233 lines
32 KiB
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233 lines
32 KiB
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<document ID-DOI="10.1206/0003-0090.457.1.1" ID-ISSN="0003-0090" ID-Zenodo-Dep="6971356" approvalRequired="59" approvalRequired_for_taxonomicNames="59" checkinTime="1659882917898" checkinUser="felipe" docAuthor="Beck, Robin M. D., Voss, Robert S. & Jansa, Sharon A." docDate="2022" docId="03EFDD5DF6D368C1DA98FF4C1930FADB" docLanguage="en" docName="BulAmeMusNatHist.2022.457.1-350.pdf" docOrigin="Bulletin of the American Museum of Natural History 2022 (457)" docSource="https://bioone.org/journals/bulletin-of-the-american-museum-of-natural-history/volume-457/issue-1/0003-0090.457.1.1/Craniodental-Morphology-and-Phylogeny-of-Marsupials/10.1206/0003-0090.457.1.1.full" docStyle="DocumentStyle:915933466F796C9C739DF4DB6B8DCFA6.8:BulAmeMusNatHis.2011-.journal_article.1cover.type1" docStyleId="915933466F796C9C739DF4DB6B8DCFA6" docStyleName="BulAmeMusNatHis.2011-.journal_article.1cover.type1" docStyleVersion="8" docTitle="Ilariidae Tedford and Woodburne 1987" docType="treatment" docVersion="9" lastPageNumber="232" masterDocId="FFD6A525F63B6829DA0BFFAE1A62FFC7" masterDocTitle="Craniodental Morphology And Phylogeny Of Marsupials" masterLastPageNumber="353" masterPageNumber="1" pageNumber="232" updateTime="1659987007554" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title>Craniodental Morphology And Phylogeny Of Marsupials</mods:title>
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<mods:namePart>Beck, Robin M. D.</mods:namePart>
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<mods:affiliation>School of Science, Engineering and Environment University of Salford, U. K. & School of Biological, Earth & Environmental Sciences University of New South Wales, Australia & Division of Vertebrate Zoology (Mammalogy) American Museum of Natural History</mods:affiliation>
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<mods:namePart>Voss, Robert S.</mods:namePart>
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<mods:affiliation>Division of Vertebrate Zoology (Mammalogy) American Museum of Natural History</mods:affiliation>
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<mods:namePart>Jansa, Sharon A.</mods:namePart>
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<mods:affiliation>Bell Museum and Department of Ecology, Evolution, and Behavior University of Minnesota</mods:affiliation>
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<mods:date>2022</mods:date>
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<mods:number>2022-06-28</mods:number>
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<mods:number>2022</mods:number>
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<mods:number>457</mods:number>
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<mods:url>https://bioone.org/journals/bulletin-of-the-american-museum-of-natural-history/volume-457/issue-1/0003-0090.457.1.1/Craniodental-Morphology-and-Phylogeny-of-Marsupials/10.1206/0003-0090.457.1.1.full</mods:url>
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<mods:identifier type="DOI">10.1206/0003-0090.457.1.1</mods:identifier>
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<mods:identifier type="ISSN">0003-0090</mods:identifier>
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<treatment LSID="urn:lsid:plazi:treatment:03EFDD5DF6D368C1DA98FF4C1930FADB" httpUri="http://treatment.plazi.org/id/03EFDD5DF6D368C1DA98FF4C1930FADB" lastPageNumber="232" pageId="232" pageNumber="232">
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<subSubSection box="[147,596,226,250]" pageId="232" pageNumber="232" type="nomenclature">
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<paragraph blockId="232.[147,596,226,250]" box="[147,596,226,250]" pageId="232" pageNumber="232">
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<heading box="[147,596,226,250]" centered="true" fontSize="10" level="2" pageId="232" pageNumber="232" reason="4">
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†
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<taxonomicName authority="Tedford and Woodburne, 1987" authorityName="Tedford and Woodburne" authorityYear="1987" box="[158,596,226,250]" class="Mammalia" family="Ilariidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="232" pageNumber="232" phylum="Chordata" rank="family">
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Ilariidae
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<bibRefCitation author="Woodburne, M. O. & R. H. Tedford & M. Archer" box="[259,596,226,250]" pageId="232" pageNumber="232" pagination="639 - 679" refId="ref235391" refString="Woodburne, M. O., R. H. Tedford, and M. Archer. 1987 b. New Miocene ringtail possums (Marsupialia: Pseudocheiridae) from South Australia. In M. Archer (editor), Possums and opossums: studies in evolution: 639 - 679. Sydney: Surrey Beatty and Sons." type="book chapter" year="1987">Tedford and Woodburne, 1987</bibRefCitation>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection box="[140,351,291,315]" pageId="232" pageNumber="232" type="synonymic_list">
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<paragraph blockId="232.[108,639,291,1737]" box="[140,351,291,315]" pageId="232" pageNumber="232">
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CONTENTS: †
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<emphasis box="[285,351,291,315]" italics="true" pageId="232" pageNumber="232">
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<taxonomicName authorityName="Tedford & Woodburne" authorityYear="1987" box="[285,348,291,315]" class="Mammalia" family="Ilariidae" genus="Ilaria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="232" pageNumber="232" phylum="Chordata" rank="genus">Ilaria</taxonomicName>
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.
