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<document id="994B97B7282AA5E95FF7377E8DAEC7A2" ID-DOI="10.11646/zootaxa.5492.3.7" ID-ISSN="1175-5326" ID-Zenodo-Dep="13234967" ID-ZooBank="47F51BFA-AC3B-42BB-BBBC-E5ACE29C3F2F" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722933112560" checkinUser="plazi" docAuthor="Naik, Santhosh &amp; Shashank, P. R." docDate="2024" docId="03A987C2FFD0DA49FF23458AFAC73C1C" docLanguage="en" docName="zootaxa.5492.3.7.pdf" docOrigin="Zootaxa 5492 (3)" docSource="http://dx.doi.org/10.11646/zootaxa.5492.3.7" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Terthreutis subrectangulus Naik &amp; Shashank, 2024, sp. nov." docType="treatment" docVersion="2" lastPageNumber="419" masterDocId="FF90FFBAFFD6DA43FFB44008FFC73A15" masterDocTitle="Discovery of two new species and three new records of the tribe Archipini (Lepidoptera: Tortricidae) from India" masterLastPageNumber="420" masterPageNumber="409" pageNumber="415" updateTime="1722933579633" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="DA3DE3E498062724BDC77CD929A6EC08">Discovery of two new species and three new records of the tribe Archipini (Lepidoptera: Tortricidae) from India</mods:title>
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<mods:name id="F0B5A21774CDE1F5F7108C32D7D3C716" type="personal">
<mods:role id="CD305767F604518363D8A1B1DF24F78D">
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<mods:namePart id="5F6048B6EFCC3CBD01F68162DE130ADD">Naik, Santhosh</mods:namePart>
<mods:affiliation id="E502280D832F522DC481C5713146020A">National Pusa Collection, Division of Entomology, ICAR- Indian Agricultural Research Institute, New Delhi, 110012, India</mods:affiliation>
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<mods:namePart id="82B9FDBC24B037C424EB3EBCE5A9E0A4">Shashank, P. R.</mods:namePart>
<mods:nameIdentifier id="0EEC9F94F96918BB4F7DD2DE2820DB6F" type="ORCID">0000-0002-8177-6091</mods:nameIdentifier>
<mods:nameIdentifier id="F5A290DF50CBDD616FE91B5FA38ACD33" type="email">spathour@gmail.com</mods:nameIdentifier>
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<mods:title id="E20079BED15EB4C78772BB768B873B90">Zootaxa</mods:title>
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<mods:date id="8A1E4E03502F8881C59C7583644C48C7">2024</mods:date>
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<mods:number id="92FFB8B9D5808461969ADB00783E3A9E">2024-08-06</mods:number>
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<mods:number id="BA881CF83A3007240CFF2124179BA32E">5492</mods:number>
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<mods:number id="C55A33147859E30DF98E13FFFD34E959">3</mods:number>
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<mods:identifier id="BFFF857BC45477A76A64C657D6AFD6BB" type="DOI">10.11646/zootaxa.5492.3.7</mods:identifier>
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<treatment id="03A987C2FFD0DA49FF23458AFAC73C1C" LSID="urn:lsid:plazi:treatment:03A987C2FFD0DA49FF23458AFAC73C1C" httpUri="http://treatment.plazi.org/id/03A987C2FFD0DA49FF23458AFAC73C1C" lastPageId="10" lastPageNumber="419" pageId="6" pageNumber="415">
<subSubSection id="C31A655FFFD0DA45FF23458AFC983F89" box="[151,863,1410,1437]" pageId="6" pageNumber="415" type="nomenclature">
<paragraph id="8BBF36D4FFD0DA45FF23458AFC983F89" blockId="6.[151,863,1410,1473]" box="[151,863,1410,1437]" pageId="6" pageNumber="415">
<heading id="D0F781B8FFD0DA45FF23458AFC983F89" bold="true" box="[151,863,1410,1437]" fontSize="11" level="1" pageId="6" pageNumber="415" reason="1">
<emphasis id="B974EAC6FFD0DA45FF23458AFC983F89" bold="true" box="[151,863,1410,1437]" pageId="6" pageNumber="415">
<taxonomicName id="4C004D57FFD0DA45FF23458AFE1A3F88" authorityName="Naik &amp; Shashank" authorityYear="2024" box="[151,477,1410,1437]" class="Insecta" family="Tortricidae" genus="Terthreutis" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="415" phylum="Arthropoda" rank="species" species="subrectangulus" status="sp. nov.">
<emphasis id="B974EAC6FFD0DA45FF23458AFE1A3F88" bold="true" box="[151,477,1410,1437]" italics="true" pageId="6" pageNumber="415">Terthreutis subrectangulus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24757BDFFD0DA45FE50458BFD853F88" box="[484,578,1411,1437]" pageId="6" pageNumber="415" rank="species">sp. nov.</taxonomicNameLabel>
Shashank &amp; Santhosh
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C31A655FFFD0DA45FF2345AFFEFA3C37" pageId="6" pageNumber="415" type="description">
<paragraph id="8BBF36D4FFD0DA45FF2345AFFE713FD4" blockId="6.[151,863,1410,1473]" box="[151,438,1446,1473]" pageId="6" pageNumber="415">
(
<figureCitation id="133B2A51FFD0DA45FF2B45AFFF363FD4" box="[159,241,1447,1473]" captionStart="FIGURE 1" captionStartId="2.[151,250,1472,1496]" captionTargetBox="[162,1425,284,1438]" captionTargetId="figure-62@2.[151,1436,273,1449]" captionTargetPageId="2" captionText="FIGURE 1. Dorsal habitus illustrations. A. Choristoneura ribhoiensis sp. nov. (holotype, male); B. Clepsis laetornata (male); C. Clepsis laetornata (female); D. Diplocalyptis ferruginimixta (male); E. Diplocalyptis operosa (male); F. Terthreutis subrectangulus sp. nov. (holotype, male). Scale bar = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.13234971" httpUri="https://zenodo.org/record/13234971/files/figure.png" pageId="6" pageNumber="415">Fig. 1F</figureCitation>
,
<figureCitation id="133B2A51FFD0DA45FF4945AFFED83FD4" box="[253,287,1447,1473]" captionStart="FIGURE 2" captionStartId="3.[151,250,1692,1716]" captionTargetBox="[151,1432,818,1662]" captionTargetId="figure-241@3.[151,1437,813,1669]" captionTargetPageId="3" captionText="FIGURE 2. Labial palpi. A. Choristoneura ribhoiensis sp. nov. (holotype, male); B. Clepsis laetornata (male); C. Diplocalyptis ferruginimixta (male); D. Diplocalyptis operosa (male); E. Terthreutis subrectangulus sp. nov. (holotype, male). Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13234975" httpUri="https://zenodo.org/record/13234975/files/figure.png" pageId="6" pageNumber="415">2E</figureCitation>
,
<figureCitation id="133B2A51FFD0DA45FE9E45AFFE9A3FD4" box="[298,349,1447,1473]" captionStart="FIGURE 3" captionStartId="7.[151,250,1915,1939]" captionTargetBox="[162,1425,192,1880]" captionTargetId="figure-17@7.[151,1436,181,1891]" captionTargetPageId="7" captionText="FIGURE 3. Wing venation; Choristoneura ribhoiensis sp. nov. (holotype, male) (A. Forewing, B. Hindwing); Clepsis laetornata (C. Forewing, D. Hindwing)); Diplocalyptis ferruginimixta (E. Forewing, F. Hindwing); Diplocalyptis operosa (G. Forewing, H. Hindwing); Terthreutis subrectangulus sp. nov. (holotype, male) (I. Forewing, J. Hindwing). Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13234977" httpUri="https://zenodo.org/record/13234977/files/figure.png" pageId="6" pageNumber="415">3I–J</figureCitation>
