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<mods:title id="E55CBD4A2371309BDD683BEAB5ADAAB2">The operculate micro land snail genus Dicharax Kobelt &amp; Moellendorff, 1900 (Caenogastropoda, Alycaeidae) in Thailand, with description of new species</mods:title>
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<mods:affiliation id="7AF0542F0182E6C0C81CE2899BDB8A64">Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand</mods:affiliation>
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<mods:namePart id="219165C11D071874A70E14846879DC63">Pall-Gergely, Barna</mods:namePart>
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<mods:affiliation id="ED45A03937A63ABCCDABF10934333257">Plant Protection Institute, Centre for Agricultural Research, Herman Otto ut 15, H- 1022 Budapest, Hungary</mods:affiliation>
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<mods:namePart id="E7F57CF37A4BB70CD29B78630E7FD010">Sutcharit, Chirasak</mods:namePart>
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<mods:namePart id="7FF97B8B9173D631538C0187F0699A23">Tongkerd, Piyoros</mods:namePart>
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<treatment id="E64839C4C6075B7A80C6684D5F22F4E4" ID-GBIF-Taxon="182236442" LSID="urn:lsid:plazi:treatment:E64839C4C6075B7A80C6684D5F22F4E4" httpUri="http://treatment.plazi.org/id/E64839C4C6075B7A80C6684D5F22F4E4" lastPageNumber="1" pageId="0" pageNumber="1" scope_family="Alycaeidae" scope_subClass="Caenogastropoda">
<subSubSection id="749CCB6B0714541DA906AA1408E73096" pageId="0" pageNumber="1" type="nomenclature">
<paragraph id="6FEB85884CB1CE5162B10BD4CC644FC0" pageId="0" pageNumber="1">
<taxonomicName id="E2A7B862BE45CDCAAA8D71E857B09554" LSID="E64839C4-C607-5B7A-80C6-684D5F22F4E4" authority="(Theobald, 1870)" baseAuthorityName="Theobald" baseAuthorityYear="1870" class="Gastropoda" family="Cyclophoridae" genus="Dicharax" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Dicharax cucullatus" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="cucullatus">Dicharax cucullatus (Theobald, 1870)</taxonomicName>
<figureCitation id="355110E35631D7A04AF7BFB3EC42790F" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Dicharax cucullatus A. Syntype of Alycaeus cucullatus, NHMUK 1888.12.4.951; B. Syntype of Alycaeus davisi, NHMUK 1903.7.1.1630; C, D. Specimens CUMZ 5071 from Tham Ban Luang, Doi Ang Khang, Chiang Mai Province. Close-up images of the aperture not to scale. Photo: NHM (A, B.)." figureDoi="10.3897/zse.97.59143.figure2" httpUri="https://binary.pensoft.net/fig/494366" pageId="0" pageNumber="1">Figs 2</figureCitation>
<figureCitation id="38C0B2A52DDC0DC01453BFB882909002" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Dicharax cucullatus from Chiang Mai Province A. Specimen CUMZ 7421 from Doi Chiang Dao; B. Specimen CUMZ 5071 from Tham Ban Luang, Doi Ang Khang; C. Specimen CUMZ 7422 from Siribhum Waterfall, Doi Inthanon; D. Specimen CUMZ 7423 from Wachirathan Waterfall, Doi Inthanon; E. Specimen CUMZ 7424 from Doi Inthanon, 2000 m a. s. l. Close-up images of the aperture not to scale." figureDoi="10.3897/zse.97.59143.figure3" httpUri="https://binary.pensoft.net/fig/494367" pageId="0" pageNumber="1">, 3</figureCitation>
<figureCitation id="A91BBE075F07E63553F3DC6309CCA69E" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. A, B. Dicharax cucullatus, specimens CUMZ 7422 from Siribhum Waterfall, Doi Inthanon, Chiang Mai Province; A. With operculum; C, D. Dicharax notus C. Syntype NHMUK 1903.7.1.3065; D. Specimen CUMZ 7425 from Tham Luang Sa Koen, Nan Province. Close-up images of the aperture not to scale. Photo: NHM (C.)." figureDoi="10.3897/zse.97.59143.figure4" httpUri="https://binary.pensoft.net/fig/494368" pageId="0" pageNumber="1">, 4A, B</figureCitation>
<figureCitation id="BE37E087B5F550F2D1468DC2E535EEAA" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Living Dicharax specimens. A. Dicharax cucullatus, specimen CUMZ 7432 from Doi Inthanon, Chiang Mai Province; B. Dicharax notus, specimen CUMZ 7425 from Tham Luang Sa Koen, Nan Province. Both photographed in the laboratory. Not to scale. Photo: A. Pholyotha." figureDoi="10.3897/zse.97.59143.figure5" httpUri="https://binary.pensoft.net/fig/494369" pageId="0" pageNumber="1">, 5A</figureCitation>
<figureCitation id="EE46AEC94D08C7730B034EB7A0557063" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Synoptic view of the eight Dicharax taxa in Thailand. A. Dicharax omissus, specimen CUMZ 7426 (gold plated); B. Dicharax pratatensis, specimen CUMZ 7427; C. Dicharax panhai sp. nov., paratype CUMZ 7429 / 2; D. Dicharax pongrati sp. nov., paratype CUMZ 7430 / 2; E. Dicharax notus, specimen CUMZ 7425; F. Dicharax burchi sp. nov., paratype CUMZ 7428 / 2; G. Dicharax borealis sp. nov., paratype CUMZ 5072 / 2; H. Dicharax cucullatus, specimen CUMZ 7421." figureDoi="10.3897/zse.97.59143.figure6" httpUri="https://binary.pensoft.net/fig/494370" pageId="0" pageNumber="1">, 6H</figureCitation>
</paragraph>
</subSubSection>
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<paragraph id="2C8BCFBCB1B5945D1919B69AAD3675F7" pageId="0" pageNumber="1">
<taxonomicName id="D17E43C4FD66B80F7FFB1A80BF6679F2" authorityName="Theobald" authorityYear="1870" class="Gastropoda" family="Cyclophoridae" genus="Alycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alycaeus cucullatus" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="cucullatus">Alycaeus cucullatus</taxonomicName>
Theobald, 1870: 396, 397, pl. 18, fig. 2. Type locality: Shan States [Myanmar].
<bibRefCitation id="A3AE004AE187E6FECE1F525581D037AD" author="Hanley, SCT" journalOrPublisher="Annals of the Lyceum of Natural History of New York" pageId="0" pageNumber="1" refId="B23" refString="Hanley, SCT, Theobald, W, 1874.] [Published in parts, dates follow Prashad (1927)]" title="] [Published in parts, dates follow Prashad (1927)]" year="1874">Hanley and Theobald 1874</bibRefCitation>
: 39, pl. 96, figs 1, 4;
<bibRefCitation id="98A011EB93F6B7BB881604594779430B" author="Reeve, LA" journalOrPublisher="Journal and Proceedings of the Asiatic Society of Bengal, new series" pageId="0" pageNumber="1" refId="B53" refString="Reeve, LA, 1878. Monograph of the genus Alycaeus. In: Reeve LA (Ed.) Conchologica Iconica: or, Illustrations of the Shells of Molluscous Animals. Reeve and Co., London, 20: 13 pp." title="Monograph of the genus Alycaeus. In: Reeve LA (Ed.) Conchologica Iconica: or, Illustrations of the Shells of Molluscous Animals. Reeve and Co., London, 20: 13 pp." year="1878">Reeve 1878</bibRefCitation>
: pl. 2, sp. 12;
<bibRefCitation id="D41EE31EDC2E79096DF6AE1173E8210D" author="Godwin-Austen, HH" journalOrPublisher="Journal of the Asiatic Society of Bengal" pageId="0" pageNumber="1" refId="B19" refString="Godwin-Austen, HH, 1914. Land and freshwater mollusca of India, including South Arabia, Baluchistan, Afghanistan, Kashmir, Nepal, Burma, Pegu, Tenasserim, Malaya Peninsula, Ceylon and other islands of the Indian Ocean. Vol. 2, Part 12: 311-442." title="Land and freshwater mollusca of India, including South Arabia, Baluchistan, Afghanistan, Kashmir, Nepal, Burma, Pegu, Tenasserim, Malaya Peninsula, Ceylon and other islands of the Indian Ocean. Vol. 2, Part 12: 311 - 442." year="1914">Godwin-Austen 1914</bibRefCitation>
: 407, pl. 155, fig. 5;
<bibRefCitation id="812E0F2D0AF2094480E15945C006A309" author="Tarruella, A" journalOrPublisher="Spira" pageId="0" pageNumber="1" pagination="71 - 76" refId="B59" refString="Tarruella, A, Domenech, JL, 2011. Listado taxonomico ilustrado de la familia Cyclophoridae J.E. Gray, 1847 (Mollusca: Gastropoda): Parte 2. La subfamilia Alycaeinae J.E. Gray, 1850. Spira 4: 71 - 76" title="Listado taxonomico ilustrado de la familia Cyclophoridae J. E. Gray, 1847 (Mollusca: Gastropoda): Parte 2. La subfamilia Alycaeinae J. E. Gray, 1850." volume="4" year="2011">
Tarruella and
<normalizedToken id="09097285EBA0E10768B17199C21EEC59" originalValue="Domènech">Domenech</normalizedToken>
2011
</bibRefCitation>
: 72.
</paragraph>
<paragraph id="3700A31C1ABDEA0EBCE27554D6AF1C14" pageId="0" pageNumber="1">
<taxonomicName id="31530A8AC74CF61D58C70917B7A56FDC" authorityName="Theobald" authorityYear="1870" class="Gastropoda" family="Cyclophoridae" genus="Alycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alycaeus (Charax) cucullatus" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="cucullatus" subGenus="Charax">Alycaeus (Charax) cucullatus</taxonomicName>
-
<bibRefCitation id="52B7023CA1F0A8622CCC4CC9D8BD571E" author="Kobelt, W" journalOrPublisher="Nachrichtsblatt der Deutschen Malakozoologischen Gesellschaft" pageId="0" pageNumber="1" pagination="137 - 152" refId="B28" refString="Kobelt, W, Moellendorff, OF von, 1897. Catalog der gegenwaertig lebend bekannten Pneumonopomen. Nachrichtsblatt der Deutschen Malakozoologischen Gesellschaft 29: 137 - 152" title="Catalog der gegenwaertig lebend bekannten Pneumonopomen." volume="29" year="1897">
Kobelt and
<normalizedToken id="D6CB85D421A40E7B1D782655699F5034" originalValue="Möllendorff">Moellendorff</normalizedToken>
1897
</bibRefCitation>
: 150.
</paragraph>
<paragraph id="E05FAA9F08B8DF5D566DFE9A99545A97" pageId="0" pageNumber="1">
<taxonomicName id="46D1A616BE367AEA32E7E6C971D18969" authorityName="Godwin-Austen" authorityYear="1914" class="Gastropoda" family="Cyclophoridae" genus="Alycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alycaeus davisi" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="davisi">Alycaeus davisi</taxonomicName>
Godwin-Austen, 1914: 408, pl. 148, fig. 9, 9a. Type locality: Siam and Shan State boundary. New synonym.
</paragraph>
<paragraph id="A2469B54EDC648646E73DA23CA8AD790" pageId="0" pageNumber="1">
<taxonomicName id="4C07B8CB890D0EDA708773CE6E3309E6" authorityName="Theobald" authorityYear="1870" class="Gastropoda" family="Cyclophoridae" genus="Alycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alycaeus (Dicharax) cucullatus" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="cucullatus" subGenus="Dicharax">Alycaeus (Dicharax) cucullatus</taxonomicName>
-
<bibRefCitation id="9528BB6B95231FF7BD074383A78A4B4A" author="Kobelt, W" journalOrPublisher="Collins, London" pageId="0" pageNumber="1" refId="B27" refString="Kobelt, W, 1902. Cyclophoridae. Das Tierreich, 662 pp." title="Cyclophoridae. Das Tierreich, 662 pp." year="1902">Kobelt 1902</bibRefCitation>
: 367, 368;
<bibRefCitation id="A68D626A2CFF17C585EF15BB4D14B4E5" author="Gude, GK" journalOrPublisher="Taylor and Francis, London" pageId="0" pageNumber="1" refId="B20" refString="Gude, GK, 1921. The Fauna of British India including Ceylon and Burma. Mollusca - III. Land Operculates. Taylor and Francis, London" title="The Fauna of British India including Ceylon and Burma. Mollusca - III. Land Operculates." year="1921">Gude 1921</bibRefCitation>
: 244, 245.
</paragraph>
<paragraph id="682EE7B9592B4B3B398485C83A6A22E4" pageId="0" pageNumber="1">
<taxonomicName id="CE4777DDAB386C8E6CE1A2FDFCB132F6" authorityName="Godwin-Austen" authorityYear="1914" class="Gastropoda" family="Cyclophoridae" genus="Alycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alycaeus (Chamalycaeus) davisi" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="davisi" subGenus="Chamalycaeus">Alycaeus (Chamalycaeus) davisi</taxonomicName>
-
<bibRefCitation id="640FB209AE5EE909E5F2BF36930F82D2" author="Gude, GK" journalOrPublisher="Taylor and Francis, London" pageId="0" pageNumber="1" refId="B20" refString="Gude, GK, 1921. The Fauna of British India including Ceylon and Burma. Mollusca - III. Land Operculates. Taylor and Francis, London" title="The Fauna of British India including Ceylon and Burma. Mollusca - III. Land Operculates." year="1921">Gude 1921</bibRefCitation>
: 226.
</paragraph>
<paragraph id="9750217D26D37E0E9F9F64E3086E72D1" pageId="0" pageNumber="1">
<taxonomicName id="AC6E73F64C56776D7BCC5307DEF86BAC" baseAuthorityName="Theobald" baseAuthorityYear="1870" class="Gastropoda" family="Cyclophoridae" genus="Dicharax" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Dicharax cucullatus" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="cucullatus">Dicharax cucullatus</taxonomicName>
-
<bibRefCitation id="8BA2C82120FD3611D0841F6140AC7DC2" DOI="https://doi.org/10.3897/zookeys.981.53583" author="Pall-Gergely, B" journalOrPublisher="ZooKeys" pageId="0" pageNumber="1" pagination="1 - 220" refId="B47" refString="Pall-Gergely, B, Sajan, S, Tripathy, B, Meng, K, Takahiro, A, Ablett, J, 2020b. Genus-level revision of the Alycaeidae (Gastropoda: Cyclophoroidea), with an annotated species catalogue. ZooKeys 981: 1 - 220, DOI: https://doi.org/10.3897/zookeys.981.53583" title="Genus-level revision of the Alycaeidae (Gastropoda: Cyclophoroidea), with an annotated species catalogue." url="https://doi.org/10.3897/zookeys.981.53583" volume="981" year="2020 b">
<normalizedToken id="D07C3E5B3753821EAD16D59301B36F5F" originalValue="Páll-Gergely">Pall-Gergely</normalizedToken>
et al. 2020b
</bibRefCitation>
: 57, 58.
</paragraph>
<paragraph id="2D272DC0C66D21DAC40E4A56C6801354" pageId="0" pageNumber="1">
<taxonomicName id="4784BBD13933AFECC201AE5C51DAE175" authorityName="Godwin-Austen" authorityYear="1914" class="Gastropoda" family="Cyclophoridae" genus="Dicharax" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Dicharax davisi" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="davisi">Dicharax davisi</taxonomicName>
-
<bibRefCitation id="3175A1458A156587FF5F2A3BB4BA7021" DOI="https://doi.org/10.3897/zookeys.981.53583" author="Pall-Gergely, B" journalOrPublisher="ZooKeys" pageId="0" pageNumber="1" pagination="1 - 220" refId="B47" refString="Pall-Gergely, B, Sajan, S, Tripathy, B, Meng, K, Takahiro, A, Ablett, J, 2020b. Genus-level revision of the Alycaeidae (Gastropoda: Cyclophoroidea), with an annotated species catalogue. ZooKeys 981: 1 - 220, DOI: https://doi.org/10.3897/zookeys.981.53583" title="Genus-level revision of the Alycaeidae (Gastropoda: Cyclophoroidea), with an annotated species catalogue." url="https://doi.org/10.3897/zookeys.981.53583" volume="981" year="2020 b">
<normalizedToken id="BE1468B90CD44D0D2759DB70BB9FC84B" originalValue="Páll-Gergely">Pall-Gergely</normalizedToken>
et al. 2020b
</bibRefCitation>
: 58.
</paragraph>
</subSubSection>
<subSubSection id="6F29DAA5B72C76966289CFF348FB5EF0" pageId="0" pageNumber="1" type="type material">
<paragraph id="BD024829113B9DE1ED1973456C13D052" pageId="0" pageNumber="1">Type material examined.</paragraph>
<paragraph id="3991DD6EC99CDFC44A2C602D691922F7" pageId="0" pageNumber="1">
<emphasis id="2E2596CCAF3A55795E849680542CA182" bold="true" italics="true" pageId="0" pageNumber="1">Syntypes</emphasis>
of
<taxonomicName id="C28355ED04812FA5B1C89F970178AC40" authorityName="Theobald" authorityYear="1870" class="Gastropoda" family="Cyclophoridae" genus="Alycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alycaeus cucullatus" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="cucullatus">
<emphasis id="84F1A037CF4886E3228E23FF2219A441" italics="true" pageId="0" pageNumber="1">Alycaeus cucullatus</emphasis>
</taxonomicName>
NHMUK 1888.12.4.951-952 from Shan States (2 shells; Fig.
<figureCitation id="0898E06D1F032CB141A29AE242C2071B" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Dicharax cucullatus A. Syntype of Alycaeus cucullatus, NHMUK 1888.12.4.951; B. Syntype of Alycaeus davisi, NHMUK 1903.7.1.1630; C, D. Specimens CUMZ 5071 from Tham Ban Luang, Doi Ang Khang, Chiang Mai Province. Close-up images of the aperture not to scale. Photo: NHM (A, B.)." figureDoi="10.3897/zse.97.59143.figure2" httpUri="https://binary.pensoft.net/fig/494366" pageId="0" pageNumber="1">2A</figureCitation>
).
<emphasis id="D06EC49531FF30A0BC810F55591890F1" bold="true" italics="true" pageId="0" pageNumber="1">Syntypes</emphasis>
of
<taxonomicName id="50E633C6FA67EF5396060199792E8BAB" authorityName="Godwin-Austen" authorityYear="1914" class="Gastropoda" family="Cyclophoridae" genus="Alycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alycaeus davisi" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="davisi">
<emphasis id="8D5874332A8378603862C9511B3EFF9B" italics="true" pageId="0" pageNumber="1">Alycaeus davisi</emphasis>
</taxonomicName>
NHMUK 1903.7.1.1630 from Shan States (4 shells; Fig.
<figureCitation id="0609DD3503A6B7D410E6393B58C1BE42" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Dicharax cucullatus A. Syntype of Alycaeus cucullatus, NHMUK 1888.12.4.951; B. Syntype of Alycaeus davisi, NHMUK 1903.7.1.1630; C, D. Specimens CUMZ 5071 from Tham Ban Luang, Doi Ang Khang, Chiang Mai Province. Close-up images of the aperture not to scale. Photo: NHM (A, B.)." figureDoi="10.3897/zse.97.59143.figure2" httpUri="https://binary.pensoft.net/fig/494366" pageId="0" pageNumber="1">2B</figureCitation>
).
</paragraph>
<caption id="B702569F40B9846F039D62F4B91A89E1" doi="10.3897/zse.97.59143.figure2" httpUri="https://binary.pensoft.net/fig/494366" pageId="0" pageNumber="1" start="Figure 2" startId="F2">
<paragraph id="88E439E3CF3FE8638FCED8B6A66EEC68" pageId="0" pageNumber="1">
<emphasis id="B5E1369D70A65AB3DD3261B8C17E5DE3" bold="true" pageId="0" pageNumber="1">Figure 2.</emphasis>
<taxonomicName id="6D6381D243292BEA3C3F151BA3A83FE8" baseAuthorityName="Theobald" baseAuthorityYear="1870" class="Gastropoda" family="Cyclophoridae" genus="Dicharax" kingdom="Animalia" lsidName="Dicharax cucullatus" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="cucullatus">
<emphasis id="67B4EF0D0C29B51457A266F196F4E7E0" italics="true" pageId="0" pageNumber="1">Dicharax cucullatus</emphasis>
</taxonomicName>
<emphasis id="6098E8F46504FB5A4C77C40394EE0E88" bold="true" pageId="0" pageNumber="1">A.</emphasis>
Syntype of
<taxonomicName id="0AEBF67623E97C0AE37391BCBBC5CCDC" authorityName="Theobald" authorityYear="1870" class="Gastropoda" family="Cyclophoridae" genus="Alycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alycaeus cucullatus" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="cucullatus">
<emphasis id="7DB331D720987E348FA53A65BD39F25C" italics="true" pageId="0" pageNumber="1">Alycaeus cucullatus</emphasis>
</taxonomicName>
, NHMUK 1888.12.4.951;
<emphasis id="6D525C39ECD4FEAF0022F25666D7040F" bold="true" pageId="0" pageNumber="1">B.</emphasis>
Syntype of
<taxonomicName id="E2B055C0C1972D1DC323E80D1D969B35" authorityName="Godwin-Austen" authorityYear="1914" class="Gastropoda" family="Cyclophoridae" genus="Alycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alycaeus davisi" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="davisi">
<emphasis id="D8D12358D28FAE18E45397E88FDB6CB3" italics="true" pageId="0" pageNumber="1">Alycaeus davisi</emphasis>
</taxonomicName>
, NHMUK 1903.7.1.1630;
<emphasis id="2EDDA221297D909D3A0EB82D3A319D1B" bold="true" pageId="0" pageNumber="1">C, D.</emphasis>
Specimens CUMZ 5071 from Tham Ban Luang, Doi Ang Khang, Chiang Mai Province. Close-up images of the aperture not to scale. Photo: NHM (
<emphasis id="62187310ACACA7B60C6E509407450CA6" bold="true" pageId="0" pageNumber="1">A, B.</emphasis>
).
</paragraph>
</caption>
</subSubSection>
<subSubSection id="ACD3712FBA3649D428D2D27A183344BE" pageId="0" pageNumber="1" type="materials_examined">
<paragraph id="2E8ABCF3301683B272067766C5968499" pageId="0" pageNumber="1">Other material examined.</paragraph>
<paragraph id="96065A216162FB345D39C220FF0D0BD1" pageId="0" pageNumber="1">
<materialsCitation id="30EE335C6DD046384D9BA675DBCD3754" collectingDate="1997-05-24" collectingDateMax="2020-10-06" collectingDateMin="1997-05-24" collectorName="S. Panha, Panha, J. B, Burch, P, Panha, P, C. Sutcharit, P. Jirapatrasilp, Jirapatrasilp, R, Srisonchai, W, Siriwut, T, Seesamut, C, Jirapatrasilp, A, Pholyotha, W, Siriwut, N, Likhitrakarn, C" country="Thailand" county="Doi Chiang Dao" elevation="2000" latitude="18.585695" location="Tham Ban Luang" longLatPrecision="1" longitude="98.484276" municipality="Chiang Mai Province" specimenCount="6" stateProvince="Chiang Mai Province">
CUMZ 7421 from
<collectingCounty id="454E8CA7FB6EB5BF9CEB1DA59969ED41">Doi Chiang Dao</collectingCounty>
,
<collectingMunicipality id="95605EF746AE09DE61BB4AB5F5B90D8D">Chiang Mai Province</collectingMunicipality>
,
<collectingCountry id="4233ECF64127C3C1B7A40E01F77F5BFA" name="Thailand">Thailand</collectingCountry>
,
<geoCoordinate id="3D7CEAB7D0245BCBA0A2D8911505B88F" degrees="19" direction="north" minutes="14" orientation="latitude" precision="1" seconds="26.7" value="19.24075">19°14'26.7&quot;N</geoCoordinate>
,
<geoCoordinate id="426A1CBC5AA9D6C65944074336B8588A" degrees="98" direction="east" minutes="52" orientation="longitude" precision="1" seconds="49.8" value="98.8805">98°52'49.8&quot;E</geoCoordinate>
,
<collectingDate id="CF84E74333DC5AC039E0A34C9138FC74" value="1997-05-24" valueMax="1997-05-28" valueMin="1997-05-24">24-28 May 1997</collectingDate>
, coll.
<collectorName id="48C226E59DD5D61B8DE97FC624B7D3FB">
S.
<collectorName id="EFA412791C955C1098D2755B46B03C97">Panha, J.B</collectorName>
</collectorName>
.
<collectorName id="3BC358AA327D8E4B6066A247F222AA44">Burch, P</collectorName>
. Dumrongrojwattana (1 shell; Figs
<figureCitation id="BD28874E394A865E032AE36BD4C33B9A" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Dicharax cucullatus from Chiang Mai Province A. Specimen CUMZ 7421 from Doi Chiang Dao; B. Specimen CUMZ 5071 from Tham Ban Luang, Doi Ang Khang; C. Specimen CUMZ 7422 from Siribhum Waterfall, Doi Inthanon; D. Specimen CUMZ 7423 from Wachirathan Waterfall, Doi Inthanon; E. Specimen CUMZ 7424 from Doi Inthanon, 2000 m a. s. l. Close-up images of the aperture not to scale." figureDoi="10.3897/zse.97.59143.figure3" httpUri="https://binary.pensoft.net/fig/494367" pageId="0" pageNumber="1">3A</figureCitation>
,
<figureCitation id="F94A24F8DBEEF664C8487F7A9A0FD0B8" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Synoptic view of the eight Dicharax taxa in Thailand. A. Dicharax omissus, specimen CUMZ 7426 (gold plated); B. Dicharax pratatensis, specimen CUMZ 7427; C. Dicharax panhai sp. nov., paratype CUMZ 7429 / 2; D. Dicharax pongrati sp. nov., paratype CUMZ 7430 / 2; E. Dicharax notus, specimen CUMZ 7425; F. Dicharax burchi sp. nov., paratype CUMZ 7428 / 2; G. Dicharax borealis sp. nov., paratype CUMZ 5072 / 2; H. Dicharax cucullatus, specimen CUMZ 7421." figureDoi="10.3897/zse.97.59143.figure6" httpUri="https://binary.pensoft.net/fig/494370" pageId="0" pageNumber="1">6H</figureCitation>
). CUMZ 5071 from
<location id="4859EADB04A22A716843C5AF5FC51BF3" LSID="urn:lsid:plazi:treatment:E64839C4C6075B7A80C6684D5F22F4E4:4859EADB04A22A716843C5AF5FC51BF3" country="Thailand" county="Doi Chiang Dao" latitude="18.585695" longLatPrecision="1" longitude="98.484276" municipality="Chiang Mai Province" name="Tham Ban Luang" stateProvince="Chiang Mai Province">Tham Ban Luang</location>
,
<location id="B949A7E353522169549EF33674BF9269" LSID="urn:lsid:plazi:treatment:E64839C4C6075B7A80C6684D5F22F4E4:B949A7E353522169549EF33674BF9269" country="Thailand" county="Doi Chiang Dao" latitude="18.585695" longLatPrecision="1" longitude="98.484276" municipality="Chiang Mai Province" name="Doi Ang Khang" stateProvince="Chiang Mai Province">Doi Ang Khang</location>
,
<collectingRegion id="ED35DCD08827DE5F7B1E104E679EDD45" country="Thailand" name="Chiang Mai">Chiang Mai Province</collectingRegion>
,
<collectingCountry id="F8F99B4CFAB0813E8D9D1FC98667A46D" name="Thailand">Thailand</collectingCountry>
,
<geoCoordinate id="9AF236669E0ECA1AEA4CA844F3F187D5" degrees="19" direction="north" minutes="52" orientation="latitude" precision="1" seconds="13.7" value="19.870472">19°52'13.7&quot;N</geoCoordinate>
,
<geoCoordinate id="1749FB8D4E0A09EB686198B87A6DCA97" degrees="99" direction="east" minutes="02" orientation="longitude" precision="1" seconds="44.1" value="99.045586">99°02'44.1&quot;E</geoCoordinate>
,
<collectingDate id="94785E7D0E7AEA99537D0C82B9B3C677" value="2000-03-17">17 Mar. 2000</collectingDate>
, coll.