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</emphasis>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="232" pageNumber="232" type="description">
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<paragraph blockId="232.[108,639,291,1737]" pageId="232" pageNumber="232">STEM AGE: 30.5 Mya (95% HPD: 27.3–34.1 Mya).</paragraph>
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<paragraph blockId="232.[108,639,291,1737]" box="[140,347,391,415]" pageId="232" pageNumber="232">CROWN AGE: N/A.</paragraph>
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<paragraph blockId="232.[108,639,291,1737]" pageId="232" pageNumber="232">UNAMBIGUOUS CRANIODENTAL AUTAPOMORPHIES: Mandibular symphysis fused (char. 97: 0→1; ci = 0.333); masseteric fossa imperforate (char. 99: 1→0; ci = 0.333); and metacone much larger than paracone (char. 137: 1→2; ci = 0.400).</paragraph>
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</subSubSection>
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<subSubSection pageId="232" pageNumber="232" type="discussion">
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<paragraph blockId="232.[108,639,291,1737]" pageId="232" pageNumber="232">
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COMMENTS: This enigmatic family is currently known from three named species. †
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<taxonomicName authorityName="Tedford & Woodburne" authorityYear="1987" class="Mammalia" family="Ilariidae" genus="Ilaria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="232" pageNumber="232" phylum="Chordata" rank="species">
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<emphasis italics="true" pageId="232" pageNumber="232">Ilaria illumidens</emphasis>
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</taxonomicName>
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and †
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<taxonomicName authorityName="Tedford & Woodburne" authorityYear="1987" box="[255,359,655,679]" class="Mammalia" family="Ilariidae" genus="Ilaria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="232" pageNumber="232" phylum="Chordata" rank="species">
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<emphasis box="[255,359,655,679]" italics="true" pageId="232" pageNumber="232">I. lawsoni</emphasis>
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</taxonomicName>
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are calf-sized taxa (~
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<quantity metricMagnitude="2" metricUnit="kg" metricValue="1.5" pageId="232" pageNumber="232" unit="kg" value="150.0">150 kg</quantity>
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;
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<bibRefCitation author="Beck, R. M. D. & M. L. Taglioretti" box="[147,323,688,712]" pageId="232" pageNumber="232" pagination="385 - 417" refId="ref192802" refString="Beck, R. M. D., and M. L. Taglioretti. 2020. A nearly complete juvenile skull of the marsupial Sparassocynus derivatus from the Pliocene of Argentina, the affinities of " sparassocynids, " and the diversification of opossums (Marsupialia; Didelphimorphia; Didelphidae). Journal of Mammalian Evolution 27: 385 - 417." type="journal article" year="2020">Beck et al., 2020</bibRefCitation>
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: supplementary information) from the late Oligocene Pinpa Local Fauna of the Namba Formation,
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<collectingRegion box="[321,489,754,778]" country="Australia" name="South Australia" pageId="232" pageNumber="232">South Australia</collectingRegion>
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(
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<bibRefCitation author="Woodburne, M. O. & R. H. Tedford & M. Archer" pageId="232" pageNumber="232" pagination="639 - 679" refId="ref235391" refString="Woodburne, M. O., R. H. Tedford, and M. Archer. 1987 b. New Miocene ringtail possums (Marsupialia: Pseudocheiridae) from South Australia. In M. Archer (editor), Possums and opossums: studies in evolution: 639 - 679. Sydney: Surrey Beatty and Sons." type="book chapter" year="1987">Tedford and Woodburne, 1987</bibRefCitation>
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), whereas †
