272 lines
20 KiB
XML
272 lines
20 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/phytokeys.138.38721" ID-PMC="PMC6969003" ID-Pensoft-Pub="1314-2003-138-203" ID-Pensoft-UUID="37174C2A6257592B83BED9D2E2CAA79B" ID-PubMed="31988614" ModsDocID="1314-2003-138-203" checkinTime="1578758105156" checkinUser="pensoft" docAuthor="Maw, Mya Bhone, Ding, Hong-Bo, Yang, Bin, Win, Pyae Pyae & Tan, Yun-Hong" docDate="2020" docId="E6C1FEBDF7065D1D9D705DF6C5CF679A" docLanguage="en" docName="PhytoKeys 138: 203-217" docOrigin="PhytoKeys 138" docSource="http://dx.doi.org/10.3897/phytokeys.138.38721" docTitle="Begonia crassitepala Y. H. Tan & M. B. Maw 2020, sp. nov." docType="treatment" docVersion="5" id="37174C2A6257592B83BED9D2E2CAA79B" lastPageNumber="203" masterDocId="37174C2A6257592B83BED9D2E2CAA79B" masterDocTitle="Taxonomic studies on Begonia (Begoniaceae) in Myanmar I: three new species and supplementary description of Begonia rheophytica from Northern Myanmar" masterLastPageNumber="217" masterPageNumber="203" pageNumber="203" updateTime="1668139614508" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Taxonomic studies on Begonia (Begoniaceae) in Myanmar I: three new species and supplementary description of Begonia rheophytica from Northern Myanmar</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Maw, Mya Bhone</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Ding, Hong-Bo</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Yang, Bin</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Win, Pyae Pyae</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Tan, Yun-Hong</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>PhytoKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2020</mods:date>
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<mods:detail type="volume">
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<mods:number>138</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>203</mods:start>
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<mods:end>217</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/phytokeys.138.38721</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/phytokeys.138.38721</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1314-2003-138-203</mods:identifier>
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<mods:identifier type="Pensoft-UUID">37174C2A6257592B83BED9D2E2CAA79B</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="161672083" LSID="urn:lsid:plazi:treatment:E6C1FEBDF7065D1D9D705DF6C5CF679A" httpUri="http://treatment.plazi.org/id/E6C1FEBDF7065D1D9D705DF6C5CF679A" lastPageNumber="203" pageId="0" pageNumber="203">
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<subSubSection pageId="0" pageNumber="203" type="nomenclature">
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<paragraph pageId="0" pageNumber="203">
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<taxonomicName LSID="E6C1FEBD-F706-5D1D-9D70-5DF6C5CF679A" authority="Y. H. Tan & M. B. Maw" authorityName="Y. H. Tan & M. B. Maw" authorityYear="2020" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Begonia crassitepala" order="Cucurbitales" pageId="0" pageNumber="203" phylum="Tracheophyta" rank="species" species="crassitepala" status="sp. nov.">Begonia crassitepala Y.H.Tan & M.B.Maw</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="203">sp. nov.</taxonomicNameLabel>
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<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Begonia crassitepala Y. H. Tan & M. B. Maw, sp. nov. (photographed by H. B. Ding) A habit B inflorescence C stipule and stem (showing the whitish soft spine-like hairs) D rhizome E single leaf (front view) F single leaf (back view) G hirsute leaf margins H flowers (front view staminate flower) I staminate flower (back view) J dried fruits K pistillate flower (front view) M staminate flower showing 4 tepals N pistillate flower (side view) L capsule and cross section of ovary." figureDoi="10.3897/phytokeys.138.38721.figure4" httpUri="https://binary.pensoft.net/fig/370292" pageId="0" pageNumber="203">Figure 4</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="203" type="diagnosis">
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<paragraph pageId="0" pageNumber="203">Diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="203">
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<taxonomicName authorityName="Y. H. Tan & M. B. Maw" authorityYear="2020" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Begonia crassitepala" order="Cucurbitales" pageId="0" pageNumber="203" phylum="Tracheophyta" rank="species" species="crassitepala">
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<emphasis italics="true" pageId="0" pageNumber="203">Begonia crassitepala</emphasis>
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</taxonomicName>
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Y.H. Tan & M.B. Maw is morphologically similar to
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<taxonomicName lsidName="B. dryadis" pageId="0" pageNumber="203" rank="species" species="dryadis">
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<emphasis italics="true" pageId="0" pageNumber="203">B. dryadis</emphasis>
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</taxonomicName>
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<bibRefCitation author="Irmscher, E" journalOrPublisher="Notes from the Royal Botanic Garden Edinburgh" pageId="0" pageNumber="203" pagination="35 - 45" refId="B9" refString="Irmscher, E, 1951. Some new Chinese species of Begonia. Notes from the Royal Botanic Garden Edinburgh 21 (1): 35 - 45" title="Some new Chinese species of Begonia." volume="21" year="1951">Irmscher (1951</bibRefCitation>
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: 41) in ovate to broadly ovate leaf, but it can be distinguished by its stem and petiole having white prickles (
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<emphasis italics="true" pageId="0" pageNumber="203">vs.</emphasis>
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puberulous), adaxially leaf having densely pinkish or grey hirsute (
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<emphasis italics="true" pageId="0" pageNumber="203">vs.</emphasis>
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subglabrous), abaxially outer 2 tepals of pistillate flower and ovary having reddish or whitish succulent strigose and tuberculate (
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<emphasis italics="true" pageId="0" pageNumber="203">vs.</emphasis>
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puberulous).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="203" type="materials_examined">
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<paragraph pageId="0" pageNumber="203">Type.</paragraph>
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<paragraph pageId="0" pageNumber="203">
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Myanmar, Kachin State, Putao District, on the way from Ratbaw to Alanga,
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<geoCoordinate degrees="27" direction="north" minutes="17" orientation="latitude" precision="1" seconds="13.73" value="27.287148">27°17'13.73"N</geoCoordinate>
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,
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<geoCoordinate degrees="97" direction="east" minutes="44" orientation="longitude" precision="1" seconds="24.28" value="97.740074">97°44'24.28"E</geoCoordinate>
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, 836 m, 15 June 2018,
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<emphasis italics="true" pageId="0" pageNumber="203">Y.H. Tan</emphasis>
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,
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<emphasis italics="true" pageId="0" pageNumber="203">B. Yang</emphasis>
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,
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<emphasis italics="true" pageId="0" pageNumber="203">H.B. Ding</emphasis>
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,
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<emphasis italics="true" pageId="0" pageNumber="203">X.D. Zeng</emphasis>
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,
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<emphasis italics="true" pageId="0" pageNumber="203">M.B. Maw & H.L. Naing M4495</emphasis>
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(holotype: HITBC!; isotypes: RAF!).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="203" type="description">
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<paragraph pageId="0" pageNumber="203">Description.</paragraph>
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<paragraph pageId="0" pageNumber="203">
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Perennial herb, monoecious, rhizomatous.
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<emphasis bold="true" pageId="0" pageNumber="203">Stem</emphasis>
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erect, 40-60 cm tall, reddish, densely rusty wooly tomentose, and sparsely covered by whitish soft spine-like hairs, internodes 1-2 cm long.
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<emphasis bold="true" pageId="0" pageNumber="203">Stipule</emphasis>
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lanceolate, 1-1.3
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<normalizedToken originalValue="×">x</normalizedToken>
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0.3-0.5 cm, slightly or densely rusty tomentose, deciduous.
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<emphasis bold="true" pageId="0" pageNumber="203">Leaf</emphasis>
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petiole 4.5-15 cm, densely rusty tomentose, slightly whitish stiff hairs;
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<emphasis bold="true" pageId="0" pageNumber="203">blade</emphasis>
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ovate to broadly ovate, base cordate with overlapping lobes, 12-20
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<normalizedToken originalValue="×">x</normalizedToken>
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15-19.5 cm, asymmetric, adaxiall
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<emphasis bold="true" pageId="0" pageNumber="203">y</emphasis>
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green, densely pinkish or grey hirsute, abaxially green, rusty puberulous, venation palmate-pinnate, densely rusty wooly tomentose along midrib and veins, margin denticulate, reddish hirsute along the margin, apex acuminate.
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<emphasis bold="true" pageId="0" pageNumber="203">Inflorescence</emphasis>
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nearly terminal, cymose, peduncle 4.4-6.0 cm long, reddish, slightly rusty wooly tomentose and whitish soft spine-like hairs.
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<emphasis bold="true" pageId="0" pageNumber="203">Staminate flower</emphasis>
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: pedicel 1.8-2.5 cm long, whitish villose, tepals 4, unequal, outer 2, ovate, 2.2-2.7
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<normalizedToken originalValue="×">x</normalizedToken>
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2.6-2.8 cm, whitish or pinkish with pink lining, thick, ca. 2 mm, inner 2, smaller, 1.7
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<normalizedToken originalValue="×">x</normalizedToken>
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2.1 cm, whitish or pinkish (sometimes with pinkish lining), densely whitish villose on the outer surface; stamen numerous, ca. 200; filaments ca. 2 mm long, fused at base; anther oblong to elliptic, 1-2 mm long.
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<emphasis bold="true" pageId="0" pageNumber="203">Pistillate flower</emphasis>
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: pedicel 2.2-2.5 cm, densely whitish or rusty villose, bracteoles absent; tepals 5, equal, obovate, outer 2, 2.0-2.9
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<normalizedToken originalValue="×">x</normalizedToken>
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1.8-2.0 cm, pure white or sometimes with pink lining, densely pinkish or whitish strigose on outer surface, margin entire, inner 3, similar to outer ones but smaller, 1.8-2.1
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<normalizedToken originalValue="×">x</normalizedToken>
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1.6-1.8 cm, ovary 2-locular, placentation axillary, placentae 2 per locule, densely reddish or whitish succulent strigose and tuberculate, styles 2, fused at base, stigma bifid with twisted bands, highly convolute, yellow or golden yellow.
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<emphasis bold="true" pageId="0" pageNumber="203">Fruit</emphasis>
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berry-like, elliptic, pale green to pink, 3 wings, unequal, with whitish or reddish succulent strigose and tuberculate (especially on wings).
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</paragraph>
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<caption doi="10.3897/phytokeys.138.38721.figure4" httpUri="https://binary.pensoft.net/fig/370292" pageId="0" pageNumber="203" start="Figure 4" startId="F4">
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<paragraph pageId="0" pageNumber="203">
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<emphasis bold="true" pageId="0" pageNumber="203">Figure 4.</emphasis>
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<taxonomicName authorityName="Y. H. Tan & M. B. Maw" authorityYear="2020" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Begonia crassitepala" order="Cucurbitales" pageId="0" pageNumber="203" phylum="Tracheophyta" rank="species" species="crassitepala">
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<emphasis italics="true" pageId="0" pageNumber="203">Begonia crassitepala</emphasis>
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</taxonomicName>
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Y.H.Tan & M.B.Maw, sp. nov. (photographed by H.B. Ding)
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<emphasis bold="true" pageId="0" pageNumber="203">A</emphasis>
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habit
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<emphasis bold="true" pageId="0" pageNumber="203">B</emphasis>
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inflorescence
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<emphasis bold="true" pageId="0" pageNumber="203">C</emphasis>
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stipule and stem (showing the whitish soft spine-like hairs)
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<emphasis bold="true" pageId="0" pageNumber="203">D</emphasis>
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rhizome
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<emphasis bold="true" pageId="0" pageNumber="203">E</emphasis>
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single leaf (front view)
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<emphasis bold="true" pageId="0" pageNumber="203">F</emphasis>
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single leaf (back view)
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<emphasis bold="true" pageId="0" pageNumber="203">G</emphasis>
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hirsute leaf margins
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<emphasis bold="true" pageId="0" pageNumber="203">H</emphasis>
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flowers (front view staminate flower)
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<emphasis bold="true" pageId="0" pageNumber="203">I</emphasis>
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staminate flower (back view)
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<emphasis bold="true" pageId="0" pageNumber="203">J</emphasis>
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dried fruits
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<emphasis bold="true" pageId="0" pageNumber="203">K</emphasis>
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pistillate flower (front view)
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<emphasis bold="true" pageId="0" pageNumber="203">M</emphasis>
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staminate flower showing 4 tepals
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<emphasis bold="true" pageId="0" pageNumber="203">N</emphasis>
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pistillate flower (side view)
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<emphasis bold="true" pageId="0" pageNumber="203">L</emphasis>
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capsule and cross section of ovary.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="203" type="phenology">
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<paragraph pageId="0" pageNumber="203">Phenology.</paragraph>
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<paragraph pageId="0" pageNumber="203">Flowering from June-July; fruiting from July-August.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="203" type="distribution">
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<paragraph pageId="0" pageNumber="203">Distribution.</paragraph>
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<paragraph pageId="0" pageNumber="203">Endemic to the type locality, Putao District, Kachin State, Northern Myanmar.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="203" type="etymology">
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<paragraph pageId="0" pageNumber="203">Etymology.</paragraph>
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<paragraph pageId="0" pageNumber="203">The species epithet refers to its thick tepals.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="203" type="ecology">
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<paragraph pageId="0" pageNumber="203">Ecology.</paragraph>
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<paragraph pageId="0" pageNumber="203">In the tropical montane forest up to about 577 m elevation, on the moist soil slope.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="203" type="conservation status">
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<paragraph pageId="0" pageNumber="203">Conservation status.</paragraph>
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<paragraph pageId="0" pageNumber="203">
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Data Deficient (DD).
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<taxonomicName authorityName="Y. H. Tan & M. B. Maw" authorityYear="2020" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Begonia crassitepala" order="Cucurbitales" pageId="0" pageNumber="203" phylum="Tracheophyta" rank="species" species="crassitepala">
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<emphasis italics="true" pageId="0" pageNumber="203">Begonia crassitepala</emphasis>
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</taxonomicName>
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have been collected along the roadside from Langsa to Gawlaw village where no signs of major anthropogenic disturbance were noticed. Further exploration is required to access the current range of the species (
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<bibRefCitation author="IUCN" journalOrPublisher="Botanical Bulletin of Academia Sinica" pageId="0" pageNumber="203" publicationUrl="http://www.iucnredlist.org/documents/RedListGuidelines.pdf" refId="B16" refString="IUCN, 2017. Guidelines for using the IUCN red list categories and criteria. version 13. Prepared by the Standards and Petitions Subcommittee. http://www.iucnredlist.org/documents/RedListGuidelines.pdf" title="Guidelines for using the IUCN red list categories and criteria. version 13. Prepared by the Standards and Petitions Subcommittee." url="http://www.iucnredlist.org/documents/RedListGuidelines.pdf" year="2017">IUCN 2017</bibRefCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="203" type="materials_examined">
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<paragraph pageId="0" pageNumber="203">Additional specimens examined</paragraph>
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<paragraph pageId="0" pageNumber="203">
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<emphasis bold="true" pageId="0" pageNumber="203">(paratypes).</emphasis>
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Myanmar. Kachin State: Putao District, along Langsa to Gawlei, tropical montane forest,
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<geoCoordinate degrees="27" direction="north" minutes="32" orientation="latitude" precision="1" seconds="28.94" value="27.541372">27°32'28.94"N</geoCoordinate>
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,
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<geoCoordinate degrees="97" direction="east" minutes="56" orientation="longitude" precision="1" seconds="36.09" value="97.94336">97°56'36.09"E</geoCoordinate>
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, 577 m, 2 June 2018,
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<emphasis italics="true" pageId="0" pageNumber="203">Myanmar Exped. M3952</emphasis>
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(HITBC!; RAF!); Kachin State, Putao District, Gathu to Tongwang Cave,
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<geoCoordinate degrees="27" direction="north" minutes="29" orientation="latitude" precision="1" seconds="53.48" value="27.49819">27°29'53.48"N</geoCoordinate>
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,