,
<figureCitation id="133B2A51FFD0DA45FEDD45AEFE693FD4" box="[361,430,1446,1473]" captionStart="FIGURE 5" captionStartId="9.[151,250,1943,1967]" captionTargetBox="[206,1381,191,1909]" captionTargetId="figure-17@9.[196,1391,181,1919]" captionTargetPageId="9" captionText="FIGURE 5. Diplocalyptis ferruginimixta (A. Male genitalia, B. Uncus, C. Phallus); Diplocalyptis operosa (D. Male genitalia, E. Uncus, F. Phallus); Terthreutis subrectangulus sp. nov. (holotype, male) (G. Male genitalia, H. Uncus &amp; Socii). Scale bar A, D, G= 0.5 mm; B, C, E, F, H= 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13234981" httpUri="https://zenodo.org/record/13234981/files/figure.png" pageId="6" pageNumber="415">5G–H</figureCitation>
)
</paragraph>
<paragraph id="8BBF36D4FFD0DA45FF2345E4FDF93C16" blockId="6.[151,574,1515,1570]" box="[151,574,1515,1540]" pageId="6" pageNumber="415">
Type locality:
<collectingCountry id="F3177644FFD0DA45FE9D45E3FE9A3C16" box="[297,349,1515,1539]" name="India" pageId="6" pageNumber="415">India</collectingCountry>
(Umiam,
<collectingRegion id="49C4F836FFD0DA45FE7145E3FDF03C16" box="[453,567,1515,1539]" country="India" name="Meghalaya" pageId="6" pageNumber="415">Meghalaya</collectingRegion>
)
</paragraph>
<paragraph id="8BBF36D4FFD0DA45FF234602FEFA3C37" blockId="6.[151,574,1515,1570]" box="[151,317,1546,1570]" pageId="6" pageNumber="415">
<materialsCitation id="3B683C89FFD0DA45FF234602FEFA3C37" box="[151,317,1546,1570]" collectionCode="INPC" pageId="6" pageNumber="415" specimenCount="1" typeStatus="holotype">
<typeStatus id="54BB8876FFD0DA45FF234602FF3E3C37" box="[151,249,1546,1570]" pageId="6" pageNumber="415" type="holotype">Holotype</typeStatus>
:
<collectionCode id="ED11AE11FFD0DA45FEB74602FEFA3C37" box="[259,317,1546,1570]" country="India" httpUri="http://grbio.org/cool/qfpa-5ik3" name="National Pusa Collections" pageId="6" pageNumber="415">INPC</collectionCode>
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C31A655FFFD0DA45FF234658FACD3DE3" pageId="6" pageNumber="415" type="diagnosis">
<paragraph id="8BBF36D4FFD0DA45FF234658FACD3DE3" blockId="6.[151,1437,1616,2039]" pageId="6" pageNumber="415">
<emphasis id="B974EAC6FFD0DA45FF234658FED53C7F" bold="true" box="[151,274,1616,1642]" pageId="6" pageNumber="415">Diagnosis.</emphasis>
<taxonomicName id="4C004D57FFD0DA45FEAE4658FD813C7F" authorityName="Naik &amp; Shashank" authorityYear="2024" box="[282,582,1616,1642]" class="Insecta" family="Tortricidae" genus="Terthreutis" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="415" phylum="Arthropoda" rank="species" species="subrectangulus" status="sp. nov.">
<emphasis id="B974EAC6FFD0DA45FEAE4658FD813C7F" box="[282,582,1616,1642]" italics="true" pageId="6" pageNumber="415">Terthreutis subrectangulus</emphasis>
</taxonomicName>
<emphasis id="B974EAC6FFD0DA45FDFB4658FD6C3C7F" bold="true" box="[591,683,1616,1642]" pageId="6" pageNumber="415">
<taxonomicNameLabel id="A24757BDFFD0DA45FDFB4658FD6C3C7F" box="[591,683,1616,1642]" pageId="6" pageNumber="415" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is closely related to
<taxonomicName id="4C004D57FFD0DA45FC284658FABA3C7F" authority="Razowski, 2008" authorityName="Razowski" authorityYear="2008" box="[924,1405,1616,1642]" class="Insecta" family="Tortricidae" genus="Terthreutis" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="415" phylum="Arthropoda" rank="species" species="chiangmaiana">
<emphasis id="B974EAC6FFD0DA45FC284658FB793C7F" box="[924,1214,1616,1642]" italics="true" pageId="6" pageNumber="415">Terthreutis chiangmaiana</emphasis>
<bibRefCitation id="EF914B25FFD0DA45FB734658FABA3C7F" author="Razowski, J." box="[1223,1405,1616,1642]" pageId="6" pageNumber="415" pagination="283 - 299" refId="ref7112" refString="Razowski, J. (2008) On two South Asian genera Ceramea Diakonoff and Terthreutis Meyrick (Lepidoptera: Tortricidae). Polish Journal of Entomology, 4 (77), 283 - 299." type="journal article" year="2008">
Razowski,
<quantity id="4CF89B31FFD0DA45FAF14658FABA3C7F" box="[1349,1405,1616,1642]" metricMagnitude="1" metricUnit="m" metricValue="5.10032" pageId="6" pageNumber="415" unit="in" value="2008.0">2008</quantity>
</bibRefCitation>
</taxonomicName>
in adult morphological resemblance but differs in male genitalia characters.
<taxonomicName id="4C004D57FFD0DA45FC6D467CFAC33C9B" authorityName="Naik &amp; Shashank" authorityYear="2024" box="[985,1284,1652,1678]" class="Insecta" family="Tortricidae" genus="Terthreutis" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="415" phylum="Arthropoda" rank="species" species="subrectangulus">
<emphasis id="B974EAC6FFD0DA45FC6D467CFAC33C9B" box="[985,1284,1652,1678]" italics="true" pageId="6" pageNumber="415">Terthreutis subrectangulus</emphasis>
</taxonomicName>
has transtilla incomplete, valva subrectangular, and smooth phallus without any serration whereas
<taxonomicName id="4C004D57FFD0DA45FBF84691FAC03CA7" authorityName="Razowski" authorityYear="2008" box="[1100,1287,1688,1714]" class="Insecta" family="Tortricidae" genus="Terthreutis" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="415" phylum="Arthropoda" rank="species" species="chiangmaiana">
<emphasis id="B974EAC6FFD0DA45FBF84691FAC03CA7" box="[1100,1287,1688,1714]" italics="true" pageId="6" pageNumber="415">T. chiangmaiana</emphasis>
</taxonomicName>
has transtilla complete and continuous, valva rounded, and serrated phallus with the presence of some denticles in the distal portion. Also, it is found that the transtilla is non-continuous or incomplete with two sclerotized lobes and this
<typeStatus id="54BB8876FFD0DA45FF23470DFF003D0A" box="[151,199,1797,1823]" pageId="6" pageNumber="415">type</typeStatus>
of transtilla is present in some species of
<taxonomicName id="4C004D57FFD0DA45FD1E470CFCE43D0B" authorityName="Meyrick" authorityYear="1918" box="[682,803,1796,1822]" class="Insecta" family="Tortricidae" genus="Terthreutis" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="415" phylum="Arthropoda" rank="genus">
<emphasis id="B974EAC6FFD0DA45FD1E470CFCE43D0B" box="[682,803,1796,1822]" italics="true" pageId="6" pageNumber="415">Terthreutis</emphasis>
</taxonomicName>
such as
<taxonomicName id="4C004D57FFD0DA45FC33470DFBDB3D0B" authorityName="Bai" authorityYear="1993" box="[903,1052,1796,1822]" class="Insecta" family="Tortricidae" genus="Terthreutis" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="415" phylum="Arthropoda" rank="species" species="bipunctata">
<emphasis id="B974EAC6FFD0DA45FC33470DFBDB3D0B" box="[903,1052,1796,1822]" italics="true" pageId="6" pageNumber="415">T. bipunctata</emphasis>
</taxonomicName>
,
<taxonomicName id="4C004D57FFD0DA45FB9F470DFB033D0B" authorityName="Bai" authorityYear="1993" box="[1067,1220,1796,1822]" class="Insecta" family="Tortricidae" genus="Terthreutis" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="415" phylum="Arthropoda" rank="species" species="orbicularis">
<emphasis id="B974EAC6FFD0DA45FB9F470DFB033D0B" box="[1067,1220,1796,1822]" italics="true" pageId="6" pageNumber="415">T. orbicularis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C004D57FFD0DA45FB60470DFABF3D0B" baseAuthorityName="Meyrick" baseAuthorityYear="1938" box="[1236,1400,1796,1822]" class="Insecta" family="Tortricidae" genus="Terthreutis" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="415" phylum="Arthropoda" rank="species" species="xanthocycla">
<emphasis id="B974EAC6FFD0DA45FB60470DFABF3D0B" box="[1236,1400,1796,1822]" italics="true" pageId="6" pageNumber="415">T. xanthocycla</emphasis>
</taxonomicName>
,
<taxonomicName id="4C004D57FFD0DA45FA3C470DFEFF3D57" class="Insecta" family="Tortricidae" genus="Terthreutis" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="415" phylum="Arthropoda" rank="species" species="sphaerocosma">
<emphasis id="B974EAC6FFD0DA45FA3C470DFEFF3D57" italics="true" pageId="6" pageNumber="415">T. sphaerocosma</emphasis>
</taxonomicName>
and
<taxonomicName id="4C004D57FFD0DA45FEDB4721FE0A3D57" authorityName="Bai" authorityYear="1993" box="[367,461,1833,1858]" class="Insecta" family="Tortricidae" genus="Terthreutis" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="415" phylum="Arthropoda" rank="species" species="series">
<emphasis id="B974EAC6FFD0DA45FEDB4721FE0A3D57" box="[367,461,1833,1858]" italics="true" pageId="6" pageNumber="415">T. series</emphasis>
</taxonomicName>
. However, there are thirteen species described worldwide (
<bibRefCitation id="EF914B25FFD0DA45FBDC4720FA503D57" author="Buchsbaum, U. &amp; Chen, M. Y." box="[1128,1431,1832,1858]" pageId="6" pageNumber="415" pagination="349 - 356" refId="ref6417" refString="Buchsbaum, U. &amp; Chen, M. Y. (2013) A new Terthreutis Meyrick, 1918 species from Taiwan (Lepidoptera, Tortricidae) Contribution to the moths of Taiwan 5. Entomofauna, 26, 349 - 356." type="journal article" year="2013">Buchsbaum &amp; Chen, 2013</bibRefCitation>
;
<bibRefCitation id="EF914B25FFD0DA45FF234744FE473D73" author="Gilligan, T. M. &amp; Baixeras, J. &amp; Brown, J. W." box="[151,384,1868,1895]" pageId="6" pageNumber="415" refId="ref6544" refString="Gilligan, T. M., Baixeras, J. &amp; Brown, J. W. (2018) T @ RTS: Online World Catalogue of the Tortricidae. Version. 4.0. Available from: http: // www. tortricid. net / Catalogue. asp (accessed 12 April 2023)" type="url" year="2018">
Gilligan
<emphasis id="B974EAC6FFD0DA45FF4F4745FEF23D73" box="[251,309,1868,1894]" italics="true" pageId="6" pageNumber="415">et al.</emphasis>
, 2018
</bibRefCitation>
), out of which three species have been described from
<collectingCountry id="F3177644FFD0DA45FC464744FBF73D73" box="[1010,1072,1868,1894]" name="India" pageId="6" pageNumber="415">India</collectingCountry>
:
<emphasis id="B974EAC6FFD0DA45FB8F4745FA473D73" box="[1083,1408,1868,1894]" italics="true" pageId="6" pageNumber="415">
<taxonomicName id="4C004D57FFD0DA45FB8F4745FB7A3D73" baseAuthorityName="Butler" baseAuthorityYear="1886" box="[1083,1213,1869,1894]" class="Insecta" family="Tortricidae" genus="Terthreutis" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="415" phylum="Arthropoda" rank="species" species="argentea">T. argentea</taxonomicName>
,
<taxonomicName id="4C004D57FFD0DA45FB7E4745FA893D73" authorityName="Meyrick" authorityYear="1928" box="[1226,1358,1868,1894]" class="Insecta" family="Tortricidae" genus="Terthreutis" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="415" phylum="Arthropoda" rank="species" species="bulligera">T. bulligera</taxonomicName>
and
</emphasis>
<taxonomicName id="4C004D57FFD0DA45FA3C4745FD863D9F" authority="(Pathania et al., 2020)" baseAuthorityName="Pathania" baseAuthorityYear="2020" class="Insecta" family="Tortricidae" genus="Terthreutis" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="415" phylum="Arthropoda" rank="species" species="sphaerocosma">
<emphasis id="B974EAC6FFD0DA45FA3C4745FEFF3D9F" italics="true" pageId="6" pageNumber="415">T. sphaerocosma</emphasis>
(
<bibRefCitation id="EF914B25FFD0DA45FEFC4778FDFE3D9F" author="Pathania, P. C. &amp; Das, A. &amp; Brown, J. &amp; Chandra, K." box="[328,569,1904,1931]" pageId="6" pageNumber="415" pagination="1 - 95" refId="ref7020" refString="Pathania, P. C., Das, A., Brown, J. &amp; Chandra, K. (2020) Catalogue of Tortricidae Latreille, 1802 (Lepidoptera: Tortricoidea) of India. Zootaxa, 4757 (1), 1 - 95. https: // doi. org / 10.11646 / zootaxa. 4757.1.1" type="journal article" year="2020">
Pathania
<emphasis id="B974EAC6FFD0DA45FE074779FE2A3D9F" box="[435,493,1904,1930]" italics="true" pageId="6" pageNumber="415">et al.</emphasis>
, 2020
</bibRefCitation>
)
</taxonomicName>
. However, most of the species of
<taxonomicName id="4C004D57FFD0DA45FC724778FBF83D9F" authorityName="Meyrick" authorityYear="1918" box="[966,1087,1904,1930]" class="Insecta" family="Tortricidae" genus="Terthreutis" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="415" phylum="Arthropoda" rank="genus">
<emphasis id="B974EAC6FFD0DA45FC724778FBF83D9F" box="[966,1087,1904,1930]" italics="true" pageId="6" pageNumber="415">Terthreutis</emphasis>
</taxonomicName>
are different but present some elements similar to the new species for example an incomplete transtilla, elongated and well sclerotised uncus, gnathal arms long and wide and well sclerotised sacculus about as long as 1/2 of the length of valva. This study adds one more species to the world fauna of
<taxonomicName id="4C004D57FFD0DA45FDE547D4FD0D3DE3" authorityName="Meyrick" authorityYear="1918" box="[593,714,2012,2038]" class="Insecta" family="Tortricidae" genus="Terthreutis" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="415" phylum="Arthropoda" rank="genus">
<emphasis id="B974EAC6FFD0DA45FDE547D4FD0D3DE3" box="[593,714,2012,2038]" italics="true" pageId="6" pageNumber="415">Terthreutis</emphasis>
</taxonomicName>
and increases the number of Indian species to four.
</paragraph>
</subSubSection>
<caption id="DF7F665CFFD1DA44FF234773FAEF3DCF" ID-DOI="http://doi.org/10.5281/zenodo.13234977" ID-Zenodo-Dep="13234977" httpUri="https://zenodo.org/record/13234977/files/figure.png" pageId="7" pageNumber="416" startId="7.[151,250,1915,1939]" targetBox="[162,1425,192,1880]" targetPageId="7" targetType="figure">
<paragraph id="8BBF36D4FFD1DA44FF234773FAEF3DCF" blockId="7.[151,1437,1915,2011]" pageId="7" pageNumber="416">
<emphasis id="B974EAC6FFD1DA44FF234773FED63D86" bold="true" box="[151,273,1915,1939]" pageId="7" pageNumber="416">FIGURE 3.</emphasis>
Wing venation;
<taxonomicName id="4C004D57FFD1DA44FE004773FD7A3D86" authorityName="Naik &amp; Shashank" authorityYear="2024" box="[436,701,1915,1939]" class="Insecta" family="Tortricidae" genus="Choristoneura" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="416" phylum="Arthropoda" rank="species" species="ribhoiensis" status="sp. nov.">
<emphasis id="B974EAC6FFD1DA44FE004773FD7A3D86" box="[436,701,1915,1939]" italics="true" pageId="7" pageNumber="416">Choristoneura ribhoiensis</emphasis>
</taxonomicName>
<emphasis id="B974EAC6FFD1DA44FD754773FCC83D86" bold="true" box="[705,783,1915,1939]" pageId="7" pageNumber="416">
<taxonomicNameLabel id="A24757BDFFD1DA44FD754773FCC83D86" box="[705,783,1915,1939]" pageId="7" pageNumber="416" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(holotype, male) (
<emphasis id="B974EAC6FFD1DA44FC724773FC183D86" bold="true" box="[966,991,1915,1939]" pageId="7" pageNumber="416">A.</emphasis>
Forewing,
<emphasis id="B974EAC6FFD1DA44FBFB4773FBA13D86" bold="true" box="[1103,1126,1915,1939]" pageId="7" pageNumber="416">B.</emphasis>
Hindwing);
<taxonomicName id="4C004D57FFD1DA44FB504773FA5A3D86" baseAuthorityName="Wang Li &amp; Wang" baseAuthorityYear="2003" box="[1252,1437,1915,1939]" class="Insecta" family="Tortricidae" genus="Clepsis" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="416" phylum="Arthropoda" rank="species" species="laetornata">
<emphasis id="B974EAC6FFD1DA44FB504773FA5A3D86" box="[1252,1437,1915,1939]" italics="true" pageId="7" pageNumber="416">Clepsis laetornata</emphasis>
</taxonomicName>
(
<emphasis id="B974EAC6FFD1DA44FF144797FF7F3DA2" bold="true" box="[160,184,1951,1975]" pageId="7" pageNumber="416">C.</emphasis>
Forewing,
<emphasis id="B974EAC6FFD1DA44FE994797FE813DA2" bold="true" box="[301,326,1951,1975]" pageId="7" pageNumber="416">D.</emphasis>
Hindwing));
<taxonomicName id="4C004D57FFD1DA44FE654797FD353DA2" authorityName="Razowski" authorityYear="2009" box="[465,754,1951,1975]" class="Insecta" family="Tortricidae" genus="Diplocalyptis" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="416" phylum="Arthropoda" rank="species" species="ferruginimixta">
<emphasis id="B974EAC6FFD1DA44FE654797FD353DA2" box="[465,754,1951,1975]" italics="true" pageId="7" pageNumber="416">Diplocalyptis ferruginimixta</emphasis>
</taxonomicName>
(
<emphasis id="B974EAC6FFD1DA44FCB64797FCDE3DA2" bold="true" box="[770,793,1951,1975]" pageId="7" pageNumber="416">E.</emphasis>
Forewing,
<emphasis id="B974EAC6FFD1DA44FC3A4797FC663DA2" bold="true" box="[910,929,1951,1975]" pageId="7" pageNumber="416">F.</emphasis>
Hindwing);
<taxonomicName id="4C004D57FFD1DA44FB914797FAC23DA2" baseAuthorityName="Meyrick" baseAuthorityYear="1908" box="[1061,1285,1951,1975]" class="Insecta" family="Tortricidae" genus="Diplocalyptis" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="416" phylum="Arthropoda" rank="species" species="operosa">
<emphasis id="B974EAC6FFD1DA44FB914797FAC23DA2" box="[1061,1285,1951,1975]" italics="true" pageId="7" pageNumber="416">Diplocalyptis operosa</emphasis>
</taxonomicName>
(
<emphasis id="B974EAC6FFD1DA44FAA04797FAE93DA2" bold="true" box="[1300,1326,1951,1975]" pageId="7" pageNumber="416">G.</emphasis>
Forewing,
<emphasis id="B974EAC6FFD1DA44FF2347CBFF763DCE" bold="true" box="[151,177,1987,2011]" pageId="7" pageNumber="416">H.</emphasis>
Hindwing);
<taxonomicName id="4C004D57FFD1DA44FE8747CBFD873DCE" authorityName="Naik &amp; Shashank" authorityYear="2024" box="[307,576,1987,2011]" class="Insecta" family="Tortricidae" genus="Terthreutis" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="416" phylum="Arthropoda" rank="species" species="subrectangulus" status="sp. nov.">
<emphasis id="B974EAC6FFD1DA44FE8747CBFD873DCE" box="[307,576,1987,2011]" italics="true" pageId="7" pageNumber="416">Terthreutis subrectangulus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24757BDFFD1DA44FDF347CBFD5E3DCF" box="[583,665,1987,2011]" pageId="7" pageNumber="416" rank="species">
<emphasis id="B974EAC6FFD1DA44FDF347CBFD553DCE" bold="true" box="[583,658,1987,2011]" pageId="7" pageNumber="416">sp. nov</emphasis>
.
</taxonomicNameLabel>
(holotype, male) (
<emphasis id="B974EAC6FFD1DA44FCE147CBFCA23DCE" bold="true" box="[853,869,1987,2011]" pageId="7" pageNumber="416">I.</emphasis>
Forewing,
<emphasis id="B974EAC6FFD1DA44FC6E47CBFC2A3DCE" bold="true" box="[986,1005,1987,2011]" pageId="7" pageNumber="416">J.</emphasis>
Hindwing). Scale bar = 1 mm.
</paragraph>
</caption>
<caption id="DF7F665CFFDEDA4BFF23479FFBB33DC6" ID-DOI="http://doi.org/10.5281/zenodo.13234979" ID-Zenodo-Dep="13234979" httpUri="https://zenodo.org/record/13234979/files/figure.png" pageId="8" pageNumber="417" startId="8.[151,250,1943,1967]" targetBox="[166,1420,192,1908]" targetPageId="8" targetType="figure">
<paragraph id="8BBF36D4FFDEDA4BFF23479FFBB33DC6" blockId="8.[151,1437,1943,2003]" pageId="8" pageNumber="417">
<emphasis id="B974EAC6FFDEDA4BFF23479FFED03DBA" bold="true" box="[151,279,1943,1967]" pageId="8" pageNumber="417">FIGURE 4.</emphasis>
<taxonomicName id="4C004D57FFDEDA4BFE96479FFDF63DBA" authorityName="Naik &amp; Shashank" authorityYear="2024" box="[290,561,1943,1967]" class="Insecta" family="Tortricidae" genus="Choristoneura" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="417" phylum="Arthropoda" rank="species" species="ribhoiensis" status="sp. nov.">
<emphasis id="B974EAC6FFDEDA4BFE96479FFDF63DBA" box="[290,561,1943,1967]" italics="true" pageId="8" pageNumber="417">Choristoneura ribhoiensis</emphasis>
</taxonomicName>
<emphasis id="B974EAC6FFDEDA4BFD8F479FFD573DBA" bold="true" box="[571,656,1943,1967]" pageId="8" pageNumber="417">
<taxonomicNameLabel id="A24757BDFFDEDA4BFD8F479FFD573DBA" box="[571,656,1943,1967]" pageId="8" pageNumber="417" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<emphasis id="B974EAC6FFDEDA4BFD17479FFD7B3DBA" bold="true" box="[675,700,1943,1967]" pageId="8" pageNumber="417">A.</emphasis>
Male genitalia,
<emphasis id="B974EAC6FFDEDA4BFCD9479FFC433DBA" bold="true" box="[877,900,1943,1967]" pageId="8" pageNumber="417">B.</emphasis>
Uncus,
<emphasis id="B974EAC6FFDEDA4BFC54479FFC3F3DBA" bold="true" box="[992,1016,1943,1967]" pageId="8" pageNumber="417">C.</emphasis>
Transtilla process,
<emphasis id="B974EAC6FFDEDA4BFB7E479FFB243DBA" bold="true" box="[1226,1251,1943,1967]" pageId="8" pageNumber="417">D.</emphasis>
Phallus);
<taxonomicName id="4C004D57FFDEDA4BFAE6479FFEC63DC6" baseAuthorityName="Wang Li &amp; Wang" baseAuthorityYear="2003" class="Insecta" family="Tortricidae" genus="Clepsis" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="417" phylum="Arthropoda" rank="species" species="laetornata">
<emphasis id="B974EAC6FFDEDA4BFAE6479FFEC63DC6" italics="true" pageId="8" pageNumber="417">Clepsis laetornata</emphasis>
</taxonomicName>
(
<emphasis id="B974EAC6FFDEDA4BFEA447B3FEE03DC6" bold="true" box="[272,295,1979,2003]" pageId="8" pageNumber="417">E.</emphasis>
Male genitalia,
<emphasis id="B974EAC6FFDEDA4BFE7F47B3FE193DC6" bold="true" box="[459,478,1979,2003]" pageId="8" pageNumber="417">F.</emphasis>
Female genitalia). Scale bar A, E, F= 0.5 mm; B, C, D= 0.1 mm.
</paragraph>
</caption>
<caption id="DF7F665CFFDFDA4AFF23479FFDE43DE2" ID-DOI="http://doi.org/10.5281/zenodo.13234981" ID-Zenodo-Dep="13234981" httpUri="https://zenodo.org/record/13234981/files/figure.png" pageId="9" pageNumber="418" startId="9.[151,250,1943,1967]" targetBox="[206,1381,191,1909]" targetPageId="9" targetType="figure">
<paragraph id="8BBF36D4FFDFDA4AFF23479FFDE43DE2" blockId="9.[151,1437,1943,2040]" pageId="9" pageNumber="418">
<emphasis id="B974EAC6FFDFDA4AFF23479FFED43DA5" bold="true" box="[151,275,1943,1968]" pageId="9" pageNumber="418">FIGURE 5.</emphasis>
<taxonomicName id="4C004D57FFDFDA4AFEAE479FFDFC3DA5" authorityName="Razowski" authorityYear="2009" box="[282,571,1943,1968]" class="Insecta" family="Tortricidae" genus="Diplocalyptis" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="418" phylum="Arthropoda" rank="species" species="ferruginimixta">
<emphasis id="B974EAC6FFDFDA4AFEAE479FFDFC3DA5" box="[282,571,1943,1968]" italics="true" pageId="9" pageNumber="418">Diplocalyptis ferruginimixta</emphasis>
</taxonomicName>
(
<emphasis id="B974EAC6FFDFDA4AFDFF479FFDA33DBA" bold="true" box="[587,612,1943,1967]" pageId="9" pageNumber="418">A.</emphasis>
Male genitalia,
<emphasis id="B974EAC6FFDFDA4AFCBD4790FCE63DA5" bold="true" box="[777,801,1944,1968]" pageId="9" pageNumber="418">B.</emphasis>
Uncus,
<emphasis id="B974EAC6FFDFDA4AFCC1479FFC493DBA" bold="true" box="[885,910,1943,1967]" pageId="9" pageNumber="418">C.</emphasis>
Phallus);
<taxonomicName id="4C004D57FFDFDA4AFC41479FFB113DBA" baseAuthorityName="Meyrick" baseAuthorityYear="1908" box="[1013,1238,1943,1967]" class="Insecta" family="Tortricidae" genus="Diplocalyptis" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="418" phylum="Arthropoda" rank="species" species="operosa">
<emphasis id="B974EAC6FFDFDA4AFC41479FFB113DBA" box="[1013,1238,1943,1967]" italics="true" pageId="9" pageNumber="418">Diplocalyptis operosa</emphasis>
</taxonomicName>
(
<emphasis id="B974EAC6FFDFDA4AFB514790FB3A3DA5" bold="true" box="[1253,1277,1944,1968]" pageId="9" pageNumber="418">D.</emphasis>
Male genitalia,
<emphasis id="B974EAC6FFDFDA4AFF2347B4FF693DC1" bold="true" box="[151,174,1980,2004]" pageId="9" pageNumber="418">E.</emphasis>
Uncus,
<emphasis id="B974EAC6FFDFDA4AFEB647B4FED23DC1" bold="true" box="[258,277,1980,2004]" pageId="9" pageNumber="418">F.</emphasis>
Phallus);
<taxonomicName id="4C004D57FFDFDA4AFECF47B3FD4F3DC6" authorityName="Naik &amp; Shashank" authorityYear="2024" box="[379,648,1979,2003]" class="Insecta" family="Tortricidae" genus="Terthreutis" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="418" phylum="Arthropoda" rank="species" species="subrectangulus" status="sp. nov.">
<emphasis id="B974EAC6FFDFDA4AFECF47B3FD4F3DC6" box="[379,648,1979,2003]" italics="true" pageId="9" pageNumber="418">Terthreutis subrectangulus</emphasis>
</taxonomicName>
<emphasis id="B974EAC6FFDFDA4AFD3A47B4FD183DC1" bold="true" box="[654,735,1980,2004]" pageId="9" pageNumber="418">
<taxonomicNameLabel id="A24757BDFFDFDA4AFD3A47B4FD183DC1" box="[654,735,1980,2004]" pageId="9" pageNumber="418" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(holotype, male) (
<emphasis id="B974EAC6FFDFDA4AFC2F47B3FC723DC6" bold="true" box="[923,949,1979,2003]" pageId="9" pageNumber="418">G.</emphasis>
Male genitalia,
<emphasis id="B974EAC6FFDFDA4AFBEC47B4FBB43DC1" bold="true" box="[1112,1139,1980,2004]" pageId="9" pageNumber="418">H.</emphasis>
Uncus &amp; Socii). Scale bar A, D, G= 0.5 mm; B, C, E, F, H= 0.1 mm.
</paragraph>
</caption>
<subSubSection id="C31A655FFFDCDA49FF73409EFE653E04" pageId="10" pageNumber="419" type="description">
<paragraph id="8BBF36D4FFDCDA49FF73409EFE533B9C" blockId="10.[151,1437,150,1545]" pageId="10" pageNumber="419">
<emphasis id="B974EAC6FFDCDA49FF73409EFE9E3AA5" bold="true" box="[199,345,150,176]" pageId="10" pageNumber="419">Description.</emphasis>
Head. Ocellus small, brown compound eyes. Head dusted cream white, face cream white. Vertex and dorsal part of frons clothed with long pale cream white rough, upwardly projected scales. Ventral part of frons region covered with small, flat, upwardly raised pale ochreous white scales. Labial palpi slender, clothed with forwardly projected whitish cream scales; first segment short, cream white, tinged with brownish cream; second segment curved, brownish white, interspersed with pale ochreous along dorsal margin from base to near apex; small third segment, porrect with large ochreous white scale. Antenna filiform, pale ochreous white with short cilia. Chaetosemata present.
</paragraph>
<paragraph id="8BBF36D4FFDCDA49FF73419BFE3938D8" blockId="10.[151,1437,150,1545]" pageId="10" pageNumber="419">
Thorax. Pronotal collar, tegulae, mesonotum, and posterior crest brownish cream mixed with ochreous yellow scales; legs unmodified, middle legs with a pair of tibial spurs; hind tibia covered with dense, cream white scales interspersed with light cream scales, inner margin with tuft of whitish cream scales. Wings: Forewing (wingspan
<quantity id="4CF89B31FFDCDA49FA3441D3FF01380C" metricMagnitude="-2" metricUnit="m" metricValue="2.0" pageId="10" pageNumber="419" unit="mm" value="20.0">20 mm</quantity>
; n= 5) uniformly broad, costa tinged ochreous, termen weakly oblique, gently sinuate; cream brownish ground color; tornal spot small, brown accompanied by more anterior cream brown elongate blotch; postmedian blotch rounded, brownish cream; some brown dots in apical and terminal area. Hindwing yellowish cream with concolor cilia along the apical and anal margin. Wing venation: Forewing with all the veins separated beyond the discal cell, R4 and R5 stalked, M3 and CuA1 arched basally, CuP indistinct. Hindwing Sc reach up to Rs and M1 stalked, M3 and CuA1 stalked, CuP distinct.
</paragraph>
<paragraph id="8BBF36D4FFDCDA49FF7342DFFE0D39F8" blockId="10.[151,1437,150,1545]" pageId="10" pageNumber="419">Abdomen. Clothed with yellowish cream. Male genitalia: Uncus long, slender, strongly scelerotised, uniform width narrow towards apex; tegumen broad, rectangular, well seclerotised with strong shoulders; socius not prominent; gnathos strong with small lateral prominence and large terminal lobe; gnathal hooks highly sclerotized, from apex of tegumen shoulder; valva large, subrectangular, symmetrical, lightly sclerotised with minute hairs; transtilla incomplete, with two sclerotised lobes; sacculus strongly sclerotized, uniform width, reach 1/2 of valva; cucullus membranous with tufts of hairs; juxta medium, dumbbell- shaped with broad and rounded base; anellus cup shaped and fused with phallus; vinculum medium well sclerotized. Phallus simple, long and slender with uniform width. No cornuti detected.</paragraph>
<paragraph id="8BBF36D4FFDCDA49FF7343FFFE653E04" blockId="10.[151,1437,150,1545]" box="[199,418,1015,1041]" pageId="10" pageNumber="419">
<emphasis id="B974EAC6FFDCDA49FF7343FFFEE23E04" bold="true" box="[199,293,1015,1041]" pageId="10" pageNumber="419">Female.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="C31A655FFFDCDA49FF734412FCF63EFC" pageId="10" pageNumber="419" type="materials_examined">
<paragraph id="8BBF36D4FFDCDA49FF734412FCF63EFC" blockId="10.[151,1437,150,1545]" pageId="10" pageNumber="419">
<emphasis id="B974EAC6FFDCDA49FF734412FE5E3E21" bold="true" box="[199,409,1050,1076]" pageId="10" pageNumber="419">Material studied.</emphasis>
<materialsCitation id="3B683C89FFDCDA49FE104413FC0D3E4C" collectingDate="2021-03-17" collectorName="Mercury &amp; Santhosh Naik G &amp; Sunil" country="India" latitude="25.05611" location="Umiam" longLatPrecision="21" longitude="91.91167" municipality="Ri Bhoi" pageId="10" pageNumber="419" specimenCount="2" specimenCount-male="2" stateProvince="Meghalaya">
<collectingCountry id="F3177644FFDCDA49FE104413FE253E20" box="[420,482,1051,1077]" name="India" pageId="10" pageNumber="419">India</collectingCountry>
:
<collectingRegion id="49C4F836FFDCDA49FE444413FDB63E20" box="[496,625,1051,1077]" country="India" name="Meghalaya" pageId="10" pageNumber="419">Meghalaya</collectingRegion>
:
<collectingMunicipality id="6BDBACAEFFDCDA49FD344413FD273E20" box="[640,736,1051,1077]" pageId="10" pageNumber="419">Ri Bhoi</collectingMunicipality>
: CPGSAS,
<location id="8EDF600FFFDCDA49FCDF4413FC783E20" LSID="urn:lsid:plazi:treatment:03A987C2FFD0DA49FF23458AFAC73C1C:8EDF600FFFDCDA49FCDF4413FC783E20" box="[875,959,1051,1077]" country="India" latitude="25.05611" longLatPrecision="21" longitude="91.91167" municipality="Ri Bhoi" name="Umiam" pageId="10" pageNumber="419" stateProvince="Meghalaya">Umiam</location>
(
<geoCoordinate id="EE345013FFDCDA49FC674412FBA43E20" box="[979,1123,1050,1077]" degrees="25" direction="north" minutes="03" orientation="latitude" pageId="10" pageNumber="419" precision="15" seconds="22" value="25.05611">25°03’22’’N</geoCoordinate>
,
<geoCoordinate id="EE345013FFDCDA49FBC74412FB383E20" box="[1139,1279,1050,1077]" degrees="91" direction="east" minutes="54" orientation="longitude" pageId="10" pageNumber="419" precision="15" seconds="42" value="91.91167">91°54’42’’E</geoCoordinate>
, 1520 MSL),
<specimenCount id="9D06FD5DFFDCDA49FF234437FF7A3E4C" box="[151,189,1087,1113]" count="2" pageId="10" pageNumber="419" type="male">2♂</specimenCount>
,
<date id="FFBE1014FFDCDA49FF784437FE8F3E4C" box="[204,328,1087,1113]" pageId="10" pageNumber="419" value="2021-03-17">
<collectingDate id="EFFAE9FCFFDCDA49FF784437FE8F3E4C" box="[204,328,1087,1113]" pageId="10" pageNumber="419" value="2021-03-17">17.iii.2021</collectingDate>
</date>
,
<collectorName id="26F55302FFDCDA49FEE34437FE703E4C" box="[343,439,1087,1113]" pageId="10" pageNumber="419">Mercury</collectorName>
vapour lamp coll.
<collectorName id="26F55302FFDCDA49FD2D4437FCA73E4C" box="[665,864,1087,1113]" pageId="10" pageNumber="419">Santhosh Naik G</collectorName>
&amp;
<collectorName id="26F55302FFDCDA49FC3F4437FC0D3E4C" box="[907,970,1087,1113]" pageId="10" pageNumber="419">Sunil</collectorName>
</materialsCitation>
;
<materialsCitation id="3B683C89FFDCDA49FC6D4437FB613E68" collectingDate="2020-02-10" collectionCode="R" collectorName="Mercury &amp; Niranjana G N &amp; Tahseen" country="India" elevation="493" latitude="24.197498" location="Kolasib" longLatPrecision="21" longitude="92.69444" pageId="10" pageNumber="419" specimenCount="1" specimenCount-male="1" stateProvince="Mizoram">
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:
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:
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(
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,
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,
<quantity id="4CF89B31FFDCDA49FE9B446BFEB43E68" box="[303,371,1123,1149]" metricMagnitude="2" metricUnit="m" metricValue="4.93" pageId="10" pageNumber="419" unit="m" value="493.0">
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</quantity>
),
<specimenCount id="9D06FD5DFFDCDA49FE32446BFE6C3E68" box="[390,427,1123,1149]" count="1" pageId="10" pageNumber="419" type="male">1♂</specimenCount>
,
<date id="FFBE1014FFDCDA49FE03446BFDEB3E68" box="[439,556,1123,1149]" pageId="10" pageNumber="419" value="2020-02-10">
<collectingDate id="EFFAE9FCFFDCDA49FE03446BFDEB3E68" box="[439,556,1123,1149]" pageId="10" pageNumber="419" value="2020-02-10">10.ii.2020</collectingDate>
</date>
,
<collectorName id="26F55302FFDCDA49FD83446BFD503E68" box="[567,663,1123,1149]" pageId="10" pageNumber="419">Mercury</collectorName>
vapour lamp coll.
<collectorName id="26F55302FFDCDA49FCDD446BFBC93E68" box="[873,1038,1123,1149]" pageId="10" pageNumber="419">Niranjana G N</collectorName>
&amp;
<collectorName id="26F55302FFDCDA49FB85446BFB4B3E68" box="[1073,1164,1123,1149]" pageId="10" pageNumber="419">Tahseen</collectorName>
<collectionCode id="ED11AE11FFDCDA49FB27446BFB613E68" box="[1171,1190,1123,1149]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="10" pageNumber="419">R</collectionCode>
</materialsCitation>
<materialsCitation id="3B683C89FFDCDA49FB19446BFAB13EB4" collectingDate="2020-02-11" collectionCode="R" collectorName="Mercury &amp; Niranjana G N &amp; Tahseen" country="India" elevation="493" latitude="24.197498" location="Kolasib" longLatPrecision="21" longitude="92.69444" pageId="10" pageNumber="419" specimenCount="1" specimenCount-male="1" stateProvince="Mizoram">
H;
<collectingRegion id="49C4F836FFDCDA49FB64446BFAFC3E68" box="[1232,1339,1123,1149]" country="India" name="Mizoram" pageId="10" pageNumber="419">Mizoram</collectingRegion>
:
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(
<geoCoordinate id="EE345013FFDCDA49FF14448EFEE83EB4" box="[160,303,1158,1185]" degrees="24" direction="north" minutes="11" orientation="latitude" pageId="10" pageNumber="419" precision="15" seconds="51" value="24.197498">24°11’51’’N</geoCoordinate>
,
<geoCoordinate id="EE345013FFDCDA49FE8A448FFE0C3EB4" box="[318,459,1159,1185]" degrees="92" direction="east" minutes="41" orientation="longitude" pageId="10" pageNumber="419" precision="15" seconds="40" value="92.69444">92°41’40’’E</geoCoordinate>
,
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<elevation id="002DD1E7FFDCDA49FE6D448FFDD93EB4" box="[473,542,1159,1185]" metricMagnitude="2" metricUnit="m" metricValue="4.93" pageId="10" pageNumber="419" unit="m" value="493.0">493m</elevation>
</quantity>
),
<specimenCount id="9D06FD5DFFDCDA49FD80448FFD9F3EB4" box="[564,600,1159,1185]" count="1" pageId="10" pageNumber="419" type="male">1♂</specimenCount>
,
<date id="FFBE1014FFDCDA49FDDC448FFD1C3EB4" box="[616,731,1159,1185]" pageId="10" pageNumber="419" value="2020-02-11">
<collectingDate id="EFFAE9FCFFDCDA49FDDC448FFD1C3EB4" box="[616,731,1159,1185]" pageId="10" pageNumber="419" value="2020-02-11">11.ii.2020</collectingDate>
</date>
,
<collectorName id="26F55302FFDCDA49FD5D448FFC8E3EB4" box="[745,841,1159,1185]" pageId="10" pageNumber="419">Mercury</collectorName>
vapour lamp coll.
<collectorName id="26F55302FFDCDA49FB9D448FFB133EB4" box="[1065,1236,1159,1185]" pageId="10" pageNumber="419">Niranjana G N</collectorName>
&amp;
<collectorName id="26F55302FFDCDA49FB4A448FFA9E3EB4" box="[1278,1369,1159,1185]" pageId="10" pageNumber="419">Tahseen</collectorName>
<collectionCode id="ED11AE11FFDCDA49FAD7448FFAB13EB4" box="[1379,1398,1159,1185]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="10" pageNumber="419">R</collectionCode>
</materialsCitation>
H;
<materialsCitation id="3B683C89FFDCDA49FF2344A3FE663EFC" collectingDate="2019-11-19" collectorName="Mercury &amp; Shivanshu G" country="India" latitude="31.682222" location="Neri" longLatPrecision="20" longitude="76.487495" municipality="Hamirpur" pageId="10" pageNumber="419" specimenCount="1" specimenCount-female="1" stateProvince="Himachal Pradesh">
<collectingCountry id="F3177644FFDCDA49FF2344A3FF123ED0" box="[151,213,1195,1221]" name="India" pageId="10" pageNumber="419">India</collectingCountry>
:
<collectingRegion id="49C4F836FFDCDA49FF5444A3FE763ED0" box="[224,433,1195,1221]" country="India" name="Himachal Pradesh" pageId="10" pageNumber="419">Himachal Pradesh</collectingRegion>
:
<collectingMunicipality id="6BDBACAEFFDCDA49FE0F44A3FDEB3ED0" box="[443,556,1195,1221]" pageId="10" pageNumber="419">Hamirpur</collectingMunicipality>
:
<location id="8EDF600FFFDCDA49FD8344A3FDAE3ED0" LSID="urn:lsid:plazi:treatment:03A987C2FFD0DA49FF23458AFAC73C1C:8EDF600FFFDCDA49FD8344A3FDAE3ED0" box="[567,617,1195,1221]" country="India" latitude="31.682222" longLatPrecision="20" longitude="76.487495" municipality="Hamirpur" name="Neri" pageId="10" pageNumber="419" stateProvince="Himachal Pradesh">Neri</location>
(
<geoCoordinate id="EE345013FFDCDA49FDCD44A2FCCE3ED0" box="[633,777,1194,1221]" degrees="31" direction="north" minutes="40" orientation="latitude" pageId="10" pageNumber="419" precision="15" seconds="56" value="31.682222">31°40’56’’N</geoCoordinate>
,
<geoCoordinate id="EE345013FFDCDA49FCA144A2FC663ED0" box="[789,929,1194,1221]" degrees="76" direction="east" minutes="29" orientation="longitude" pageId="10" pageNumber="419" precision="15" seconds="15" value="76.487495">76°29’15’’E</geoCoordinate>
, 733 MSL),
<specimenCount id="9D06FD5DFFDCDA49FB8444A3FB943ED0" box="[1072,1107,1195,1221]" count="1" pageId="10" pageNumber="419" type="female">1♀</specimenCount>
,
<date id="FFBE1014FFDCDA49FBD544A3FB1C3ED0" box="[1121,1243,1195,1221]" pageId="10" pageNumber="419" value="2019-11-19">
<collectingDate id="EFFAE9FCFFDCDA49FBD544A3FB1C3ED0" box="[1121,1243,1195,1221]" pageId="10" pageNumber="419" value="2019-11-19">19.xi.2019</collectingDate>
</date>
,
<collectorName id="26F55302FFDCDA49FB5344A3FA803ED0" box="[1255,1351,1195,1221]" pageId="10" pageNumber="419">Mercury</collectorName>
vapour lamp coll.
<collectorName id="26F55302FFDCDA49FEBA44C7FE663EFC" box="[270,417,1231,1257]" pageId="10" pageNumber="419">Shivanshu G</collectorName>
</materialsCitation>
; genitalia slide no. L00050946 (
<specimenCount id="9D06FD5DFFDCDA49FCB844C7FCE43EFC" box="[780,803,1231,1257]" count="1" pageId="10" pageNumber="419" type="male">♂</specimenCount>
).
</paragraph>
</subSubSection>
<subSubSection id="C31A655FFFDCDA49FF7344FBFC6D3F24" pageId="10" pageNumber="419" type="etymology">
<paragraph id="8BBF36D4FFDCDA49FF7344FBFC6D3F24" blockId="10.[151,1437,150,1545]" pageId="10" pageNumber="419">
<emphasis id="B974EAC6FFDCDA49FF7344FBFE893F18" bold="true" box="[199,334,1267,1293]" pageId="10" pageNumber="419">Etymology.</emphasis>
The new species
<taxonomicName id="4C004D57FFDCDA49FDA644FBFCFC3F18" authorityName="Naik &amp; Shashank" authorityYear="2024" box="[530,827,1267,1293]" class="Insecta" family="Tortricidae" genus="Terthreutis" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="419" phylum="Arthropoda" rank="species" species="subrectangulus" status="sp. nov.">
<emphasis id="B974EAC6FFDCDA49FDA644FBFCFC3F18" box="[530,827,1267,1293]" italics="true" pageId="10" pageNumber="419">Terthreutis subrectangulus</emphasis>
</taxonomicName>
<emphasis id="B974EAC6FFDCDA49FCF544FBFC5D3F18" bold="true" box="[833,922,1267,1293]" pageId="10" pageNumber="419">
<taxonomicNameLabel id="A24757BDFFDCDA49FCF544FBFC5D3F18" box="[833,922,1267,1293]" pageId="10" pageNumber="419" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is named after the subrectangular shape of the valva, which is distinct from all other described species of
<taxonomicName id="4C004D57FFDCDA49FC9E451FFC643F24" authorityName="Meyrick" authorityYear="1918" box="[810,931,1303,1329]" class="Insecta" family="Tortricidae" genus="Terthreutis" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="419" phylum="Arthropoda" rank="genus">
<emphasis id="B974EAC6FFDCDA49FC9E451FFC643F24" box="[810,931,1303,1329]" italics="true" pageId="10" pageNumber="419">Terthreutis</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C31A655FFFDCDA49FF734532FC413F40" box="[199,902,1338,1365]" pageId="10" pageNumber="419" type="distribution">
<paragraph id="8BBF36D4FFDCDA49FF734532FC413F40" blockId="10.[151,1437,150,1545]" box="[199,902,1338,1365]" pageId="10" pageNumber="419">
<emphasis id="B974EAC6FFDCDA49FF734532FEA63F41" bold="true" box="[199,353,1338,1364]" pageId="10" pageNumber="419">Distribution.</emphasis>
<collectingCountry id="F3177644FFDCDA49FED34533FE663F40" box="[359,417,1339,1365]" name="India" pageId="10" pageNumber="419">India</collectingCountry>
(
<collectingRegion id="49C4F836FFDCDA49FE1B4533FD463F40" box="[431,641,1339,1365]" country="India" name="Himachal Pradesh" pageId="10" pageNumber="419">Himachal Pradesh</collectingRegion>
,
<collectingRegion id="49C4F836FFDCDA49FD384533FCCC3F40" box="[652,779,1339,1365]" country="India" name="Meghalaya" pageId="10" pageNumber="419">Meghalaya</collectingRegion>
,
<collectingRegion id="49C4F836FFDCDA49FCA24533FCB83F40" box="[790,895,1339,1365]" country="India" name="Mizoram" pageId="10" pageNumber="419">Mizoram</collectingRegion>
)
</paragraph>
</subSubSection>
<subSubSection id="C31A655FFFDCDA49FF734557FAC73C1C" pageId="10" pageNumber="419" type="discussion">
<paragraph id="8BBF36D4FFDCDA49FF734557FAC73C1C" blockId="10.[151,1437,150,1545]" pageId="10" pageNumber="419">
<emphasis id="B974EAC6FFDCDA49FF734557FEFC3F6C" bold="true" box="[199,315,1375,1401]" pageId="10" pageNumber="419">Remarks.</emphasis>
Genus
<taxonomicName id="4C004D57FFDCDA49FE3B4557FDCF3F6C" authorityName="Meyrick" authorityYear="1918" box="[399,520,1375,1401]" class="Insecta" family="Tortricidae" genus="Terthreutis" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="419" phylum="Arthropoda" rank="genus">
<emphasis id="B974EAC6FFDCDA49FE3B4557FDCF3F6C" box="[399,520,1375,1401]" italics="true" pageId="10" pageNumber="419">Terthreutis</emphasis>
</taxonomicName>
is described by Meyrick (1918), and species of the genus were studied by
<bibRefCitation id="EF914B25FFDCDA49FAF44557FEE93F88" author="Kawabe, A. &amp; Komai, F. &amp; Razwoski, J." pageId="10" pageNumber="419" pagination="103 - 109" refId="ref6698" refString="Kawabe, A., Komai, F. &amp; Razwoski, J. (1992) Tortricidae. In: Heppner, J. B. &amp; Inoue, H. (Eds.), Lepidoptera of Taiwan. Vol. 1. Part 2. Checklist. The Association for Tropical Lepidoptera, Gainesville, Florida, pp. 103 - 109." type="book chapter" year="1992">
Kawabe
<emphasis id="B974EAC6FFDCDA49FF23458BFF113F88" box="[151,214,1411,1437]" italics="true" pageId="10" pageNumber="419">et al.</emphasis>
(1992)
</bibRefCitation>
and
<bibRefCitation id="EF914B25FFDCDA49FEC5458BFDFC3F88" author="Razowski, J." box="[369,571,1411,1437]" pageId="10" pageNumber="419" pagination="283 - 299" refId="ref7112" refString="Razowski, J. (2008) On two South Asian genera Ceramea Diakonoff and Terthreutis Meyrick (Lepidoptera: Tortricidae). Polish Journal of Entomology, 4 (77), 283 - 299." type="journal article" year="2008">Razowski (2008)</bibRefCitation>
.
<taxonomicName id="4C004D57FFDCDA49FDF9458BFCBA3F88" authorityName="Naik &amp; Shashank" authorityYear="2024" box="[589,893,1411,1437]" class="Insecta" family="Tortricidae" genus="Terthreutis" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="419" phylum="Arthropoda" rank="species" species="subrectangulus" status="sp. nov.">
<emphasis id="B974EAC6FFDCDA49FDF9458BFCBA3F88" box="[589,893,1411,1437]" italics="true" pageId="10" pageNumber="419">Terthreutis subrectangulus</emphasis>
</taxonomicName>
<emphasis id="B974EAC6FFDCDA49FC3F458BFC2C3F88" bold="true" box="[907,1003,1411,1437]" pageId="10" pageNumber="419">
<taxonomicNameLabel id="A24757BDFFDCDA49FC3F458BFC2C3F88" box="[907,1003,1411,1437]" pageId="10" pageNumber="419" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
resembles
<taxonomicName id="4C004D57FFDCDA49FBC2458BFE8C3FD4" authority="Razowski, 2008" authorityName="Razowski" authorityYear="2008" class="Insecta" family="Tortricidae" genus="Terthreutis" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="419" phylum="Arthropoda" rank="species" species="chiangmaiana">
<emphasis id="B974EAC6FFDCDA49FBC2458BFA5B3F88" box="[1142,1436,1411,1437]" italics="true" pageId="10" pageNumber="419">Terthreutis chiangmaiana</emphasis>
<bibRefCitation id="EF914B25FFDCDA49FF2345AFFE8C3FD4" author="Razowski, J." box="[151,331,1447,1473]" pageId="10" pageNumber="419" pagination="283 - 299" refId="ref7112" refString="Razowski, J. (2008) On two South Asian genera Ceramea Diakonoff and Terthreutis Meyrick (Lepidoptera: Tortricidae). Polish Journal of Entomology, 4 (77), 283 - 299." type="journal article" year="2008">
Razowski,
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</bibRefCitation>
</taxonomicName>
in the appearance, but differs in the male genitalia, transtilla incomplete, valva subrectangular, and phallus is smooth without any serration; whereas in
<taxonomicName id="4C004D57FFDCDA49FD5645C3FC593FF0" authorityName="Razowski" authorityYear="2008" box="[738,926,1483,1509]" class="Insecta" family="Tortricidae" genus="Terthreutis" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="419" phylum="Arthropoda" rank="species" species="chiangmaiana">
<emphasis id="B974EAC6FFDCDA49FD5645C3FC593FF0" box="[738,926,1483,1509]" italics="true" pageId="10" pageNumber="419">T. chiangmaiana</emphasis>
</taxonomicName>
the transtilla is complete and continuous, the valva is rounded, and the phallus is serrated with the presence of some denticles in the distal portion.
</paragraph>
</subSubSection>
</treatment>
</document>
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