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<taxonomicName authorityName="Pledge" authorityYear="1987" box="[427,613,788,811]" class="Mammalia" family="Ilariidae" genus="Kuterintja" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="232" pageNumber="232" phylum="Chordata" rank="species">
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<emphasis box="[427,613,788,811]" italics="true" pageId="232" pageNumber="232">Kuterintja ngama</emphasis>
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</taxonomicName>
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is a much smaller taxon (~
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<quantity box="[376,440,820,844]" metricMagnitude="1" metricUnit="kg" metricValue="1.6" pageId="232" pageNumber="232" unit="kg" value="16.0">16 kg</quantity>
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;
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<bibRefCitation author="Beck, R. M. D. & M. L. Taglioretti" box="[452,634,820,845]" pageId="232" pageNumber="232" pagination="385 - 417" refId="ref192802" refString="Beck, R. M. D., and M. L. Taglioretti. 2020. A nearly complete juvenile skull of the marsupial Sparassocynus derivatus from the Pliocene of Argentina, the affinities of " sparassocynids, " and the diversification of opossums (Marsupialia; Didelphimorphia; Didelphidae). Journal of Mammalian Evolution 27: 385 - 417." type="journal article" year="2020">Beck et al., 2020</bibRefCitation>
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: supplementary information) from sites in
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<collectingRegion country="Australia" name="South Australia" pageId="232" pageNumber="232">South Australia</collectingRegion>
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and
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<collectingRegion box="[262,390,886,910]" country="Australia" name="Queensland" pageId="232" pageNumber="232">Queensland</collectingRegion>
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(
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<bibRefCitation author="Pledge" box="[405,558,886,911]" firstAuthor="Pledge" pageId="232" pageNumber="232" pagination="419 - 422" refId="ref221569" refString="Pledge, N. S. 1987 a. Kuterintja ngama, a new genus and species of primitive vombatoid marsupial from the medial Miocene Ngama Local Fauna of South Australia. In M. Archer (editor), Possums and opossums: studies in evolution: 419 - 422. Sydney: Surrey Beatty and Sons." type="book chapter" year="1987" yearSuffix="a">Pledge, 1987a</bibRefCitation>
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;
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<bibRefCitation author="Myers, T. J. & M. Archer" pageId="232" pageNumber="232" pagination="379 - 392" refId="ref219166" refString="Myers, T. J., and M. Archer. 1997. Kuterintja ngama (Marsupialia, Ilariidae): a revised systematic analysis based on material from the late Oligocene of Riversleigh, northwestern Queensland. Memoirs of the Queensland Museum 41 (2): 379 - 392." type="journal article" year="1997">Myers and Archer, 1997</bibRefCitation>
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) that are also late Oligocene (
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<bibRefCitation author="Black, K. H. & M. Archer & S. J. Hand & H. Godthelp" box="[115,302,953,977]" pageId="232" pageNumber="232" pagination="679 - 706" refId="ref194192" refString="Black, K. H., M. Archer, S. J. Hand, and H. Godthelp. 2013. Revision in the marsupial diprotodontid genus Neohelos: systematics and biostratigraphy. Acta Palaeontologica Polonica 58 (4): 679 - 706." type="journal article" year="2013">Black et al., 2013</bibRefCitation>
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;
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<bibRefCitation author="Woodhead" box="[314,559,953,977]" etAl="et al." firstAuthor="Woodhead" pageId="232" pageNumber="232" pagination="153 - 167" refId="ref235654" refString="Woodhead, J., et al. 2014. Developing a radiometricallydated chronologic sequence for Neogene biotic change in Australia, from the Riversleigh World Heritage Area of Queensland. Gondwana Research 29 (1): 153 - 167." type="journal article" year="2014">Woodhead et al., 2014</bibRefCitation>
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;
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<bibRefCitation author="Arena, D. A." pageId="232" pageNumber="232" pagination="43 - 60" refId="ref191468" refString="Arena, D. A., et al. 2015. Mammalian lineages and the biostratigraphy and biochronology of Cenozoic faunas from the Riversleigh World Heritage Area, Australia. Lethaia 49 (1): 43 - 60." type="journal article" year="2015">Arena et al., 2015</bibRefCitation>
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). Additionally, an unnamed ilariid, intermediate in size between †
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<taxonomicName authorityName="Tedford & Woodburne" authorityYear="1987" box="[441,499,1019,1043]" class="Mammalia" family="Ilariidae" genus="Ilaria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="232" pageNumber="232" phylum="Chordata" rank="genus">
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<emphasis box="[441,499,1019,1043]" italics="true" pageId="232" pageNumber="232">Ilaria</emphasis>
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</taxonomicName>
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and †
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<taxonomicName authorityName="Pledge" authorityYear="1987" class="Mammalia" family="Ilariidae" genus="Kuterintja" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="232" pageNumber="232" phylum="Chordata" rank="genus">
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<emphasis italics="true" pageId="232" pageNumber="232">Kuterintja</emphasis>
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</taxonomicName>
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, is known from the late Oligocene Pwerte Marnte Marnte Fauna of the
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<collectingRegion box="[430,636,1085,1109]" country="Australia" name="Northern Territory" pageId="232" pageNumber="232">Northern Territory</collectingRegion>
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(
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<bibRefCitation author="Murray, P. F. & D. Megirian" box="[115,461,1118,1142]" pageId="232" pageNumber="232" pagination="211 - 228" refId="ref218942" refString="Murray, P. F., and D. Megirian. 2006 b. The Pwerte Marnte Marnte Local Fauna: a new vertebrate assemblage of presumed Oligocene age from the Northern Territory of Australia. Alcheringa: an Australasian Journal of Palaeontology Special Issue 1: 211 - 228." type="book chapter" year="2006">Murray and Megirian, 2006b</bibRefCitation>
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). The apparent absence of representatives of this morphologically distinctive family from sites younger than the late Oligocene suggests that ilariids probably went extinct around the Oligocene-Miocene boundary (
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<bibRefCitation author="Black, K. H. & M. Archer & S. J. Hand & H. Godthelp" box="[116,317,1283,1307]" pageId="232" pageNumber="232" pagination="983 - 1078" refId="ref194115" refString="Black, K. H., M. Archer, S. J. Hand, and H. Godthelp. 2012 b. The rise of Australian marsupials: a synopsis of biostratigraphic, phylogenetic, palaeoecologic and palaeobiogeographic understanding. In J. A. Talent (editor), Earth and life: global biodiversity, extinction intervals and biogeographic perturbations through time: 983 - 1078. Dordrecht: Springer Verlag." type="book chapter" year="2012">Black et al., 2012b</bibRefCitation>
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).
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</paragraph>
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<paragraph blockId="232.[108,639,291,1737]" lastBlockId="232.[684,1213,226,1308]" pageId="232" pageNumber="232">
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Only †
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<taxonomicName authorityName="Tedford & Woodburne" authorityYear="1987" box="[216,277,1316,1340]" class="Mammalia" family="Ilariidae" genus="Ilaria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="232" pageNumber="232" phylum="Chordata" rank="genus">
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<emphasis box="[216,277,1316,1340]" italics="true" pageId="232" pageNumber="232">Ilaria</emphasis>
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</taxonomicName>
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has been included as a terminal here, because it is the sole ilariid known from cranial material (
