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<mods:title id="9E61CD2993C3E8F70B7CE7E8BEC5A559">The genusCultrobates (Acari: Oribatida: Ceratokalummidae) in the Galapagos Islands and Central</mods:title>
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<mods:namePart id="F55B1EAC7FAF8BCDF859DDB986A4FAD9">K, Heinrich Schatz</mods:namePart>
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<paragraph id="9C5F5C77FF95FFBEFE24FEED4F59FEED" blockId="16.[491,1040,328,354]" box="[491,1040,328,354]" pageId="16" pageNumber="444">
<heading id="C717EB1BFF95FFBEFE24FEED4F59FEED" bold="true" box="[491,1040,328,354]" fontSize="11" level="2" pageId="16" pageNumber="444" reason="3">
<taxonomicName id="5BE027F4FF95FFBEFE24FEED4F59FEED" ID-CoL="32DSM" authority="Willmann, 1930" authorityName="Willmann" authorityYear="1930" box="[491,1040,328,354]" class="Arachnida" family="Ceratokalummidae" genus="Cultrobates" kingdom="Animalia" order="Sarcoptiformes" pageId="16" pageNumber="444" phylum="Arthropoda" rank="species" species="heterodactylus">
<emphasis id="AE948065FF95FFBEFE24FEED4F59FEED" bold="true" box="[491,1040,328,354]" pageId="16" pageNumber="444">
<emphasis id="AE948065FF95FFBEFE24FEED480FFEED" bold="true" box="[491,838,328,354]" italics="true" pageId="16" pageNumber="444">Cultrobates heterodactylus</emphasis>
<bibRefCitation id="F8712186FF95FFBEFC82FEED4F59FEED" author="Willmann C." box="[845,1040,328,354]" pageId="16" pageNumber="444" pagination="239 - 246" refId="ref14696" refString="Willmann C. 1930. Neue Oribatiden aus Guatemala. Zoologischer Anzeiger, 88 (9 - 10): 239 - 246." type="journal article" year="1930">Willmann, 1930</bibRefCitation>
</emphasis>
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</paragraph>
<paragraph id="9C5F5C77FF95FFBEFE25FED84ECDFE1A" blockId="16.[490,1559,381,1061]" box="[490,1412,381,405]" pageId="16" pageNumber="444">
<bibRefCitation id="F8712186FF95FFBEFE25FED849C3FE1A" author="Willmann C." box="[490,650,381,405]" pageId="16" pageNumber="444" pagination="239 - 246" refId="ref14696" refString="Willmann C. 1930. Neue Oribatiden aus Guatemala. Zoologischer Anzeiger, 88 (9 - 10): 239 - 246." type="journal article" year="1930">Willmann 1930</bibRefCitation>
: p. 242, fig. 4;
<bibRefCitation id="F8712186FF95FFBEFCE7FED848FCFE1A" author="Mahunka S." box="[808,949,381,405]" pageId="16" pageNumber="444" pagination="709 - 724" refId="ref14015" refString="Mahunka S. 1983. Neue und interessante Milben aus dem Genfer Museum. 47. Oribatida Americana 7: Guatemala II (Acari). Revue suisse de Zoologie, 90 (3): 709 - 724. https: // doi. org / 10.5962 / bhl. part. 82009" type="journal article" year="1983">Mahunka 1983</bibRefCitation>
: p. 723, figs 32, 33;
<bibRefCitation id="F8712186FF95FFBEFBBAFED84ECDFE1A" author="Balogh J. &amp; Balogh P." box="[1141,1412,381,405]" pageId="16" pageNumber="444" pagination="333" refId="ref13393" refString="Balogh J., Balogh P. 1990. Oribatid Mites of the Neotropical Region II. In: Balogh J. (Ed.). The soil mites of the world, vol. 3. Amsterdam: Elsevier. pp. 333." type="book chapter" year="1990">Balogh and Balogh 1990:</bibRefCitation>
</paragraph>
</subSubSection>
<subSubSection id="D4FA0FFCFF95FFBEFE24FE3A4805FBAA" pageId="16" pageNumber="444" type="description">
<paragraph id="9C5F5C77FF95FFBEFE24FE3A4EFFFE38" blockId="16.[490,1559,381,1061]" box="[491,1462,415,439]" pageId="16" pageNumber="444">
133, figs 122 D, E;
<bibRefCitation id="F8712186FF95FFBEFD7FFE3A4882FE38" author="Balogh J. &amp; Balogh P." box="[688,971,415,439]" pageId="16" pageNumber="444" pagination="453" refId="ref13472" refString="Balogh J., Balogh P. 2002 a. Identification Keys to the oribatid mites of the Extra-Holarctic Regions, vol. I. Miskolc: Well-Press Publishing Limited. pp. 453." type="book chapter" year="2002">Balogh and Balogh 2002a</bibRefCitation>
: p. 359;
<bibRefCitation id="F8712186FF95FFBEFBD2FE3A4E76FE38" author="Balogh J. &amp; Balogh P." box="[1053,1343,415,439]" pageId="16" pageNumber="444" refId="ref13507" refString="Balogh J., Balogh P. 2002 b. Identification Keys to the oribatid mites of the Extra-Holarctic Regions," type="book" year="2002">Balogh and Balogh 2002b</bibRefCitation>
: fig. 473:6;
</paragraph>
<paragraph id="9C5F5C77FF95FFBEFE24FE6749FAFE55" blockId="16.[490,1559,381,1061]" box="[491,691,450,474]" pageId="16" pageNumber="444">
<bibRefCitation id="F8712186FF95FFBEFE24FE674922FE55" author="Schatz H." box="[491,619,450,474]" pageId="16" pageNumber="444" pagination="209 - 288" refId="ref14403" refString="Schatz H. 2006. Catalogue of known oribatid mite species (Acari: Oribatida) from the Central American landbridge (First part). Tropical Zoology, 19 (2): 209 - 288." type="journal article" year="2006">Schatz 2006</bibRefCitation>
: p. 282.
</paragraph>
<paragraph id="9C5F5C77FF95FFBEFDDCFE41484BFC9E" blockId="16.[490,1559,381,1061]" pageId="16" pageNumber="444">
The original description (
<bibRefCitation id="F8712186FF95FFBEFCE9FE41488CFE73" author="Willmann C." box="[806,965,484,508]" pageId="16" pageNumber="444" pagination="239 - 246" refId="ref14696" refString="Willmann C. 1930. Neue Oribatiden aus Guatemala. Zoologischer Anzeiger, 88 (9 - 10): 239 - 246." type="journal article" year="1930">Willmann 1930</bibRefCitation>
), based on
<specimenCount id="8AE697FEFF95FFBEFB86FE404FAAFE73" box="[1097,1251,484,508]" count="2" pageId="16" pageNumber="444" type="generic">two specimens</specimenCount>
from
<collectingCountry id="E4F71CE7FF95FFBEFAEDFE414EDCFE73" box="[1314,1429,484,508]" name="Guatemala" pageId="16" pageNumber="444">Guatemala</collectingCountry>
is short and lacks information about some morphological structures.
<bibRefCitation id="F8712186FF95FFBEFB4AFDA24E56FD90" author="Mahunka S." box="[1157,1311,519,543]" pageId="16" pageNumber="444" pagination="709 - 724" refId="ref14015" refString="Mahunka S. 1983. Neue und interessante Milben aus dem Genfer Museum. 47. Oribatida Americana 7: Guatemala II (Acari). Revue suisse de Zoologie, 90 (3): 709 - 724. https: // doi. org / 10.5962 / bhl. part. 82009" type="journal article" year="1983">Mahunka (1983)</bibRefCitation>
found another specimen in Tikal,
<collectingCountry id="E4F71CE7FF95FFBEFD7EFD8F486EFDCD" box="[689,807,554,578]" name="Guatemala" pageId="16" pageNumber="444">Guatemala</collectingCountry>
, and provided additional morphological information. I was able to study this specimen. Until further findings in
<collectingCountry id="E4F71CE7FF95FFBEFBCCFDE84F1EFDEB" box="[1027,1111,589,612]" name="Panama" pageId="16" pageNumber="444">Panama</collectingCountry>
