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<document id="7AF2B706A8AB4739DE6647A9FE1D5D23" ID-DOI="10.11646/zootaxa.3962.1.6" ID-GBIF-Dataset="56c8905d-2efb-4336-be6f-20a7670e193e" ID-ISSN="1175-5326" ID-Zenodo-Dep="243914" ID-ZooBank="5872B302-2C4E-4102-B6EC-BC679B3645D4" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461068752453" checkinUser="plazi" docAuthor="Hoang, Huy Duc, Pham, Hung Manh &amp; Tran, Ngan Trong" docDate="2015" docId="03EFB841B56DFFBA06FEF9D4FCC0D41B" docLanguage="en" docName="zt03962p138.pdf" docOrigin="Zootaxa 3962 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Onychostoma dongnaiensis Hoang, Pham &amp; Tran, 2015, sp. nov." docType="treatment" docVersion="8" lastPageNumber="136" masterDocId="FFD6C039B56AFFB70669FF8AFFFFD31E" masterDocTitle="Two new species of shovel-jaw carp Onychostoma (Teleostei: Cyprinidae) from southern Vietnam" masterLastPageNumber="138" masterPageNumber="123" pageNumber="130" updateTime="1698583857553" updateUser="plazi">
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<mods:title id="00E68670971C9E9EB75E5AAD26703717">Two new species of shovel-jaw carp Onychostoma (Teleostei: Cyprinidae) from southern Vietnam</mods:title>
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<mods:namePart id="D216692C3522B461BA8D699329AD7DD0">Hoang, Huy Duc</mods:namePart>
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<mods:namePart id="337EE17A600575367F116925AD41BD3A">Pham, Hung Manh</mods:namePart>
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<mods:namePart id="BBA43A7FD5EF4C5C973E205DCFE1AF3A">Tran, Ngan Trong</mods:namePart>
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<mods:date id="40344D5935ED46CE9649B877CF212BE7">2015</mods:date>
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<treatment id="03EFB841B56DFFBA06FEF9D4FCC0D41B" ID-DOI="http://doi.org/10.5281/zenodo.5628884" ID-GBIF-Taxon="119615493" ID-Zenodo-Dep="5628884" LSID="urn:lsid:plazi:treatment:03EFB841B56DFFBA06FEF9D4FCC0D41B" httpUri="http://treatment.plazi.org/id/03EFB841B56DFFBA06FEF9D4FCC0D41B" lastPageId="13" lastPageNumber="136" pageId="7" pageNumber="130">
<subSubSection id="C35C5ADCB56DFFB006FEF9D4FDB6D566" box="[151,585,1630,1656]" pageId="7" pageNumber="130" type="nomenclature">
<paragraph id="8BF90957B56DFFB006FEF9D4FDB6D566" blockId="7.[151,585,1630,1656]" box="[151,585,1630,1656]" pageId="7" pageNumber="130">
<heading id="D0B1BE3BB56DFFB006FEF9D4FDB6D566" bold="true" box="[151,585,1630,1656]" fontSize="11" level="1" pageId="7" pageNumber="130" reason="1">
<emphasis id="B932D545B56DFFB006FEF9D4FDB6D566" bold="true" box="[151,585,1630,1656]" pageId="7" pageNumber="130">
<taxonomicName id="4C4672D4B56DFFB006FEF9D4FE1BD566" ID-CoL="49RHB" box="[151,484,1630,1656]" class="Actinopterygii" family="Cyprinidae" genus="Onychostoma" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="130" phylum="Chordata" rank="species" species="dongnaiensis" status="sp. nov.">
<emphasis id="B932D545B56DFFB006FEF9D4FE1BD566" bold="true" box="[151,484,1630,1656]" italics="true" pageId="7" pageNumber="130">Onychostoma dongnaiensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A201683EB56DFFB00782F9D4FDB6D566" box="[491,585,1630,1656]" pageId="7" pageNumber="130" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="C35C5ADCB56DFFB006FEF92FFE4CD4A4" pageId="7" pageNumber="130" type="materials_examined">
<paragraph id="8BF90957B56DFFB006FEF92FFCE9D418" blockId="7.[151,1437,1701,2014]" pageId="7" pageNumber="130">
<emphasis id="B932D545B56DFFB006FEF92FFEF2D5A0" bold="true" box="[151,269,1701,1726]" pageId="7" pageNumber="130">
<typeStatus id="54FDB7F5B56DFFB006FEF92FFEF7D5A0" box="[151,264,1701,1726]" pageId="7" pageNumber="130" type="holotype">Holotype</typeStatus>
:
</emphasis>
<collectionCode id="ED579192B56DFFB00770F92FFEACD5A0" box="[281,339,1701,1726]" httpUri="http://grbio.org/cool/0nv9-p2k7" name="University of Science, Ho Chi Minh City, Vietnam" pageId="7" pageNumber="130">UNS</collectionCode>
00851, adult female,
<quantity id="4CBEA4B2B56DFFB00424F92CFD4FD5A3" box="[589,688,1702,1726]" metricMagnitude="-1" metricUnit="m" metricValue="1.72" pageId="7" pageNumber="130" unit="mm" value="172.0">172 mm</quantity>
SL; upper Da Teh main stream: middle Dong Nai drainage in montane evergreen forest in Lam Dong Province,
<collectingCountry id="F35149C7B56DFFB004B5F943FCC4D5FC" box="[732,827,1737,1762]" name="Vietnam" pageId="7" pageNumber="130">Vietnam</collectingCountry>
(
<geoCoordinate id="EE726F90B56DFFB00524F943FC04D5FC" box="[845,1019,1737,1762]" direction="north" orientation="latitude" pageId="7" pageNumber="130" precision="1" value="11.603834">11°3613.80”N</geoCoordinate>
,
<geoCoordinate id="EE726F90B56DFFB00263F943FB3CD5FC" box="[1034,1219,1737,1762]" direction="east" orientation="longitude" pageId="7" pageNumber="130" precision="1" value="107.597115">107°3549.60”E</geoCoordinate>
,
<quantity id="4CBEA4B2B56DFFB002BBF940FAE3D5FC" box="[1234,1308,1738,1762]" metricMagnitude="2" metricUnit="m" metricValue="1.72" pageId="7" pageNumber="130" unit="m" value="172.0">172 m</quantity>
)
<date id="FFF82F97B56DFFB00347F940FF30D418" pageId="7" pageNumber="130" value="2014-03-11">11 March 2014</date>
by Pham Manh Hung and Hoang
<collectingCountry id="F35149C7B56DFFB00427F964FD82D41B" box="[590,637,1774,1797]" name="Germany" pageId="7" pageNumber="130">Duc</collectingCountry>
Huy (
<figureCitation id="137D15D2B56DFFB004AAF967FCFAD418" box="[707,773,1773,1798]" captionStart="FIGURE 1" captionStartId="3.[151,250,1672,1694]" captionTargetBox="[160,1429,203,1638]" captionTargetId="figure@3.[151,1436,195,1648]" captionTargetPageId="3" captionText="FIGURE 1. Type localities of Onychostoma krongnoensis (black star) and O. dongnaiensis (white star)." httpUri="https://zenodo.org/record/243915/files/figure.png" pageId="7" pageNumber="130">Fig. 1</figureCitation>
).
</paragraph>
<paragraph id="8BF90957B56DFFB006AEF89BFE4CD4A4" blockId="7.[151,1437,1701,2014]" pageId="7" pageNumber="130">
<emphasis id="B932D545B56DFFB006AEF89BFEB7D434" bold="true" box="[199,328,1809,1834]" pageId="7" pageNumber="130">
<typeStatus id="54FDB7F5B56DFFB006AEF89BFEBCD434" box="[199,323,1809,1834]" pageId="7" pageNumber="130" type="paratype">Paratypes</typeStatus>
:
</emphasis>
NSMT-P
<date id="FFF82F97B56DFFB007ACF898FDE6D434" box="[453,537,1810,1834]" pageId="7" pageNumber="130" value="49-12-12">121249</date>
(adult female,
<quantity id="4CBEA4B2B56DFFB004BDF89BFCC8D437" box="[724,823,1809,1834]" metricMagnitude="-1" metricUnit="m" metricValue="1.55" pageId="7" pageNumber="130" unit="mm" value="155.0">155 mm</quantity>
SL),
<date id="FFF82F97B56DFFB005E8F898FBC6D434" box="[897,1081,1809,1834]" pageId="7" pageNumber="130" value="2014-03-11">11 March 2014</date>
, collected at same locality as
<typeStatus id="54FDB7F5B56DFFB006FEF8BFFF02D450" box="[151,253,1845,1870]" pageId="7" pageNumber="130" type="holotype">holotype</typeStatus>
;
<collectionCode id="ED579192B56DFFB00764F8BFFEBBD450" box="[269,324,1845,1870]" httpUri="http://grbio.org/cool/0nv9-p2k7" name="University of Science, Ho Chi Minh City, Vietnam" pageId="7" pageNumber="130">UNS</collectionCode>
00852 (adult female,
<quantity id="4CBEA4B2B56DFFB0042DF8BFFD59D453" box="[580,678,1845,1870]" metricMagnitude="-1" metricUnit="m" metricValue="1.96" pageId="7" pageNumber="130" unit="mm" value="196.0">196 mm</quantity>
SL),
<collectionCode id="ED579192B56DFFB00486F8BFFCD8D450" box="[751,807,1845,1870]" httpUri="http://grbio.org/cool/0nv9-p2k7" name="University of Science, Ho Chi Minh City, Vietnam" pageId="7" pageNumber="130">UNS</collectionCode>
00853 (adult female,
<quantity id="4CBEA4B2B56DFFB0024FF8BCFB77D453" box="[1062,1160,1846,1870]" metricMagnitude="-1" metricUnit="m" metricValue="1.8" pageId="7" pageNumber="130" unit="mm" value="180.0">180 mm</quantity>
SL),
<collectionCode id="ED579192B56DFFB002B8F8BFFAF7D450" box="[1233,1288,1845,1870]" httpUri="http://grbio.org/cool/0nv9-p2k7" name="University of Science, Ho Chi Minh City, Vietnam" pageId="7" pageNumber="130">UNS</collectionCode>
00854 (adult female,
<quantity id="4CBEA4B2B56DFFB00698F8D3FEB0D46F" box="[241,335,1881,1906]" metricMagnitude="-1" metricUnit="m" metricValue="1.66" pageId="7" pageNumber="130" unit="mm" value="166.0">166 mm</quantity>
SL),
<collectionCode id="ED579192B56DFFB007E7F8D0FE3BD46C" box="[398,452,1882,1906]" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="7" pageNumber="130">ZRC</collectionCode>
54625 (adult male,
<quantity id="4CBEA4B2B56DFFB004CFF8D3FCFBD46F" box="[678,772,1881,1906]" metricMagnitude="-1" metricUnit="m" metricValue="1.5699999999999998" pageId="7" pageNumber="130" unit="mm" value="157.0">157 mm</quantity>
SL),
<date id="FFF82F97B56DFFB0052AF8D0FC1BD46C" box="[835,996,1881,1906]" pageId="7" pageNumber="130" value="2014-03-07">7 March 2014</date>
, collected at same locality as
<typeStatus id="54FDB7F5B56DFFB0035AF8D3FA67D46C" box="[1331,1432,1881,1906]" pageId="7" pageNumber="130" type="holotype">holotype</typeStatus>
;
<collectionCode id="ED579192B56DFFB006FEF8F4FF32D488" box="[151,205,1918,1942]" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="7" pageNumber="130">ZRC</collectionCode>
54626 (adult female,
<quantity id="4CBEA4B2B56DFFB007A6F8F4FDD0D48B" box="[463,559,1918,1942]" metricMagnitude="-1" metricUnit="m" metricValue="1.73" pageId="7" pageNumber="130" unit="mm" value="173.0">173 mm</quantity>
SL),
<date id="FFF82F97B56DFFB0041AF8F4FCE7D488" box="[627,792,1917,1942]" pageId="7" pageNumber="130" value="2014-04-11">11 April 2014</date>
, collected at same locality as
<typeStatus id="54FDB7F5B56DFFB0021CF8F7FB28D488" box="[1141,1239,1917,1942]" pageId="7" pageNumber="130" type="holotype">holotype</typeStatus>
(
<geoCoordinate id="EE726F90B56DFFB00280F8F4FA68D488" box="[1257,1431,1918,1942]" direction="north" orientation="latitude" pageId="7" pageNumber="130" precision="1" value="11.637164">11°3813.79”N</geoCoordinate>
,
<geoCoordinate id="EE726F90B56DFFB006FEF82BFEAED4A4" box="[151,337,1953,1978]" direction="east" orientation="longitude" pageId="7" pageNumber="130" precision="1" value="107.62393">107°3726.14”E</geoCoordinate>
,
<quantity id="4CBEA4B2B56DFFB00735F82BFE58D4A4" box="[348,423,1953,1978]" metricMagnitude="2" metricUnit="m" metricValue="1.95" pageId="7" pageNumber="130" unit="m" value="195.0">195 m</quantity>
).
</paragraph>
</subSubSection>
<subSubSection id="C35C5ADCB56DFFBF06AEF84FFEFDD3E6" lastPageId="8" lastPageNumber="131" pageId="7" pageNumber="130" type="etymology">
<paragraph id="8BF90957B56DFFBF06AEF84FFEFDD3E6" blockId="7.[151,1437,1701,2014]" lastBlockId="8.[151,1436,151,392]" lastPageId="8" lastPageNumber="131" pageId="7" pageNumber="130">
<emphasis id="B932D545B56DFFB006AEF84FFEB1D4C0" bold="true" box="[199,334,1989,2014]" pageId="7" pageNumber="130">Etymology.</emphasis>
Specific epithet is in reference to the
<typeStatus id="54FDB7F5B56DFFB00577F84CFCB1D4C0" box="[798,846,1990,2014]" pageId="7" pageNumber="130">type</typeStatus>
locality of middle Dong Nai drainage. Suggested common name: Dongnai srang (English), Cá srang Đồng Nai (Vietnamese). Srang is the vernacular name of this species of
<taxonomicName id="4C4672D4B562FFBF077AFF36FE52D3CB" box="[275,429,188,213]" class="Actinopterygii" family="Cyprinidae" genus="Onychostoma" kingdom="Animalia" order="Cypriniformes" pageId="8" pageNumber="131" phylum="Chordata" rank="genus">
<emphasis id="B932D545B562FFBF077AFF36FE52D3CB" box="[275,429,188,213]" italics="true" pageId="8" pageNumber="131">Onychostoma</emphasis>
</taxonomicName>
by the Mạ people in the Da Teh catchment of the middle Dong Nai River, Lam Dong Province.
</paragraph>
</subSubSection>
<subSubSection id="C35C5ADCB562FFBF06AEFE8EFB86D296" pageId="8" pageNumber="131" type="diagnosis">
<paragraph id="8BF90957B562FFBF06AEFE8EFB86D296" blockId="8.[151,1436,151,392]" pageId="8" pageNumber="131">
<emphasis id="B932D545B562FFBF06AEFE8EFEBED203" bold="true" box="[199,321,260,285]" pageId="8" pageNumber="131">Diagnosis.</emphasis>
<taxonomicName id="4C4672D4B562FFBF072EFE8EFD84D203" box="[327,635,260,285]" class="Actinopterygii" family="Cyprinidae" genus="Onychostoma" kingdom="Animalia" order="Cypriniformes" pageId="8" pageNumber="131" phylum="Chordata" rank="species" species="dongnaiensis">
<emphasis id="B932D545B562FFBF072EFE8EFD84D203" box="[327,635,260,285]" italics="true" pageId="8" pageNumber="131">Onychostoma dongnaiensis</emphasis>
</taxonomicName>
is distinguished from its congeners by a combination of (1) mouth wide (width 1.41.6 times in HW), (2) predorsal scales 1415, (3) body depth high (3.13.4 times in SL), (4) eye diameter small (4.24.9 times in HL), (5) no barbels in adults and juveniles, (6) a strong serrated last simple ray of dorsal fin, and (7) dorsal-fin length high (3.84.5 times in SL) (
<figureCitation id="137D15D2B562FFBF0509FEE5FC52D296" box="[864,941,367,392]" captionStart="FIGURE 5" captionStartId="8.[151,250,1965,1987]" captionTargetBox="[163,1419,473,1929]" captionTargetId="figure@8.[151,1435,447,1940]" captionTargetPageId="8" captionText="FIGURE 5. Lateral view of Onychostoma dongnaiensis specimens (a) female holotype UNS 00851 in life, (b) female holotype UNS 00851 in preservative, (c) male paratype ZRC 54625 in preservative. Scale 10 mm." httpUri="https://zenodo.org/record/243919/files/figure.png" pageId="8" pageNumber="131">Figs. 5</figureCitation>
a, b, c; 3e, f, g, h).
</paragraph>
</subSubSection>
<caption id="DF3959DFB562FFBF06FEF827FBECD4FC" httpUri="https://zenodo.org/record/243919/files/figure.png" pageId="8" pageNumber="131" targetBox="[163,1419,473,1929]" targetPageId="8">
<paragraph id="8BF90957B562FFBF06FEF827FBECD4FC" blockId="8.[151,1436,1965,2018]" pageId="8" pageNumber="131">
<emphasis id="B932D545B562FFBF06FEF827FEEDD4DD" bold="true" box="[151,274,1965,1987]" pageId="8" pageNumber="131">FIGURE 5.</emphasis>
Lateral view of
<taxonomicName id="4C4672D4B562FFBF07D3F827FD2ED4DD" box="[442,721,1965,1987]" class="Actinopterygii" family="Cyprinidae" genus="Onychostoma" kingdom="Animalia" order="Cypriniformes" pageId="8" pageNumber="131" phylum="Chordata" rank="species" species="dongnaiensis">
<emphasis id="B932D545B562FFBF07D3F827FD2ED4DD" box="[442,721,1965,1987]" italics="true" pageId="8" pageNumber="131">Onychostoma dongnaiensis</emphasis>
</taxonomicName>
specimens (a) female holotype UNS00851 in life, (b) female holotype UNS00851 in preservative, (c) male paratype ZRC 54625 in preservative. Scale 10 mm.
</paragraph>
</caption>
<subSubSection id="C35C5ADCB563FFBB06AEFF1DFF17D55B" lastPageId="12" lastPageNumber="135" pageId="9" pageNumber="132" type="description">
<paragraph id="8BF90957B563FFBE06AEFF1DFEFFD296" blockId="9.[151,1437,151,392]" pageId="9" pageNumber="132">
<emphasis id="B932D545B563FFBE06AEFF1DFEA6D3AE" bold="true" box="[199,345,151,176]" pageId="9" pageNumber="132">Description.</emphasis>
General appearance shown in
<figureCitation id="137D15D2B563FFBE04AEFF1DFCD4D3AE" box="[711,811,151,176]" captionStart="FIGURE 5" captionStartId="8.[151,250,1965,1987]" captionTargetBox="[163,1419,473,1929]" captionTargetId="figure@8.[151,1435,447,1940]" captionTargetPageId="8" captionText="FIGURE 5. Lateral view of Onychostoma dongnaiensis specimens (a) female holotype UNS 00851 in life, (b) female holotype UNS 00851 in preservative, (c) male paratype ZRC 54625 in preservative. Scale 10 mm." httpUri="https://zenodo.org/record/243919/files/figure.png" pageId="9" pageNumber="132">Figure 5</figureCitation>
; meristic and morphometric data of seven adult
<typeStatus id="54FDB7F5B563FFBE0302FF12FA63D3AE" box="[1387,1436,152,176]" pageId="9" pageNumber="132">type</typeStatus>
specimens given in
<tableCitation id="C6C43CECB563FFBE071AFF36FE37D3CA" box="[371,456,188,213]" captionStart="TABLE 2" captionStartId="9.[151,235,440,462]" captionTargetBox="[165,1398,493,1948]" captionTargetPageId="9" captionText="TABLE 2. Morphometric and meristic characters of Onychostoma dongnaiensis sp. nov. Range and mean include the holotype." httpUri="http://table.plazi.org/id/DF3959DFB563FFBE06FEFE32FA62D2D0" pageId="9" pageNumber="132" tableUuid="DF3959DFB563FFBE06FEFE32FA62D2D0">Table 2</tableCitation>
. Head longer than deep, dorsal profile strongly convex. Snout stout and rounded, longer than eye diameter in adults. Interorbital area slightly convex. Mouth subterminal. Maxillary reaching vertical of anterior margin of orbit. Upper lips thick, upper region covered by rostral fold (rostral cap) completely and exposed at corners. Epidermal tubercles on snout numerous but small to medium-sized and sparse in all
<typeStatus id="54FDB7F5B563FFBE0284FEA2FAE2D25E" box="[1261,1309,296,320]" pageId="9" pageNumber="132">type</typeStatus>
specimens except
<typeStatus id="54FDB7F5B563FFBE0681FEC6FEB2D27B" box="[232,333,332,357]" pageId="9" pageNumber="132" type="holotype">holotype</typeStatus>
, which has no tubercles (
<figureCitation id="137D15D2B563FFBE0403FEC6FD51D27A" box="[618,686,332,357]" captionStart="FIGURE 3" captionStartId="5.[151,250,1842,1864]" captionTargetBox="[172,1412,529,1810]" captionTargetId="figure@5.[163,1423,519,1819]" captionTargetPageId="5" captionText="FIGURE 3. Front view of head of Onychostoma krongnoensis (a) female holotype UNS 00805, 205 mm SL, (b) immature paratype UNS 00821, 123 mm SL, (c) male paratype UNS 00825, 214 mm SL; O. dongnaiensis (e) female holotype UNS 00851, 172 mm SL, (f) female paratype UNS 00852, 196 mm SL, (g) male paratype ZRC 54625, 157 mm SL; O. leptura (i) female UNS 00684, 186 mm SL. Ventral view of head of O. krongnoensis, wide mouth (d) male paratype UNS 00825; O. dongnaiensis, wide mouth (h) female holotype UNS 00851; O. leptura, wide mouth (j) female UNS 00684; O. meridionale, moderate mouth (k) immature UNS 00826, 108 mm SL; O. uniforme, narrow mouth (l) female UNS 00689, 130 mm SL. Scale 10 mm." httpUri="https://zenodo.org/record/243917/files/figure.png" pageId="9" pageNumber="132">Fig. 3</figureCitation>
e). Mouth width about 1.41.6 times in maximum HW; no barbels (
<figureCitation id="137D15D2B563FFBE06F6FEE5FF1DD296" box="[159,226,367,392]" captionStart="FIGURE 3" captionStartId="5.[151,250,1842,1864]" captionTargetBox="[172,1412,529,1810]" captionTargetId="figure@5.[163,1423,519,1819]" captionTargetPageId="5" captionText="FIGURE 3. Front view of head of Onychostoma krongnoensis (a) female holotype UNS 00805, 205 mm SL, (b) immature paratype UNS 00821, 123 mm SL, (c) male paratype UNS 00825, 214 mm SL; O. dongnaiensis (e) female holotype UNS 00851, 172 mm SL, (f) female paratype UNS 00852, 196 mm SL, (g) male paratype ZRC 54625, 157 mm SL; O. leptura (i) female UNS 00684, 186 mm SL. Ventral view of head of O. krongnoensis, wide mouth (d) male paratype UNS 00825; O. dongnaiensis, wide mouth (h) female holotype UNS 00851; O. leptura, wide mouth (j) female UNS 00684; O. meridionale, moderate mouth (k) immature UNS 00826, 108 mm SL; O. uniforme, narrow mouth (l) female UNS 00689, 130 mm SL. Scale 10 mm." httpUri="https://zenodo.org/record/243917/files/figure.png" pageId="9" pageNumber="132">Fig. 3</figureCitation>
h).