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<bibRefCitation author="Woodburne, M. O. & R. H. Tedford & M. Archer" pageId="232" pageNumber="232" pagination="639 - 679" refId="ref235391" refString="Woodburne, M. O., R. H. Tedford, and M. Archer. 1987 b. New Miocene ringtail possums (Marsupialia: Pseudocheiridae) from South Australia. In M. Archer (editor), Possums and opossums: studies in evolution: 639 - 679. Sydney: Surrey Beatty and Sons." type="book chapter" year="1987">Tedford and Woodburne, 1987</bibRefCitation>
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); other ilariids are known primarily from fragmentary dental material. Ilariids have been universally accepted to be vombatiforms since their original description, but their position within Vombatiformes is controversial (
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<bibRefCitation author="Marshall, L. G. & J. A. Case & M. O. Woodburne" pageId="232" pageNumber="232" pagination="433 - 505" refId="ref215725" refString="Marshall, L. G., J. A. Case, and M. O. Woodburne. 1990. Phylogenetic relationships of the families of marsupials. Current Mammalogy 2: 433 - 505." type="journal article" year="1990">Marshall et al., 1990</bibRefCitation>
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;
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<bibRefCitation author="Murray, P. F." box="[315,470,1581,1605]" pageId="232" pageNumber="232" pagination="1 - 33" refId="ref218685" refString="Murray, P. F. 1998. Palaeontology and palaeobiology of wombats. In R. T. Wells and P. A. Pridmore (editors), Wombats: 1 - 33. Chipping Norton: Surrey Beatty and Sons." type="book chapter" year="1998">Murray, 1998</bibRefCitation>
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;
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<bibRefCitation author="Bassarova, M. & M. Archer" pageId="232" pageNumber="232" pagination="25 - 27" refId="ref192343" refString="Bassarova, M., and M. Archer. 1999. Living and extinct pseudocheirids (Marsupialia, Pseudocheiridae): Phylogenetic relationships and changes in diversity through time. Australian Mammalogy 21: 25 - 27." type="journal article" year="1999">Archer et al., 1999</bibRefCitation>
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;
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<bibRefCitation author="Black, K. H. & M. Archer & S. J. Hand & H. Godthelp" box="[174,377,1614,1638]" pageId="232" pageNumber="232" pagination="983 - 1078" refId="ref194115" refString="Black, K. H., M. Archer, S. J. Hand, and H. Godthelp. 2012 b. The rise of Australian marsupials: a synopsis of biostratigraphic, phylogenetic, palaeoecologic and palaeobiogeographic understanding. In J. A. Talent (editor), Earth and life: global biodiversity, extinction intervals and biogeographic perturbations through time: 983 - 1078. Dordrecht: Springer Verlag." type="book chapter" year="2012">Black et al., 2012b</bibRefCitation>
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) and has varied among phylogenetic analyses (
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<bibRefCitation author="Munson, C. J." box="[361,523,1647,1671]" pageId="232" pageNumber="232" pagination="1 - 99" refId="ref218350" refString="Munson, C. J. 1992. Postcranial descriptions of Ilaria and Ngapakaldia (Vombatiformes, Marsupialia) and the phylogeny of the vombatiforms based on postcranial morphology. University of California Publications in Zoology 125: 1 - 99." type="journal article" year="1992">Munson, 1992</bibRefCitation>
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;
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<bibRefCitation author="Gillespie, A. K." pageId="232" pageNumber="232" refId="ref205118" refString="Gillespie, A. K. 2007. Diversity and systematics of marsupial lions from the Riversleigh World Heritage Area and the evolution of the Thylacoleonidae. Ph. D. dissertation, School of Biological, Earth, and Environmental Sciences, University of New South Wales, Sydney." type="book" year="2007">Gillespie, 2007</bibRefCitation>
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||
;
|
||
<bibRefCitation author="Black, K. H." box="[175,304,1680,1704]" pageId="232" pageNumber="232" refId="ref193722" refString="Black, K. H. 2008. Diversity, phylogeny and biostratigraphy of diprotodontoids (Marsupialia: Diprotodontidae, Palorchestidae) from the Riversleigh World Heritage Area. Ph. D. dissertation, School of Biological, Earth, and Environmental Sciences, University of New South Wales, Sydney." type="book" year="2008">Black, 2008</bibRefCitation>
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||
;
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||
<bibRefCitation author="Black, K. H. & M. Archer & S. J. Hand" box="[316,519,1680,1704]" pageId="232" pageNumber="232" pagination="125 - 138" refId="ref194056" refString="Black, K. H., M. Archer, and S. J. Hand. 2012 a. New Tertiary koala (Marsupialia, Phascolarctidae) from Riversleigh, Australia, with a revision of phascolarctid phylogenetics, paleoecology, and paleobiodiversity. Journal of Vertebrate Paleontology 32 (1): 125 - 138." type="journal article" year="2012">Black et al., 2012a</bibRefCitation>
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;
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||
<bibRefCitation author="Brewer, P. & M. Archer & S. J. Hand & R. Abel" pageId="232" pageNumber="232" pagination="1 - 40" refId="ref195250" refString="Brewer, P., M. Archer, S. J. Hand, and R. Abel. 2015. New genus of primitive wombat (Vombatidae, Marsupialia) from Miocene deposits in the Riversleigh World Heritage Area (Queensland, Australia). Palaeontologia Electronica 8.1.9 A: 1 - 40." type="book chapter" year="2015">Brewer et al., 2015</bibRefCitation>
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;
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||
<bibRefCitation author="Gillespie, A. K. & M. Archer & S. J. Hand" box="[212,434,1713,1737]" pageId="232" pageNumber="232" pagination="1 - 26" refId="ref205243" refString="Gillespie, A. K., M. Archer, and S. J. Hand. 2016. A tiny new marsupial lion (Marsupialia, Thylacoleonidae) from the early Miocene of Australia. Palaeontologia Electronica 9.2.26 A: 1 - 26." type="book chapter" year="2016">Gillespie et al., 2016</bibRefCitation>
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;
|
||
<bibRefCitation author="Beck, R. M. D. & M. L. Taglioretti" box="[444,626,1713,1737]" pageId="232" pageNumber="232" pagination="385 - 417" refId="ref192802" refString="Beck, R. M. D., and M. L. Taglioretti. 2020. A nearly complete juvenile skull of the marsupial Sparassocynus derivatus from the Pliocene of Argentina, the affinities of " sparassocynids, " and the diversification of opossums (Marsupialia; Didelphimorphia; Didelphidae). Journal of Mammalian Evolution 27: 385 - 417." type="journal article" year="2020">Beck et al., 2020</bibRefCitation>
|
||
). Our undated total-evidence analysis (fig. 32) places †
|
||
<taxonomicName authorityName="Tedford & Woodburne" authorityYear="1987" box="[788,852,259,283]" class="Mammalia" family="Ilariidae" genus="Ilaria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="232" pageNumber="232" phylum="Chordata" rank="genus">
|
||
<emphasis box="[788,852,259,283]" italics="true" pageId="232" pageNumber="232">Ilaria</emphasis>
|
||
</taxonomicName>
|
||
as sister to †
|
||
<taxonomicName box="[1047,1211,259,283]" class="Mammalia" family="Wynyardiidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="232" pageNumber="232" phylum="Chordata" rank="family">Wynyardiidae</taxonomicName>
|
||
(†
|
||
<taxonomicName box="[708,829,292,315]" class="Mammalia" family="Wynyardiidae" genus="Muramura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="232" pageNumber="232" phylum="Chordata" rank="genus">
|
||
<emphasis box="[708,829,292,315]" italics="true" pageId="232" pageNumber="232">Muramura</emphasis>
|
||
</taxonomicName>
|
||
+ †
|
||
<taxonomicName baseAuthorityName="Pledge" baseAuthorityYear="2005" box="[880,1044,292,316]" class="Mammalia" family="Wynyardiidae" genus="Namilamadeta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="232" pageNumber="232" phylum="Chordata" rank="genus">
|
||
<emphasis box="[880,1044,292,316]" italics="true" pageId="232" pageNumber="232">Namilamadeta</emphasis>
|
||
</taxonomicName>
|
||
), a result also reported by
|
||
<bibRefCitation author="Black, K. H." box="[826,971,325,349]" pageId="232" pageNumber="232" refId="ref193722" refString="Black, K. H. 2008. Diversity, phylogeny and biostratigraphy of diprotodontoids (Marsupialia: Diprotodontidae, Palorchestidae) from the Riversleigh World Heritage Area. Ph. D. dissertation, School of Biological, Earth, and Environmental Sciences, University of New South Wales, Sydney." type="book" year="2008">Black (2008)</bibRefCitation>
|
||
,
|
||
<bibRefCitation author="Black, K. H. & M. Archer & S. J. Hand" box="[985,1208,325,349]" pageId="232" pageNumber="232" pagination="125 - 138" refId="ref194056" refString="Black, K. H., M. Archer, and S. J. Hand. 2012 a. New Tertiary koala (Marsupialia, Phascolarctidae) from Riversleigh, Australia, with a revision of phascolarctid phylogenetics, paleoecology, and paleobiodiversity. Journal of Vertebrate Paleontology 32 (1): 125 - 138." type="journal article" year="2012">Black et al. (2012a)</bibRefCitation>