(
<bibRefCitation id="F8712186FF95FFBEFBAAFDE94FACFDEB" author="Schatz H." box="[1125,1253,588,612]" pageId="16" pageNumber="444" pagination="209 - 288" refId="ref14403" refString="Schatz H. 2006. Catalogue of known oribatid mite species (Acari: Oribatida) from the Central American landbridge (First part). Tropical Zoology, 19 (2): 209 - 288." type="journal article" year="2006">Schatz 2006</bibRefCitation>
) no other records
<emphasis id="AE948065FF95FFBEFA6FFDE949C7FD08" italics="true" pageId="16" pageNumber="444">
of
<taxonomicName id="5BE027F4FF95FFBEFA74FDE849C7FD08" authorityName="Willmann" authorityYear="1930" class="Arachnida" family="Ceratokalummidae" genus="Cultrobates" kingdom="Animalia" order="Sarcoptiformes" pageId="16" pageNumber="444" phylum="Arthropoda" rank="species" species="heterodactylus">C. heterodactylus</taxonomicName>
</emphasis>
were known (for erroneous and unproven reports see chapter “Corrections to previously stated distribution data”). I found this species in four different sites in the lowlands of Central and Northwest
<collectingCountry id="E4F71CE7FF95FFBEFCDFFD114821FD44" box="[784,872,692,715]" name="Panama" pageId="16" pageNumber="444">Panama</collectingCountry>
. Later, additional reports from
<collectingCountry id="E4F71CE7FF95FFBEFB15FD114E67FD44" box="[1242,1326,692,715]" name="Panama" pageId="16" pageNumber="444">Panama</collectingCountry>
were given by
<bibRefCitation id="F8712186FF95FFBEFE24FD7249FEFD61" author="Basset Y." box="[491,695,726,750]" pageId="16" pageNumber="444" pagination="1481 - 1484" refId="ref13639" refString="Basset Y. et al. 2012. Arthropod Diversity in a Tropical Forest. Science, 338, 1481 - 1484. https: // doi. org / 10.1126 / science. 1226727" type="journal article" year="2012">
Basset
<emphasis id="AE948065FF95FFBEFDF8FD724924FD61" box="[567,621,726,750]" italics="true" pageId="16" pageNumber="444">et al.</emphasis>
(2012)
</bibRefCitation>
. Recently,
<taxonomicName id="5BE027F4FF95FFBEFCFAFD734F13FD61" authorityName="Willmann" authorityYear="1930" box="[821,1114,726,750]" class="Arachnida" family="Ceratokalummidae" genus="Cultrobates" kingdom="Animalia" order="Sarcoptiformes" pageId="16" pageNumber="444" phylum="Arthropoda" rank="species" species="heterodactylus">
<emphasis id="AE948065FF95FFBEFCFAFD734F13FD61" box="[821,1114,726,750]" italics="true" pageId="16" pageNumber="444">Cultrobates heterodactylus</emphasis>
</taxonomicName>
has also been reported from
<collectingCountry id="E4F71CE7FF95FFBEFA5CFD734EADFD61" box="[1427,1508,726,750]" name="Mexico" pageId="16" pageNumber="444">Mexico</collectingCountry>
(
<bibRefCitation id="F8712186FF95FFBEFE3DFD5C49B0FC9E" author="Revelo-Tobar H. &amp; Estrada-Venegas E. G. &amp; Martinez A. E." box="[498,761,761,785]" pageId="16" pageNumber="444" pagination="24 - 38" refId="ref14265" refString="Revelo-Tobar H., Estrada-Venegas E. G., Martinez A. E. 2022. New records of oribatid mites from Mexico with description of a new species. Zootaxa, 5168 (1): 24 - 38. https: // doi. org / 10.11646 / zootaxa. 5168.1.2" type="journal article" year="2022">
Revelo-Tobar
<emphasis id="AE948065FF95FFBEFD43FD5F4988FC9E" box="[652,705,761,785]" italics="true" pageId="16" pageNumber="444">et al.</emphasis>
2022
</bibRefCitation>
).
</paragraph>
<paragraph id="9C5F5C77FF95FFBEFDDCFCBE48E2FC6F" blockId="16.[490,1559,381,1061]" pageId="16" pageNumber="444">
The examined specimens from the populations in
<collectingCountry id="E4F71CE7FF95FFBEFBEDFCB94F3FFCBC" box="[1058,1142,796,819]" name="Panama" pageId="16" pageNumber="444">Panama</collectingCountry>
were morphologically consistent with the specimen from
<collectingCountry id="E4F71CE7FF95FFBEFD11FC9B4818FCD9" box="[734,849,830,854]" name="Guatemala" pageId="16" pageNumber="444">Guatemala</collectingCountry>
and with the descriptions by
<bibRefCitation id="F8712186FF95FFBEFB59FC9B4E09FCD9" author="Willmann C." box="[1174,1344,830,854]" pageId="16" pageNumber="444" pagination="239 - 246" refId="ref14696" refString="Willmann C. 1930. Neue Oribatiden aus Guatemala. Zoologischer Anzeiger, 88 (9 - 10): 239 - 246." type="journal article" year="1930">Willmann (1930)</bibRefCitation>
and
<bibRefCitation id="F8712186FF95FFBEFAB1FC9B496CFCF7" author="Mahunka S." pageId="16" pageNumber="444" pagination="709 - 724" refId="ref14015" refString="Mahunka S. 1983. Neue und interessante Milben aus dem Genfer Museum. 47. Oribatida Americana 7: Guatemala II (Acari). Revue suisse de Zoologie, 90 (3): 709 - 724. https: // doi. org / 10.5962 / bhl. part. 82009" type="journal article" year="1983">Mahunka (1983)</bibRefCitation>
. Some additional morphological features which were not given in detail in previous descriptions are presented here on the basis of specimens from
<collectingCountry id="E4F71CE7FF95FFBEFB6EFC214FB0FC14" box="[1185,1273,900,923]" name="Panama" pageId="16" pageNumber="444">Panama</collectingCountry>
. Comparable speciesspecific differential characters among the species
<taxonomicName id="5BE027F4FF95FFBEFBCFFC034FF3FC32" authorityName="Willmann" authorityYear="1930" box="[1024,1210,933,957]" class="Arachnida" family="Ceratokalummidae" genus="Cultrobates" kingdom="Animalia" order="Sarcoptiformes" pageId="16" pageNumber="444" phylum="Arthropoda" rank="species" species="heterodactylus">
<emphasis id="AE948065FF95FFBEFBCFFC034FF3FC32" box="[1024,1210,933,957]" italics="true" pageId="16" pageNumber="444">C. heterodactylus</emphasis>
</taxonomicName>
,
<taxonomicName id="5BE027F4FF95FFBEFB06FC034E08FC32" authority="K, 2023" authorityName="K" authorityYear="2023" box="[1225,1345,933,957]" class="Arachnida" family="Ceratokalummidae" genus="Cultrobates" kingdom="Animalia" order="Sarcoptiformes" pageId="16" pageNumber="444" phylum="Arthropoda" rank="species" species="ermilovi" status="n. sp.">
<emphasis id="AE948065FF95FFBEFB06FC034E08FC32" box="[1225,1345,933,957]" italics="true" pageId="16" pageNumber="444">C. ermilovi</emphasis>
</taxonomicName>
<emphasis id="AE948065FF95FFBEFA87FC034ECDFC31" bold="true" box="[1352,1412,934,958]" pageId="16" pageNumber="444">
<taxonomicNameLabel id="B5A73D1EFF95FFBEFA87FC034ECDFC31" box="[1352,1412,934,958]" pageId="16" pageNumber="444" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="5BE027F4FF95FFBEFA73FC03497CFC6F" authority="K, 2023" authorityName="K" authorityYear="2023" class="Arachnida" family="Ceratokalummidae" genus="Cultrobates" kingdom="Animalia" order="Sarcoptiformes" pageId="16" pageNumber="444" phylum="Arthropoda" rank="species" species="subiasi" status="n. sp.">
<emphasis id="AE948065FF95FFBEFA73FC03497CFC6F" italics="true" pageId="16" pageNumber="444">C. subiasi</emphasis>
</taxonomicName>
<emphasis id="AE948065FF95FFBEFDF2FC6D4930FC6F" bold="true" box="[573,633,968,992]" pageId="16" pageNumber="444">
<taxonomicNameLabel id="B5A73D1EFF95FFBEFDF2FC6D4930FC6F" box="[573,633,968,992]" pageId="16" pageNumber="444" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
are given in the remark (#1).
</paragraph>
<paragraph id="9C5F5C77FF95FFBEFDDCFC4E4E3DFB8D" blockId="16.[490,1559,381,1061]" box="[531,1396,1003,1027]" pageId="16" pageNumber="444">
<emphasis id="AE948065FF95FFBEFDDCFC4E4932FB8C" box="[531,635,1003,1027]" italics="true" pageId="16" pageNumber="444">Body size</emphasis>
. females (n=25): L 265(277)290, W 153(162)175, males (n=13): L
</paragraph>
<paragraph id="9C5F5C77FF95FFBEFE24FBA84805FBAA" blockId="16.[490,1559,381,1061]" box="[491,844,1037,1061]" pageId="16" pageNumber="444">250(267)270, W 145(152)165.</paragraph>
</subSubSection>
</treatment>
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