</paragraph>
<caption id="DF3959DFB563FFBE06FEFE32FA62D2D0" ID-Table-UUID="DF3959DFB563FFBE06FEFE32FA62D2D0" box="[151,1437,440,462]" httpUri="http://table.plazi.org/id/DF3959DFB563FFBE06FEFE32FA62D2D0" pageId="9" pageNumber="132" targetBox="[165,1398,493,1948]" targetIsTable="true" targetPageId="9">
<paragraph id="8BF90957B563FFBE06FEFE32FA62D2D0" blockId="9.[151,1437,440,462]" box="[151,1437,440,462]" pageId="9" pageNumber="132">
<emphasis id="B932D545B563FFBE06FEFE32FEFCD2D0" bold="true" box="[151,259,440,462]" pageId="9" pageNumber="132">TABLE 2.</emphasis>
Morphometric and meristic characters of
<taxonomicName id="4C4672D4B563FFBE04C2FE32FC3DD2D0" box="[683,962,440,462]" class="Actinopterygii" family="Cyprinidae" genus="Onychostoma" kingdom="Animalia" order="Cypriniformes" pageId="9" pageNumber="132" phylum="Chordata" rank="species" species="dongnaiensis" status="sp. nov.">
<emphasis id="B932D545B563FFBE04C2FE32FC3DD2D0" box="[683,962,440,462]" italics="true" pageId="9" pageNumber="132">Onychostoma dongnaiensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A201683EB563FFBE05AEFE33FBEAD2D0" box="[967,1045,441,462]" pageId="9" pageNumber="132" rank="species">sp. nov.</taxonomicNameLabel>
Range and mean include the holotype.
</paragraph>
</caption>
<paragraph id="8BF90957B563FFBE06CCFD9EFA8CD482" pageId="9" pageNumber="132">
<table id="F946FBF7B563004806CCFE67FA89D482" box="[165,1398,493,1948]" gridcols="5" gridrows="35" pageId="9" pageNumber="132">
<tr id="35760B15B563004806CCFE67FA89D135" box="[165,1398,493,555]" gridrow="0" pageId="9" pageNumber="132">
<th id="76A76269B563004806CCFE67FD43D135" box="[165,700,493,555]" gridcol="0" gridrow="0" pageId="9" pageNumber="132">
<emphasis id="B932D545B563FFBE06CCFD9EFE5CD135" bold="true" box="[165,419,532,555]" pageId="9" pageNumber="132">Standard length (mm)</emphasis>
</th>
<th id="76A76269B56300480540FE67FC6ED135" box="[809,913,493,555]" gridcol="1" gridrow="0" pageId="9" pageNumber="132">
<emphasis id="B932D545B563FFBE0540FE67FC8ED135" bold="true" box="[809,913,493,555]" pageId="9" pageNumber="132">Holotype 172</emphasis>
</th>
<th id="76A76269B563004805B5FE67FBB7D135" box="[988,1096,493,555]" gridcol="2" gridrow="0" pageId="9" pageNumber="132">
<emphasis id="B932D545B563FFBE0584FE67FBCAD11A" bold="true" box="[1005,1077,493,516]" pageId="9" pageNumber="132">Range</emphasis>
15.519.6
</th>
<th id="76A76269B563004802C1FE67FAE6D135" box="[1192,1305,493,555]" gridcol="3" gridrow="0" pageId="9" pageNumber="132">
<emphasis id="B932D545B563FFBE02C1FE67FAE6D11A" bold="true" box="[1192,1305,493,516]" pageId="9" pageNumber="132">Mean±SD</emphasis>
17.1±1.4
</th>
<th id="76A76269B5630048030AFE67FA89D135" box="[1379,1398,493,555]" gridcol="4" gridrow="0" pageId="9" pageNumber="132">
<emphasis id="B932D545B563FFBE030AFE67FA89D11A" bold="true" box="[1379,1398,493,516]" pageId="9" pageNumber="132">N</emphasis>
7
</th>
</tr>
<tr id="35760B15B563004806CCFDB4FA89D14B" box="[165,1398,574,597]" gridrow="1" pageId="9" pageNumber="132" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1">
<th id="76A76269B563004806CCFDB4FD43D14B" box="[165,700,574,597]" gridcol="0" gridrow="1" pageId="9" pageNumber="132">
<emphasis id="B932D545B563FFBE06CCFDB4FEA8D14B" bold="true" box="[165,343,574,597]" pageId="9" pageNumber="132">Morphometrics</emphasis>
</th>
</tr>
<tr id="35760B15B563004806CCFDE1FA89D19C" box="[165,1398,619,642]" gridrow="2" pageId="9" pageNumber="132" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1">
<th id="76A76269B563004806CCFDE1FD43D19C" box="[165,700,619,642]" gridcol="0" gridrow="2" pageId="9" pageNumber="132">
<emphasis id="B932D545B563FFBE06D0FDE1FF04D19C" bold="true" box="[185,251,619,642]" pageId="9" pageNumber="132">% SL</emphasis>
</th>
</tr>
<tr id="35760B15B563004806CCFD1EFA89D1B5" box="[165,1398,660,683]" gridrow="3" pageId="9" pageNumber="132">
<th id="76A76269B563004806CCFD1EFD43D1B5" box="[165,700,660,683]" gridcol="0" gridrow="3" pageId="9" pageNumber="132">Head length</th>
<td id="76A76269B56300480540FD1EFC6ED1B5" box="[809,913,660,683]" gridcol="1" gridrow="3" pageId="9" pageNumber="132">19.8</td>
<td id="76A76269B563004805B5FD1EFBB7D1B5" box="[988,1096,660,683]" gridcol="2" gridrow="3" pageId="9" pageNumber="132">19.421</td>
<td id="76A76269B563004802C1FD1EFAE6D1B5" box="[1192,1305,660,683]" gridcol="3" gridrow="3" pageId="9" pageNumber="132">20.3±0.7</td>
<td id="76A76269B5630048030AFD1EFA89D1B5" box="[1379,1398,660,683]" gridcol="4" gridrow="3" pageId="9" pageNumber="132">7</td>
</tr>
<tr id="35760B15B563004806CCFD36FA89D1CD" box="[165,1398,700,723]" gridrow="4" pageId="9" pageNumber="132">
<th id="76A76269B563004806CCFD36FD43D1CD" box="[165,700,700,723]" gridcol="0" gridrow="4" pageId="9" pageNumber="132">Depth of body</th>
<td id="76A76269B56300480540FD36FC6ED1CD" box="[809,913,700,723]" gridcol="1" gridrow="4" pageId="9" pageNumber="132">29.2</td>
<td id="76A76269B563004805B5FD36FBB7D1CD" box="[988,1096,700,723]" gridcol="2" gridrow="4" pageId="9" pageNumber="132">29.232.7</td>
<td id="76A76269B563004802C1FD36FAE6D1CD" box="[1192,1305,700,723]" gridcol="3" gridrow="4" pageId="9" pageNumber="132">30.8±1.1</td>
<td id="76A76269B5630048030AFD36FA89D1CD" box="[1379,1398,700,723]" gridcol="4" gridrow="4" pageId="9" pageNumber="132">7</td>
</tr>
<tr id="35760B15B563004806CCFD6EFA89D1E5" box="[165,1398,740,763]" gridrow="5" pageId="9" pageNumber="132">
<th id="76A76269B563004806CCFD6EFD43D1E5" box="[165,700,740,763]" gridcol="0" gridrow="5" pageId="9" pageNumber="132">Body width at dorsal-fin origin</th>
<td id="76A76269B56300480540FD6EFC6ED1E5" box="[809,913,740,763]" gridcol="1" gridrow="5" pageId="9" pageNumber="132">16.0</td>
<td id="76A76269B563004805B5FD6EFBB7D1E5" box="[988,1096,740,763]" gridcol="2" gridrow="5" pageId="9" pageNumber="132">12.716</td>
<td id="76A76269B563004802C1FD6EFAE6D1E5" box="[1192,1305,740,763]" gridcol="3" gridrow="5" pageId="9" pageNumber="132">14.5±1.3</td>
<td id="76A76269B5630048030AFD6EFA89D1E5" box="[1379,1398,740,763]" gridcol="4" gridrow="5" pageId="9" pageNumber="132">7</td>
</tr>
<tr id="35760B15B563004806CCFC86FA89D03D" box="[165,1398,780,803]" gridrow="6" pageId="9" pageNumber="132">
<th id="76A76269B563004806CCFC86FD43D03D" box="[165,700,780,803]" gridcol="0" gridrow="6" pageId="9" pageNumber="132">Predorsal length</th>
<td id="76A76269B56300480540FC86FC6ED03D" box="[809,913,780,803]" gridcol="1" gridrow="6" pageId="9" pageNumber="132">43.7</td>
<td id="76A76269B563004805B5FC86FBB7D03D" box="[988,1096,780,803]" gridcol="2" gridrow="6" pageId="9" pageNumber="132">40.845.8</td>
<td id="76A76269B563004802C1FC86FAE6D03D" box="[1192,1305,780,803]" gridcol="3" gridrow="6" pageId="9" pageNumber="132">43.3±1.5</td>
<td id="76A76269B5630048030AFC86FA89D03D" box="[1379,1398,780,803]" gridcol="4" gridrow="6" pageId="9" pageNumber="132">7</td>
</tr>
<tr id="35760B15B563004806CCFCBEFA89D055" box="[165,1398,820,843]" gridrow="7" pageId="9" pageNumber="132">
<th id="76A76269B563004806CCFCBEFD43D055" box="[165,700,820,843]" gridcol="0" gridrow="7" pageId="9" pageNumber="132">Prepectoral length</th>
<td id="76A76269B56300480540FCBEFC6ED055" box="[809,913,820,843]" gridcol="1" gridrow="7" pageId="9" pageNumber="132">18.7</td>
<td id="76A76269B563004805B5FCBEFBB7D055" box="[988,1096,820,843]" gridcol="2" gridrow="7" pageId="9" pageNumber="132">18.721.7</td>
<td id="76A76269B563004802C1FCBEFAE6D055" box="[1192,1305,820,843]" gridcol="3" gridrow="7" pageId="9" pageNumber="132">20.0±1.1</td>
<td id="76A76269B5630048030AFCBEFA89D055" box="[1379,1398,820,843]" gridcol="4" gridrow="7" pageId="9" pageNumber="132">7</td>
</tr>
<tr id="35760B15B563004806CCFCD6FA89D06D" box="[165,1398,860,883]" gridrow="8" pageId="9" pageNumber="132">
<th id="76A76269B563004806CCFCD6FD43D06D" box="[165,700,860,883]" gridcol="0" gridrow="8" pageId="9" pageNumber="132">Prepelvic length</th>
<td id="76A76269B56300480540FCD6FC6ED06D" box="[809,913,860,883]" gridcol="1" gridrow="8" pageId="9" pageNumber="132">46.9</td>
<td id="76A76269B563004805B5FCD6FBB7D06D" box="[988,1096,860,883]" gridcol="2" gridrow="8" pageId="9" pageNumber="132">46.951.2</td>
<td id="76A76269B563004802C1FCD6FAE6D06D" box="[1192,1305,860,883]" gridcol="3" gridrow="8" pageId="9" pageNumber="132">48.6±1.5</td>
<td id="76A76269B5630048030AFCD6FA89D06D" box="[1379,1398,860,883]" gridcol="4" gridrow="8" pageId="9" pageNumber="132">7</td>
</tr>
<tr id="35760B15B563004806CCFC0EFA89D085" box="[165,1398,900,923]" gridrow="9" pageId="9" pageNumber="132">
<th id="76A76269B563004806CCFC0EFD43D085" box="[165,700,900,923]" gridcol="0" gridrow="9" pageId="9" pageNumber="132">Preanal length</th>
<td id="76A76269B56300480540FC0EFC6ED085" box="[809,913,900,923]" gridcol="1" gridrow="9" pageId="9" pageNumber="132">70.0</td>
<td id="76A76269B563004805B5FC0EFBB7D085" box="[988,1096,900,923]" gridcol="2" gridrow="9" pageId="9" pageNumber="132">69.773.5</td>
<td id="76A76269B563004802C1FC0EFAE6D085" box="[1192,1305,900,923]" gridcol="3" gridrow="9" pageId="9" pageNumber="132">70.8±1.3</td>
<td id="76A76269B5630048030AFC0EFA89D085" box="[1379,1398,900,923]" gridcol="4" gridrow="9" pageId="9" pageNumber="132">7</td>
</tr>
<tr id="35760B15B563004806CCFC26FA89D0DD" box="[165,1398,940,963]" gridrow="10" pageId="9" pageNumber="132">
<th id="76A76269B563004806CCFC26FD43D0DD" box="[165,700,940,963]" gridcol="0" gridrow="10" pageId="9" pageNumber="132">Distance between pectoral- and pelvic-fin origins</th>
<td id="76A76269B56300480540FC26FC6ED0DD" box="[809,913,940,963]" gridcol="1" gridrow="10" pageId="9" pageNumber="132">24.5</td>
<td id="76A76269B563004805B5FC26FBB7D0DD" box="[988,1096,940,963]" gridcol="2" gridrow="10" pageId="9" pageNumber="132">24.528.7</td>
<td id="76A76269B563004802C1FC26FAE6D0DD" box="[1192,1305,940,963]" gridcol="3" gridrow="10" pageId="9" pageNumber="132">26.4±1.8</td>
<td id="76A76269B5630048030AFC26FA89D0DD" box="[1379,1398,940,963]" gridcol="4" gridrow="10" pageId="9" pageNumber="132">7</td>
</tr>
<tr id="35760B15B563004806CCFC5EFA89D0F5" box="[165,1398,980,1003]" gridrow="11" pageId="9" pageNumber="132">
<th id="76A76269B563004806CCFC5EFD43D0F5" box="[165,700,980,1003]" gridcol="0" gridrow="11" pageId="9" pageNumber="132">Distance between pelvic- and anal-fin origins</th>
<td id="76A76269B56300480540FC5EFC6ED0F5" box="[809,913,980,1003]" gridcol="1" gridrow="11" pageId="9" pageNumber="132">19.2</td>
<td id="76A76269B563004805B5FC5EFBB7D0F5" box="[988,1096,980,1003]" gridcol="2" gridrow="11" pageId="9" pageNumber="132">17.822.4</td>
<td id="76A76269B563004802C1FC5EFAE6D0F5" box="[1192,1305,980,1003]" gridcol="3" gridrow="11" pageId="9" pageNumber="132">19.2±1.6</td>
<td id="76A76269B5630048030AFC5EFA89D0F5" box="[1379,1398,980,1003]" gridcol="4" gridrow="11" pageId="9" pageNumber="132">7</td>
</tr>
<tr id="35760B15B563004806CCFC76FA89D70D" box="[165,1398,1020,1043]" gridrow="12" pageId="9" pageNumber="132">
<th id="76A76269B563004806CCFC76FD43D70D" box="[165,700,1020,1043]" gridcol="0" gridrow="12" pageId="9" pageNumber="132">Depth of caudal peduncle</th>
<td id="76A76269B56300480540FC76FC6ED70D" box="[809,913,1020,1043]" gridcol="1" gridrow="12" pageId="9" pageNumber="132">9.3</td>
<td id="76A76269B563004805B5FC76FBB7D70D" box="[988,1096,1020,1043]" gridcol="2" gridrow="12" pageId="9" pageNumber="132">9.210</td>
<td id="76A76269B563004802C1FC76FAE6D70D" box="[1192,1305,1020,1043]" gridcol="3" gridrow="12" pageId="9" pageNumber="132">9.6±0.3</td>
<td id="76A76269B5630048030AFC76FA89D70D" box="[1379,1398,1020,1043]" gridcol="4" gridrow="12" pageId="9" pageNumber="132">7</td>
</tr>
<tr id="35760B15B563004806CCFBAEFA89D725" box="[165,1398,1060,1083]" gridrow="13" pageId="9" pageNumber="132">
<th id="76A76269B563004806CCFBAEFD43D725" box="[165,700,1060,1083]" gridcol="0" gridrow="13" pageId="9" pageNumber="132">Length of caudal peduncle</th>
<td id="76A76269B56300480540FBAEFC6ED725" box="[809,913,1060,1083]" gridcol="1" gridrow="13" pageId="9" pageNumber="132">23.6</td>
<td id="76A76269B563004805B5FBAEFBB7D725" box="[988,1096,1060,1083]" gridcol="2" gridrow="13" pageId="9" pageNumber="132">2224.3</td>
<td id="76A76269B563004802C1FBAEFAE6D725" box="[1192,1305,1060,1083]" gridcol="3" gridrow="13" pageId="9" pageNumber="132">23.2±0.7</td>
<td id="76A76269B5630048030AFBAEFA89D725" box="[1379,1398,1060,1083]" gridcol="4" gridrow="13" pageId="9" pageNumber="132">7</td>
</tr>
<tr id="35760B15B563004806CCFBC6FA89D77D" box="[165,1398,1100,1123]" gridrow="14" pageId="9" pageNumber="132">
<th id="76A76269B563004806CCFBC6FD43D77D" box="[165,700,1100,1123]" gridcol="0" gridrow="14" pageId="9" pageNumber="132">Length of dorsal fin</th>
<td id="76A76269B56300480540FBC6FC6ED77D" box="[809,913,1100,1123]" gridcol="1" gridrow="14" pageId="9" pageNumber="132">23.3</td>
<td id="76A76269B563004805B5FBC6FBB7D77D" box="[988,1096,1100,1123]" gridcol="2" gridrow="14" pageId="9" pageNumber="132">22.326</td>
<td id="76A76269B563004802C1FBC6FAE6D77D" box="[1192,1305,1100,1123]" gridcol="3" gridrow="14" pageId="9" pageNumber="132">23.5±1.2</td>
<td id="76A76269B5630048030AFBC6FA89D77D" box="[1379,1398,1100,1123]" gridcol="4" gridrow="14" pageId="9" pageNumber="132">7</td>
</tr>
<tr id="35760B15B563004806CCFBFEFA89D795" box="[165,1398,1140,1163]" gridrow="15" pageId="9" pageNumber="132">
<th id="76A76269B563004806CCFBFEFD43D795" box="[165,700,1140,1163]" gridcol="0" gridrow="15" pageId="9" pageNumber="132">Length of dorsal-fin base</th>
<td id="76A76269B56300480540FBFEFC6ED795" box="[809,913,1140,1163]" gridcol="1" gridrow="15" pageId="9" pageNumber="132">16.3</td>
<td id="76A76269B563004805B5FBFEFBB7D795" box="[988,1096,1140,1163]" gridcol="2" gridrow="15" pageId="9" pageNumber="132">15.617.6</td>
<td id="76A76269B563004802C1FBFEFAE6D795" box="[1192,1305,1140,1163]" gridcol="3" gridrow="15" pageId="9" pageNumber="132">16.7±0.8</td>
<td id="76A76269B5630048030AFBFEFA89D795" box="[1379,1398,1140,1163]" gridcol="4" gridrow="15" pageId="9" pageNumber="132">7</td>
</tr>
<tr id="35760B15B563004806CCFB16FA89D7AD" box="[165,1398,1180,1203]" gridrow="16" pageId="9" pageNumber="132">
<th id="76A76269B563004806CCFB16FD43D7AD" box="[165,700,1180,1203]" gridcol="0" gridrow="16" pageId="9" pageNumber="132">Length of pectoral fin</th>
<td id="76A76269B56300480540FB16FC6ED7AD" box="[809,913,1180,1203]" gridcol="1" gridrow="16" pageId="9" pageNumber="132">18.1</td>
<td id="76A76269B563004805B5FB16FBB7D7AD" box="[988,1096,1180,1203]" gridcol="2" gridrow="16" pageId="9" pageNumber="132">17.819.4</td>
<td id="76A76269B563004802C1FB16FAE6D7AD" box="[1192,1305,1180,1203]" gridcol="3" gridrow="16" pageId="9" pageNumber="132">18.3±0.6</td>
<td id="76A76269B5630048030AFB16FA89D7AD" box="[1379,1398,1180,1203]" gridcol="4" gridrow="16" pageId="9" pageNumber="132">7</td>
</tr>
<tr id="35760B15B563004806CCFB4EFA89D7C5" box="[165,1398,1220,1243]" gridrow="17" pageId="9" pageNumber="132">
<th id="76A76269B563004806CCFB4EFD43D7C5" box="[165,700,1220,1243]" gridcol="0" gridrow="17" pageId="9" pageNumber="132">Length of pelvic fin</th>
<td id="76A76269B56300480540FB4EFC6ED7C5" box="[809,913,1220,1243]" gridcol="1" gridrow="17" pageId="9" pageNumber="132">19.2</td>
<td id="76A76269B563004805B5FB4EFBB7D7C5" box="[988,1096,1220,1243]" gridcol="2" gridrow="17" pageId="9" pageNumber="132">17.419.2</td>
<td id="76A76269B563004802C1FB4EFAE6D7C5" box="[1192,1305,1220,1243]" gridcol="3" gridrow="17" pageId="9" pageNumber="132">18.4±0.6</td>
<td id="76A76269B5630048030AFB4EFA89D7C5" box="[1379,1398,1220,1243]" gridcol="4" gridrow="17" pageId="9" pageNumber="132">7</td>
</tr>
<tr id="35760B15B563004806CCFB66FA89D61D" box="[165,1398,1260,1283]" gridrow="18" pageId="9" pageNumber="132">
<th id="76A76269B563004806CCFB66FD43D61D" box="[165,700,1260,1283]" gridcol="0" gridrow="18" pageId="9" pageNumber="132">Length of anal fin</th>
<td id="76A76269B56300480540FB66FC6ED61D" box="[809,913,1260,1283]" gridcol="1" gridrow="18" pageId="9" pageNumber="132">21.0</td>
<td id="76A76269B563004805B5FB66FBB7D61D" box="[988,1096,1260,1283]" gridcol="2" gridrow="18" pageId="9" pageNumber="132">17.821.7</td>
<td id="76A76269B563004802C1FB66FAE6D61D" box="[1192,1305,1260,1283]" gridcol="3" gridrow="18" pageId="9" pageNumber="132">20.6±1.3</td>
<td id="76A76269B5630048030AFB66FA89D61D" box="[1379,1398,1260,1283]" gridcol="4" gridrow="18" pageId="9" pageNumber="132">7</td>
</tr>
<tr id="35760B15B563004806CCFA9EFA89D635" box="[165,1398,1300,1323]" gridrow="19" pageId="9" pageNumber="132">
<th id="76A76269B563004806CCFA9EFD43D635" box="[165,700,1300,1323]" gridcol="0" gridrow="19" pageId="9" pageNumber="132">Length of anal-fin base</th>
<td id="76A76269B56300480540FA9EFC6ED635" box="[809,913,1300,1323]" gridcol="1" gridrow="19" pageId="9" pageNumber="132">8.7</td>
<td id="76A76269B563004805B5FA9EFBB7D635" box="[988,1096,1300,1323]" gridcol="2" gridrow="19" pageId="9" pageNumber="132">7.711.1</td>