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<geoCoordinate degrees="97" direction="east" minutes="58" orientation="longitude" precision="1" seconds="30.84" value="97.975235">97°58'30.84"E</geoCoordinate>
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, 664 m, 4 June 2018,
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<emphasis italics="true" pageId="0" pageNumber="203">Myanmar Exped. M4008</emphasis>
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(HITBC!; RAF!); Kachin State, Putao District, Gathu to Tangsa,
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<geoCoordinate degrees="27" direction="north" minutes="28" orientation="latitude" precision="1" seconds="41.17" value="27.478104">27°28'41.17"N</geoCoordinate>
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,
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<geoCoordinate degrees="97" direction="east" minutes="56" orientation="longitude" precision="1" seconds="46.40" value="97.94622">97°56'46.40"E</geoCoordinate>
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, 550 m, 31 May 2018,
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<emphasis italics="true" pageId="0" pageNumber="203">Myanmar Exped. M3797</emphasis>
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(HITBC!; RAF!); Kachin State, Putao District, Putao Township, Maliraing area, buffer zone of Hkakaborazi National Park, between camp 1 and camp 2,
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<geoCoordinate degrees="27" direction="north" minutes="38" orientation="latitude" precision="1" seconds="03.6" value="27.634333">27°38'03.6"N</geoCoordinate>
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,
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<geoCoordinate degrees="97" direction="east" minutes="22" orientation="longitude" precision="1" seconds="11.2" value="97.36978">97°22'11.2"E</geoCoordinate>
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, 552 m, 6 November 2016,
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<emphasis italics="true" pageId="0" pageNumber="203">Kate et al. 2253</emphasis>
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(NY02653741!); Kachin State, Putao District, Naungmung Township, buffer zone of Hkakaborazi National Park, Hill next to Naungmung village,
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<geoCoordinate degrees="27" direction="north" minutes="31" orientation="latitude" precision="1" seconds="02.2" value="27.517277">27°31'02.2"N</geoCoordinate>
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,
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<geoCoordinate degrees="97" direction="east" minutes="50" orientation="longitude" precision="1" seconds="46.2" value="97.84617">97°50'46.2"E</geoCoordinate>
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, 845 m, 12 June 2017,
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<emphasis italics="true" pageId="0" pageNumber="203">Kate et al. 2880</emphasis>
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(NY02653917!); Between N Dung Ga and Ting Pru Ting Sar, 27 August 1953,
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<emphasis italics="true" pageId="0" pageNumber="203">Thar Hla & Chit Ko Ko 4447</emphasis>
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(RAF!).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="203" type="affinities">
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<paragraph pageId="0" pageNumber="203">Affinities.</paragraph>
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<paragraph pageId="0" pageNumber="203">
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<taxonomicName authorityName="Y. H. Tan & M. B. Maw" authorityYear="2020" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Begonia crassitepala" order="Cucurbitales" pageId="0" pageNumber="203" phylum="Tracheophyta" rank="species" species="crassitepala">
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<emphasis italics="true" pageId="0" pageNumber="203">Begonia crassitepala</emphasis>
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</taxonomicName>
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is the most distinct species in the
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||
<taxonomicName class="Magnoliopsida" family="Begoniaceae" genus="Begonia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="" order="Cucurbitales" pageId="0" pageNumber="203" phylum="Tracheophyta" rank="section" section="Platycentrum">section Platycentrum</taxonomicName>
|
||
thanks to its thicken tepals and succulent strigose and tuberculate ovary. The new species shares similar characteristics with
|
||
<taxonomicName lsidName="B. dryadis" pageId="0" pageNumber="203" rank="species" species="dryadis">
|
||
<emphasis italics="true" pageId="0" pageNumber="203">B. dryadis</emphasis>
|
||
</taxonomicName>
|
||
in ovate to broadly ovate leaf and 4 tepals of staminate flower. However, it can be easily distinguished by its stem and petiole having rusty tomentose and whitish soft spine-like hairs (
|
||
<emphasis italics="true" pageId="0" pageNumber="203">vs.</emphasis>
|
||
puberulous), adaxially leaf having densely pinkish or grey hirsute (
|
||
<emphasis italics="true" pageId="0" pageNumber="203">vs.</emphasis>
|
||
subglabrous), adaxially outer 2 tepals of pistillate flower having densely pinkish or whitish strigose (
|
||
<emphasis italics="true" pageId="0" pageNumber="203">vs.</emphasis>
|
||
puberulous), ovary having densely reddish or whitish succulent strigose and tuberculate (
|
||
<emphasis italics="true" pageId="0" pageNumber="203">vs.</emphasis>
|
||
puberulous).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |