treatments-xml/data/BB/2F/1B/BB2F1B21FF87C915FF6D7E252C413DD1.xml

<document id="E21519BC4C521E739619E37D73FF05EE" ID-CLB-Dataset="32776" ID-DOI="10.11646/zootaxa.4282.3.3" ID-GBIF-Dataset="b62507f4-92ae-4f0c-ad05-4146c2915009" ID-ISSN="1175-5326" ID-Zenodo-Dep="827922" ID-ZooBank="45F9968A-02B2-46A5-91F1-E3E801C3ABB2" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1500107202169" checkinUser="plazi" docAuthor="Fathinia, Behzad, Rastegar-Pouyani, Eskandar, Rastegar-Pouyani, Nasrullah &amp; Darvishnia, Hamid" docDate="2017" docId="BB2F1B21FF87C915FF6D7E252C413DD1" docLanguage="en" docName="zootaxa.4282.3.3.pdf" docOrigin="Zootaxa 4282 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Rhynchocalamus ilamensis Fathinia, Rastegar-Pouyani, Rastegar-Pouyani &amp; Darvishnia, 2017, sp. nov." docType="treatment" docVersion="7" lastPageNumber="481" masterDocId="47166359FF83C91DFFFA7F3A29713C3E" masterDocTitle="A new species of the genus Rhynchocalamus Günther, 1864 (Reptilia: Squamata: Colubridae) from Ilam province in western Iran" masterLastPageNumber="486" masterPageNumber="473" pageNumber="477" updateTime="1698465004800" updateUser="ExternalLinkService">
<mods:mods id="AC1AA33638B8945E3F73C8681CD2447C" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="37511B20673F9721D7629BB9562D4A27">
<mods:title id="7D89982B8A088FA14447A6BF50B3E8D9">A new species of the genus Rhynchocalamus Günther, 1864 (Reptilia: Squamata: Colubridae) from Ilam province in western Iran</mods:title>
</mods:titleInfo>
<mods:name id="2F6B91D2F5C4BBE708795F8DC5637AF6" type="personal">
<mods:role id="BE858AED9A14BA7719C773EE546EAD49">
<mods:roleTerm id="C2B6B2C59D0A8BE4E6FDB8978EA6A3B8">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="58355707074E770089B5F18C2D2610B1">Fathinia, Behzad</mods:namePart>
</mods:name>
<mods:name id="C7B140075CA7AD39DB015BE30D1E1C90" type="personal">
<mods:role id="C621F833F2E7E603A1D0FC429FD4A771">
<mods:roleTerm id="DE588EB5578569655A0BF8D3A1528535">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="226B9C8A1A5F2B0A23D0B08E124DD0A0">Rastegar-Pouyani, Eskandar</mods:namePart>
</mods:name>
<mods:name id="177D32A6E269A866424FD4FEA43C57DD" type="personal">
<mods:role id="EA86DEE1C74835D4322586FF00AF71DA">
<mods:roleTerm id="612124C2892247BBC73B5A01974F4131">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="A5769E6566200E20F2852CDCE83386A5">Rastegar-Pouyani, Nasrullah</mods:namePart>
</mods:name>
<mods:name id="00D67917FB5F6A35C838FA419255AA5E" type="personal">
<mods:role id="D4912861D55DDEC41F301E44C863906B">
<mods:roleTerm id="A522F99B121E9D96AABC11A830382239">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="D7C1A942BD8A3D1DC00146C59BB03403">Darvishnia, Hamid</mods:namePart>
</mods:name>
<mods:typeOfResource id="D6E53AA75ECBF311109AA7527BAC926F">text</mods:typeOfResource>
<mods:relatedItem id="AE5519C083A069D184422D5C687291D1" type="host">
<mods:titleInfo id="D3C8FE1EF4FD67940394352C9CC26DD3">
<mods:title id="BD461D50A66987E7764070F8CB49257A">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="1AAAFA525FD938E76F35DA8D485544BC">
<mods:date id="041086B4981F073301A79A641BE03A2E">2017</mods:date>
<mods:detail id="33115999F41604A201EE9EBB32656C75" type="volume">
<mods:number id="690277CF651EBF1285A088AE8D154082">4282</mods:number>
</mods:detail>
<mods:detail id="78A88790EFA6CDF8CE74A86B0451A6C4" type="issue">
<mods:number id="061A51A7B8C1767EEBB5FCFAE98D92DD">3</mods:number>
</mods:detail>
<mods:extent id="F3B6211B180A4751466A8F3297F8F681" unit="page">
<mods:start id="9F65E9971163D675FC760AD4CD9C7582">473</mods:start>
<mods:end id="879F1C18C83DFCBB67452BEB78B80E49">486</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="ABDBB8BF348203B36AE271E67FA05F24">journal article</mods:classification>
<mods:identifier id="00657BBC9F4F3C3132778D0CEB59063B" type="CLB-Dataset">32776</mods:identifier>
<mods:identifier id="99C8FAA1340B5CC4CF9A2635C7B8D711" type="DOI">10.11646/zootaxa.4282.3.3</mods:identifier>
<mods:identifier id="C83626B2409DD8B2C9E7F3FBFDD445D2" type="GBIF-Dataset">b62507f4-92ae-4f0c-ad05-4146c2915009</mods:identifier>
<mods:identifier id="37B6EFEF95BE19610CD08BC267E475F3" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="4DEFB3E4D6AD252EC99D00486007CEC9" type="Zenodo-Dep">827922</mods:identifier>
<mods:identifier id="6AD9BA956DC6C82CD3FF2AA6157ED626" type="ZooBank">45F9968A-02B2-46A5-91F1-E3E801C3ABB2</mods:identifier>
</mods:mods>
<treatment id="BB2F1B21FF87C915FF6D7E252C413DD1" ID-DOI="http://doi.org/10.5281/zenodo.6003701" ID-GBIF-Taxon="132548389" ID-Zenodo-Dep="6003701" LSID="urn:lsid:plazi:treatment:BB2F1B21FF87C915FF6D7E252C413DD1" httpUri="http://treatment.plazi.org/id/BB2F1B21FF87C915FF6D7E252C413DD1" lastPageId="8" lastPageNumber="481" pageId="4" pageNumber="477">
<subSubSection id="7B9CF9BCFF87C919FF6D7E2528EB3D65" pageId="4" pageNumber="477" type="nomenclature">
<paragraph id="3339AA37FF87C919FF6D7E252B363D04" blockId="4.[151,583,287,347]" box="[151,583,287,314]" pageId="4" pageNumber="477">
<heading id="68711D5BFF87C919FF6D7E252B363D04" bold="true" box="[151,583,287,314]" fontSize="11" level="1" pageId="4" pageNumber="477" reason="1">
<emphasis id="01F27625FF87C919FF6D7E252B363D04" bold="true" box="[151,583,287,314]" pageId="4" pageNumber="477">
<taxonomicName id="F486D1B4FF87C919FF6D7E2528933D07" box="[151,482,287,313]" class="Reptilia" family="Colubridae" genus="Rhynchocalamus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="477" phylum="Chordata" rank="species" species="ilamensis" status="sp. nov.">
<emphasis id="01F27625FF87C919FF6D7E2528933D07" bold="true" box="[151,482,287,313]" italics="true" pageId="4" pageNumber="477">Rhynchocalamus ilamensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="1AC1CB5EFF87C919FE137E1A2B363D04" box="[489,583,288,314]" pageId="4" pageNumber="477" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="3339AA37FF87C919FF6D7E7828EB3D65" blockId="4.[151,583,287,347]" box="[151,410,322,347]" pageId="4" pageNumber="477">
(
<tableCitation id="7E049F8CFF87C919FF647E78286C3D65" box="[158,285,322,347]" captionStart-0="TABLE 1" captionStart-1="TABLE 2" captionStart-2="TABLE 3" captionStart-3="TABLE 4" captionStartId-0="2.[151,239,152,175]" captionStartId-1="3.[151,239,1403,1426]" captionStartId-2="4.[151,239,1048,1071]" captionStartId-3="7.[151,239,1755,1778]" captionText-0="TABLE 1. Pholidosis characters and morphometric measurements (in mm) of Rhynchocalamus ilamensis sp. nov. 1 — Sex; 2 — Loreals (left / right); 3 — Preoculars (left / right); 4 — Postoculars (left / right); 5 — Temporals (left / right); 6 — Posttemporals (left / right); 7 — Supralabials (left / right); 8 — Infralabials (left / right); 9 — Gular scales surrounding the last infralabials (left / right); 10 — Gular scales in a row between posterior inframaxillars; 11 — Dorsal plus temporal scales surrounding the posterior margin of the parietals; 12 — Ventrals; 13 — Longitudinal dorsal scale rows at mid-trunk (between ventrals no. 85 - 90); 14 — Subcaudals; 15 — Rostrum height; 16 — Rostrum width; 17 — Distance between nostrils; 18 — Diameter of eyes; 19 — Pileus length; 20 — Pileus width; 21 — Head length; 22 — Head width; 23 — Head height; 24 — Supraocular length; 25 — Frontal width; 26 — Frontal length; 27 — Anterior inframaxillar length; 28 — Posterior inframaxillar length; 29 — Snout-vent length; 30 — Tail length; 31 — Internasal triangular (d) or trapezoid shaped (t); 32 — Suture length of internasal much shorter (--), shorter (-), equal to (=) or longer than (+) prefrontal suture; 33 — Parietals shorter (-), equal to (=) or longer than (+) the distance from posterior tip of rostral to the posterior tip of frontal; 34 — pairs of infralabials in contact with anterior chin shield." captionText-1="TABLE 2. Definition of weighted multistate characters of the four species of Rhynchocalamus (R. arabicus, R. melanocephalus, R. satunini and R. ilamensis sp. nov.) used in the hierarchical cluster analysis. Abbreviations: CH., character; OBC, overall body color; DC, dorsum color; VC, ventral color; HBP, headband presence; NBP, neckband presence; NBS, neckband shape; NB. V, neckband in contact with ventrals; NB. HB, neckband in contact with headband; NBT, neckband thickness." captionText-2="TABLE 3. Specimens of Rhynchocalamus ilamensis sp. nov. included in this study with respective voucher reference, geographical distribution (latitude and longitude) and GenBank accession numbers." captionText-3="TABLE 4. Genetic distance matrix among the four species of Rhynchocalamus based on Kimura 2 - parameter (K 2 P) method in concatenated (below, grey cells) and Cytb (above, white cells) sequences." pageId="4" pageNumber="477">TableS 1–4</tableCitation>
; 
<figureCitation id="ABBDB6B2FF87C919FED37E7828E33D65" box="[297,402,322,347]" captionStart-0="FIGURE 2" captionStart-1="FIGURE 3" captionStart-2="FIGURE 4" captionStartId-0="4.[151,250,1976,1998]" captionStartId-1="5.[151,250,1249,1271]" captionStartId-2="6.[151,250,1004,1026]" captionTargetBox-0="[160,1421,1392,1940]" captionTargetBox-1="[190,1396,203,1228]" captionTargetBox-2="[172,1417,204,983]" captionTargetId-0="figure@4.[151,1436,1383,1955]" captionTargetId-1="figure@5.[150,1436,193,1228]" captionTargetId-2="figure@6.[151,1435,193,983]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionTargetPageId-2="6" captionText-0="FIGURE 2. Dorsal (a) and Ventral (b) Views of the preserVed holotype specimen of Rhynchocalamus ilamensis sp. nov. (YUZM-CRh. 1)." captionText-1="FIGURE 3. Dorsal head and neck View of Rhynchocalamus ilamensis sp. nov. (a) holotye (YUZM-CRh. 1); (b) paratype (YUZM-CRh. 2); (c) paratype (YUZM-CRh. 3)." captionText-2="FIGURE 4. Ventral head and neck View of Rhynchocalamus ilamensis sp. nov. (a) holotye (YUZM-CRh. 1); (b) paratype (YUZM-CRh. 2); (c) paratype (YUZM-CRh. 3)." httpUri-0="https://zenodo.org/record/827926/files/figure.png" httpUri-1="https://zenodo.org/record/827928/files/figure.png" httpUri-2="https://zenodo.org/record/827930/files/figure.png" pageId="4" pageNumber="477">FigS. 2–4</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="7B9CF9BCFF87C919FF6D7EBF2BB43E64" pageId="4" pageNumber="477" type="materials_examined">
<paragraph id="3339AA37FF87C919FF6D7EBF2A3F3DFA" blockId="4.[151,1436,389,1007]" pageId="4" pageNumber="477">
<materialsCitation id="83EEA06AFF87C919FF6D7EBF2A3F3DFA" ID-GBIF-Occurrence="1572324447" collectingDate="2011-06-06" country="Iran" elevation="724" latitude="33.640667" location="Bijar No-hunting area" longLatPrecision="1" longitude="46.038948" pageId="4" pageNumber="477" specimenCount="1" stateProvince="Ilam Province" typeStatus="holotype">
<emphasis id="01F27625FF87C919FF6D7EBF287C3DA0" bold="true" box="[151,269,389,414]" pageId="4" pageNumber="477">
<typeStatus id="EC3D1495FF87C919FF6D7EBF28793DA0" box="[151,264,389,414]" pageId="4" pageNumber="477" type="holotype">Holotype</typeStatus>
:
</emphasis>
YUZM-CRh.1: road-kill, adult male, collected at Bina &amp; 
<location id="3659FCECFF87C919FC137EBF2D813DA3" LSID="urn:lsid:plazi:treatment:BB2F1B21FF87C915FF6D7E252C413DD1:3659FCECFF87C919FC137EBF2D813DA3" box="[1001,1264,389,414]" country="Iran" latitude="33.640667" longLatPrecision="1" longitude="46.038948" name="Bijar No-hunting area" pageId="4" pageNumber="477" stateProvince="Ilam Province">Bijar No-hunting area</location>
(
<geoCoordinate id="56B2CCF0FF87C919FAF27EBF2CE93DA0" box="[1288,1432,389,414]" direction="north" orientation="latitude" pageId="4" pageNumber="477" precision="1" value="33.640667">33.640668N</geoCoordinate>
, 
<geoCoordinate id="56B2CCF0FF87C919FF6D7E9628533DFB" box="[151,290,428,453]" direction="east" orientation="longitude" pageId="4" pageNumber="477" precision="1" value="46.038948">46.038949E</geoCoordinate>
, 
<quantity id="F47E07D2FF87C919FED77E9628053DFA" box="[301,372,428,452]" metricMagnitude="2" metricUnit="m" metricValue="7.24" pageId="4" pageNumber="477" unit="m" value="724.0">
<elevation id="B8AB4D04FF87C919FED77E9628053DFA" box="[301,372,428,452]" metricMagnitude="2" metricUnit="m" metricValue="7.24" pageId="4" pageNumber="477" unit="m" value="724.0">724 m</elevation>
</quantity>
aSl), 
<collectingRegion id="F14264D5FF87C919FE4B7E962B233DFB" box="[433,594,428,453]" country="Iran" name="Ilam" pageId="4" pageNumber="477">Ilam Province</collectingRegion>
, 
<collectingCountry id="4B91EAA7FF87C919FDA77E972BFB3DFA" box="[605,650,429,452]" name="Iran" pageId="4" pageNumber="477">Iran</collectingCountry>
on 
<date id="47388CF7FF87C919FD4E7E972A383DFA" box="[692,841,424,453]" pageId="4" pageNumber="477" value="2011-06-06">
<collectingDate id="577C751FFF87C919FD4E7E972A383DFA" box="[692,841,424,453]" pageId="4" pageNumber="477" value="2011-06-06">June 6th 2011</collectingDate>
</date>
.
</materialsCitation>
</paragraph>
<paragraph id="3339AA37FF87C919FF3D7EEA2A2E3E31" blockId="4.[151,1436,389,1007]" pageId="4" pageNumber="477">
<materialsCitation id="83EEA06AFF87C919FF3D7EEA2A2E3E31" ID-GBIF-Occurrence="1572324446" collectingDate="2013-08-03" country="Iran" elevation="1383" latitude="32.948154" location="Abdanan" longLatPrecision="1" longitude="47.333237" pageId="4" pageNumber="477" specimenCount="1" stateProvince="Ilam Province" typeStatus="paratype">
<emphasis id="01F27625FF87C919FF3D7EEA28393DD7" bold="true" box="[199,328,464,489]" pageId="4" pageNumber="477">
<typeStatus id="EC3D1495FF87C919FF3D7EEA28323DD7" box="[199,323,464,489]" pageId="4" pageNumber="477" type="paratype">Paratypes</typeStatus>
:
</emphasis>
YUZM-CRh.2: adult female, collected at Dinaar-Kooh preServed area (
<geoCoordinate id="56B2CCF0FF87C919FB8B7EEA2C733DD7" box="[1137,1282,464,489]" direction="north" orientation="latitude" pageId="4" pageNumber="477" precision="1" value="32.948154">32.948156N</geoCoordinate>
, 
<geoCoordinate id="56B2CCF0FF87C919FAF77EEA2CE93DD7" box="[1293,1432,464,489]" direction="east" orientation="longitude" pageId="4" pageNumber="477" precision="1" value="47.333237">47.333236E</geoCoordinate>
, 
<quantity id="F47E07D2FF87C919FF6D7ECD299D3E31" box="[151,236,503,527]" metricMagnitude="3" metricUnit="m" metricValue="1.383" pageId="4" pageNumber="477" unit="m" value="1383.0">
<elevation id="B8AB4D04FF87C919FF6D7ECD299D3E31" box="[151,236,503,527]" metricMagnitude="3" metricUnit="m" metricValue="1.383" pageId="4" pageNumber="477" unit="m" value="1383.0">1383 m</elevation>
</quantity>
aSl), 
<location id="3659FCECFF87C919FED37ECC28E03E31" LSID="urn:lsid:plazi:treatment:BB2F1B21FF87C915FF6D7E252C413DD1:3659FCECFF87C919FED37ECC28E03E31" box="[297,401,502,527]" country="Iran" latitude="32.948154" longLatPrecision="1" longitude="47.333237" name="Abdanan" pageId="4" pageNumber="477" stateProvince="Ilam Province">Abdanan</location>
, 
<collectingRegion id="F14264D5FF87C919FE667ECC2B4F3E31" box="[412,574,502,527]" country="Iran" name="Ilam" pageId="4" pageNumber="477">Ilam Province</collectingRegion>
, 
<collectingCountry id="4B91EAA7FF87C919FDB27EC22B043E31" box="[584,629,504,527]" name="Iran" pageId="4" pageNumber="477">Iran</collectingCountry>
on 
<date id="47388CF7FF87C919FD657ECD2A2D3E31" box="[671,860,499,527]" pageId="4" pageNumber="477" value="2013-08-03">
<collectingDate id="577C751FFF87C919FD657ECD2A2D3E31" box="[671,860,499,527]" pageId="4" pageNumber="477" value="2013-08-03">AuguSt 3rd, 2013</collectingDate>
</date>
.
</materialsCitation>
</paragraph>
<paragraph id="3339AA37FF87C919FF3D7D202BB43E64" blockId="4.[151,1436,389,1007]" pageId="4" pageNumber="477">
<materialsCitation id="83EEA06AFF87C919FF3D7D202BB43E64" ID-GBIF-Occurrence="1572324445" collectingDate="2013-08-03" country="Iran" elevation="1180" latitude="32.95259" location="Abdanan" longLatPrecision="1" longitude="47.344166" pageId="4" pageNumber="477" specimenCount="1" stateProvince="Ilam Province">
YUZM-CRh.3: juvenile, collected at Dinaar-Kooh preServed area (
<geoCoordinate id="56B2CCF0FF87C919FC2B7D202D103E0D" box="[977,1121,538,563]" direction="north" orientation="latitude" pageId="4" pageNumber="477" precision="1" value="32.95259">32.952591N</geoCoordinate>
, 
<geoCoordinate id="56B2CCF0FF87C919FB8A7D202D8A3E0D" box="[1136,1275,538,563]" direction="east" orientation="longitude" pageId="4" pageNumber="477" precision="1" value="47.344166">47.344166E</geoCoordinate>
, 
<quantity id="F47E07D2FF87C919FAF07D212C133E0C" box="[1290,1378,539,563]" metricMagnitude="3" metricUnit="m" metricValue="1.18" pageId="4" pageNumber="477" unit="m" value="1180.0">
<elevation id="B8AB4D04FF87C919FAF07D212C133E0C" box="[1290,1378,539,563]" metricMagnitude="3" metricUnit="m" metricValue="1.18" pageId="4" pageNumber="477" unit="m" value="1180.0">1180 m</elevation>
</quantity>
aSl), 
<location id="3659FCECFF87C919FF6D7D7B298E3E64" LSID="urn:lsid:plazi:treatment:BB2F1B21FF87C915FF6D7E252C413DD1:3659FCECFF87C919FF6D7D7B298E3E64" box="[151,255,577,602]" country="Iran" latitude="32.95259" longLatPrecision="1" longitude="47.344166" name="Abdanan" pageId="4" pageNumber="477" stateProvince="Ilam Province">Abdanan</location>
, 
<collectingRegion id="F14264D5FF87C919FEF07D7B28DA3E64" box="[266,427,577,602]" country="Iran" name="Ilam" pageId="4" pageNumber="477">Ilam Province</collectingRegion>
, 
<collectingCountry id="4B91EAA7FF87C919FE4C7D7828923E67" box="[438,483,578,601]" name="Iran" pageId="4" pageNumber="477">Iran</collectingCountry>
on 
<date id="47388CF7FF87C919FDF77D782BB43E64" box="[525,709,574,602]" pageId="4" pageNumber="477" value="2013-08-03">
<collectingDate id="577C751FFF87C919FDF77D782BB43E64" box="[525,709,574,602]" pageId="4" pageNumber="477" value="2013-08-03">AuguSt 3rd, 2013</collectingDate>
</date>
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="7B9CF9BCFF87C918FF3D7D5F2A2A3BBC" lastPageId="5" lastPageNumber="478" pageId="4" pageNumber="477" type="description">
<paragraph id="3339AA37FF87C919FF3D7D5F2BAB3E40" blockId="4.[151,1436,389,1007]" box="[199,730,613,638]" pageId="4" pageNumber="477">
<emphasis id="01F27625FF87C919FF3D7D5F28AC3E40" bold="true" box="[199,477,613,638]" pageId="4" pageNumber="477">
Description of 
<typeStatus id="EC3D1495FF87C919FE8C7D5F28AC3E40" box="[374,477,613,638]" pageId="4" pageNumber="477" type="holotype">holotype</typeStatus>
</emphasis>
(
<figureCitation id="ABBDB6B2FF87C919FE167D5F2B4B3E40" box="[492,570,613,638]" captionStart="FIGURE 2" captionStartId="4.[151,250,1976,1998]" captionTargetBox="[160,1421,1392,1940]" captionTargetId="figure@4.[151,1436,1383,1955]" captionTargetPageId="4" captionText="FIGURE 2. Dorsal (a) and Ventral (b) Views of the preserVed holotype specimen of Rhynchocalamus ilamensis sp. nov. (YUZM-CRh. 1)." httpUri="https://zenodo.org/record/827926/files/figure.png" pageId="4" pageNumber="477">FigS. 2</figureCitation>
a &amp; b, 3a, 4a):
</paragraph>
<paragraph id="3339AA37FF87C919FF3D7DB3286B3FD1" blockId="4.[151,1436,389,1007]" pageId="4" pageNumber="477">
<emphasis id="01F27625FF87C919FF3D7DB328363E9C" bold="true" box="[199,327,649,674]" pageId="4" pageNumber="477">Pholidosis.</emphasis>
Elongate, Slender body; head Small, not diStinct from the neck; pupil round; roStral clearly offSet from adjacent ScaleS, wider than height, extending backward, wedging between the internaSalS; naSal undivided; internaSalS trapezoid in Shape; internaSal Suture Shorter than that between the prefrontalS; frontal length greater than width; SupraocularS narrow, narrower anteriorly than poSteriorly; parietalS large, longer than the frontal; right and left lorealS pentagonal in Shape; one preocular and one poStocular, much deeper than long; temporalS 1+1 on each Side; 7 SupralabialS on each Side (3rd and 4th in contact with eye); 9 infralabialS on each Side, the firSt pair in contact behind the mental; a large pair of anterior chin ShieldS, larger than poSterior oneS, in contact with the firSt four infralabialS; poSterior chin ShieldS Separated by one Scale; 13 dorSal ScaleS at level of the 10th ventral plate and 15 ScaleS at midbody and 15 ScaleS at level of the 10th plate before the cloaca; 211 ventralS; anal divided; 70 divided SubcaudalS.
</paragraph>
<caption id="67F9FABFFF87C919FF6D7B222D633871" pageId="4" pageNumber="477">
<paragraph id="3339AA37FF87C919FF6D7B222D633871" blockId="4.[151,1436,1047,1103]" pageId="4" pageNumber="477">
<emphasis id="01F27625FF87C919FF6D7B22287B3811" bold="true" box="[151,266,1048,1071]" pageId="4" pageNumber="477">TABLE 3.</emphasis>
Specimens of 
<taxonomicName id="F486D1B4FF87C919FE577B2D2BBE3811" box="[429,719,1047,1071]" class="Reptilia" family="Colubridae" genus="Rhynchocalamus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="477" phylum="Chordata" rank="species" species="ilamensis" status="sp. nov.">
<emphasis id="01F27625FF87C919FE577B2D2BBE3811" box="[429,719,1047,1071]" italics="true" pageId="4" pageNumber="477">Rhynchocalamus ilamensis</emphasis>
</taxonomicName>
<emphasis id="01F27625FF87C919FD227B222A413811" bold="true" box="[728,816,1048,1071]" pageId="4" pageNumber="477">
<taxonomicNameLabel id="1AC1CB5EFF87C919FD227B222A413811" box="[728,816,1048,1071]" pageId="4" pageNumber="477" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
included in this study with respective voucher reference, geographical distribution (latitude and longitude) and GenBank accession numbers.
</paragraph>
</caption>
<paragraph id="3339AA37FF87C919FF657B512C0238C3" pageId="4" pageNumber="477">
<table id="41865897FF8736E2FF657B512C0238C3" box="[159,1395,1131,1277]" gridcols="6" gridrows="4" pageId="4" pageNumber="477">
<tr id="8DB6A875FF8736E2FF657B512C0238BF" box="[159,1395,1131,1153]" gridrow="0" pageId="4" pageNumber="477">
<th id="CE67C109FF8736E2FF657B51281D38BF" box="[159,364,1131,1153]" gridcol="0" gridrow="0" pageId="4" pageNumber="477">Species</th>
<th id="CE67C109FF8736E2FE447B512B0738BF" box="[446,630,1131,1153]" gridcol="1" gridrow="0" pageId="4" pageNumber="477">Voucher reference</th>
<th id="CE67C109FF8736E2FD2A7B512A5E38BF" box="[720,815,1131,1153]" gridcol="2" gridrow="0" pageId="4" pageNumber="477">Latitude</th>
<th id="CE67C109FF8736E2FC6F7B512A8C38BF" box="[917,1021,1131,1153]" gridcol="3" gridrow="0" pageId="4" pageNumber="477">Longitude</th>
<th id="CE67C109FF8736E2FBA17B512DBA38BF" box="[1115,1227,1131,1153]" gridcol="4" gridrow="0" pageId="4" pageNumber="477">Cytb</th>
<th id="CE67C109FF8736E2FAF87B512C0238BF" box="[1282,1395,1131,1153]" gridcol="5" gridrow="0" pageId="4" pageNumber="477">C-mos</th>
</tr>
<tr id="8DB6A875FF8736E2FF657BAE2C023895" box="[159,1395,1172,1195]" gridrow="1" pageId="4" pageNumber="477">
<th id="CE67C109FF8736E2FF657BAE281D3895" box="[159,364,1172,1195]" gridcol="0" gridrow="1" pageId="4" pageNumber="477">
<taxonomicName id="F486D1B4FF87C919FF657BAC286A3894" box="[159,283,1172,1195]" class="Reptilia" family="Colubridae" genus="Rhynchocalamus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="477" phylum="Chordata" rank="species" species="ilamensis" status="sp. nov.">
<emphasis id="01F27625FF87C919FF657BAC286A3894" box="[159,283,1172,1195]" italics="true" pageId="4" pageNumber="477">R. ilamensis</emphasis>
</taxonomicName>
<emphasis id="01F27625FF87C919FEDB7BAF281D3894" bold="true" box="[289,364,1173,1194]" pageId="4" pageNumber="477">
<taxonomicNameLabel id="1AC1CB5EFF87C919FEDB7BAF281D3894" box="[289,364,1173,1194]" pageId="4" pageNumber="477" rank="species">sp.nov.</taxonomicNameLabel>
</emphasis>
</th>
<td id="CE67C109FF8736E2FE447BAE2B073895" box="[446,630,1172,1195]" gridcol="1" gridrow="1" pageId="4" pageNumber="477">YUZM-CRh.1</td>
<td id="CE67C109FF8736E2FD2A7BAE2A5E3895" box="[720,815,1172,1195]" gridcol="2" gridrow="1" pageId="4" pageNumber="477">33.64067</td>
<td id="CE67C109FF8736E2FC6F7BAE2A8C3895" box="[917,1021,1172,1195]" gridcol="3" gridrow="1" pageId="4" pageNumber="477">46.03895</td>
<td id="CE67C109FF8736E2FBA17BAE2DBA3895" box="[1115,1227,1172,1195]" gridcol="4" gridrow="1" pageId="4" pageNumber="477">
<accessionNumber id="2CD537D4FF87C919FBA17BAE2DBA3894" box="[1115,1227,1172,1194]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KY777448" pageId="4" pageNumber="477">KY777448</accessionNumber>
</td>
<td id="CE67C109FF8736E2FAF87BAE2C023895" box="[1282,1395,1172,1195]" gridcol="5" gridrow="1" pageId="4" pageNumber="477">-----</td>
</tr>
<tr id="8DB6A875FF8736E2FF657B842C0238EA" box="[159,1395,1214,1236]" gridrow="2" pageId="4" pageNumber="477">
<th id="CE67C109FF8736E2FF657B84281D38EA" box="[159,364,1214,1236]" gridcol="0" gridrow="2" pageId="4" pageNumber="477">
<taxonomicName id="F486D1B4FF87C919FF657B85286A38EA" box="[159,283,1214,1236]" class="Reptilia" family="Colubridae" genus="Rhynchocalamus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="477" phylum="Chordata" rank="species" species="ilamensis" status="sp. nov.">
<emphasis id="01F27625FF87C919FF657B85286A38EA" box="[159,283,1214,1236]" italics="true" pageId="4" pageNumber="477">R. ilamensis</emphasis>
</taxonomicName>
<emphasis id="01F27625FF87C919FEDB7B84281D38ED" bold="true" box="[289,364,1214,1235]" pageId="4" pageNumber="477">
<taxonomicNameLabel id="1AC1CB5EFF87C919FEDB7B84281D38ED" box="[289,364,1214,1235]" pageId="4" pageNumber="477" rank="species">sp.nov.</taxonomicNameLabel>
</emphasis>
</th>
<td id="CE67C109FF8736E2FE447B842B0738EA" box="[446,630,1214,1236]" gridcol="1" gridrow="2" pageId="4" pageNumber="477">YUZM-CRh.2</td>
<td id="CE67C109FF8736E2FD2A7B842A5E38EA" box="[720,815,1214,1236]" gridcol="2" gridrow="2" pageId="4" pageNumber="477">32.94816</td>
<td id="CE67C109FF8736E2FC6F7B842A8C38EA" box="[917,1021,1214,1236]" gridcol="3" gridrow="2" pageId="4" pageNumber="477">47.33324</td>
<td id="CE67C109FF8736E2FBA17B842DBA38EA" box="[1115,1227,1214,1236]" gridcol="4" gridrow="2" pageId="4" pageNumber="477">
<accessionNumber id="2CD537D4FF87C919FBA17B842DBA38EA" box="[1115,1227,1214,1236]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KY777450" pageId="4" pageNumber="477">KY777450</accessionNumber>
</td>
<td id="CE67C109FF8736E2FAF87B842C0238EA" box="[1282,1395,1214,1236]" gridcol="5" gridrow="2" pageId="4" pageNumber="477">
<accessionNumber id="2CD537D4FF87C919FAF87B842C0338EA" box="[1282,1394,1214,1236]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KY777447" pageId="4" pageNumber="477">KY777447</accessionNumber>
</td>
</tr>
<tr id="8DB6A875FF8736E2FF657BDD2C0238C3" box="[159,1395,1255,1277]" gridrow="3" pageId="4" pageNumber="477">
<th id="CE67C109FF8736E2FF657BDD281D38C3" box="[159,364,1255,1277]" gridcol="0" gridrow="3" pageId="4" pageNumber="477">
<taxonomicName id="F486D1B4FF87C919FF657BD2286A38C3" box="[159,283,1255,1277]" class="Reptilia" family="Colubridae" genus="Rhynchocalamus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="477" phylum="Chordata" rank="species" species="ilamensis" status="sp. nov.">
<emphasis id="01F27625FF87C919FF657BD2286A38C3" box="[159,283,1255,1277]" italics="true" pageId="4" pageNumber="477">R. ilamensis</emphasis>
</taxonomicName>
<emphasis id="01F27625FF87C919FEDB7BDD281D38C2" bold="true" box="[289,364,1255,1276]" pageId="4" pageNumber="477">
<taxonomicNameLabel id="1AC1CB5EFF87C919FEDB7BDD281D38C2" box="[289,364,1255,1276]" pageId="4" pageNumber="477" rank="species">sp.nov.</taxonomicNameLabel>
</emphasis>
</th>
<td id="CE67C109FF8736E2FE447BDD2B0738C3" box="[446,630,1255,1277]" gridcol="1" gridrow="3" pageId="4" pageNumber="477">YUZM-CRh.3</td>
<td id="CE67C109FF8736E2FD2A7BDD2A5E38C3" box="[720,815,1255,1277]" gridcol="2" gridrow="3" pageId="4" pageNumber="477">32.95259</td>
<td id="CE67C109FF8736E2FC6F7BDD2A8C38C3" box="[917,1021,1255,1277]" gridcol="3" gridrow="3" pageId="4" pageNumber="477">47.34417</td>
<td id="CE67C109FF8736E2FBA17BDD2DBA38C3" box="[1115,1227,1255,1277]" gridcol="4" gridrow="3" pageId="4" pageNumber="477">
<accessionNumber id="2CD537D4FF87C919FBA17BDD2DBA38C3" box="[1115,1227,1255,1277]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KY777449" pageId="4" pageNumber="477">KY777449</accessionNumber>
</td>
<td id="CE67C109FF8736E2FAF87BDD2C0238C3" box="[1282,1395,1255,1277]" gridcol="5" gridrow="3" pageId="4" pageNumber="477">
<accessionNumber id="2CD537D4FF87C919FAF87BDD2C0238C3" box="[1282,1395,1255,1277]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KY777446" pageId="4" pageNumber="477">KY777446</accessionNumber>
</td>
</tr>
</table>
</paragraph>
<caption id="67F9FABFFF87C919FF6D788228323BD3" httpUri="https://zenodo.org/record/827926/files/figure.png" pageId="4" pageNumber="477" targetBox="[160,1421,1392,1940]" targetPageId="4">
<paragraph id="3339AA37FF87C919FF6D788228323BD3" blockId="4.[151,1435,1976,2029]" pageId="4" pageNumber="477">
<emphasis id="01F27625FF87C919FF6D788228663BF3" bold="true" box="[151,279,1976,1998]" pageId="4" pageNumber="477">FIGURE 2.</emphasis>
Dorsal (a) and Ventral (b) Views of the preserVed holotype specimen of 
<taxonomicName id="F486D1B4FF87C919FBDF78822C4C3BF0" box="[1061,1341,1976,1998]" class="Reptilia" family="Colubridae" genus="Rhynchocalamus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="477" phylum="Chordata" rank="species" species="ilamensis" status="sp. nov.">
<emphasis id="01F27625FF87C919FBDF78822C4C3BF0" box="[1061,1341,1976,1998]" italics="true" pageId="4" pageNumber="477">Rhynchocalamus ilamensis</emphasis>
</taxonomicName>
<emphasis id="01F27625FF87C919FABD78822CEA3BF3" bold="true" box="[1351,1435,1976,1997]" pageId="4" pageNumber="477">
<taxonomicNameLabel id="1AC1CB5EFF87C919FABD78822CEA3BF3" box="[1351,1435,1976,1997]" pageId="4" pageNumber="477" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(YUZM-CRh.1).
</paragraph>
</caption>
<caption id="67F9FABFFF86C918FF6D7BDB2B073924" httpUri="https://zenodo.org/record/827928/files/figure.png" pageId="5" pageNumber="478" targetBox="[190,1396,203,1228]" targetPageId="5">
<paragraph id="3339AA37FF86C918FF6D7BDB2B073924" blockId="5.[151,1436,1249,1306]" pageId="5" pageNumber="478">
<emphasis id="01F27625FF86C918FF6D7BDB286738C8" bold="true" box="[151,278,1249,1271]" pageId="5" pageNumber="478">FIGURE 3.</emphasis>
Dorsal head and neck View of 
<taxonomicName id="F486D1B4FF86C918FD917BDB2AF338C9" box="[619,898,1249,1271]" class="Reptilia" family="Colubridae" genus="Rhynchocalamus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="478" phylum="Chordata" rank="species" species="ilamensis" status="sp. nov.">
<emphasis id="01F27625FF86C918FD917BDB2AF338C9" box="[619,898,1249,1271]" italics="true" pageId="5" pageNumber="478">Rhynchocalamus ilamensis</emphasis>
</taxonomicName>
<emphasis id="01F27625FF86C918FC767BD82A9138C9" bold="true" box="[908,992,1250,1271]" pageId="5" pageNumber="478">
<taxonomicNameLabel id="1AC1CB5EFF86C918FC767BD82A9138C9" box="[908,992,1250,1271]" pageId="5" pageNumber="478" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(a) holotye (YUZM-CRh.1); (b) paratype (YUZM-CRh.2); (c) paratype (YUZM-CRh.3).
</paragraph>
</caption>
<paragraph id="3339AA37FF86C918FF3D7A712B2039B9" blockId="5.[151,1437,1355,2030]" pageId="5" pageNumber="478">
<emphasis id="01F27625FF86C918FF3D7A712B77395A" bold="true" box="[199,518,1355,1380]" pageId="5" pageNumber="478">Morphometric characters.</emphasis>
Snout-vent length 
<quantity id="F47E07D2FF86C918FD1C7A712A3B395D" box="[742,842,1355,1380]" metricMagnitude="-1" metricUnit="m" metricValue="3.15" pageId="5" pageNumber="478" unit="mm" value="315.0">315 mm</quantity>
; tail length 
<quantity id="F47E07D2FF86C918FC2D7A712D3E395D" box="[983,1103,1355,1379]" metricMagnitude="-2" metricUnit="m" metricValue="8.020999999999999" pageId="5" pageNumber="478" unit="mm" value="80.21">80.21 mm</quantity>
; head length 
<quantity id="F47E07D2FF86C918FB147A712C28395D" box="[1262,1369,1355,1380]" metricMagnitude="-3" metricUnit="m" metricValue="9.45" pageId="5" pageNumber="478" unit="mm" value="9.45">9.45 mm</quantity>
; head width 
<quantity id="F47E07D2FF86C918FF277A54283539B9" box="[221,324,1390,1415]" metricMagnitude="-3" metricUnit="m" metricValue="5.72" pageId="5" pageNumber="478" unit="mm" value="5.72">5.72 mm</quantity>
; and head height 
<quantity id="F47E07D2FF86C918FDF27A552B3D39B9" box="[520,588,1391,1415]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="5" pageNumber="478" unit="mm" value="3.0">3 mm</quantity>
.
</paragraph>
<paragraph id="3339AA37FF86C918FF3D7AA928653A29" blockId="5.[151,1437,1355,2030]" pageId="5" pageNumber="478">
<emphasis id="01F27625FF86C918FF3D7AA928053992" bold="true" box="[199,372,1427,1452]" pageId="5" pageNumber="478">Color pattern.</emphasis>
Upper Surface of head uniformly lemon, with no darker markingS or bandS; a thin U-Shaped dark marking on neck, with a thickneSS equal that of two adjacent ScaleS; dorSal Surface of body and tail uniformly lemon; ventral SurfaceS from mental to tail tip uniformly white; color pattern paler in alcohol preServed than live SpecimenS.
</paragraph>
<paragraph id="3339AA37FF86C918FF3D79192D3E3AB8" blockId="5.[151,1437,1355,2030]" pageId="5" pageNumber="478">
<emphasis id="01F27625FF86C918FF3D79192BF43A02" bold="true" box="[199,645,1571,1596]" pageId="5" pageNumber="478">
Description of 
<typeStatus id="EC3D1495FF86C918FE86791928873A02" box="[380,502,1571,1596]" pageId="5" pageNumber="478" type="paratype">Paratypes</typeStatus>
. Pholidosis.
</emphasis>
<typeStatus id="EC3D1495FF86C918FD74791E2B813A02" box="[654,752,1572,1596]" pageId="5" pageNumber="478" type="paratype">Paratype</typeStatus>
SpecimenS are Similar to 
<typeStatus id="EC3D1495FF86C918FBE179192D0C3A02" box="[1051,1149,1571,1596]" pageId="5" pageNumber="478" type="holotype">holotype</typeStatus>
except for the following few variationS: 12 dorSal ScaleS at level of the 10th ventral plate in YUZM-CRh.3; 228 ventralS in YUZM-CRh.3 and 
<quantity id="F47E07D2FF86C918FF3C7954287C3AB8" box="[198,269,1645,1670]" metricMagnitude="0" metricUnit="m" metricValue="5.7658" pageId="5" pageNumber="478" unit="in" value="227.0">227 in</quantity>
YUZM-CRh.2; 64 SubcaudalS in YUZM-CRh.3 and 
<quantity id="F47E07D2FF86C918FC9979572AED3AB8" box="[867,924,1645,1670]" metricMagnitude="0" metricUnit="m" metricValue="1.5493999999999999" pageId="5" pageNumber="478" unit="in" value="61.0">61 in</quantity>
YUZM-CRh.2.
</paragraph>
<paragraph id="3339AA37FF86C918FF3D79AB2A333B28" blockId="5.[151,1437,1355,2030]" pageId="5" pageNumber="478">
<emphasis id="01F27625FF86C918FF3D79AB2B733A94" bold="true" box="[199,514,1681,1706]" pageId="5" pageNumber="478">Morphometric characters.</emphasis>
Snout-vent length 
<quantity id="F47E07D2FF86C918FD2D79AB2A423A97" box="[727,819,1681,1706]" metricMagnitude="-1" metricUnit="m" metricValue="1.63" pageId="5" pageNumber="478" unit="mm" value="163.0">163 mm</quantity>
for YUZM-CRh.3 and 
<quantity id="F47E07D2FF86C918FBC179AB2DE93A97" box="[1083,1176,1681,1706]" metricMagnitude="-1" metricUnit="m" metricValue="2.6" pageId="5" pageNumber="478" unit="mm" value="260.0">260 mm</quantity>
for YUZM-CRh.2; tail length 
<quantity id="F47E07D2FF86C918FF1E798F28383AF0" box="[228,329,1717,1742]" metricMagnitude="-2" metricUnit="m" metricValue="3.51" pageId="5" pageNumber="478" unit="mm" value="35.1">35.1 mm</quantity>
for YUZM-CRh.3 and 
<quantity id="F47E07D2FF86C918FDAE798F2BB63AF0" box="[596,711,1717,1742]" metricMagnitude="-2" metricUnit="m" metricValue="5.612" pageId="5" pageNumber="478" unit="mm" value="56.12">56.12 mm</quantity>
for YUZM-CRh.2; head length 
<quantity id="F47E07D2FF86C918FBCE798F2DE83AF0" box="[1076,1177,1717,1742]" metricMagnitude="-3" metricUnit="m" metricValue="6.33" pageId="5" pageNumber="478" unit="mm" value="6.33">6.33 mm</quantity>
for YUZM-CRh.3 and 
<quantity id="F47E07D2FF86C918FF6D79E3298A3ACF" box="[151,251,1753,1778]" metricMagnitude="-3" metricUnit="m" metricValue="7.15" pageId="5" pageNumber="478" unit="mm" value="7.15">7.15 mm</quantity>
for YUZM-CRh.2; head width 
<quantity id="F47E07D2FF86C918FD9879E02BB63ACF" box="[610,711,1754,1778]" metricMagnitude="-3" metricUnit="m" metricValue="3.73" pageId="5" pageNumber="478" unit="mm" value="3.73">3.73 mm</quantity>
for YUZM-CRh.3 and 
<quantity id="F47E07D2FF86C918FC2879E32D463ACF" box="[978,1079,1753,1778]" metricMagnitude="-3" metricUnit="m" metricValue="3.57" pageId="5" pageNumber="478" unit="mm" value="3.57">3.57 mm</quantity>
for YUZM-CRh.2; head height 
<quantity id="F47E07D2FF86C918FF6D79C7298A3B28" box="[151,251,1789,1814]" metricMagnitude="-3" metricUnit="m" metricValue="2.05" pageId="5" pageNumber="478" unit="mm" value="2.05">2.05 mm</quantity>
for YUZM-CRh.3 and 
<quantity id="F47E07D2FF86C918FDFE79C42B193B28" box="[516,616,1790,1814]" metricMagnitude="-3" metricUnit="m" metricValue="2.34" pageId="5" pageNumber="478" unit="mm" value="2.34">2.34 mm</quantity>
for YUZM-CRh.2.
</paragraph>
<paragraph id="3339AA37FF86C918FF3D781B2A2A3BBC" blockId="5.[151,1437,1355,2030]" pageId="5" pageNumber="478">
<emphasis id="01F27625FF86C918FF3D781B28063B04" bold="true" box="[199,375,1825,1850]" pageId="5" pageNumber="478">Color pattern.</emphasis>
In general, body color of the 
<typeStatus id="EC3D1495FF86C918FD1278182A253B04" box="[744,852,1826,1850]" pageId="5" pageNumber="478" type="paratype">paratypeS</typeStatus>
iS Similar to that of the 
<typeStatus id="EC3D1495FF86C918FB79781B2D993B04" box="[1155,1256,1825,1850]" pageId="5" pageNumber="478" type="holotype">holotype</typeStatus>
, except for the following variationS: a Small dark Spot between parietalS in YUZM-CRh.2; neck band not U-Shaped, divided into two Separate blotcheS on either SideS of neck in both 
<typeStatus id="EC3D1495FF86C918FD1D78502A273BBC" box="[743,854,1898,1922]" pageId="5" pageNumber="478" type="paratype">paratypeS</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection id="7B9CF9BCFF86C91BFF3D78B72C7739F0" lastPageId="6" lastPageNumber="479" pageId="5" pageNumber="478" type="etymology">
<paragraph id="3339AA37FF86C918FF3D78B72C533BD0" blockId="5.[151,1437,1355,2030]" pageId="5" pageNumber="478">
<emphasis id="01F27625FF86C918FF3D78B7283F3B98" bold="true" box="[199,334,1933,1958]" pageId="5" pageNumber="478">Etymology.</emphasis>
<taxonomicName id="F486D1B4FF86C918FEAF78B72BF73B98" box="[341,646,1933,1958]" class="Reptilia" family="Colubridae" genus="Rhynchocalamus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="478" phylum="Chordata" rank="species" species="ilamensis" status="sp. nov.">
<emphasis id="01F27625FF86C918FEAF78B72BF73B98" box="[341,646,1933,1958]" italics="true" pageId="5" pageNumber="478">Rhynchocalamus ilamensis</emphasis>
</taxonomicName>
<emphasis id="01F27625FF86C918FD7678B72B973B98" bold="true" box="[652,742,1933,1958]" pageId="5" pageNumber="478">
<taxonomicNameLabel id="1AC1CB5EFF86C918FD7678B72B973B98" box="[652,742,1933,1958]" pageId="5" pageNumber="478" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
iS named after 
<collectingRegion id="F14264D5FF86C918FC6F78B72D473B98" box="[917,1078,1933,1958]" country="Iran" name="Ilam" pageId="5" pageNumber="478">Ilam Province</collectingRegion>
, a Small province in the weSt of 
<collectingCountry id="4B91EAA7FF86C918FF6D788829B53BF7" box="[151,196,1970,1993]" name="Iran" pageId="5" pageNumber="478">Iran</collectingCountry>
having diverSe habitatS and ecoSyStemS Such 
<collectingCountry id="4B91EAA7FF86C918FD2A78882B963BF4" box="[720,743,1970,1994]" name="American Samoa" pageId="5" pageNumber="478">aS</collectingCountry>
high mountainS covered with oak foreStS, 
<collectingRegion id="F14264D5FF86C918FB2A78882C563BF4" box="[1232,1319,1970,1994]" country="American Samoa" name="Western" pageId="5" pageNumber="478">weStern</collectingRegion>
limeStone foothillS of the ZagroS MountainS, alluvial plainS and Sand duneS, Supporting a rich diverSity of reptileS.
</paragraph>
<caption id="67F9FABFFF85C91BFF6D7CD62B07381F" httpUri="https://zenodo.org/record/827930/files/figure.png" pageId="6" pageNumber="479" targetBox="[172,1417,204,983]" targetPageId="6">
<paragraph id="3339AA37FF85C91BFF6D7CD62B07381F" blockId="6.[151,1435,1004,1057]" pageId="6" pageNumber="479">
<emphasis id="01F27625FF85C91BFF6D7CD62867383F" bold="true" box="[151,278,1004,1026]" pageId="6" pageNumber="479">FIGURE 4.</emphasis>
Ventral head and neck View of 
<taxonomicName id="F486D1B4FF85C91BFD947CD62AF5383C" box="[622,900,1004,1026]" class="Reptilia" family="Colubridae" genus="Rhynchocalamus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="479" phylum="Chordata" rank="species" species="ilamensis" status="sp. nov.">
<emphasis id="01F27625FF85C91BFD947CD62AF5383C" box="[622,900,1004,1026]" italics="true" pageId="6" pageNumber="479">Rhynchocalamus ilamensis</emphasis>
</taxonomicName>
<emphasis id="01F27625FF85C91BFC747CD72A93383C" bold="true" box="[910,994,1005,1026]" pageId="6" pageNumber="479">
<taxonomicNameLabel id="1AC1CB5EFF85C91BFC747CD72A93383C" box="[910,994,1005,1026]" pageId="6" pageNumber="479" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(a) holotye (YUZM-CRh.1); (b) paratype (YUZM-CRh.2); (c) paratype (YUZM-CRh.3).
</paragraph>
</caption>
<paragraph id="3339AA37FF85C91BFF3D7B772C7739F0" blockId="6.[151,1437,1101,2026]" pageId="6" pageNumber="479">
<emphasis id="01F27625FF85C91BFF3D7B7728133858" bold="true" box="[199,354,1101,1126]" pageId="6" pageNumber="479">Comparison.</emphasis>
A number of meriStic and multiState characterS differentiate 
<taxonomicName id="F486D1B4FF85C91BFBF67B772C4D3858" box="[1036,1340,1101,1126]" class="Reptilia" family="Colubridae" genus="Rhynchocalamus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="479" phylum="Chordata" rank="species" species="ilamensis" status="sp. nov.">
<emphasis id="01F27625FF85C91BFBF67B772C4D3858" box="[1036,1340,1101,1126]" italics="true" pageId="6" pageNumber="479">Rhynchocalamus ilamensis</emphasis>
</taxonomicName>
<emphasis id="01F27625FF85C91BFAB97B772CED3858" bold="true" box="[1347,1436,1101,1126]" pageId="6" pageNumber="479">
<taxonomicNameLabel id="1AC1CB5EFF85C91BFAB97B772CED3858" box="[1347,1436,1101,1126]" pageId="6" pageNumber="479" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
from itS congenerS (
<taxonomicName id="F486D1B4FF85C91BFE827B49288838B4" box="[376,505,1137,1162]" class="Reptilia" family="Colubridae" genus="Rhynchocalamus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="479" phylum="Chordata" rank="species" species="arabicus">
<emphasis id="01F27625FF85C91BFE827B49288838B4" box="[376,505,1137,1162]" italics="true" pageId="6" pageNumber="479">R. arabicus</emphasis>
</taxonomicName>
, 
<taxonomicName id="F486D1B4FF85C91BFDFD7B492BAD38B4" box="[519,732,1137,1162]" class="Reptilia" family="Colubridae" genus="Rhynchocalamus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="479" phylum="Chordata" rank="species" species="melanocephalus">
<emphasis id="01F27625FF85C91BFDFD7B492BAD38B4" box="[519,732,1137,1162]" italics="true" pageId="6" pageNumber="479">R. melanocephalus</emphasis>
</taxonomicName>
and 
<taxonomicName id="F486D1B4FF85C91BFCE97B492AFD38B4" box="[787,908,1139,1162]" class="Reptilia" family="Colubridae" genus="Rhynchocalamus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="479" phylum="Chordata" rank="species" species="satunini">
<emphasis id="01F27625FF85C91BFCE97B492AFD38B4" box="[787,908,1139,1162]" italics="true" pageId="6" pageNumber="479">R. satunini</emphasis>
</taxonomicName>
): dorSum lemon in color (black in 
<taxonomicName id="F486D1B4FF85C91BFAEE7B492CE438B4" box="[1300,1429,1137,1162]" class="Reptilia" family="Colubridae" genus="Rhynchocalamus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="479" phylum="Chordata" rank="species" species="arabicus">
<emphasis id="01F27625FF85C91BFAEE7B492CE438B4" box="[1300,1429,1137,1162]" italics="true" pageId="6" pageNumber="479">R. arabicus</emphasis>
</taxonomicName>
, yellowiSh brown in 
<taxonomicName id="F486D1B4FF85C91BFE807BAD2B213890" box="[378,592,1173,1198]" class="Reptilia" family="Colubridae" genus="Rhynchocalamus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="479" phylum="Chordata" rank="species" species="melanocephalus">
<emphasis id="01F27625FF85C91BFE807BAD2B213890" box="[378,592,1173,1198]" italics="true" pageId="6" pageNumber="479">R. melanocephalus</emphasis>
</taxonomicName>
and pinkiSh in 
<taxonomicName id="F486D1B4FF85C91BFCFB7BAD2A0D3890" box="[769,892,1175,1198]" class="Reptilia" family="Colubridae" genus="Rhynchocalamus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="479" phylum="Chordata" rank="species" species="satunini">
<emphasis id="01F27625FF85C91BFCFB7BAD2A0D3890" box="[769,892,1175,1198]" italics="true" pageId="6" pageNumber="479">R. satunini</emphasis>
</taxonomicName>
); ventral white (black in 
<taxonomicName id="F486D1B4FF85C91BFB677BAD2C513890" box="[1181,1312,1173,1198]" class="Reptilia" family="Colubridae" genus="Rhynchocalamus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="479" phylum="Chordata" rank="species" species="arabicus">
<emphasis id="01F27625FF85C91BFB677BAD2C513890" box="[1181,1312,1173,1198]" italics="true" pageId="6" pageNumber="479">R. arabicus</emphasis>
</taxonomicName>
, yellowiSh white in 
<taxonomicName id="F486D1B4FF85C91BFEFB7B8128AB38EC" box="[257,474,1209,1234]" class="Reptilia" family="Colubridae" genus="Rhynchocalamus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="479" phylum="Chordata" rank="species" species="melanocephalus">
<emphasis id="01F27625FF85C91BFEFB7B8128AB38EC" box="[257,474,1209,1234]" italics="true" pageId="6" pageNumber="479">R. melanocephalus</emphasis>
</taxonomicName>
and pinkiSh in 
<taxonomicName id="F486D1B4FF85C91BFD6D7B812A6538EC" box="[663,788,1211,1234]" class="Reptilia" family="Colubridae" genus="Rhynchocalamus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="479" phylum="Chordata" rank="species" species="satunini">
<emphasis id="01F27625FF85C91BFD6D7B812A6538EC" box="[663,788,1211,1234]" italics="true" pageId="6" pageNumber="479">R. satunini</emphasis>
</taxonomicName>
); head uniformly lemon without marking(S) (uniformly black in 
<taxonomicName id="F486D1B4FF85C91BFF0D7BE5280638C8" box="[247,375,1245,1270]" class="Reptilia" family="Colubridae" genus="Rhynchocalamus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="479" phylum="Chordata" rank="species" species="arabicus">
<emphasis id="01F27625FF85C91BFF0D7BE5280638C8" box="[247,375,1245,1270]" italics="true" pageId="6" pageNumber="479">R. arabicus</emphasis>
</taxonomicName>
and 
<taxonomicName id="F486D1B4FF85C91BFE577BE52BF338C8" box="[429,642,1245,1270]" class="Reptilia" family="Colubridae" genus="Rhynchocalamus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="479" phylum="Chordata" rank="species" species="melanocephalus">
<emphasis id="01F27625FF85C91BFE577BE52BF338C8" box="[429,642,1245,1270]" italics="true" pageId="6" pageNumber="479">R. melanocephalus</emphasis>
</taxonomicName>
, and black tranSverS StripeS on parietalS and prefrontalS in 
<taxonomicName id="F486D1B4FF85C91BFAE87BE52CFA38C8" box="[1298,1419,1247,1270]" class="Reptilia" family="Colubridae" genus="Rhynchocalamus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="479" phylum="Chordata" rank="species" species="satunini">
<emphasis id="01F27625FF85C91BFAE87BE52CFA38C8" box="[1298,1419,1247,1270]" italics="true" pageId="6" pageNumber="479">R. satunini</emphasis>
</taxonomicName>
); dark neck marking 
<collectingCountry id="4B91EAA7FF85C91BFE737A3828D13924" box="[393,416,1282,1306]" name="American Samoa" pageId="6" pageNumber="479">aS</collectingCountry>
a thin U-Shape or two Separated blotcheS (uniformly black in 
<taxonomicName id="F486D1B4FF85C91BFB497A392C4C3924" box="[1203,1341,1281,1306]" class="Reptilia" family="Colubridae" genus="Rhynchocalamus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="479" phylum="Chordata" rank="species" species="arabicus">
<emphasis id="01F27625FF85C91BFB497A392C4C3924" box="[1203,1341,1281,1306]" italics="true" pageId="6" pageNumber="479">R. arabicus</emphasis>
</taxonomicName>
and 
<taxonomicName id="F486D1B4FF85C91BFA7E7A39283C3900" class="Reptilia" family="Colubridae" genus="Rhynchocalamus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="479" phylum="Chordata" rank="species" species="melanocephalus">
<emphasis id="01F27625FF85C91BFA7E7A39283C3900" italics="true" pageId="6" pageNumber="479">R. melanocephalus</emphasis>
</taxonomicName>
, a thick tranSverSe black marking in 
<taxonomicName id="F486D1B4FF85C91BFD137A1D2A133900" box="[745,866,1319,1342]" class="Reptilia" family="Colubridae" genus="Rhynchocalamus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="479" phylum="Chordata" rank="species" species="satunini">
<emphasis id="01F27625FF85C91BFD137A1D2A133900" box="[745,866,1319,1342]" italics="true" pageId="6" pageNumber="479">R. satunini</emphasis>
</taxonomicName>
); neck band not in contact with ventralS (unknown in 
<taxonomicName id="F486D1B4FF85C91BFF497A712845395C" box="[179,308,1353,1378]" class="Reptilia" family="Colubridae" genus="Rhynchocalamus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="479" phylum="Chordata" rank="species" species="arabicus">
<emphasis id="01F27625FF85C91BFF497A712845395C" box="[179,308,1353,1378]" italics="true" pageId="6" pageNumber="479">R. arabicus</emphasis>
</taxonomicName>
; in contact with ventralS in 
<taxonomicName id="F486D1B4FF85C91BFD917A712A31395C" box="[619,832,1353,1378]" class="Reptilia" family="Colubridae" genus="Rhynchocalamus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="479" phylum="Chordata" rank="species" species="melanocephalus">
<emphasis id="01F27625FF85C91BFD917A712A31395C" box="[619,832,1353,1378]" italics="true" pageId="6" pageNumber="479">R. melanocephalus</emphasis>
</taxonomicName>
, not in contact with ventralS in 
<taxonomicName id="F486D1B4FF85C91BFB5A7A712C68395C" box="[1184,1305,1355,1378]" class="Reptilia" family="Colubridae" genus="Rhynchocalamus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="479" phylum="Chordata" rank="species" species="satunini">
<emphasis id="01F27625FF85C91BFB5A7A712C68395C" box="[1184,1305,1355,1378]" italics="true" pageId="6" pageNumber="479">R. satunini</emphasis>
</taxonomicName>
); neckband not in contact with head or headband (unknown in 
<taxonomicName id="F486D1B4FF85C91BFD367A552A3C39B8" box="[716,845,1389,1414]" class="Reptilia" family="Colubridae" genus="Rhynchocalamus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="479" phylum="Chordata" rank="species" species="arabicus">
<emphasis id="01F27625FF85C91BFD367A552A3C39B8" box="[716,845,1389,1414]" italics="true" pageId="6" pageNumber="479">R. arabicus</emphasis>
</taxonomicName>
; in contact with headband in 
<taxonomicName id="F486D1B4FF85C91BFB6F7A552C1B39B8" box="[1173,1386,1389,1414]" class="Reptilia" family="Colubridae" genus="Rhynchocalamus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="479" phylum="Chordata" rank="species" species="melanocephalus">
<emphasis id="01F27625FF85C91BFB6F7A552C1B39B8" box="[1173,1386,1389,1414]" italics="true" pageId="6" pageNumber="479">R. melanocephalus</emphasis>
</taxonomicName>
; not in contact with headband in 
<taxonomicName id="F486D1B4FF85C91BFE2E7AA92B3C3994" box="[468,589,1427,1450]" class="Reptilia" family="Colubridae" genus="Rhynchocalamus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="479" phylum="Chordata" rank="species" species="satunini">
<emphasis id="01F27625FF85C91BFE2E7AA92B3C3994" box="[468,589,1427,1450]" italics="true" pageId="6" pageNumber="479">R. satunini</emphasis>
</taxonomicName>
); Seven SupralabialS (
<quantity id="F47E07D2FF85C91BFCB87AAB2AFB3994" box="[834,906,1425,1450]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" metricValueMax="1.524" metricValueMin="1.016" pageId="6" pageNumber="479" unit="in" value="5.0" valueMax="6.0" valueMin="4.0">4–6 in</quantity>
<taxonomicName id="F486D1B4FF85C91BFC6B7AA92D633994" box="[913,1042,1425,1450]" class="Reptilia" family="Colubridae" genus="Rhynchocalamus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="479" phylum="Chordata" rank="species" species="arabicus">
<emphasis id="01F27625FF85C91BFC6B7AA92D633994" box="[913,1042,1425,1450]" italics="true" pageId="6" pageNumber="479">R. arabicus</emphasis>
</taxonomicName>
, 
<quantity id="F47E07D2FF85C91BFBDA7AAB2D3A3994" box="[1056,1099,1425,1450]" metricMagnitude="-1" metricUnit="m" metricValue="1.524" pageId="6" pageNumber="479" unit="in" value="6.0">6 in</quantity>
<taxonomicName id="F486D1B4FF85C91BFBA87AA92C593994" box="[1106,1320,1425,1450]" class="Reptilia" family="Colubridae" genus="Rhynchocalamus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="479" phylum="Chordata" rank="species" species="melanocephalus">
<emphasis id="01F27625FF85C91BFBA87AA92C593994" box="[1106,1320,1425,1450]" italics="true" pageId="6" pageNumber="479">R. melanocephalus</emphasis>
</taxonomicName>
, 
<quantity id="F47E07D2FF85C91BFACF7AA82C0D3994" box="[1333,1404,1425,1450]" metricMagnitude="-1" metricUnit="m" metricValue="1.905" metricValueMax="2.032" metricValueMin="1.778" pageId="6" pageNumber="479" unit="in" value="7.5" valueMax="8.0" valueMin="7.0">7–8 in</quantity>
<taxonomicName id="F486D1B4FF85C91BFA7E7AA9298039F0" class="Reptilia" family="Colubridae" genus="Rhynchocalamus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="479" phylum="Chordata" rank="species" species="satunini">
<emphasis id="01F27625FF85C91BFA7E7AA9298039F0" italics="true" pageId="6" pageNumber="479">R. satunini</emphasis>
</taxonomicName>
); nine infralabialS (
<quantity id="F47E07D2FF85C91BFE357A8C288B39F0" box="[463,506,1461,1486]" metricMagnitude="-1" metricUnit="m" metricValue="2.032" pageId="6" pageNumber="479" unit="in" value="8.0">8 in</quantity>
<taxonomicName id="F486D1B4FF85C91BFDFA7A8D2BF039F0" box="[512,641,1461,1486]" class="Reptilia" family="Colubridae" genus="Rhynchocalamus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="479" phylum="Chordata" rank="species" species="arabicus">
<emphasis id="01F27625FF85C91BFDFA7A8D2BF039F0" box="[512,641,1461,1486]" italics="true" pageId="6" pageNumber="479">R. arabicus</emphasis>
</taxonomicName>
, moStly 
<quantity id="F47E07D2FF85C91BFD187A8D2A7C39F0" box="[738,781,1461,1486]" metricMagnitude="-1" metricUnit="m" metricValue="1.778" pageId="6" pageNumber="479" unit="in" value="7.0">7 in</quantity>
<taxonomicName id="F486D1B4FF85C91BFCEE7A8D2A9B39F0" box="[788,1002,1461,1486]" class="Reptilia" family="Colubridae" genus="Rhynchocalamus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="479" phylum="Chordata" rank="species" species="melanocephalus">
<emphasis id="01F27625FF85C91BFCEE7A8D2A9B39F0" box="[788,1002,1461,1486]" italics="true" pageId="6" pageNumber="479">R. melanocephalus</emphasis>
</taxonomicName>
, moStly 
<quantity id="F47E07D2FF85C91BFBB17A8C2D0739F0" box="[1099,1142,1461,1486]" metricMagnitude="-1" metricUnit="m" metricValue="2.032" pageId="6" pageNumber="479" unit="in" value="8.0">8 in</quantity>
<taxonomicName id="F486D1B4FF85C91BFB867A8D2D8739F0" box="[1148,1270,1463,1486]" class="Reptilia" family="Colubridae" genus="Rhynchocalamus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="479" phylum="Chordata" rank="species" species="satunini">
<emphasis id="01F27625FF85C91BFB867A8D2D8739F0" box="[1148,1270,1463,1486]" italics="true" pageId="6" pageNumber="479">R. satunini</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="7B9CF9BCFF85C915FF3D7AE32C413DD1" lastPageId="8" lastPageNumber="481" pageId="6" pageNumber="479" type="description">
<paragraph id="3339AA37FF85C91BFF3D7AE32C4F3A04" blockId="6.[151,1437,1101,2026]" pageId="6" pageNumber="479">
<emphasis id="01F27625FF85C91BFF3D7AE328FE39CC" bold="true" box="[199,399,1497,1522]" pageId="6" pageNumber="479">Cluster analysis.</emphasis>
ThiS analySiS baSed on 13 characterS (nine multiState and four meriStic) reSulted in a tree topography, with 
<taxonomicName id="F486D1B4FF85C91BFEA17AC528AD3A28" box="[347,476,1533,1558]" class="Reptilia" family="Colubridae" genus="Rhynchocalamus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="479" phylum="Chordata" rank="species" species="arabicus">
<emphasis id="01F27625FF85C91BFEA17AC528AD3A28" box="[347,476,1533,1558]" italics="true" pageId="6" pageNumber="479">R. arabicus</emphasis>
</taxonomicName>
at the baSe of the tree, SiSter to a clade compoSing of 
<taxonomicName id="F486D1B4FF85C91BFBCB7AC52C773A28" box="[1073,1286,1533,1558]" class="Reptilia" family="Colubridae" genus="Rhynchocalamus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="479" phylum="Chordata" rank="species" species="melanocephalus">
<emphasis id="01F27625FF85C91BFBCB7AC52C773A28" box="[1073,1286,1533,1558]" italics="true" pageId="6" pageNumber="479">R. melanocephalus</emphasis>
</taxonomicName>
, 
<taxonomicName id="F486D1B4FF85C91BFAE97AC52CED3A28" box="[1299,1436,1533,1558]" class="Reptilia" family="Colubridae" genus="Rhynchocalamus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="479" phylum="Chordata" rank="species" species="ilamensis" status="sp. nov.">
<emphasis id="01F27625FF85C91BFAE97AC52CED3A28" box="[1299,1436,1533,1558]" italics="true" pageId="6" pageNumber="479">R. ilamensis</emphasis>
</taxonomicName>
<emphasis id="01F27625FF85C91BFF6D791B29813A04" bold="true" box="[151,240,1569,1594]" pageId="6" pageNumber="479">
<taxonomicNameLabel id="1AC1CB5EFF85C91BFF6D791B29813A04" box="[151,240,1569,1594]" pageId="6" pageNumber="479" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and 
<taxonomicName id="F486D1B4FF85C91BFEDD791928D13A04" box="[295,416,1571,1594]" class="Reptilia" family="Colubridae" genus="Rhynchocalamus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="479" phylum="Chordata" rank="species" species="satunini">
<emphasis id="01F27625FF85C91BFEDD791928D13A04" box="[295,416,1571,1594]" italics="true" pageId="6" pageNumber="479">R. satunini</emphasis>
</taxonomicName>
. The tree topology ShowS that 
<taxonomicName id="F486D1B4FF85C91BFD0079192A023A04" box="[762,883,1571,1594]" class="Reptilia" family="Colubridae" genus="Rhynchocalamus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="479" phylum="Chordata" rank="species" species="satunini">
<emphasis id="01F27625FF85C91BFD0079192A023A04" box="[762,883,1571,1594]" italics="true" pageId="6" pageNumber="479">R. satunini</emphasis>
</taxonomicName>
iS SiSter to 
<taxonomicName id="F486D1B4FF85C91BFC0979192D0D3A04" box="[1011,1148,1569,1594]" class="Reptilia" family="Colubridae" genus="Rhynchocalamus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="479" phylum="Chordata" rank="species" species="ilamensis" status="sp. nov.">
<emphasis id="01F27625FF85C91BFC0979192D0D3A04" box="[1011,1148,1569,1594]" italics="true" pageId="6" pageNumber="479">R. ilamensis</emphasis>
</taxonomicName>
<emphasis id="01F27625FF85C91BFB79791B2DAC3A04" bold="true" box="[1155,1245,1569,1594]" pageId="6" pageNumber="479">
<taxonomicNameLabel id="1AC1CB5EFF85C91BFB79791B2DAC3A04" box="[1155,1245,1569,1594]" pageId="6" pageNumber="479" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="ABBDB6B2FF85C91BFB16791B2C413A04" box="[1260,1328,1569,1594]" captionStart="FIGURE 5" captionStartId="7.[151,250,1230,1252]" captionTargetBox="[297,1278,459,1201]" captionTargetId="figure@7.[278,1298,445,1215]" captionTargetPageId="7" captionText="FIGURE 5. Hierarchical cluster analysis of four species of Rhynchocalamus (R. arabicus, R. melanocephalus, R. satunini and R. ilamensis sp. nov.) based on multisate and meristic characters." httpUri="https://zenodo.org/record/827932/files/figure.png" pageId="6" pageNumber="479">Fig. 5</figureCitation>
).
</paragraph>
<paragraph id="3339AA37FF85C91BFF3D797F2DFE3AD0" blockId="6.[151,1437,1101,2026]" pageId="6" pageNumber="479">
<emphasis id="01F27625FF85C91BFF3D797F285B3A60" bold="true" box="[199,298,1605,1630]" pageId="6" pageNumber="479">Habitat.</emphasis>
<taxonomicName id="F486D1B4FF85C91BFECA797F2B113A60" box="[304,608,1605,1630]" class="Reptilia" family="Colubridae" genus="Rhynchocalamus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="479" phylum="Chordata" rank="species" species="ilamensis" status="sp. nov.">
<emphasis id="01F27625FF85C91BFECA797F2B113A60" box="[304,608,1605,1630]" italics="true" pageId="6" pageNumber="479">Rhynchocalamus ilamensis</emphasis>
</taxonomicName>
<emphasis id="01F27625FF85C91BFD9C797F2BCE3A60" bold="true" box="[614,703,1605,1630]" pageId="6" pageNumber="479">
<taxonomicNameLabel id="1AC1CB5EFF85C91BFD9C797F2BCE3A60" box="[614,703,1605,1630]" pageId="6" pageNumber="479" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
inhabitS the limeStone 
<collectingRegion id="F14264D5FF85C91BFC3B797C2D693A60" box="[961,1048,1606,1630]" country="American Samoa" name="Western" pageId="6" pageNumber="479">weStern</collectingRegion>
foothillS of the ZagroS MountainS. Habitat cover iS made of annual to perennial plantS Such 
<collectingCountry id="4B91EAA7FF85C91BFCB179502A133ABC" box="[843,866,1642,1666]" name="American Samoa" pageId="6" pageNumber="479">aS</collectingCountry>
<taxonomicName id="F486D1B4FF85C91BFC9579532A9B3ABC" box="[879,1002,1641,1666]" class="Magnoliopsida" family="Gramineae" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cyperales" pageId="6" pageNumber="479" phylum="Tracheophyta" rank="family">Gramineae</taxonomicName>
(annual graSSeS), buSheS and ShrubS (
<taxonomicName id="F486D1B4FF85C91BFF6479B728383A98" box="[158,329,1677,1702]" class="Magnoliopsida" family="Capparidaceae" higherTaxonomySource="GBIF,IPNI" kingdom="Plantae" order="Brassicales" pageId="6" pageNumber="479" phylum="Tracheophyta" rank="family">Capparidaceae</taxonomicName>
: 
<taxonomicName id="F486D1B4FF85C91BFEA179B42B563A98" box="[347,551,1678,1702]" class="Magnoliopsida" family="Capparaceae" genus="Capparis" kingdom="Plantae" order="Capparales" pageId="6" pageNumber="479" phylum="Tracheophyta" rank="species" species="spinosa">
<emphasis id="01F27625FF85C91BFEA179B42B563A98" box="[347,551,1678,1702]" italics="true" pageId="6" pageNumber="479">Capparis spinosa</emphasis>
</taxonomicName>
, 
<taxonomicName id="F486D1B4FF85C91BFDC779B72A7F3A98" box="[573,782,1677,1702]" class="Magnoliopsida" family="Cleomaceae" genus="Cleome" kingdom="Plantae" order="Capparales" pageId="6" pageNumber="479" phylum="Tracheophyta" rank="species" species="oxypetala">
<emphasis id="01F27625FF85C91BFDC779B72A7F3A98" box="[573,782,1677,1702]" italics="true" pageId="6" pageNumber="479">Cleome oxypetala</emphasis>
</taxonomicName>
; 
<taxonomicName id="F486D1B4FF85C91BFCDE79B72A943A98" box="[804,997,1677,1702]" class="Magnoliopsida" family="Caryophyllaceae" kingdom="Plantae" order="Caryophyllales" pageId="6" pageNumber="479" phylum="Tracheophyta" rank="family">Caryophyllaceae</taxonomicName>
: 
<taxonomicName id="F486D1B4FF85C91BFC0D79B72C783A98" box="[1015,1289,1677,1702]" class="Magnoliopsida" family="Caryophyllaceae" genus="Gypsophyla" higherTaxonomySource="GBIF" kingdom="Plantae" order="Caryophyllales" pageId="6" pageNumber="479" phylum="Tracheophyta" rank="species" species="linearifolia">
<emphasis id="01F27625FF85C91BFC0D79B72C783A98" box="[1015,1289,1677,1702]" italics="true" pageId="6" pageNumber="479">Gypsophyla linearifolia</emphasis>
</taxonomicName>
, 
<taxonomicName id="F486D1B4FF85C91BFAE479B42CE23A98" box="[1310,1427,1677,1702]" class="Magnoliopsida" family="Caryophyllaceae" genus="Gypsophyla" higherTaxonomySource="GBIF" kingdom="Plantae" order="Caryophyllales" pageId="6" pageNumber="479" phylum="Tracheophyta" rank="species" species="pallida">
<emphasis id="01F27625FF85C91BFAE479B42CE23A98" box="[1310,1427,1677,1702]" italics="true" pageId="6" pageNumber="479">G. pallida</emphasis>
</taxonomicName>
; 
<taxonomicName id="F486D1B4FF85C91BFF6D798B28243AF4" box="[151,341,1713,1738]" class="Insecta" family="Chenopodiaceae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="479" phylum="Arthropoda" rank="family">Chenopodiaceae</taxonomicName>
: 
<taxonomicName id="F486D1B4FF85C91BFE9E798B2B2F3AF4" box="[356,606,1713,1738]" class="Magnoliopsida" family="Amaranthaceae" genus="Halocharis" kingdom="Animalia" order="Caryophyllales" pageId="6" pageNumber="479" phylum="Tracheophyta" rank="species" species="sulphurea">
<emphasis id="01F27625FF85C91BFE9E798B2B2F3AF4" box="[356,606,1713,1738]" italics="true" pageId="6" pageNumber="479">Halocharis sulphurea</emphasis>
</taxonomicName>
, 
<taxonomicName id="F486D1B4FF85C91BFD8879892A4E3AF4" box="[626,831,1715,1738]" class="Magnoliopsida" family="Amaranthaceae" genus="Noaea" kingdom="Plantae" order="Caryophyllales" pageId="6" pageNumber="479" phylum="Tracheophyta" rank="species" species="mucronata">
<emphasis id="01F27625FF85C91BFD8879892A4E3AF4" box="[626,831,1715,1738]" italics="true" pageId="6" pageNumber="479">Noaea mucronata</emphasis>
</taxonomicName>
, 
<taxonomicName id="F486D1B4FF85C91BFCAE798B2D6E3AF4" box="[852,1055,1713,1738]" class="Magnoliopsida" family="Amaranthaceae" genus="Salsola" kingdom="Plantae" order="Caryophyllales" pageId="6" pageNumber="479" phylum="Tracheophyta" rank="species" species="imbricate">
<emphasis id="01F27625FF85C91BFCAE798B2D6E3AF4" box="[852,1055,1713,1738]" italics="true" pageId="6" pageNumber="479">Salsola imbricate</emphasis>
</taxonomicName>
; CompoSitae: 
<taxonomicName id="F486D1B4FF85C91BFB37798B2CE53AF4" box="[1229,1428,1713,1738]" class="Magnoliopsida" family="Asteraceae" genus="Achillea" kingdom="Plantae" order="Asterales" pageId="6" pageNumber="479" phylum="Tracheophyta" rank="species" species="conferta">
<emphasis id="01F27625FF85C91BFB37798B2CE53AF4" box="[1229,1428,1713,1738]" italics="true" pageId="6" pageNumber="479">Achillea conferta</emphasis>
</taxonomicName>
; 
<taxonomicName id="F486D1B4FF85C91BFF6D79EC28743AD0" box="[151,261,1750,1774]" class="Magnoliopsida" family="Rosaceae" kingdom="Plantae" order="Rosales" pageId="6" pageNumber="479" phylum="Tracheophyta" rank="family">RoSaceae</taxonomicName>
: 
<taxonomicName id="F486D1B4FF85C91BFEF579EF289B3AD0" box="[271,490,1749,1774]" class="Magnoliopsida" family="Rosaceae" genus="Amygdalus" higherTaxonomySource="CoL" kingdom="Plantae" order="Rosales" pageId="6" pageNumber="479" phylum="Tracheophyta" rank="species" species="arabica">
<emphasis id="01F27625FF85C91BFEF579EF289B3AD0" box="[271,490,1749,1774]" italics="true" pageId="6" pageNumber="479">Amygdalus arabica</emphasis>
</taxonomicName>
), and SparSe treeS (
<taxonomicName id="F486D1B4FF85C91BFD3979EC2A023AD0" box="[707,883,1749,1774]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" pageId="6" pageNumber="479" phylum="Tracheophyta" rank="species" species="brantii">
<emphasis id="01F27625FF85C91BFD3979EC2A023AD0" box="[707,883,1749,1774]" italics="true" pageId="6" pageNumber="479">Quercus brantii</emphasis>
</taxonomicName>
and 
<taxonomicName id="F486D1B4FF85C91BFC5079EF2D0E3AD0" box="[938,1151,1749,1774]" class="Magnoliopsida" family="Anacardiaceae" genus="Pistachia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Sapindales" pageId="6" pageNumber="479" phylum="Tracheophyta" rank="species" species="atlantica">
<emphasis id="01F27625FF85C91BFC5079EF2D0E3AD0" box="[938,1151,1749,1774]" italics="true" pageId="6" pageNumber="479">Pistachia atlantica</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="3339AA37FF85C91AFF3D79C328303CEA" blockId="6.[151,1437,1101,2026]" lastBlockId="7.[151,1437,151,392]" lastPageId="7" lastPageNumber="480" pageId="6" pageNumber="479">
<emphasis id="01F27625FF85C91BFF3D79C328DC3B2C" bold="true" box="[199,429,1785,1810]" pageId="6" pageNumber="479">Sympatric reptiles.</emphasis>
Some reptile SpecieS Sympatric to 
<taxonomicName id="F486D1B4FF85C91BFCB379C32DFB3B2C" box="[841,1162,1785,1810]" class="Magnoliopsida" family="Rosaceae" genus="Rhynchochalamus" kingdom="Plantae" order="Rosales" pageId="6" pageNumber="479" phylum="Tracheophyta" rank="species" species="ilamensis" status="sp. nov.">
<emphasis id="01F27625FF85C91BFCB379C32DFB3B2C" box="[841,1162,1785,1810]" italics="true" pageId="6" pageNumber="479">Rhynchochalamus ilamensis</emphasis>
</taxonomicName>
<emphasis id="01F27625FF85C91BFB6F79C32D833B2C" bold="true" box="[1173,1266,1785,1810]" pageId="6" pageNumber="479">
<taxonomicNameLabel id="1AC1CB5EFF85C91BFB6F79C32D833B2C" box="[1173,1266,1785,1810]" pageId="6" pageNumber="479" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
are 
<collectingCountry id="4B91EAA7FF85C91BFAD379C02C313B2C" box="[1321,1344,1786,1810]" name="American Samoa" pageId="6" pageNumber="479">aS</collectingCountry>
follow: 
<taxonomicName id="F486D1B4FF85C91BFF6D7827287E3B08" box="[151,271,1821,1846]" class="Reptilia" family="Agamidae" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="479" phylum="Chordata" rank="family">Agamidae</taxonomicName>
: 
<taxonomicName id="F486D1B4FF85C91BFEDB782728AA3B08" box="[289,475,1821,1846]" class="Reptilia" family="Agamidae" genus="Laudakia" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="479" phylum="Chordata" rank="species" species="nupta">
<emphasis id="01F27625FF85C91BFEDB782728AA3B08" box="[289,475,1821,1846]" italics="true" pageId="6" pageNumber="479">Laudakia nupta</emphasis>
</taxonomicName>
, 
<taxonomicName id="F486D1B4FF85C91BFE0B78272BA13B08" box="[497,720,1821,1846]" class="Reptilia" family="Agamidae" genus="Trapelus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="479" phylum="Chordata" rank="species" species="ruderatus">
<emphasis id="01F27625FF85C91BFE0B78272BA13B08" box="[497,720,1821,1846]" italics="true" pageId="6" pageNumber="479">Trapelus ruderatus</emphasis>
</taxonomicName>
; 
<taxonomicName id="F486D1B4FF85C91BFD1D78272A133B08" box="[743,866,1821,1846]" class="Reptilia" family="Lacertidae" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="479" phylum="Chordata" rank="family">Lacertidae</taxonomicName>
: 
<taxonomicName id="F486D1B4FF85C91BFC8E78272D323B08" box="[884,1091,1821,1846]" class="Reptilia" family="Lacertidae" genus="Ophisops" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="479" phylum="Chordata" rank="species" species="elegans">
<emphasis id="01F27625FF85C91BFC8E78272D323B08" box="[884,1091,1821,1846]" italics="true" pageId="6" pageNumber="479">Ophisops elegans</emphasis>
</taxonomicName>
, 
<taxonomicName id="F486D1B4FF85C91BFBA078272CE53B08" box="[1114,1428,1821,1846]" class="Reptilia" family="Lacertidae" genus="Acanthodactylus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="479" phylum="Chordata" rank="species" species="boskianus">
<emphasis id="01F27625FF85C91BFBA078272CE53B08" box="[1114,1428,1821,1846]" italics="true" pageId="6" pageNumber="479">Acanthodactylus boskianus</emphasis>
</taxonomicName>
; 
<taxonomicName id="F486D1B4FF85C91BFF6D787B284C3B64" box="[151,317,1857,1882]" class="Reptilia" family="Eublepharidae" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="479" phylum="Chordata" rank="family">Eublepharidae</taxonomicName>
: 
<taxonomicName id="F486D1B4FF85C91BFEB1787B2B023B64" box="[331,627,1857,1882]" class="Reptilia" family="Eublepharidae" genus="Eublepharis" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="479" phylum="Chordata" rank="species" species="angramainyu">
<emphasis id="01F27625FF85C91BFEB1787B2B023B64" box="[331,627,1857,1882]" italics="true" pageId="6" pageNumber="479">Eublepharis angramainyu</emphasis>
</taxonomicName>
; 
<taxonomicName id="F486D1B4FF85C91BFD7F787B2A603B64" box="[645,785,1857,1882]" class="Reptilia" family="Gekkonidae" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="479" phylum="Chordata" rank="family">Gekkonidae</taxonomicName>
: 
<taxonomicName id="F486D1B4FF85C91BFCE5787B2D513B64" box="[799,1056,1857,1882]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="479" phylum="Chordata" rank="species" species="persicus">
<emphasis id="01F27625FF85C91BFCE5787B2D513B64" box="[799,1056,1857,1882]" italics="true" pageId="6" pageNumber="479">Hemidactylus persicus</emphasis>
</taxonomicName>
, 
<taxonomicName id="F486D1B4FF85C91BFBC8787B2C1D3B64" box="[1074,1388,1857,1882]" class="Reptilia" family="Gekkonidae" genus="Mediodactylus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="479" phylum="Chordata" rank="species" species="heteropholis">
<emphasis id="01F27625FF85C91BFBC8787B2C1D3B64" box="[1074,1388,1857,1882]" italics="true" pageId="6" pageNumber="479">Mediodactylus heteropholis</emphasis>
</taxonomicName>
, 
<taxonomicName id="F486D1B4FF85C91BFA87787928703B40" class="Magnoliopsida" family="Rosaceae" genus="Rhynchochalamus" kingdom="Plantae" order="Rosales" pageId="6" pageNumber="479" phylum="Tracheophyta" rank="species" species="ilamensis">
<emphasis id="01F27625FF85C91BFA8778792CE53B64" box="[1405,1428,1859,1882]" italics="true" pageId="6" pageNumber="479">M</emphasis>
. 
<emphasis id="01F27625FF85C91BFF6D785F28703B40" box="[151,257,1893,1918]" italics="true" pageId="6" pageNumber="479">ilamensis</emphasis>
</taxonomicName>
; 
<taxonomicName id="F486D1B4FF85C91BFEEF785F28A73B40" box="[277,470,1893,1918]" class="Reptilia" family="Phyllodactylidae" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="479" phylum="Chordata" rank="family">Phyllodactylidae</taxonomicName>
: 
<taxonomicName id="F486D1B4FF85C91BFE1C785C2BE13B40" box="[486,656,1893,1918]" class="Reptilia" family="Phyllodactylidae" genus="Asaccus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="479" phylum="Chordata" rank="species" species="elisae">
<emphasis id="01F27625FF85C91BFE1C785C2BE13B40" box="[486,656,1893,1918]" italics="true" pageId="6" pageNumber="479">Asaccus elisae</emphasis>
</taxonomicName>
; 
<taxonomicName id="F486D1B4FF85C91BFD5E785F2A673B40" box="[676,790,1893,1918]" class="Reptilia" family="Scincidae" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="479" phylum="Chordata" rank="family">Scincidae</taxonomicName>
: 
<taxonomicName id="F486D1B4FF85C91BFCDC785F2D563B40" box="[806,1063,1893,1918]" class="Reptilia" family="Scincidae" genus="Ablepharus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="479" phylum="Chordata" rank="species" species="panonicus">
<emphasis id="01F27625FF85C91BFCDC785F2D563B40" box="[806,1063,1893,1918]" italics="true" pageId="6" pageNumber="479">Ablepharus panonicus</emphasis>
</taxonomicName>
, 
<taxonomicName id="F486D1B4FF85C91BFBC1785D2C7E3B40" box="[1083,1295,1895,1918]" class="Reptilia" family="Scincidae" genus="Heremites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="479" phylum="Chordata" rank="species" species="auratus">
<emphasis id="01F27625FF85C91BFBC1785D2C7E3B40" box="[1083,1295,1895,1918]" italics="true" pageId="6" pageNumber="479">Heremites auratus</emphasis>
</taxonomicName>
; 
<taxonomicName id="F486D1B4FF85C91BFADE785F2CE83B40" box="[1316,1433,1893,1918]" class="Reptilia" family="Varanidae" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="479" phylum="Chordata" rank="family">Varanidae</taxonomicName>
: 
<taxonomicName id="F486D1B4FF85C91BFF6D78B1281B3B9C" box="[151,362,1931,1954]" class="Reptilia" family="Varanidae" genus="Varanus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="479" phylum="Chordata" rank="species" species="nesterovi">
<emphasis id="01F27625FF85C91BFF6D78B1281B3B9C" box="[151,362,1931,1954]" italics="true" pageId="6" pageNumber="479">Varanus nesterovi</emphasis>
</taxonomicName>
; 
<taxonomicName id="F486D1B4FF85C91BFE7E78B32B653B9C" box="[388,532,1929,1954]" class="Reptilia" family="Typhlopidae" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="479" phylum="Chordata" rank="family">Typhlopidae</taxonomicName>
: 
<taxonomicName id="F486D1B4FF85C91BFDD078B32A2C3B9C" box="[554,861,1929,1954]" class="Reptilia" family="Typhlopidae" genus="Xerotyphlops" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="479" phylum="Chordata" rank="species" species="vermicularis">
<emphasis id="01F27625FF85C91BFDD078B32A2C3B9C" box="[554,861,1929,1954]" italics="true" pageId="6" pageNumber="479">Xerotyphlops vermicularis</emphasis>
</taxonomicName>
; 
<taxonomicName id="F486D1B4FF85C91BFC8278B32D333B9C" box="[888,1090,1929,1954]" class="Reptilia" family="Leptotyphlopidae" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="479" phylum="Chordata" rank="family">Leptotyphlopidae</taxonomicName>
: 
<taxonomicName id="F486D1B4FF85C91BFBAD78B32CE53B9C" box="[1111,1428,1929,1954]" class="Reptilia" family="Leptotyphlopidae" genus="Myriopholis" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="479" phylum="Chordata" rank="species" species="macrorhyncha">
<emphasis id="01F27625FF85C91BFBAD78B32CE53B9C" box="[1111,1428,1929,1954]" italics="true" pageId="6" pageNumber="479">Myriopholis macrorhyncha</emphasis>
</taxonomicName>
; 
<taxonomicName id="F486D1B4FF85C91BFF6D7897299A3BF8" box="[151,235,1965,1990]" class="Reptilia" family="Boidae" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="479" phylum="Chordata" rank="family">Boidae</taxonomicName>
: 
<taxonomicName id="F486D1B4FF85C91BFF0D789528F73BF8" box="[247,390,1965,1990]" class="Reptilia" family="Boidae" genus="Eryx" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="479" phylum="Chordata" rank="species" species="jaculus">
<emphasis id="01F27625FF85C91BFF0D789528F73BF8" box="[247,390,1965,1990]" italics="true" pageId="6" pageNumber="479">Eryx jaculus</emphasis>
</taxonomicName>
; 
<taxonomicName id="F486D1B4FF85C91BFE6D78972B693BF8" box="[407,536,1965,1990]" class="Reptilia" family="Colubridae" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="479" phylum="Chordata" rank="family">Colubridae</taxonomicName>
: 
<taxonomicName id="F486D1B4FF85C91BFDDE78952BAB3BF8" box="[548,730,1967,1990]" class="Reptilia" family="Colubridae" genus="Eirenis" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="479" phylum="Chordata" rank="species" species="persicus">
<emphasis id="01F27625FF85C91BFDDE78952BAB3BF8" box="[548,730,1967,1990]" italics="true" pageId="6" pageNumber="479">Eirenis persicus</emphasis>
</taxonomicName>
, 
<taxonomicName id="F486D1B4FF85C91BFD1178952AB63BF8" box="[747,967,1965,1990]" class="Reptilia" family="Colubridae" genus="Eirenis" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="479" phylum="Chordata" rank="species" species="punctatolineatus">
<emphasis id="01F27625FF85C91BFD1178952AB63BF8" box="[747,967,1965,1990]" italics="true" pageId="6" pageNumber="479">E. punctatolineatus</emphasis>
</taxonomicName>
, 
<taxonomicName id="F486D1B4FF85C91BFC2C78952DFF3BF8" box="[982,1166,1965,1990]" class="Reptilia" family="Colubridae" genus="Eirenis" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="479" phylum="Chordata" rank="species" species="coronelloides">
<emphasis id="01F27625FF85C91BFC2C78952DFF3BF8" box="[982,1166,1965,1990]" italics="true" pageId="6" pageNumber="479">E. coronelloides</emphasis>
</taxonomicName>
, 
<taxonomicName id="F486D1B4FF85C91BFB6578972CE53BF8" box="[1183,1428,1965,1990]" class="Reptilia" family="Colubridae" genus="Hemorhois" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="479" phylum="Chordata" rank="species" species="ravergieri">
<emphasis id="01F27625FF85C91BFB6578972CE53BF8" box="[1183,1428,1965,1990]" italics="true" pageId="6" pageNumber="479">Hemorhois ravergieri</emphasis>
</taxonomicName>
, 
<taxonomicName id="F486D1B4FF85C91BFF6D78EB28EE3BD4" box="[151,415,2001,2026]" class="Reptilia" family="Colubridae" genus="Platyceps" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="479" phylum="Chordata" rank="species" species="rhodorachis">
<emphasis id="01F27625FF85C91BFF6D78EB28EE3BD4" box="[151,415,2001,2026]" italics="true" pageId="6" pageNumber="479">Platyceps rhodorachis</emphasis>
</taxonomicName>
, 
<taxonomicName id="F486D1B4FF85C91BFE4178EB2B903BD4" box="[443,737,2001,2026]" class="Reptilia" family="Colubridae" genus="Spalerosophis" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="479" phylum="Chordata" rank="species" species="microlepis">
<emphasis id="01F27625FF85C91BFE4178EB2B903BD4" box="[443,737,2001,2026]" italics="true" pageId="6" pageNumber="479">Spalerosophis microlepis</emphasis>
</taxonomicName>
, 
<taxonomicName id="F486D1B4FF85C91BFD0678EB2A983BD4" box="[764,1001,2001,2026]" class="Reptilia" family="Lamprophiidae" genus="Malpolon" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="479" phylum="Chordata" rank="species" species="insignitus">
<emphasis id="01F27625FF85C91BFD0678EB2A983BD4" box="[764,1001,2001,2026]" italics="true" pageId="6" pageNumber="479">Malpolon insignitus</emphasis>
</taxonomicName>
, 
<taxonomicName id="F486D1B4FF85C91BFBFE78EB2C763BD4" box="[1028,1287,2001,2026]" class="Reptilia" family="Lamprophiidae" genus="Psammophis" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="479" phylum="Chordata" rank="species" species="schokari">
<emphasis id="01F27625FF85C91BFBFE78EB2C763BD4" box="[1028,1287,2001,2026]" italics="true" pageId="6" pageNumber="479">Psammophis schokari</emphasis>
</taxonomicName>
, 
<taxonomicName id="F486D1B4FF85C91AFAD878EB287D3C8E" class="Reptilia" family="Colubridae" genus="Telescopus" kingdom="Animalia" lastPageId="7" lastPageNumber="480" order="Squamata" pageId="6" pageNumber="479" phylum="Chordata" rank="species" species="tessellatus">
<emphasis id="01F27625FF85C91AFAD878EB287D3C8E" italics="true" lastPageId="7" lastPageNumber="480" pageId="6" pageNumber="479">Telescopus tessellatus</emphasis>
</taxonomicName>
; 
<taxonomicName id="F486D1B4FF84C91AFED97FAD28FB3C8E" box="[291,394,151,176]" class="Reptilia" family="Elapidae" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="480" phylum="Chordata" rank="family">Elapidae</taxonomicName>
: 
<taxonomicName id="F486D1B4FF84C91AFE677FAD2BD73C8E" box="[413,678,151,176]" class="Reptilia" family="Elapidae" genus="Walterinnesia" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="480" phylum="Chordata" rank="species" species="morgani">
<emphasis id="01F27625FF84C91AFE677FAD2BD73C8E" box="[413,678,151,176]" italics="true" pageId="7" pageNumber="480">Walterinnesia morgani</emphasis>
</taxonomicName>
; 
<taxonomicName id="F486D1B4FF84C91AFD477FAD2A5F3C8E" box="[701,814,151,176]" class="Reptilia" family="Viperidae" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="480" phylum="Chordata" rank="family">Viperidae</taxonomicName>
: 
<taxonomicName id="F486D1B4FF84C91AFCB87FA32D4C3C8E" box="[834,1085,151,176]" class="Reptilia" family="Viperidae" genus="Macrovipera" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="480" phylum="Chordata" rank="species" species="lebetina">
<emphasis id="01F27625FF84C91AFCB87FA32D4C3C8E" box="[834,1085,151,176]" italics="true" pageId="7" pageNumber="480">Macrovipera lebetina</emphasis>
</taxonomicName>
, 
<taxonomicName id="F486D1B4FF84C91AFBAE7FAD2C013C8E" box="[1108,1392,151,176]" class="Reptilia" family="Viperidae" genus="Pseudocerastes" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="480" phylum="Chordata" rank="species" species="persicus">
<emphasis id="01F27625FF84C91AFBAE7FAD2C013C8E" box="[1108,1392,151,176]" italics="true" pageId="7" pageNumber="480">Pseudocerastes persicus</emphasis>
</taxonomicName>
, 
<taxonomicName id="F486D1B4FF84C91AFA7D7FA3284B3CEB" class="Reptilia" family="Viperidae" genus="Pseudocerastes" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="480" phylum="Chordata" rank="species" species="urarachnoides">
<emphasis id="01F27625FF84C91AFA7D7FA3284B3CEB" italics="true" pageId="7" pageNumber="480">P. urarachnoides</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="3339AA37FF84C91AFF3D7FE52D2F3DB6" blockId="7.[151,1437,151,392]" pageId="7" pageNumber="480">
<emphasis id="01F27625FF84C91AFF3D7FE52B513CC6" bold="true" box="[199,544,223,248]" pageId="7" pageNumber="480">Phylogenetic reconstruction.</emphasis>
The concatenated dataSet compriSed 1182 aligned SiteS of which 221 were parSimony informative. Nucleotide frequency diStribution iS 30.7 % for A, 24.2% for C, 28.9% for T and 16.2% for G. Saturation analySeS revealed that the concatenated data-Set haS little SubStitution Saturation, i.e., the ISS value of 0.29 (the obServed index of SubStitution Saturation) iS Significantly lower than ISSc value of 0.77 (critical index of SubStitution Saturation), indicating that our data are Suitable for phylogenetic analySeS.
</paragraph>
<caption id="67F9FABFFF84C91AFF6D7BF42A5D393D" httpUri="https://zenodo.org/record/827932/files/figure.png" pageId="7" pageNumber="480" targetBox="[297,1278,459,1201]" targetPageId="7">
<paragraph id="3339AA37FF84C91AFF6D7BF42A5D393D" blockId="7.[151,1436,1230,1283]" pageId="7" pageNumber="480">
<emphasis id="01F27625FF84C91AFF6D7BF4286238DA" bold="true" box="[151,275,1230,1252]" pageId="7" pageNumber="480">FIGURE 5.</emphasis>
Hierarchical cluster analysis of four species of 
<taxonomicName id="F486D1B4FF84C91AFD0D7BF42AD538DA" box="[759,932,1230,1252]" class="Reptilia" family="Colubridae" genus="Rhynchocalamus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="480" phylum="Chordata" rank="genus">
<emphasis id="01F27625FF84C91AFD0D7BF42AD538DA" box="[759,932,1230,1252]" italics="true" pageId="7" pageNumber="480">Rhynchocalamus</emphasis>
</taxonomicName>
(
<taxonomicName id="F486D1B4FF84C91AFC497BEA2D5938DA" box="[947,1064,1230,1253]" class="Reptilia" family="Colubridae" genus="Rhynchocalamus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="480" phylum="Chordata" rank="species" species="arabicus">
<emphasis id="01F27625FF84C91AFC497BEA2D5938DA" box="[947,1064,1230,1253]" italics="true" pageId="7" pageNumber="480">R. arabicus</emphasis>
</taxonomicName>
, 
<taxonomicName id="F486D1B4FF84C91AFBCF7BEA2D8438DA" box="[1077,1269,1230,1253]" class="Reptilia" family="Colubridae" genus="Rhynchocalamus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="480" phylum="Chordata" rank="species" species="melanocephalus">
<emphasis id="01F27625FF84C91AFBCF7BEA2D8438DA" box="[1077,1269,1230,1253]" italics="true" pageId="7" pageNumber="480">R. melanocephalus</emphasis>
</taxonomicName>
, 
<taxonomicName id="F486D1B4FF84C91AFAF97BEA2C0038DB" box="[1283,1393,1232,1253]" class="Reptilia" family="Colubridae" genus="Rhynchocalamus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="480" phylum="Chordata" rank="species" species="satunini">
<emphasis id="01F27625FF84C91AFAF97BEA2C0038DB" box="[1283,1393,1232,1253]" italics="true" pageId="7" pageNumber="480">R. satunini</emphasis>
</taxonomicName>
and 
<taxonomicName id="F486D1B4FF84C91AFF6D7BD42862393D" box="[151,275,1261,1283]" class="Magnoliopsida" family="Rosaceae" genus="Rhynchochalamus" kingdom="Plantae" order="Rosales" pageId="7" pageNumber="480" phylum="Tracheophyta" rank="species" species="ilamensis" status="sp. nov.">
<emphasis id="01F27625FF84C91AFF6D7BD42862393D" box="[151,275,1261,1283]" italics="true" pageId="7" pageNumber="480">R. ilamensis</emphasis>
</taxonomicName>
<emphasis id="01F27625FF84C91AFEE37BD42818393D" bold="true" box="[281,361,1262,1283]" pageId="7" pageNumber="480">
<taxonomicNameLabel id="1AC1CB5EFF84C91AFEE37BD42818393D" box="[281,361,1262,1283]" pageId="7" pageNumber="480" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
) based on multisate and meristic characters.
</paragraph>
</caption>
<paragraph id="3339AA37FF84C91AFF3D7A102AB03A24" blockId="7.[151,1437,1322,1707]" pageId="7" pageNumber="480">
The ML phylogenetic tree (
<figureCitation id="ABBDB6B2FF84C91AFDDF7A102B03397D" box="[549,626,1322,1347]" captionStart="FIGURE 6" captionStartId="8.[151,250,1419,1441]" captionTargetBox="[166,1440,574,1385]" captionTargetId="figure@8.[161,1446,565,1393]" captionTargetPageId="8" captionText="FIGURE 6. Maximum likelihood tree of Rhynchocalamus based on the concatenated dataset of partial mitochondrial (Cytb) and nuclear (C-mos) genes. Numbers aboVe and below the lines denote ML bootstrap and posterior probability support Values, respectiVely. Photos of the different species include R. arabicus, R. satunini and R. melanocephalus from Šmíd et al. (2015), Muhtarophis barani from AVcı et al. (2015) and Lytorhynchus diadema from the Reptile-Database (http: // www. reptiledatabase. org /)." httpUri="https://zenodo.org/record/827934/files/figure.png" pageId="7" pageNumber="480">Fig. 6</figureCitation>
) ShowS that 
<taxonomicName id="F486D1B4FF84C91AFCE57A162AC3397D" box="[799,946,1322,1347]" class="Magnoliopsida" family="Rosaceae" genus="Rhynchochalamus" kingdom="Plantae" order="Rosales" pageId="7" pageNumber="480" phylum="Tracheophyta" rank="species" species="ilamensis" status="sp. nov.">
<emphasis id="01F27625FF84C91AFCE57A162AC3397D" box="[799,946,1322,1347]" italics="true" pageId="7" pageNumber="480">R. ilamensis</emphasis>
</taxonomicName>
<emphasis id="01F27625FF84C91AFC397A102D59397D" bold="true" box="[963,1064,1322,1347]" pageId="7" pageNumber="480">
<taxonomicNameLabel id="1AC1CB5EFF84C91AFC397A102D59397D" box="[963,1064,1322,1347]" pageId="7" pageNumber="480" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
iS phylogenetically related to 
<taxonomicName id="F486D1B4FF84C91AFF6D7A7428273959" box="[151,342,1358,1383]" class="Reptilia" family="Colubridae" genus="Rhynchocalamus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="480" phylum="Chordata" rank="genus">
<emphasis id="01F27625FF84C91AFF6D7A7428273959" box="[151,342,1358,1383]" italics="true" pageId="7" pageNumber="480">Rhynchocalamus</emphasis>
</taxonomicName>
. 
<taxonomicName id="F486D1B4FF84C91AFE9E7A742B523959" box="[356,547,1358,1383]" class="Reptilia" family="Colubridae" genus="Rhynchocalamus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="480" phylum="Chordata" rank="genus">
<emphasis id="01F27625FF84C91AFE9E7A742B523959" box="[356,547,1358,1383]" italics="true" pageId="7" pageNumber="480">Rhynchocalamus</emphasis>
</taxonomicName>
began diverging with 
<taxonomicName id="F486D1B4FF84C91AFCDB7A6A2A863959" box="[801,1015,1358,1383]" class="Reptilia" family="Colubridae" genus="Rhynchocalamus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="480" phylum="Chordata" rank="species" species="melanocephalus">
<emphasis id="01F27625FF84C91AFCDB7A6A2A863959" box="[801,1015,1358,1383]" italics="true" pageId="7" pageNumber="480">R. melanocephalus</emphasis>
</taxonomicName>
which formS a SiSter taxon to a clade compoSed of the other three SpecieS (
<taxonomicName id="F486D1B4FF84C91AFDBF7A4E2BB939B5" box="[581,712,1394,1419]" class="Reptilia" family="Colubridae" genus="Rhynchocalamus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="480" phylum="Chordata" rank="species" species="arabicus">
<emphasis id="01F27625FF84C91AFDBF7A4E2BB939B5" box="[581,712,1394,1419]" italics="true" pageId="7" pageNumber="480">R. arabicus</emphasis>
</taxonomicName>
, 
<taxonomicName id="F486D1B4FF84C91AFD227A4E2A2539B5" box="[728,852,1396,1419]" class="Reptilia" family="Colubridae" genus="Rhynchocalamus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="480" phylum="Chordata" rank="species" species="satunini">
<emphasis id="01F27625FF84C91AFD227A4E2A2539B5" box="[728,852,1396,1419]" italics="true" pageId="7" pageNumber="480">R. satunini</emphasis>
</taxonomicName>
and 
<taxonomicName id="F486D1B4FF84C91AFC757A4E2D6B39B5" box="[911,1050,1394,1419]" class="Magnoliopsida" family="Rosaceae" genus="Rhynchochalamus" kingdom="Plantae" order="Rosales" pageId="7" pageNumber="480" phylum="Tracheophyta" rank="species" species="ilamensis" status="sp. nov.">
<emphasis id="01F27625FF84C91AFC757A4E2D6B39B5" box="[911,1050,1394,1419]" italics="true" pageId="7" pageNumber="480">R. ilamensis</emphasis>
</taxonomicName>
<emphasis id="01F27625FF84C91AFBD97A482D0E39B5" bold="true" box="[1059,1151,1394,1419]" pageId="7" pageNumber="480">
<taxonomicNameLabel id="1AC1CB5EFF84C91AFBD97A482D0E39B5" box="[1059,1151,1394,1419]" pageId="7" pageNumber="480" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
). The Second dichotomy SeparateS 
<taxonomicName id="F486D1B4FF84C91AFEFE7AA228F53991" box="[260,388,1430,1455]" class="Reptilia" family="Colubridae" genus="Rhynchocalamus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="480" phylum="Chordata" rank="species" species="arabicus">
<emphasis id="01F27625FF84C91AFEFE7AA228F53991" box="[260,388,1430,1455]" italics="true" pageId="7" pageNumber="480">R. arabicus</emphasis>
</taxonomicName>
from a clade compoSed of 
<taxonomicName id="F486D1B4FF84C91AFD487AA22A4B3991" box="[690,826,1430,1455]" class="Magnoliopsida" family="Rosaceae" genus="Rhynchochalamus" kingdom="Plantae" order="Rosales" pageId="7" pageNumber="480" phylum="Tracheophyta" rank="species" species="ilamensis" status="sp. nov.">
<emphasis id="01F27625FF84C91AFD487AA22A4B3991" box="[690,826,1430,1455]" italics="true" pageId="7" pageNumber="480">R. ilamensis</emphasis>
</taxonomicName>
<emphasis id="01F27625FF84C91AFCBA7AAC2AE83991" bold="true" box="[832,921,1430,1455]" pageId="7" pageNumber="480">
<taxonomicNameLabel id="1AC1CB5EFF84C91AFCBA7AAC2AE83991" box="[832,921,1430,1455]" pageId="7" pageNumber="480" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and 
<taxonomicName id="F486D1B4FF84C91AFC347AA22D363991" box="[974,1095,1432,1455]" class="Reptilia" family="Colubridae" genus="Rhynchocalamus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="480" phylum="Chordata" rank="species" species="satunini">
<emphasis id="01F27625FF84C91AFC347AA22D363991" box="[974,1095,1432,1455]" italics="true" pageId="7" pageNumber="480">R. satunini</emphasis>
</taxonomicName>
. ReconStructed ML tree baSed on 
<emphasis id="01F27625FF84C91AFF447A80298239ED" box="[190,243,1466,1491]" italics="true" pageId="7" pageNumber="480">Cytb</emphasis>
SequenceS Showed a Similar topology to the concatenated dataSet, except for a polytomy at the baSal diviSion of 
<taxonomicName id="F486D1B4FF84C91AFEE37AE428A939C9" box="[281,472,1502,1527]" class="Reptilia" family="Colubridae" genus="Rhynchocalamus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="480" phylum="Chordata" rank="genus">
<emphasis id="01F27625FF84C91AFEE37AE428A939C9" box="[281,472,1502,1527]" italics="true" pageId="7" pageNumber="480">Rhynchocalamus</emphasis>
</taxonomicName>
clade leading to an ambiguity in determining the moSt baSal clade within thiS genuS. The BI concatenated tree reSulted in a topology identical to the ML tree.
</paragraph>
<paragraph id="3339AA37FF84C91AFF3D791C289B3A95" blockId="7.[151,1437,1322,1707]" pageId="7" pageNumber="480">
Kimura 2-parameter (K2P) method of meaSuring genetic diStance uSed for both the concatenated and 
<emphasis id="01F27625FF84C91AFA9D791C2CED3A01" box="[1383,1436,1574,1599]" italics="true" pageId="7" pageNumber="480">Cytb</emphasis>
dataSetS ShowS that 
<taxonomicName id="F486D1B4FF84C91AFE8579762B7A3A5D" box="[383,523,1610,1635]" class="Magnoliopsida" family="Rosaceae" genus="Rhynchochalamus" kingdom="Plantae" order="Rosales" pageId="7" pageNumber="480" phylum="Tracheophyta" rank="species" species="ilamensis" status="sp. nov.">
<emphasis id="01F27625FF84C91AFE8579762B7A3A5D" box="[383,523,1610,1635]" italics="true" pageId="7" pageNumber="480">R. ilamensis</emphasis>
</taxonomicName>
<emphasis id="01F27625FF84C91AFDEF79702B033A5D" bold="true" box="[533,626,1610,1635]" pageId="7" pageNumber="480">
<taxonomicNameLabel id="1AC1CB5EFF84C91AFDEF79702B033A5D" box="[533,626,1610,1635]" pageId="7" pageNumber="480" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Separated from itS cloSeSt relative, 
<taxonomicName id="F486D1B4FF84C91AFBE979762DFE3A5D" box="[1043,1167,1612,1635]" class="Reptilia" family="Colubridae" genus="Rhynchocalamus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="480" phylum="Chordata" rank="species" species="satunini">
<emphasis id="01F27625FF84C91AFBE979762DFE3A5D" box="[1043,1167,1612,1635]" italics="true" pageId="7" pageNumber="480">R. satunini</emphasis>
</taxonomicName>
, by 6% and 9% in the concatenated and the 
<emphasis id="01F27625FF84C91AFE76795428B03AB9" box="[396,449,1646,1671]" italics="true" pageId="7" pageNumber="480">Cytb</emphasis>
SequenceS, reSpectively (
<tableCitation id="7E049F8CFF84C91AFD1B79542A4B3AB9" box="[737,826,1646,1671]" captionStart="TABLE 4" captionStartId="7.[151,239,1755,1778]" captionText="TABLE 4. Genetic distance matrix among the four species of Rhynchocalamus based on Kimura 2 - parameter (K 2 P) method in concatenated (below, grey cells) and Cytb (above, white cells) sequences." pageId="7" pageNumber="480">Table 4</tableCitation>
). ThiS method alSo revealS the low genetic variability within theSe SpecieS (
<tableCitation id="7E049F8CFF84C91AFE7F79A828AD3A95" box="[389,476,1682,1707]" captionStart="TABLE 5" captionStartId="8.[151,239,152,175]" captionText="TABLE 5. Genetic distance within the four species of Rhynchocalamus based on Kimura 2 - parameter (K 2 P) method in Cytb and concatenated sequences. This parameter was not calculated for R. arabicus due to inadequate samples (only one sample was available). Abbreviations: D., distance; S. E., standard error of mean." pageId="7" pageNumber="480">Table 5</tableCitation>
).
</paragraph>
<caption id="67F9FABFFF84C91AFF6D79E12D693B2D" pageId="7" pageNumber="480">
<paragraph id="3339AA37FF84C91AFF6D79E12D693B2D" blockId="7.[151,1436,1754,1811]" pageId="7" pageNumber="480">
<emphasis id="01F27625FF84C91AFF6D79E1287D3ACC" bold="true" box="[151,268,1755,1778]" pageId="7" pageNumber="480">TABLE 4.</emphasis>
Genetic distance matrix among the four species of 
<taxonomicName id="F486D1B4FF84C91AFCB179E02D703ACC" box="[843,1025,1754,1778]" class="Reptilia" family="Colubridae" genus="Rhynchocalamus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="480" phylum="Chordata" rank="genus">
<emphasis id="01F27625FF84C91AFCB179E02D703ACC" box="[843,1025,1754,1778]" italics="true" pageId="7" pageNumber="480">Rhynchocalamus</emphasis>
</taxonomicName>
based on Kimura 2-parameter (K2P) method in concatenated (below, grey cells) and 
<emphasis id="01F27625FF84C91AFD6E79C12BB63B2D" box="[660,711,1787,1811]" italics="true" pageId="7" pageNumber="480">Cytb</emphasis>
(above, white cells) sequences.
</paragraph>
</caption>
<paragraph id="3339AA37FF84C91AFE27780A2A8F3BD4" pageId="7" pageNumber="480">
<table id="41865897FF8436E2FF6578142C113BD4" box="[159,1376,1838,2026]" gridcols="6" gridrows="5" pageId="7" pageNumber="480">
<tr id="8DB6A875FF8436E2FF6578142C113B7B" box="[159,1376,1838,1861]" gridrow="0" pageId="7" pageNumber="480" rowspan-0="1">
<th id="CE67C109FF8436E2FE2778142B233B7B" box="[477,594,1838,1861]" gridcol="1" gridrow="0" pageId="7" pageNumber="480">
<taxonomicName id="F486D1B4FF84C91AFE27780A2B233B7A" box="[477,594,1838,1861]" class="Reptilia" family="Colubridae" genus="Rhynchocalamus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="480" phylum="Chordata" rank="species" species="arabicus">
<emphasis id="01F27625FF84C91AFE27780A2B233B7A" box="[477,594,1838,1861]" italics="true" pageId="7" pageNumber="480">R. arabicus</emphasis>
</taxonomicName>
</th>
<th id="CE67C109FF8436E2FD6278142A653B7B" box="[664,788,1838,1861]" gridcol="2" gridrow="0" pageId="7" pageNumber="480">
<taxonomicName id="F486D1B4FF84C91AFD62780A2A653B7A" box="[664,788,1838,1861]" class="Magnoliopsida" family="Rosaceae" genus="Rhynchochalamus" kingdom="Plantae" order="Rosales" pageId="7" pageNumber="480" phylum="Tracheophyta" rank="species" species="ilamensis">
<emphasis id="01F27625FF84C91AFD62780A2A653B7A" box="[664,788,1838,1861]" italics="true" pageId="7" pageNumber="480">R. ilamensis</emphasis>
</taxonomicName>
</th>
<th id="CE67C109FF8436E2FCE078142A1B3B7B" box="[794,874,1838,1861]" gridcol="3" gridrow="0" pageId="7" pageNumber="480">
<emphasis id="01F27625FF84C91AFCE078152A1B3B7A" bold="true" box="[794,874,1839,1860]" pageId="7" pageNumber="480">sp. nov.</emphasis>
</th>
<th id="CE67C109FF8436E2FC2878142DE23B7B" box="[978,1171,1838,1861]" gridcol="4" gridrow="0" pageId="7" pageNumber="480">
<taxonomicName id="F486D1B4FF84C91AFC28780A2DE23B7A" box="[978,1171,1838,1861]" class="Reptilia" family="Colubridae" genus="Rhynchocalamus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="480" phylum="Chordata" rank="species" species="melanocephalus">
<emphasis id="01F27625FF84C91AFC28780A2DE23B7A" box="[978,1171,1838,1861]" italics="true" pageId="7" pageNumber="480">R. melanocephalus</emphasis>
</taxonomicName>
</th>
<th id="CE67C109FF8436E2FB0878142C113B7B" box="[1266,1376,1838,1861]" gridcol="5" gridrow="0" pageId="7" pageNumber="480">
<taxonomicName id="F486D1B4FF84C91AFB08780A2C113B7B" box="[1266,1376,1840,1861]" class="Reptilia" family="Colubridae" genus="Rhynchocalamus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="480" phylum="Chordata" rank="species" species="satunini">
<emphasis id="01F27625FF84C91AFB08780A2C113B7B" box="[1266,1376,1840,1861]" italics="true" pageId="7" pageNumber="480">R. satunini</emphasis>
</taxonomicName>
</th>
</tr>
<tr id="8DB6A875FF8436E2FF6578622C113B50" box="[159,1376,1880,1902]" gridrow="1" pageId="7" pageNumber="480" rowspan-1="1" rowspan-3="1">
<th id="CE67C109FF8436E2FF65786228003B50" box="[159,369,1880,1902]" gridcol="0" gridrow="1" pageId="7" pageNumber="480">
<taxonomicName id="F486D1B4FF84C91AFF65786328653B50" box="[159,276,1880,1902]" class="Reptilia" family="Colubridae" genus="Rhynchocalamus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="480" phylum="Chordata" rank="species" species="arabicus">
<emphasis id="01F27625FF84C91AFF65786328653B50" box="[159,276,1880,1902]" italics="true" pageId="7" pageNumber="480">R. arabicus</emphasis>
</taxonomicName>
</th>
<td id="CE67C109FF8436E2FD6278622A653B50" box="[664,788,1880,1902]" gridcol="2" gridrow="1" pageId="7" pageNumber="480">0.12</td>
<td id="CE67C109FF8436E2FC2878622DE23B50" box="[978,1171,1880,1902]" gridcol="4" gridrow="1" pageId="7" pageNumber="480">0.12</td>
<td id="CE67C109FF8436E2FB0878622C113B50" box="[1266,1376,1880,1902]" gridcol="5" gridrow="1" pageId="7" pageNumber="480">0.13</td>
</tr>
<tr id="8DB6A875FF8436E2FF6578BB2C113BA9" box="[159,1376,1921,1943]" gridrow="2" pageId="7" pageNumber="480" rowspan-2="1" rowspan-3="1">
<th id="CE67C109FF8436E2FF6578BB28003BA9" box="[159,369,1921,1943]" gridcol="0" gridrow="2" pageId="7" pageNumber="480">
<taxonomicName id="F486D1B4FF84C91AFF6578B8286A3BA9" box="[159,283,1921,1943]" class="Magnoliopsida" family="Rosaceae" genus="Rhynchochalamus" kingdom="Plantae" order="Rosales" pageId="7" pageNumber="480" phylum="Tracheophyta" rank="species" species="ilamensis" status="sp. nov.">
<emphasis id="01F27625FF84C91AFF6578B8286A3BA9" box="[159,283,1921,1943]" italics="true" pageId="7" pageNumber="480">R. ilamensis</emphasis>
</taxonomicName>
<emphasis id="01F27625FF84C91AFEDB78BB28003BA8" bold="true" box="[289,369,1921,1942]" pageId="7" pageNumber="480">
<taxonomicNameLabel id="1AC1CB5EFF84C91AFEDB78BB28003BA8" box="[289,369,1921,1942]" pageId="7" pageNumber="480" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</th>
<td id="CE67C109FF8436E2FE2778BB2B233BA9" box="[477,594,1921,1943]" gridcol="1" gridrow="2" pageId="7" pageNumber="480">0.09</td>
<td id="CE67C109FF8436E2FC2878BB2DE23BA9" box="[978,1171,1921,1943]" gridcol="4" gridrow="2" pageId="7" pageNumber="480">0.13</td>
<td id="CE67C109FF8436E2FB0878BB2C113BA9" box="[1266,1376,1921,1943]" gridcol="5" gridrow="2" pageId="7" pageNumber="480">0.09</td>
</tr>
<tr id="8DB6A875FF8436E2FF6578912C113BFF" box="[159,1376,1963,1985]" gridrow="3" pageId="7" pageNumber="480" rowspan-3="1" rowspan-4="1">
<th id="CE67C109FF8436E2FF65789128003BFF" box="[159,369,1963,1985]" gridcol="0" gridrow="3" pageId="7" pageNumber="480">
<taxonomicName id="F486D1B4FF84C91AFF657896282E3BFF" box="[159,351,1963,1985]" class="Reptilia" family="Colubridae" genus="Rhynchocalamus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="480" phylum="Chordata" rank="species" species="melanocephalus">
<emphasis id="01F27625FF84C91AFF657896282E3BFF" box="[159,351,1963,1985]" italics="true" pageId="7" pageNumber="480">R. melanocephalus</emphasis>
</taxonomicName>
</th>
<td id="CE67C109FF8436E2FE2778912B233BFF" box="[477,594,1963,1985]" gridcol="1" gridrow="3" pageId="7" pageNumber="480">0.08</td>
<td id="CE67C109FF8436E2FD6278912A653BFF" box="[664,788,1963,1985]" gridcol="2" gridrow="3" pageId="7" pageNumber="480">0.10</td>
<td id="CE67C109FF8436E2FB0878912C113BFF" box="[1266,1376,1963,1985]" gridcol="5" gridrow="3" pageId="7" pageNumber="480">0.13</td>
</tr>
<tr id="8DB6A875FF8436E2FF6578EE2C113BD4" box="[159,1376,2004,2026]" gridrow="4" pageId="7" pageNumber="480" rowspan-3="1" rowspan-5="1">
<th id="CE67C109FF8436E2FF6578EE28003BD4" box="[159,369,2004,2026]" gridcol="0" gridrow="4" pageId="7" pageNumber="480">
<taxonomicName id="F486D1B4FF84C91AFF6578EF287C3BD4" box="[159,269,2005,2026]" class="Reptilia" family="Colubridae" genus="Rhynchocalamus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="480" phylum="Chordata" rank="species" species="satunini">
<emphasis id="01F27625FF84C91AFF6578EF287C3BD4" box="[159,269,2005,2026]" italics="true" pageId="7" pageNumber="480">R. satunini</emphasis>
</taxonomicName>
</th>
<td id="CE67C109FF8436E2FE2778EE2B233BD4" box="[477,594,2004,2026]" gridcol="1" gridrow="4" pageId="7" pageNumber="480">0.08</td>
<td id="CE67C109FF8436E2FD6278EE2A653BD4" box="[664,788,2004,2026]" gridcol="2" gridrow="4" pageId="7" pageNumber="480">0.06</td>
<td id="CE67C109FF8436E2FC2878EE2DE23BD4" box="[978,1171,2004,2026]" gridcol="4" gridrow="4" pageId="7" pageNumber="480">0.07</td>
</tr>
</table>
</paragraph>
<caption id="67F9FABFFF8BC915FF6D7FA22A853CD1" pageId="8" pageNumber="481">
<paragraph id="3339AA37FF8BC915FF6D7FA22A853CD1" blockId="8.[151,1436,151,239]" pageId="8" pageNumber="481">
<emphasis id="01F27625FF8BC915FF6D7FA2287A3C91" bold="true" box="[151,267,152,175]" pageId="8" pageNumber="481">TABLE 5.</emphasis>
Genetic distance within the four species of 
<taxonomicName id="F486D1B4FF8BC915FD197FAD2AE83C91" box="[739,921,151,175]" class="Reptilia" family="Colubridae" genus="Rhynchocalamus" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="481" phylum="Chordata" rank="genus">
<emphasis id="01F27625FF8BC915FD197FAD2AE83C91" box="[739,921,151,175]" italics="true" pageId="8" pageNumber="481">Rhynchocalamus</emphasis>
</taxonomicName>
based on Kimura 2-parameter (K2P) method in 
<emphasis id="01F27625FF8BC915FF6D7F8D29BB3CF1" box="[151,202,183,207]" italics="true" pageId="8" pageNumber="481">Cytb</emphasis>
and concatenated sequences. This parameter was not calculated for 
<taxonomicName id="F486D1B4FF8BC915FC607F822D653CF1" box="[922,1044,183,207]" class="Reptilia" family="Colubridae" genus="Rhynchocalamus" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="481" phylum="Chordata" rank="species" species="arabicus">
<emphasis id="01F27625FF8BC915FC607F822D653CF1" box="[922,1044,183,207]" italics="true" pageId="8" pageNumber="481">R. arabicus</emphasis>
</taxonomicName>
due to inadequate samples (only one sample was available). Abbreviations: D., distance; S.E., standard error of mean.
</paragraph>
</caption>
<paragraph id="3339AA37FF8BC915FE247E302D113D1E" blockId="8.[159,1328,266,495]" box="[478,1120,266,288]" pageId="8" pageNumber="481">Cytb Concatenated</paragraph>
<paragraph id="3339AA37FF8BC915FE247E0F2C413DD1" blockId="8.[159,1328,266,495]" pageId="8" pageNumber="481">
D. S.E. D. S.E. 
<taxonomicName id="F486D1B4FF8BC915FF657E6428653D4D" box="[159,276,349,371]" class="Reptilia" family="Colubridae" genus="Rhynchocalamus" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="481" phylum="Chordata" rank="species" species="arabicus">
<emphasis id="01F27625FF8BC915FF657E6428653D4D" box="[159,276,349,371]" italics="true" pageId="8" pageNumber="481">R. arabicus</emphasis>
</taxonomicName>
---- ---- ---- ---- 
<taxonomicName id="F486D1B4FF8BC915FF657EB2286A3DA2" box="[159,283,390,413]" class="Magnoliopsida" family="Rosaceae" genus="Rhynchochalamus" kingdom="Plantae" order="Rosales" pageId="8" pageNumber="481" phylum="Tracheophyta" rank="species" species="ilamensis" status="sp. nov.">
<emphasis id="01F27625FF8BC915FF657EB2286A3DA2" box="[159,283,390,413]" italics="true" pageId="8" pageNumber="481">R. ilamensis</emphasis>
</taxonomicName>
<emphasis id="01F27625FF8BC915FEDB7EBD28003DA2" bold="true" box="[289,369,391,412]" pageId="8" pageNumber="481">
<taxonomicNameLabel id="1AC1CB5EFF8BC915FEDB7EBD28003DA2" box="[289,369,391,412]" pageId="8" pageNumber="481" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
0.030 0.006 0.007 0.003 
<taxonomicName id="F486D1B4FF8BC915FF657E8B282E3DF8" box="[159,351,432,454]" class="Reptilia" family="Colubridae" genus="Rhynchocalamus" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="481" phylum="Chordata" rank="species" species="melanocephalus">
<emphasis id="01F27625FF8BC915FF657E8B282E3DF8" box="[159,351,432,454]" italics="true" pageId="8" pageNumber="481">R. melanocephalus</emphasis>
</taxonomicName>
0.003 0.002 0.002 0.001 
<taxonomicName id="F486D1B4FF8BC915FF657EE0287C3DD1" box="[159,269,474,495]" class="Reptilia" family="Colubridae" genus="Rhynchocalamus" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="481" phylum="Chordata" rank="species" species="satunini">
<emphasis id="01F27625FF8BC915FF657EE0287C3DD1" box="[159,269,474,495]" italics="true" pageId="8" pageNumber="481">R. satunini</emphasis>
</taxonomicName>
0.022 0.005 0.014 0.003
</paragraph>
</subSubSection>
</treatment>
</document>