<td id="76A76269B563004802C1FA9EFAE6D635" box="[1192,1305,1300,1323]" gridcol="3" gridrow="19" pageId="9" pageNumber="132">9.2±1.3</td>
<td id="76A76269B5630048030AFA9EFA89D635" box="[1379,1398,1300,1323]" gridcol="4" gridrow="19" pageId="9" pageNumber="132">7</td>
</tr>
<tr id="35760B15B563004806CCFAB5FA89D648" box="[165,1398,1343,1366]" gridrow="20" pageId="9" pageNumber="132" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1">
<th id="76A76269B563004806CCFAB5FD43D648" box="[165,700,1343,1366]" gridcol="0" gridrow="20" pageId="9" pageNumber="132">
<emphasis id="B932D545B563FFBE06D0FAB5FEFED648" bold="true" box="[185,257,1343,1366]" pageId="9" pageNumber="132">% HL</emphasis>
</th>
</tr>
<tr id="35760B15B563004806CCFAFEFA89D695" box="[165,1398,1396,1419]" gridrow="21" pageId="9" pageNumber="132">
<th id="76A76269B563004806CCFAFEFD43D695" box="[165,700,1396,1419]" gridcol="0" gridrow="21" pageId="9" pageNumber="132">Head depth</th>
<td id="76A76269B56300480540FAFEFC6ED695" box="[809,913,1396,1419]" gridcol="1" gridrow="21" pageId="9" pageNumber="132">88.2</td>
<td id="76A76269B563004805B5FAFEFBB7D695" box="[988,1096,1396,1419]" gridcol="2" gridrow="21" pageId="9" pageNumber="132">71.489.5</td>
<td id="76A76269B563004802C1FAFEFAE6D695" box="[1192,1305,1396,1419]" gridcol="3" gridrow="21" pageId="9" pageNumber="132">81.1±6.6</td>
<td id="76A76269B5630048030AFAFEFA89D695" box="[1379,1398,1396,1419]" gridcol="4" gridrow="21" pageId="9" pageNumber="132">7</td>
</tr>
<tr id="35760B15B563004806CCFA2EFA89D6A5" box="[165,1398,1444,1467]" gridrow="22" pageId="9" pageNumber="132">
<th id="76A76269B563004806CCFA2EFD43D6A5" box="[165,700,1444,1467]" gridcol="0" gridrow="22" pageId="9" pageNumber="132">Head depth at eye</th>
<td id="76A76269B56300480540FA2EFC6ED6A5" box="[809,913,1444,1467]" gridcol="1" gridrow="22" pageId="9" pageNumber="132">67.6</td>
<td id="76A76269B563004805B5FA2EFBB7D6A5" box="[988,1096,1444,1467]" gridcol="2" gridrow="22" pageId="9" pageNumber="132">57.170.3</td>
<td id="76A76269B563004802C1FA2EFAE6D6A5" box="[1192,1305,1444,1467]" gridcol="3" gridrow="22" pageId="9" pageNumber="132">64.3±5.0</td>
<td id="76A76269B5630048030AFA2EFA89D6A5" box="[1379,1398,1444,1467]" gridcol="4" gridrow="22" pageId="9" pageNumber="132">7</td>
</tr>
<tr id="35760B15B563004806CCFA46FA89D6FD" box="[165,1398,1484,1507]" gridrow="23" pageId="9" pageNumber="132">
<th id="76A76269B563004806CCFA46FD43D6FD" box="[165,700,1484,1507]" gridcol="0" gridrow="23" pageId="9" pageNumber="132">Maximum head width</th>
<td id="76A76269B56300480540FA46FC6ED6FD" box="[809,913,1484,1507]" gridcol="1" gridrow="23" pageId="9" pageNumber="132">67.6</td>
<td id="76A76269B563004805B5FA46FBB7D6FD" box="[988,1096,1484,1507]" gridcol="2" gridrow="23" pageId="9" pageNumber="132">57.167.6</td>
<td id="76A76269B563004802C1FA46FAE6D6FD" box="[1192,1305,1484,1507]" gridcol="3" gridrow="23" pageId="9" pageNumber="132">62.3±3.2</td>
<td id="76A76269B5630048030AFA46FA89D6FD" box="[1379,1398,1484,1507]" gridcol="4" gridrow="23" pageId="9" pageNumber="132">7</td>
</tr>
<tr id="35760B15B563004806CCFA7EFA89D515" box="[165,1398,1524,1547]" gridrow="24" pageId="9" pageNumber="132">
<th id="76A76269B563004806CCFA7EFD43D515" box="[165,700,1524,1547]" gridcol="0" gridrow="24" pageId="9" pageNumber="132">Snout length</th>
<td id="76A76269B56300480540FA7EFC6ED515" box="[809,913,1524,1547]" gridcol="1" gridrow="24" pageId="9" pageNumber="132">35.3</td>
<td id="76A76269B563004805B5FA7EFBB7D515" box="[988,1096,1524,1547]" gridcol="2" gridrow="24" pageId="9" pageNumber="132">33.336.8</td>
<td id="76A76269B563004802C1FA7EFAE6D515" box="[1192,1305,1524,1547]" gridcol="3" gridrow="24" pageId="9" pageNumber="132">34.7±1.3</td>
<td id="76A76269B5630048030AFA7EFA89D515" box="[1379,1398,1524,1547]" gridcol="4" gridrow="24" pageId="9" pageNumber="132">7</td>
</tr>
<tr id="35760B15B563004806CCF996FA89D52D" box="[165,1398,1564,1587]" gridrow="25" pageId="9" pageNumber="132">
<th id="76A76269B563004806CCF996FD43D52D" box="[165,700,1564,1587]" gridcol="0" gridrow="25" pageId="9" pageNumber="132">Interorbital width</th>
<td id="76A76269B56300480540F996FC6ED52D" box="[809,913,1564,1587]" gridcol="1" gridrow="25" pageId="9" pageNumber="132">47.1</td>
<td id="76A76269B563004805B5F996FBB7D52D" box="[988,1096,1564,1587]" gridcol="2" gridrow="25" pageId="9" pageNumber="132">40.555.3</td>
<td id="76A76269B563004802C1F996FAE6D52D" box="[1192,1305,1564,1587]" gridcol="3" gridrow="25" pageId="9" pageNumber="132">47.1±4.6</td>
<td id="76A76269B5630048030AF996FA89D52D" box="[1379,1398,1564,1587]" gridcol="4" gridrow="25" pageId="9" pageNumber="132">7</td>
</tr>
<tr id="35760B15B563004806CCF9CEFA89D545" box="[165,1398,1604,1627]" gridrow="26" pageId="9" pageNumber="132">
<th id="76A76269B563004806CCF9CEFD43D545" box="[165,700,1604,1627]" gridcol="0" gridrow="26" pageId="9" pageNumber="132">Eye diameter</th>
<td id="76A76269B56300480540F9CEFC6ED545" box="[809,913,1604,1627]" gridcol="1" gridrow="26" pageId="9" pageNumber="132">20.6</td>
<td id="76A76269B563004805B5F9CEFBB7D545" box="[988,1096,1604,1627]" gridcol="2" gridrow="26" pageId="9" pageNumber="132">20.323.8</td>
<td id="76A76269B563004802C1F9CEFAE6D545" box="[1192,1305,1604,1627]" gridcol="3" gridrow="26" pageId="9" pageNumber="132">21.7±1.5</td>
<td id="76A76269B5630048030AF9CEFA89D545" box="[1379,1398,1604,1627]" gridcol="4" gridrow="26" pageId="9" pageNumber="132">7</td>
</tr>
<tr id="35760B15B563004806CCF9E6FA89D59D" box="[165,1398,1644,1667]" gridrow="27" pageId="9" pageNumber="132">
<th id="76A76269B563004806CCF9E6FD43D59D" box="[165,700,1644,1667]" gridcol="0" gridrow="27" pageId="9" pageNumber="132">Mouth width</th>
<td id="76A76269B56300480540F9E6FC6ED59D" box="[809,913,1644,1667]" gridcol="1" gridrow="27" pageId="9" pageNumber="132">41.8</td>
<td id="76A76269B563004805B5F9E6FBB7D59D" box="[988,1096,1644,1667]" gridcol="2" gridrow="27" pageId="9" pageNumber="132">33.344.7</td>
<td id="76A76269B563004802C1F9E6FAE6D59D" box="[1192,1305,1644,1667]" gridcol="3" gridrow="27" pageId="9" pageNumber="132">40.5±3.6</td>
<td id="76A76269B5630048030AF9E6FA89D59D" box="[1379,1398,1644,1667]" gridcol="4" gridrow="27" pageId="9" pageNumber="132">7</td>
</tr>
<tr id="35760B15B563004806CCF91EFA89D5B5" box="[165,1398,1684,1707]" gridrow="28" pageId="9" pageNumber="132" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1">
<th id="76A76269B563004806CCF91EFD43D5B5" box="[165,700,1684,1707]" gridcol="0" gridrow="28" pageId="9" pageNumber="132">
<emphasis id="B932D545B563FFBE06D0F91EFEF5D5B5" bold="true" box="[185,266,1684,1707]" pageId="9" pageNumber="132">Counts</emphasis>
</th>
</tr>
<tr id="35760B15B563004806CCF937FA89D5CA" box="[165,1398,1725,1748]" gridrow="29" pageId="9" pageNumber="132">
<th id="76A76269B563004806CCF937FD43D5CA" box="[165,700,1725,1748]" gridcol="0" gridrow="29" pageId="9" pageNumber="132">Lateral-line scales</th>
<td id="76A76269B56300480540F937FC6ED5CA" box="[809,913,1725,1748]" gridcol="1" gridrow="29" pageId="9" pageNumber="132">47</td>
<td id="76A76269B563004805B5F937FBB7D5CA" box="[988,1096,1725,1748]" gridcol="2" gridrow="29" pageId="9" pageNumber="132">4447</td>
<td id="76A76269B563004802C1F937FAE6D5CA" box="[1192,1305,1725,1748]" gridcol="3" gridrow="29" pageId="9" pageNumber="132">46.1±1.2</td>
<td id="76A76269B5630048030AF937FA89D5CA" box="[1379,1398,1725,1748]" gridcol="4" gridrow="29" pageId="9" pageNumber="132">7</td>
</tr>
<tr id="35760B15B563004806CCF96FFA89D5E2" box="[165,1398,1765,1788]" gridrow="30" pageId="9" pageNumber="132">
<th id="76A76269B563004806CCF96FFD43D5E2" box="[165,700,1765,1788]" gridcol="0" gridrow="30" pageId="9" pageNumber="132">Scales between lateral line and origin of dorsal fin</th>
<td id="76A76269B56300480540F96FFC6ED5E2" box="[809,913,1765,1788]" gridcol="1" gridrow="30" pageId="9" pageNumber="132">7</td>
<td id="76A76269B563004805B5F96FFBB7D5E2" box="[988,1096,1765,1788]" gridcol="2" gridrow="30" pageId="9" pageNumber="132">7</td>
<td id="76A76269B563004802C1F96FFAE6D5E2" box="[1192,1305,1765,1788]" gridcol="3" gridrow="30" pageId="9" pageNumber="132">7.0±0.0</td>
<td id="76A76269B5630048030AF96FFA89D5E2" box="[1379,1398,1765,1788]" gridcol="4" gridrow="30" pageId="9" pageNumber="132">7</td>
</tr>
<tr id="35760B15B563004806CCF887FA89D43A" box="[165,1398,1805,1828]" gridrow="31" pageId="9" pageNumber="132">
<th id="76A76269B563004806CCF887FD43D43A" box="[165,700,1805,1828]" gridcol="0" gridrow="31" pageId="9" pageNumber="132">Scales between lateral line and origin of pelvic fin</th>
<td id="76A76269B56300480540F887FC6ED43A" box="[809,913,1805,1828]" gridcol="1" gridrow="31" pageId="9" pageNumber="132">5</td>
<td id="76A76269B563004805B5F887FBB7D43A" box="[988,1096,1805,1828]" gridcol="2" gridrow="31" pageId="9" pageNumber="132">5</td>
<td id="76A76269B563004802C1F887FAE6D43A" box="[1192,1305,1805,1828]" gridcol="3" gridrow="31" pageId="9" pageNumber="132">5.0±0.0</td>
<td id="76A76269B5630048030AF887FA89D43A" box="[1379,1398,1805,1828]" gridcol="4" gridrow="31" pageId="9" pageNumber="132">7</td>
</tr>
<tr id="35760B15B563004806CCF8BFFA89D452" box="[165,1398,1845,1868]" gridrow="32" pageId="9" pageNumber="132">
<th id="76A76269B563004806CCF8BFFD43D452" box="[165,700,1845,1868]" gridcol="0" gridrow="32" pageId="9" pageNumber="132">Scales between lateral line and origin of anal fin</th>
<td id="76A76269B56300480540F8BFFC6ED452" box="[809,913,1845,1868]" gridcol="1" gridrow="32" pageId="9" pageNumber="132">5</td>
<td id="76A76269B563004805B5F8BFFBB7D452" box="[988,1096,1845,1868]" gridcol="2" gridrow="32" pageId="9" pageNumber="132">5</td>
<td id="76A76269B563004802C1F8BFFAE6D452" box="[1192,1305,1845,1868]" gridcol="3" gridrow="32" pageId="9" pageNumber="132">5.6±0.5</td>
<td id="76A76269B5630048030AF8BFFA89D452" box="[1379,1398,1845,1868]" gridcol="4" gridrow="32" pageId="9" pageNumber="132">7</td>
</tr>
<tr id="35760B15B563004806CCF8D7FA89D46A" box="[165,1398,1885,1908]" gridrow="33" pageId="9" pageNumber="132">
<th id="76A76269B563004806CCF8D7FD43D46A" box="[165,700,1885,1908]" gridcol="0" gridrow="33" pageId="9" pageNumber="132">Circumpeduncular scales</th>
<td id="76A76269B56300480540F8D7FC6ED46A" box="[809,913,1885,1908]" gridcol="1" gridrow="33" pageId="9" pageNumber="132">16</td>
<td id="76A76269B563004805B5F8D7FBB7D46A" box="[988,1096,1885,1908]" gridcol="2" gridrow="33" pageId="9" pageNumber="132">1616</td>
<td id="76A76269B563004802C1F8D7FAE6D46A" box="[1192,1305,1885,1908]" gridcol="3" gridrow="33" pageId="9" pageNumber="132">16.0±0.0</td>
<td id="76A76269B5630048030AF8D7FA89D46A" box="[1379,1398,1885,1908]" gridcol="4" gridrow="33" pageId="9" pageNumber="132">7</td>
</tr>
<tr id="35760B15B563004806CCF80FFA89D482" box="[165,1398,1925,1948]" gridrow="34" pageId="9" pageNumber="132">
<th id="76A76269B563004806CCF80FFD43D482" box="[165,700,1925,1948]" gridcol="0" gridrow="34" pageId="9" pageNumber="132">Predorsal scales</th>
<td id="76A76269B56300480540F80FFC6ED482" box="[809,913,1925,1948]" gridcol="1" gridrow="34" pageId="9" pageNumber="132">15</td>
<td id="76A76269B563004805B5F80FFBB7D482" box="[988,1096,1925,1948]" gridcol="2" gridrow="34" pageId="9" pageNumber="132">1415</td>
<td id="76A76269B563004802C1F80FFAE6D482" box="[1192,1305,1925,1948]" gridcol="3" gridrow="34" pageId="9" pageNumber="132">14.7±0.5</td>
<td id="76A76269B5630048030AF80FFA89D482" box="[1379,1398,1925,1948]" gridcol="4" gridrow="34" pageId="9" pageNumber="132">7</td>
</tr>
</table>
</paragraph>
<paragraph id="8BF90957B560FFBD06AEFF1DFCFED3E6" blockId="10.[151,1437,151,1979]" pageId="10" pageNumber="133">Body high, moderately compressed; caudal peduncle slender, about 2.42.6 times longer than deep. Dorsal body profile convex, ventral profile rounded. Lateral line complete, 4547 scales; 1415 predorsal scales; 7/1/5 scales in transverse row anterior to pelvic-fin insertion.</paragraph>
<paragraph id="8BF90957B560FFBD06AEFE8EFAE9D2CE" blockId="10.[151,1437,151,1979]" pageId="10" pageNumber="133">
Dorsal fin high with 4 simple and 8 branched rays; first simple ray as a tiny process, vestigial in a few specimens; last simple ray strongly serrated; dorsal-fin origin inserted slightly in front of vertical with pelvic-fin origin; distal margin anteriorly concave and posteriorly convex. Pectoral fin pointed with 1 simple and 16 branched rays. Pelvic fin pointed, with 1 simple and 8 branched rays; axillary scale present. Anus immediately in front of anal fin. Anal fin with 3 simple and 5 branched rays; short (17.8% SL) and small in male; long (20.021.7% SL) and large in female (
<figureCitation id="137D15D2B560FFBD0714FE3DFE40D2CE" box="[381,447,439,464]" captionStart="FIGURE 5" captionStartId="8.[151,250,1965,1987]" captionTargetBox="[163,1419,473,1929]" captionTargetId="figure@8.[151,1435,447,1940]" captionTargetPageId="8" captionText="FIGURE 5. Lateral view of Onychostoma dongnaiensis specimens (a) female holotype UNS 00851 in life, (b) female holotype UNS 00851 in preservative, (c) male paratype ZRC 54625 in preservative. Scale 10 mm." httpUri="https://zenodo.org/record/243919/files/figure.png" pageId="10" pageNumber="133">Fig. 5</figureCitation>
b, c). Caudal fin deeply forked with 9+8 principal rays, 8+7 being branched.
</paragraph>
<paragraph id="8BF90957B560FFBD06AEFE56FE5AD1EE" blockId="10.[151,1437,151,1979]" pageId="10" pageNumber="133">
<emphasis id="B932D545B560FFBD06AEFE56FE92D2EB" bold="true" box="[199,365,476,501]" pageId="10" pageNumber="133">Colour in life.</emphasis>
Head dark turquoise on back, light turquoise around orbital and on side, white on opercula and lower jaw. Body light turquoise on back, silver white on lateroventral surface, snowy white belly with a longitudinal black stripe running along lateral line. Scales silverly bluish between back and lateral line; lateral-line scales and row of scales just above lateral line in anterior half of body pigmented at center of scale bases. Fins hyaline. Dorsal fin dark turquoise at origin, bold on rays; distal margin concave anteriorly and convex posteriorly; reddish at distal part. Pectoral fin and pelvic fin yellowish green and reddish at first rays and distal parts. Anal fin yellowish green on rays, pinkish distally and hyaline on distal margin. Caudal fin near peduncle dark turquoise, yellowish to red on tips.
</paragraph>
<paragraph id="8BF90957B560FFBD06AEFD76FE0DD09E" blockId="10.[151,1437,151,1979]" pageId="10" pageNumber="133">
<emphasis id="B932D545B560FFBD06AEFD76FE25D00B" bold="true" box="[199,474,764,789]" pageId="10" pageNumber="133">Colour in preservative.</emphasis>
Similar to that of fresh condition except as noted below. Upper half of body including head brown. Lower half of body including head pale ivory colour except anterior half of body darker. Pigments at scale bases on almost whole body outstanding. All turquoise, yellowish, silverly bluish, reddish and snowy whitish colors disappeared (
<figureCitation id="137D15D2B560FFBD071EFCEDFE45D09E" box="[375,442,871,896]" captionStart="FIGURE 5" captionStartId="8.[151,250,1965,1987]" captionTargetBox="[163,1419,473,1929]" captionTargetId="figure@8.[151,1435,447,1940]" captionTargetPageId="8" captionText="FIGURE 5. Lateral view of Onychostoma dongnaiensis specimens (a) female holotype UNS 00851 in life, (b) female holotype UNS 00851 in preservative, (c) male paratype ZRC 54625 in preservative. Scale 10 mm." httpUri="https://zenodo.org/record/243919/files/figure.png" pageId="10" pageNumber="133">Fig. 5</figureCitation>
b, c).
</paragraph>
<paragraph id="8BF90957B560FFBD06AEFC06FC02D7BC" blockId="10.[151,1437,151,1979]" pageId="10" pageNumber="133">
<emphasis id="B932D545B560FFBD06AEFC06FED4D0BB" bold="true" box="[199,299,908,933]" pageId="10" pageNumber="133">Ecology.</emphasis>
All specimens of the new species were found in the Da Teh catchment of middle Dong Nai drainage in evergreen forest between
<quantity id="4CBEA4B2B560FFBD0783FC3AFD8FD0D9" box="[490,624,944,968]" metricMagnitude="2" metricUnit="m" metricValue="1.7" metricValueMax="2.0" metricValueMin="1.4" pageId="10" pageNumber="133" unit="m" value="170.0" valueMax="200.0" valueMin="140.0">140200 m</quantity>
(
<figureCitation id="137D15D2B560FFBD04ECFC25FD30D0D6" box="[645,719,943,968]" captionStart="FIGURE 4" captionStartId="6.[151,250,1617,1639]" captionTargetBox="[161,1426,711,1582]" captionTargetId="figure@6.[151,1435,700,1593]" captionTargetPageId="6" captionText="FIGURE 4. High elevation main river at type locality of Onychostoma krongnoensis. Habitats altering from fast flowing pool (a) with bed of detritus and falling leaves, running to riffles (c) and swift current (b) over bedrock, large boulder and cobble substrates covered with periphyton (sites continuous); low elevation main river at type locality of Onychostoma dongnaiensis (d) running to riffles." httpUri="https://zenodo.org/record/243918/files/figure.png" pageId="10" pageNumber="133">Fig. 4</figureCitation>
d).
