treatments-xml/data/83/20/68/832068A34ABC538787CBBB5D16AE81F4.xml

<document ID-DOI="http://dx.doi.org/10.3897/zookeys.948.54618" ID-GBIF-Dataset="a84dce14-406e-405b-859c-00224bc9a26f" ID-PMC="PMC7381435" ID-Pensoft-Pub="1313-2970-948-47" ID-Pensoft-UUID="F9484984535F535DA14F339DBD319546" ID-PubMed="32765171" ID-ZooBank="1B7C483E56E1418DA816142EFEE8D925" ModsDocID="1313-2970-948-47" checkinTime="1594659716415" checkinUser="pensoft" docAuthor="Schneider, Scott A., Claps, Lucia E., Wei, Jiufeng, Normark, Roxanna D. &amp; Normark, Benjamin B." docDate="2020" docId="832068A34ABC538787CBBB5D16AE81F4" docLanguage="en" docName="ZooKeys 948: 47-73" docOrigin="ZooKeys 948" docSource="http://dx.doi.org/10.3897/zookeys.948.54618" docTitle="Melanaspis lilloi Schneider, Claps, Wei, Normark &amp; Normark 2020, sp. nov." docType="treatment" docUuid="1559ADD4-4074-4033-B94B-4B1E4441A974" docUuidSource="ZooBank" docVersion="4" id="F9484984535F535DA14F339DBD319546" lastPageNumber="47" masterDocId="F9484984535F535DA14F339DBD319546" masterDocTitle="Five new species of Aspidiotini (Hemiptera, Diaspididae, Aspidiotinae) from Argentina, with a key to Argentine species" masterLastPageNumber="73" masterPageNumber="47" pageNumber="47" updateTime="1668169090276" updateUser="ExternalLinkService">
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<mods:title>Five new species of Aspidiotini (Hemiptera, Diaspididae, Aspidiotinae) from Argentina, with a key to Argentine species</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Schneider, Scott A.</mods:namePart>
<mods:affiliation>USDA, Agricultural Research Service, Henry A. Wallace Beltsville Agricultural Research Center, Systematic Entomology Laboratory, Building 005 - Room 004, 10300 Baltimore Avenue, Beltsville, MD 20705, USA</mods:affiliation>
<mods:nameIdentifier type="orcid">https://orcid.org/0000-0002-9679-8437</mods:nameIdentifier>
<mods:nameIdentifier type="email">scott.schneider@ars.usda.gov</mods:nameIdentifier>
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<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Claps, Lucia E.</mods:namePart>
<mods:affiliation>Universidad Nacional de Tucuman. Facultad de Ciencias Naturales e Instituto Miguel Lillo, Instituto Superior de Entomologia &quot; Dr. Abraham Willink &quot;, Batalla de Ayacucho 491, T 4000 San Miguel de Tucuman, Tucuman, Argentina</mods:affiliation>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Wei, Jiufeng</mods:namePart>
<mods:affiliation>College of Agriculture, Shanxi Agricultural University, Taigu, Shanxi, 030801, China</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Normark, Roxanna D.</mods:namePart>
<mods:affiliation>Department of Biology, University of Massachusetts, 221 Morrill Science Center III 611 North Pleasant Street, Amherst, MA 01003, USA</mods:affiliation>
</mods:name>
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<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Normark, Benjamin B.</mods:namePart>
<mods:affiliation>Department of Biology, University of Massachusetts, 221 Morrill Science Center III 611 North Pleasant Street, Amherst, MA 01003, USA &amp; Graduate Program in Organismic and Evolutionary Biology, University of Massachusetts, 204 C French Hall, 230 Stockbridge Road Amherst, MA 01003, USA</mods:affiliation>
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<mods:date>2020</mods:date>
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<mods:number>948</mods:number>
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<mods:start>47</mods:start>
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<mods:url>http://dx.doi.org/10.3897/zookeys.948.54618</mods:url>
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<treatment ID-GBIF-Taxon="165073695" LSID="urn:lsid:zoobank.org:act:1559ADD4-4074-4033-B94B-4B1E4441A974" httpUri="http://treatment.plazi.org/id/832068A34ABC538787CBBB5D16AE81F4" lastPageNumber="47" pageId="0" pageNumber="47">
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<paragraph pageId="0" pageNumber="47">
<taxonomicName LSID="832068A3-4ABC-5387-87CB-BB5D16AE81F4" authority="Schneider, Claps, Wei, Normark &amp; Normark" authorityName="Schneider, Claps, Wei, Normark &amp; Normark" authorityYear="2020" class="Insecta" family="Diaspididae" genus="Melanaspis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Melanaspis lilloi" order="Hemiptera" pageId="0" pageNumber="47" phylum="Arthropoda" rank="species" species="lilloi" status="sp. nov.">Melanaspis lilloi Schneider, Claps, Wei, Normark &amp; Normark</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="47">sp. nov.</taxonomicNameLabel>
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Melanaspis lilloi sp. nov. Adult female, full body view, illustrated from the holotype (D 0275 L)." figureDoi="10.3897/zookeys.948.54618.figure7" httpUri="https://binary.pensoft.net/fig/431028" pageId="0" pageNumber="47">Figs 7</figureCitation>
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Melanaspis lilloi sp. nov. Adult female, expanded view of pygidium, illustrated from the holotype (D 0275 L)." figureDoi="10.3897/zookeys.948.54618.figure8" httpUri="https://binary.pensoft.net/fig/431029" pageId="0" pageNumber="47">, 8</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="47" type="materials_examined">
<paragraph pageId="0" pageNumber="47">Material examined.</paragraph>
<paragraph pageId="0" pageNumber="47">
<emphasis bold="true" italics="true" pageId="0" pageNumber="47">Holotype</emphasis>
: Argentina • 1 adult female; Jujuy, 30 km N Humahuaca; 
<geoCoordinate degrees="22.97" direction="south" orientation="latitude" precision="555" value="-22.97">22.97S</geoCoordinate>
, 
<geoCoordinate degrees="65.39" direction="west" orientation="longitude" precision="555" value="-65.39">65.39W</geoCoordinate>
; 12.II.2002; L. E. Claps, P. Zamudio, L. Diaz-Briz, &amp; P. Cabrera leg.; IFML, L. E. Claps catalog #5-02, #1092 (D0275L). 
<emphasis bold="true" italics="true" pageId="0" pageNumber="47">Paratypes</emphasis>
: Argentina • 3 adult females; same data as holotype; USNM (D0275H) • 1 adult female; same data as holotype; USNM (D0275G) • 1 adult female; same data as holotype; USNM (D0275K) • 4 adult females; same data as holotype; UMEC (D0275I) • 1 adult female; same data as holotype; UMEC (D0275M) • 4 adult females; same data as holotype; IFML (D0275J) • 1 adult female; Jujuy, Humahuaca, entrada a Iruya; 22.997S, 65.356W; 12.II.2002; L. E. Claps, P. Zamudio, L. Diaz-Briz, &amp; P. Cabrera leg.; IFML, L. E. Claps catalog #15-02 (D0297C).
</paragraph>
<caption doi="10.3897/zookeys.948.54618.figure7" httpUri="https://binary.pensoft.net/fig/431028" pageId="0" pageNumber="47" start="Figure 7" startId="F7">
<paragraph pageId="0" pageNumber="47">
<emphasis bold="true" pageId="0" pageNumber="47">Figure 7.</emphasis>
<taxonomicName authorityName="Schneider, Claps, Wei, Normark &amp; Normark" authorityYear="2020" class="Insecta" family="Diaspididae" genus="Melanaspis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Melanaspis lilloi" order="Hemiptera" pageId="0" pageNumber="47" phylum="Arthropoda" rank="species" species="lilloi">
<emphasis italics="true" pageId="0" pageNumber="47">Melanaspis lilloi</emphasis>
</taxonomicName>
sp. nov. Adult female, full body view, illustrated from the holotype (D0275L).
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="47" type="description">
<paragraph pageId="0" pageNumber="47">Description</paragraph>
<paragraph pageId="0" pageNumber="47">
(
<emphasis italics="true" pageId="0" pageNumber="47">N</emphasis>
= 16). Adult female presumed to secrete scale cover, not pupillarial. Appearance in life not recorded. Slide-mounted adult female 930-1610 (median 1350, holotype 1610) 
<normalizedToken originalValue="μm">μm</normalizedToken>
long, 820-1350 (median 1160, holotype 1330) 
<normalizedToken originalValue="μm">μm</normalizedToken>
wide; broadest near mesothorax. Body outline turbinate. Derm membranous throughout at maturity except for pygidium, which has characteristic dorsal sclerotized areas; sclerotization of these areas unusually heavy, such that paraphyses and basal scleroses of lobes difficult to discern clearly on some specimens. Antennae, simple, each with 1 long seta, distance between antennae 200-330 
<normalizedToken originalValue="μm">μm</normalizedToken>
(median 260). Without disc 
<taxonomicName class="Insecta" family="Diaspididae" genus="Melanaspis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="pores" order="Hemiptera" pageId="0" pageNumber="47" phylum="Arthropoda" rank="species" species="pores">pores</taxonomicName>
near spiracles. 
<emphasis bold="true" italics="true" pageId="0" pageNumber="47">Lobes</emphasis>
: With 4 pairs of well-developed pygidial lobes, L1-L3 apically rounded and L4 truncate or pointed, notches absent from lobes; L1 slightly wider than long, median lobes separated by narrow space 0.15 times width of L1, with basal sclerosis about 1/2 width of L1 arising from mesal edge; L2 and L3 similar in size and shape, shorter and broader than L1; L4 somewhat variable in shape, truncate or with sloping edges. 
<emphasis bold="true" italics="true" pageId="0" pageNumber="47">Paraphyses</emphasis>
: Short and clavate, scarcely longer than L1; absent between L1, paraphysis formula 2-2-3 or 2-2-4; 1 interlobular paraphysis near outer corner of L1, 1 attached to inner corner of L2, 1 in interlobular space between L2 and L3, 1 attached to inner and outer corners of L3, 1 narrow paraphysis attached to inner corner of L4 and 2-3 narrow paraphyses in interlobular space between L3 and L4, these often fused into a single complex mass and difficult to count; several paraphysis-like sclerotizations surrounding macroduct orifices present beyond L4. 
<emphasis bold="true" italics="true" pageId="0" pageNumber="47">Plates</emphasis>
: Apparently absent. 
<emphasis bold="true" italics="true" pageId="0" pageNumber="47">Ducts</emphasis>
: Dorsal pygidial macroducts of 1-barred type, nearly uniform in size, with minute orifices and long slender ducts, most arranged in distinct furrows between sclerotized areas arising from interlobular spaces; 1 submarginal macroduct orifice immediately anterior to each L1, with ducts extending beyond posterior margin of anal opening; 10-21 (median 16) duct orifices in furrow of first space, originating between L1 and L2 and extending in elongate cluster anteriorly 60-85% of distance to anus, each duct about 120-130 
<normalizedToken originalValue="μm">μm</normalizedToken>
in length; 18-40 (median 29) in furrow of second space, originating between L2 and L3 and extending anteriorly to a point laterad or anterolaterad of anus; 3-9 (median 6) on sclerotized area arising from L3; 2-24 (median 15) in furrow of third space, originating between L3 and L4 and extending anteriorly to a point anterolaterad of anus; duct orifices in furrows of second and third spaces membranous, especially towards anterolateral corner of furrow, or surrounded by partial or complete sclerotized ring, especially near posterior end and along medial margin of furrow; submedial clusters of dorsal macroducts present on each pre-pygidial abdominal segment, shorter and narrower than pygidial ducts. Ventral pygidial microducts similar to dorsal macroducts in size and shape and similarly arranged in rows on segments V-VII, 21-44 (median 33) on each side; ventral duct orifices on segment V each surrounded by conspicuous sclerotized ring, degree of sclerotization decreasing towards anterolateral corner of segment; microducts also distributed along head, thorax, and pre-pygidial margins, as well as rows extending from marginal area toward each spiracle. 
<emphasis bold="true" italics="true" pageId="0" pageNumber="47">Anal opening</emphasis>
: Oval, 20-28 (median 24) 
<normalizedToken originalValue="μm">μm</normalizedToken>
long, positioned 3.5-6.3 (median 4.6) anal lengths (85-124, median 107 
<normalizedToken originalValue="μm">μm</normalizedToken>
) from base of L1, near midpoint of pygidium. 
<taxonomicName genus="Perivulvar" lsidName="Perivulvar pores" pageId="0" pageNumber="47" rank="species" species="pores">
<emphasis bold="true" italics="true" pageId="0" pageNumber="47">Perivulvar pores</emphasis>
</taxonomicName>
: Absent.
</paragraph>
<caption doi="10.3897/zookeys.948.54618.figure8" httpUri="https://binary.pensoft.net/fig/431029" pageId="0" pageNumber="47" start="Figure 8" startId="F8">
<paragraph pageId="0" pageNumber="47">
<emphasis bold="true" pageId="0" pageNumber="47">Figure 8.</emphasis>
<taxonomicName authorityName="Schneider, Claps, Wei, Normark &amp; Normark" authorityYear="2020" class="Insecta" family="Diaspididae" genus="Melanaspis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Melanaspis lilloi" order="Hemiptera" pageId="0" pageNumber="47" phylum="Arthropoda" rank="species" species="lilloi">
<emphasis italics="true" pageId="0" pageNumber="47">Melanaspis lilloi</emphasis>
</taxonomicName>
sp. nov. Adult female, expanded view of pygidium, illustrated from the holotype (D0275L).
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="47" type="dna sequences">
<paragraph pageId="0" pageNumber="47">DNA sequences.</paragraph>
<paragraph pageId="0" pageNumber="47">
Several DNA sequences of 
<taxonomicName authorityName="Schneider, Claps, Wei, Normark &amp; Normark" authorityYear="2020" class="Insecta" family="Diaspididae" genus="Melanaspis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Melanaspis lilloi" order="Hemiptera" pageId="0" pageNumber="47" phylum="Arthropoda" rank="species" species="lilloi">
<emphasis italics="true" pageId="0" pageNumber="47">Melanaspis lilloi</emphasis>
</taxonomicName>
sp. nov. have been published, including fragments of 4 loci from a paratype (D0275G): 28S, GenBank accession number KY218989.1; EF-1α, MH915714.1 and KY221286.1; CAD, MH915988.1; and COI-II, KY22069.1 and MH916394.1. DNA sequences have also been published for other members of the type series that were ground to powder during DNA preparation; these include sequences of 28S (D0275D, DQ145363.2; D0297A, DQ145362.2 and KY219142.1), EF-1α (D0275D, DQ145475.1; D0297A, DQ145474.1 and KY221295.1), and COI-II (D0275, GQ445417.1; D0297A, GQ425005.1). DNA sequences of the primary bacterial endosymbiont, 
<taxonomicName class="Bacteroidia" family="Blattabacteriaceae" genus="Uzinura" higherTaxonomySource="GBIF" kingdom="Bacteria" lsidName="Uzinura diaspidicola" order="Flavobacteriales" pageId="0" pageNumber="47" phylum="Bacteroidota" rank="species" species="diaspidicola">
<emphasis italics="true" pageId="0" pageNumber="47">Uzinura diaspidicola</emphasis>
</taxonomicName>
, of 
<taxonomicName lsidName="M. lilloi" pageId="0" pageNumber="47" rank="species" species="lilloi">
<emphasis italics="true" pageId="0" pageNumber="47">M. lilloi</emphasis>
</taxonomicName>
sp. nov. have also been published for ground-up individuals of the type series, including fragments of 16S rDNA (D0275, DQ133558.1 and DQ868836.1; D0297A, GQ424858.1) and 23S rDNA (D0275, DQ873248.1).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="47" type="informal synonyms">
<paragraph pageId="0" pageNumber="47">Informal synonyms.</paragraph>
<paragraph pageId="0" pageNumber="47">
Specimens from the type series have appeared in several published phylogenetic trees, and have been referred to variously as &quot; 
<taxonomicName authorityName="Cockerell" authorityYear="1897" class="Insecta" family="Diaspididae" genus="Melanaspis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Melanaspis" order="Hemiptera" pageId="0" pageNumber="47" phylum="Arthropoda" rank="genus">Melanaspis</taxonomicName>
sp. nov.&quot; (
<bibRefCitation author="Gruwell, ME" journalOrPublisher="Molecular Phylogenetics and Evolution" pageId="0" pageNumber="47" refId="B12" refString="Gruwell, ME, von Dohlen, CD, Patch, K, Normark, BB, 2005. Preliminary PCR survey of bacteria associated with scale insects (Hemiptera: Coccoidea). In: Erkilic LB, Kaydan MB (Eds) Proceedings of the Tenth International Symposium on Scale Insect Studies. Scientific and Technical Research Council of Turkey, Ankara, Adana, Turkey, 101-116 pp." title="Preliminary PCR survey of bacteria associated with scale insects (Hemiptera: Coccoidea). In: Erkilic LB, Kaydan MB (Eds) Proceedings of the Tenth International Symposium on Scale Insect Studies. Scientific and Technical Research Council of Turkey, Ankara, Adana, Turkey, 101 - 116 pp." year="2005">Gruwell et al. 2005</bibRefCitation>
) &quot; 
<taxonomicName authorityName="Cockerell" authorityYear="1897" class="Insecta" family="Diaspididae" genus="Melanaspis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Melanaspis" order="Hemiptera" pageId="0" pageNumber="47" phylum="Arthropoda" rank="genus">Melanaspis</taxonomicName>
sp undesc #2&quot; and &quot; 
<taxonomicName authorityName="Cockerell" authorityYear="1897" class="Insecta" family="Diaspididae" genus="Melanaspis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Melanaspis" order="Hemiptera" pageId="0" pageNumber="47" phylum="Arthropoda" rank="genus">Melanaspis</taxonomicName>
sp undesc #3&quot; (
<bibRefCitation DOI="https://doi.org/10.1016/j.ympev.2007.01.014" author="Gruwell, ME" journalOrPublisher="Molecular Phylogenetics and Evolution" pageId="0" pageNumber="47" pagination="267 - 280" refId="B11" refString="Gruwell, ME, Morse, GE, Normark, BB, 2007. Phylogenetic congruence of armored scale insects (Hemiptera: Diaspididae) and their primary endosymbionts from the phylum Bacteroidetes. Molecular Phylogenetics and Evolution 44: 267 - 280, DOI: https://doi.org/10.1016/j.ympev.2007.01.014" title="Phylogenetic congruence of armored scale insects (Hemiptera: Diaspididae) and their primary endosymbionts from the phylum Bacteroidetes." url="https://doi.org/10.1016/j.ympev.2007.01.014" volume="44" year="2007">Gruwell et al. 2007</bibRefCitation>
; 
<bibRefCitation DOI="https://doi.org/10.1111/j.1365-3113.2005.00316.x" author="Morse, GE" journalOrPublisher="Systematic Entomology" pageId="0" pageNumber="47" pagination="338 - 349" refId="B16" refString="Morse, GE, Normark, BB, 2006. A molecular phylogenetic study of armoured scale insects (Hemiptera: Diaspididae). Systematic Entomology 31: 338 - 349, DOI: https://doi.org/10.1111/j.1365-3113.2005.00316.x" title="A molecular phylogenetic study of armoured scale insects (Hemiptera: Diaspididae)." url="https://doi.org/10.1111/j.1365-3113.2005.00316.x" volume="31" year="2006">Morse and Normark 2006</bibRefCitation>
; 
<bibRefCitation DOI="https://doi.org/10.1093/aesa/103.1.30" author="Rugman-Jones, PF" journalOrPublisher="Annals of the Entomological Society of America" pageId="0" pageNumber="47" pagination="30 - 38" refId="B19" refString="Rugman-Jones, PF, Andersen, JC, Morse, GE, Normark, BB, Stouthamer, R, 2010. Molecular phylogenetic placement of the recently described armored scale insect Abgrallaspis aguacatae and several congeners (Hemiptera: Diaspididae). Annals of the Entomological Society of America 103: 30 - 38, DOI: https://doi.org/10.1093/aesa/103.1.30" title="Molecular phylogenetic placement of the recently described armored scale insect Abgrallaspis aguacatae and several congeners (Hemiptera: Diaspididae)." url="https://doi.org/10.1093/aesa/103.1.30" volume="103" year="2010">Rugman-Jones et al. 2010</bibRefCitation>
), &quot; 
<taxonomicName authorityName="Cockerell" authorityYear="1897" class="Insecta" family="Diaspididae" genus="Melanaspis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Melanaspis" order="Hemiptera" pageId="0" pageNumber="47" phylum="Arthropoda" rank="genus">Melanaspis</taxonomicName>
sp. undesc.&quot; (
<bibRefCitation DOI="https://doi.org/10.1603/008.102.0613" author="Gruwell, ME" journalOrPublisher="Annals of the Entomological Society of America" pageId="0" pageNumber="47" pagination="1050 - 1061" refId="B13" refString="Gruwell, ME, Wu, J, Normark, BB, 2009. Diversity and phylogeny of Cardinium (Bacteroidetes) in armored scale insects (Hemiptera: Diaspididae). Annals of the Entomological Society of America 102: 1050 - 1061, DOI: https://doi.org/10.1603/008.102.0613" title="Diversity and phylogeny of Cardinium (Bacteroidetes) in armored scale insects (Hemiptera: Diaspididae)." url="https://doi.org/10.1603/008.102.0613" volume="102" year="2009">Gruwell et al. 2009</bibRefCitation>
), &quot; 
<taxonomicName authorityName="Cockerell" authorityYear="1897" class="Insecta" family="Diaspididae" genus="Melanaspis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Melanaspis" order="Hemiptera" pageId="0" pageNumber="47" phylum="Arthropoda" rank="genus">Melanaspis</taxonomicName>
sp nov 1&quot; and &quot; 
<taxonomicName authorityName="Cockerell" authorityYear="1897" class="Insecta" family="Diaspididae" genus="Melanaspis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Melanaspis" order="Hemiptera" pageId="0" pageNumber="47" phylum="Arthropoda" rank="genus">Melanaspis</taxonomicName>
sp nov 2&quot; (
<bibRefCitation DOI="https://doi.org/10.1016/j.ympev.2010.05.002" author="Andersen, JC" journalOrPublisher="Molecular Phylogenetics and Evolution" pageId="0" pageNumber="47" pagination="992 - 1003" refId="B1" refString="Andersen, JC, Wu, J, Gruwell, ME, Gwiazdowski, R, Santana, SE, Feliciano, NM, Morse, GE, Normark, BB, 2010. A phylogenetic analysis of armored scale insects (Hemiptera: Diaspididae), based upon nuclear, mitochondrial, and endosymbiont gene sequences. Molecular Phylogenetics and Evolution 57: 992 - 1003, DOI: https://doi.org/10.1016/j.ympev.2010.05.002" title="A phylogenetic analysis of armored scale insects (Hemiptera: Diaspididae), based upon nuclear, mitochondrial, and endosymbiont gene sequences." url="https://doi.org/10.1016/j.ympev.2010.05.002" volume="57" year="2010">Andersen et al. 2010</bibRefCitation>
), and &quot; 
<taxonomicName authorityName="Cockerell" authorityYear="1897" class="Insecta" family="Diaspididae" genus="Melanaspis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Melanaspis" order="Hemiptera" pageId="0" pageNumber="47" phylum="Arthropoda" rank="genus">Melanaspis</taxonomicName>
ud0276&quot; (
<bibRefCitation DOI="https://doi.org/10.1016/j.ympev.2018.09.003" author="Schneider, SA" journalOrPublisher="Molecular Phylogenetics and Evolution" pageId="0" pageNumber="47" pagination="291 - 303" refId="B21" refString="Schneider, SA, Okusu, A, Normark, BB, 2018. Molecular phylogenetics of Aspidiotini armored scale insects (Hemiptera: Diaspididae) reveals rampant paraphyly, curious species radiations, and multiple origins of association with Melissotarsus ants (Hymenoptera: Formicidae). Molecular Phylogenetics and Evolution 129: 291 - 303, DOI: https://doi.org/10.1016/j.ympev.2018.09.003" title="Molecular phylogenetics of Aspidiotini armored scale insects (Hemiptera: Diaspididae) reveals rampant paraphyly, curious species radiations, and multiple origins of association with Melissotarsus ants (Hymenoptera: Formicidae)." url="https://doi.org/10.1016/j.ympev.2018.09.003" volume="129" year="2018">Schneider et al. 2018</bibRefCitation>
; 
<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.4616.1.1" author="Normark, BB" journalOrPublisher="Zootaxa" pageId="0" pageNumber="47" pagination="001 - 098" refId="B18" refString="Normark, BB, Okusu, A, Morse, GE, Peterson, DA, Itioka, T, Schneider, SA, 2019. Phylogeny and classification of armored scale insects (Hemiptera: Coccomorpha: Diaspididae). Zootaxa 4616: 001 - 098, DOI: https://doi.org/10.11646/zootaxa.4616.1.1" title="Phylogeny and classification of armored scale insects (Hemiptera: Coccomorpha: Diaspididae)." url="https://doi.org/10.11646/zootaxa.4616.1.1" volume="4616" year="2019">Normark et al. 2019</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="47" type="remarks">
<paragraph pageId="0" pageNumber="47">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="47">
This species is very similar to 
<taxonomicName lsidName="M. targionoides" pageId="0" pageNumber="47" rank="species" species="targionoides">
<emphasis italics="true" pageId="0" pageNumber="47">M. targionoides</emphasis>
</taxonomicName>
sp. nov. The diagnosis and affinities of 
<taxonomicName lsidName="M. lilloi" pageId="0" pageNumber="47" rank="species" species="lilloi">
<emphasis italics="true" pageId="0" pageNumber="47">M. lilloi</emphasis>
</taxonomicName>
sp. nov. are discussed below under the remarks for 
<taxonomicName lsidName="M. targionoides" pageId="0" pageNumber="47" rank="species" species="targionoides">
<emphasis italics="true" pageId="0" pageNumber="47">M. targionoides</emphasis>
</taxonomicName>
sp. nov.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="47" type="host">
<paragraph pageId="0" pageNumber="47">Host plant.</paragraph>
<paragraph pageId="0" pageNumber="47">Not recorded.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="47" type="etymology">
<paragraph pageId="0" pageNumber="47">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="47">The specific epithet is a noun in the genitive case, meaning &quot;of Lillo&quot;. It honors the Instituto Miguel Lillo, academic home of Lucia Claps and the other scientists who first collected the species described in this manuscript.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="47" type="distribution">
<paragraph pageId="0" pageNumber="47">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="47">Argentina (Jujuy).</paragraph>
</subSubSection>
</treatment>
</document>