|
||
, and
|
||
<bibRefCitation author="Gillespie, A. K. & M. Archer & S. J. Hand" box="[734,978,358,382]" pageId="232" pageNumber="232" pagination="1 - 26" refId="ref205243" refString="Gillespie, A. K., M. Archer, and S. J. Hand. 2016. A tiny new marsupial lion (Marsupialia, Thylacoleonidae) from the early Miocene of Australia. Palaeontologia Electronica 9.2.26 A: 1 - 26." type="book chapter" year="2016">Gillespie et al. (2016)</bibRefCitation>
|
||
. However, our dated total-evidence analysis (
|
||
<figureCitation box="[949,1020,391,415]" captionStart="FIG" captionStartId="181.[108,150,938,959]" captionTargetId="figure-12@181.[241,1216,224,1485]" captionTargetPageId="181" captionText="FIG. 33. (opposite page and at right) Fifty-per- cent majority rule consensus of post-burn-in trees that results from dated Bayesian analysis (using combined tip-and-node dating and separate Independent Gamma Rates [IGR] clock models for the molecular and morpho- logical partitions) of our total evidence data- set. Black dots at nodes indicate ≥0.95 Bayesian posterior probability (“strong support”); dark gray dots indicate 0.75–0.94 Bayesian poste- rior probability (“moderate support”); light gray dots indicate 0.50–0.74 Bayesian poste- rior probability (“weak support”). Nodes with- out dots were constrained a priori so that their ages could be calibrated (see tables 6, 13). Orange bars represent 95% Highest Posterior Density (HPD) intervals on the ages of nodes. For clarity, 95% HPD intervals are not shown for the ages of fossil terminals." pageId="232" pageNumber="232">fig. 33</figureCitation>
|
||
) places †
|
||
<taxonomicName authorityName="Tedford & Woodburne" authorityYear="1987" box="[1122,1183,391,415]" class="Mammalia" family="Ilariidae" genus="Ilaria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="232" pageNumber="232" phylum="Chordata" rank="genus">
|
||
<emphasis box="[1122,1183,391,415]" italics="true" pageId="232" pageNumber="232">Ilaria</emphasis>
|
||
</taxonomicName>
|
||
in a trichotomy at the base of Vombatomorphia with †
|
||
<taxonomicName authorityName="Gill" authorityYear="1872" box="[765,962,457,481]" class="Mammalia" family="Diprotodontidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="232" pageNumber="232" phylum="Chordata" rank="family">Diprotodontidae</taxonomicName>
|
||
and
|
||
<taxonomicName authorityName="Burnett" authorityYear="1830" box="[1040,1179,457,481]" class="Mammalia" family="Vombatidae" kingdom="Animalia" order="Diprotodontia" pageId="232" pageNumber="232" phylum="Chordata" rank="family">Vombatidae</taxonomicName>
|
||
+ †
|
||
<taxonomicName baseAuthorityName="Pledge" baseAuthorityYear="2005" box="[698,860,490,514]" class="Mammalia" family="Wynyardiidae" genus="Namilamadeta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="232" pageNumber="232" phylum="Chordata" rank="genus">
|
||
<emphasis box="[698,860,490,514]" italics="true" pageId="232" pageNumber="232">Namilamadeta</emphasis>
|
||
</taxonomicName>
|
||
+ †
|
||
<taxonomicName box="[906,1025,491,514]" class="Mammalia" family="Wynyardiidae" genus="Muramura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="232" pageNumber="232" phylum="Chordata" rank="genus">
|
||
<emphasis box="[906,1025,491,514]" italics="true" pageId="232" pageNumber="232">Muramura</emphasis>
|
||
</taxonomicName>
|
||
(†
|
||
<taxonomicName box="[1052,1212,490,514]" class="Mammalia" family="Wynyardiidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="232" pageNumber="232" phylum="Chordata" rank="family">Wynyardiidae</taxonomicName>
|
||
is paraphyletic in this analysis). The morphological analysis of
|
||
<bibRefCitation author="Beck, R. M. D. & M. L. Taglioretti" box="[883,1075,556,580]" pageId="232" pageNumber="232" pagination="385 - 417" refId="ref192802" refString="Beck, R. M. D., and M. L. Taglioretti. 2020. A nearly complete juvenile skull of the marsupial Sparassocynus derivatus from the Pliocene of Argentina, the affinities of " sparassocynids, " and the diversification of opossums (Marsupialia; Didelphimorphia; Didelphidae). Journal of Mammalian Evolution 27: 385 - 417." type="journal article" year="2020">Beck et al. (2020)</bibRefCitation>
|
||
, meanwhile, placed ilariids as sister to all other vombatomorphians.
|
||
</paragraph>
|
||
<paragraph blockId="232.[684,1213,226,1308]" pageId="232" pageNumber="232">
|
||
The lower molars of ilariids are distinctive (
|
||
<bibRefCitation author="Pledge" box="[692,849,688,713]" firstAuthor="Pledge" pageId="232" pageNumber="232" pagination="419 - 422" refId="ref221569" refString="Pledge, N. S. 1987 a. Kuterintja ngama, a new genus and species of primitive vombatoid marsupial from the medial Miocene Ngama Local Fauna of South Australia. In M. Archer (editor), Possums and opossums: studies in evolution: 419 - 422. Sydney: Surrey Beatty and Sons." type="book chapter" year="1987" yearSuffix="a">Pledge, 1987a</bibRefCitation>
|
||
;
|
||
<bibRefCitation author="Woodburne, M. O. & R. H. Tedford & M. Archer" box="[861,1207,688,713]" pageId="232" pageNumber="232" pagination="639 - 679" refId="ref235391" refString="Woodburne, M. O., R. H. Tedford, and M. Archer. 1987 b. New Miocene ringtail possums (Marsupialia: Pseudocheiridae) from South Australia. In M. Archer (editor), Possums and opossums: studies in evolution: 639 - 679. Sydney: Surrey Beatty and Sons." type="book chapter" year="1987">Tedford and Woodburne, 1987</bibRefCitation>
|
||
;
|
||
<bibRefCitation author="Myers, T. J. & M. Archer" box="[685,954,722,746]" pageId="232" pageNumber="232" pagination="379 - 392" refId="ref219166" refString="Myers, T. J., and M. Archer. 1997. Kuterintja ngama (Marsupialia, Ilariidae): a revised systematic analysis based on material from the late Oligocene of Riversleigh, northwestern Queensland. Memoirs of the Queensland Museum 41 (2): 379 - 392." type="journal article" year="1997">Myers and Archer, 1997</bibRefCitation>
|
||
;
|
||
<bibRefCitation author="Murray, P. F. & D. Megirian" pageId="232" pageNumber="232" pagination="211 - 228" refId="ref218942" refString="Murray, P. F., and D. Megirian. 2006 b. The Pwerte Marnte Marnte Local Fauna: a new vertebrate assemblage of presumed Oligocene age from the Northern Territory of Australia. Alcheringa: an Australasian Journal of Palaeontology Special Issue 1: 211 - 228." type="book chapter" year="2006">Murray and Megirian, 2006b</bibRefCitation>
|
||
). An apparently neomorphic cuspid is present between the protoconid and metaconid, and another is present between the hypoconid and entoconid. We did not score presence of the anterior neomorphic cuspid due to difficulties in defining clearly discrete states (particularly when taking into account the complex trigonid morphology seen in pseudocheirids;
|
||
<bibRefCitation author="Archer, M." pageId="232" pageNumber="232" pagination="633 - 808" refId="ref190081" refString="Archer, M. 1984 c. The Australian marsupial radiation. In M. Archer and G. Clayton (editors), Vertebrate zoogeography and evolution in Australasia: 633 - 808. Perth: Hesperian Press." type="book chapter" year="1984">Archer, 1984c</bibRefCitation>
|
||
), but we did score presence/absence and morphology of the posterior neomorphic cuspid, which we refer to as an entostylid (see char. 174). However, presence of a cusplike (versus crestlike) entostylid was not identified as an unambiguous autapomorphy of †
|
||
<taxonomicName authorityName="Tedford and Woodburne" authorityYear="1987" box="[1078,1176,1184,1208]" class="Mammalia" family="Ilariidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="232" pageNumber="232" phylum="Chordata" rank="family">Ilariidae</taxonomicName>
|
||
in our analyses, presumably because similar structures are also present in †
|
||
<taxonomicName baseAuthorityName="Pledge" baseAuthorityYear="2005" box="[994,1159,1251,1275]" class="Mammalia" family="Wynyardiidae" genus="Namilamadeta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="232" pageNumber="232" phylum="Chordata" rank="genus">
|
||
<emphasis box="[994,1159,1251,1275]" italics="true" pageId="232" pageNumber="232">Namilamadeta</emphasis>
|
||
</taxonomicName>
|
||
and phascolarctids.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |