumber="551">leg</emphasis>
., 17,127500'N, -88,732222'W, (
<specimenCount id="9D54DEA9FFCFFFA7B811FC35FE71FC55" box="[335,435,998,1023]" pageId="6" pageNumber="551" type="female">1 female</specimenCount>
, HUJINVAMB112); Footprint Cave, Karst cave, in cave,
<quantity id="4CAAB8C5FFCFFFA7BD12FC35FB4EFC55" box="[1100,1164,999,1023]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="6" pageNumber="551" unit="m" value="200.0">200m</quantity>
from entrance, on sand next to cave walls,
<date id="FFEC33E0FFCFFFA7B8D0FBD8FDCBFB89" box="[398,521,1034,1059]" pageId="6" pageNumber="551" value="2014-09-02">02.ix.2014</date>
, Wizen G.
<emphasis id="B926C932FFCFFFA7BBC7FBD9FD79FB89" box="[665,699,1035,1059]" italics="true" pageId="6" pageNumber="551">leg</emphasis>
., 17,127500'N, -88,732222'W (
<specimenCount id="9D54DEA9FFCFFFA7BD68FBD9FB6CFB89" box="[1078,1198,1034,1059]" pageId="6" pageNumber="551" type="female">2 females</specimenCount>
, HUJINVAMB113); Footprint Cave, Karst cave, in cave,
<quantity id="4CAAB8C5FFCFFFA7BB62FBFDFDBEFBED" box="[572,636,1071,1095]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="6" pageNumber="551" unit="m" value="200.0">200m</quantity>
from entrance, on sand next to cave walls,
<date id="FFEC33E0FFCFFFA7BD2DFBFCFB2CFBED" box="[1139,1262,1070,1095]" pageId="6" pageNumber="551" value="2014-09-02">02.ix.2014</date>
, Wizen G.
<emphasis id="B926C932FFCFFFA7BC32FBFCFA4CFBEC" box="[1388,1422,1070,1094]" italics="true" pageId="6" pageNumber="551">leg</emphasis>
., 17,127500'N, -88,732222'W (
<specimenCount id="9D54DEA9FFCFFFA7B8BAFB81FD96FBC1" box="[484,596,1106,1131]" pageId="6" pageNumber="551" type="juvenile">2 juvenile</specimenCount>
females, HUJINVAMB114).
</paragraph>
</subSubSection>
<subSubSection id="C34846ABFFCFFFA8B999FBA4FDB9FE22" lastPageId="9" lastPageNumber="554" pageId="6" pageNumber="551" type="description">
<paragraph id="8BED1520FFCFFFA7B999FBA4FD01FB19" blockId="6.[151,1437,710,2031]" pageId="6" pageNumber="551">
<emphasis id="B926C932FFCFFFA7B999FBA4FD9CFB25" bold="true" box="[199,606,1142,1167]" pageId="6" pageNumber="551">
Description of the female
<typeStatus id="54E9AB82FFCFFFA7B8A6FBA4FD9CFB25" box="[504,606,1142,1167]" pageId="6" pageNumber="551" type="holotype">holotype</typeStatus>
</emphasis>
(variations found in the
<typeStatus id="54E9AB82FFCFFFA7BA2FFBA5FC1FFB25" box="[881,989,1143,1167]" pageId="6" pageNumber="551" type="paratype">paratypes</typeStatus>
are indicated in brackets; description of the chelicerae and gonopods based on
<typeStatus id="54E9AB82FFCFFFA7BB18FB49FD75FB19" box="[582,695,1179,1203]" pageId="6" pageNumber="551" type="paratype">paratypes</typeStatus>
):
</paragraph>
<paragraph id="8BED1520FFCFFFA7B999FB6CFA93FA05" blockId="6.[151,1437,710,2031]" pageId="6" pageNumber="551">
<emphasis id="B926C932FFCFFFA7B999FB6CFEF8FB7D" bold="true" box="[199,314,1214,1239]" pageId="6" pageNumber="551">Carapace</emphasis>
(
<figureCitation id="136909A5FFCFFFA7B817FB6CFE4AFB7D" box="[329,392,1214,1239]" captionStart="FIGURE 1" captionStartId="3.[151,250,1623,1645]" captionTargetBox="[151,1436,531,1601]" captionTargetId="figure@3.[151,1436,517,1602]" captionTargetPageId="3" captionText="FIGURE 1. Habitus (A C) and sternum (D F) of Charinus belizensis sp. nov. (holotype: A, D) and C. reddelli sp. nov. (holotype: B and E; paratype from Footprint cave: C and F). The distinctly short tristosternum of the holotype (E) is an abnormality and is not found in any other specimen of the species. Scale bars: 1 mm." httpUri="https://zenodo.org/record/264775/files/figure.png" pageId="6" pageNumber="551">Fig. 1</figureCitation>
B, C). Carapace flattened, wider than long (1.4 times), slightly bent downwards below lateral eyes; a thin median furrow reaches the fovea starting from the depression that replace the median eye and tubercle. Anterior margin straight, with six frontal setae. Frontal process large, triangular, not visible from above, with a rounded tip. Three pairs of shallow furrows in the lateral of the carapace, and a triangular and deep fovea. First pair of furrows placed just behind the lateral boss behind the lateral eyes; any of the furrows reaches the middle line. Median eyes and tubercle completely absent, a deep depression instead; no setae present in the depression. Lateral eyes well developed, pale, one large setae behind each triad; lenses directed upwards and slightly anteriorly.
</paragraph>
<paragraph id="8BED1520FFCFFFA7B999FA68FF2BF965" blockId="6.[151,1437,710,2031]" pageId="6" pageNumber="551">
<emphasis id="B926C932FFCFFFA7B999FA68FEECFA79" bold="true" box="[199,302,1466,1491]" pageId="6" pageNumber="551">Sternum</emphasis>
(
<figureCitation id="136909A5FFCFFFA7B81DFA68FE4AFA79" box="[323,392,1466,1491]" captionStart="FIGURE 1" captionStartId="3.[151,250,1623,1645]" captionTargetBox="[151,1436,531,1601]" captionTargetId="figure@3.[151,1436,517,1602]" captionTargetPageId="3" captionText="FIGURE 1. Habitus (A C) and sternum (D F) of Charinus belizensis sp. nov. (holotype: A, D) and C. reddelli sp. nov. (holotype: B and E; paratype from Footprint cave: C and F). The distinctly short tristosternum of the holotype (E) is an abnormality and is not found in any other specimen of the species. Scale bars: 1 mm." httpUri="https://zenodo.org/record/264775/files/figure.png" pageId="6" pageNumber="551">Fig. 1</figureCitation>
E, F): tetra-segmented, all pieces well sclerotized. Tritosternum with a round basis and projected anteriorly in a small blunt tubercle, reaching the base of the pedipalp coxae, with two apical, two median and several smaller ones on the base (the
<typeStatus id="54E9AB82FFCFFFA7BBDDF9D1FD2CF9B1" box="[643,750,1539,1563]" pageId="6" pageNumber="551" type="paratype">paratypes</typeStatus>
have a longer tritosternum and longer basal pair of setae). Middle piece (tetrasternum) in one convex piece, with a pair of large setae in its apex, and four small setae in its base. Third piece (pentasternum) formed by one convex piece, smaller than the middle piece, with two long setae at its top and with one small setae on its base. Sternites separated from each other by length of the third piece. Metasternum not paired (i.e., one single piece), with one pair of setae on an elevation at the posterior region of the plaque.
</paragraph>
<paragraph id="8BED1520FFCFFFA7B999F908FA97F959" blockId="6.[151,1437,710,2031]" box="[199,1365,1754,1779]" pageId="6" pageNumber="551">
<emphasis id="B926C932FFCFFFA7B999F908FEFEF959" bold="true" box="[199,316,1754,1779]" pageId="6" pageNumber="551">Abdomen</emphasis>
(
<figureCitation id="136909A5FFCFFFA7B815F908FE48F959" box="[331,394,1754,1779]" captionStart="FIGURE 1" captionStartId="3.[151,250,1623,1645]" captionTargetBox="[151,1436,531,1601]" captionTargetId="figure@3.[151,1436,517,1602]" captionTargetPageId="3" captionText="FIGURE 1. Habitus (A C) and sternum (D F) of Charinus belizensis sp. nov. (holotype: A, D) and C. reddelli sp. nov. (holotype: B and E; paratype from Footprint cave: C and F). The distinctly short tristosternum of the holotype (E) is an abnormality and is not found in any other specimen of the species. Scale bars: 1 mm." httpUri="https://zenodo.org/record/264775/files/figure.png" pageId="6" pageNumber="551">Fig. 1</figureCitation>
B, C): oblong, with almost indistinguishable punctuations. Ventral sacs not developed.
</paragraph>
<paragraph id="8BED1520FFCFFFA7B999F92CFD53F829" blockId="6.[151,1437,710,2031]" pageId="6" pageNumber="551">
<emphasis id="B926C932FFCFFFA7B999F92CFEF8F8BD" bold="true" box="[199,314,1790,1815]" pageId="6" pageNumber="551">Chelicera</emphasis>
(
<figureCitation id="136909A5FFCFFFA7B814F92CFE4FF8BD" box="[330,397,1790,1815]" captionStart="FIGURE 5" captionStartId="7.[151,250,1661,1683]" captionTargetBox="[191,1395,193,1639]" captionTargetId="figure@7.[191,1395,193,1639]" captionTargetPageId="7" captionText="FIGURE 5. Leg IV trichobothriotaxy and chelicerae of Charinus belizensis sp. nov. (A B, G) and C. reddeli sp. nov. (C F, H). A: left leg of C. belizensis sp. nov. male holotype (HUJINVAMB 117). B: left leg of C. belizensis sp. nov. female paratype (HUJINVAMB 118). C: left leg of C. reddelli sp. nov. female holotype (HUJINVAMB 115). D, E, F: left leg of C. reddelli sp. nov. female paratypes from Waterfall cave (HUJINVAMB 116) and Footprint Cave (HUJINVAMB 114, HUJINVAMB 113), prespectively. G: right chelicerae of C. belizensis sp. nov. (HUJINVAMB 118). H: right cheliceae of C. reddelli sp. nov. (HUJINVAMB 116). Scale bars: 1 mm." httpUri="https://zenodo.org/record/264779/files/figure.png" pageId="6" pageNumber="551">Fig. 5</figureCitation>
H): Cheliceral furrow with four internal teeth; first tooth (upper) bifid, proximal cusp of the same size as distal cusp. Third tooth slightly thinner and shorter than second tooth. Fourth tooth one third larger than the third. No tooth in the external row of the basal segment. Mesal face with a longitudinal row of seven (five in
<typeStatus id="54E9AB82FFCFFFA7B9EAF8B9FEE3F829" box="[180,289,1899,1923]" pageId="6" pageNumber="551" type="paratype">paratypes</typeStatus>
) setae. Claw with four denticles.
</paragraph>
<paragraph id="8BED1520FFCFFFA9B999F85CFECDFE7A" blockId="6.[151,1437,710,2031]" lastBlockId="8.[151,1437,151,2013]" lastPageId="8" lastPageNumber="553" pageId="6" pageNumber="551">
<emphasis id="B926C932FFCFFFA7B999F85CFEF3F80D" bold="true" box="[199,305,1934,1959]" pageId="6" pageNumber="551">Pedipalp</emphasis>
:
<emphasis id="B926C932FFCFFFA7B81FF85CFE05F80D" bold="true" box="[321,455,1934,1959]" pageId="6" pageNumber="551">Trochanter</emphasis>
(
<figureCitation id="136909A5FFCFFFA7B886F85CFDD9F80D" box="[472,539,1934,1959]" captionStart="FIGURE 2" captionStartId="4.[151,250,1090,1112]" captionTargetBox="[156,1417,302,1065]" captionTargetId="figure@4.[151,1436,301,1068]" captionTargetPageId="4" captionText="FIGURE 2. Dorsal (A, C, E) and ventral (B, D, F) view of the pedipalp of Charinus belizensis sp. nov. (holotype: A, B) and C. reddelli sp. nov. (holotype: C, D; paratype from Footprint Cave: E, F). G: right femur of the holotype showing the extra dorsal spine. H: right femur of a paratype from Waterfall cave with an arrow on dorsal spine 4. Scale bars: 1 mm." httpUri="https://zenodo.org/record/264776/files/figure.png" pageId="6" pageNumber="551">Fig. 2</figureCitation>
CF): large ventral apophysis, located in the posterior border of the trochanter, spiniform, bearing 1314 large setae, and with a blunt tip pointed forward; two subequal spines, one in about the center of the anterior row of setiferous tubercles (three setae on each side), the other at the external border, below the apophysis, a bit curved inwards.
<emphasis id="B926C932FFCEFFA6BB73F881FDBCF8C6" bold="true" box="[557,638,1875,1900]" pageId="7" pageNumber="552">Femur</emphasis>
(
<figureCitation id="136909A5FFCEFFA6BBD3F881FD0DF8C6" box="[653,719,1875,1900]" captionStart="FIGURE 2" captionStartId="4.[151,250,1090,1112]" captionTargetBox="[156,1417,302,1065]" captionTargetId="figure@4.[151,1436,301,1068]" captionTargetPageId="4" captionText="FIGURE 2. Dorsal (A, C, E) and ventral (B, D, F) view of the pedipalp of Charinus belizensis sp. nov. (holotype: A, B) and C. reddelli sp. nov. (holotype: C, D; paratype from Footprint Cave: E, F). G: right femur of the holotype showing the extra dorsal spine. H: right femur of a paratype from Waterfall cave with an arrow on dorsal spine 4. Scale bars: 1 mm." httpUri="https://zenodo.org/record/264776/files/figure.png" pageId="7" pageNumber="552">Fig. 2</figureCitation>
CH, 3B): three dorsal spines in the left pedipalp and four in the right pedipalp (the last spine very small) [in one male (HUJINVAM 116) the left pedipalp have four spines and the right three; two female
<typeStatus id="54E9AB82FFCEFFA6B8C0F84EFDCBF81E" box="[414,521,1948,1972]" pageId="7" pageNumber="552" type="paratype">paratypes</typeStatus>
(HUJINVAM 116) have four spines in both pedipalps, the last one very tiny (
<figureCitation id="136909A5FFCEFFA6BC31F849FF67F872" captionStart="FIGURE 2" captionStartId="4.[151,250,1090,1112]" captionTargetBox="[156,1417,302,1065]" captionTargetId="figure@4.[151,1436,301,1068]" captionTargetPageId="4" captionText="FIGURE 2. Dorsal (A, C, E) and ventral (B, D, F) view of the pedipalp of Charinus belizensis sp. nov. (holotype: A, B) and C. reddelli sp. nov. (holotype: C, D; paratype from Footprint Cave: E, F). G: right femur of the holotype showing the extra dorsal spine. H: right femur of a paratype from Waterfall cave with an arrow on dorsal spine 4. Scale bars: 1 mm." httpUri="https://zenodo.org/record/264776/files/figure.png" pageId="7" pageNumber="552">Fig. 2</figureCitation>
G, H)] with two prominent setiferous tubercle before the first spine (I&gt;II&gt;III&gt;IV); three ventral spines (I&gt;II&gt;III) with one small setiferous tubercle before the first spine.
<emphasis id="B926C932FFC1FFA9BA4AFF45FC91FF1A" bold="true" box="[788,851,151,176]" pageId="8" pageNumber="553">Tibia</emphasis>
(
<figureCitation id="136909A5FFC1FFA9BA3DFF45FC64FF1A" box="[867,934,151,176]" captionStart="FIGURE 2" captionStartId="4.[151,250,1090,1112]" captionTargetBox="[156,1417,302,1065]" captionTargetId="figure@4.[151,1436,301,1068]" captionTargetPageId="4" captionText="FIGURE 2. Dorsal (A, C, E) and ventral (B, D, F) view of the pedipalp of Charinus belizensis sp. nov. (holotype: A, B) and C. reddelli sp. nov. (holotype: C, D; paratype from Footprint Cave: E, F). G: right femur of the holotype showing the extra dorsal spine. H: right femur of a paratype from Waterfall cave with an arrow on dorsal spine 4. Scale bars: 1 mm." httpUri="https://zenodo.org/record/264776/files/figure.png" pageId="8" pageNumber="553">Fig. 2</figureCitation>
CF; 3B): three dorsal spines (I&gt;II&gt;III); one spine distal to I (about one fourth the size of I); one small setiferous tubercle proximal to spine III; spine II two thirds spine I and spine III one third spine I; spine I and II with two setiferous tubercle on its basal third; spine III with one setiferous tubercle in its half. Two ventral spines [the female
<typeStatus id="54E9AB82FFC1FFA9BAEBFED7FBD7FEB7" box="[949,1045,261,285]" pageId="8" pageNumber="553" type="paratype">paratype</typeStatus>
(HUJINVAM 116) have three, the last (proximal) very tiny]; second spine half size of the first.
<emphasis id="B926C932FFC1FFA9BA67FEF5FC77FEEA" bold="true" box="[825,949,295,320]" pageId="8" pageNumber="553">Basitarsus</emphasis>
(
<figureCitation id="136909A5FFC1FFA9BA9BFEF5FBC4FEEA" box="[965,1030,295,320]" captionStart="FIGURE 2" captionStartId="4.[151,250,1090,1112]" captionTargetBox="[156,1417,302,1065]" captionTargetId="figure@4.[151,1436,301,1068]" captionTargetPageId="4" captionText="FIGURE 2. Dorsal (A, C, E) and ventral (B, D, F) view of the pedipalp of Charinus belizensis sp. nov. (holotype: A, B) and C. reddelli sp. nov. (holotype: C, D; paratype from Footprint Cave: E, F). G: right femur of the holotype showing the extra dorsal spine. H: right femur of a paratype from Waterfall cave with an arrow on dorsal spine 4. Scale bars: 1 mm." httpUri="https://zenodo.org/record/264776/files/figure.png" pageId="8" pageNumber="553">Fig. 2</figureCitation>
CF): two dorsal spines, the basal 2/ 3 the size of the distal. One ventral spine at the distal half, 2/3 the basal spine dorsal.
<emphasis id="B926C932FFC1FFA9BD30FE9EFB2DFECF" bold="true" box="[1134,1263,332,357]" pageId="8" pageNumber="553">Distitarsus</emphasis>
(
<figureCitation id="136909A5FFC1FFA9BC5EFE9EFA86FECE" box="[1280,1348,332,357]" captionStart="FIGURE 3" captionStartId="5.[151,250,1645,1667]" captionTargetBox="[151,1436,195,1622]" captionTargetId="figure@5.[151,1436,195,1623]" captionTargetPageId="5" captionText="FIGURE 3. Frontal view of pedipalp tibia and distitibia of C. belizensis sp. nov. (A, C), and C. reddelli sp. nov. (B, D). Scale bars: A and B: 1 mm; C and D: 0.5 mm." httpUri="https://zenodo.org/record/264777/files/figure.png" pageId="8" pageNumber="553">Fig. 3</figureCitation>
D): two large curved spines, the distal half the size of the article and pointed forward; the proximal one third the size of the distal and pointed upward. Cleaning organ about half of the article length.
<emphasis id="B926C932FFC1FFA9BAABFE46FBF6FE07" bold="true" box="[1013,1076,404,429]" pageId="8" pageNumber="553">Claw</emphasis>
(
<figureCitation id="136909A5FFC1FFA9BD1BFE46FB48FE06" box="[1093,1162,404,429]" captionStart="FIGURE 3" captionStartId="5.[151,250,1645,1667]" captionTargetBox="[151,1436,195,1622]" captionTargetId="figure@5.[151,1436,195,1623]" captionTargetPageId="5" captionText="FIGURE 3. Frontal view of pedipalp tibia and distitibia of C. belizensis sp. nov. (A, C), and C. reddelli sp. nov. (B, D). Scale bars: A and B: 1 mm; C and D: 0.5 mm." httpUri="https://zenodo.org/record/264777/files/figure.png" pageId="8" pageNumber="553">Fig. 3</figureCitation>
D): long, with an acute, curved tip.
</paragraph>
<caption id="DF2D45A8FFCEFFA6B9C9F9AFFDEAF886" httpUri="https://zenodo.org/record/264779/files/figure.png" pageId="7" pageNumber="552" targetBox="[191,1395,193,1639]" targetPageId="7">
<paragraph id="8BED1520FFCEFFA6B9C9F9AFFDEAF886" blockId="7.[151,1436,1661,1836]" pageId="7" pageNumber="552">
<emphasis id="B926C932FFCEFFA6B9C9F9AFFECEF938" bold="true" box="[151,268,1661,1683]" pageId="7" pageNumber="552">FIGURE 5</emphasis>
. Leg IV trichobothriotaxy and chelicerae of
<taxonomicName id="4C526EA3FFCEFFA6BB92F9AFFC51F938" box="[716,915,1661,1682]" class="Arachnida" family="Charinidae" genus="Charinus" kingdom="Animalia" order="Amblypygi" pageId="7" pageNumber="552" phylum="Arthropoda" rank="species" species="belizensis" status="sp. nov.">
<emphasis id="B926C932FFCEFFA6BB92F9AFFC51F938" box="[716,915,1661,1682]" italics="true" pageId="7" pageNumber="552">Charinus belizensis</emphasis>
</taxonomicName>
<emphasis id="B926C932FFCEFFA6BAC6F9AFFC2AF938" bold="true" box="[920,1000,1661,1682]" pageId="7" pageNumber="552">
<taxonomicNameLabel id="A2157449FFCEFFA6BAC6F9AFFC2AF938" box="[920,1000,1661,1682]" pageId="7" pageNumber="552" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(AB, G) and
<taxonomicName id="4C526EA3FFCEFFA6BD23F9AFFB22F938" box="[1149,1248,1661,1682]" class="Arachnida" family="Charinidae" genus="Charinus" kingdom="Animalia" order="Amblypygi" pageId="7" pageNumber="552" phylum="Arthropoda" rank="species" species="reddeli" status="sp. nov.">
<emphasis id="B926C932FFCEFFA6BD23F9AFFB22F938" box="[1149,1248,1661,1682]" italics="true" pageId="7" pageNumber="552">C. reddeli</emphasis>
</taxonomicName>
<emphasis id="B926C932FFCEFFA6BDB9F9AFFAF5F938" bold="true" box="[1255,1335,1661,1682]" pageId="7" pageNumber="552">
<taxonomicNameLabel id="A2157449FFCEFFA6BDB9F9AFFAF5F938" box="[1255,1335,1661,1682]" pageId="7" pageNumber="552" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(CF, H). A: left leg of
<taxonomicName id="4C526EA3FFCEFFA6B879F94EFE6BF91B" box="[295,425,1692,1713]" class="Arachnida" family="Charinidae" genus="Charinus" kingdom="Animalia" order="Amblypygi" pageId="7" pageNumber="552" phylum="Arthropoda" rank="species" species="belizensis" status="sp. nov.">
<emphasis id="B926C932FFCEFFA6B879F94EFE6BF91B" box="[295,425,1692,1713]" italics="true" pageId="7" pageNumber="552">C. belizensis</emphasis>
</taxonomicName>
<emphasis id="B926C932FFCEFFA6B8EFF94EFDC1F91B" bold="true" box="[433,515,1692,1713]" pageId="7" pageNumber="552">
<taxonomicNameLabel id="A2157449FFCEFFA6B8EFF94EFDC1F91B" box="[433,515,1692,1713]" pageId="7" pageNumber="552" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
male holotype (HUJINVAMB 117). B: left leg of
<taxonomicName id="4C526EA3FFCEFFA6BD4DF94EFB57F91B" box="[1043,1173,1692,1713]" class="Arachnida" family="Charinidae" genus="Charinus" kingdom="Animalia" order="Amblypygi" pageId="7" pageNumber="552" phylum="Arthropoda" rank="species" species="belizensis" status="sp. nov.">
<emphasis id="B926C932FFCEFFA6BD4DF94EFB57F91B" box="[1043,1173,1692,1713]" italics="true" pageId="7" pageNumber="552">C. belizensis</emphasis>
</taxonomicName>
<emphasis id="B926C932FFCEFFA6BDC3F94EFB2DF91B" bold="true" box="[1181,1263,1692,1713]" pageId="7" pageNumber="552">
<taxonomicNameLabel id="A2157449FFCEFFA6BDC3F94EFB2DF91B" box="[1181,1263,1692,1713]" pageId="7" pageNumber="552" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
female paratype (HUJINVAMB 118). C: left leg of
<taxonomicName id="4C526EA3FFCEFFA6B8A4F969FDA4F965" box="[506,614,1722,1744]" class="Arachnida" family="Charinidae" genus="Charinus" kingdom="Animalia" order="Amblypygi" pageId="7" pageNumber="552" phylum="Arthropoda" rank="species" species="reddelli" status="sp. nov.">
<emphasis id="B926C932FFCEFFA6B8A4F969FDA4F965" box="[506,614,1722,1744]" italics="true" pageId="7" pageNumber="552">C. reddelli</emphasis>
</taxonomicName>
<emphasis id="B926C932FFCEFFA6BB32F968FD7FF965" bold="true" box="[620,701,1722,1743]" pageId="7" pageNumber="552">
<taxonomicNameLabel id="A2157449FFCEFFA6BB32F968FD7FF965" box="[620,701,1722,1743]" pageId="7" pageNumber="552" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
female holotype (HUJINVAMB 115). D, E, F: left leg of
<taxonomicName id="4C526EA3FFCEFFA6BC55F969FAB5F965" box="[1291,1399,1722,1744]" class="Arachnida" family="Charinidae" genus="Charinus" kingdom="Animalia" order="Amblypygi" pageId="7" pageNumber="552" phylum="Arthropoda" rank="species" species="reddelli" status="sp. nov.">
<emphasis id="B926C932FFCEFFA6BC55F969FAB5F965" box="[1291,1399,1722,1744]" italics="true" pageId="7" pageNumber="552">C. reddelli</emphasis>
</taxonomicName>
<emphasis id="B926C932FFCEFFA6BC23F968FF01F944" bold="true" pageId="7" pageNumber="552">
<taxonomicNameLabel id="A2157449FFCEFFA6BC23F968FF01F944" pageId="7" pageNumber="552" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
female paratypes from Waterfall cave (HUJINVAMB 116) and Footprint Cave (HUJINVAMB 114, HUJINVAMB 113), prespectively. G: right chelicerae of
<taxonomicName id="4C526EA3FFCEFFA6BB44F92AFD5CF8A7" box="[538,670,1784,1805]" class="Arachnida" family="Charinidae" genus="Charinus" kingdom="Animalia" order="Amblypygi" pageId="7" pageNumber="552" phylum="Arthropoda" rank="species" species="belizensis" status="sp. nov.">
<emphasis id="B926C932FFCEFFA6BB44F92AFD5CF8A7" box="[538,670,1784,1805]" italics="true" pageId="7" pageNumber="552">C. belizensis</emphasis>
</taxonomicName>
<emphasis id="B926C932FFCEFFA6BBF6F92AFD39F8A7" bold="true" box="[680,763,1784,1805]" pageId="7" pageNumber="552">
<taxonomicNameLabel id="A2157449FFCEFFA6BBF6F92AFD39F8A7" box="[680,763,1784,1805]" pageId="7" pageNumber="552" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(HUJINVAMB 118). H: right cheliceae of
<taxonomicName id="4C526EA3FFCEFFA6BD90F92AFAFFF8A7" box="[1230,1341,1784,1805]" class="Arachnida" family="Charinidae" genus="Charinus" kingdom="Animalia" order="Amblypygi" pageId="7" pageNumber="552" phylum="Arthropoda" rank="species" species="reddelli" status="sp. nov.">
<emphasis id="B926C932FFCEFFA6BD90F92AFAFFF8A7" box="[1230,1341,1784,1805]" italics="true" pageId="7" pageNumber="552">C. reddelli</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2157449FFCEFFA6BC19F92AFA59F8A7" box="[1351,1435,1784,1805]" pageId="7" pageNumber="552" rank="species">
<emphasis id="B926C932FFCEFFA6BC19F92AFA59F8A7" bold="true" box="[1351,1435,1784,1805]" pageId="7" pageNumber="552">sp. nov.</emphasis>
</taxonomicNameLabel>
(HUJINVAMB 116). Scale bars: 1 mm.
</paragraph>
</caption>
<paragraph id="8BED1520FFC1FFA9B999FE0EFC6EFD66" blockId="8.[151,1437,151,2013]" pageId="8" pageNumber="553">
<emphasis id="B926C932FFC1FFA9B999FE0EFF3DFE5F" bold="true" box="[199,255,476,501]" pageId="8" pageNumber="553">Legs</emphasis>
: All setose. Ventral corner of the prolateral face of femora IIIV projecting in a distinct spiniform process.
<emphasis id="B926C932FFC1FFA9B9A7FE2DFE5FFDB2" bold="true" box="[249,413,511,536]" pageId="8" pageNumber="553">Femur length</emphasis>
: I&gt;III&gt;II&gt;IV. Tibia I with 23 articles; distal segments with two small trichobothria, one on the dorsal and one in the lateral (ectal) side of the segment; on the left leg, one trichobothria in the second, third and fourth (from distal to proximal) segments, close to the distal border, the trichobothria on the third segment more lateral, the others more dorsal [in the
<typeStatus id="54E9AB82FFC1FFA9BB14FDBFFD74FD2F" box="[586,694,621,645]" pageId="8" pageNumber="553" type="paratype">paratypes</typeStatus>
they are ventral]; the right leg has two trichobothria in the third segment, one dorsal and one mesal; no trichobothria on the other segments. Tarsus (basitarsus+distitarsus) I with 41 articles; proximal segment 1.7 times longer than the next (
<figureCitation id="136909A5FFC1FFA9BA18FD66FC48FD66" box="[838,906,692,717]" captionStart="FIGURE 4" captionStartId="6.[151,250,582,604]" captionTargetBox="[151,1436,193,560]" captionTargetId="figure@6.[151,1436,193,561]" captionTargetPageId="6" captionText="FIGURE 4. Part of tibia and tarsus I and last segments of tarsus IV, all lateral view. Tibia and tarsus I of C. belizensis sp. nov. (A) and C. reddelli sp. nov. (B). Tarsus IV of C. belizensis sp. nov. (C) and C. reddelli sp. nov. (D). Arrows points to the membranose ring in the second segment of the tarsus. Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/264778/files/figure.png" pageId="8" pageNumber="553">Fig. 4</figureCitation>
B).
</paragraph>
<paragraph id="8BED1520FFC1FFA9B999FD05FC71FC2A" blockId="8.[151,1437,151,2013]" pageId="8" pageNumber="553">
<emphasis id="B926C932FFC1FFA9B999FD05FED9FD5A" bold="true" box="[199,283,727,752]" pageId="8" pageNumber="553">Leg IV</emphasis>
:
<emphasis id="B926C932FFC1FFA9B873FD05FE55FD5A" bold="true" box="[301,407,727,752]" pageId="8" pageNumber="553">Basitibia</emphasis>
: divided into three pseudo-articles, with one trichobothrium on the first third of the last pseudo-segments (trichobothrium
<emphasis id="B926C932FFC1FFA9BB42FD2EFDF3FCBE" box="[540,561,764,788]" italics="true" pageId="8" pageNumber="553">bt</emphasis>
).
<emphasis id="B926C932FFC1FFA9BB15FD2EFD7BFCBF" bold="true" box="[587,697,764,789]" pageId="8" pageNumber="553">Distitibia</emphasis>
(
<figureCitation id="136909A5FFC1FFA9BB95FD2EFCCCFCBF" box="[715,782,764,789]" captionStart="FIGURE 5" captionStartId="7.[151,250,1661,1683]" captionTargetBox="[191,1395,193,1639]" captionTargetId="figure@7.[191,1395,193,1639]" captionTargetPageId="7" captionText="FIGURE 5. Leg IV trichobothriotaxy and chelicerae of Charinus belizensis sp. nov. (A B, G) and C. reddeli sp. nov. (C F, H). A: left leg of C. belizensis sp. nov. male holotype (HUJINVAMB 117). B: left leg of C. belizensis sp. nov. female paratype (HUJINVAMB 118). C: left leg of C. reddelli sp. nov. female holotype (HUJINVAMB 115). D, E, F: left leg of C. reddelli sp. nov. female paratypes from Waterfall cave (HUJINVAMB 116) and Footprint Cave (HUJINVAMB 114, HUJINVAMB 113), prespectively. G: right chelicerae of C. belizensis sp. nov. (HUJINVAMB 118). H: right cheliceae of C. reddelli sp. nov. (HUJINVAMB 116). Scale bars: 1 mm." httpUri="https://zenodo.org/record/264779/files/figure.png" pageId="8" pageNumber="553">Fig. 5</figureCitation>
CF): three proximal and 13 distal trichobothria (total of 16); trichobothrium
<emphasis id="B926C932FFC1FFA9B820FCF2FE5AFC92" box="[382,408,800,824]" italics="true" pageId="8" pageNumber="553">bc</emphasis>
midway to
<emphasis id="B926C932FFC1FFA9BB7AFCF2FDF8FC92" box="[548,570,800,824]" italics="true" pageId="8" pageNumber="553">bf</emphasis>
and
<emphasis id="B926C932FFC1FFA9BB2AFCF2FD57FC92" box="[628,661,800,824]" italics="true" pageId="8" pageNumber="553">sbf</emphasis>
[in the
<typeStatus id="54E9AB82FFC1FFA9BBAFFCF2FC9DFC92" box="[753,863,800,824]" pageId="8" pageNumber="553" type="paratype">paratypes</typeStatus>
,
<emphasis id="B926C932FFC1FFA9BA32FCF2FC45FC92" box="[876,903,800,824]" italics="true" pageId="8" pageNumber="553">bc</emphasis>
is closer to
<emphasis id="B926C932FFC1FFA9BD48FCF2FBF5FC92" box="[1046,1079,800,824]" italics="true" pageId="8" pageNumber="553">sbf</emphasis>
than to
<emphasis id="B926C932FFC1FFA9BDC6FCF2FB6CFC92" box="[1176,1198,800,824]" italics="true" pageId="8" pageNumber="553">bf</emphasis>
];
<emphasis id="B926C932FFC1FFA9BD96FCF2FB19FC92" box="[1224,1243,800,824]" italics="true" pageId="8" pageNumber="553">sf</emphasis>
and
<emphasis id="B926C932FFC1FFA9BC4BFCF3FAEEFC92" box="[1301,1324,801,824]" italics="true" pageId="8" pageNumber="553">sc</emphasis>
with five trichobotrhia.
<emphasis id="B926C932FFC1FFA9B869FC96FDA5FCF7" bold="true" box="[311,615,836,861]" pageId="8" pageNumber="553">Basitibia-distitibia length</emphasis>
DT&gt;BT1&gt;BT4&gt;BT3&gt;BT2.
<emphasis id="B926C932FFC1FFA9BAF2FC96FC3EFCF7" bold="true" box="[940,1020,836,861]" pageId="8" pageNumber="553">Tarsus</emphasis>
: with very strong mark of the white ring in the distal part of the second segment of distitarsus IV (
<figureCitation id="136909A5FFC1FFA9BA12FCB5FC52FC2A" box="[844,912,871,896]" captionStart="FIGURE 4" captionStartId="6.[151,250,582,604]" captionTargetBox="[151,1436,193,560]" captionTargetId="figure@6.[151,1436,193,561]" captionTargetPageId="6" captionText="FIGURE 4. Part of tibia and tarsus I and last segments of tarsus IV, all lateral view. Tibia and tarsus I of C. belizensis sp. nov. (A) and C. reddelli sp. nov. (B). Tarsus IV of C. belizensis sp. nov. (C) and C. reddelli sp. nov. (D). Arrows points to the membranose ring in the second segment of the tarsus. Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/264778/files/figure.png" pageId="8" pageNumber="553">Fig. 4</figureCitation>
D).
</paragraph>
<paragraph id="8BED1520FFC1FFA9B999FC5EFB74FB0A" blockId="8.[151,1437,151,2013]" pageId="8" pageNumber="553">
<emphasis id="B926C932FFC1FFA9B999FC5EFEB4FC0F" bold="true" box="[199,374,908,933]" pageId="8" pageNumber="553">Measurements</emphasis>
(in mm):
<emphasis id="B926C932FFC1FFA9B8A2FC5EFD91FC0F" bold="true" box="[508,595,908,933]" pageId="8" pageNumber="553">Female</emphasis>
(n=3):
<emphasis id="B926C932FFC1FFA9BBEAFC5EFCE5FC0F" bold="true" box="[692,807,908,933]" pageId="8" pageNumber="553">Carapace</emphasis>
: Length: 2.84 (2.483.05), Width: 4.02 (3.404.40).
<emphasis id="B926C932FFC1FFA9B9C9FC7DFEC3FC62" bold="true" box="[151,257,943,968]" pageId="8" pageNumber="553">Pedipalp</emphasis>
: Femur 2.74 (2.133.09), Tibia 2.72 (1.883.28), Basitarsus 1.54 (1.131.88), Distitarsus 0.99 (0.80 1.10), Tarsal claw 0.94 (0.731.08).
<emphasis id="B926C932FFC1FFA9BB1CFC06FD44FC47" bold="true" box="[578,646,980,1005]" pageId="8" pageNumber="553">Leg I</emphasis>
: Femur 9.35 (6.9011.41), Tibia 15.60 (15.4015.80), Tarsus 15.25 (14.0016.50).
<emphasis id="B926C932FFC1FFA9B81AFC25FE4DFBBA" bold="true" box="[324,399,1015,1040]" pageId="8" pageNumber="553">Leg II</emphasis>
: Femur 6.27 (5.126.96), Basitibia 4.44 (3.605.00), Distitibia 3.14 (2.643.50), Basitarsus 1.67 (1.281.92), Other tarsal articles 0.95 (0.781.12).
<emphasis id="B926C932FFC1FFA9BA41FBCEFCBAFB9F" bold="true" box="[799,888,1052,1077]" pageId="8" pageNumber="553">Leg III</emphasis>
: Femur 7.12 (6.007.76), Basitibia 5.63 (4.80 6.08), Distitibia 3.51 (3.123.80), Basitarsus 1.97 (1.442.40), Other tarsal articles 0.93 (0.801.00).
<emphasis id="B926C932FFC1FFA9BC60FBEDFA56FBF2" bold="true" box="[1342,1428,1087,1112]" pageId="8" pageNumber="553">Leg IV</emphasis>
: Femur 6.36 (5.257.04), Basitibia I 3.32 (2.813.75), Basitibia II 1.04 (0.811.20), Basitibia III 1.63 (1.251.85), Distitibia 3.43 (2.694.00), Basitarsus 2.66 (1.584.30), Other tarsal articles 1.28 (0.852.00).
</paragraph>
<paragraph id="8BED1520FFC1FFA9B999FB7EFCDBFAFF" blockId="8.[151,1437,151,2013]" pageId="8" pageNumber="553">
<emphasis id="B926C932FFC1FFA9B999FB7EFEB4FB6F" bold="true" box="[199,374,1196,1221]" pageId="8" pageNumber="553">Measurements</emphasis>
:
<emphasis id="B926C932FFC1FFA9B8D2FB7EFE0BFB6F" bold="true" box="[396,457,1196,1221]" pageId="8" pageNumber="553">Male</emphasis>
(n=1):
<emphasis id="B926C932FFC1FFA9BB75FB7EFD5CFB6F" bold="true" box="[555,670,1196,1221]" pageId="8" pageNumber="553">Carapace</emphasis>
: Length: 2.64, Width: 3.80.
<emphasis id="B926C932FFC1FFA9BAA1FB7EFBABFB6F" bold="true" box="[1023,1129,1196,1221]" pageId="8" pageNumber="553">Pedipalp</emphasis>
: Femur 2.75, Tibia 2.59, Basitarsus 1.44, Distitarsus 0.93, Tarsal claw 0.90.
<emphasis id="B926C932FFC1FFA9BBBEFB1DFCE3FB42" bold="true" box="[736,801,1231,1256]" pageId="8" pageNumber="553">Leg I</emphasis>
: Femur 9.30, Tibia 17.18, Tarsus 15.40.
<emphasis id="B926C932FFC1FFA9BDA8FB1DFA80FB42" bold="true" box="[1270,1346,1231,1256]" pageId="8" pageNumber="553">Leg II</emphasis>
: Femur 6.16, Basitibia 4.24, Distitibia 3.04, Basitarsus 1.53, Other tarsal articles 0.80.
<emphasis id="B926C932FFC1FFA9BD59FB26FB9EFAA7" bold="true" box="[1031,1116,1268,1293]" pageId="8" pageNumber="553">Leg III</emphasis>
: Femur 6.80, Basitibia 5.36, Distitibia 3.44, Basitarsus 1.84, Other tarsal articles 0.84.
<emphasis id="B926C932FFC1FFA9BA74FAC5FCBFFA9A" bold="true" box="[810,893,1303,1328]" pageId="8" pageNumber="553">Leg IV</emphasis>
: Femur 6.00, Basitibia I 4.00, Basitibia II 1.35, Distitibia 3.45, Basitarsus 1.75, Other tarsal articles 0.85.
</paragraph>
<paragraph id="8BED1520FFC1FFA9B999FA8DFB1FFA37" blockId="8.[151,1437,151,2013]" pageId="8" pageNumber="553">
<emphasis id="B926C932FFC1FFA9B999FA8DFEADFAD2" bold="true" box="[199,367,1375,1400]" pageId="8" pageNumber="553">Color Pattern</emphasis>
(in alcohol): Chelicerae, pedipalps, carapace and abdomen yellowish-brown. Legs tibia and tarsus lighter colored. Color in live animals is similar, except for the chelicerae that are burgundy.
</paragraph>
<paragraph id="8BED1520FFC1FFA9B999FA75FABEF917" blockId="8.[151,1437,151,2013]" pageId="8" pageNumber="553">
<emphasis id="B926C932FFC1FFA9B999FA75FEF7FA6A" bold="true" box="[199,309,1447,1472]" pageId="8" pageNumber="553">Genitalia</emphasis>
: Female gonopod (
<figureCitation id="136909A5FFC1FFA9BB4BFA75FDA4FA6A" box="[533,614,1447,1472]" captionStart="FIGURE 6" captionStartId="9.[151,250,1447,1469]" captionTargetBox="[183,1406,445,1426]" captionTargetId="figure@9.[183,1407,445,1427]" captionTargetPageId="9" captionText="FIGURE 6. Female gonopod of Charinus belizensis sp. nov. (A, D, G) and C. reddelli sp. nov. (B, C, E, F, H). A: dorsal view of Charinus belizensis sp. nov.. B: dorsal view of C. reddelli sp. nov. from Waterfall cave. C: dorsal view of C. reddelli sp. nov. from Footprint cave. D: detail of the left side of the gonopod of C. belizensis sp. nov.. E: detail of the left side of the gonopod of C. reddelli sp. nov. from WC. F: detail of the left side of the gonopod of C. reddelli sp. nov. from FC. G: detail of the internal border of the gonopod of C. belizensis sp. nov.. H: detail of the internal border of the gonopod of C. reddelli sp. nov.. Scale bars: A, B: 150 Μm; C, D, detail of figure B: 50 Μm; E, F: 100 Μm; G: 20 Μm; H: 15 Μm." httpUri="https://zenodo.org/record/264780/files/figure.png" pageId="8" pageNumber="553">Figs. 6</figureCitation>
B, C, E, F, H): posterior margin of genital operculum straight, with few setae along its margin and on its surface. Gonopods oval, cushion-like, placed close to the border of the genital operculum, with a soft projection in the shape of a claw-like flap that covers the genital operculum. Internal border of the external flap is serrated, with abundant cusps close to each other. The gonopod of the female from FC (
<figureCitation id="136909A5FFC1FFA9BC31F9C6FF67F9FA" captionStart="FIGURE 6" captionStartId="9.[151,250,1447,1469]" captionTargetBox="[183,1406,445,1426]" captionTargetId="figure@9.[183,1407,445,1427]" captionTargetPageId="9" captionText="FIGURE 6. Female gonopod of Charinus belizensis sp. nov. (A, D, G) and C. reddelli sp. nov. (B, C, E, F, H). A: dorsal view of Charinus belizensis sp. nov.. B: dorsal view of C. reddelli sp. nov. from Waterfall cave. C: dorsal view of C. reddelli sp. nov. from Footprint cave. D: detail of the left side of the gonopod of C. belizensis sp. nov.. E: detail of the left side of the gonopod of C. reddelli sp. nov. from WC. F: detail of the left side of the gonopod of C. reddelli sp. nov. from FC. G: detail of the internal border of the gonopod of C. belizensis sp. nov.. H: detail of the internal border of the gonopod of C. reddelli sp. nov.. Scale bars: A, B: 150 Μm; C, D, detail of figure B: 50 Μm; E, F: 100 Μm; G: 20 Μm; H: 15 Μm." httpUri="https://zenodo.org/record/264780/files/figure.png" pageId="8" pageNumber="553">Fig. 6</figureCitation>
F) is retracted to hold the sperm sac. A layer of sediment was present in the border of genital operculum of one specimen (detail of
<figureCitation id="136909A5FFC1FFA9B82DF98EFE77F9DF" box="[371,437,1628,1653]" captionStart="FIGURE 6" captionStartId="9.[151,250,1447,1469]" captionTargetBox="[183,1406,445,1426]" captionTargetId="figure@9.[183,1407,445,1427]" captionTargetPageId="9" captionText="FIGURE 6. Female gonopod of Charinus belizensis sp. nov. (A, D, G) and C. reddelli sp. nov. (B, C, E, F, H). A: dorsal view of Charinus belizensis sp. nov.. B: dorsal view of C. reddelli sp. nov. from Waterfall cave. C: dorsal view of C. reddelli sp. nov. from Footprint cave. D: detail of the left side of the gonopod of C. belizensis sp. nov.. E: detail of the left side of the gonopod of C. reddelli sp. nov. from WC. F: detail of the left side of the gonopod of C. reddelli sp. nov. from FC. G: detail of the internal border of the gonopod of C. belizensis sp. nov.. H: detail of the internal border of the gonopod of C. reddelli sp. nov.. Scale bars: A, B: 150 Μm; C, D, detail of figure B: 50 Μm; E, F: 100 Μm; G: 20 Μm; H: 15 Μm." httpUri="https://zenodo.org/record/264780/files/figure.png" pageId="8" pageNumber="553">Fig. 6</figureCitation>
B); part of this sediment was removed to the observation of the gonopods; the presence of this cover may have a biological purpose (e.g. maintenance of moist to the book lungs or the gonopods), but cannot be inferred by now. Male gonopod with distal border of fistula sclerotized; PI curved; Lol1 long and fimbriated.
</paragraph>
<paragraph id="8BED1520FFC1FFA9B999F915FA5FF876" blockId="8.[151,1437,151,2013]" pageId="8" pageNumber="553">
<emphasis id="B926C932FFC1FFA9B999F915FEBCF94A" bold="true" box="[199,382,1735,1760]" pageId="8" pageNumber="553">Natural history</emphasis>
.
<taxonomicName id="4C526EA3FFC1FFA9B8D0F91AFDC4F94A" box="[398,518,1736,1760]" class="Arachnida" family="Charinidae" genus="Charinus" kingdom="Animalia" order="Amblypygi" pageId="8" pageNumber="553" phylum="Arthropoda" rank="species" species="reddelli" status="sp. nov.">
<emphasis id="B926C932FFC1FFA9B8D0F91AFDC4F94A" box="[398,518,1736,1760]" italics="true" pageId="8" pageNumber="553">C. reddelli</emphasis>
</taxonomicName>
<emphasis id="B926C932FFC1FFA9BB50F915FDABF94A" bold="true" box="[526,617,1735,1760]" pageId="8" pageNumber="553">
<taxonomicNameLabel id="A2157449FFC1FFA9BB50F915FDABF94A" box="[526,617,1735,1760]" pageId="8" pageNumber="553" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
was found in two karst caves located in the valley of the Caves Branch river in central
<collectingCountry id="F34555B0FFC1FFA9B80FF93EFE5FF8AF" box="[337,413,1772,1797]" name="Belize" pageId="8" pageNumber="553">Belize</collectingCountry>
: Footprint Cave, and Waterfall cave (Actun Lubul Ha). Both caves are decorated with stalagmites, stalactites and columns of dense flowstone. Footprint cave has a stream flowing straight through it, confined to the lower passage of the cave. It emerges out of the caves entrance and joins the Caves Branch river about
<quantity id="4CAAB8C5FFC1FFA9B982F885FECFF8DA" box="[220,269,1879,1904]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="8" pageNumber="553" unit="km" value="2.0">2km</quantity>
away. Only one entrance to the cave is known. Although we found several small-sized arthropods in this cave (isopods, diplurans), the most frequently encountered prey items were nymphs of
<taxonomicName id="4C526EA3FFC1FFA9BDF3F8AEFD9BF812" authority="Desutter-Grandcolas and Hubbell, 1993" authorityName="Desutter-Grandcolas and Hubbell" authorityYear="1993" class="Insecta" family="Haglotettigoniidae" genus="Mayagryllus" kingdom="Animalia" order="Orthoptera" pageId="8" pageNumber="553" phylum="Arthropoda" rank="species" species="apterus">
<emphasis id="B926C932FFC1FFA9BDF3F8AEFA5EF83E" box="[1197,1436,1916,1940]" italics="true" pageId="8" pageNumber="553">Mayagryllus apterus</emphasis>
Desutter-Grandcolas and Hubbell, 1993
</taxonomicName>
(
<taxonomicName id="4C526EA3FFC1FFA9BB37F872FD2BF813" box="[617,745,1952,1977]" class="Insecta" kingdom="Animalia" order="Orthoptera" pageId="8" pageNumber="553" phylum="Arthropoda" rank="order">Orthoptera</taxonomicName>
:
<taxonomicName id="4C526EA3FFC1FFA9BBAAF872FCA2F813" box="[756,864,1952,1977]" class="Insecta" family="Gryllidae" kingdom="Animalia" order="Orthoptera" pageId="8" pageNumber="553" phylum="Arthropoda" rank="family">Gryllidae</taxonomicName>
) and
<taxonomicName id="4C526EA3FFC1FFA9BAFFF872FF0DF876" authority="Armas and Viquez, 2010" authorityName="Armas and Viquez" authorityYear="2010" class="Arachnida" family="Hubbardiidae" genus="Belicenochrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Schizomida" pageId="8" pageNumber="553" phylum="Arthropoda" rank="species" species="peckorum">
<emphasis id="B926C932FFC1FFA9BAFFF872FB75F812" box="[929,1207,1952,1976]" italics="true" pageId="8" pageNumber="553">Belicenochrus peckorum</emphasis>
<bibRefCitation id="EFC368D1FFC1FFA9BD9EF873FF0DF876" author="Armas" pageId="8" pageNumber="553" refString="Armas, L. F. de (2010) Nuevos aracnidos de Puerto Rico (Arachnida: Amblypygi, Araneae, Opiliones, Parasitiformes, Schizomida, Scorpiones). Boletin de la Sociedad Entomologica Aragonesa, 47, 55 - 64." type="journal article" year="2010">Armas and Víquez, 2010</bibRefCitation>
</taxonomicName>
(
<taxonomicName id="4C526EA3FFC1FFA9B983F816FEA5F877" box="[221,359,1988,2013]" class="Arachnida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Schizomida" pageId="8" pageNumber="553" phylum="Arthropoda" rank="order">Schizomida</taxonomicName>
:
<taxonomicName id="4C526EA3FFC1FFA9B82FF816FDCDF877" box="[369,527,1988,2013]" class="Arachnida" family="Hubbardiidae" kingdom="Animalia" order="Schizomida" pageId="8" pageNumber="553" phylum="Arthropoda" rank="family">Hubbardiidae</taxonomicName>
). The Waterfall cave has a stream flowing between two entrances separated by ca.
</paragraph>
<paragraph id="8BED1520FFC0FFA8B9C9FF45FDB9FE22" blockId="9.[151,1437,151,392]" pageId="9" pageNumber="554">
<quantity id="4CAAB8C5FFC0FFA8B9C9FF45FF0BFF1A" box="[151,201,151,176]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="9" pageNumber="554" unit="km" value="2.0">2km</quantity>
of passage and divided by a series of cascades. This cave contains many dry cavities, and has more abundance of insects (
<taxonomicName id="4C526EA3FFC0FFA8B845FF6FFE5BFF7E" box="[283,409,189,212]" class="Insecta" family="Haglotettigoniidae" genus="Mayagryllus" kingdom="Animalia" order="Orthoptera" pageId="9" pageNumber="554" phylum="Arthropoda" rank="species" species="apterus">
<emphasis id="B926C932FFC0FFA8B845FF6FFE5BFF7E" box="[283,409,189,212]" italics="true" pageId="9" pageNumber="554">M. apterus</emphasis>
</taxonomicName>
, cockroaches) and other arthropods (isopods, spiders, soft ticks). The population of
<taxonomicName id="4C526EA3FFC0FFA8BCDCFF6EFF2EFF52" class="Arachnida" family="Charinidae" genus="Charinus" kingdom="Animalia" order="Amblypygi" pageId="9" pageNumber="554" phylum="Arthropoda" rank="species" species="reddelli" status="sp. nov.">
<emphasis id="B926C932FFC0FFA8BCDCFF6EFF2EFF52" italics="true" pageId="9" pageNumber="554">C. reddelli</emphasis>
</taxonomicName>
<emphasis id="B926C932FFC0FFA8B9A7FF0DFE9AFF52" bold="true" box="[249,344,223,248]" pageId="9" pageNumber="554">
<taxonomicNameLabel id="A2157449FFC0FFA8B9A7FF0DFE9AFF52" box="[249,344,223,248]" pageId="9" pageNumber="554" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
in the Waterfall caves appears larger (more specimens were recorded) in comparison to the Footprint cave. Individuals were never found close to each other. The egg sac contains 4
<specimenCount id="9D54DEA9FFC0FFA8BDCFFED6FB6EFEB6" box="[1169,1196,260,284]" pageId="9" pageNumber="554" type="egg">10</specimenCount>
eggs measuring
<quantity id="4CAAB8C5FFC0FFA8BC35FED6FF31FEEA" metricMagnitude="-3" metricUnit="m" metricValue="1.5699999999999998" metricValueMax="1.64" metricValueMin="1.5" pageId="9" pageNumber="554" unit="mm" value="1.5699999999999998" valueMax="1.64" valueMin="1.5">1.5 1.64mm</quantity>
in diameter. At 25°C, egg development takes ca. 150 days. The hatching praenymphae are white and measure
<quantity id="4CAAB8C5FFC0FFA8B85DFE9EFE44FECF" box="[259,390,332,357]" metricMagnitude="-3" metricUnit="m" metricValue="2.35" metricValueMax="2.5" metricValueMin="2.2" pageId="9" pageNumber="554" unit="mm" value="2.35" valueMax="2.5" valueMin="2.2">2.22.5mm</quantity>
. They climb and stay on the mothers back for 14 days, after which they molt into protonymphae measuring
<quantity id="4CAAB8C5FFC0FFA8B8E5FEA2FDC8FE22" box="[443,522,368,392]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" pageId="9" pageNumber="554" unit="mm" value="2.8">2.8mm</quantity>
in length.
</paragraph>
</subSubSection>
</treatment>
</document>