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<document id="FCBD4D633807690B1DC998D9F8184A67" ID-DOI="10.5281/zenodo.13643699" ID-ISSN="1732-2421" ID-Zenodo-Dep="13643699" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="karina" checkinTime="1723142205826" checkinUser="felipe" docAuthor="Soriano, Carmen &amp; Delclòs, Xavier" docDate="2006" docId="455787BCFF91245FFFDEFB4C6AFCFA64" docLanguage="en" docName="ActaPalaeontolPol.51.1.185-200.pdf" docOrigin="Acta Palaeontologica Polonica 51 (1)" docStyle="DocumentStyle:93F5927C23D90727DE855278A6F17B37.1:ActaPalaeontolPol.2002-2009.journal_article" docStyleId="93F5927C23D90727DE855278A6F17B37" docStyleName="ActaPalaeontolPol.2002-2009.journal_article" docStyleVersion="1" docTitle="Priacma sanzii Soriano &amp; Delclòs, 2006, sp. nov." docType="treatment" docVersion="3" lastPageNumber="194" masterDocId="B96EFFC4FF982456FFB8FF876E36FFDB" masterDocTitle="New cupedid beetles from the Lower Cretaceous of Spain and the palaeogeography of the family" masterLastPageNumber="200" masterPageNumber="185" pageNumber="194" updateTime="1725365340826" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="28CE668E63E8834DEF6EE0BCD20B2111">New cupedid beetles from the Lower Cretaceous of Spain and the palaeogeography of the family</mods:title>
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<mods:namePart id="0B88E20646BC621DFA8894CC0FC6C41A">Soriano, Carmen</mods:namePart>
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<mods:namePart id="AE64F52774C32D70BB75ACCA3658FCD9">Delclòs, Xavier</mods:namePart>
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<mods:title id="E7FC40E5F3BAD871A347738F73DCD62D">Acta Palaeontologica Polonica</mods:title>
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<mods:date id="6A43DE492496D29F46F0C9A9DD046E70">2006</mods:date>
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<treatment id="455787BCFF91245FFFDEFB4C6AFCFA64" ID-DOI="http://doi.org/10.5281/zenodo.13643752" ID-Zenodo-Dep="13643752" LSID="urn:lsid:plazi:treatment:455787BCFF91245FFFDEFB4C6AFCFA64" httpUri="http://treatment.plazi.org/id/455787BCFF91245FFFDEFB4C6AFCFA64" lastPageNumber="194" pageId="9" pageNumber="194">
<subSubSection id="85E46521FF91245FFFDEFB4C6FA5FB3D" box="[102,403,1226,1255]" pageId="9" pageNumber="194" type="nomenclature">
<paragraph id="CD4136AAFF91245FFFDEFB4C6FA5FB3D" blockId="9.[102,403,1226,1288]" box="[102,403,1226,1255]" pageId="9" pageNumber="194">
<heading id="960981C6FF91245FFFDEFB4C6FA5FB3D" box="[102,403,1226,1255]" fontSize="12" level="2" pageId="9" pageNumber="194" reason="9">
<taxonomicName id="0AFE4D29FF91245FFFDEFB4C6F11FB3C" authorityName="Soriano &amp; Delclòs" authorityYear="2006" box="[102,295,1227,1255]" class="Insecta" family="Cupedidae" genus="Priacma" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="194" phylum="Arthropoda" rank="species" species="sanzii" status="sp. nov.">
<emphasis id="FF8AEAB8FF91245FFFDEFB4C6F11FB3C" box="[102,295,1227,1255]" italics="true" pageId="9" pageNumber="194">Priacma sanzii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E4B957C3FF91245FFE97FB4D6FA5FB3D" box="[303,403,1226,1254]" pageId="9" pageNumber="194" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="85E46521FF91245FFFDEFB746AFCFA64" pageId="9" pageNumber="194" type="description">
<paragraph id="CD4136AAFF91245FFFDEFB746ED2FAD3" blockId="9.[102,403,1226,1288]" box="[102,228,1267,1288]" pageId="9" pageNumber="194">
<figureCitation id="55C52A2FFF91245FFFDEFB746E8EFAD3" box="[102,184,1267,1288]" captionStart="Fig" captionStartId="7.[102,134,1379,1399]" captionTargetBox="[160,1427,225,1352]" captionTargetId="figure-233@7.[160,1427,225,1352]" captionTargetPageId="7" captionText="Fig. 6. Barremian (Early Cretaceous) ommatins Brochocoleus (Brochocoleini) and Zygadenia (Notocupedini), and cupedins Priacma (Priacmini) and Anaglyphites (Mesocupedini) from Spain. A. Zygadenia diazromerali sp. nov. Holotype MCCM LH 12001a, part, Las Hoyas, Cuenca Province. B. Zygadenia martinclosas sp. nov. Holotype IEI LC 2663a, part, La Cabrúa, Lleida Province. C. Zygadenia longicoxa sp. nov. Holotype MCCM LH 908030a, part, Las Hoyas, Cuenca Province, Lower Cretaceous. D. Zygadenia siniestri sp. nov. Holotype MCCM LH 23515a, part, Las Hoyas, Cuenca Province. E. Priacma sanzii sp. nov. Holotype MCCM LH 24524, part, Las Hoyas, Cuenca Province. F. Anaglyphites zherikhini sp. nov. Holotype MCCM LH 15122a, part, Las Hoyas, Cuenca Province. G.?Anaglyphites pluricavus sp. nov. Holotype IEI LP 97Ga, part, La Pedrera, Lleida Province. Scale bars 2 mm." figureDoi="http://doi.org/10.5281/zenodo.13643715" httpUri="https://zenodo.org/record/13643715/files/figure.png" pageId="9" pageNumber="194">Figs. 6E</figureCitation>
,
<figureCitation id="55C52A2FFF91245FFF7AFB746ED6FAD3" box="[194,224,1267,1288]" captionStart="Fig" captionStartId="9.[102,135,813,833]" captionTargetBox="[112,1483,237,782]" captionTargetId="figure-486@9.[102,1486,228,785]" captionTargetPageId="9" captionText="Fig. 8. Camera lucida drawings of Barremian (Early Cretaceous) ommatin Zygadenia (Ommatini) and cupedin Priacma (Priacmini) from Las Hoyas, Cuenca Province, Spain. A. Zygadenia siniestri sp. nov. Holotype MCCM LH 23515a, part, dorsal (A1) and ventral (A2) aspects. B. Priacma sanzii sp. nov. Holotype MCCM LH 24524, part, dorsal (B1) and ventral (B2) aspects. Scale bars 2 mm." figureDoi="http://doi.org/10.5281/zenodo.13643719" httpUri="https://zenodo.org/record/13643719/files/figure.png" pageId="9" pageNumber="194">8B</figureCitation>
.
</paragraph>
<paragraph id="CD4136AAFF91245FFFDEFA9B6F65FA96" blockId="9.[102,775,1308,1511]" pageId="9" pageNumber="194">
<materialsCitation id="7D963CF7FF91245FFFDEFA9B6F79FA96" collectorName="La Cierva" country="Spain" location="Serrania de Cuenca" municipality="Cuenca Province" pageId="9" pageNumber="194" specimenCount="1" typeStatus="holotype">
<emphasis id="FF8AEAB8FF91245FFFDEFA9B6ED3FAEA" box="[102,229,1308,1329]" italics="true" pageId="9" pageNumber="194">
<typeStatus id="12458808FF91245FFFDEFA9B6EA3FAEA" box="[102,149,1308,1329]" pageId="9" pageNumber="194">Type</typeStatus>
locality
</emphasis>
: Las Hoyas fossil site,
<collectorName id="600B537CFF91245FFE69FA9B6C03FAEA" box="[465,565,1308,1329]" pageId="9" pageNumber="194">La Cierva</collectorName>
,
<location id="C8216071FF91245FFDF9FA9B6D35FAEA" LSID="urn:lsid:plazi:treatment:455787BCFF91245FFFDEFB4C6AFCFA64:C8216071FF91245FFDF9FA9B6D35FAEA" box="[577,771,1308,1329]" country="Spain" municipality="Cuenca Province" name="Serrania de Cuenca" pageId="9" pageNumber="194">Serranía de Cuenca</location>
,
<collectingMunicipality id="2D25ACD0FF91245FFFDEFABF6F3AFA96" box="[102,268,1336,1357]" pageId="9" pageNumber="194">Cuenca Province</collectingMunicipality>
,
<collectingCountry id="B5E9763AFF91245FFEADFABF6F79FA96" box="[277,335,1336,1357]" name="Spain" pageId="9" pageNumber="194">Spain</collectingCountry>
</materialsCitation>
.
</paragraph>
<paragraph id="CD4136AAFF91245FFFDEFADC6FB8FA73" blockId="9.[102,775,1308,1511]" pageId="9" pageNumber="194">
<emphasis id="FF8AEAB8FF91245FFFDEFADC6F04FAAB" box="[102,306,1371,1392]" italics="true" pageId="9" pageNumber="194">Stratigraphic horizon</emphasis>
: Second lithosome of finely laminated limestones of La Huérguina Formation, Lower Cretaceous, Barremian (FregenalMartínez and Meléndez 2000).
</paragraph>
<paragraph id="CD4136AAFF91245FFFDEFA306EADFA3C" blockId="9.[102,775,1308,1511]" pageId="9" pageNumber="194">
<emphasis id="FF8AEAB8FF91245FFFDEFA306F7CFA17" box="[102,330,1463,1484]" italics="true" pageId="9" pageNumber="194">Derivation of the name</emphasis>
: Named after the Spanish paleontologist J.L. Sanz.
</paragraph>
<paragraph id="CD4136AAFF91245FFFDEFA7C6C06F9EF" blockId="9.[102,774,1531,1588]" pageId="9" pageNumber="194">
<emphasis id="FF8AEAB8FF91245FFFDEFA7C6EFEF9C8" box="[102,200,1531,1555]" italics="true" pageId="9" pageNumber="194">Material</emphasis>
.—
<materialsCitation id="7D963CF7FF91245FFF50FA7C6C06F9EF" collectionCode="MCCM" pageId="9" pageNumber="194" specimenCode="LH 24524" specimenCount="1" typeStatus="holotype">
<typeStatus id="12458808FF91245FFF50FA7C6F64F9CF" box="[232,338,1531,1556]" pageId="9" pageNumber="194" type="holotype">Holotype</typeStatus>
<collectionCode id="ABEFAE6FFF91245FFEE1FA7C6F86F9CF" box="[345,432,1531,1556]" country="India" httpUri="http://grbio.org/cool/stc7-amit" name="Madras Christian College" pageId="9" pageNumber="194">MCCM</collectionCode>
<specimenCode id="9D589ED1FF91245FFE0FFA7B6C1AF9CF" box="[439,556,1531,1556]" collectionCode="MCCM" country="India" httpUri="http://grbio.org/cool/stc7-amit" name="Madras Christian College" pageId="9" pageNumber="194">LH 24524</specimenCode>
, only the part. Well preserved specimen, lacking part of legs.
</materialsCitation>
</paragraph>
<paragraph id="CD4136AAFF91245FFFDEF9C06C7FF924" blockId="9.[102,775,1606,1791]" pageId="9" pageNumber="194">
<emphasis id="FF8AEAB8FF91245FFFDEF9C06EEEF985" box="[102,216,1607,1630]" italics="true" pageId="9" pageNumber="194">Diagnosis</emphasis>
.—From the
<typeStatus id="12458808FF91245FFEDCF9C06FA2F984" box="[356,404,1607,1631]" pageId="9" pageNumber="194">type</typeStatus>
species of the genus differs in the absence of spines on edge of elytra. From other fossil species (
<taxonomicName id="0AFE4D29FF91245FFFD7F9006FFFF944" authority="Ponomarenko, 2000" authorityName="Ponomarenko" authorityYear="2000" box="[111,457,1670,1695]" class="Insecta" family="Cupedidae" genus="Priacma" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="194" phylum="Arthropoda" rank="species" species="striata">
<emphasis id="FF8AEAB8FF91245FFFD7F9006EEFF945" box="[111,217,1671,1694]" italics="true" pageId="9" pageNumber="194">P. striata</emphasis>
Ponomarenko, 2000
</taxonomicName>
,
<taxonomicName id="0AFE4D29FF91245FFE6FF9006E97F964" authority="Ponomarenko, 1986" authorityName="Ponomarenko" authorityYear="1986" class="Insecta" family="Cupedidae" genus="Priacma" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="194" phylum="Arthropoda" rank="species" species="corrupta">
<emphasis id="FF8AEAB8FF91245FFE6FF9006C6CF945" box="[471,602,1671,1694]" italics="true" pageId="9" pageNumber="194">P. corrupta</emphasis>
<bibRefCitation id="A96F4B5BFF91245FFDDBF9016E97F964" author="Ponomarenko, A. G." pageId="9" pageNumber="194" pagination="108 - 112" refId="ref10255" refString="Ponomarenko, A. G. 1986. Description of fossils. Coleoptera, Myrmeleontida (Neuroptera, Panorpidae, Phryganeidea) [in Russian]. In: A. P. Rasnicyn (ed.), Nasekomye v rannemelovyh ekosistemah zapadnoj Mongolii. Trudy Sovmestnoj Sovetsko-Mongol'skoj Paleontologiceskoj Ekspedicii 28: 108 - 112." type="journal article" year="1986">Ponomarenko, 1986</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="0AFE4D29FF91245FFF08F9206C2AF964" authority="Ponomarenko, 1997" authorityName="Ponomarenko" authorityYear="1997" box="[176,540,1702,1727]" class="Insecta" family="Ommatidae" genus="Zygadenia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="194" phylum="Arthropoda" rank="species" species="oculata">
<emphasis id="FF8AEAB8FF91245FFF08F9206F11F965" box="[176,295,1702,1726]" italics="true" pageId="9" pageNumber="194">P. oculata</emphasis>
<bibRefCitation id="A96F4B5BFF91245FFE8BF9216C2AF964" author="Ponomarenko, A. G." box="[307,540,1702,1727]" pageId="9" pageNumber="194" pagination="45 - 55" refId="ref10494" refString="Ponomarenko, A. G. 1997. New beetles of the Family Cupedidae from the Mesozoic of Mongolia. Ommatini, Mesocupedini, Priacmini. Paleontological Journal 31: 45 - 55." type="journal article" year="1997">Ponomarenko, 1997</bibRefCitation>
</taxonomicName>
, and
<taxonomicName id="0AFE4D29FF91245FFDE7F9206F7CF904" authority="Ponomarenko, 1997" authorityName="Ponomarenko" authorityYear="1997" class="Insecta" family="Ommatidae" genus="Cionocoleus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="194" phylum="Arthropoda" rank="species" species="longicapitis">
<emphasis id="FF8AEAB8FF91245FFDE7F9206D30F965" box="[607,774,1702,1726]" italics="true" pageId="9" pageNumber="194">P. longicapitis</emphasis>
<bibRefCitation id="A96F4B5BFF91245FFFDEF9416F7CF904" author="Ponomarenko, A. G." box="[102,330,1734,1759]" pageId="9" pageNumber="194" pagination="45 - 55" refId="ref10494" refString="Ponomarenko, A. G. 1997. New beetles of the Family Cupedidae from the Mesozoic of Mongolia. Ommatini, Mesocupedini, Priacmini. Paleontological Journal 31: 45 - 55." type="journal article" year="1997">Ponomarenko, 1997</bibRefCitation>
</taxonomicName>
) differs in having a smaller size, elongated head and comparatively shorter eyes.
</paragraph>
<paragraph id="CD4136AAFF91245FFFDEF8956AFCFA64" blockId="9.[102,775,1809,2026]" lastBlockId="9.[812,1486,934,1471]" pageId="9" pageNumber="194">
<emphasis id="FF8AEAB8FF91245FFFDEF8956EDCF8F2" box="[102,234,1810,1833]" italics="true" pageId="9" pageNumber="194">Description</emphasis>
.—Measurements: length
<quantity id="0A069B4FFF91245FFDBDF8966C6BF8F2" box="[517,605,1809,1834]" metricMagnitude="-3" metricUnit="m" metricValue="8.7" pageId="9" pageNumber="194" unit="mm" value="8.7">8.7 mm</quantity>
; width
<quantity id="0A069B4FFF91245FFD13F8966D35F8F2" box="[683,771,1809,1834]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" pageId="9" pageNumber="194" unit="mm" value="2.3">2.3 mm</quantity>
; elytra length
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. Small and comparatively elongated beetle. Head slightly longer than wider (without mandibles), abruptly narrowed anteriorly towards the eyes. Eyes large, almost as long as temples, occiput truncated. Antennae moniliform, slightly longer than pronotum, scape and pedicel 1.5 times wider than rest of antennomeres, last antennomere elongated and rounded at apex. Cervical constriction indistinct. Pronotum as long as wide, rounded at its 3/4 length, then narrowed posteriorly, about 1.4 times as wide as long at posterior edge. Procoxae rounded, with comparatively wide prosternal process. Mesocoxae transverse, about 1.5 times longer than wider. Mesosternum nearly quadrate. Metasternum transverse, about 1.7 times longer than wider, with well expressed longitudinal and transverse sutures. Abdomen with 5 visible flat ventrites not superimposed each others. Last ventrite 1.7 times as long as previous one. Elytra 4.5 times as long as wide, with rounded apex and indistict cells on surface; epipleura rather wide; main veins poorly distinct from the intercalaries; disc of the elytra with numerous small rectangle cells (about 40 per row). Profemora about three times longer than wider at its medium part. Protibiae somewhat longer than profemora. Whole surface of body covered with dense puntuation.
</paragraph>
</subSubSection>
</treatment>
</document>