<taxonomicName id="4C4672D4B560FFBD0491FC25FBCDD0D6" box="[760,1074,943,968]" class="Actinopterygii" family="Cyprinidae" genus="Onychostoma" kingdom="Animalia" order="Cypriniformes" pageId="10" pageNumber="133" phylum="Chordata" rank="species" species="dongnaiensis">
<emphasis id="B932D545B560FFBD0491FC25FBCDD0D6" box="[760,1074,943,968]" italics="true" pageId="10" pageNumber="133">Onychostoma dongnaiensis</emphasis>
</taxonomicName>
lives in clear water but some specimens were collected in silty water. Water conditions of 26.428.1°C,
<collectingCountry id="F35149C7B560FFBD0243FC5FFBB3D0F2" box="[1066,1100,981,1004]" name="Philippines" pageId="10" pageNumber="133">pH</collectingCountry>
7.027.6, DO 82.791.4%, conductivity 1225 µS.cm -1 and flow velocity
<quantity id="4CBEA4B2B560FFBD04F6FC71FCD1D70D" box="[671,814,1019,1043]" metricMagnitude="-1" metricUnit="m" metricValue="4.45" metricValueMax="7.7" metricValueMin="1.2" pageId="10" pageNumber="133" unit="m" value="0.445" valueMax="0.77" valueMin="0.12">0.120.77 m</quantity>
/s were recorded. The species occurred in swift currents over bedrock, large boulders and cobble substrates covered with periphyton (
<figureCitation id="137D15D2B560FFBD0590FB94FBC2D729" box="[1017,1085,1054,1079]" captionStart="FIGURE 4" captionStartId="6.[151,250,1617,1639]" captionTargetBox="[161,1426,711,1582]" captionTargetId="figure@6.[151,1435,700,1593]" captionTargetPageId="6" captionText="FIGURE 4. High elevation main river at type locality of Onychostoma krongnoensis. Habitats altering from fast flowing pool (a) with bed of detritus and falling leaves, running to riffles (c) and swift current (b) over bedrock, large boulder and cobble substrates covered with periphyton (sites continuous); low elevation main river at type locality of Onychostoma dongnaiensis (d) running to riffles." httpUri="https://zenodo.org/record/243918/files/figure.png" pageId="10" pageNumber="133">Fig. 4</figureCitation>
d). This is similar to the habitat of
<taxonomicName id="4C4672D4B560FFBD06D2FBC9FE87D745" box="[187,376,1090,1115]" class="Actinopterygii" family="Cyprinidae" genus="Onychostoma" kingdom="Animalia" order="Cypriniformes" pageId="10" pageNumber="133" phylum="Chordata" rank="species" species="krongnoensis">
<emphasis id="B932D545B560FFBD06D2FBC9FE87D745" box="[187,376,1090,1115]" italics="true" pageId="10" pageNumber="133">
<collectingCountry id="F35149C7B560FFBD06D2FBC9FF2DD744" box="[187,210,1091,1114]" name="Iceland" pageId="10" pageNumber="133">O</collectingCountry>
. krongnoensis
</emphasis>
</taxonomicName>
. When feeding,
<taxonomicName id="4C4672D4B560FFBD042DFBC9FCFFD745" box="[580,768,1090,1115]" class="Actinopterygii" family="Cyprinidae" genus="Onychostoma" kingdom="Animalia" order="Cypriniformes" pageId="10" pageNumber="133" phylum="Chordata" rank="species" species="dongnaiensis">
<emphasis id="B932D545B560FFBD042DFBC9FCFFD745" box="[580,768,1090,1115]" italics="true" pageId="10" pageNumber="133">
<collectingCountry id="F35149C7B560FFBD042DFBC9FDA4D744" box="[580,603,1091,1114]" name="Iceland" pageId="10" pageNumber="133">O</collectingCountry>
. dongnaiensis
</emphasis>
</taxonomicName>
flips rocks and scrapes algae from them.
<taxonomicName id="4C4672D4B560FFBD036BFBC8FED4D761" class="Actinopterygii" family="Cyprinidae" genus="Onychostoma" kingdom="Animalia" order="Cypriniformes" pageId="10" pageNumber="133" phylum="Chordata" rank="species" species="dongnaiensis">
<emphasis id="B932D545B560FFBD036BFBC8FED4D761" italics="true" pageId="10" pageNumber="133">Onychostoma dongnaiensis</emphasis>
</taxonomicName>
is found with
<taxonomicName id="4C4672D4B560FFBD0782FBECFCD3D761" box="[491,812,1126,1151]" class="Actinopterygii" family="Cyprinidae" genus="Mystacoleucus" kingdom="Animalia" order="Cypriniformes" pageId="10" pageNumber="133" phylum="Chordata" rank="species" species="obtusirostris">
<emphasis id="B932D545B560FFBD0782FBECFCD3D761" box="[491,812,1126,1151]" italics="true" pageId="10" pageNumber="133">Mystacoleucus obtusirostris</emphasis>
</taxonomicName>
, the bonylip barb
<taxonomicName id="4C4672D4B560FFBD027EFBECFB02D761" box="[1047,1277,1126,1151]" class="Actinopterygii" family="Cyprinidae" genus="Osteochilus" kingdom="Animalia" order="Cypriniformes" pageId="10" pageNumber="133" phylum="Chordata" rank="species" species="vittatus">
<emphasis id="B932D545B560FFBD027EFBECFB02D761" box="[1047,1277,1126,1151]" italics="true" pageId="10" pageNumber="133">Osteochilus vittatus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C4672D4B560FFBD037AFBE2FEF8D7BD" class="Actinopterygii" family="Cyprinidae" genus="Poropuntius" kingdom="Animalia" order="Cypriniformes" pageId="10" pageNumber="133" phylum="Chordata" rank="species" species="deauratus">
<emphasis id="B932D545B560FFBD037AFBE2FEF8D7BD" italics="true" pageId="10" pageNumber="133">Poropuntius deauratus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C4672D4B560FFBD077CFB00FE38D7BD" box="[277,455,1162,1187]" class="Actinopterygii" family="Channidae" genus="Channa" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="133" phylum="Chordata" rank="species" species="gachua">
<emphasis id="B932D545B560FFBD077CFB00FE38D7BD" box="[277,455,1162,1187]" italics="true" pageId="10" pageNumber="133">Channa gachua</emphasis>
</taxonomicName>
,
<taxonomicName id="4C4672D4B560FFBD07BCFB00FD36D7BD" box="[469,713,1162,1187]" class="Actinopterygii" family="Cobitidae" genus="Yasuhikotakia" kingdom="Animalia" order="Cypriniformes" pageId="10" pageNumber="133" phylum="Chordata" rank="species" species="morleti">
<emphasis id="B932D545B560FFBD07BCFB00FD36D7BD" box="[469,713,1162,1187]" italics="true" pageId="10" pageNumber="133">Yasuhikotakia morleti</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C4672D4B560FFBD056FFB01FC08D7BD" box="[774,1015,1162,1187]" class="Actinopterygii" family="Cobitidae" genus="Acantopsis" kingdom="Animalia" order="Cypriniformes" pageId="10" pageNumber="133" phylum="Chordata" rank="species" species="dialuzona">
<emphasis id="B932D545B560FFBD056FFB01FC08D7BD" box="[774,1015,1162,1187]" italics="true" pageId="10" pageNumber="133">Acantopsis dialuzona</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BF90957B560FFBD06AEFB24FEE4D515" blockId="10.[151,1437,151,1979]" pageId="10" pageNumber="133">
<emphasis id="B932D545B560FFBD06AEFB24FE44D7D9" bold="true" box="[199,443,1198,1223]" pageId="10" pageNumber="133">Conservation status.</emphasis>
The conservation status of this new species requires particular attention. It has been long recognized by the local Mạ minority people. It is likely to be restricted to relatively small stretches of low-elevation forest streams of the middle Dong Nai drainage and hence particularly vulnerable to threatening processes such as siltation and overfishing. Siltation (from road construction, deforestation, agriculture etc.) which covers rocks and stones is a particular threat to this species as it covers the algae growing on the stones (
<figureCitation id="137D15D2B560FFBD0201FAB4FB53D649" box="[1128,1196,1342,1367]" captionStart="FIGURE 4" captionStartId="6.[151,250,1617,1639]" captionTargetBox="[161,1426,711,1582]" captionTargetId="figure@6.[151,1435,700,1593]" captionTargetPageId="6" captionText="FIGURE 4. High elevation main river at type locality of Onychostoma krongnoensis. Habitats altering from fast flowing pool (a) with bed of detritus and falling leaves, running to riffles (c) and swift current (b) over bedrock, large boulder and cobble substrates covered with periphyton (sites continuous); low elevation main river at type locality of Onychostoma dongnaiensis (d) running to riffles." httpUri="https://zenodo.org/record/243918/files/figure.png" pageId="10" pageNumber="133">Fig. 4</figureCitation>
d). This species feeds on the algae and animals living in the algae. Anthropogenic modification of stream morphology, logging, deforestation, agriculture and overfishing frequently occur in the Da Teh catchment. These activities impact the aquatic environment of this species so its populations are potentially threatened. Given the available information, we suggest the species should be considered Data Deficient following IUCNs Red List categories (IUCN 2014, version 11).
</paragraph>
<paragraph id="8BF90957B560FFBD06AEF99CFC46D4A5" blockId="10.[151,1437,151,1979]" pageId="10" pageNumber="133">
<emphasis id="B932D545B560FFBD06AEF99CFE93D531" bold="true" box="[199,364,1558,1583]" pageId="10" pageNumber="133">Comparisons.</emphasis>
In general,
<taxonomicName id="4C4672D4B560FFBD079BF99DFD57D531" box="[498,680,1558,1583]" class="Actinopterygii" family="Cyprinidae" genus="Onychostoma" kingdom="Animalia" order="Cypriniformes" pageId="10" pageNumber="133" phylum="Chordata" rank="species" species="dongnaiensis">
<collectingCountry id="F35149C7B560FFBD079BF99DFDF9D530" box="[498,518,1559,1582]" name="Iceland" pageId="10" pageNumber="133">
<emphasis id="B932D545B560FFBD079BF99DFDF9D530" box="[498,518,1559,1582]" italics="true" pageId="10" pageNumber="133">O</emphasis>
</collectingCountry>
.
<emphasis id="B932D545B560FFBD047DF99CFD57D531" box="[532,680,1558,1583]" italics="true" pageId="10" pageNumber="133">dongnaiensis</emphasis>
</taxonomicName>
most closely resembles
<taxonomicName id="4C4672D4B560FFBD05D3F99DFBD2D531" box="[954,1069,1558,1583]" class="Actinopterygii" family="Cyprinidae" genus="Onychostoma" kingdom="Animalia" order="Cypriniformes" pageId="10" pageNumber="133" phylum="Chordata" rank="species" species="leptura">
<collectingCountry id="F35149C7B560FFBD05D3F99DFC31D530" box="[954,974,1559,1582]" name="Iceland" pageId="10" pageNumber="133">
<emphasis id="B932D545B560FFBD05D3F99DFC31D530" box="[954,974,1559,1582]" italics="true" pageId="10" pageNumber="133">O</emphasis>
</collectingCountry>
.
<emphasis id="B932D545B560FFBD05B5F99CFBD2D531" box="[988,1069,1558,1583]" italics="true" pageId="10" pageNumber="133">leptura</emphasis>
</taxonomicName>
and
<taxonomicName id="4C4672D4B560FFBD020AF99DFB1DD531" box="[1123,1250,1558,1583]" class="Actinopterygii" family="Cyprinidae" genus="Barbus" kingdom="Animalia" order="Cypriniformes" pageId="10" pageNumber="133" phylum="Chordata" rank="species" species="gerlachi">
<collectingCountry id="F35149C7B560FFBD020AF99DFB88D530" box="[1123,1143,1559,1582]" name="Iceland" pageId="10" pageNumber="133">
<emphasis id="B932D545B560FFBD020AF99DFB88D530" box="[1123,1143,1559,1582]" italics="true" pageId="10" pageNumber="133">O</emphasis>
</collectingCountry>
.
<emphasis id="B932D545B560FFBD02ECF99CFB1DD531" box="[1157,1250,1558,1583]" italics="true" pageId="10" pageNumber="133">gerlachi</emphasis>
</taxonomicName>
when compared with congeners in the Mekong basin and Red river basin. From detailed measurements and comparisons of specimens of both species,
<taxonomicName id="4C4672D4B560FFBD07AEF9D5FD82D569" box="[455,637,1630,1655]" class="Actinopterygii" family="Cyprinidae" genus="Onychostoma" kingdom="Animalia" order="Cypriniformes" pageId="10" pageNumber="133" phylum="Chordata" rank="species" species="dongnaiensis">
<collectingCountry id="F35149C7B560FFBD07AEF9D5FE24D568" box="[455,475,1631,1654]" name="Iceland" pageId="10" pageNumber="133">
<emphasis id="B932D545B560FFBD07AEF9D5FE24D568" box="[455,475,1631,1654]" italics="true" pageId="10" pageNumber="133">O</emphasis>
</collectingCountry>
.
<emphasis id="B932D545B560FFBD0780F9D4FD82D569" box="[489,637,1630,1655]" italics="true" pageId="10" pageNumber="133">dongnaiensis</emphasis>
</taxonomicName>
differs from
<taxonomicName id="4C4672D4B560FFBD0579F9D5FC7CD569" box="[784,899,1630,1655]" class="Actinopterygii" family="Cyprinidae" genus="Onychostoma" kingdom="Animalia" order="Cypriniformes" pageId="10" pageNumber="133" phylum="Chordata" rank="species" species="leptura">
<emphasis id="B932D545B560FFBD0579F9D5FC7CD569" box="[784,899,1630,1655]" italics="true" pageId="10" pageNumber="133">
<collectingCountry id="F35149C7B560FFBD0579F9D5FCD8D568" box="[784,807,1631,1654]" name="Iceland" pageId="10" pageNumber="133">O</collectingCountry>
. leptura
</emphasis>
</taxonomicName>
in having scales in transverse row 7/1/5 vs. 7/1/ 4, predorsal scales 1415 vs. 1213, eye small with diameter 20.323.8 vs. 22.930.3% HL and serrated last simple dorsal ray vs. smooth last simple dorsal ray.
<taxonomicName id="4C4672D4B560FFBD04B5F92CFBEED5A1" box="[732,1041,1702,1727]" class="Actinopterygii" family="Cyprinidae" genus="Onychostoma" kingdom="Animalia" order="Cypriniformes" pageId="10" pageNumber="133" phylum="Chordata" rank="species" species="dongnaiensis">
<emphasis id="B932D545B560FFBD04B5F92CFBEED5A1" box="[732,1041,1702,1727]" italics="true" pageId="10" pageNumber="133">Onychostoma dongnaiensis</emphasis>
</taxonomicName>
differs from
<taxonomicName id="4C4672D4B560FFBD02CCF92DFADAD5A1" box="[1189,1317,1702,1727]" class="Actinopterygii" family="Cyprinidae" genus="Barbus" kingdom="Animalia" order="Cypriniformes" pageId="10" pageNumber="133" phylum="Chordata" rank="species" species="gerlachi">
<collectingCountry id="F35149C7B560FFBD02CCF92DFB46D5A0" box="[1189,1209,1703,1726]" name="Iceland" pageId="10" pageNumber="133">
<emphasis id="B932D545B560FFBD02CCF92DFB46D5A0" box="[1189,1209,1703,1726]" italics="true" pageId="10" pageNumber="133">O</emphasis>
</collectingCountry>
.
<emphasis id="B932D545B560FFBD02A1F92CFADAD5A1" box="[1224,1317,1702,1727]" italics="true" pageId="10" pageNumber="133">gerlachi</emphasis>
</taxonomicName>
by having predorsal scales 1415 vs. 1215, caudal peduncle 23.2 vs. 21.2% SL, prepectoral length 18.7 21.7 vs. 21.426.5% SL, eye small with diameter 20.323.8% vs. 27.435% HL.
<taxonomicName id="4C4672D4B560FFBD05BEF964FAF2D419" box="[983,1293,1774,1799]" class="Actinopterygii" family="Cyprinidae" genus="Onychostoma" kingdom="Animalia" order="Cypriniformes" pageId="10" pageNumber="133" phylum="Chordata" rank="species" species="dongnaiensis">
<emphasis id="B932D545B560FFBD05BEF964FAF2D419" box="[983,1293,1774,1799]" italics="true" pageId="10" pageNumber="133">Onychostoma dongnaiensis</emphasis>
</taxonomicName>
differs from
<taxonomicName id="4C4672D4B560FFBD06FEF899FEBFD435" box="[151,320,1810,1835]" class="Actinopterygii" family="Cyprinidae" genus="Onychostoma" kingdom="Animalia" order="Cypriniformes" pageId="10" pageNumber="133" phylum="Chordata" rank="species" species="meridionale">
<emphasis id="B932D545B560FFBD06FEF899FEBFD435" box="[151,320,1810,1835]" italics="true" pageId="10" pageNumber="133">
<collectingCountry id="F35149C7B560FFBD06FEF899FF51D434" box="[151,174,1811,1834]" name="Iceland" pageId="10" pageNumber="133">O</collectingCountry>
. meridionale
</emphasis>
</taxonomicName>
in having mouth 1.41.6 vs. 2.22.3 times in maximum HW (
<figureCitation id="137D15D2B560FFBD0596F898FBBDD435" box="[1023,1090,1810,1835]" captionStart="FIGURE 3" captionStartId="5.[151,250,1842,1864]" captionTargetBox="[172,1412,529,1810]" captionTargetId="figure@5.[163,1423,519,1819]" captionTargetPageId="5" captionText="FIGURE 3. Front view of head of Onychostoma krongnoensis (a) female holotype UNS 00805, 205 mm SL, (b) immature paratype UNS 00821, 123 mm SL, (c) male paratype UNS 00825, 214 mm SL; O. dongnaiensis (e) female holotype UNS 00851, 172 mm SL, (f) female paratype UNS 00852, 196 mm SL, (g) male paratype ZRC 54625, 157 mm SL; O. leptura (i) female UNS 00684, 186 mm SL. Ventral view of head of O. krongnoensis, wide mouth (d) male paratype UNS 00825; O. dongnaiensis, wide mouth (h) female holotype UNS 00851; O. leptura, wide mouth (j) female UNS 00684; O. meridionale, moderate mouth (k) immature UNS 00826, 108 mm SL; O. uniforme, narrow mouth (l) female UNS 00689, 130 mm SL. Scale 10 mm." httpUri="https://zenodo.org/record/243917/files/figure.png" pageId="10" pageNumber="133">Fig. 3</figureCitation>
h, k), the last simple dorsal ray strongly serrated vs. not serrated posteriorly, eye diameter 4.24.9 vs. 3.74.5 times in HL, and scales in transverse row 7/1/5 vs. 6.5/1/6.5.
<taxonomicName id="4C4672D4B560FFBD07CDF8D0FD26D46D" box="[420,729,1882,1907]" class="Actinopterygii" family="Cyprinidae" genus="Onychostoma" kingdom="Animalia" order="Cypriniformes" pageId="10" pageNumber="133" phylum="Chordata" rank="species" species="dongnaiensis">
<emphasis id="B932D545B560FFBD07CDF8D0FD26D46D" box="[420,729,1882,1907]" italics="true" pageId="10" pageNumber="133">Onychostoma dongnaiensis</emphasis>
</taxonomicName>
differs from
<taxonomicName id="4C4672D4B560FFBD0507F8D1FC04D46D" box="[878,1019,1883,1907]" class="Actinopterygii" family="Cyprinidae" genus="Onychostoma" kingdom="Animalia" order="Cypriniformes" pageId="10" pageNumber="133" phylum="Chordata" rank="species" species="fusiforme">
<emphasis id="B932D545B560FFBD0507F8D1FC04D46D" box="[878,1019,1883,1907]" italics="true" pageId="10" pageNumber="133">
<collectingCountry id="F35149C7B560FFBD0507F8D1FC7AD46C" box="[878,901,1883,1906]" name="Iceland" pageId="10" pageNumber="133">O</collectingCountry>
. fusiforme
</emphasis>
</taxonomicName>
in having mouth 1.41.6 vs. 2.02.2 times in maximum HW, caudal peduncle 2.22.6 vs. 3.03.6 times longer than deep, eye diameter 4.24.9 vs. 3.74.4 times in HL, and scales in transverse row 7/1/5 vs. 6.5/1/44.5.
</paragraph>
<paragraph id="8BF90957B561FFBC06AEFF1DFC7AD203" blockId="11.[151,1437,151,752]" pageId="11" pageNumber="134">
<bibRefCitation id="EFD774A6B561FFBC06AEFF1DFE7BD3AE" box="[199,388,151,176]" pageId="11" pageNumber="134" refString="Mai, D. Y., Nguyen, V. T., Nguyen, V. T., Le, H. Y. &amp; Hua, B. L. (1992) Identification of Freshwater Fishes of southern Vietnam. Nha Xuat Ban Khoa Hoc va Ky Thuat, Ha Noi, 351 pp. [in Vietnamese]" type="book">
Mai
<emphasis id="B932D545B561FFBC0692FF13FED3D3AE" box="[251,300,151,176]" italics="true" pageId="11" pageNumber="134">et al</emphasis>
. (1992)
</bibRefCitation>
described one specimen as the name
<emphasis id="B932D545B561FFBC0541FF1DFC11D3AE" box="[808,1006,151,176]" italics="true" pageId="11" pageNumber="134">Scaphidonichthys</emphasis>
sp. from Phu Lap, Tan Phu, Dong Nai province near the Dong Nai River at elevation (
<geoCoordinate id="EE726F90B561FFBC04C5FF36FC9DD3CA" box="[684,866,188,212]" direction="north" orientation="latitude" pageId="11" pageNumber="134" precision="1" value="11.454786">11°2717.23” N</geoCoordinate>
,
<geoCoordinate id="EE726F90B561FFBC0504FF36FBDFD3CA" box="[877,1056,188,212]" direction="east" orientation="longitude" pageId="11" pageNumber="134" precision="1" value="107.48455">107°294.40” E</geoCoordinate>
,
<quantity id="4CBEA4B2B561FFBC0245FF36FB88D3CA" box="[1068,1143,188,213]" metricMagnitude="2" metricUnit="m" metricValue="1.55" pageId="11" pageNumber="134" unit="m" value="155.0">155 m</quantity>
). We consider this species conspecific with
<taxonomicName id="4C4672D4B561FFBC0733FF6AFDEED3E6" box="[346,529,223,248]" class="Actinopterygii" family="Cyprinidae" genus="Onychostoma" kingdom="Animalia" order="Cypriniformes" pageId="11" pageNumber="134" phylum="Chordata" rank="species" species="dongnaiensis">
<collectingCountry id="F35149C7B561FFBC0733FF6AFE91D3E9" box="[346,366,224,247]" name="Iceland" pageId="11" pageNumber="134">
<emphasis id="B932D545B561FFBC0733FF6AFE91D3E9" box="[346,366,224,247]" italics="true" pageId="11" pageNumber="134">O</emphasis>
</collectingCountry>
.
<emphasis id="B932D545B561FFBC0714FF55FDEED3E6" box="[381,529,223,248]" italics="true" pageId="11" pageNumber="134">dongnaiensis</emphasis>
</taxonomicName>
based on examination of its morphometrics (
<bibRefCitation id="EFD774A6B561FFBC0270FF55FB33D3E6" author="Mai" box="[1049,1228,223,248]" pageId="11" pageNumber="134" refString="Mai, D. Y., Nguyen, V. T., Nguyen, V. T., Le, H. Y. &amp; Hua, B. L. (1992) Identification of Freshwater Fishes of southern Vietnam. Nha Xuat Ban Khoa Hoc va Ky Thuat, Ha Noi, 351 pp. [in Vietnamese]" type="book" year="1992">
Mai
<emphasis id="B932D545B561FFBC0239FF6BFB7CD3E6" box="[1104,1155,223,248]" italics="true" pageId="11" pageNumber="134">et al</emphasis>
. 1992
</bibRefCitation>
) similar to our
<taxonomicName id="4C4672D4B561FFBC03E8FF6AFED4D203" class="Actinopterygii" family="Cyprinidae" genus="Onychostoma" kingdom="Animalia" order="Cypriniformes" pageId="11" pageNumber="134" phylum="Chordata" rank="species" species="dongnaiensis">
<collectingCountry id="F35149C7B561FFBC03E8FF6AFA6AD3E9" box="[1409,1429,224,247]" name="Iceland" pageId="11" pageNumber="134">
<emphasis id="B932D545B561FFBC03E8FF6AFA6AD3E9" box="[1409,1429,224,247]" italics="true" pageId="11" pageNumber="134">O</emphasis>
</collectingCountry>
.