<collectorName id="E022509003AED616228AA39BF56A0631">
S.
<collectorName id="56B92F0F5564D0E9320D9F9E0796587C">Panha, P</collectorName>
</collectorName>
. Tongkerd (34 shells; Figs
<figureCitation id="4F4AD5101F3E33A7CBDE97F922CF9D1F" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Dicharax cucullatus A. Syntype of Alycaeus cucullatus, NHMUK 1888.12.4.951; B. Syntype of Alycaeus davisi, NHMUK 1903.7.1.1630; C, D. Specimens CUMZ 5071 from Tham Ban Luang, Doi Ang Khang, Chiang Mai Province. Close-up images of the aperture not to scale. Photo: NHM (A, B.)." figureDoi="10.3897/zse.97.59143.figure2" httpUri="https://binary.pensoft.net/fig/494366" pageId="0" pageNumber="1">2C, D</figureCitation>
,
<figureCitation id="DBDEDD2A8512DBC50580BD59C6BA2658" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Dicharax cucullatus from Chiang Mai Province A. Specimen CUMZ 7421 from Doi Chiang Dao; B. Specimen CUMZ 5071 from Tham Ban Luang, Doi Ang Khang; C. Specimen CUMZ 7422 from Siribhum Waterfall, Doi Inthanon; D. Specimen CUMZ 7423 from Wachirathan Waterfall, Doi Inthanon; E. Specimen CUMZ 7424 from Doi Inthanon, 2000 m a. s. l. Close-up images of the aperture not to scale." figureDoi="10.3897/zse.97.59143.figure3" httpUri="https://binary.pensoft.net/fig/494367" pageId="0" pageNumber="1">3B</figureCitation>
). CUMZ 7422 from
<location id="F54C3D2CB721F4CDC62866D7AF8D860A" LSID="urn:lsid:plazi:treatment:E64839C4C6075B7A80C6684D5F22F4E4:F54C3D2CB721F4CDC62866D7AF8D860A" country="Thailand" county="Doi Chiang Dao" latitude="18.585695" longLatPrecision="1" longitude="98.484276" municipality="Chiang Mai Province" name="Siribhum Waterfall" stateProvince="Chiang Mai Province">Siribhum Waterfall</location>
,
<location id="DC809DF3F63AFC6D589BCC3E205228D5" LSID="urn:lsid:plazi:treatment:E64839C4C6075B7A80C6684D5F22F4E4:DC809DF3F63AFC6D589BCC3E205228D5" country="Thailand" county="Doi Chiang Dao" latitude="18.585695" longLatPrecision="1" longitude="98.484276" municipality="Chiang Mai Province" name="Doi Inthanon" stateProvince="Chiang Mai Province">Doi Inthanon</location>
,
<collectingRegion id="33B781423CFD856D51CF0FDA36AF9D5C" country="Thailand" name="Chiang Mai">Chiang Mai Province</collectingRegion>
,
<collectingCountry id="EE74552747A19DCA0D8E257AE176CC31" name="Thailand">Thailand</collectingCountry>
,
<geoCoordinate id="F821116613EFEEB189A65C0BF2494CFC" degrees="18" direction="north" minutes="32" orientation="latitude" precision="1" seconds="48.5" value="18.546804">18°32'48.5&quot;N</geoCoordinate>
,
<geoCoordinate id="97B63DDC04FAA7A353E177814C2BFA34" degrees="98" direction="east" minutes="30" orientation="longitude" precision="1" seconds="44.0" value="98.51222">98°30'44.0&quot;E</geoCoordinate>
,
<collectingDate id="5628503EA90DFD87993C23A6F726F1D1" value="2009-10-13" valueMax="2009-10-15" valueMin="2009-10-13">13-15 Oct. 2009</collectingDate>
, coll.
<collectorName id="56028F75456ED1073C49F7745A6481AB">C. Sutcharit</collectorName>
(71 shells; Figs
<figureCitation id="7D8201E1C5AC2E3D6E4EFAAF4704702C" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Dicharax cucullatus from Chiang Mai Province A. Specimen CUMZ 7421 from Doi Chiang Dao; B. Specimen CUMZ 5071 from Tham Ban Luang, Doi Ang Khang; C. Specimen CUMZ 7422 from Siribhum Waterfall, Doi Inthanon; D. Specimen CUMZ 7423 from Wachirathan Waterfall, Doi Inthanon; E. Specimen CUMZ 7424 from Doi Inthanon, 2000 m a. s. l. Close-up images of the aperture not to scale." figureDoi="10.3897/zse.97.59143.figure3" httpUri="https://binary.pensoft.net/fig/494367" pageId="0" pageNumber="1">3C</figureCitation>
,
<figureCitation id="4B19E9941EE980DE375CAFE3C3C932CD" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. A, B. Dicharax cucullatus, specimens CUMZ 7422 from Siribhum Waterfall, Doi Inthanon, Chiang Mai Province; A. With operculum; C, D. Dicharax notus C. Syntype NHMUK 1903.7.1.3065; D. Specimen CUMZ 7425 from Tham Luang Sa Koen, Nan Province. Close-up images of the aperture not to scale. Photo: NHM (C.)." figureDoi="10.3897/zse.97.59143.figure4" httpUri="https://binary.pensoft.net/fig/494368" pageId="0" pageNumber="1">4A, B</figureCitation>
). CUMZ 7423 from
<location id="B2600674D13F60E53D8872EF833881AA" LSID="urn:lsid:plazi:treatment:E64839C4C6075B7A80C6684D5F22F4E4:B2600674D13F60E53D8872EF833881AA" country="Thailand" county="Doi Chiang Dao" latitude="18.585695" longLatPrecision="1" longitude="98.484276" municipality="Chiang Mai Province" name="Wachirathan Waterfall" stateProvince="Chiang Mai Province">Wachirathan Waterfall</location>
,
<location id="F773450A9508A7BE1B23F1E2F01ED6DE" LSID="urn:lsid:plazi:treatment:E64839C4C6075B7A80C6684D5F22F4E4:F773450A9508A7BE1B23F1E2F01ED6DE" country="Thailand" county="Doi Chiang Dao" latitude="18.585695" longLatPrecision="1" longitude="98.484276" municipality="Chiang Mai Province" name="Doi Inthanon" stateProvince="Chiang Mai Province">Doi Inthanon</location>
,
<collectingRegion id="048C5CFAA37644C6D6FC4BC1F4F36877" country="Thailand" name="Chiang Mai">Chiang Mai Province</collectingRegion>
,
<collectingCountry id="780FC8E29A46D9CF179B08A482591A78" name="Thailand">Thailand</collectingCountry>
,
<geoCoordinate id="9A4A59B02FD6CE6D9B4DB06B15B6186E" degrees="18" direction="north" minutes="32" orientation="latitude" precision="1" seconds="31.0" value="18.541945">18°32'31.0&quot;N</geoCoordinate>
,
<geoCoordinate id="FA9055F2582138D53BA0CC34D53855F5" degrees="98" direction="east" minutes="35" orientation="longitude" precision="1" seconds="53.8" value="98.59828">98°35'53.8&quot;E</geoCoordinate>
,
<collectingDate id="CB0884B3AB9910CC9A3E39F02CDEB792" value="2015-01-21">21 Jan. 2015</collectingDate>
, coll.
<collectorName id="D14248F1E8D3FDB82045756207D9487F">
P.
<collectorName id="C655BA9BF86E653F26D22DC6B7FFF80D">Jirapatrasilp, R</collectorName>
</collectorName>
.
<collectorName id="E93DEF8510B7253F4BDC96C29A1DFB26">Srisonchai, W</collectorName>
.
<collectorName id="C3D0ACFE6A45366EE423FAA544ED6093">Siriwut, T</collectorName>
.
<collectorName id="B93EBE9E63D27261E05283430ED4FFE6">Seesamut, C</collectorName>
. Sutcharit (10 shells; Fig.
<figureCitation id="47A80911AEA07E4733B6D5802674D317" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Dicharax cucullatus from Chiang Mai Province A. Specimen CUMZ 7421 from Doi Chiang Dao; B. Specimen CUMZ 5071 from Tham Ban Luang, Doi Ang Khang; C. Specimen CUMZ 7422 from Siribhum Waterfall, Doi Inthanon; D. Specimen CUMZ 7423 from Wachirathan Waterfall, Doi Inthanon; E. Specimen CUMZ 7424 from Doi Inthanon, 2000 m a. s. l. Close-up images of the aperture not to scale." figureDoi="10.3897/zse.97.59143.figure3" httpUri="https://binary.pensoft.net/fig/494367" pageId="0" pageNumber="1">3D</figureCitation>
). CUMZ 7424 from
<location id="2AABCADB7F62F1F3A58D1BA5FF00DCE4" LSID="urn:lsid:plazi:treatment:E64839C4C6075B7A80C6684D5F22F4E4:2AABCADB7F62F1F3A58D1BA5FF00DCE4" country="Thailand" county="Doi Chiang Dao" latitude="18.585695" longLatPrecision="1" longitude="98.484276" municipality="Chiang Mai Province" name="Doi Inthanon" stateProvince="Chiang Mai Province">Doi Inthanon</location>
,
<quantity id="AF3C24F058390ECAD80F09C138B5305A" metricMagnitude="3" metricUnit="m" metricValue="2.0" unit="m" value="2000.0">
<elevation id="50E9B7EA69E80396B5A552CF9C86CA02" metricMagnitude="3" metricUnit="m" metricValue="2.0" unit="m" value="2000.0">2,000 m</elevation>
</quantity>
a.s.l,
<collectingRegion id="4FD150FB6E0EAC46E436F68663A532CE" country="Thailand" name="Chiang Mai">Chiang Mai Province</collectingRegion>
,
<collectingCountry id="DA3CD007B5A292DEC10F4CE7585DE357" name="Thailand">Thailand</collectingCountry>
,
<geoCoordinate id="A2A37C934CED3E1CD51E6CA1E7B7C106" degrees="18" direction="north" minutes="35" orientation="latitude" precision="1" seconds="08.5" value="18.585695">18°35'08.5&quot;N</geoCoordinate>
,
<geoCoordinate id="E69183292583C7467FC0C5FE5EA618F7" degrees="98" direction="east" minutes="29" orientation="longitude" precision="1" seconds="03.4" value="98.484276">98°29'03.4&quot;E</geoCoordinate>
,
<collectingDate id="202CE88140CC07C09221B572FD09BD89" value="2015-01-21">21 Jan. 2015</collectingDate>
, coll.
<collectorName id="A043B1358E020623C323433CE30EC357">
P.
<collectorName id="D2DC2370F1D04CDC1BED95DD53EB464D">Jirapatrasilp, R</collectorName>
</collectorName>
.
<collectorName id="C3D33208CF0B28FBFF0E849BD00DD103">Srisonchai, W</collectorName>
.
<collectorName id="ABAC10114FBD1FE8FC47D507CB9137AB">Siriwut, T</collectorName>
.
<collectorName id="B65C7B37620E350C027DEBFD13C4A5FE">Seesamut, C</collectorName>
. Sutcharit (3 shells; Fig.
<figureCitation id="C5E9FF53AC8A5CE7216F60DCD17C5D08" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Dicharax cucullatus from Chiang Mai Province A. Specimen CUMZ 7421 from Doi Chiang Dao; B. Specimen CUMZ 5071 from Tham Ban Luang, Doi Ang Khang; C. Specimen CUMZ 7422 from Siribhum Waterfall, Doi Inthanon; D. Specimen CUMZ 7423 from Wachirathan Waterfall, Doi Inthanon; E. Specimen CUMZ 7424 from Doi Inthanon, 2000 m a. s. l. Close-up images of the aperture not to scale." figureDoi="10.3897/zse.97.59143.figure3" httpUri="https://binary.pensoft.net/fig/494367" pageId="0" pageNumber="1">3E</figureCitation>
). CUMZ 7432 from Doi Inthanon, ~
<quantity id="9D873D9C68E708EBD22AC8C98E708DA3" metricMagnitude="3" metricUnit="m" metricValue="1.85" metricValueMax="2.0" metricValueMin="1.7" unit="m" value="1850.0" valueMax="2000.0" valueMin="1700.0">
<elevation id="D69E531961B52F544088E3AFC026D21D" metricMagnitude="3" metricUnit="m" metricValue="1.85" metricValueMax="2.0" metricValueMin="1.7" unit="m" value="1850.0" valueMax="2000.0" valueMin="1700.0">1,700-2,000 m</elevation>
</quantity>
a.s.l,
<collectingRegion id="B2D2EA86B8F0EAFB116ED382951C7032" country="Thailand" name="Chiang Mai">Chiang Mai Province</collectingRegion>
,
<collectingCountry id="7AEDF67AC757BDE8396B9292FB954306" name="Thailand">Thailand</collectingCountry>
,
<geoCoordinate id="21A11E1A15E88A9056CF3819E50944C6" degrees="18" direction="north" minutes="35" orientation="latitude" precision="1" seconds="08.5" value="18.585695">18°35'08.5&quot;N</geoCoordinate>
,
<geoCoordinate id="EBBD81CD7AE7C3BCF77E463645C8803C" degrees="98" direction="east" minutes="29" orientation="longitude" precision="1" seconds="03.4" value="98.484276">98°29'03.4&quot;E</geoCoordinate>
,
<collectingDate id="7605F68F3033C6CE3A50F479EE7186C8" value="2020-10-05" valueMax="2020-10-06" valueMin="2020-10-05">5-6 Oct. 2020</collectingDate>
, coll.
<collectorName id="871CFFE7D2780D436410A9718453FB4D">
P.
<collectorName id="1CD4105EE546D4CB62D1D2A7FA8F998B">Jirapatrasilp, A</collectorName>
</collectorName>
.
<collectorName id="B5361581A7344428DADFD37FE2F03583">Pholyotha, W</collectorName>
.
<collectorName id="362A49C1058458E86BF9E179321023CB">Siriwut, N</collectorName>
.
<collectorName id="8D046665670E6B16681316BCD73CE7D3">Likhitrakarn, C</collectorName>
. Sutcharit (
<specimenCount id="E4AE768456192CB2EEB3BAD920EA0F60" type="generic">6 specimens</specimenCount>
in ethanol; Fig.
<figureCitation id="884B3929128ADA6459EBD4700E623700" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Living Dicharax specimens. A. Dicharax cucullatus, specimen CUMZ 7432 from Doi Inthanon, Chiang Mai Province; B. Dicharax notus, specimen CUMZ 7425 from Tham Luang Sa Koen, Nan Province. Both photographed in the laboratory. Not to scale. Photo: A. Pholyotha." figureDoi="10.3897/zse.97.59143.figure5" httpUri="https://binary.pensoft.net/fig/494369" pageId="0" pageNumber="1">5A</figureCitation>
)
</materialsCitation>
.
</paragraph>
<caption id="1DC9142BBEDC1B56BEFBEF1C69AAD5D3" doi="10.3897/zse.97.59143.figure3" httpUri="https://binary.pensoft.net/fig/494367" pageId="0" pageNumber="1" start="Figure 3" startId="F3">
<paragraph id="B1017BE612FA356216B27B9EA08D4B14" pageId="0" pageNumber="1">
<emphasis id="69B597369668E43B29A0966B52AC816A" bold="true" pageId="0" pageNumber="1">Figure 3.</emphasis>
<taxonomicName id="0D50597EAE3BF8D959955C3728CFD426" baseAuthorityName="Theobald" baseAuthorityYear="1870" class="Gastropoda" family="Cyclophoridae" genus="Dicharax" kingdom="Animalia" lsidName="Dicharax cucullatus" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="cucullatus">
<emphasis id="E635EFFD545A41CE4A926F1EDB2D488C" italics="true" pageId="0" pageNumber="1">Dicharax cucullatus</emphasis>
</taxonomicName>
from
<collectingRegion id="7D1E31E5A316840EA9E0C8CE91B2FDDD" country="Thailand" name="Chiang Mai">Chiang Mai Province</collectingRegion>
<emphasis id="2B60A44620548BA0480D635A4817C9DC" bold="true" pageId="0" pageNumber="1">A.</emphasis>
Specimen CUMZ 7421 from Doi Chiang Dao;
<emphasis id="813598FD2983B282177C1F2C85C8035E" bold="true" pageId="0" pageNumber="1">B.</emphasis>
Specimen CUMZ 5071 from Tham Ban Luang, Doi Ang Khang;
<emphasis id="E30C8958DCDB78823FC578D9B53A0C50" bold="true" pageId="0" pageNumber="1">C.</emphasis>
Specimen CUMZ 7422 from Siribhum Waterfall, Doi Inthanon;
<emphasis id="6BC0E00111AEFC832676132E29739E6D" bold="true" pageId="0" pageNumber="1">D.</emphasis>
Specimen CUMZ 7423 from Wachirathan Waterfall, Doi Inthanon;
<emphasis id="423A9CDC58A57CC144F760D111EFE46D" bold="true" pageId="0" pageNumber="1">E.</emphasis>
Specimen CUMZ 7424 from Doi Inthanon,
<quantity id="B65109081FAE58D911891E298AF7FB62" metricMagnitude="3" metricUnit="m" metricValue="2.0" unit="m" value="2000.0">2000 m</quantity>
a.s.l. Close-up images of the aperture not to scale.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="24B39AFB558F45162057CAABEED24A64" pageId="0" pageNumber="1" type="diagnosis">
<paragraph id="F235330CF6E2963F676D880DFBBF9F1E" pageId="0" pageNumber="1">Diagnosis.</paragraph>
<paragraph id="5296DB93B37E916CE8A35486800EAD69" pageId="0" pageNumber="1">Shell medium-sized (SH up to 3.5 mm, SW up to 5.5 mm), depressed-conical. Spire ca. 1/5 of shell height. R1 with regular, strong ribs; R2 longer than R3; R2 with 49-57 ribs. R3 with sharp swelling near aperture. Aperture round. Inner peristome with four to five indentations, always with parieto-columellar indentation, sometimes weak at basal. Outer peristome fringed, slightly reflected but not conspicuous. Umbilicus round. Operculum multispiral with elevated lamella.</paragraph>
</subSubSection>
<subSubSection id="B1538777C56384BA28F7027B652E9998" pageId="0" pageNumber="1" type="distribution">
<paragraph id="1C044AB2AE6D2E6F45FDE50AEE66DCA1" pageId="0" pageNumber="1">Distribution.</paragraph>
<paragraph id="395F3C273251AE1975092EC9EF4AABA8" pageId="0" pageNumber="1">
Shan State, Myanmar (
<bibRefCitation id="79F3A2E95754E3DBE1BBAE5D688FF7E1" author="Gude, GK" journalOrPublisher="Taylor and Francis, London" pageId="0" pageNumber="1" refId="B20" refString="Gude, GK, 1921. The Fauna of British India including Ceylon and Burma. Mollusca - III. Land Operculates. Taylor and Francis, London" title="The Fauna of British India including Ceylon and Burma. Mollusca - III. Land Operculates." year="1921">Gude 1921</bibRefCitation>
), and Chiang Mai Province, Thailand.
</paragraph>
</subSubSection>
<subSubSection id="D80E3C84ED92EF93E68FC500051DF80B" pageId="0" pageNumber="1" type="remarks">
<paragraph id="F88BD10892C45AA69A400E768B7B59F8" pageId="0" pageNumber="1">Remarks.</paragraph>
<paragraph id="426C58E726D0389A750AF508FA06CFE3" pageId="0" pageNumber="1">
This species was originally described from Shan State and diagnosed by its fringed peristome with five indentations (
<bibRefCitation id="8F79AFF02BA533C26121577BAE82AA8A" author="Theobald, W" journalOrPublisher="Journal of the Asiatic Society of Bengal" pageId="0" pageNumber="1" pagination="395 - 402" refId="B61" refString="Theobald, W, 1870. Descriptions of new species of land shells from the Shan States and Pegu. Journal of the Asiatic Society of Bengal 39: 395 - 402" title="Descriptions of new species of land shells from the Shan States and Pegu." volume="39" year="1870">Theobald 1870</bibRefCitation>
). Later,
<bibRefCitation id="6C582FE699230CADE1F27A27ABF4963F" author="Godwin-Austen, HH" journalOrPublisher="Journal of the Asiatic Society of Bengal" pageId="0" pageNumber="1" refId="B19" refString="Godwin-Austen, HH, 1914. Land and freshwater mollusca of India, including South Arabia, Baluchistan, Afghanistan, Kashmir, Nepal, Burma, Pegu, Tenasserim, Malaya Peninsula, Ceylon and other islands of the Indian Ocean. Vol. 2, Part 12: 311-442." title="Land and freshwater mollusca of India, including South Arabia, Baluchistan, Afghanistan, Kashmir, Nepal, Burma, Pegu, Tenasserim, Malaya Peninsula, Ceylon and other islands of the Indian Ocean. Vol. 2, Part 12: 311 - 442." year="1914">Godwin-Austen (1914)</bibRefCitation>
described
<taxonomicName id="6CFD3A4A155B48F17F58C8756A83D0AE" authorityName="Godwin-Austen" authorityYear="1914" class="Gastropoda" family="Cyclophoridae" genus="Alycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alycaeus davisi" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="davisi">
<emphasis id="7AC89493440E8C2B1DFDB79BCBEB5E7F" italics="true" pageId="0" pageNumber="1">Alycaeus davisi</emphasis>
</taxonomicName>
with a less fringed peristome with four indentations from Siam and Shan State boundary. Upon examining the type specimens, both species agree in having a depressed-conical shell shape, a long R2 and a sharp swelling of R3 near the aperture (Fig.
<figureCitation id="354606A92FDFD8DD612E754F0FD0773B" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Dicharax cucullatus A. Syntype of Alycaeus cucullatus, NHMUK 1888.12.4.951; B. Syntype of Alycaeus davisi, NHMUK 1903.7.1.1630; C, D. Specimens CUMZ 5071 from Tham Ban Luang, Doi Ang Khang, Chiang Mai Province. Close-up images of the aperture not to scale. Photo: NHM (A, B.)." figureDoi="10.3897/zse.97.59143.figure2" httpUri="https://binary.pensoft.net/fig/494366" pageId="0" pageNumber="1">2A, B</figureCitation>
). The types of
<taxonomicName id="6D43D3482D1796A4F5C57E5568071C32" lsidName="A. cucullatus" pageId="0" pageNumber="1" rank="species" species="cucullatus">
<emphasis id="3DA4325257BBD34B87A65B591952ABCD" italics="true" pageId="0" pageNumber="1">A. cucullatus</emphasis>
</taxonomicName>
differ from those of
<taxonomicName id="2DDA259826E6B03F79DB88CDDA75403D" lsidName="A. davisi" pageId="0" pageNumber="1" rank="species" species="davisi">
<emphasis id="2E19C559B90F2833BEC3408187AB2EE2" italics="true" pageId="0" pageNumber="1">A. davisi</emphasis>
</taxonomicName>
in having a more fringed peristome and more indentations. However, based on recently collected specimens, we conclude that this variation is to be interpreted as intraspecific as we found that this crenulation may vary even within the same population from nearly smooth (Fig.
<figureCitation id="7E68A852374A24E404EA442BF70D9369" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. A, B. Dicharax cucullatus, specimens CUMZ 7422 from Siribhum Waterfall, Doi Inthanon, Chiang Mai Province; A. With operculum; C, D. Dicharax notus C. Syntype NHMUK 1903.7.1.3065; D. Specimen CUMZ 7425 from Tham Luang Sa Koen, Nan Province. Close-up images of the aperture not to scale. Photo: NHM (C.)." figureDoi="10.3897/zse.97.59143.figure4" httpUri="https://binary.pensoft.net/fig/494368" pageId="0" pageNumber="1">4A</figureCitation>
) via slight (Fig.
<figureCitation id="B724A3A8C113C2BF8615B756E2CCB3FE" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. A, B. Dicharax cucullatus, specimens CUMZ 7422 from Siribhum Waterfall, Doi Inthanon, Chiang Mai Province; A. With operculum; C, D. Dicharax notus C. Syntype NHMUK 1903.7.1.3065; D. Specimen CUMZ 7425 from Tham Luang Sa Koen, Nan Province. Close-up images of the aperture not to scale. Photo: NHM (C.)." figureDoi="10.3897/zse.97.59143.figure4" httpUri="https://binary.pensoft.net/fig/494368" pageId="0" pageNumber="1">4B</figureCitation>
) to strong (Fig.
<figureCitation id="E2C62C5BCD20040EAA3512AD97394DFE" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Dicharax cucullatus A. Syntype of Alycaeus cucullatus, NHMUK 1888.12.4.951; B. Syntype of Alycaeus davisi, NHMUK 1903.7.1.1630; C, D. Specimens CUMZ 5071 from Tham Ban Luang, Doi Ang Khang, Chiang Mai Province. Close-up images of the aperture not to scale. Photo: NHM (A, B.)." figureDoi="10.3897/zse.97.59143.figure2" httpUri="https://binary.pensoft.net/fig/494366" pageId="0" pageNumber="1">2C, D</figureCitation>
). Because both species were described from the same geographical region,
<taxonomicName id="8726AC16ADB898A7ED1981EEF9E09FAF" authorityName="Godwin-Austen" authorityYear="1914" class="Gastropoda" family="Cyclophoridae" genus="Alycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alycaeus davisi" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="davisi">
<emphasis id="B85DC0802E2CAF8D2776216D3653FA1D" italics="true" pageId="0" pageNumber="1">Alycaeus davisi</emphasis>
</taxonomicName>
is herein regarded as a junior subjective synonym of
<taxonomicName id="0CA015B2E87D5F2DD15DB7A3FD1ACCD1" authorityName="Theobald" authorityYear="1870" class="Gastropoda" family="Cyclophoridae" genus="Alycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alycaeus cucullatus" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="cucullatus">
<emphasis id="F5751154B9DE7B1FD1039D224CCB45EF" italics="true" pageId="0" pageNumber="1">Alycaeus cucullatus</emphasis>
</taxonomicName>
. Refer to Discussion regarding the color of living specimen and variation in spiral striation.
</paragraph>
<caption id="1311E3AAA119AFEA782B55554ADCF1B0" doi="10.3897/zse.97.59143.figure4" httpUri="https://binary.pensoft.net/fig/494368" pageId="0" pageNumber="1" start="Figure 4" startId="F4">
<paragraph id="6367FEE033798B6F3518923765999876" pageId="0" pageNumber="1">
<emphasis id="B9A7B36687DC3878B4E9B77D5E8A6D0D" bold="true" pageId="0" pageNumber="1">Figure 4.</emphasis>
<emphasis id="94D431DD233A8B9390E113C5E2D7BDF6" bold="true" pageId="0" pageNumber="1">A, B.</emphasis>
<taxonomicName id="B3D08047E3E8226BCAA9CA230619F2EF" baseAuthorityName="Theobald" baseAuthorityYear="1870" class="Gastropoda" family="Cyclophoridae" genus="Dicharax" kingdom="Animalia" lsidName="Dicharax cucullatus" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="cucullatus">
<emphasis id="16E7E4C00B91C62617DCA369F5764F8D" italics="true" pageId="0" pageNumber="1">Dicharax cucullatus</emphasis>
</taxonomicName>
, specimens CUMZ 7422 from Siribhum Waterfall, Doi Inthanon, Chiang Mai Province;
<emphasis id="07583093DB9F1C15890E0D47070C7718" bold="true" pageId="0" pageNumber="1">A.</emphasis>
With operculum;
<emphasis id="738460E25EA8E7B51C0A8E2F9F33F7EC" bold="true" pageId="0" pageNumber="1">C, D.</emphasis>
<taxonomicName id="58E7EB9AB020D30E8F9F4F10B08E9138" baseAuthorityName="Godwin-Austen" baseAuthorityYear="1914" class="Gastropoda" family="Cyclophoridae" genus="Dicharax" kingdom="Animalia" lsidName="Dicharax notus" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="notus">
<emphasis id="01D83F705BB850F1C40563976503BD04" italics="true" pageId="0" pageNumber="1">Dicharax notus</emphasis>
</taxonomicName>
<emphasis id="284BE27DA64651776D97244CD085A918" bold="true" pageId="0" pageNumber="1">C.</emphasis>
Syntype NHMUK 1903.7.1.3065;
<emphasis id="88383ED87CDCD1994CA817D2968BE2E2" bold="true" pageId="0" pageNumber="1">D.</emphasis>
Specimen CUMZ 7425 from Tham Luang Sa Koen, Nan Province. Close-up images of the aperture not to scale. Photo: NHM (
<emphasis id="4C9195E1C697883731396142F5093E7E" bold="true" pageId="0" pageNumber="1">C.</emphasis>
).
</paragraph>
</caption>
<caption id="0DD7216DDCD797B0124FA8612520EA38" doi="10.3897/zse.97.59143.figure5" httpUri="https://binary.pensoft.net/fig/494369" pageId="0" pageNumber="1" start="Figure 5" startId="F5">
<paragraph id="072631760FD6F918BD47780BBA91D917" pageId="0" pageNumber="1">
<emphasis id="5ACCD732B10CACF5FC60FD8CF4ED0FEB" bold="true" pageId="0" pageNumber="1">Figure 5.</emphasis>
Living
<taxonomicName id="12B4D301492B90B035906A6358867471" authorityName="Kobelt &amp; Moellendorff" authorityYear="1900" class="Gastropoda" family="Cyclophoridae" genus="Dicharax" kingdom="Animalia" lsidName="Dicharax" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis id="64F4C308694C6EB81F5FEF804FED57BC" italics="true" pageId="0" pageNumber="1">Dicharax</emphasis>
</taxonomicName>
specimens.
<emphasis id="CD1C4BCB886B8F16572935A9EF8FF9A0" bold="true" pageId="0" pageNumber="1">A.</emphasis>
<taxonomicName id="DFDAD543BC45F7BE833583D843FF9ED3" baseAuthorityName="Theobald" baseAuthorityYear="1870" class="Gastropoda" family="Cyclophoridae" genus="Dicharax" kingdom="Animalia" lsidName="Dicharax cucullatus" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="cucullatus">
<emphasis id="7C646891230E44408F7327002D4386A6" italics="true" pageId="0" pageNumber="1">Dicharax cucullatus</emphasis>
</taxonomicName>
, specimen CUMZ 7432 from Doi Inthanon, Chiang Mai Province;
<emphasis id="0FC6EDC310B981E7B84D88510C2A5019" bold="true" pageId="0" pageNumber="1">B.</emphasis>
<taxonomicName id="E847DB286F908DEB1DE0E8F74B3E2E29" baseAuthorityName="Godwin-Austen" baseAuthorityYear="1914" class="Gastropoda" family="Cyclophoridae" genus="Dicharax" kingdom="Animalia" lsidName="Dicharax notus" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="notus">
<emphasis id="BBBDA133B25BF9DB0C5DD54CD11616E7" italics="true" pageId="0" pageNumber="1">Dicharax notus</emphasis>
</taxonomicName>
, specimen CUMZ 7425 from Tham Luang Sa Koen, Nan Province. Both photographed in the laboratory. Not to scale. Photo: A. Pholyotha.
</paragraph>
</caption>
<paragraph id="F8FDD78F0A10AD2C3224351D4A2004EE" pageId="0" pageNumber="1">
<taxonomicName id="ABCC4711527166FCCBE67F5CBC6BE09C" baseAuthorityName="Theobald" baseAuthorityYear="1870" class="Gastropoda" family="Cyclophoridae" genus="Dicharax" kingdom="Animalia" lsidName="Dicharax cucullatus" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="cucullatus">
<emphasis id="9F31ADB9705C7D2215FC4DA360AD64F7" italics="true" pageId="0" pageNumber="1">Dicharax cucullatus</emphasis>
</taxonomicName>
differs from other species with similar shell shape and fringed peristome, i.e.
<taxonomicName id="367B466CDA968AD4900FDC4672067B1F" lsidName="D. anthostoma" pageId="0" pageNumber="1" rank="species" species="anthostoma">
<emphasis id="2F0408971F20CA141C6A99536D3F8592" italics="true" pageId="0" pageNumber="1">D. anthostoma</emphasis>
</taxonomicName>
(
<normalizedToken id="75CD2F97BBA922CDC9571E293D5BB881" originalValue="Möllendorff">Moellendorff</normalizedToken>
, 1885) (
<bibRefCitation id="85AC9B275D3CD5085E5C1B13B523FC84" DOI="https://doi.org/10.11646/zootaxa.4331.1.1" author="Pall-Gergely, B" journalOrPublisher="Zootaxa" pageId="0" pageNumber="1" pagination="1 - 124" refId="B45" refString="Pall-Gergely, B, Hunyadi, A, Do, DS, Naggs, F, Asami, T, 2017. Revision of the Alycaeidae of China, Laos and Vietnam (Gastropoda: Cyclophoroidea) I: The genera Dicharax and Metalycaeus. Zootaxa 4331: 1 - 124, DOI: https://doi.org/10.11646/zootaxa.4331.1.1" title="Revision of the Alycaeidae of China, Laos and Vietnam (Gastropoda: Cyclophoroidea) I: The genera Dicharax and Metalycaeus." url="https://doi.org/10.11646/zootaxa.4331.1.1" volume="4331" year="2017">
<normalizedToken id="928CE1B147629BB5DD2CC58742908B72" originalValue="Páll-Gergely">Pall-Gergely</normalizedToken>
et al. 2017
</bibRefCitation>
: fig. 19a, b) from China,
<taxonomicName id="86A00D7E5BB8FA89CCE222ABB52230F6" lsidName="D. ataranensis" pageId="0" pageNumber="1" rank="species" species="ataranensis">
<emphasis id="6DA3B7700A34067DD8D40D5B47DA7650" italics="true" pageId="0" pageNumber="1">D. ataranensis</emphasis>
</taxonomicName>
(Godwin-Austen, 1914) (
<bibRefCitation id="068E377A2553DD30EAFC1C35DFC388A1" DOI="https://doi.org/10.3897/zookeys.981.53583" author="Pall-Gergely, B" journalOrPublisher="ZooKeys" pageId="0" pageNumber="1" pagination="1 - 220" refId="B47" refString="Pall-Gergely, B, Sajan, S, Tripathy, B, Meng, K, Takahiro, A, Ablett, J, 2020b. Genus-level revision of the Alycaeidae (Gastropoda: Cyclophoroidea), with an annotated species catalogue. ZooKeys 981: 1 - 220, DOI: https://doi.org/10.3897/zookeys.981.53583" title="Genus-level revision of the Alycaeidae (Gastropoda: Cyclophoroidea), with an annotated species catalogue." url="https://doi.org/10.3897/zookeys.981.53583" volume="981" year="2020 b">
<normalizedToken id="EFEB9E7A88CDD458211C24E520D6F30E" originalValue="Páll-Gergely">Pall-Gergely</normalizedToken>
et al. 2020b
</bibRefCitation>
: figs 11, 12) from Myanmar and
<taxonomicName id="6EFEB53B7BF8D3274ACE42B9E684B7E1" lsidName="D. fimbriatus" pageId="0" pageNumber="1" rank="species" species="fimbriatus">
<emphasis id="E24F9B6B2D7F65852BC9A27E3F5954A5" italics="true" pageId="0" pageNumber="1">D. fimbriatus</emphasis>
</taxonomicName>
(
<bibRefCitation id="F19ABA5845C43EC8239EE368033C7762" DOI="https://doi.org/10.11646/zootaxa.4331.1.1" author="Pall-Gergely, B" journalOrPublisher="Zootaxa" pageId="0" pageNumber="1" pagination="1 - 124" refId="B45" refString="Pall-Gergely, B, Hunyadi, A, Do, DS, Naggs, F, Asami, T, 2017. Revision of the Alycaeidae of China, Laos and Vietnam (Gastropoda: Cyclophoroidea) I: The genera Dicharax and Metalycaeus. Zootaxa 4331: 1 - 124, DOI: https://doi.org/10.11646/zootaxa.4331.1.1" title="Revision of the Alycaeidae of China, Laos and Vietnam (Gastropoda: Cyclophoroidea) I: The genera Dicharax and Metalycaeus." url="https://doi.org/10.11646/zootaxa.4331.1.1" volume="4331" year="2017">
<normalizedToken id="1B932450D79B6AC5EEFDB555B12EDE99" originalValue="Páll-Gergely">Pall-Gergely</normalizedToken>
et al. 2017
</bibRefCitation>
: figs 35-37) from China and Vietnam in possessing a sharp swelling nearer to the aperture. In addition,
<taxonomicName id="4ADE109D3E35A1A7EF646FAA68BBAB2E" lsidName="D. cucullatus" pageId="0" pageNumber="1" rank="species" species="cucullatus">
<emphasis id="DE0713F39E90335042054A08F72BE56A" italics="true" pageId="0" pageNumber="1">D. cucullatus</emphasis>
</taxonomicName>
differs from
<taxonomicName id="8D36B937D73CD83B8DE5E790E01CEE43" lsidName="D. anthostoma" pageId="0" pageNumber="1" rank="species" species="anthostoma">
<emphasis id="BA45C36946D8193AFA6FDA67C650C079" italics="true" pageId="0" pageNumber="1">D. anthostoma</emphasis>
</taxonomicName>
in having shorter distance between the inner and outer peristomes and from
<taxonomicName id="354259B0DFFC599A200A1F7CE0385DDF" lsidName="D. ataranensis" pageId="0" pageNumber="1" rank="species" species="ataranensis">
<emphasis id="E631A2FF5B9B16E13ACA523E37B20BC6" italics="true" pageId="0" pageNumber="1">D. ataranensis</emphasis>
</taxonomicName>
in possessing a shorter R3. Although
<taxonomicName id="91C01614348CEC03E67C02E93B34BEB7" lsidName="D. tangmaiensis" pageId="0" pageNumber="1" rank="species" species="tangmaiensis">
<emphasis id="4F528832411F37A0935A28BF7E5D3F70" italics="true" pageId="0" pageNumber="1">D. tangmaiensis</emphasis>
</taxonomicName>
(Chen &amp; Zhang, 2001) (
<bibRefCitation id="D36CC0E2CEA6548E24370AD7A84E88F8" DOI="https://doi.org/10.3897/zookeys.981.53583" author="Pall-Gergely, B" journalOrPublisher="ZooKeys" pageId="0" pageNumber="1" pagination="1 - 220" refId="B47" refString="Pall-Gergely, B, Sajan, S, Tripathy, B, Meng, K, Takahiro, A, Ablett, J, 2020b. Genus-level revision of the Alycaeidae (Gastropoda: Cyclophoroidea), with an annotated species catalogue. ZooKeys 981: 1 - 220, DOI: https://doi.org/10.3897/zookeys.981.53583" title="Genus-level revision of the Alycaeidae (Gastropoda: Cyclophoroidea), with an annotated species catalogue." url="https://doi.org/10.3897/zookeys.981.53583" volume="981" year="2020 b">
<normalizedToken id="6FD5CA19B4BA5BC6E4F280828340F984" originalValue="Páll-Gergely">Pall-Gergely</normalizedToken>
et al. 2020b
</bibRefCitation>
: fig. 14) from Tongmai Town, Bomi County, Tibet Autonomous Region, China is very similar to
<taxonomicName id="E0059982E50D9956AFD3AF65CBD5AD2E" lsidName="D. cucullatus" pageId="0" pageNumber="1" rank="species" species="cucullatus">
<emphasis id="9A21BF6F58C9BFD12CF286C986F8FEA9" italics="true" pageId="0" pageNumber="1">D. cucullatus</emphasis>
</taxonomicName>
, the distance between both
<normalizedToken id="57BD25B7FDE9AB6220CC082FA8DBD92A" originalValue="species’">species'</normalizedToken>
type localities is approximately 1,100 km. Thus, the status of
<taxonomicName id="0B0801638F8F3D11C56A15C1CFE0A3F0" lsidName="D. tangmaiensis" pageId="0" pageNumber="1" rank="species" species="tangmaiensis">
<emphasis id="28FB2256E5190A1479CB16B5540F9EF4" italics="true" pageId="0" pageNumber="1">D. tangmaiensis</emphasis>
</taxonomicName>
needs further scrutiny.
</paragraph>
<paragraph id="577A62707C29F426EE9CDF137D84C371" pageId="0" pageNumber="1">
Some specimens from Doi Inthanon possess a more depressed shell with a lower spire, a very low swelling of R3 and a nearly smooth inner peristome (Fig.
<figureCitation id="C9196C3F236DDA17E3368AB1DAAD4A4E" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. A, B. Dicharax cucullatus, specimens CUMZ 7422 from Siribhum Waterfall, Doi Inthanon, Chiang Mai Province; A. With operculum; C, D. Dicharax notus C. Syntype NHMUK 1903.7.1.3065; D. Specimen CUMZ 7425 from Tham Luang Sa Koen, Nan Province. Close-up images of the aperture not to scale. Photo: NHM (C.)." figureDoi="10.3897/zse.97.59143.figure4" httpUri="https://binary.pensoft.net/fig/494368" pageId="0" pageNumber="1">4A</figureCitation>
) very similar to
<taxonomicName id="A7EB14A22C7CD4C1F7DDE8C789EDDBEF" lsidName="D. borealis" pageId="0" pageNumber="1" rank="species" species="borealis">
<emphasis id="83552369F253114BF951FEF048934534" italics="true" pageId="0" pageNumber="1">D. borealis</emphasis>
</taxonomicName>
sp. nov. However,
<taxonomicName id="9437B6084749904EBCD738C076AA86AB" lsidName="D. cucullatus" pageId="0" pageNumber="1" rank="species" species="cucullatus">
<emphasis id="7024C7F90118F3ADF43E85A19AF495EB" italics="true" pageId="0" pageNumber="1">D. cucullatus</emphasis>
</taxonomicName>
could be distinguished from
<taxonomicName id="9FC4DC51420A99CC8B5CBCAA8DCA28A0" lsidName="D. borealis" pageId="0" pageNumber="1" rank="species" species="borealis">
<emphasis id="E8B97159CED5A6FDAD86DDAB38AA5227" italics="true" pageId="0" pageNumber="1">D. borealis</emphasis>
</taxonomicName>
sp. nov. by a round aperture and always having a parieto-columellar indentation.
</paragraph>
</subSubSection>
</treatment>
</document>
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