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<document id="60A1BADF83DC359FDC35DE8456A85B81" ID-DOI="10.11646/zootaxa.3635.5.3" ID-GBIF-Dataset="012c682c-6953-4e8e-a3fe-8af53403d142" ID-ISSN="1175-5326" ID-Zenodo-Dep="220591" ID-ZooBank="40372BBF-2235-4770-AEB8-41E342DFF770" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460587444626" checkinUser="plazi" docAuthor="Mikhaljova, Elena V., Burkitbaeva, Ulzhan D., Tuf, Ivan H. &amp; Ulykpan, Kaman" docDate="2013" docId="03AC87CDFFC6DC7FFF52FF0B064A01A7" docLanguage="en" docName="zt03635p544.pdf" docOrigin="Zootaxa 3635 (5)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Altajosoma bukhtarma Mikhaljova, sp. nov." docType="treatment" docVersion="8" lastPageNumber="538" masterDocId="FF95FFB5FFC2DC7AFFC5FF9C046F024B" masterDocTitle="The millipede order Chordeumatida (Diplopoda) in Kazakhstan, with descriptions of three new species" masterLastPageNumber="544" masterPageNumber="533" pageNumber="537" updateTime="1698325424133" updateUser="plazi">
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<mods:titleInfo id="C97FD56EFDB7E75B1209E75E5421FDA2">
<mods:title id="BC97780E0C0039B02BE821250906DD6C">The millipede order Chordeumatida (Diplopoda) in Kazakhstan, with descriptions of three new species</mods:title>
</mods:titleInfo>
<mods:name id="759E67C0751B347E5A2EE88F7A556C5C" type="personal">
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<mods:namePart id="3715DF2CBF7B0C2783E8469AFE94A2CD">Mikhaljova, Elena V.</mods:namePart>
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<mods:namePart id="F6599EED2172370E964C3361BD2341A2">Burkitbaeva, Ulzhan D.</mods:namePart>
</mods:name>
<mods:name id="BAED0527FFAEB4C6059D35E475426028" type="personal">
<mods:role id="F8EEB5C8F83A7C17C6C9D68B6494BE74">
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<mods:namePart id="DF4569AA2A13C7107DEFBB73780D8165">Tuf, Ivan H.</mods:namePart>
</mods:name>
<mods:name id="B0FE7B95DAC7564F43B7F1EB21D7A33E" type="personal">
<mods:role id="2EC17BEBC1C0EBE9B7FAE07C7EB41D41">
<mods:roleTerm id="2A3BD609818F41CC9AC2187BEDE7638E">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="391889B33CE138EB49AAC59BFF4A9879">Ulykpan, Kaman</mods:namePart>
</mods:name>
<mods:typeOfResource id="934FA2B8E554F51072A75DC722EE1E49">text</mods:typeOfResource>
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<mods:titleInfo id="7749B01F89E8266C4B58D22E8EDB6364">
<mods:title id="06C8C89E52A58E0C515615E95DFCA8E7">Zootaxa</mods:title>
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<mods:part id="C35DDFCD1D0D393B4FCFA90A7FAE73F8">
<mods:date id="13949228BF33194165C7785728EC0AFD">2013</mods:date>
<mods:detail id="AAE04F11C46E95744CE7F77170192414" type="volume">
<mods:number id="1557679C604E5A348FBC392385F9C621">3635</mods:number>
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<mods:detail id="BF953F662A1A4EA54DCBEAFE65DC1E3D" type="issue">
<mods:number id="F37154F81BFFDDF1B5B870EBC0D02284">5</mods:number>
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<mods:identifier id="D29612AF050CBEC48266FF38275B5DBA" type="DOI">10.11646/zootaxa.3635.5.3</mods:identifier>
<mods:identifier id="648E69873874A311098882ADDADD0D65" type="GBIF-Dataset">012c682c-6953-4e8e-a3fe-8af53403d142</mods:identifier>
<mods:identifier id="7774D78960F21D10BF3CEBFBF26693A9" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="7AB1A4B9A02C25F7078571BF76F58A69" type="Zenodo-Dep">220591</mods:identifier>
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<treatment id="03AC87CDFFC6DC7FFF52FF0B064A01A7" ID-DOI="http://doi.org/10.5281/zenodo.6145104" ID-GBIF-Taxon="119565225" ID-Zenodo-Dep="6145104" LSID="urn:lsid:plazi:treatment:03AC87CDFFC6DC7FFF52FF0B064A01A7" httpUri="http://treatment.plazi.org/id/03AC87CDFFC6DC7FFF52FF0B064A01A7" lastPageId="5" lastPageNumber="538" pageId="4" pageNumber="537">
<subSubSection id="C31F6550FFC6DC7EFF52FF0B04970299" pageId="4" pageNumber="537" type="nomenclature">
<paragraph id="8BBA36DBFFC6DC7EFF52FF0B06C702FA" blockId="4.[151,680,151,210]" box="[151,680,151,177]" pageId="4" pageNumber="537">
<heading id="D0F281B7FFC6DC7EFF52FF0B06C702FA" bold="true" box="[151,680,151,177]" fontSize="11" level="1" pageId="4" pageNumber="537" reason="1">
<emphasis id="B971EAC9FFC6DC7EFF52FF0B06C702FA" bold="true" box="[151,680,151,177]" pageId="4" pageNumber="537">
<taxonomicName id="4C054D58FFC6DC7EFF52FF0B062B02FA" ID-CoL="C992" authority="Mikhaljova" authorityName="Mikhaljova" box="[151,580,151,177]" class="Diplopoda" family="Diplomaragnidae" genus="Altajosoma" kingdom="Animalia" order="Chordeumatida" pageId="4" pageNumber="537" phylum="Arthropoda" rank="species" species="bukhtarma" status="sp. nov.">
<emphasis id="B971EAC9FFC6DC7EFF52FF0B05C402FA" bold="true" box="[151,427,151,177]" italics="true" pageId="4" pageNumber="537">Altajosoma bukhtarma</emphasis>
Mikhaljova
</taxonomicName>
<taxonomicNameLabel id="A24257B2FFC6DC7EFD8EFF0B06C702FA" box="[587,680,151,177]" pageId="4" pageNumber="537" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BBA36DBFFC6DC7EFF52FF2604970299" blockId="4.[151,680,151,210]" box="[151,248,186,210]" pageId="4" pageNumber="537">
<figureCitation id="133E2A5EFFC6DC7EFF52FF2604970299" box="[151,248,186,210]" captionStart-0="FIGURES 5 7" captionStart-1="FIGURES 8 9" captionStartId-0="3.[151,264,1898,1921]" captionStartId-1="4.[151,264,1904,1927]" captionTargetBox-0="[335,1254,195,1841]" captionTargetBox-1="[249,1341,632,1876]" captionTargetId-0="figure@3.[322,1268,193,1843]" captionTargetId-1="figure@4.[241,1346,623,1883]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionText-0="FIGURES 5 7. Altajosoma bukhtarma sp. nov., male paratype. 5, gonopods, caudal view; 6, gonopods, front view; 7, basal parts of telopodites of anterior gonopods; lp, lateral sheath process of posterior gonopod colpocoxite; pp, posterior process of posterior gonopod angiocoxite; ms, mesal sheath processes of posterior gonopod colpocoxites; ap, anterior process of posterior gonopod angiocoxite." captionText-1="FIGURES 8 9. Altajosoma bukhtarma sp. nov., male paratype. 8, gonopods, caudal view; 9, gonopods, front view; lp, lateral sheath process of posterior gonopod colpocoxite; pp, posterior process of posterior gonopod angiocoxite; ap, anterior process of posterior gonopod angiocoxite; ms, mesal sheath processes of posterior gonopod colpocoxites. Scale in mm." httpUri-0="https://zenodo.org/record/220593/files/figure.png" httpUri-1="https://zenodo.org/record/220594/files/figure.png" pageId="4" pageNumber="537">Figs 59</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C31F6550FFC6DC7EFF52FE9E00210329" pageId="4" pageNumber="537" type="materials_examined">
<paragraph id="8BBA36DBFFC6DC7EFF52FE9E00210329" blockId="4.[151,1436,258,570]" pageId="4" pageNumber="537">
<emphasis id="B971EAC9FFC6DC7EFF52FE9E05EE0350" bold="true" box="[151,385,258,283]" pageId="4" pageNumber="537">Material examined.</emphasis>
<typeStatus id="54BE8879FFC6DC7EFE4DFE9F05800350" box="[392,495,259,283]" pageId="4" pageNumber="537" type="holotype">
<emphasis id="B971EAC9FFC6DC7EFE4DFE9F05800350" box="[392,495,259,283]" italics="true" pageId="4" pageNumber="537">Holotype</emphasis>
</typeStatus>
:
<specimenCount id="9D03FD52FFC6DC7EFE3AFE9E06250351" box="[511,586,258,282]" pageId="4" pageNumber="537" type="male">1 male</specimenCount>
(IBSS),
<collectingCountry id="F312764BFFC6DC7EFD6AFE9E075B0351" box="[687,820,258,282]" name="Kazakhstan" pageId="4" pageNumber="537">Kazakhstan</collectingCountry>
, Vostochno Kazakhstanskaya Area, Katon-Karagaiskii District,
<quantity id="4CFD9B3EFFC6DC7EFF39FEBA05390375" box="[252,342,294,318]" metricMagnitude="3" metricUnit="m" metricValue="1.39" pageId="4" pageNumber="537" unit="m" value="1390.0">1390 m</quantity>
, 49°08´499´´ N, 085°57´670´´ E,
<taxonomicName id="4C054D58FFC6DC7EFD25FEBB07730375" box="[736,796,295,318]" class="Pinopsida" family="Pinaceae" genus="Larix" kingdom="Plantae" order="Pinales" pageId="4" pageNumber="537" phylum="Tracheophyta" rank="genus">
<emphasis id="B971EAC9FFC6DC7EFD25FEBB07730375" box="[736,796,295,318]" italics="true" pageId="4" pageNumber="537">Larix</emphasis>
</taxonomicName>
forest,
<date id="FFBB101BFFC6DC7EFCB2FEBA005C0375" box="[887,1075,294,319]" pageId="4" pageNumber="537" value="2010-08-22">22 August 2010</date>
, leg. K. Ulykpan.
<typeStatus id="54BE8879FFC6DC7EFACEFEB401120374" box="[1291,1405,296,319]" pageId="4" pageNumber="537" type="paratype">
<emphasis id="B971EAC9FFC6DC7EFACEFEB401120374" box="[1291,1405,296,319]" italics="true" pageId="4" pageNumber="537">Paratypes</emphasis>
</typeStatus>
:
<specimenCount id="9D03FD52FFC6DC7EFA4BFEBA04A20329" pageId="4" pageNumber="537" type="male">1 male</specimenCount>
(PU),
<specimenCount id="9D03FD52FFC6DC7EFEDCFED6050B0329" box="[281,356,330,354]" pageId="4" pageNumber="537" type="male">1 male</specimenCount>
(IBSS), together with
<typeStatus id="54BE8879FFC6DC7EFDA7FED606A80329" box="[610,711,330,354]" pageId="4" pageNumber="537" type="holotype">holotype</typeStatus>
,
<date id="FFBB101BFFC6DC7EFD17FED607E70329" box="[722,904,330,355]" pageId="4" pageNumber="537" value="2010-08-22">22 August 2010</date>
, leg. K. Ulykpan.