<emphasis id="B932D545B561FFBC06FEFE8EFED4D203" box="[151,299,260,285]" italics="true" pageId="11" pageNumber="134">dongnaiensis</emphasis>
</taxonomicName>
specimen (UNS00856) collected at the same locality.
</paragraph>
<paragraph id="8BF90957B561FFBC06AEFEADFA99D296" blockId="11.[151,1437,151,752]" pageId="11" pageNumber="134">
<emphasis id="B932D545B561FFBC06AEFEADFE44D25E" bold="true" box="[199,443,295,320]" pageId="11" pageNumber="134">Morphological data.</emphasis>
Predorsal scales and eye diameter are considered reliable taxonomic characters, and to our knowledge, the short eye diameter (4.25.6 times in HL) in the two new species is extreme among all
<taxonomicName id="4C4672D4B561FFBC06FEFEE5FECED296" box="[151,305,367,392]" class="Actinopterygii" family="Cyprinidae" genus="Onychostoma" kingdom="Animalia" order="Cypriniformes" pageId="11" pageNumber="134" phylum="Chordata" rank="genus">
<emphasis id="B932D545B561FFBC06FEFEE5FECED296" box="[151,305,367,392]" italics="true" pageId="11" pageNumber="134">Onychostoma</emphasis>
</taxonomicName>
species from southeast Asia (
<bibRefCitation id="EFD774A6B561FFBC0417FEE5FD0DD296" author="Taki" box="[638,754,367,392]" pageId="11" pageNumber="134" refString="Taki, Y. (1975) Cyprinid fishes of the genera Onychostoma and Scaphiodonichthys from upper Laos, with remarks on the dispersal of the genera and their allies. Japanese Journal of Ichthyology, 22 (3), 143 - 150." type="journal article" year="1975">Taki 1975</bibRefCitation>
,
<bibRefCitation id="EFD774A6B561FFBC0497FEE5FC65D296" author="Kottelat" box="[766,922,367,392]" pageId="11" pageNumber="134" refString="Kottelat, M. (1998) Fishes of the Nam Theun and Xe Bangfai basins, Laos, with diagnoses of twenty-two new species (Teleostei: Cyprinidae, Balitoridae, Cobitidae, Coiidae and Odontobutidae). Ichthyological Exploration of Freshwaters, 9, 1 - 128." type="journal article" year="1998">Kottelat 1998</bibRefCitation>
,
<bibRefCitation id="EFD774A6B561FFBC05CFFEE5FAA5D296" author="Lothongkham" box="[934,1370,367,392]" pageId="11" pageNumber="134" refString="Lothongkham, A. &amp; Musikasinthorn, P (2005) First record of the cyprinid fish, Onychostoma gerlachi (Cypriniformes: Cyprinidae), from the Nan river basin of the Chao Phraya river system in northern Thailand. The Natural History Bulletin of the Siam Society, 53, 237 - 243." type="journal article" year="2005">Lothongkham &amp; Musikasinthorn 2005</bibRefCitation>
).
</paragraph>
<paragraph id="8BF90957B561FFBC06AEFE1EFED4D19B" blockId="11.[151,1437,151,752]" pageId="11" pageNumber="134">
<taxonomicName id="4C4672D4B561FFBC06AEFE1EFE02D2B3" box="[199,509,404,429]" class="Actinopterygii" family="Cyprinidae" genus="Onychostoma" kingdom="Animalia" order="Cypriniformes" pageId="11" pageNumber="134" phylum="Chordata" rank="species" species="krongnoensis">
<emphasis id="B932D545B561FFBC06AEFE1EFE02D2B3" box="[199,509,404,429]" italics="true" pageId="11" pageNumber="134">Onychostoma krongnoensis</emphasis>
</taxonomicName>
is most similar to
<taxonomicName id="4C4672D4B561FFBC04B8FE1FFC74D2B3" box="[721,907,404,429]" class="Actinopterygii" family="Cyprinidae" genus="Onychostoma" kingdom="Animalia" order="Cypriniformes" pageId="11" pageNumber="134" phylum="Chordata" rank="species" species="dongnaiensis">
<collectingCountry id="F35149C7B561FFBC04B8FE1FFD1AD2B2" box="[721,741,405,428]" name="Iceland" pageId="11" pageNumber="134">
<emphasis id="B932D545B561FFBC04B8FE1FFD1AD2B2" box="[721,741,405,428]" italics="true" pageId="11" pageNumber="134">O</emphasis>
</collectingCountry>
.
<emphasis id="B932D545B561FFBC049DFE1EFC74D2B3" box="[756,907,404,429]" italics="true" pageId="11" pageNumber="134">dongnaiensis</emphasis>
</taxonomicName>
, but differs most dramatically in colour in life.
<taxonomicName id="4C4672D4B561FFBC06FEFE3DFE30D2CE" box="[151,463,439,464]" class="Actinopterygii" family="Cyprinidae" genus="Onychostoma" kingdom="Animalia" order="Cypriniformes" pageId="11" pageNumber="134" phylum="Chordata" rank="species" species="krongnoensis">
<emphasis id="B932D545B561FFBC06FEFE3DFE30D2CE" box="[151,463,439,464]" italics="true" pageId="11" pageNumber="134">Onychostoma krongnoensis</emphasis>
</taxonomicName>
is dark turquoise on the back and yellow to pinkish orange on lateroventral surface (
<figureCitation id="137D15D2B561FFBC06F6FE56FF19D2EA" box="[159,230,476,501]" captionStart="FIGURE 2" captionStartId="4.[151,244,1973,1995]" captionTargetBox="[221,1385,712,1940]" captionTargetId="figure@4.[207,1400,702,1949]" captionTargetPageId="4" captionText="FIGURE 2. Lateral view of Onychostoma krongnoensis specimens (a) female holotype UNS 00805 in life, (b) female holotype UNS 00805 in preservative, (c) male paratype UNS 00825 in preservative. Scale 10 mm." httpUri="https://zenodo.org/record/243916/files/figure.png" pageId="11" pageNumber="134">Fig. 2</figureCitation>
), and
<taxonomicName id="4C4672D4B561FFBC0759FE57FE16D2EB" box="[304,489,476,501]" class="Actinopterygii" family="Cyprinidae" genus="Onychostoma" kingdom="Animalia" order="Cypriniformes" pageId="11" pageNumber="134" phylum="Chordata" rank="species" species="dongnaiensis">
<emphasis id="B932D545B561FFBC0759FE57FE16D2EB" box="[304,489,476,501]" italics="true" pageId="11" pageNumber="134">
<collectingCountry id="F35149C7B561FFBC0759FE57FEB8D2EA" box="[304,327,477,500]" name="Iceland" pageId="11" pageNumber="134">O</collectingCountry>
. dongnaiensis
</emphasis>
</taxonomicName>
is light turquoise on the back, and bluish silver-white on the lateroventral surface (
<figureCitation id="137D15D2B561FFBC06F6FE75FF1CD106" box="[159,227,511,536]" captionStart="FIGURE 5" captionStartId="8.[151,250,1965,1987]" captionTargetBox="[163,1419,473,1929]" captionTargetId="figure@8.[151,1435,447,1940]" captionTargetPageId="8" captionText="FIGURE 5. Lateral view of Onychostoma dongnaiensis specimens (a) female holotype UNS 00851 in life, (b) female holotype UNS 00851 in preservative, (c) male paratype ZRC 54625 in preservative. Scale 10 mm." httpUri="https://zenodo.org/record/243919/files/figure.png" pageId="11" pageNumber="134">Fig. 5</figureCitation>
).
<taxonomicName id="4C4672D4B561FFBC0691FE75FDD3D106" box="[248,556,511,536]" class="Actinopterygii" family="Cyprinidae" genus="Onychostoma" kingdom="Animalia" order="Cypriniformes" pageId="11" pageNumber="134" phylum="Chordata" rank="species" species="krongnoensis">
<emphasis id="B932D545B561FFBC0691FE75FDD3D106" box="[248,556,511,536]" italics="true" pageId="11" pageNumber="134">Onychostoma krongnoensis</emphasis>
</taxonomicName>
also has a slightly deeper body (3.54.3 vs. 3.13.4 times in SL) and a slightly shorter caudal-peduncle (3.84.2 vs. 4.14.5 times in SL). The number of epidermal tubercles on the snout also may vary in the two species with more on
<taxonomicName id="4C4672D4B561FFBC04EDFDC2FCBFD17E" box="[644,832,583,608]" class="Actinopterygii" family="Cyprinidae" genus="Onychostoma" kingdom="Animalia" order="Cypriniformes" pageId="11" pageNumber="134" phylum="Chordata" rank="species" species="krongnoensis">
<collectingCountry id="F35149C7B561FFBC04EDFDC2FD67D141" box="[644,664,584,607]" name="Iceland" pageId="11" pageNumber="134">
<emphasis id="B932D545B561FFBC04EDFDC2FD67D141" box="[644,664,584,607]" italics="true" pageId="11" pageNumber="134">O</emphasis>
</collectingCountry>
.
<emphasis id="B932D545B561FFBC04C0FDCDFCBFD17E" box="[681,832,583,608]" italics="true" pageId="11" pageNumber="134">krongnoensis</emphasis>
</taxonomicName>
, although seasonal and ontogenetic variation has not been studied.
</paragraph>
<paragraph id="8BF90957B561FFBC06AEFD05FC97D1EE" blockId="11.[151,1437,151,752]" pageId="11" pageNumber="134">
<emphasis id="B932D545B561FFBC06AEFD05FE7BD1B6" bold="true" box="[199,388,655,680]" pageId="11" pageNumber="134">Molecular data.</emphasis>
The newly collected specimens from the Krong No and Dong Nai rivers are embedded within a clade containing all Mekong
<taxonomicName id="4C4672D4B561FFBC047AFD3EFD52D1D3" box="[531,685,692,717]" class="Actinopterygii" family="Cyprinidae" genus="Onychostoma" kingdom="Animalia" order="Cypriniformes" pageId="11" pageNumber="134" phylum="Chordata" rank="genus">
<emphasis id="B932D545B561FFBC047AFD3EFD52D1D3" box="[531,685,692,717]" italics="true" pageId="11" pageNumber="134">Onychostoma</emphasis>
</taxonomicName>
species (
<figureCitation id="137D15D2B561FFBC054BFD3EFC92D1D3" box="[802,877,692,717]" captionStart="FIGURE 6" captionStartId="11.[151,250,1584,1606]" captionTargetBox="[155,1429,816,1553]" captionTargetId="figure@11.[151,1436,807,1559]" captionTargetPageId="11" captionText="FIGURE 6. Maximum-likelihood tree based on 16 S ribosomal RNA mitochondrial gene sequences for species of Onychostoma (Wang et al. 2012) and for the outgroup Percocypris pingi. Numbers on branches are ML bootstrap values (values ≥ 45 % shown)." httpUri="https://zenodo.org/record/243920/files/figure.png" pageId="11" pageNumber="134">Fig. 6</figureCitation>
). This clade receives 100% bootstrap support.
<taxonomicName id="4C4672D4B561FFBC06FEFD5DFE40D1EE" box="[151,447,727,752]" class="Actinopterygii" family="Cyprinidae" genus="Onychostoma" kingdom="Animalia" order="Cypriniformes" pageId="11" pageNumber="134" phylum="Chordata" rank="species" species="meridionale">
<emphasis id="B932D545B561FFBC06FEFD5DFE40D1EE" box="[151,447,727,752]" italics="true" pageId="11" pageNumber="134">Onychostoma meridionale</emphasis>
</taxonomicName>
was also embedded within this clade.
</paragraph>
<caption id="DF3959DFB561FFBC06FEF9BAFE7ED59D" httpUri="https://zenodo.org/record/243920/files/figure.png" pageId="11" pageNumber="134" targetBox="[155,1429,816,1553]" targetPageId="11">
<paragraph id="8BF90957B561FFBC06FEF9BAFE7ED59D" blockId="11.[151,1436,1584,1667]" pageId="11" pageNumber="134">
<emphasis id="B932D545B561FFBC06FEF9BAFEE3D558" bold="true" box="[151,284,1584,1606]" pageId="11" pageNumber="134">FIGURE 6.</emphasis>
Maximum-likelihood tree based on 16S ribosomal RNA mitochondrial gene sequences for species of
<taxonomicName id="4C4672D4B561FFBC06FEF9C4FE0AD57A" authority="Wang et al. 2012" authorityName="Wang et al." authorityYear="2012" box="[151,501,1614,1637]" class="Actinopterygii" family="Cyprinidae" genus="Onychostoma" kingdom="Animalia" order="Cypriniformes" pageId="11" pageNumber="134" phylum="Chordata" rank="genus">
<emphasis id="B932D545B561FFBC06FEF9C4FEDCD57A" box="[151,291,1614,1636]" italics="true" pageId="11" pageNumber="134">Onychostoma</emphasis>
(Wang
<emphasis id="B932D545B561FFBC0710F9DAFE56D57A" box="[377,425,1614,1637]" italics="true" pageId="11" pageNumber="134">et al</emphasis>
. 2012)
</taxonomicName>
and for the outgroup
<taxonomicName id="4C4672D4B561FFBC048DF9DAFC64D57B" box="[740,923,1616,1637]" class="Actinopterygii" family="Cyprinidae" genus="Percocypris" kingdom="Animalia" order="Cypriniformes" pageId="11" pageNumber="134" phylum="Chordata" rank="species" species="pingi">
<emphasis id="B932D545B561FFBC048DF9DAFC64D57B" box="[740,923,1616,1637]" italics="true" pageId="11" pageNumber="134">Percocypris pingi</emphasis>
</taxonomicName>
. Numbers on branches are ML bootstrap values (values ≥ 45% shown).
</paragraph>
</caption>
<paragraph id="8BF90957B561FFBC06AEF95EFCB2D4BE" blockId="11.[151,1436,1748,1952]" pageId="11" pageNumber="134">
Our molecular phylogeny suggests that the Krong No and Dong Nai river specimens are most closely related to
<taxonomicName id="4C4672D4B561FFBC06FEF972FEE4D40E" box="[151,283,1783,1808]" class="Actinopterygii" family="Cyprinidae" genus="Barbus" kingdom="Animalia" order="Cypriniformes" pageId="11" pageNumber="134" phylum="Chordata" rank="species" species="gerlachi">
<emphasis id="B932D545B561FFBC06FEF972FEE4D40E" box="[151,283,1783,1808]" italics="true" pageId="11" pageNumber="134">
<collectingCountry id="F35149C7B561FFBC06FEF972FF51D411" box="[151,174,1784,1807]" name="Iceland" pageId="11" pageNumber="134">O</collectingCountry>
. gerlachi
</emphasis>
</taxonomicName>
, differing by 2.0% and 2.1%, respectively, to
<taxonomicName id="4C4672D4B561FFBC052FF972FC32D40E" box="[838,973,1783,1808]" class="Actinopterygii" family="Cyprinidae" genus="Barbus" kingdom="Animalia" order="Cypriniformes" pageId="11" pageNumber="134" phylum="Chordata" rank="species" species="gerlachi">
<collectingCountry id="F35149C7B561FFBC052FF972FCA5D411" box="[838,858,1784,1807]" name="Iceland" pageId="11" pageNumber="134">
<emphasis id="B932D545B561FFBC052FF972FCA5D411" box="[838,858,1784,1807]" italics="true" pageId="11" pageNumber="134">O</emphasis>
</collectingCountry>
.
<emphasis id="B932D545B561FFBC0504F97DFC32D40E" box="[877,973,1783,1808]" italics="true" pageId="11" pageNumber="134">gerlachi</emphasis>
</taxonomicName>
, and&gt;2.0% to all
<taxonomicName id="4C4672D4B561FFBC02DBF97DFAB3D40E" box="[1202,1356,1783,1808]" class="Actinopterygii" family="Cyprinidae" genus="Onychostoma" kingdom="Animalia" order="Cypriniformes" pageId="11" pageNumber="134" phylum="Chordata" rank="genus">
<emphasis id="B932D545B561FFBC02DBF97DFAB3D40E" box="[1202,1356,1783,1808]" italics="true" pageId="11" pageNumber="134">Onychostoma</emphasis>
</taxonomicName>
in the analysis. The Krong No and Dong Nai specimens are closely related to each other and differ by 0.2% sequence divergence, which is larger than the difference between
<taxonomicName id="4C4672D4B561FFBC0559F8CAFBFCD446" box="[816,1027,1856,1880]" class="Actinopterygii" family="Cyprinidae" genus="Onychostoma" kingdom="Animalia" order="Cypriniformes" pageId="11" pageNumber="134" phylum="Chordata" rank="species" species="angustistomata">
<collectingCountry id="F35149C7B561FFBC0559F8CAFCBBD449" box="[816,836,1856,1879]" name="Iceland" pageId="11" pageNumber="134">
<emphasis id="B932D545B561FFBC0559F8CAFCBBD449" box="[816,836,1856,1879]" italics="true" pageId="11" pageNumber="134">O</emphasis>
</collectingCountry>
.
<emphasis id="B932D545B561FFBC0531F8CBFBFCD446" box="[856,1027,1857,1880]" italics="true" pageId="11" pageNumber="134">angustistomata</emphasis>
</taxonomicName>
and
<taxonomicName id="4C4672D4B561FFBC022AF8CAFB4FD446" box="[1091,1200,1856,1880]" class="Actinopterygii" family="Cyprinidae" genus="Onychostoma" kingdom="Animalia" order="Cypriniformes" pageId="11" pageNumber="134" phylum="Chordata" rank="species" species="rarum">
<collectingCountry id="F35149C7B561FFBC022AF8CAFBA8D449" box="[1091,1111,1856,1879]" name="Iceland" pageId="11" pageNumber="134">
<emphasis id="B932D545B561FFBC022AF8CAFBA8D449" box="[1091,1111,1856,1879]" italics="true" pageId="11" pageNumber="134">O</emphasis>
</collectingCountry>
.
<emphasis id="B932D545B561FFBC0203F8CBFB4FD446" box="[1130,1200,1857,1880]" italics="true" pageId="11" pageNumber="134">rarum</emphasis>
</taxonomicName>
by 0.1% (Table 3).
<taxonomicName id="4C4672D4B561FFBC06FEF8EEFE3ED463" box="[151,449,1892,1917]" class="Actinopterygii" family="Cyprinidae" genus="Onychostoma" kingdom="Animalia" order="Cypriniformes" pageId="11" pageNumber="134" phylum="Chordata" rank="species" species="meridionale">
<emphasis id="B932D545B561FFBC06FEF8EEFE3ED463" box="[151,449,1892,1917]" italics="true" pageId="11" pageNumber="134">Onychostoma meridionale</emphasis>
</taxonomicName>
appears most closely related to
<taxonomicName id="4C4672D4B561FFBC055AF8EFFC4BD463" box="[819,948,1892,1917]" class="Actinopterygii" family="Cyprinidae" genus="Barbus" kingdom="Animalia" order="Cypriniformes" pageId="11" pageNumber="134" phylum="Chordata" rank="species" species="gerlachi">
<collectingCountry id="F35149C7B561FFBC055AF8EFFCB8D462" box="[819,839,1893,1916]" name="Iceland" pageId="11" pageNumber="134">
<emphasis id="B932D545B561FFBC055AF8EFFCB8D462" box="[819,839,1893,1916]" italics="true" pageId="11" pageNumber="134">O</emphasis>
</collectingCountry>
.
<emphasis id="B932D545B561FFBC053EF8EEFC4BD463" box="[855,948,1892,1917]" italics="true" pageId="11" pageNumber="134">gerlachi</emphasis>
</taxonomicName>
, but differs by 2.1% from this species and&gt;2.0% from all other
<taxonomicName id="4C4672D4B561FFBC07E0F80DFDDCD4BE" box="[393,547,1927,1952]" class="Actinopterygii" family="Cyprinidae" genus="Onychostoma" kingdom="Animalia" order="Cypriniformes" pageId="11" pageNumber="134" phylum="Chordata" rank="genus">
<emphasis id="B932D545B561FFBC07E0F80DFDDCD4BE" box="[393,547,1927,1952]" italics="true" pageId="11" pageNumber="134">Onychostoma</emphasis>
</taxonomicName>
specimens in the analysis.
</paragraph>
<paragraph id="8BF90957B566FFBB061EFE1BF9F5D2B9" blockId="12.[119,1546,401,423]" box="[119,1546,401,423]" pageId="12" pageNumber="135">
<emphasis id="B932D545B566FFBB061EFE1BFF32D2B9" bold="true" box="[119,205,401,423]" pageId="12" pageNumber="135">Table 3.</emphasis>
Samples used in the phylogenetic analysis of
<taxonomicName id="4C4672D4B566FFBB04F0FE1BFCDDD2B8" box="[665,802,401,422]" class="Actinopterygii" family="Cyprinidae" genus="Onychostoma" kingdom="Animalia" order="Cypriniformes" pageId="12" pageNumber="135" phylum="Chordata" rank="genus">
<emphasis id="B932D545B566FFBB04F0FE1BFCDDD2B8" box="[665,802,401,422]" italics="true" pageId="12" pageNumber="135">Onychostoma</emphasis>
</taxonomicName>
with the outgroup
<taxonomicName id="4C4672D4B566FFBB0589FE1EFB7DD2B8" box="[992,1154,404,422]" class="Actinopterygii" family="Cyprinidae" genus="Percocypris" kingdom="Animalia" order="Cypriniformes" pageId="12" pageNumber="135" phylum="Chordata" rank="species" species="pingi">
<emphasis id="B932D545B566FFBB0589FE1EFB7DD2B8" box="[992,1154,404,422]" italics="true" pageId="12" pageNumber="135">Percocypris pingi</emphasis>
</taxonomicName>
and genetic distances between species.
</paragraph>
<paragraph id="8BF90957B566FFBB0469FE41F9CDD2C0" blockId="12.[132,1600,459,1605]" box="[512,1586,459,479]" pageId="12" pageNumber="135">
1 2 3 4 5 6 7
<date id="FFF82F97B566FFBB0250FE41FB12D2C0" box="[1081,1261,459,478]" pageId="12" pageNumber="135" value="2010-09-08">8 9 10</date>
<date id="FFF82F97B566FFBB034EFE41FA1ED2C0" box="[1319,1505,459,478]" pageId="12" pageNumber="135" value="2013-12-11">11 12 13</date>
14
</paragraph>
<paragraph id="8BF90957B566FFBB06F9FE62FEECD2E5" blockId="12.[132,1600,459,1605]" box="[144,275,488,507]" pageId="12" pageNumber="135">
.
<taxonomicName id="4C4672D4B566FFBB06F2FE62FEECD2E5" box="[155,275,488,507]" class="Actinopterygii" family="Cyprinidae" genus="Onychostoma" kingdom="Animalia" order="Cypriniformes" pageId="12" pageNumber="135" phylum="Chordata" rank="species" species="alticorpus">
<emphasis id="B932D545B566FFBB06F2FE62FEECD2E5" box="[155,275,488,507]" italics="true" pageId="12" pageNumber="135">
<collectingCountry id="F35149C7B566FFBB06F2FE62FF51D2E5" box="[155,174,488,507]" name="Iceland" pageId="12" pageNumber="135">O</collectingCountry>
. alticorpus
</emphasis>
</taxonomicName>
</paragraph>
<paragraph id="8BF90957B566FFBB06EDFD8BFF37D10B" blockId="12.[132,1600,459,1605]" box="[132,200,513,533]" pageId="12" pageNumber="135">
<collectingCountry id="F35149C7B566FFBB06EDFD8BFF37D10B" box="[132,200,513,533]" name="Taiwan" pageId="12" pageNumber="135">Taiwan</collectingCountry>
</paragraph>
<paragraph id="8BF90957B566FFBB06EDFD91FF17D131" blockId="12.[132,1600,459,1605]" box="[132,232,539,559]" pageId="12" pageNumber="135">
<accessionNumber id="941594B4B566FFBB06EDFD91FF17D131" box="[132,232,539,559]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KC791686" pageId="12" pageNumber="135">KC791686</accessionNumber>
</paragraph>
<paragraph id="8BF90957B566FFBB06FBFDBCFEBDD156" blockId="12.[132,1600,459,1605]" box="[146,322,566,585]" pageId="12" pageNumber="135">
.
<taxonomicName id="4C4672D4B566FFBB06F2FDBCFEBDD156" box="[155,322,566,585]" class="Actinopterygii" family="Cyprinidae" genus="Onychostoma" kingdom="Animalia" order="Cypriniformes" pageId="12" pageNumber="135" phylum="Chordata" rank="species" species="angustistomata">
<emphasis id="B932D545B566FFBB06F2FDBCFEBDD156" box="[155,322,566,585]" italics="true" pageId="12" pageNumber="135">
<collectingCountry id="F35149C7B566FFBB06F2FDBCFF51D157" box="[155,174,566,585]" name="Iceland" pageId="12" pageNumber="135">O</collectingCountry>
. angustistomata
</emphasis>
</taxonomicName>
</paragraph>
<paragraph id="8BF90957B566FFBB06EDFDDAFF16D160" blockId="12.[132,1600,459,1605]" pageId="12" pageNumber="135">
<collectingCountry id="F35149C7B566FFBB06EDFDDAFF45D17A" box="[132,186,592,612]" name="China" pageId="12" pageNumber="135">China</collectingCountry>
0.045
<accessionNumber id="941594B4B566FFBB06EDFDE0FF16D160" box="[132,233,618,638]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/HQ235714" pageId="12" pageNumber="135">HQ235714</accessionNumber>
</paragraph>
<paragraph id="8BF90957B566FFBB06F8FD0EFEDED189" blockId="12.[132,1600,459,1605]" box="[145,289,644,663]" pageId="12" pageNumber="135">
.
<taxonomicName id="4C4672D4B566FFBB06F2FD0EFEDED189" box="[155,289,644,663]" class="Actinopterygii" family="Cyprinidae" genus="Onychostoma" kingdom="Animalia" order="Cypriniformes" pageId="12" pageNumber="135" phylum="Chordata" rank="species" species="barbatulum">
<emphasis id="B932D545B566FFBB06F2FD0EFEDED189" box="[155,289,644,663]" italics="true" pageId="12" pageNumber="135">
<collectingCountry id="F35149C7B566FFBB06F2FD0EFF51D189" box="[155,174,644,663]" name="Iceland" pageId="12" pageNumber="135">O</collectingCountry>
. barbatulum
</emphasis>
</taxonomicName>
</paragraph>
<paragraph id="8BF90957B566FFBB06EDFD14FF17D1D2" blockId="12.[132,1600,459,1605]" pageId="12" pageNumber="135">
<collectingCountry id="F35149C7B566FFBB06EDFD14FF37D1AC" box="[132,200,670,690]" name="Taiwan" pageId="12" pageNumber="135">Taiwan</collectingCountry>
0.063 0.052
<accessionNumber id="941594B4B566FFBB06EDFD32FF17D1D2" box="[132,232,696,716]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KC896762" pageId="12" pageNumber="135">KC896762</accessionNumber>
</paragraph>
<paragraph id="8BF90957B566FFBB06FBFD59FF1DD01E" blockId="12.[132,1600,459,1605]" pageId="12" pageNumber="135">
.
<taxonomicName id="4C4672D4B566FFBB06F2FD59FED1D1FB" box="[155,302,722,742]" class="Actinopterygii" family="Cyprinidae" genus="Onychostoma" kingdom="Animalia" order="Cypriniformes" pageId="12" pageNumber="135" phylum="Chordata" rank="species" species="dongnaiensis" status="sp. nov.">
<emphasis id="B932D545B566FFBB06F2FD59FED1D1FB" box="[155,302,722,742]" italics="true" pageId="12" pageNumber="135">
<collectingCountry id="F35149C7B566FFBB06F2FD59FF51D1F8" box="[155,174,723,742]" name="Iceland" pageId="12" pageNumber="135">O</collectingCountry>
. dongnaiensis
</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A201683EB566FFBB075DFD59FE83D1F8" box="[308,380,723,742]" pageId="12" pageNumber="135" rank="species">sp. nov.</taxonomicNameLabel>
0.063 0.066 0.070
<accessionNumber id="941594B4B566FFBB06EDFD66FF1DD01E" box="[132,226,748,768]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KJ754099" pageId="12" pageNumber="135">KJ754099</accessionNumber>
</paragraph>
<paragraph id="8BF90957B566FFBB06FBFC8CFEE8D007" blockId="12.[132,1600,459,1605]" box="[146,279,774,794]" pageId="12" pageNumber="135">
.
<taxonomicName id="4C4672D4B566FFBB06F2FC8DFEE8D007" box="[155,279,774,794]" class="Actinopterygii" family="Cyprinidae" genus="Onychostoma" kingdom="Animalia" order="Cypriniformes" pageId="12" pageNumber="135" phylum="Chordata" rank="species" species="elongatum">
<emphasis id="B932D545B566FFBB06F2FC8DFEE8D007" box="[155,279,774,794]" italics="true" pageId="12" pageNumber="135">
<collectingCountry id="F35149C7B566FFBB06F2FC8DFF51D004" box="[155,174,775,794]" name="Iceland" pageId="12" pageNumber="135">O</collectingCountry>
. elongatum
</emphasis>
</taxonomicName>
</paragraph>
<paragraph id="8BF90957B566FFBB06EDFCAAFF00D050" blockId="12.[132,1600,459,1605]" pageId="12" pageNumber="135">
<collectingCountry id="F35149C7B566FFBB06EDFCAAFF42D02A" box="[132,189,800,820]" name="China" pageId="12" pageNumber="135">China</collectingCountry>
: Rongan, Guangxi Zhuang 0.072 0.068 0.065 0.084 Auto, Region
</paragraph>
<paragraph id="8BF90957B566FFBB06EDFCDEFF16D076" blockId="12.[132,1600,459,1605]" box="[132,233,852,872]" pageId="12" pageNumber="135">
<accessionNumber id="941594B4B566FFBB06EDFCDEFF16D076" box="[132,233,852,872]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/GQ406254" pageId="12" pageNumber="135">GQ406254</accessionNumber>
</paragraph>
<paragraph id="8BF90957B566FFBB06FBFCE4FEFDD09C" blockId="12.[132,1600,459,1605]" box="[146,258,878,898]" pageId="12" pageNumber="135">
.
<taxonomicName id="4C4672D4B566FFBB06F2FCE5FEFDD09C" box="[155,258,879,898]" class="Actinopterygii" family="Cyprinidae" genus="Barbus" kingdom="Animalia" order="Cypriniformes" pageId="12" pageNumber="135" phylum="Chordata" rank="species" species="gerlachi">
<emphasis id="B932D545B566FFBB06F2FCE5FEFDD09C" box="[155,258,879,898]" italics="true" pageId="12" pageNumber="135">
<collectingCountry id="F35149C7B566FFBB06F2FCE5FF51D09C" box="[155,174,879,898]" name="Iceland" pageId="12" pageNumber="135">O</collectingCountry>
. gerlachi
</emphasis>
</taxonomicName>
</paragraph>
<paragraph id="8BF90957B566FFBB06EDFC02FF16D0A9" blockId="12.[132,1600,459,1605]" pageId="12" pageNumber="135">
<collectingCountry id="F35149C7B566FFBB06EDFC02FF42D082" box="[132,189,904,924]" name="China" pageId="12" pageNumber="135">China</collectingCountry>
: Jinghong, Yunnan Prov. 0.067 0.064 0.076 0.021 0.089
<accessionNumber id="941594B4B566FFBB06EDFC29FF16D0A9" box="[132,233,931,951]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/DQ845862" pageId="12" pageNumber="135">DQ845862</accessionNumber>
</paragraph>
<paragraph id="8BF90957B566FFBB06FBFC37FF1ED0F4" blockId="12.[132,1600,459,1605]" pageId="12" pageNumber="135">
.
<taxonomicName id="4C4672D4B566FFBB06F2FC37FECFD0CE" box="[155,304,957,976]" class="Actinopterygii" family="Cyprinidae" genus="Onychostoma" kingdom="Animalia" order="Cypriniformes" pageId="12" pageNumber="135" phylum="Chordata" rank="species" species="krongnoensis" status="sp. nov.">
<emphasis id="B932D545B566FFBB06F2FC37FECFD0CE" box="[155,304,957,976]" italics="true" pageId="12" pageNumber="135">
<collectingCountry id="F35149C7B566FFBB06F2FC37FF51D0CE" box="[155,174,957,976]" name="Iceland" pageId="12" pageNumber="135">O</collectingCountry>
. krongnoensis
</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A201683EB566FFBB075CFC37FE82D0CE" box="[309,381,957,976]" pageId="12" pageNumber="135" rank="species">sp. nov.</taxonomicNameLabel>
0.062 0.064 0.067 0.002 0.083 0.020
<accessionNumber id="941594B4B566FFBB06EDFC5CFF1ED0F4" box="[132,225,982,1002]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KJ754097" pageId="12" pageNumber="135">KJ754097</accessionNumber>
</paragraph>
<paragraph id="8BF90957B566FFBB06F8FC7BFF07D71A" blockId="12.[132,1600,459,1605]" box="[145,248,1009,1028]" pageId="12" pageNumber="135">
.
<taxonomicName id="4C4672D4B566FFBB06F2FC7BFF07D71A" box="[155,248,1009,1028]" class="Actinopterygii" family="Cyprinidae" genus="Onychostoma" kingdom="Animalia" order="Cypriniformes" pageId="12" pageNumber="135" phylum="Chordata" rank="species" species="leptura">
<emphasis id="B932D545B566FFBB06F2FC7BFF07D71A" box="[155,248,1009,1028]" italics="true" pageId="12" pageNumber="135">
<collectingCountry id="F35149C7B566FFBB06F2FC7BFF51D71A" box="[155,174,1009,1028]" name="Iceland" pageId="12" pageNumber="135">O</collectingCountry>
. leptura
</emphasis>
</taxonomicName>
</paragraph>
<paragraph id="8BF90957B566FFBB06EDFB81FF00D727" blockId="12.[132,1600,459,1605]" pageId="12" pageNumber="135">
<collectingCountry id="F35149C7B566FFBB06EDFB81FF42D701" box="[132,189,1035,1055]" name="China" pageId="12" pageNumber="135">China</collectingCountry>
: Xilin, Guangxi Zhuang 0.060 0.058 0.063 0.030 0.080 0.033 0.028 Auto, Region
</paragraph>
<paragraph id="8BF90957B566FFBB06EDFBB5FF16D74D" blockId="12.[132,1600,459,1605]" box="[132,233,1087,1107]" pageId="12" pageNumber="135">
<accessionNumber id="941594B4B566FFBB06EDFBB5FF16D74D" box="[132,233,1087,1107]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/GQ406257" pageId="12" pageNumber="135">GQ406257</accessionNumber>
</paragraph>
<paragraph id="8BF90957B566FFBB06FBFBD0FF2AD772" blockId="12.[132,1600,459,1605]" box="[146,213,1113,1133]" pageId="12" pageNumber="135">
.
<taxonomicName id="4C4672D4B566FFBB06F2FBD0FF2AD772" box="[155,213,1113,1133]" class="Actinopterygii" family="Cyprinidae" genus="Onychostoma" kingdom="Animalia" order="Cypriniformes" pageId="12" pageNumber="135" phylum="Chordata" rank="species" species="lini">
<emphasis id="B932D545B566FFBB06F2FBD0FF2AD772" box="[155,213,1113,1133]" italics="true" pageId="12" pageNumber="135">
<collectingCountry id="F35149C7B566FFBB06F2FBD0FF51D773" box="[155,174,1114,1133]" name="Iceland" pageId="12" pageNumber="135">O</collectingCountry>
. lini
</emphasis>
</taxonomicName>
</paragraph>
<paragraph id="8BF90957B566FFBB06EDFBF9FF1ED7BF" blockId="12.[132,1600,459,1605]" pageId="12" pageNumber="135">
<collectingCountry id="F35149C7B566FFBB06EDFBF9FF42D799" box="[132,189,1139,1159]" name="China" pageId="12" pageNumber="135">China</collectingCountry>
: Youyang, Chongqing 0.048 0.044 0.041 0.052 0.065 0.059 0.050 0.046
<accessionNumber id="941594B4B566FFBB06EDFB04FF1ED7BF" box="[132,225,1166,1185]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/JQ343982" pageId="12" pageNumber="135">JQ343982</accessionNumber>
</paragraph>
<paragraph id="8BF90957B566FFBB06EDFB22FED9D7A4" blockId="12.[132,1600,459,1605]" box="[132,294,1191,1211]" pageId="12" pageNumber="135">
10.
<taxonomicName id="4C4672D4B566FFBB06CFFB22FED9D7A4" box="[166,294,1191,1211]" class="Actinopterygii" family="Cyprinidae" genus="Onychostoma" kingdom="Animalia" order="Cypriniformes" pageId="12" pageNumber="135" phylum="Chordata" rank="species" species="macrolepis">
<emphasis id="B932D545B566FFBB06CFFB22FED9D7A4" box="[166,294,1191,1211]" italics="true" pageId="12" pageNumber="135">
<collectingCountry id="F35149C7B566FFBB06CFFB22FF46D7A5" box="[166,185,1192,1211]" name="Iceland" pageId="12" pageNumber="135">O</collectingCountry>
. macrolepis
</emphasis>
</taxonomicName>
</paragraph>
<paragraph id="8BF90957B566FFBB06EDFB4BFF15D7F1" blockId="12.[132,1600,459,1605]" pageId="12" pageNumber="135">
<collectingCountry id="F35149C7B566FFBB06EDFB4BFF42D7CB" box="[132,189,1217,1237]" name="China" pageId="12" pageNumber="135">China</collectingCountry>
: Taian, Shandong Prov. 0.054 0.050 0.051 0.057 0.073 0.060 0.055 0.053 0.019
<accessionNumber id="941594B4B566FFBB06EDFB51FF15D7F1" box="[132,234,1243,1263]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/GQ406258" pageId="12" pageNumber="135">GQ406258</accessionNumber>
</paragraph>
<paragraph id="8BF90957B566FFBB06EDFB7DFF0CD63B" blockId="12.[132,1600,459,1605]" pageId="12" pageNumber="135">
11.
<taxonomicName id="4C4672D4B566FFBB06CFFB7DFED0D614" box="[166,303,1271,1290]" class="Actinopterygii" family="Cyprinidae" genus="Onychostoma" kingdom="Animalia" order="Cypriniformes" pageId="12" pageNumber="135" phylum="Chordata" rank="species" species="meridionale">
<emphasis id="B932D545B566FFBB06CFFB7DFED0D614" box="[166,303,1271,1290]" italics="true" pageId="12" pageNumber="135">
<collectingCountry id="F35149C7B566FFBB06CFFB7DFF46D614" box="[166,185,1271,1290]" name="Iceland" pageId="12" pageNumber="135">O</collectingCountry>
. meridionale
</emphasis>
</taxonomicName>
0.074 0.067 0.082 0.018 0.092 0.021 0.020 0.037 0.064 0.065 12.
<taxonomicName id="4C4672D4B566FFBB06CFFA99FF0CD63B" box="[166,243,1298,1318]" class="Actinopterygii" family="Cyprinidae" genus="Onychostoma" kingdom="Animalia" order="Cypriniformes" pageId="12" pageNumber="135" phylum="Chordata" rank="species" species="ovale">
<emphasis id="B932D545B566FFBB06CFFA99FF0CD63B" box="[166,243,1298,1318]" italics="true" pageId="12" pageNumber="135">
<collectingCountry id="F35149C7B566FFBB06CFFA99FF46D638" box="[166,185,1299,1318]" name="Iceland" pageId="12" pageNumber="135">O</collectingCountry>
. ovale
</emphasis>
</taxonomicName>
</paragraph>
<paragraph id="8BF90957B566FFBB06EDFAA6FF1ED644" blockId="12.[132,1600,459,1605]" pageId="12" pageNumber="135">
<collectingCountry id="F35149C7B566FFBB06EDFAA6FF42D65E" box="[132,189,1324,1344]" name="China" pageId="12" pageNumber="135">China</collectingCountry>
: Tiane, Guangxi Prov. 0.035 0.014 0.029 0.047 0.050 0.049 0.047 0.039 0.029 0.051 0.056
<accessionNumber id="941594B4B566FFBB06EDFACDFF1ED644" box="[132,225,1351,1370]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/JX074089" pageId="12" pageNumber="135">JX074089</accessionNumber>
</paragraph>
<paragraph id="8BF90957B566FFBB06EDFAEBFF05D66D" blockId="12.[132,1600,459,1605]" box="[132,250,1377,1396]" pageId="12" pageNumber="135">
13.
<taxonomicName id="4C4672D4B566FFBB06CFFAEBFF05D66D" box="[166,250,1377,1396]" class="Actinopterygii" family="Cyprinidae" genus="Onychostoma" kingdom="Animalia" order="Cypriniformes" pageId="12" pageNumber="135" phylum="Chordata" rank="species" species="rarum">
<emphasis id="B932D545B566FFBB06CFFAEBFF05D66D" box="[166,250,1377,1396]" italics="true" pageId="12" pageNumber="135">
<collectingCountry id="F35149C7B566FFBB06CFFAEBFF46D66A" box="[166,185,1377,1396]" name="Iceland" pageId="12" pageNumber="135">O</collectingCountry>
. rarum
</emphasis>
</taxonomicName>
</paragraph>
<paragraph id="8BF90957B566FFBB06EDFAF0FF16D6B7" blockId="12.[132,1600,459,1605]" pageId="12" pageNumber="135">
<collectingCountry id="F35149C7B566FFBB06EDFAF0FF45D690" box="[132,186,1402,1422]" name="China" pageId="12" pageNumber="135">China</collectingCountry>
0.044 0.001 0.051 0.064 0.068 0.063 0.063 0.056 0.043 0.050 0.066 0.012
<accessionNumber id="941594B4B566FFBB06EDFA1FFF16D6B7" box="[132,233,1429,1449]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/HQ235715" pageId="12" pageNumber="135">HQ235715</accessionNumber>
</paragraph>
<paragraph id="8BF90957B566FFBB06EDFA25FF02D6DF" blockId="12.[132,1600,459,1605]" box="[132,253,1455,1474]" pageId="12" pageNumber="135">
14.
<taxonomicName id="4C4672D4B566FFBB06CFFA25FF02D6DF" box="[166,253,1455,1474]" class="Actinopterygii" family="Cyprinidae" genus="Onychostoma" kingdom="Animalia" order="Cypriniformes" pageId="12" pageNumber="135" phylum="Chordata" rank="species" species="simum">
<emphasis id="B932D545B566FFBB06CFFA25FF02D6DF" box="[166,253,1455,1474]" italics="true" pageId="12" pageNumber="135">
<collectingCountry id="F35149C7B566FFBB06CFFA25FF46D6DC" box="[166,185,1455,1474]" name="Iceland" pageId="12" pageNumber="135">O</collectingCountry>
. simum
</emphasis>
</taxonomicName>
</paragraph>
<paragraph id="8BF90957B566FFBB06EDFA43FF15D6E9" blockId="12.[132,1600,459,1605]" pageId="12" pageNumber="135">
<collectingCountry id="F35149C7B566FFBB06EDFA43FF42D6C3" box="[132,189,1481,1501]" name="China" pageId="12" pageNumber="135">China</collectingCountry>
: Hejiang, Sichuan Prov. 0.024 0.044 0.050 0.056 0.066 0.062 0.055 0.054 0.041 0.047 0.066 0.033 0.042
<accessionNumber id="941594B4B566FFBB06EDFA69FF15D6E9" box="[132,234,1507,1527]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/DQ845861" pageId="12" pageNumber="135">DQ845861</accessionNumber>
</paragraph>
<paragraph id="8BF90957B566FFBB06EDFA77FEB7D50E" blockId="12.[132,1600,459,1605]" box="[132,328,1533,1553]" pageId="12" pageNumber="135">
15.
<taxonomicName id="4C4672D4B566FFBB06CFFA74FEB7D50E" box="[166,328,1534,1552]" class="Actinopterygii" family="Cyprinidae" genus="Percocypris" kingdom="Animalia" order="Cypriniformes" pageId="12" pageNumber="135" phylum="Chordata" rank="species" species="pingi">
<emphasis id="B932D545B566FFBB06CFFA74FEB7D50E" box="[166,328,1534,1552]" italics="true" pageId="12" pageNumber="135">Percocypris pingi</emphasis>
</taxonomicName>
</paragraph>
<paragraph id="8BF90957B566FFBB06EDF99DFF17D55B" blockId="12.[132,1600,459,1605]" pageId="12" pageNumber="135">
<collectingCountry id="F35149C7B566FFBB06EDF99DFF42D535" box="[132,189,1559,1579]" name="China" pageId="12" pageNumber="135">China</collectingCountry>
: Hejiang, Sichuan Prov. 0.073 0.067 0.074 0.089 0.085 0.089 0.087 0.076 0.070 0.072 0.095 0.052 0.066 0.070
<accessionNumber id="941594B4B566FFBB06EDF9BBFF17D55B" box="[132,232,1585,1605]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/GQ406259" pageId="12" pageNumber="135">GQ406259</accessionNumber>
</paragraph>
</subSubSection>
<subSubSection id="C35C5ADCB566FFBA06FEF8B0FCC0D41B" lastPageId="13" lastPageNumber="136" pageId="12" pageNumber="135" type="discussion">
<paragraph id="8BF90957B566FFBB06FEF8B0FEE4D44A" blockId="12.[151,283,1850,1876]" box="[151,283,1850,1876]" pageId="12" pageNumber="135">
<emphasis id="B932D545B566FFBB06FEF8B0FEE4D44A" bold="true" box="[151,283,1850,1876]" pageId="12" pageNumber="135">Discussion</emphasis>
</paragraph>
<paragraph id="8BF90957B566FFBA06FEF80AFB57D3E9" blockId="12.[151,1436,1920,2017]" lastBlockId="13.[151,1436,151,1005]" lastPageId="13" lastPageNumber="136" pageId="12" pageNumber="135">
With two new species described here, a total of 22 species of
<taxonomicName id="4C4672D4B566FFBB053BF80AFC13D487" box="[850,1004,1920,1945]" class="Actinopterygii" family="Cyprinidae" genus="Onychostoma" kingdom="Animalia" order="Cypriniformes" pageId="12" pageNumber="135" phylum="Chordata" rank="genus">
<emphasis id="B932D545B566FFBB053BF80AFC13D487" box="[850,1004,1920,1945]" italics="true" pageId="12" pageNumber="135">Onychostoma</emphasis>
</taxonomicName>
are now recognised, with five species occurring in the Mekong drainage.
<taxonomicName id="4C4672D4B566FFBB045DF82EFC93D4A3" box="[564,876,1956,1981]" class="Actinopterygii" family="Cyprinidae" genus="Onychostoma" kingdom="Animalia" order="Cypriniformes" pageId="12" pageNumber="135" phylum="Chordata" rank="species" species="krongnoensis">
<emphasis id="B932D545B566FFBB045DF82EFC93D4A3" box="[564,876,1956,1981]" italics="true" pageId="12" pageNumber="135">Onychostoma krongnoensis</emphasis>
</taxonomicName>
inhabits the Ea Krong No river drainage of the Langbiang Plateau. These mountain rivers are perennial, shallow water bodies characterized by low temperature, high turbulent current, and rocky substratum.
<collectingCountry id="F35149C7B567FFBA04C0FF12FD37D3AE" box="[681,712,152,176]" name="American Samoa" pageId="13" pageNumber="136">As</collectingCountry>
adaptions to strong water currents,
<taxonomicName id="4C4672D4B567FFBA0203FF12FADBD3AE" box="[1130,1316,151,176]" class="Actinopterygii" family="Cyprinidae" genus="Onychostoma" kingdom="Animalia" order="Cypriniformes" pageId="13" pageNumber="136" phylum="Chordata" rank="species" species="krongnoensis">
<emphasis id="B932D545B567FFBA0203FF12FADBD3AE" box="[1130,1316,151,176]" italics="true" pageId="13" pageNumber="136">
<collectingCountry id="F35149C7B567FFBA0203FF12FB7ED3B1" box="[1130,1153,152,175]" name="Iceland" pageId="13" pageNumber="136">O</collectingCountry>
. krongnoensis
</emphasis>
</taxonomicName>
has dense epidermal tubercles on the snout tip, a slender, streamlined body with an increased number of predorsal scales (mostly 17 scales), and a longer caudal peduncle compared to other species of
<taxonomicName id="4C4672D4B567FFBA026EFF55FB5ED3E6" box="[1031,1185,223,248]" class="Actinopterygii" family="Cyprinidae" genus="Onychostoma" kingdom="Animalia" order="Cypriniformes" pageId="13" pageNumber="136" phylum="Chordata" rank="genus">
<emphasis id="B932D545B567FFBA026EFF55FB5ED3E6" box="[1031,1185,223,248]" italics="true" pageId="13" pageNumber="136">Onychostoma</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BF90957B567FFBA06AEFE8EFB56D122" blockId="13.[151,1436,151,1005]" pageId="13" pageNumber="136">
Tubercles of cyprinids exhibit sexual dimorphism and ontogenetic variation in the size, shape and distribution (
<bibRefCitation id="EFD774A6B567FFBA06F7FEADFE4DD25E" author="Wiley" box="[158,434,295,320]" pageId="13" pageNumber="136" refString="Wiley, M. L. &amp; Collette, B. B. (1970) Breeding tubercles and contact organa in fishes: their occurrence structure, and significance. Bulletin of the American Museum of Natural History, 143, 143 - 216." type="journal article" year="1970">Wiley &amp; Collette, 1970</bibRefCitation>
). In contrast to other species of
<taxonomicName id="4C4672D4B567FFBA0541FEADFC3DD25E" box="[808,962,295,320]" class="Actinopterygii" family="Cyprinidae" genus="Onychostoma" kingdom="Animalia" order="Cypriniformes" pageId="13" pageNumber="136" phylum="Chordata" rank="genus">
<emphasis id="B932D545B567FFBA0541FEADFC3DD25E" box="[808,962,295,320]" italics="true" pageId="13" pageNumber="136">Onychostoma</emphasis>
</taxonomicName>
, in
<taxonomicName id="4C4672D4B567FFBA059BFEA2FB53D25E" box="[1010,1196,295,320]" class="Actinopterygii" family="Cyprinidae" genus="Onychostoma" kingdom="Animalia" order="Cypriniformes" pageId="13" pageNumber="136" phylum="Chordata" rank="species" species="krongnoensis">
<collectingCountry id="F35149C7B567FFBA059BFEA2FBF9D221" box="[1010,1030,296,319]" name="Iceland" pageId="13" pageNumber="136">
<emphasis id="B932D545B567FFBA059BFEA2FBF9D221" box="[1010,1030,296,319]" italics="true" pageId="13" pageNumber="136">O</emphasis>
</collectingCountry>
.
<emphasis id="B932D545B567FFBA027EFEADFB53D25E" box="[1047,1196,295,320]" italics="true" pageId="13" pageNumber="136">krongnoensis</emphasis>
</taxonomicName>
the epidermis of the snout possesses 23 irregular transverse rows of 1343 tubercles in juveniles and adults of both sexes (
<figureCitation id="137D15D2B567FFBA0350FEC6FA81D27B" box="[1337,1406,332,357]" captionStart="FIGURE 3" captionStartId="5.[151,250,1842,1864]" captionTargetBox="[172,1412,529,1810]" captionTargetId="figure@5.[163,1423,519,1819]" captionTargetPageId="5" captionText="FIGURE 3. Front view of head of Onychostoma krongnoensis (a) female holotype UNS 00805, 205 mm SL, (b) immature paratype UNS 00821, 123 mm SL, (c) male paratype UNS 00825, 214 mm SL; O. dongnaiensis (e) female holotype UNS 00851, 172 mm SL, (f) female paratype UNS 00852, 196 mm SL, (g) male paratype ZRC 54625, 157 mm SL; O. leptura (i) female UNS 00684, 186 mm SL. Ventral view of head of O. krongnoensis, wide mouth (d) male paratype UNS 00825; O. dongnaiensis, wide mouth (h) female holotype UNS 00851; O. leptura, wide mouth (j) female UNS 00684; O. meridionale, moderate mouth (k) immature UNS 00826, 108 mm SL; O. uniforme, narrow mouth (l) female UNS 00689, 130 mm SL. Scale 10 mm." httpUri="https://zenodo.org/record/243917/files/figure.png" pageId="13" pageNumber="136">Fig. 3</figureCitation>
b). Variation in the surface of the snout in hill-stream fishes has been suggested to be an adaptation to life in torrential streams (
<bibRefCitation id="EFD774A6B567FFBA0694FE1EFE12D2B2" author="Hoshiyar" box="[253,493,404,429]" pageId="13" pageNumber="136" refString="Hoshiyar, S., Joshi S. C., Bisht, I. &amp; Agarwal, S. K. (2013) The adaptive modifications of snout epidermis of a hill-stream fish Schizothorax richardsonii of Kumaun Himalaya: A SEM Investigation. Research Journal of Animal, Veterinary and Fishery Sciences, 1, 1 - 4." type="journal article" year="2013">
Hoshiyar
<emphasis id="B932D545B567FFBA0707FE1FFE5FD2B3" box="[366,416,404,429]" italics="true" pageId="13" pageNumber="136">et al</emphasis>
. 2013
</bibRefCitation>
). The snout epidermis of
<taxonomicName id="4C4672D4B567FFBA057BFE1FFC35D2B3" box="[786,970,404,429]" class="Actinopterygii" family="Cyprinidae" genus="Onychostoma" kingdom="Animalia" order="Cypriniformes" pageId="13" pageNumber="136" phylum="Chordata" rank="species" species="krongnoensis">
<collectingCountry id="F35149C7B567FFBA057BFE1FFCD9D2B2" box="[786,806,405,428]" name="Iceland" pageId="13" pageNumber="136">
<emphasis id="B932D545B567FFBA057BFE1FFCD9D2B2" box="[786,806,405,428]" italics="true" pageId="13" pageNumber="136">O</emphasis>
</collectingCountry>
.
<emphasis id="B932D545B567FFBA055FFE1EFC35D2B3" box="[822,970,404,429]" italics="true" pageId="13" pageNumber="136">krongnoensis</emphasis>
</taxonomicName>
is subject to frictional stress as it comes into contact with water current, thus supporting this hypothesis. In contrast, species that inhabit low elevation streams with slower velocity, such as
<taxonomicName id="4C4672D4B567FFBA042DFE57FD04D2EB" box="[580,763,476,501]" class="Actinopterygii" family="Cyprinidae" genus="Onychostoma" kingdom="Animalia" order="Cypriniformes" pageId="13" pageNumber="136" phylum="Chordata" rank="species" species="dongnaiensis">
<collectingCountry id="F35149C7B567FFBA042DFE57FDA7D2EA" box="[580,600,477,500]" name="Iceland" pageId="13" pageNumber="136">
<emphasis id="B932D545B567FFBA042DFE57FDA7D2EA" box="[580,600,477,500]" italics="true" pageId="13" pageNumber="136">O</emphasis>
</collectingCountry>
.
<emphasis id="B932D545B567FFBA040EFE56FD04D2EB" box="[615,763,476,501]" italics="true" pageId="13" pageNumber="136">dongnaiensis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C4672D4B567FFBA0562FE57FC80D2EB" box="[779,895,476,501]" class="Actinopterygii" family="Cyprinidae" genus="Onychostoma" kingdom="Animalia" order="Cypriniformes" pageId="13" pageNumber="136" phylum="Chordata" rank="species" species="leptura">
<collectingCountry id="F35149C7B567FFBA0562FE57FCE0D2EA" box="[779,799,477,500]" name="Iceland" pageId="13" pageNumber="136">
<emphasis id="B932D545B567FFBA0562FE57FCE0D2EA" box="[779,799,477,500]" italics="true" pageId="13" pageNumber="136">O</emphasis>
</collectingCountry>
.
<emphasis id="B932D545B567FFBA0547FE56FC80D2EB" box="[814,895,476,501]" italics="true" pageId="13" pageNumber="136">leptura</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C4672D4B567FFBA05D6FE57FBB4D2EA" box="[959,1099,476,500]" class="Actinopterygii" family="Cyprinidae" genus="Onychostoma" kingdom="Animalia" order="Cypriniformes" pageId="13" pageNumber="136" phylum="Chordata" rank="species" species="uniforme">
<collectingCountry id="F35149C7B567FFBA05D6FE57FC2CD2EA" box="[959,979,477,500]" name="Iceland" pageId="13" pageNumber="136">
<emphasis id="B932D545B567FFBA05D6FE57FC2CD2EA" box="[959,979,477,500]" italics="true" pageId="13" pageNumber="136">O</emphasis>
</collectingCountry>
.
<emphasis id="B932D545B567FFBA058AFE56FBB4D2EA" box="[995,1099,476,500]" italics="true" pageId="13" pageNumber="136">uniforme</emphasis>
</taxonomicName>
, have fewer smaller tubercles on the snout (Hoang
<emphasis id="B932D545B567FFBA07C8FD8BFE25D106" box="[417,474,511,536]" italics="true" pageId="13" pageNumber="136">et al</emphasis>
. pers.obs.). Breeding tubercles also occur on the anal-fin rays of males of
<taxonomicName id="4C4672D4B567FFBA03E8FD8AFED3D123" class="Actinopterygii" family="Cyprinidae" genus="Onychostoma" kingdom="Animalia" order="Cypriniformes" pageId="13" pageNumber="136" phylum="Chordata" rank="species" species="krongnoensis">
<collectingCountry id="F35149C7B567FFBA03E8FD8AFA6AD109" box="[1409,1429,512,535]" name="Iceland" pageId="13" pageNumber="136">
<emphasis id="B932D545B567FFBA03E8FD8AFA6AD109" box="[1409,1429,512,535]" italics="true" pageId="13" pageNumber="136">O</emphasis>
</collectingCountry>
.
<emphasis id="B932D545B567FFBA06FEFDAEFED3D123" box="[151,300,548,573]" italics="true" pageId="13" pageNumber="136">krongnoensis</emphasis>
</taxonomicName>
and other stream cyprinids (
<bibRefCitation id="EFD774A6B567FFBA0403FDAEFC3DD122" author="Witkowski" box="[618,962,548,573]" pageId="13" pageNumber="136" refString="Witkowski, A. &amp; Rogowska, M. W. (1991) Breeding tubercles in some European cyprinid fishes (Osteichthyes, Cypriniformes: Cyprinidae). Zoologische Abhandlungen, 9, 129 - 155." type="journal article" year="1991">Witkowski &amp; Rogowska 1991</bibRefCitation>
;
<bibRefCitation id="EFD774A6B567FFBA05A7FDAEFB62D122" author="Poncin" box="[974,1181,548,573]" pageId="13" pageNumber="136" refString="Poncin, P., Nzau Matondo, B., Termol, C., Kestemont, P. &amp; Philippart, J. C. (2011) Relationships between circulating androgens, aggressive behaviour and breeding tubercles in males of the common bream Abramis brama L. in an aquarium environment. Fish Physiology and Biochemistry, 37, 533 - 542. http: // dx. doi. org / 10.1007 / s 10695 - 010 - 9455 - y" type="journal article" year="2011">
Poncin
<emphasis id="B932D545B567FFBA024BFDAFFBADD123" box="[1058,1106,548,573]" italics="true" pageId="13" pageNumber="136">et al</emphasis>
. 2011
</bibRefCitation>
).
</paragraph>
<paragraph id="8BF90957B567FFBA06AEFDCDFC74D043" blockId="13.[151,1436,151,1005]" pageId="13" pageNumber="136">
Body shape affects movement in stream fishes (
<bibRefCitation id="EFD774A6B567FFBA0485FDCDFC34D17E" author="Chuang" box="[748,971,583,608]" pageId="13" pageNumber="136" refString="Chuang, L. C, Lin, Y. S. &amp; Liang, S. H. (2006) Ecomorphological comparison and habitat preference of 2 cyprinid fishes, Varicorhinus barbatulus and Candidia barbatus, in Hapen Creek of northern Taiwan. Zoological Studies 45, 114 - 123." type="journal article" year="2006">
Chuang
<emphasis id="B932D545B567FFBA0527FDC3FC7ED17E" box="[846,897,583,608]" italics="true" pageId="13" pageNumber="136">et al</emphasis>
. 2006
</bibRefCitation>
) and may be a useful tool for predicting habitat preferences (
<bibRefCitation id="EFD774A6B567FFBA0710FDE6FD92D19A" author="Stolbunov" box="[377,621,620,645]" pageId="13" pageNumber="136" refString="Stolbunov, I. A., Hai Than, N. T. &amp; Pavlov, D. S. (2011) Morphological and behavioral variation of Rasbora paviei (Cyprinidae, Cypriniformes) from lotic and limnic habitats (Central Vietnam). Journal of Ichthyology, 51, 352 - 357. http: // dx. doi. org / 10.1134 / S 003294521102010 X" type="journal article" year="2011">
Stolbunov
<emphasis id="B932D545B567FFBA079AFDE7FDDBD19B" box="[499,548,620,645]" italics="true" pageId="13" pageNumber="136">et al</emphasis>
. 2011
</bibRefCitation>
). For
<taxonomicName id="4C4672D4B567FFBA04C5FDE7FC9DD19B" box="[684,866,620,645]" class="Actinopterygii" family="Cyprinidae" genus="Onychostoma" kingdom="Animalia" order="Cypriniformes" pageId="13" pageNumber="136" phylum="Chordata" rank="species" species="krongnoensis">
<collectingCountry id="F35149C7B567FFBA04C5FDE7FD3FD19A" box="[684,704,621,644]" name="Iceland" pageId="13" pageNumber="136">
<emphasis id="B932D545B567FFBA04C5FDE7FD3FD19A" box="[684,704,621,644]" italics="true" pageId="13" pageNumber="136">O</emphasis>
</collectingCountry>
.
<emphasis id="B932D545B567FFBA04A7FDE6FC9DD19B" box="[718,866,620,645]" italics="true" pageId="13" pageNumber="136">krongnoensis</emphasis>
</taxonomicName>
, the slender, streamlined body with a large number of predorsal scales (1517) and long caudal peduncle may provide greater swimming ability and allow it to inhabit rapid rivers and streams at high elevations such as those in the Ea Krong No drainage. In contrast,
<taxonomicName id="4C4672D4B567FFBA028FFD3FFA63D1D3" box="[1254,1436,692,717]" class="Actinopterygii" family="Cyprinidae" genus="Onychostoma" kingdom="Animalia" order="Cypriniformes" pageId="13" pageNumber="136" phylum="Chordata" rank="species" species="dongnaiensis">
<collectingCountry id="F35149C7B567FFBA028FFD3FFB05D1D2" box="[1254,1274,693,716]" name="Iceland" pageId="13" pageNumber="136">
<emphasis id="B932D545B567FFBA028FFD3FFB05D1D2" box="[1254,1274,693,716]" italics="true" pageId="13" pageNumber="136">O</emphasis>
</collectingCountry>
.
<emphasis id="B932D545B567FFBA0361FD3EFA63D1D3" box="[1288,1436,692,717]" italics="true" pageId="13" pageNumber="136">dongnaiensis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C4672D4B567FFBA06ACFD52FEC7D1EE" box="[197,312,727,752]" class="Actinopterygii" family="Cyprinidae" genus="Onychostoma" kingdom="Animalia" order="Cypriniformes" pageId="13" pageNumber="136" phylum="Chordata" rank="species" species="leptura">
<emphasis id="B932D545B567FFBA06ACFD52FF27D1F1" box="[197,216,728,751]" italics="true" pageId="13" pageNumber="136">
<collectingCountry id="F35149C7B567FFBA06ACFD52FF27D1F1" box="[197,216,728,751]" name="Iceland" pageId="13" pageNumber="136">O</collectingCountry>
</emphasis>
.
<emphasis id="B932D545B567FFBA068EFD5DFEC7D1EE" box="[231,312,727,752]" italics="true" pageId="13" pageNumber="136">leptura</emphasis>
</taxonomicName>
have a deeper body, fewer predorsal scales (1215) and a short caudal peduncle, which may be more suitable features for slow, low elevation streams such as the middle of the Dong Nai and Red river drainages (Hoang
<emphasis id="B932D545B567FFBA069BFCABFEDBD026" box="[242,292,799,824]" italics="true" pageId="13" pageNumber="136">et al</emphasis>
. pers.obs.). Further studies on the capabilities and physiological mechanisms of
<taxonomicName id="4C4672D4B567FFBA02DAFCAAFA94D026" box="[1203,1387,799,824]" class="Actinopterygii" family="Cyprinidae" genus="Onychostoma" kingdom="Animalia" order="Cypriniformes" pageId="13" pageNumber="136" phylum="Chordata" rank="species" species="krongnoensis">
<collectingCountry id="F35149C7B567FFBA02DAFCAAFB38D029" box="[1203,1223,800,823]" name="Iceland" pageId="13" pageNumber="136">
<emphasis id="B932D545B567FFBA02DAFCAAFB38D029" box="[1203,1223,800,823]" italics="true" pageId="13" pageNumber="136">O</emphasis>
</collectingCountry>
.
<emphasis id="B932D545B567FFBA02BFFC95FA94D026" box="[1238,1387,799,824]" italics="true" pageId="13" pageNumber="136">krongnoensis</emphasis>
</taxonomicName>
that allow the species to live in such rapid streams would be interesting.
</paragraph>
<paragraph id="8BF90957B567FFBA06AEFCEDFD57D0F3" blockId="13.[151,1436,151,1005]" pageId="13" pageNumber="136">
Pronounced sexual dimorphism in the anal-fin size has been observed for several species of
<taxonomicName id="4C4672D4B567FFBA036BFCEDFA63D09E" box="[1282,1436,871,896]" class="Actinopterygii" family="Cyprinidae" genus="Onychostoma" kingdom="Animalia" order="Cypriniformes" pageId="13" pageNumber="136" phylum="Chordata" rank="genus">
<emphasis id="B932D545B567FFBA036BFCEDFA63D09E" box="[1282,1436,871,896]" italics="true" pageId="13" pageNumber="136">Onychostoma</emphasis>
</taxonomicName>
(Hoang
<emphasis id="B932D545B567FFBA069CFC07FED6D0BB" box="[245,297,908,933]" italics="true" pageId="13" pageNumber="136">et al</emphasis>
. pers. obs.), but has not been reported. This morphological character highlights the importance of making intraspecific comparisons separately for each sex. The functional role of the sexual dimorphism in
<taxonomicName id="4C4672D4B567FFBA03E8FC3AFED3D0F3" class="Actinopterygii" family="Cyprinidae" genus="Onychostoma" kingdom="Animalia" order="Cypriniformes" pageId="13" pageNumber="136" phylum="Chordata" rank="species" species="krongnoensis">
<emphasis id="B932D545B567FFBA03E8FC3AFED3D0F3" italics="true" pageId="13" pageNumber="136">
<collectingCountry id="F35149C7B567FFBA03E8FC3AFA67D0D9" box="[1409,1432,944,967]" name="Iceland" pageId="13" pageNumber="136">O</collectingCountry>
. krongnoensis
</emphasis>
</taxonomicName>
and
<taxonomicName id="4C4672D4B567FFBA0708FC5FFDE7D0F3" box="[353,536,980,1005]" class="Actinopterygii" family="Cyprinidae" genus="Onychostoma" kingdom="Animalia" order="Cypriniformes" pageId="13" pageNumber="136" phylum="Chordata" rank="species" species="dongnaiensis">
<emphasis id="B932D545B567FFBA0708FC5FFDE7D0F3" box="[353,536,980,1005]" italics="true" pageId="13" pageNumber="136">
<collectingCountry id="F35149C7B567FFBA0708FC5FFE87D0F2" box="[353,376,981,1004]" name="Iceland" pageId="13" pageNumber="136">O</collectingCountry>
. dongnaiensis
</emphasis>
</taxonomicName>
is unknown.
</paragraph>
<paragraph id="8BF90957B567FFBA06FEFB96FE25D6FB" blockId="13.[151,1437,1051,1797]" pageId="13" pageNumber="136">
<emphasis id="B932D545B567FFBA06FEFB96FCFCD72B" bold="true" box="[151,771,1051,1077]" pageId="13" pageNumber="136">
Species diversity of
<taxonomicName id="4C4672D4B567FFBA07E0FB91FDD7D72A" box="[393,552,1051,1076]" class="Actinopterygii" family="Cyprinidae" genus="Onychostoma" kingdom="Animalia" order="Cypriniformes" pageId="13" pageNumber="136" phylum="Chordata" rank="genus">
<emphasis id="B932D545B567FFBA07E0FB91FDD7D72A" bold="true" box="[393,552,1051,1076]" italics="true" pageId="13" pageNumber="136">Onychostoma</emphasis>
</taxonomicName>
in Mekong basin.
</emphasis>
<taxonomicName id="4C4672D4B567FFBA0565FB96FBBBD72B" box="[780,1092,1052,1077]" class="Actinopterygii" family="Cyprinidae" genus="Onychostoma" kingdom="Animalia" order="Cypriniformes" pageId="13" pageNumber="136" phylum="Chordata" rank="species" species="krongnoensis">
<emphasis id="B932D545B567FFBA0565FB96FBBBD72B" box="[780,1092,1052,1077]" italics="true" pageId="13" pageNumber="136">Onychostoma krongnoensis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C4672D4B567FFBA0217FB97FAC9D72B" box="[1150,1334,1052,1077]" class="Actinopterygii" family="Cyprinidae" genus="Onychostoma" kingdom="Animalia" order="Cypriniformes" pageId="13" pageNumber="136" phylum="Chordata" rank="species" species="dongnaiensis">
<collectingCountry id="F35149C7B567FFBA0217FB97FB6DD72A" box="[1150,1170,1053,1076]" name="Iceland" pageId="13" pageNumber="136">
<emphasis id="B932D545B567FFBA0217FB97FB6DD72A" box="[1150,1170,1053,1076]" italics="true" pageId="13" pageNumber="136">O</emphasis>
</collectingCountry>
.
<emphasis id="B932D545B567FFBA02CBFB96FAC9D72B" box="[1186,1334,1052,1077]" italics="true" pageId="13" pageNumber="136">dongnaiensis</emphasis>
</taxonomicName>
are very similar to
<taxonomicName id="4C4672D4B567FFBA077FFBCAFE62D746" box="[278,413,1087,1112]" class="Actinopterygii" family="Cyprinidae" genus="Barbus" kingdom="Animalia" order="Cypriniformes" pageId="13" pageNumber="136" phylum="Chordata" rank="species" species="gerlachi">
<collectingCountry id="F35149C7B567FFBA077FFBCAFED5D749" box="[278,298,1088,1111]" name="Iceland" pageId="13" pageNumber="136">
<emphasis id="B932D545B567FFBA077FFBCAFED5D749" box="[278,298,1088,1111]" italics="true" pageId="13" pageNumber="136">O</emphasis>
</collectingCountry>
.
<emphasis id="B932D545B567FFBA0729FBB5FE62D746" box="[320,413,1087,1112]" italics="true" pageId="13" pageNumber="136">gerlachi</emphasis>
</taxonomicName>
,
<taxonomicName id="4C4672D4B567FFBA07DBFBCAFDB9D746" box="[434,582,1088,1112]" class="Actinopterygii" family="Cyprinidae" genus="Onychostoma" kingdom="Animalia" order="Cypriniformes" pageId="13" pageNumber="136" phylum="Chordata" rank="species" species="fusiforme">
<collectingCountry id="F35149C7B567FFBA07DBFBCAFE39D749" box="[434,454,1088,1111]" name="Iceland" pageId="13" pageNumber="136">
<emphasis id="B932D545B567FFBA07DBFBCAFE39D749" box="[434,454,1088,1111]" italics="true" pageId="13" pageNumber="136">O</emphasis>
</collectingCountry>
.
<emphasis id="B932D545B567FFBA07B5FBCAFDB9D746" box="[476,582,1088,1112]" italics="true" pageId="13" pageNumber="136">fusiforme</emphasis>
</taxonomicName>
and
<taxonomicName id="4C4672D4B567FFBA04E3FBCAFCC2D746" box="[650,829,1087,1112]" class="Actinopterygii" family="Cyprinidae" genus="Onychostoma" kingdom="Animalia" order="Cypriniformes" pageId="13" pageNumber="136" phylum="Chordata" rank="species" species="meridionale">
<collectingCountry id="F35149C7B567FFBA04E3FBCAFD61D749" box="[650,670,1088,1111]" name="Iceland" pageId="13" pageNumber="136">
<emphasis id="B932D545B567FFBA04E3FBCAFD61D749" box="[650,670,1088,1111]" italics="true" pageId="13" pageNumber="136">O</emphasis>
</collectingCountry>
.
<emphasis id="B932D545B567FFBA04DAFBB5FCC2D746" box="[691,829,1087,1112]" italics="true" pageId="13" pageNumber="136">meridionale</emphasis>
</taxonomicName>
, and have probably been misidentified due to the similarities in their adult overall morphology and geographic distributions.
<bibRefCitation id="EFD774A6B567FFBA026EFBEEFB20D762" box="[1031,1247,1124,1149]" pageId="13" pageNumber="136" refString="Shan, X., Lin, R., Yuem, P. &amp; Chu, X. (2000) Onychostoma. In: Yue, P (Ed.), Fauna Sinica, Osteichthyes. Cypriniformes III. Science Press, Beijing, pp. 126 - 147." type="book chapter">
Shan
<emphasis id="B932D545B567FFBA0223FBEFFB80D763" box="[1098,1151,1124,1149]" italics="true" pageId="13" pageNumber="136">et al</emphasis>
. (2000)
</bibRefCitation>
, Kottelat (2001, 2009),
<bibRefCitation id="EFD774A6B567FFBA0683FB0DFD46D7BE" author="Lothongkham" box="[234,697,1159,1184]" pageId="13" pageNumber="136" refString="Lothongkham, A. &amp; Musikasinthorn, P (2005) First record of the cyprinid fish, Onychostoma gerlachi (Cypriniformes: Cyprinidae), from the Nan river basin of the Chao Phraya river system in northern Thailand. The Natural History Bulletin of the Siam Society, 53, 237 - 243." type="journal article" year="2005">Lothongkham &amp; Musikasinthorn (2005)</bibRefCitation>
and
<bibRefCitation id="EFD774A6B567FFBA0491FB02FC2DD7BE" box="[760,978,1159,1184]" pageId="13" pageNumber="136" refString="Kano, Y., Adnan, M. S. B., Grudpan, C., Grudpan, J., Magtoon, W., Musikasinthorn, P., Natori, Y., Ottomanski, S., Praxaysonbath, B., Phongsa, K., Rangsiruji, A., Shibukawa, K., Shimatani, Y., So, N., Suvarnaraksha, A., Thach, P., Thanh, P. N., Tran, D. D., Utsugi, K. &amp; Yamashita, T. (2013) An online database on freshwater fish diversity and distribution in Mainland Southeast Asia. Ichthyological Research, 60, 293 - 295. Available from: http: // ffish. asia / (accessed 26 March 2014)" type="journal article">
Kano
<emphasis id="B932D545B567FFBA0529FB03FC8AD7BE" box="[832,885,1159,1184]" italics="true" pageId="13" pageNumber="136">et al</emphasis>
. (2013)
</bibRefCitation>
described
<taxonomicName id="4C4672D4B567FFBA023AFB02FB28D7BE" box="[1107,1239,1159,1184]" class="Actinopterygii" family="Cyprinidae" genus="Barbus" kingdom="Animalia" order="Cypriniformes" pageId="13" pageNumber="136" phylum="Chordata" rank="species" species="gerlachi">
<collectingCountry id="F35149C7B567FFBA023AFB02FB98D781" box="[1107,1127,1160,1183]" name="Iceland" pageId="13" pageNumber="136">
<emphasis id="B932D545B567FFBA023AFB02FB98D781" box="[1107,1127,1160,1183]" italics="true" pageId="13" pageNumber="136">O</emphasis>
</collectingCountry>
.
<emphasis id="B932D545B567FFBA0213FB0DFB28D7BE" box="[1146,1239,1159,1184]" italics="true" pageId="13" pageNumber="136">gerlachi</emphasis>
</taxonomicName>
as a widespread species from the Pearl River, upper Red River to the Langcang River, Nam Ou, Se Banghiang River-Mekong basin, and the Nan River-Chao Phraya basin.
<bibRefCitation id="EFD774A6B567FFBA040FFB45FCE7D7F6" author="Kottelat" box="[614,792,1231,1256]" pageId="13" pageNumber="136" refString="Kottelat, M. (1998) Fishes of the Nam Theun and Xe Bangfai basins, Laos, with diagnoses of twenty-two new species (Teleostei: Cyprinidae, Balitoridae, Cobitidae, Coiidae and Odontobutidae). Ichthyological Exploration of Freshwaters, 9, 1 - 128." type="journal article" year="1998">Kottelat (1998)</bibRefCitation>
,
<bibRefCitation id="EFD774A6B567FFBA0541FB45FC10D7F6" box="[808,1007,1231,1256]" pageId="13" pageNumber="136" refString="Xin, Q., Zhang, E. &amp; Cao, W. X. (2009) Onychostoma virgulatum, a new species of cyprinid fish from southern Anhui Province, South China. Ichthyological Exploration of Freshwaters, 20, 255 - 266." type="journal article">
Xin
<emphasis id="B932D545B567FFBA0534FB5BFC6CD7F6" box="[861,915,1231,1256]" italics="true" pageId="13" pageNumber="136">et al</emphasis>
. (2009)
</bibRefCitation>
and
<bibRefCitation id="EFD774A6B567FFBA0247FB5AFAF7D7F6" box="[1070,1288,1231,1256]" pageId="13" pageNumber="136" refString="Kano, Y., Adnan, M. S. B., Grudpan, C., Grudpan, J., Magtoon, W., Musikasinthorn, P., Natori, Y., Ottomanski, S., Praxaysonbath, B., Phongsa, K., Rangsiruji, A., Shibukawa, K., Shimatani, Y., So, N., Suvarnaraksha, A., Thach, P., Thanh, P. N., Tran, D. D., Utsugi, K. &amp; Yamashita, T. (2013) An online database on freshwater fish diversity and distribution in Mainland Southeast Asia. Ichthyological Research, 60, 293 - 295. Available from: http: // ffish. asia / (accessed 26 March 2014)" type="journal article">
Kano
<emphasis id="B932D545B567FFBA021FFB5BFB54D7F6" box="[1142,1195,1231,1256]" italics="true" pageId="13" pageNumber="136">et al</emphasis>
. (2013)
</bibRefCitation>
recorded
<taxonomicName id="4C4672D4B567FFBA03E8FB5AFEFED612" class="Actinopterygii" family="Cyprinidae" genus="Onychostoma" kingdom="Animalia" order="Cypriniformes" pageId="13" pageNumber="136" phylum="Chordata" rank="species" species="fusiforme">
<collectingCountry id="F35149C7B567FFBA03E8FB5AFA6AD7F9" box="[1409,1429,1232,1255]" name="Iceland" pageId="13" pageNumber="136">
<emphasis id="B932D545B567FFBA03E8FB5AFA6AD7F9" box="[1409,1429,1232,1255]" italics="true" pageId="13" pageNumber="136">O</emphasis>
</collectingCountry>
.
<emphasis id="B932D545B567FFBA06FEFB7EFEFED612" box="[151,257,1268,1292]" italics="true" pageId="13" pageNumber="136">fusiforme</emphasis>
</taxonomicName>
as occurring from the Langcang River (Yunnan,
<collectingCountry id="F35149C7B567FFBA0542FB7EFC8DD613" box="[811,882,1268,1293]" name="China" pageId="13" pageNumber="136">China</collectingCountry>
), Ing River, Kok River (
<collectingCountry id="F35149C7B567FFBA02EAFB7EFB15D613" box="[1155,1258,1268,1293]" name="Thailand" pageId="13" pageNumber="136">Thailand</collectingCountry>
) to Nam Theun River (
<collectingCountry id="F35149C7B567FFBA068EFA92FEDCD62E" box="[231,291,1304,1328]" name="Laos" pageId="13" pageNumber="136">Laos</collectingCountry>
).
<bibRefCitation id="EFD774A6B567FFBA0751FA9DFE20D62E" author="Kottelat" box="[312,479,1303,1328]" pageId="13" pageNumber="136" refString="Kottelat, M. (1998) Fishes of the Nam Theun and Xe Bangfai basins, Laos, with diagnoses of twenty-two new species (Teleostei: Cyprinidae, Balitoridae, Cobitidae, Coiidae and Odontobutidae). Ichthyological Exploration of Freshwaters, 9, 1 - 128." type="journal article" year="1998">Kottelat (1998</bibRefCitation>
, 2007, 2011),
<bibRefCitation id="EFD774A6B567FFBA04EEFA92FCA3D62E" box="[647,860,1303,1328]" pageId="13" pageNumber="136" refString="Kano, Y., Adnan, M. S. B., Grudpan, C., Grudpan, J., Magtoon, W., Musikasinthorn, P., Natori, Y., Ottomanski, S., Praxaysonbath, B., Phongsa, K., Rangsiruji, A., Shibukawa, K., Shimatani, Y., So, N., Suvarnaraksha, A., Thach, P., Thanh, P. N., Tran, D. D., Utsugi, K. &amp; Yamashita, T. (2013) An online database on freshwater fish diversity and distribution in Mainland Southeast Asia. Ichthyological Research, 60, 293 - 295. Available from: http: // ffish. asia / (accessed 26 March 2014)" type="journal article">
Kano
<emphasis id="B932D545B567FFBA04A4FA93FCFED62E" box="[717,769,1303,1328]" italics="true" pageId="13" pageNumber="136">et al</emphasis>
. (2013)
</bibRefCitation>
and our survey (2013) described and recorded
<taxonomicName id="4C4672D4B567FFBA03E8FA92FEE1D64B" class="Actinopterygii" family="Cyprinidae" genus="Onychostoma" kingdom="Animalia" order="Cypriniformes" pageId="13" pageNumber="136" phylum="Chordata" rank="species" species="meridionale">
<collectingCountry id="F35149C7B567FFBA03E8FA92FA6AD631" box="[1409,1429,1304,1327]" name="Iceland" pageId="13" pageNumber="136">
<emphasis id="B932D545B567FFBA03E8FA92FA6AD631" box="[1409,1429,1304,1327]" italics="true" pageId="13" pageNumber="136">O</emphasis>
</collectingCountry>
.
<emphasis id="B932D545B567FFBA06FEFAB6FEE1D64B" box="[151,286,1340,1365]" italics="true" pageId="13" pageNumber="136">meridionale</emphasis>
</taxonomicName>
from the Se Bangfai River, Sekong River (
<collectingCountry id="F35149C7B567FFBA056BFAB6FCC2D64A" box="[770,829,1340,1364]" name="Laos" pageId="13" pageNumber="136">Laos</collectingCountry>
), Sesan River (
<collectingCountry id="F35149C7B567FFBA058FFAB6FBA0D64B" box="[998,1119,1340,1365]" name="Cambodia" pageId="13" pageNumber="136">Cambodia</collectingCountry>
), Sa Thay River-tributary of the Sesan River partly in the
<collectingCountry id="F35149C7B567FFBA0793FAD5FDA6D666" box="[506,601,1375,1400]" name="Vietnam" pageId="13" pageNumber="136">Vietnam</collectingCountry>
Central Highlands to the Tonle Sap River (
<collectingCountry id="F35149C7B567FFBA020EFAD5FB1ED666" box="[1127,1249,1375,1400]" name="Cambodia" pageId="13" pageNumber="136">Cambodia</collectingCountry>
). Even amongst geographically close river basins such as the Sekong River, Sesan River and Sre Pok River in the middle Mekong basin, there are distinct species of
<taxonomicName id="4C4672D4B567FFBA0448FA2DFD44D6DE" box="[545,699,1447,1472]" class="Actinopterygii" family="Cyprinidae" genus="Onychostoma" kingdom="Animalia" order="Cypriniformes" pageId="13" pageNumber="136" phylum="Chordata" rank="genus">
<emphasis id="B932D545B567FFBA0448FA2DFD44D6DE" box="[545,699,1447,1472]" italics="true" pageId="13" pageNumber="136">Onychostoma</emphasis>
</taxonomicName>
present:
<taxonomicName id="4C4672D4B567FFBA054DFA22FC23D6DE" box="[804,988,1447,1472]" class="Actinopterygii" family="Cyprinidae" genus="Onychostoma" kingdom="Animalia" order="Cypriniformes" pageId="13" pageNumber="136" phylum="Chordata" rank="species" species="krongnoensis">
<collectingCountry id="F35149C7B567FFBA054DFA22FCC7D6A1" box="[804,824,1448,1471]" name="Iceland" pageId="13" pageNumber="136">
<emphasis id="B932D545B567FFBA054DFA22FCC7D6A1" box="[804,824,1448,1471]" italics="true" pageId="13" pageNumber="136">O</emphasis>
</collectingCountry>
.
<emphasis id="B932D545B567FFBA0521FA2DFC23D6DE" box="[840,988,1447,1472]" italics="true" pageId="13" pageNumber="136">krongnoensis</emphasis>
</taxonomicName>
in the Srepok River,
<taxonomicName id="4C4672D4B567FFBA02BAFA22FA82D6DE" box="[1235,1405,1447,1472]" class="Actinopterygii" family="Cyprinidae" genus="Onychostoma" kingdom="Animalia" order="Cypriniformes" pageId="13" pageNumber="136" phylum="Chordata" rank="species" species="meridionale">
<collectingCountry id="F35149C7B567FFBA02BAFA22FB18D6A1" box="[1235,1255,1448,1471]" name="Iceland" pageId="13" pageNumber="136">
<emphasis id="B932D545B567FFBA02BAFA22FB18D6A1" box="[1235,1255,1448,1471]" italics="true" pageId="13" pageNumber="136">O</emphasis>
</collectingCountry>
.
<emphasis id="B932D545B567FFBA029FFA2DFA82D6DE" box="[1270,1405,1447,1472]" italics="true" pageId="13" pageNumber="136">meridionale</emphasis>
</taxonomicName>
in the Sekong and Sesan rivers.
</paragraph>
<paragraph id="8BF90957B567FFBA06AEFA65FCC0D41B" blockId="13.[151,1437,1051,1797]" pageId="13" pageNumber="136">
<taxonomicName id="4C4672D4B567FFBA06AEFA65FE01D516" box="[199,510,1519,1544]" class="Actinopterygii" family="Cyprinidae" genus="Onychostoma" kingdom="Animalia" order="Cypriniformes" pageId="13" pageNumber="136" phylum="Chordata" rank="species" species="krongnoensis">
<emphasis id="B932D545B567FFBA06AEFA65FE01D516" box="[199,510,1519,1544]" italics="true" pageId="13" pageNumber="136">Onychostoma krongnoensis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C4672D4B567FFBA0451FA7AFD0FD516" box="[568,752,1519,1544]" class="Actinopterygii" family="Cyprinidae" genus="Onychostoma" kingdom="Animalia" order="Cypriniformes" pageId="13" pageNumber="136" phylum="Chordata" rank="species" species="dongnaiensis">
<collectingCountry id="F35149C7B567FFBA0451FA7AFDB3D519" box="[568,588,1520,1543]" name="Iceland" pageId="13" pageNumber="136">
<emphasis id="B932D545B567FFBA0451FA7AFDB3D519" box="[568,588,1520,1543]" italics="true" pageId="13" pageNumber="136">O</emphasis>
</collectingCountry>
.
<emphasis id="B932D545B567FFBA0435FA65FD0FD516" box="[604,752,1519,1544]" italics="true" pageId="13" pageNumber="136">dongnaiensis</emphasis>
</taxonomicName>
are the first records of the genus in the upper Srepok River and middle Dong Nai River on the Langbiang Plateau, southern
<collectingCountry id="F35149C7B567FFBA050EF99EFC35D533" box="[871,970,1556,1581]" name="Vietnam" pageId="13" pageNumber="136">Vietnam</collectingCountry>
. Among species of the Mekong group,
<taxonomicName id="4C4672D4B567FFBA03E8F99FFF0BD54E" class="Actinopterygii" family="Cyprinidae" genus="Barbus" kingdom="Animalia" order="Cypriniformes" pageId="13" pageNumber="136" phylum="Chordata" rank="species" species="gerlachi">
<emphasis id="B932D545B567FFBA03E8F99FFF0BD54E" italics="true" pageId="13" pageNumber="136">
<collectingCountry id="F35149C7B567FFBA03E8F99FFA67D532" box="[1409,1432,1557,1580]" name="Iceland" pageId="13" pageNumber="136">O</collectingCountry>
. gerlachi
</emphasis>
</taxonomicName>
occurs the furthest north. This species could be the ancestral lineage of the Mekong clade. Despite the lack of
<taxonomicName id="4C4672D4B567FFBA06DCF9D7FEBDD56A" box="[181,322,1628,1652]" class="Actinopterygii" family="Cyprinidae" genus="Onychostoma" kingdom="Animalia" order="Cypriniformes" pageId="13" pageNumber="136" phylum="Chordata" rank="species" species="fusiforme">
<collectingCountry id="F35149C7B567FFBA06DCF9D7FF36D56A" box="[181,201,1629,1652]" name="Iceland" pageId="13" pageNumber="136">
<emphasis id="B932D545B567FFBA06DCF9D7FF36D56A" box="[181,201,1629,1652]" italics="true" pageId="13" pageNumber="136">O</emphasis>
</collectingCountry>
.
<emphasis id="B932D545B567FFBA06B1F9D6FEBDD56A" box="[216,322,1628,1652]" italics="true" pageId="13" pageNumber="136">fusiforme</emphasis>
</taxonomicName>
in our molecular data, phylogenetic analysis and geographical distribution suggest that this lineage may have migrated towards the southern Mekong basin. The geological event that separated the Dong Nai dranage from the Mekong drainage would have isolated
<taxonomicName id="4C4672D4B567FFBA056CF92FFC37D5A3" box="[773,968,1700,1725]" class="Actinopterygii" family="Cyprinidae" genus="Onychostoma" kingdom="Animalia" order="Cypriniformes" pageId="13" pageNumber="136" phylum="Chordata" rank="species" species="dongnaiensis">
<collectingCountry id="F35149C7B567FFBA056CF92FFCE6D5A2" box="[773,793,1701,1724]" name="Iceland" pageId="13" pageNumber="136">
<emphasis id="B932D545B567FFBA056CF92FFCE6D5A2" box="[773,793,1701,1724]" italics="true" pageId="13" pageNumber="136">O</emphasis>
</collectingCountry>
.
<emphasis id="B932D545B567FFBA055DF92EFC37D5A3" box="[820,968,1700,1725]" italics="true" pageId="13" pageNumber="136">dongnaiensis</emphasis>
</taxonomicName>
from
<taxonomicName id="4C4672D4B567FFBA024DF92FFB18D5A3" box="[1060,1255,1700,1725]" class="Actinopterygii" family="Cyprinidae" genus="Onychostoma" kingdom="Animalia" order="Cypriniformes" pageId="13" pageNumber="136" phylum="Chordata" rank="species" species="krongnoensis">
<collectingCountry id="F35149C7B567FFBA024DF92FFBC7D5A2" box="[1060,1080,1701,1724]" name="Iceland" pageId="13" pageNumber="136">
<emphasis id="B932D545B567FFBA024DF92FFBC7D5A2" box="[1060,1080,1701,1724]" italics="true" pageId="13" pageNumber="136">O</emphasis>
</collectingCountry>
.
<emphasis id="B932D545B567FFBA023AF92EFB18D5A3" box="[1107,1255,1700,1725]" italics="true" pageId="13" pageNumber="136">krongnoensis</emphasis>
</taxonomicName>
. Detailed and comprehensive surveys to gain a better understanding of geographical distributions, and molecular phylogenetic analysis for
<taxonomicName id="4C4672D4B567FFBA0749F966FE45D41B" box="[288,442,1772,1797]" class="Actinopterygii" family="Cyprinidae" genus="Onychostoma" kingdom="Animalia" order="Cypriniformes" pageId="13" pageNumber="136" phylum="Chordata" rank="genus">
<emphasis id="B932D545B567FFBA0749F966FE45D41B" box="[288,442,1772,1797]" italics="true" pageId="13" pageNumber="136">Onychostoma</emphasis>
</taxonomicName>
in the Mekong basin are essential.
</paragraph>
</subSubSection>
</treatment>
</document>