</paragraph>
</subSubSection>
<subSubSection id="C31F6550FFC6DC7EFF02FEF207510385" pageId="4" pageNumber="537" type="diagnosis">
<paragraph id="8BBA36DBFFC6DC7EFF02FEF207510385" blockId="4.[151,1436,258,570]" pageId="4" pageNumber="537">
<emphasis id="B971EAC9FFC6DC7EFF02FEF2052E03CC" bold="true" box="[199,321,366,391]" pageId="4" pageNumber="537">Diagnosis.</emphasis>
Differs from congeners mainly by the large, straight, lateral sheath processes of the posterior gonopod colpocoxites, coupled with large posterior angiocoxal processes and small mesal sheath processes of the colpocoxite fused mesally into a single cyathiform structure.
</paragraph>
</subSubSection>
<subSubSection id="C31F6550FFC6DC7FFF0CFE46064A01A7" lastPageId="5" lastPageNumber="538" pageId="4" pageNumber="537" type="description">
<paragraph id="8BBA36DBFFC6DC7EFF0CFE4606520071" blockId="4.[151,1436,258,570]" pageId="4" pageNumber="537">
<emphasis id="B971EAC9FFC6DC7EFF0CFE46053403B8" bold="true" box="[201,347,474,499]" pageId="4" pageNumber="537">Description.</emphasis>
Male. Length
<quantity id="4CFD9B3EFFC6DC7EFDC6FE4606EF03B9" box="[515,640,474,498]" metricMagnitude="-2" metricUnit="m" metricValue="1.15" metricValueMax="1.2" metricValueMin="1.1" pageId="4" pageNumber="537" unit="mm" value="11.5" valueMax="12.0" valueMin="11.0">1112 mm</quantity>
, width with paraterga 0.91.0 mm. Coloration in alcohol brown. Legs brown with marbled distal parts. Ocellaria black. Antennae brown. Protruding telopodites of posterior gonopods dark brown. Forehead with light spot.
</paragraph>
<caption id="DF7A6653FFC6DC7EFF52F8EC0092058F" httpUri="https://zenodo.org/record/220594/files/figure.png" pageId="4" pageNumber="537" targetBox="[249,1341,632,1876]" targetPageId="4">
<paragraph id="8BBA36DBFFC6DC7EFF52F8EC0092058F" blockId="4.[151,1436,1904,1988]" pageId="4" pageNumber="537">
<emphasis id="B971EAC9FFC6DC7EFF52F8EC055505CD" bold="true" box="[151,314,1904,1927]" pageId="4" pageNumber="537">FIGURES 89.</emphasis>
<taxonomicName id="4C054D58FFC6DC7EFE84F8ED064605CC" box="[321,553,1905,1927]" class="Diplopoda" family="Diplomaragnidae" genus="Altajosoma" kingdom="Animalia" order="Chordeumatida" pageId="4" pageNumber="537" phylum="Arthropoda" rank="species" species="bukhtarma" status="sp. nov.">
<emphasis id="B971EAC9FFC6DC7EFE84F8ED064605CC" box="[321,553,1905,1927]" italics="true" pageId="4" pageNumber="537">Altajosoma bukhtarma</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24257B2FFC6DC7EFDEAF8ED06EF05CD" box="[559,640,1905,1926]" pageId="4" pageNumber="537" rank="species">
<emphasis id="B971EAC9FFC6DC7EFDEAF8ED06EF05CD" bold="true" box="[559,640,1905,1926]" pageId="4" pageNumber="537">sp. nov.</emphasis>
</taxonomicNameLabel>
, male paratype. 8, gonopods, caudal view; 9, gonopods, front view;
<emphasis id="B971EAC9FFC6DC7EFAFEF8ED013F05CD" bold="true" box="[1339,1360,1905,1926]" pageId="4" pageNumber="537">lp</emphasis>
, lateral sheath process of posterior gonopod colpocoxite;
<emphasis id="B971EAC9FFC6DC7EFD56F80C06C005EE" bold="true" box="[659,687,1936,1957]" pageId="4" pageNumber="537">pp</emphasis>
, posterior process of posterior gonopod angiocoxite;
<emphasis id="B971EAC9FFC6DC7EFB17F80C008205EE" bold="true" box="[1234,1261,1936,1957]" pageId="4" pageNumber="537">ap</emphasis>
, anterior process of posterior gonopod angiocoxite;
<emphasis id="B971EAC9FFC6DC7EFE30F832067B0588" bold="true" box="[501,532,1966,1987]" pageId="4" pageNumber="537">ms</emphasis>
, mesal sheath processes of posterior gonopod colpocoxites. Scale in mm.
</paragraph>
</caption>
<paragraph id="8BBA36DBFFC7DC7FFF0CFF0B077B02BC" blockId="5.[151,1437,151,1005]" pageId="5" pageNumber="538">Body with 32 segments. Head covered with short and relatively long setae. Eye patches subtriangular, each composed of at least 25 ocelli. Collum semi-circular. Body width gradually increasing until somite 7, body parallelsided on somites 824(25), thereafter gradually tapering.</paragraph>
<paragraph id="8BBA36DBFFC7DC7FFF0CFE9900E003CC" blockId="5.[151,1437,151,1005]" pageId="5" pageNumber="538">Paraterga beginning on somite 2, well developed on somites 326, reduced on somite 27, onward missing. Paraterga of pregonopodal somites smaller. Metazonital macrochaetae in a transverse row on somites 3031, like an elongate (to different degrees) triangle on preceding somites. Numerous macrochaetae broken off, remaining ones long, pointed apically; anterolateral macrochaetae shortest, caudolateral ones longest.</paragraph>
<paragraph id="8BBA36DBFFC7DC7FFF02FE0905A00014" blockId="5.[151,1437,151,1005]" pageId="5" pageNumber="538">Legs long and slender. Leg pairs 37 somewhat enlarged, femora swollen. Legs (including leg pairs 10 and 11) with a small group of funnel-shaped tarsal papillae apically near claw, however tarsal papillae gradually missing toward middle part of body. Claws of pregonopodal legs at base with long setoid filament ventrally but without well-developed additional claws dorsally. Claws of postgonopodal legs (including leg pairs 10 and 11) at base with two small additional claws dorsally and long setoid filament ventrally, but small additional claws gradually missing toward middle part of body.</paragraph>
<paragraph id="8BBA36DBFFC7DC7FFF02FDF107C000CF" blockId="5.[151,1437,151,1005]" box="[199,943,620,645]" pageId="5" pageNumber="538">Legs 10 and 11 with coxal glands but without other modifications.</paragraph>
<paragraph id="8BBA36DBFFC7DC7FFF02FD1301220134" blockId="5.[151,1437,151,1005]" pageId="5" pageNumber="538">
Anterior gonopod telopodite 1-segmented, flagelliform (Fis. 5, 7, 8). Telopodite base and distal part of coxosternum attached to adjacent mesal portion of posterior gonopod. Posterior gonopod colpocoxites fused to 2/3 extent, arcuate, curved caudad. Colpocoxite distal part narrowed, apex hook-shaped. Lateral sheath process of colpocoxite (
<emphasis id="B971EAC9FFC7DC7FFEEDFD600550015F" bold="true" box="[296,319,764,788]" pageId="5" pageNumber="538">lp</emphasis>
) straight with prominence in its middle part, pointed apically. Mesal sheath processes of colpocoxite not large, fused mesally into single cyathiform structure (
<emphasis id="B971EAC9FFC7DC7FFC86FCBC070A0173" bold="true" box="[835,869,800,824]" pageId="5" pageNumber="538">ms</emphasis>
). Angiocoxite with a globule in posterior view. Posterior angiocoxal process (
<emphasis id="B971EAC9FFC7DC7FFE22FCD806690117" bold="true" box="[487,518,836,860]" pageId="5" pageNumber="538">pp</emphasis>
) large, curved forward with beak-shaped pointed lateral margin and teeth apically. Angiocoxites with depressions and ridges in anterior view, each supplied with small process (
<emphasis id="B971EAC9FFC7DC7FFB6CFCF400A901CB" bold="true" box="[1193,1222,872,896]" pageId="5" pageNumber="538">ap</emphasis>
) (
<figureCitation id="133E2A5EFFC7DC7FFB1AFCFB014C0134" box="[1247,1315,871,895]" captionStart="FIGURES 5 7" captionStartId="3.[151,264,1898,1921]" captionTargetBox="[335,1254,195,1841]" captionTargetId="figure@3.[322,1268,193,1843]" captionTargetPageId="3" captionText="FIGURES 5 7. Altajosoma bukhtarma sp. nov., male paratype. 5, gonopods, caudal view; 6, gonopods, front view; 7, basal parts of telopodites of anterior gonopods; lp, lateral sheath process of posterior gonopod colpocoxite; pp, posterior process of posterior gonopod angiocoxite; ms, mesal sheath processes of posterior gonopod colpocoxites; ap, anterior process of posterior gonopod angiocoxite." httpUri="https://zenodo.org/record/220593/files/figure.png" pageId="5" pageNumber="538">Fig. 6</figureCitation>
,
<figureCitation id="133E2A5EFFC7DC7FFAEAFCFB01500134" box="[1327,1343,871,895]" captionStart="FIGURES 8 9" captionStartId="4.[151,264,1904,1927]" captionTargetBox="[249,1341,632,1876]" captionTargetId="figure@4.[241,1346,623,1883]" captionTargetPageId="4" captionText="FIGURES 8 9. Altajosoma bukhtarma sp. nov., male paratype. 8, gonopods, caudal view; 9, gonopods, front view; lp, lateral sheath process of posterior gonopod colpocoxite; pp, posterior process of posterior gonopod angiocoxite; ap, anterior process of posterior gonopod angiocoxite; ms, mesal sheath processes of posterior gonopod colpocoxites. Scale in mm." httpUri="https://zenodo.org/record/220594/files/figure.png" pageId="5" pageNumber="538">9</figureCitation>
).
</paragraph>
<paragraph id="8BBA36DBFFC7DC7FFF02FC1005FF01EF" blockId="5.[151,1437,151,1005]" box="[199,400,908,932]" pageId="5" pageNumber="538">Female unknown.</paragraph>
<paragraph id="8BBA36DBFFC7DC7FFF02FC2C064A01A7" blockId="5.[151,1437,151,1005]" pageId="5" pageNumber="538">
<emphasis id="B971EAC9FFC7DC7FFF02FC2C057B0183" bold="true" box="[199,276,944,968]" pageId="5" pageNumber="538">Name.</emphasis>
The specific epithet refers to the
<typeStatus id="54BE8879FFC7DC7FFD4DFC2C06D70183" box="[648,696,944,968]" pageId="5" pageNumber="538">type</typeStatus>
locality (
<taxonomicName id="4C054D58FFC7DC7FFCE4FC3307F0018C" box="[801,927,943,967]" class="Diplopoda" family="Diplomaragnidae" genus="Altajosoma" kingdom="Animalia" order="Chordeumatida" pageId="5" pageNumber="543" phylum="Arthropoda" rank="species" species="bukhtarma" status="sp. nov.">Bukhtarma</taxonomicName>
is the name of the river, which flows near the
<typeStatus id="54BE8879FFC7DC7FFF52FC4904A801A6" box="[151,199,981,1005]" pageId="5" pageNumber="538">type</typeStatus>
locality), a noun in apposition.
</paragraph>
</subSubSection>
</treatment>
</document>