771 lines
99 KiB
XML
771 lines
99 KiB
XML
<document id="3786D6CC6108254BA9B42506C0294A5D" ID-DOI="10.11646/zootaxa.4034.3.3" ID-GBIF-Dataset="2a0f9261-82b9-473c-8f0c-e071218395a2" ID-ISSN="1175-5326" ID-Zenodo-Dep="235573" ID-ZooBank="BBA4A435-BE77-4573-B5C3-2746BEE51A60" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461302839758" checkinUser="plazi" docAuthor="Shih, Hsi-Te, Ng, Peter K. L. & Christy, John H." docDate="2015" docId="646187D2FFD370701EDCEE34FB527293" docLanguage="en" docName="zt04034p494.pdf" docOrigin="Zootaxa 4034 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Uca Petruca" docType="treatment" docVersion="8" lastPageNumber="486" masterDocId="9858FFAAFFD6707F1E4BEA72FF697246" masterDocTitle="Uca (Petruca), a new subgenus for the rock fiddler crab Uca panamensis (Stimpson, 1859) from Central America, with comments on some species of the American broad-fronted subgenera" masterLastPageNumber="494" masterPageNumber="471" pageNumber="476" updateTime="1698617574764" updateUser="plazi">
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<mods:title id="5312C7E0A5E9028D4534D08CB76E6B30">Uca (Petruca), a new subgenus for the rock fiddler crab Uca panamensis (Stimpson, 1859) from Central America, with comments on some species of the American broad-fronted subgenera</mods:title>
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<mods:namePart id="7B75A231CF9F2C72B9D7F90FC3E94A3D">Shih, Hsi-Te</mods:namePart>
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<mods:namePart id="5A3BE71BE0067D76823A5F819AC6C8EB">Ng, Peter K. L.</mods:namePart>
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<mods:namePart id="9E43490C904277034EB772A52DDD3EAE">Christy, John H.</mods:namePart>
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<mods:typeOfResource id="4C1543A4B0DCB42D6FF52D357F78DFFA">text</mods:typeOfResource>
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<mods:title id="0B83A4567E633CE7463AA1D5499F8150">Zootaxa</mods:title>
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<mods:date id="960D7D34A753F8579B1B5C2D8AA92EAC">2015</mods:date>
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<mods:number id="A8AD2C958C190685248ABE7FE570F74E">4034</mods:number>
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<treatment id="646187D2FFD370701EDCEE34FB527293" ID-DOI="http://doi.org/10.5281/zenodo.6113941" ID-GBIF-Taxon="119643628" ID-Zenodo-Dep="6113941" LSID="urn:lsid:plazi:treatment:646187D2FFD370701EDCEE34FB527293" httpUri="http://treatment.plazi.org/id/646187D2FFD370701EDCEE34FB527293" lastPageId="15" lastPageNumber="486" pageId="5" pageNumber="476">
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<subSubSection id="A4D2654FFFD3707A1EDCEE34FE8B7626" box="[151,482,1094,1120]" pageId="5" pageNumber="476" type="nomenclature">
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<paragraph id="EC7736C4FFD3707A1EDCEE34FE8B7626" blockId="5.[151,482,1094,1120]" box="[151,482,1094,1120]" pageId="5" pageNumber="476">
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<heading id="B73F81A8FFD3707A1EDCEE34FE8B7626" bold="true" box="[151,482,1094,1120]" fontSize="11" level="1" pageId="5" pageNumber="476" reason="1">
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<emphasis id="DEBCEAD6FFD3707A1EDCEE34FE8B7626" bold="true" box="[151,482,1094,1120]" pageId="5" pageNumber="476">
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<taxonomicName id="2BC84D47FFD3707A1EDCEE34FE287626" ID-CoL="7NXYV" authority="Petruca" authorityName="Petruca" box="[151,321,1094,1120]" class="Malacostraca" family="Ocypodidae" genus="Uca" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="476" phylum="Arthropoda" rank="genus">
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<emphasis id="DEBCEAD6FFD3707A1EDCEE34FFA17626" bold="true" box="[151,200,1094,1120]" italics="true" pageId="5" pageNumber="476">Uca</emphasis>
|
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(
|
||
<emphasis id="DEBCEAD6FFD3707A1E92EE34FE5E7626" bold="true" box="[217,311,1094,1120]" italics="true" pageId="5" pageNumber="476">Petruca</emphasis>
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)
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</taxonomicName>
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subgen. nov.
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</emphasis>
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</heading>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="A4D2654FFFD3707A1EDCEEFCFC4676A6" pageId="5" pageNumber="476" type="reference_group">
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<paragraph id="EC7736C4FFD3707A1EDCEEFCFCF676E5" blockId="5.[151,1138,1165,1248]" box="[151,927,1165,1187]" pageId="5" pageNumber="476">
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<treatmentCitationGroup id="CCD811EAFFD3707A1EDCEEFCFCF676E5" box="[151,927,1165,1187]" pageId="5" pageNumber="476">
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<taxonomicName id="2BC84D47FFD3707A1EDCEEFCFEE076E5" ID-CoL="7PP9B" authority="Bott, 1954: 160" authorityName="Bott" authorityPageNumber="160" authorityYear="1954" box="[151,393,1165,1187]" class="Malacostraca" family="Ocypodidae" genus="Minuca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="476" phylum="Arthropoda" rank="genus">
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<emphasis id="DEBCEAD6FFD3707A1EDCEEFCFF8D76E5" box="[151,228,1166,1187]" italics="true" pageId="5" pageNumber="476">Minuca</emphasis>
|
||
<treatmentCitation id="6D6910D5FFD3707A1EA0EEFCFEE076E5" author="Bott" box="[235,393,1165,1187]" page="160" pageId="5" pageNumber="476" year="1954">
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<bibRefCitation id="88594B35FFD3707A1EA0EEFCFE3376E5" author="Bott" box="[235,346,1165,1187]" pageId="5" pageNumber="476" refString="Bott, R. (1954) Dekapoden (Crustacea) aus El Salvador. 1. Winkerkrabben (Uca). Senckenbergiana Biologica, 35 (3 / 4), 155 - 180." type="journal article" year="1954">Bott, 1954</bibRefCitation>
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||
: 160
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||
</treatmentCitation>
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||
</taxonomicName>
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||
(part);
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<treatmentCitation id="6D6910D5FFD3707A1F9FEEFCFD0576E5" author="Bott" box="[468,620,1165,1187]" page="323" pageId="5" pageNumber="476" year="1973">
|
||
<bibRefCitation id="88594B35FFD3707A1F9FEEFCFD5576E5" author="Bott" box="[468,572,1165,1187]" pageId="5" pageNumber="476" refString="Bott, R. (1973) Die verwandtschaftlichen Beziehungen der Uca Arten. Senckenbergiana Biologica, 54, 315 - 325." type="journal article" year="1973">Bott 1973</bibRefCitation>
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||
: 323
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||
</treatmentCitation>
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||
(part);
|
||
<treatmentCitation id="6D6910D5FFD3707A1CF3EEFFFC0876E5" author="Crane" box="[696,865,1165,1187]" page="154" pageId="5" pageNumber="476" year="1975">
|
||
<bibRefCitation id="88594B35FFD3707A1CF3EEFFFC5876E5" author="Crane" box="[696,817,1165,1187]" pageId="5" pageNumber="476" refString="Crane, J. (1975) Fiddler Crabs of the World (Ocypodidae: Genus Uca). Princeton University Press, Princeton, New Jersey, xxiii + 736 pp." type="book" year="1975">Crane 1975</bibRefCitation>
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||
: 154
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||
</treatmentCitation>
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||
(part)
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||
</treatmentCitationGroup>
|
||
</paragraph>
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||
<paragraph id="EC7736C4FFD3707A1EDCEEDFFC4676A6" blockId="5.[151,1138,1165,1248]" pageId="5" pageNumber="476">
|
||
<treatmentCitationGroup id="CCD811EAFFD3707A1EDCEEDFFC4676A6" pageId="5" pageNumber="476">
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<taxonomicName id="2BC84D47FFD3707A1EDCEEDFFEE47684" ID-CoL="7PMPX" authority="Bott, 1973: 324" authorityName="Bott" authorityPageNumber="324" authorityYear="1973" box="[151,397,1196,1218]" class="Malacostraca" family="Ocypodidae" genus="Leptuca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="476" phylum="Arthropoda" rank="genus">
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||
<emphasis id="DEBCEAD6FFD3707A1EDCEEDFFF807684" box="[151,233,1197,1218]" italics="true" pageId="5" pageNumber="476">Leptuca</emphasis>
|
||
<treatmentCitation id="6D6910D5FFD3707A1EA4EEDFFEE47684" author="Bott" box="[239,397,1196,1218]" page="324" pageId="5" pageNumber="476" year="1973">
|
||
<bibRefCitation id="88594B35FFD3707A1EA4EEDFFE347684" author="Bott" box="[239,349,1196,1218]" pageId="5" pageNumber="476" refString="Bott, R. (1973) Die verwandtschaftlichen Beziehungen der Uca Arten. Senckenbergiana Biologica, 54, 315 - 325." type="journal article" year="1973">Bott, 1973</bibRefCitation>
|
||
: 324
|
||
</treatmentCitation>
|
||
</taxonomicName>
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||
(part); Beinlich & von
|
||
<treatmentCitation id="6D6910D5FFD3707A1C30EEDFFC757684" author="Hagen" box="[635,796,1196,1218]" page="26" pageId="5" pageNumber="476" year="2006">
|
||
<bibRefCitation id="88594B35FFD3707A1C30EEDFFD937684" author="Hagen" box="[635,762,1196,1218]" pageId="5" pageNumber="476" refString="Beinlich, B. & von Hagen, H. O. (2006) Materials for a more stable subdivision of the genus Uca Leach. Zoologische Mededelingen, 80, 9 - 32." type="journal article" year="2006">Hagen 2006</bibRefCitation>
|
||
: 26
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||
</treatmentCitation>
|
||
(part); Ng
|
||
<emphasis id="DEBCEAD6FFD3707A1DC5EEDFFCA87684" box="[910,961,1196,1218]" italics="true" pageId="5" pageNumber="476">et al.</emphasis>
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||
2008: 241 (part).
|
||
<taxonomicName id="2BC84D47FFD3707A1EDCEEB8FEFA76A6" ID-CoL="442cf5d0-c970-418e-b256-3cf9823a9c9f" authority="Crane, 1975: 211" authorityName="Crane" authorityPageNumber="211" authorityYear="1975" box="[151,403,1226,1248]" class="Malacostraca" family="Ocypodidae" genus="Celuca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="476" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="DEBCEAD6FFD3707A1EDCEEB8FFB676A6" box="[151,223,1226,1248]" italics="true" pageId="5" pageNumber="476">Celuca</emphasis>
|
||
<treatmentCitation id="6D6910D5FFD3707A1EAEEEB8FEFA76A6" author="Crane" box="[229,403,1226,1248]" page="211" pageId="5" pageNumber="476" year="1975">
|
||
<bibRefCitation id="88594B35FFD3707A1EAEEEB8FE0C76A6" author="Crane" box="[229,357,1226,1248]" pageId="5" pageNumber="476" refString="Crane, J. (1975) Fiddler Crabs of the World (Ocypodidae: Genus Uca). Princeton University Press, Princeton, New Jersey, xxiii + 736 pp." type="book" year="1975">Crane, 1975</bibRefCitation>
|
||
: 211
|
||
</treatmentCitation>
|
||
</taxonomicName>
|
||
(part);
|
||
<treatmentCitation id="6D6910D5FFD3707A1F94EEB8FDD076A6" author="Rosenberg" box="[479,697,1226,1248]" page="848" pageId="5" pageNumber="476" year="2001">
|
||
<bibRefCitation id="88594B35FFD3707A1F94EEB8FDEE76A6" author="Rosenberg" box="[479,647,1226,1248]" pageId="5" pageNumber="476" refString="Rosenberg, M. S. (2001) The systematics and taxonomy of fiddler crabs: a phylogeny of the genus Uca. Journal of Crustacean Biology, 21, 839 - 869. http: // dx. doi. org / 10.1163 / 20021975 - 99990176" type="journal article" year="2001">Rosenberg 2001</bibRefCitation>
|
||
: 848
|
||
</treatmentCitation>
|
||
, 852 (part).
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="A4D2654FFFD3707A1EDCEF7FFB6B7760" box="[151,1026,1293,1318]" pageId="5" pageNumber="476" type="materials_examined">
|
||
<paragraph id="EC7736C4FFD3707A1EDCEF7FFB6B7760" blockId="5.[151,1437,1293,2038]" box="[151,1026,1293,1318]" pageId="5" pageNumber="476">
|
||
<emphasis id="DEBCEAD6FFD3707A1EDCEF7FFE427760" bold="true" box="[151,299,1293,1318]" pageId="5" pageNumber="476">
|
||
<typeStatus id="33738866FFD3707A1EDCEF7FFFBB7760" box="[151,210,1293,1318]" pageId="5" pageNumber="476">Type</typeStatus>
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||
species
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||
</emphasis>
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||
.
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||
<taxonomicName id="2BC84D47FFD3707A1F72EF7FFD9A7760" authority="Stimpson, 1859" authorityName="Stimpson" authorityYear="1859" box="[313,755,1293,1318]" class="Malacostraca" family="Ocypodidae" genus="Gelasimus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="476" phylum="Arthropoda" rank="species" species="panamensis">
|
||
<emphasis id="DEBCEAD6FFD3707A1F72EF7FFD537763" box="[313,570,1293,1318]" italics="true" pageId="5" pageNumber="476">Gelasimus panamensis</emphasis>
|
||
<bibRefCitation id="88594B35FFD3707A1C0AEF7FFD9A7760" author="Stimpson" box="[577,755,1293,1318]" pageId="5" pageNumber="476" refString="Stimpson, W. (1859) Notes on North American Crustacea, No. 1. Annals of the Lyceum of Natural History of New York, 7 (4 - 5), 49 - 93, pl. I." type="journal article" year="1859">Stimpson, 1859</bibRefCitation>
|
||
</taxonomicName>
|
||
, by present designation.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="A4D2654FFFD3707A1E8CEF43FCE97728" pageId="5" pageNumber="476" type="etymology">
|
||
<paragraph id="EC7736C4FFD3707A1E8CEF43FCE97728" blockId="5.[151,1437,1293,2038]" pageId="5" pageNumber="476">
|
||
<emphasis id="DEBCEAD6FFD3707A1E8CEF43FE2F770C" bold="true" box="[199,326,1329,1354]" pageId="5" pageNumber="476">Etymology</emphasis>
|
||
. From the Greek
|
||
<emphasis id="DEBCEAD6FFD3707A1C5FEF40FD26770F" box="[532,591,1330,1353]" italics="true" pageId="5" pageNumber="476">petra</emphasis>
|
||
for “rock or stone”, for the stony habitat (
|
||
<figureCitation id="74F32A41FFD3707A1A73EF43FB12770C" box="[1080,1147,1329,1354]" captionStart="FIGURE 8" captionStartId="13.[151,250,1719,1741]" captionTargetBox="[163,1425,205,1689]" captionTargetId="figure@13.[151,1436,193,1698]" captionTargetPageId="13" captionText="FIGURE 8. Uca panamensis ♂ (A, B) and its habitat (C). Photographs taken in Panamá. B, courtesy of P. Backwell." httpUri="https://zenodo.org/record/235581/files/figure.png" pageId="5" pageNumber="476">Fig. 8</figureCitation>
|
||
C) of the
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||
<typeStatus id="33738866FFD3707A1AA7EF40FA75770C" box="[1260,1308,1330,1354]" pageId="5" pageNumber="476">type</typeStatus>
|
||
species, in arbitrary combination with the genus name
|
||
<taxonomicName id="2BC84D47FFD3707A1C34EF24FDC7772B" box="[639,686,1366,1389]" class="Malacostraca" family="Ocypodidae" genus="Uca" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="476" phylum="Arthropoda" rank="genus">
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||
<emphasis id="DEBCEAD6FFD3707A1C34EF24FDC7772B" box="[639,686,1366,1389]" italics="true" pageId="5" pageNumber="476">Uca</emphasis>
|
||
</taxonomicName>
|
||
. Gender feminine.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="A4D2654FFFD3707A1E8CEF0BFAF475CC" pageId="5" pageNumber="476" type="diagnosis">
|
||
<paragraph id="EC7736C4FFD3707A1E8CEF0BFAF475CC" blockId="5.[151,1437,1293,2038]" pageId="5" pageNumber="476">
|
||
<emphasis id="DEBCEAD6FFD3707A1E8CEF0BFE5377D4" bold="true" box="[199,314,1401,1426]" pageId="5" pageNumber="476">Diagnosis</emphasis>
|
||
. Carapace flat, widest distance of carapace between anterolateral angles, angles acute, produced obliquely outwards, anteriorly; anterolateral margins relatively long (
|
||
<figureCitation id="74F32A41FFD3707A1DE7EFEFFC8477F0" box="[940,1005,1437,1462]" captionStart="FIGURE 1" captionStartId="6.[151,250,1911,1933]" captionTargetBox="[163,1423,215,1877]" captionTargetId="figure@6.[151,1436,193,1890]" captionTargetPageId="6" captionText="FIGURE 1. Uca panamensis (neotype, USNM 1294205, CW 16.8 mm). A, dorsal view; B, boss on carapace behind right dorsolateral margin (arrowed); C, ventral view. Scale, B = 2 mm." httpUri="https://zenodo.org/record/235574/files/figure.png" pageId="5" pageNumber="476">Fig. 1</figureCitation>
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||
A); boss (raised tuberculate ridge) on carapace behind dorsolateral margin (
|
||
<figureCitation id="74F32A41FFD3707A1C1BEFB3FDFD779C" box="[592,660,1473,1498]" captionStart="FIGURE 1" captionStartId="6.[151,250,1911,1933]" captionTargetBox="[163,1423,215,1877]" captionTargetId="figure@6.[151,1436,193,1890]" captionTargetPageId="6" captionText="FIGURE 1. Uca panamensis (neotype, USNM 1294205, CW 16.8 mm). A, dorsal view; B, boss on carapace behind right dorsolateral margin (arrowed); C, ventral view. Scale, B = 2 mm." httpUri="https://zenodo.org/record/235574/files/figure.png" pageId="5" pageNumber="476">Fig. 1</figureCitation>
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||
B); front wide, about 1/3 carapace width (
|
||
<figureCitation id="74F32A41FFD3707A1AC5EFB3FBBF779C" box="[1166,1238,1473,1498]" captionStart="FIGURE 2" captionStartId="7.[151,250,1925,1947]" captionTargetBox="[159,1425,203,1891]" captionTargetId="figure@7.[151,1436,193,1903]" captionTargetPageId="7" captionText="FIGURE 2. Uca panamensis (neotype, USNM 1294205, CW 16.8 mm). A, orbital region; B, inner side of orbital region, showing the tubercle of orbital armature (arrowed); C, frontal view; D, front; E, third maxillipeds. Scale, A, E = 2 mm; B = 1 mm; C = 10 mm; D = 5 mm." httpUri="https://zenodo.org/record/235575/files/figure.png" pageId="5" pageNumber="476">Fig. 2</figureCitation>
|
||
C, D); suborbital crenulations strong, each truncate, separate, increasing in size towards antero-external angle (
|
||
<figureCitation id="74F32A41FFD3707A1A98EF97FA4877B8" box="[1235,1313,1509,1534]" captionStart="FIGURE 1" captionStartId="6.[151,250,1911,1933]" captionTargetBox="[163,1423,215,1877]" captionTargetId="figure@6.[151,1436,193,1890]" captionTargetPageId="6" captionText="FIGURE 1. Uca panamensis (neotype, USNM 1294205, CW 16.8 mm). A, dorsal view; B, boss on carapace behind right dorsolateral margin (arrowed); C, ventral view. Scale, B = 2 mm." httpUri="https://zenodo.org/record/235574/files/figure.png" pageId="5" pageNumber="476">Figs. 1</figureCitation>
|
||
C, 2A–C); orbital floor without mound or pile, but with tubercle at inner corner adjacent to antennule (
|
||
<figureCitation id="74F32A41FFD3707A1AD0EC7BFBB47464" box="[1179,1245,1545,1570]" captionStart="FIGURE 2" captionStartId="7.[151,250,1925,1947]" captionTargetBox="[159,1425,203,1891]" captionTargetId="figure@7.[151,1436,193,1903]" captionTargetPageId="7" captionText="FIGURE 2. Uca panamensis (neotype, USNM 1294205, CW 16.8 mm). A, orbital region; B, inner side of orbital region, showing the tubercle of orbital armature (arrowed); C, frontal view; D, front; E, third maxillipeds. Scale, A, E = 2 mm; B = 1 mm; C = 10 mm; D = 5 mm." httpUri="https://zenodo.org/record/235575/files/figure.png" pageId="5" pageNumber="476">Fig. 2</figureCitation>
|
||
A–C). Male with pleonal clasping apparatus in abdominal cavity. Urocardiac ossicles of gastric mill (
|
||
<figureCitation id="74F32A41FFD3707A1A1DEC5FFBF37400" box="[1110,1178,1581,1606]" captionStart="FIGURE 5" captionStartId="10.[151,250,1927,1949]" captionTargetBox="[159,1427,205,1895]" captionTargetId="figure@10.[151,1436,193,1906]" captionTargetPageId="10" captionText="FIGURE 5. Urocardiac ossicle of Uca panamensis. A, B, neotype, USNM 1294205 (16.8 mm); C, ZRC (CW 12.6 mm, ♂); D, ZRC (CW 13.7 mm, ♀). A, C, D, ventral view; B, lateroventral view. Scale, A, B = 1 mm; C, D = 0.5 mm." httpUri="https://zenodo.org/record/235578/files/figure.png" pageId="5" pageNumber="476">Fig. 5</figureCitation>
|
||
) simple, median tooth with 3 pairs of similar transverse ridges, separated by gaps, not reaching central ridge on posterior tooth plate; first pair of ridges strongest, next 2 lower, with 2 or 3 weak pairs of cusps on stem region; stem region with long, wide median part (
|
||
<figureCitation id="74F32A41FFD3707A1F60ECEBFE0474F4" box="[299,365,1689,1714]" captionStart="FIGURE 5" captionStartId="10.[151,250,1927,1949]" captionTargetBox="[159,1427,205,1895]" captionTargetId="figure@10.[151,1436,193,1906]" captionTargetPageId="10" captionText="FIGURE 5. Urocardiac ossicle of Uca panamensis. A, B, neotype, USNM 1294205 (16.8 mm); C, ZRC (CW 12.6 mm, ♂); D, ZRC (CW 13.7 mm, ♀). A, C, D, ventral view; B, lateroventral view. Scale, A, B = 1 mm; C, D = 0.5 mm." httpUri="https://zenodo.org/record/235578/files/figure.png" pageId="5" pageNumber="476">Fig. 5</figureCitation>
|
||
A).
|
||
<taxonomicName id="2BC84D47FFD3707A1FD2ECEBFEB474F4" box="[409,477,1689,1714]" class="Malacostraca" family="Ocypodidae" genus="Uca" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="493" phylum="Arthropoda" rank="species" species="major">Major</taxonomicName>
|
||
pollex, dactylus with tips blunt, without or with weak tooth on cutting margins (
|
||
<figureCitation id="74F32A41FFD3707A1B2FECEBFFCD7490" captionStart="FIGURE 3" captionStartId="8.[151,250,1887,1909]" captionTargetBox="[161,1423,207,1857]" captionTargetId="figure@8.[151,1436,193,1866]" captionTargetPageId="8" captionText="FIGURE 3. Major and minor chelipeds of Uca panamensis (neotype, USNM 1294205, CW 16.8 mm). A, B, outer and inner views of major cheliped; C, D, outer and inner views of minor cheliped. Scale, C, D = 2 mm." httpUri="https://zenodo.org/record/235576/files/figure.png" pageId="5" pageNumber="476">Figs. 3</figureCitation>
|
||
A, B, 8A, B). Adult male
|
||
<taxonomicName id="2BC84D47FFD3707A1F9BECCFFD787490" box="[464,529,1725,1750]" class="Malacostraca" family="Ocypodidae" genus="Uca" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="476" phylum="Arthropoda" rank="species" species="major">major</taxonomicName>
|
||
manus with outer surface smooth, proximal outer part disproportionately thick, with posterior extension beyond distal part of carpus (
|
||
<figureCitation id="74F32A41FFD3707A1D42EC93FC3174BC" box="[777,856,1761,1786]" captionStart="FIGURE 3" captionStartId="8.[151,250,1887,1909]" captionTargetBox="[161,1423,207,1857]" captionTargetId="figure@8.[151,1436,193,1866]" captionTargetPageId="8" captionText="FIGURE 3. Major and minor chelipeds of Uca panamensis (neotype, USNM 1294205, CW 16.8 mm). A, B, outer and inner views of major cheliped; C, D, outer and inner views of minor cheliped. Scale, C, D = 2 mm." httpUri="https://zenodo.org/record/235576/files/figure.png" pageId="5" pageNumber="476">Figs. 3</figureCitation>
|
||
A, 8A); inner surface with vestigial oblique ridge, without or with low, rounded tubercles (
|
||
<figureCitation id="74F32A41FFD3707A1C1CED77FDF37558" box="[599,666,1797,1822]" captionStart="FIGURE 3" captionStartId="8.[151,250,1887,1909]" captionTargetBox="[161,1423,207,1857]" captionTargetId="figure@8.[151,1436,193,1866]" captionTargetPageId="8" captionText="FIGURE 3. Major and minor chelipeds of Uca panamensis (neotype, USNM 1294205, CW 16.8 mm). A, B, outer and inner views of major cheliped; C, D, outer and inner views of minor cheliped. Scale, C, D = 2 mm." httpUri="https://zenodo.org/record/235576/files/figure.png" pageId="5" pageNumber="476">Fig. 3</figureCitation>
|
||
B). Adult minor cheliped with manus rounded, broad almost as long; gape wide; fingers with numerous long stiff setae on tip (
|
||
<figureCitation id="74F32A41FFD3707A1D53ED5BFC0F7504" box="[792,870,1833,1858]" captionStart="FIGURE 3" captionStartId="8.[151,250,1887,1909]" captionTargetBox="[161,1423,207,1857]" captionTargetId="figure@8.[151,1436,193,1866]" captionTargetPageId="8" captionText="FIGURE 3. Major and minor chelipeds of Uca panamensis (neotype, USNM 1294205, CW 16.8 mm). A, B, outer and inner views of major cheliped; C, D, outer and inner views of minor cheliped. Scale, C, D = 2 mm." httpUri="https://zenodo.org/record/235576/files/figure.png" pageId="5" pageNumber="476">Figs. 3</figureCitation>
|
||
C, D, 8B). G1 (
|
||
<figureCitation id="74F32A41FFD3707A1A58ED5BFB3E7504" box="[1043,1111,1833,1858]" captionStart="FIGURE 4" captionStartId="9.[151,250,1917,1939]" captionTargetBox="[159,1422,203,1887]" captionTargetId="figure@9.[151,1436,193,1896]" captionTargetPageId="9" captionText="FIGURE 4. Uca panamensis, right G 1 (neotype, USNM 1294205, CW 16.8 mm). A, dorsal view; B, dorsal view of terminal segment; C, D, ventral view of terminal segment. Scale, A = 2 mm; B, C = 0.5 mm." httpUri="https://zenodo.org/record/235577/files/figure.png" pageId="5" pageNumber="476">Fig. 4</figureCitation>
|
||
) with anterior flange (before genital pore) broader than posterior one (after genital pore); inner process large, broad, becoming thicker distally, bent forwards at right angles, extending beyond tip of posterior flange; thumb slender, ending below base of flange.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="A4D2654FFFD370701E8CEDE7FB527293" lastPageId="15" lastPageNumber="486" pageId="5" pageNumber="476" type="discussion">
|
||
<paragraph id="EC7736C4FFD3707A1E8CEDE7FAE475B0" blockId="5.[151,1437,1293,2038]" pageId="5" pageNumber="476">
|
||
<emphasis id="DEBCEAD6FFD3707A1E8CEDE7FE5D75E8" bold="true" box="[199,308,1941,1966]" pageId="5" pageNumber="476">Remarks</emphasis>
|
||
. One of the diagnostic characters of
|
||
<emphasis id="DEBCEAD6FFD3707A1C99EDE4FC4275EB" box="[722,811,1942,1965]" italics="true" pageId="5" pageNumber="476">Petruca</emphasis>
|
||
<emphasis id="DEBCEAD6FFD3707A1D79EDE7FCAC75E8" bold="true" box="[818,965,1941,1966]" pageId="5" pageNumber="476">subgen. nov.</emphasis>
|
||
is that the carapace is widest between the tips of the anterolateral angles (
|
||
<figureCitation id="74F32A41FFD3707A1FB7EDCBFD557594" box="[508,572,1977,2002]" captionStart="FIGURE 1" captionStartId="6.[151,250,1911,1933]" captionTargetBox="[163,1423,215,1877]" captionTargetId="figure@6.[151,1436,193,1890]" captionTargetPageId="6" captionText="FIGURE 1. Uca panamensis (neotype, USNM 1294205, CW 16.8 mm). A, dorsal view; B, boss on carapace behind right dorsolateral margin (arrowed); C, ventral view. Scale, B = 2 mm." httpUri="https://zenodo.org/record/235574/files/figure.png" pageId="5" pageNumber="476">Fig. 1</figureCitation>
|
||
A). In this feature, it is similar to, but more pronounced than in any
|
||
<taxonomicName id="2BC84D47FFD3707A1B09EDC8FAF47597" box="[1346,1437,1978,2001]" class="Malacostraca" family="Ocypodidae" genus="Leptuca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="476" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="DEBCEAD6FFD3707A1B09EDC8FAF47597" box="[1346,1437,1978,2001]" italics="true" pageId="5" pageNumber="476">Leptuca</emphasis>
|
||
</taxonomicName>
|
||
species. The maximum carapace width in
|
||
<taxonomicName id="2BC84D47FFD3707A1C26EDACFDAA75B3" box="[621,707,2014,2037]" class="Malacostraca" family="Ocypodidae" genus="Minuca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="476" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="DEBCEAD6FFD3707A1C26EDACFDAA75B3" box="[621,707,2014,2037]" italics="true" pageId="5" pageNumber="476">Minuca</emphasis>
|
||
</taxonomicName>
|
||
species is usually behind the anterolateral angles (
|
||
<bibRefCitation id="88594B35FFD3707A1AB2EDACFAE975B0" author="Crane" box="[1273,1408,2013,2038]" pageId="5" pageNumber="476" refString="Crane, J. (1975) Fiddler Crabs of the World (Ocypodidae: Genus Uca). Princeton University Press, Princeton, New Jersey, xxiii + 736 pp." type="book" year="1975">Crane 1975</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="B8B7664CFFD070791EDCED05FC4375EA" httpUri="https://zenodo.org/record/235574/files/figure.png" pageId="6" pageNumber="477" targetBox="[163,1423,215,1877]" targetPageId="6">
|
||
<paragraph id="EC7736C4FFD070791EDCED05FC4375EA" blockId="6.[151,1436,1911,1964]" pageId="6" pageNumber="477">
|
||
<emphasis id="DEBCEAD6FFD070791EDCED05FE6675CA" bold="true" box="[151,271,1911,1933]" pageId="6" pageNumber="477">FIGURE 1</emphasis>
|
||
.
|
||
<taxonomicName id="2BC84D47FFD070791F55ED0AFEA075CB" box="[286,457,1912,1933]" class="Malacostraca" family="Ocypodidae" genus="Uca" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="477" phylum="Arthropoda" rank="species" species="panamensis">
|
||
<emphasis id="DEBCEAD6FFD070791F55ED0AFEA075CB" box="[286,457,1912,1933]" italics="true" pageId="6" pageNumber="477">Uca panamensis</emphasis>
|
||
</taxonomicName>
|
||
(neotype, USNM 1294205, CW 16.8 mm). A, dorsal view; B, boss on carapace behind right dorsolateral margin (arrowed); C, ventral view. Scale, B = 2 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="B8B7664CFFD170781EDCEDF7FED3759E" httpUri="https://zenodo.org/record/235575/files/figure.png" pageId="7" pageNumber="478" targetBox="[159,1425,203,1891]" targetPageId="7">
|
||
<paragraph id="EC7736C4FFD170781EDCEDF7FED3759E" blockId="7.[151,1436,1924,2008]" pageId="7" pageNumber="478">
|
||
<emphasis id="DEBCEAD6FFD170781EDCEDF7FE7975DC" bold="true" box="[151,272,1925,1947]" pageId="7" pageNumber="478">FIGURE 2</emphasis>
|
||
.
|
||
<taxonomicName id="2BC84D47FFD170781F54EDF4FEA375DD" box="[287,458,1926,1947]" class="Malacostraca" family="Ocypodidae" genus="Uca" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="478" phylum="Arthropoda" rank="species" species="panamensis">
|
||
<emphasis id="DEBCEAD6FFD170781F54EDF4FEA375DD" box="[287,458,1926,1947]" italics="true" pageId="7" pageNumber="478">Uca panamensis</emphasis>
|
||
</taxonomicName>
|
||
(neotype, USNM 1294205, CW 16.8 mm). A, orbital region; B, inner side of orbital region, showing the tubercle of orbital armature (arrowed); C, frontal view; D, front; E, third maxillipeds. Scale, A, E = 2 mm; B = 1 mm; C = 10 mm; D = 5 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="B8B7664CFFDE70771EDCED2DFB2F75D2" httpUri="https://zenodo.org/record/235576/files/figure.png" pageId="8" pageNumber="479" targetBox="[161,1423,207,1857]" targetPageId="8">
|
||
<paragraph id="EC7736C4FFDE70771EDCED2DFB2F75D2" blockId="8.[151,1435,1887,1940]" pageId="8" pageNumber="479">
|
||
<emphasis id="DEBCEAD6FFDE70771EDCED2DFE677532" bold="true" box="[151,270,1887,1909]" pageId="8" pageNumber="479">FIGURE 3</emphasis>
|
||
.
|
||
<taxonomicName id="2BC84D47FFDE70771F50ED2DFE307533" box="[283,345,1887,1909]" class="Malacostraca" family="Ocypodidae" genus="Uca" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="493" phylum="Arthropoda" rank="species" species="major">Major</taxonomicName>
|
||
and minor chelipeds of
|
||
<taxonomicName id="2BC84D47FFDE70771C18ED12FD947533" box="[595,765,1888,1909]" class="Malacostraca" family="Ocypodidae" genus="Uca" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="479" phylum="Arthropoda" rank="species" species="panamensis">
|
||
<emphasis id="DEBCEAD6FFDE70771C18ED12FD947533" box="[595,765,1888,1909]" italics="true" pageId="8" pageNumber="479">Uca panamensis</emphasis>
|
||
</taxonomicName>
|
||
(neotype, USNM 1294205, CW 16.8 mm). A, B, outer and inner views of
|
||
<taxonomicName id="2BC84D47FFDE70771EBFED0CFE4675D2" box="[244,303,1918,1940]" class="Malacostraca" family="Ocypodidae" genus="Uca" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="479" phylum="Arthropoda" rank="species" species="major">major</taxonomicName>
|
||
cheliped; C, D, outer and inner views of minor cheliped. Scale, C, D = 2 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="B8B7664CFFDF70761EDCED0FFC8F75F4" httpUri="https://zenodo.org/record/235577/files/figure.png" pageId="9" pageNumber="480" targetBox="[159,1422,203,1887]" targetPageId="9">
|
||
<paragraph id="EC7736C4FFDF70761EDCED0FFC8F75F4" blockId="9.[151,1435,1917,1970]" pageId="9" pageNumber="480">
|
||
<emphasis id="DEBCEAD6FFDF70761EDCED0FFE6475D4" bold="true" box="[151,269,1917,1939]" pageId="9" pageNumber="480">FIGURE 4</emphasis>
|
||
.
|
||
<taxonomicName id="2BC84D47FFDF70761F50ED0CFEAD75D5" box="[283,452,1918,1939]" class="Malacostraca" family="Ocypodidae" genus="Uca" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="480" phylum="Arthropoda" rank="species" species="panamensis">
|
||
<emphasis id="DEBCEAD6FFDF70761F50ED0CFEAD75D5" box="[283,452,1918,1939]" italics="true" pageId="9" pageNumber="480">Uca panamensis</emphasis>
|
||
</taxonomicName>
|
||
, right G1 (neotype, USNM 1294205, CW 16.8 mm). A, dorsal view; B, dorsal view of terminal segment; C, D, ventral view of terminal segment. Scale, A = 2 mm; B, C = 0.5 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="B8B7664CFFDC70751EDCEDF5FBB975FA" httpUri="https://zenodo.org/record/235578/files/figure.png" pageId="10" pageNumber="481" targetBox="[159,1427,205,1895]" targetPageId="10">
|
||
<paragraph id="EC7736C4FFDC70751EDCEDF5FBB975FA" blockId="10.[151,1435,1927,1980]" pageId="10" pageNumber="481">
|
||
<emphasis id="DEBCEAD6FFDC70751EDCEDF5FE6575DB" bold="true" box="[151,268,1927,1949]" pageId="10" pageNumber="481">FIGURE 5</emphasis>
|
||
. Urocardiac ossicle of
|
||
<taxonomicName id="2BC84D47FFDC70751FBEEDFBFDF475D8" box="[501,669,1929,1950]" class="Malacostraca" family="Ocypodidae" genus="Uca" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="481" phylum="Arthropoda" rank="species" species="panamensis">
|
||
<emphasis id="DEBCEAD6FFDC70751FBEEDFBFDF475D8" box="[501,669,1929,1950]" italics="true" pageId="10" pageNumber="481">Uca panamensis</emphasis>
|
||
</taxonomicName>
|
||
. A, B, neotype, USNM 1294205 (16.8 mm); C, ZRC (CW 12.6 mm, ♂); D, ZRC (CW 13.7 mm, ♀). A, C, D, ventral view; B, lateroventral view. Scale, A, B = 1 mm; C, D = 0.5 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="B8B7664CFFDD70741EDCED2CFCDB75A9" httpUri="https://zenodo.org/record/235579/files/figure.png" pageId="11" pageNumber="482" targetBox="[173,1413,207,1861]" targetPageId="11">
|
||
<paragraph id="EC7736C4FFDD70741EDCED2CFCDB75A9" blockId="11.[151,1437,1886,2031]" pageId="11" pageNumber="482">
|
||
<emphasis id="DEBCEAD6FFDD70741EDCED2CFE677535" bold="true" box="[151,270,1886,1908]" pageId="11" pageNumber="482">FIGURE 6</emphasis>
|
||
. Urocardiac ossicle of the subgenus
|
||
<taxonomicName id="2BC84D47FFDD70741CC3ED2DFDBC7532" box="[648,725,1887,1908]" class="Malacostraca" family="Ocypodidae" genus="Minuca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="482" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="DEBCEAD6FFDD70741CC3ED2DFDBC7532" box="[648,725,1887,1908]" italics="true" pageId="11" pageNumber="482">Minuca</emphasis>
|
||
</taxonomicName>
|
||
species. A,
|
||
<taxonomicName id="2BC84D47FFDD70741D1FED2DFC857532" box="[852,1004,1886,1908]" class="Malacostraca" family="Ocypodidae" genus="Uca" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="482" phylum="Arthropoda" rank="species" species="brevifrons">
|
||
<emphasis id="DEBCEAD6FFDD70741D1FED2DFC857532" box="[852,1004,1886,1908]" italics="true" pageId="11" pageNumber="482">Uca brevifrons</emphasis>
|
||
</taxonomicName>
|
||
(ZRC 2012.0126, CW 22.1, ♂); B,
|
||
<taxonomicName id="2BC84D47FFDD70741B29ED2DFFBD75D5" class="Malacostraca" family="Ocypodidae" genus="Uca" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="482" phylum="Arthropoda" rank="species" species="virens">
|
||
<emphasis id="DEBCEAD6FFDD70741B29ED2DFA1D7532" box="[1378,1396,1887,1908]" italics="true" pageId="11" pageNumber="482">U</emphasis>
|
||
. cf.
|
||
<emphasis id="DEBCEAD6FFDD70741EDCED0CFFBD75D5" box="[151,212,1918,1939]" italics="true" pageId="11" pageNumber="482">virens</emphasis>
|
||
</taxonomicName>
|
||
(NCHUZOOL 13946, CW 17.0 mm, ♂); C,
|
||
<taxonomicName id="2BC84D47FFDD70741CF0ED0CFC2F75D5" box="[699,838,1918,1939]" class="Malacostraca" family="Ocypodidae" genus="Uca" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="482" phylum="Arthropoda" rank="species" species="victoriana">
|
||
<emphasis id="DEBCEAD6FFDD70741CF0ED0CFC2F75D5" box="[699,838,1918,1939]" italics="true" pageId="11" pageNumber="482">U. victoriana</emphasis>
|
||
</taxonomicName>
|
||
(NCHUZOOL 13945, CW 12.2 mm, ♀); D,
|
||
<taxonomicName id="2BC84D47FFDD70741B65ED0CFAF575D5" box="[1326,1436,1918,1939]" class="Malacostraca" family="Ocypodidae" genus="Uca" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="482" phylum="Arthropoda" rank="species" species="pugnax">
|
||
<emphasis id="DEBCEAD6FFDD70741B65ED0CFAF575D5" box="[1326,1436,1918,1939]" italics="true" pageId="11" pageNumber="482">U. pugnax</emphasis>
|
||
</taxonomicName>
|
||
(NCHUZOOL 13941, CW 15.2 mm, ♂); E,
|
||
<taxonomicName id="2BC84D47FFDD70741C24EDEFFD8F75F4" box="[623,742,1948,1970]" class="Malacostraca" family="Ocypodidae" genus="Uca" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="482" phylum="Arthropoda" rank="species" species="burgersi">
|
||
<emphasis id="DEBCEAD6FFDD70741C24EDEFFD8F75F4" box="[623,742,1948,1970]" italics="true" pageId="11" pageNumber="482">U. burgersi</emphasis>
|
||
</taxonomicName>
|
||
(NCHUZOOL 13956, CW 13.0 mm); F,
|
||
<taxonomicName id="2BC84D47FFDD70741AEAEDEFFB9675F4" box="[1185,1279,1949,1970]" class="Malacostraca" family="Ocypodidae" genus="Uca" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="482" phylum="Arthropoda" rank="species" species="rapax">
|
||
<emphasis id="DEBCEAD6FFDD70741AEAEDEFFB9675F4" box="[1185,1279,1949,1970]" italics="true" pageId="11" pageNumber="482">U. rapax</emphasis>
|
||
</taxonomicName>
|
||
(NCHUZOOL 13942, CW 23.4 mm, ♂); G,
|
||
<taxonomicName id="2BC84D47FFDD70741FF4EDC9FD437596" box="[447,554,1978,2000]" class="Malacostraca" family="Ocypodidae" genus="Uca" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="482" phylum="Arthropoda" rank="species" species="mordax">
|
||
<emphasis id="DEBCEAD6FFDD70741FF4EDC9FD437596" box="[447,554,1978,2000]" italics="true" pageId="11" pageNumber="482">U. mordax</emphasis>
|
||
</taxonomicName>
|
||
(NCHUZOOL 13940, CW 18.0 mm, ♀); H,
|
||
<taxonomicName id="2BC84D47FFDD70741DBDEDC9FB3A7596" box="[1014,1107,1979,2000]" class="Malacostraca" family="Ocypodidae" genus="Uca" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="482" phylum="Arthropoda" rank="species" species="minax">
|
||
<emphasis id="DEBCEAD6FFDD70741DBDEDC9FB3A7596" box="[1014,1107,1979,2000]" italics="true" pageId="11" pageNumber="482">U. minax</emphasis>
|
||
</taxonomicName>
|
||
(NCHUZOOL 13957, CW 23.2 mm, ♂); I,
|
||
<taxonomicName id="2BC84D47FFDD70741F42EDA8FE1C75A9" box="[265,373,2010,2031]" class="Malacostraca" family="Ocypodidae" genus="Uca" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="482" phylum="Arthropoda" rank="species" species="vocator">
|
||
<emphasis id="DEBCEAD6FFDD70741F42EDA8FE1C75A9" box="[265,373,2010,2031]" italics="true" pageId="11" pageNumber="482">U. vocator</emphasis>
|
||
</taxonomicName>
|
||
(NCHUZOOL 13948, CW 21.9 mm, ♀). Scale = 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="B8B7664CFFDA70731EDCED28FB9D75AC" httpUri="https://zenodo.org/record/235580/files/figure.png" pageId="12" pageNumber="483" targetBox="[176,1408,203,1846]" targetPageId="12">
|
||
<paragraph id="EC7736C4FFDA70731EDCED28FB9D75AC" blockId="12.[151,1436,1882,2027]" pageId="12" pageNumber="483">
|
||
<emphasis id="DEBCEAD6FFDA70731EDCED28FE677529" bold="true" box="[151,270,1882,1904]" pageId="12" pageNumber="483">FIGURE 7</emphasis>
|
||
. Urocardiac ossicle of the subgenus
|
||
<taxonomicName id="2BC84D47FFDA70731CC0ED29FDB47536" box="[651,733,1883,1904]" class="Malacostraca" family="Ocypodidae" genus="Leptuca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="483" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="DEBCEAD6FFDA70731CC0ED29FDB47536" box="[651,733,1883,1904]" italics="true" pageId="12" pageNumber="483">Leptuca</emphasis>
|
||
</taxonomicName>
|
||
species. A,
|
||
<taxonomicName id="2BC84D47FFDA70731D17ED29FB717536" box="[860,1048,1882,1904]" class="Malacostraca" family="Ocypodidae" genus="Uca" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="483" phylum="Arthropoda" rank="species" species="subcylindrica">
|
||
<emphasis id="DEBCEAD6FFDA70731D17ED29FB717536" box="[860,1048,1882,1904]" italics="true" pageId="12" pageNumber="483">Uca subcylindrica</emphasis>
|
||
</taxonomicName>
|
||
(NCHUZOOL 13952, CW 15.2 mm, ♂); B,
|
||
<taxonomicName id="2BC84D47FFDA70731E96ED08FE1E75C8" box="[221,375,1912,1935]" class="Malacostraca" family="Ocypodidae" genus="Uca" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="483" phylum="Arthropoda" rank="species" species="terpsichores">
|
||
<emphasis id="DEBCEAD6FFDA70731E96ED08FE1E75C8" box="[221,375,1912,1935]" italics="true" pageId="12" pageNumber="483">U. terpsichores</emphasis>
|
||
</taxonomicName>
|
||
(NCHUZOOL 13582, CW 9.9 mm, ♂); C,
|
||
<taxonomicName id="2BC84D47FFDA70731D78ED08FCB075C8" box="[819,985,1913,1935]" class="Malacostraca" family="Ocypodidae" genus="Uca" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="483" phylum="Arthropoda" rank="species" species="uruguayensis">
|
||
<emphasis id="DEBCEAD6FFDA70731D78ED08FCB075C8" box="[819,985,1913,1935]" italics="true" pageId="12" pageNumber="483">U. uruguayensis</emphasis>
|
||
</taxonomicName>
|
||
(NCHUZOOL 13577, CW 14.1 mm, ♂); D,
|
||
<taxonomicName id="2BC84D47FFDA70731EDCEDEAFE4B75EB" box="[151,290,1944,1965]" class="Malacostraca" family="Ocypodidae" genus="Uca" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="483" phylum="Arthropoda" rank="species" species="spinicarpa">
|
||
<emphasis id="DEBCEAD6FFDA70731EDCEDEAFFC075EB" box="[151,169,1944,1965]" italics="true" pageId="12" pageNumber="483">U</emphasis>
|
||
.
|
||
<emphasis id="DEBCEAD6FFDA70731EFEEDEAFE4B75EB" box="[181,290,1944,1965]" italics="true" pageId="12" pageNumber="483">spinicarpa</emphasis>
|
||
</taxonomicName>
|
||
(NCHUZOOL 13947, CW 13.3 mm, ♀); E,
|
||
<taxonomicName id="2BC84D47FFDA70731CA3EDEAFC3475EB" box="[744,861,1944,1965]" class="Malacostraca" family="Ocypodidae" genus="Uca" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="483" phylum="Arthropoda" rank="species" species="panacea">
|
||
<emphasis id="DEBCEAD6FFDA70731CA3EDEAFC3475EB" box="[744,861,1944,1965]" italics="true" pageId="12" pageNumber="483">U. panacea</emphasis>
|
||
</taxonomicName>
|
||
(NCHUZOOL 13950, CW 14.8 mm, ♂); F,
|
||
<taxonomicName id="2BC84D47FFDA70731B54EDEAFAF575EB" box="[1311,1436,1943,1965]" class="Malacostraca" family="Ocypodidae" genus="Uca" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="483" phylum="Arthropoda" rank="species" species="pugilator">
|
||
<emphasis id="DEBCEAD6FFDA70731B54EDEAFAF575EB" box="[1311,1436,1943,1965]" italics="true" pageId="12" pageNumber="483">U. pugilator</emphasis>
|
||
</taxonomicName>
|
||
(NCHUZOOL 13586, CW 15.9 mm, ♂); G,
|
||
<taxonomicName id="2BC84D47FFDA70731CC6EDC5FC63758A" box="[653,778,1975,1996]" class="Malacostraca" family="Ocypodidae" genus="Uca" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="483" phylum="Arthropoda" rank="species" species="speciosa">
|
||
<emphasis id="DEBCEAD6FFDA70731CC6EDC5FC63758A" box="[653,778,1975,1996]" italics="true" pageId="12" pageNumber="483">U. speciosa</emphasis>
|
||
</taxonomicName>
|
||
(NCHUZOOL 13951, CW 12.6 mm, ♂); H,
|
||
<taxonomicName id="2BC84D47FFDA70731B5AEDC5FAF5758A" box="[1297,1436,1974,1996]" class="Malacostraca" family="Ocypodidae" genus="Uca" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="483" phylum="Arthropoda" rank="species" species="umbratila">
|
||
<emphasis id="DEBCEAD6FFDA70731B5AEDC5FAF5758A" box="[1297,1436,1974,1996]" italics="true" pageId="12" pageNumber="483">U. umbratila</emphasis>
|
||
</taxonomicName>
|
||
(NCHUZOOL 13579, CW 16.2 mm, ♂); I,
|
||
<taxonomicName id="2BC84D47FFDA70731C1BEDA4FDDE75AC" box="[592,695,2004,2027]" class="Malacostraca" family="Ocypodidae" genus="Uca" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="483" phylum="Arthropoda" rank="species" species="thayeri">
|
||
<emphasis id="DEBCEAD6FFDA70731C1BEDA4FD0B75AD" box="[592,610,2006,2027]" italics="true" pageId="12" pageNumber="483">U</emphasis>
|
||
.
|
||
<emphasis id="DEBCEAD6FFDA70731C24EDA6FDDE75AC" box="[623,695,2004,2026]" italics="true" pageId="12" pageNumber="483">thayeri</emphasis>
|
||
</taxonomicName>
|
||
(NCHUZOOL 13953, CW 19.0 mm, ♂). Scale = 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="B8B7664CFFDB70721EDCECC5FA5E748B" box="[151,1335,1719,1742]" httpUri="https://zenodo.org/record/235581/files/figure.png" pageId="13" pageNumber="484" targetBox="[163,1425,205,1689]" targetPageId="13">
|
||
<paragraph id="EC7736C4FFDB70721EDCECC5FA5E748B" blockId="13.[151,1335,1719,1742]" box="[151,1335,1719,1742]" pageId="13" pageNumber="484">
|
||
<emphasis id="DEBCEAD6FFDB70721EDCECC5FE64748A" bold="true" box="[151,269,1719,1741]" pageId="13" pageNumber="484">FIGURE 8</emphasis>
|
||
.
|
||
<taxonomicName id="2BC84D47FFDB70721F52ECCBFEA87488" box="[281,449,1721,1742]" class="Malacostraca" family="Ocypodidae" genus="Uca" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="484" phylum="Arthropoda" rank="species" species="panamensis">
|
||
<emphasis id="DEBCEAD6FFDB70721F52ECCBFEA87488" box="[281,449,1721,1742]" italics="true" pageId="13" pageNumber="484">Uca panamensis</emphasis>
|
||
</taxonomicName>
|
||
♂ (A, B) and its habitat (C). Photographs taken in Panamá. B, courtesy of P. Backwell.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="EC7736C4FFDB70711E8CEC84FDB5735B" blockId="13.[151,1436,1782,2023]" lastBlockId="14.[151,1437,151,2013]" lastPageId="14" lastPageNumber="485" pageId="13" pageNumber="484">
|
||
Beinlich & von
|
||
<bibRefCitation id="88594B35FFDB70721FCFEC85FD487549" author="Hagen" box="[388,545,1782,1807]" pageId="13" pageNumber="484" refString="Beinlich, B. & von Hagen, H. O. (2006) Materials for a more stable subdivision of the genus Uca Leach. Zoologische Mededelingen, 80, 9 - 32." type="journal article" year="2006">Hagen (2006)</bibRefCitation>
|
||
suggested that the function of the orbital armature at the inner corner of the orbital floor is to secure the eyestalk in its depressed position. This character of
|
||
<taxonomicName id="2BC84D47FFDB70721A6BED69FBA17574" box="[1056,1224,1819,1842]" class="Malacostraca" family="Ocypodidae" genus="Uca" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="484" phylum="Arthropoda" rank="species" species="panamensis">
|
||
<emphasis id="DEBCEAD6FFDB70721A6BED69FBA17574" box="[1056,1224,1819,1842]" italics="true" pageId="13" pageNumber="484">U. panamensis</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="74F32A41FFDB70721A93ED68FA757575" box="[1240,1308,1818,1843]" captionStart="FIGURE 2" captionStartId="7.[151,250,1925,1947]" captionTargetBox="[159,1425,203,1891]" captionTargetId="figure@7.[151,1436,193,1903]" captionTargetPageId="7" captionText="FIGURE 2. Uca panamensis (neotype, USNM 1294205, CW 16.8 mm). A, orbital region; B, inner side of orbital region, showing the tubercle of orbital armature (arrowed); C, frontal view; D, front; E, third maxillipeds. Scale, A, E = 2 mm; B = 1 mm; C = 10 mm; D = 5 mm." httpUri="https://zenodo.org/record/235575/files/figure.png" pageId="13" pageNumber="484">Fig. 2</figureCitation>
|
||
B; Beinlich & von
|
||
<bibRefCitation id="88594B35FFDB70721EA2ED4DFE1E7511" author="Hagen" box="[233,375,1854,1879]" pageId="13" pageNumber="484" refString="Beinlich, B. & von Hagen, H. O. (2006) Materials for a more stable subdivision of the genus Uca Leach. Zoologische Mededelingen, 80, 9 - 32." type="journal article" year="2006">Hagen 2006</bibRefCitation>
|
||
: fig. 7j) is considered to be ancestral because it is only present in the subgenera
|
||
<taxonomicName id="2BC84D47FFDB70721B53ED4DFA2E7510" box="[1304,1351,1855,1878]" class="Malacostraca" family="Ocypodidae" genus="Uca" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="484" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="DEBCEAD6FFDB70721B53ED4DFA2E7510" box="[1304,1351,1855,1878]" italics="true" pageId="13" pageNumber="484">Uca</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="88594B35FFDB70721B1CED4DFFBA753D" author="Crane" pageId="13" pageNumber="484" refString="Crane, J. (1975) Fiddler Crabs of the World (Ocypodidae: Genus Uca). Princeton University Press, Princeton, New Jersey, xxiii + 736 pp." type="book" year="1975">Crane 1975</bibRefCitation>
|
||
: figs. 26E, 31D, 32J–K; Beinlich & von
|
||
<bibRefCitation id="88594B35FFDB70721CEBED11FC44753D" author="Hagen" box="[672,813,1890,1915]" pageId="13" pageNumber="484" refString="Beinlich, B. & von Hagen, H. O. (2006) Materials for a more stable subdivision of the genus Uca Leach. Zoologische Mededelingen, 80, 9 - 32." type="journal article" year="2006">Hagen 2006</bibRefCitation>
|
||
: fig. 7d) and
|
||
<taxonomicName id="2BC84D47FFDB70721D88ED11FB66753D" box="[963,1039,1891,1915]" class="Malacostraca" family="Ocypodidae" genus="Afruca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="484" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="DEBCEAD6FFDB70721D88ED11FB66753D" box="[963,1039,1891,1915]" italics="true" pageId="13" pageNumber="484">Afruca</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="88594B35FFDB70721A55ED11FBCE753D" author="Crane" box="[1054,1191,1890,1915]" pageId="13" pageNumber="484" refString="Crane, J. (1975) Fiddler Crabs of the World (Ocypodidae: Genus Uca). Princeton University Press, Princeton, New Jersey, xxiii + 736 pp." type="book" year="1975">Crane 1975</bibRefCitation>
|
||
: fig. 27D–F; Beinlich & von
|
||
<bibRefCitation id="88594B35FFDB70721EAFEDF5FE0675D9" author="Hagen" box="[228,367,1926,1951]" pageId="13" pageNumber="484" refString="Beinlich, B. & von Hagen, H. O. (2006) Materials for a more stable subdivision of the genus Uca Leach. Zoologische Mededelingen, 80, 9 - 32." type="journal article" year="2006">Hagen 2006</bibRefCitation>
|
||
: fig.
|
||
<date id="98761004FFDB70721FEDEDF4FED775D9" box="[422,446,1926,1951]" pageId="13" pageNumber="484">7i</date>
|
||
), and is also seen in
|
||
<taxonomicName id="2BC84D47FFDB70721CE8EDF4FC6E75D9" box="[675,775,1926,1951]" class="Malacostraca" family="Ocypodidae" genus="Ocypode" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="484" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="DEBCEAD6FFDB70721CE8EDF4FC6E75D9" box="[675,775,1926,1951]" italics="true" pageId="13" pageNumber="484">Ocypode</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<taxonomicName id="2BC84D47FFDB70721D5FEDF4FCCB75D9" box="[788,930,1926,1951]" class="Malacostraca" family="Ocypodidae" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="484" phylum="Arthropoda" rank="family">Ocypodidae</taxonomicName>
|
||
:
|
||
<taxonomicName id="2BC84D47FFDB70721DE7EDF4FB5E75D9" box="[940,1079,1926,1951]" class="Malacostraca" family="Ocypodidae" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="484" phylum="Arthropoda" rank="subFamily" subFamily="Ocypodinae">Ocypodinae</taxonomicName>
|
||
;
|
||
<bibRefCitation id="88594B35FFDB70721A0AEDF4FBB675D9" author="Orchard" box="[1089,1247,1926,1951]" pageId="13" pageNumber="484" refString="Orchard, M. (2012) Crabs of Christmas Island. Christmas Island Natural History Association, Christmas Island, Australia. 288 pp." type="book" year="2012">Orchard 2012</bibRefCitation>
|
||
: 232–251;
|
||
<bibRefCitation id="88594B35FFDB70721B11EDF5FE7D7585" author="Wong" pageId="13" pageNumber="484" refString="Wong, K. J. H., Shih, H. - T. & Chan, B. K. K. (2012) The ghost crab Ocypode mortoni George, 1982 (Crustacea: Decapoda: Ocypodidae): redescription, distribution at its type locality, and the phylogeny of East Asian Ocypode species. Zootaxa, 3550, 71 - 87." type="journal article" year="2012">
|
||
Wong
|
||
<emphasis id="DEBCEAD6FFDB70721EDCEDD9FFB97585" box="[151,208,1962,1987]" italics="true" pageId="13" pageNumber="484">et al.</emphasis>
|
||
2012
|
||
</bibRefCitation>
|
||
: fig. 6b) and
|
||
<taxonomicName id="2BC84D47FFDB70721FE6EDD8FD747585" box="[429,541,1962,1987]" class="Malacostraca" family="Heloeciidae" genus="Heloecius" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="484" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="DEBCEAD6FFDB70721FE6EDD8FD747585" box="[429,541,1962,1987]" italics="true" pageId="13" pageNumber="484">Heloecius</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<taxonomicName id="2BC84D47FFDB70721C67EDD8FDDE7585" box="[556,695,1962,1987]" class="Malacostraca" family="Heloeciidae" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="484" phylum="Arthropoda" rank="family">Heloeciidae</taxonomicName>
|
||
; Beinlich & von
|
||
<bibRefCitation id="88594B35FFDB70721D30EDD9FB607585" author="Hagen" box="[891,1033,1962,1987]" pageId="13" pageNumber="484" refString="Beinlich, B. & von Hagen, H. O. (2006) Materials for a more stable subdivision of the genus Uca Leach. Zoologische Mededelingen, 80, 9 - 32." type="journal article" year="2006">Hagen 2006</bibRefCitation>
|
||
: fig. 7c). Other fiddler crabs do not have this character, or have instead evolved other structures that may secure the eyestalks (such as broadening of the margins of the orbits, and one or more rows of tubercles, partly on raised “mounds” behind the lower orbital border) (see Beinlich & von
|
||
<bibRefCitation id="88594B35FFD870711F9EEACFFD0B7293" author="Hagen" box="[469,610,188,213]" pageId="14" pageNumber="485" refString="Beinlich, B. & von Hagen, H. O. (2006) Materials for a more stable subdivision of the genus Uca Leach. Zoologische Mededelingen, 80, 9 - 32." type="journal article" year="2006">Hagen 2006</bibRefCitation>
|
||
). The supposedly more plesiomorphic orbital characters of
|
||
<taxonomicName id="2BC84D47FFD870711ABDEACFFAF57292" box="[1270,1436,189,212]" class="Malacostraca" family="Ocypodidae" genus="Uca" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="485" phylum="Arthropoda" rank="species" species="panamensis">
|
||
<emphasis id="DEBCEAD6FFD870711ABDEACFFAF57292" box="[1270,1436,189,212]" italics="true" pageId="14" pageNumber="485">U. panamensis</emphasis>
|
||
</taxonomicName>
|
||
supports our hypothesis that
|
||
<emphasis id="DEBCEAD6FFD870711FA8EA93FD5572BE" box="[483,572,225,248]" italics="true" pageId="14" pageNumber="485">Petruca</emphasis>
|
||
<emphasis id="DEBCEAD6FFD870711C0EEAADFDB372BE" bold="true" box="[581,730,223,248]" pageId="14" pageNumber="485">subgen. nov.</emphasis>
|
||
separated earlier from the other two American broad-fronted subgenera,
|
||
<taxonomicName id="2BC84D47FFD870711F5DEB77FE05735A" box="[278,364,261,284]" class="Malacostraca" family="Ocypodidae" genus="Minuca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="485" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="DEBCEAD6FFD870711F5DEB77FE05735A" box="[278,364,261,284]" italics="true" pageId="14" pageNumber="485">Minuca</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="2BC84D47FFD870711FE9EB77FE94735A" box="[418,509,261,284]" class="Malacostraca" family="Ocypodidae" genus="Leptuca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="485" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="DEBCEAD6FFD870711FE9EB77FE94735A" box="[418,509,261,284]" italics="true" pageId="14" pageNumber="485">Leptuca</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="74F32A41FFD870711C47EB76FD39735A" box="[524,592,260,285]" captionStart="FIGURE 9. A" captionStartId="17.[151,250,1908,1930]" captionTargetBox="[160,1392,213,1872]" captionTargetId="figure@17.[151,1436,193,1886]" captionTargetPageId="17" captionText="FIGURE 9. A Bayesian inference (BI) tree of Petruca subgen. nov. and other related subgenera, based on the combined 16 S rDNA, cytochrome oxidase subunit I genes (COI) and 28 S rDNA. See Table 1 for details of the specimens. Probability values at the nodes represent support values for BI, as well as two methods of maximum likelihood (ML), with ML * from RAxML and ML ** from GARLI." httpUri="https://zenodo.org/record/235582/files/figure.png" pageId="14" pageNumber="485">Fig. 9</figureCitation>
|
||
, see below).
|
||
</paragraph>
|
||
<paragraph id="EC7736C4FFD870711E8CEB55FC577153" blockId="14.[151,1437,151,2013]" pageId="14" pageNumber="485">
|
||
The posterior extension of the
|
||
<taxonomicName id="2BC84D47FFD870711C73EB55FD107306" box="[568,633,295,320]" class="Malacostraca" family="Ocypodidae" genus="Uca" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="485" phylum="Arthropoda" rank="species" species="major">major</taxonomicName>
|
||
manus (
|
||
<figureCitation id="74F32A41FFD870711CA8EB55FC4F7306" box="[739,806,295,320]" captionStart="FIGURE 1" captionStartId="6.[151,250,1911,1933]" captionTargetBox="[163,1423,215,1877]" captionTargetId="figure@6.[151,1436,193,1890]" captionTargetPageId="6" captionText="FIGURE 1. Uca panamensis (neotype, USNM 1294205, CW 16.8 mm). A, dorsal view; B, boss on carapace behind right dorsolateral margin (arrowed); C, ventral view. Scale, B = 2 mm." httpUri="https://zenodo.org/record/235574/files/figure.png" pageId="14" pageNumber="485">Fig. 1</figureCitation>
|
||
A, C) is a unique character of the new subgenus, as previously noted by
|
||
<bibRefCitation id="88594B35FFD870711F36EB3EFD7C7323" author="Crane" box="[381,533,332,357]" pageId="14" pageNumber="485" refString="Crane, J. (1975) Fiddler Crabs of the World (Ocypodidae: Genus Uca). Princeton University Press, Princeton, New Jersey, xxiii + 736 pp." type="book" year="1975">Crane (1975)</bibRefCitation>
|
||
. The posteriorly extended rounded protuberance prevents a full lateral extension of the manus and
|
||
<bibRefCitation id="88594B35FFD870711F29EB02FD5873CE" author="Crane" box="[354,561,367,392]" pageId="14" pageNumber="485" refString="Crane, J. (1975) Fiddler Crabs of the World (Ocypodidae: Genus Uca). Princeton University Press, Princeton, New Jersey, xxiii + 736 pp." type="book" year="1975">Crane (1975: 158)</bibRefCitation>
|
||
suggested that males strike the ground with this part of their claw when they court producing a seismic signal. Subsequent observations (J. Christy & U. Schober unpublished), however, revealed that the claw is held well above the substrate during the entire wave. Crane also seldom saw courtship waving. Indeed, courtship waving by this species on a cobble beach near the Pacific entrance of the
|
||
<collectingCountry id="94DF7654FFD870711AB0EBAEFA3D73B2" box="[1275,1364,476,500]" name="Panama" pageId="14" pageNumber="485">Panamá</collectingCountry>
|
||
Canal occurs during a relatively short interval of approximately an hour beginning late in the tidal cycle well after low tide and after the crabs have fed, and on only a few days each semi-lunar breeding cycle. Males, which typically are very dark in colour at this location, lighten on the carapace to a khaki or cream colour when they court (
|
||
<figureCitation id="74F32A41FFD870711B7DE835FA107026" box="[1334,1401,583,608]" captionStart="FIGURE 8" captionStartId="13.[151,250,1719,1741]" captionTargetBox="[163,1425,205,1689]" captionTargetId="figure@13.[151,1436,193,1698]" captionTargetPageId="13" captionText="FIGURE 8. Uca panamensis ♂ (A, B) and its habitat (C). Photographs taken in Panamá. B, courtesy of P. Backwell." httpUri="https://zenodo.org/record/235581/files/figure.png" pageId="14" pageNumber="485">Fig. 8</figureCitation>
|
||
B), typically from the highest point on a stone, one male per perch. Males that court from elevated positions may be seen more easily by prospective mates but also by their predators including great-tailed grackles [
|
||
<taxonomicName id="2BC84D47FFD870711B67E8FDFEAC708A" authority="Gmelin, 1788" authorityName="Gmelin" authorityYear="1788" class="Aves" family="Icteridae" genus="Quiscalus" kingdom="Animalia" order="Passeriformes" pageId="14" pageNumber="485" phylum="Chordata" rank="species" species="mexicanus">
|
||
<emphasis id="DEBCEAD6FFD870711B67E8FDFE64708A" italics="true" pageId="14" pageNumber="485">Quiscalus mexicanus</emphasis>
|
||
(Gmelin, 1788)
|
||
</taxonomicName>
|
||
] and the furtive but quick grapsid crab
|
||
<taxonomicName id="2BC84D47FFD870711DC3E8C7FAF1708A" authority="H. Milne Edwards, 1837" authorityName="H. Milne Edwards" authorityYear="1837" box="[904,1432,692,717]" class="Malacostraca" family="Grapsidae" genus="Geograpsus" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="485" phylum="Arthropoda" rank="species" species="lividus">
|
||
<emphasis id="DEBCEAD6FFD870711DC3E8C7FB0A708B" box="[904,1123,692,717]" italics="true" pageId="14" pageNumber="485">Geograpsus lividus</emphasis>
|
||
(H. Milne Edwards, 1837)
|
||
</taxonomicName>
|
||
. When disturbed, courting males drop from their perch landing at the seam between the stone and the sand or shell hash matrix where the openings to their burrows are located.
|
||
</paragraph>
|
||
<paragraph id="EC7736C4FFD870711E8CE96DFB267683" blockId="14.[151,1437,151,2013]" pageId="14" pageNumber="485">
|
||
The significance of the smooth inner and outer surfaces of the
|
||
<taxonomicName id="2BC84D47FFD870711DC1E96DFCA2717E" box="[906,971,799,824]" class="Malacostraca" family="Ocypodidae" genus="Uca" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="485" phylum="Arthropoda" rank="species" species="major">major</taxonomicName>
|
||
chela (
|
||
<figureCitation id="74F32A41FFD870711A55E96DFB36717E" box="[1054,1119,799,824]" captionStart="FIGURE 3" captionStartId="8.[151,250,1887,1909]" captionTargetBox="[161,1423,207,1857]" captionTargetId="figure@8.[151,1436,193,1866]" captionTargetPageId="8" captionText="FIGURE 3. Major and minor chelipeds of Uca panamensis (neotype, USNM 1294205, CW 16.8 mm). A, B, outer and inner views of major cheliped; C, D, outer and inner views of minor cheliped. Scale, C, D = 2 mm." httpUri="https://zenodo.org/record/235576/files/figure.png" pageId="14" pageNumber="485">Fig. 3</figureCitation>
|
||
B) is unknown. This feature is also present in species inhabiting sandy or muddy habitats, e.g.,
|
||
<taxonomicName id="2BC84D47FFD870711DEAE937FA1B711A" authority="Hoffmann, 1874" authorityName="Hoffmann" authorityYear="1874" box="[929,1394,836,861]" class="Malacostraca" family="Ocypodidae" genus="Uca" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="485" phylum="Arthropoda" rank="species" species="inversa" subGenus="Cranuca">
|
||
<emphasis id="DEBCEAD6FFD870711DEAE937FCDC711A" box="[929,949,837,860]" italics="true" pageId="14" pageNumber="485">U</emphasis>
|
||
. (
|
||
<emphasis id="DEBCEAD6FFD870711D9BE937FB5B711A" box="[976,1074,837,860]" italics="true" pageId="14" pageNumber="485">Cranuca</emphasis>
|
||
)
|
||
<emphasis id="DEBCEAD6FFD870711A0EE937FBF1711A" box="[1093,1176,837,860]" italics="true" pageId="14" pageNumber="485">inversa</emphasis>
|
||
(Hoffmann, 1874)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="2BC84D47FFD870711BCAE937FD4471C6" authority="Alcock, 1900" authorityName="Alcock" authorityYear="1900" class="Malacostraca" family="Ocypodidae" genus="Uca" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="485" phylum="Arthropoda" rank="species" species="sindensis" subGenus="Austruca">
|
||
<emphasis id="DEBCEAD6FFD870711BCAE937FAFC711A" box="[1409,1429,837,860]" italics="true" pageId="14" pageNumber="485">U</emphasis>
|
||
. (
|
||
<emphasis id="DEBCEAD6FFD870711EEBE91AFE6C7139" box="[160,261,872,895]" italics="true" pageId="14" pageNumber="485">Austruca</emphasis>
|
||
)
|
||
<emphasis id="DEBCEAD6FFD870711F5EE915FE1571C6" box="[277,380,871,896]" italics="true" pageId="14" pageNumber="485">sindensis</emphasis>
|
||
(Alcock, 1900)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="2BC84D47FFD870711C73E91BFCA771C6" authority="Crane, 1941" authorityName="Crane" authorityYear="1941" box="[568,974,871,896]" class="Malacostraca" family="Ocypodidae" genus="Uca" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="485" phylum="Arthropoda" rank="species" species="argillicola" subGenus="Minuca">
|
||
<emphasis id="DEBCEAD6FFD870711C73E91BFD2571C6" box="[568,588,873,896]" italics="true" pageId="14" pageNumber="485">U</emphasis>
|
||
. (
|
||
<emphasis id="DEBCEAD6FFD870711C28E91BFDD171C6" box="[611,696,873,896]" italics="true" pageId="14" pageNumber="485">Minuca</emphasis>
|
||
)
|
||
<emphasis id="DEBCEAD6FFD870711C83E915FC5471C6" box="[712,829,871,896]" italics="true" pageId="14" pageNumber="485">argillicola</emphasis>
|
||
<bibRefCitation id="88594B35FFD870711D0FE91AFCA771C6" author="Crane" box="[836,974,872,896]" pageId="14" pageNumber="485" refString="Crane, J. (1941) Crabs of the genus Uca from the west coast of Central America. Eastern Pacific Expeditions of the New York Zoological Society. XXVI. Zoologica, 26 (19), 145 - 208." type="journal article" year="1941">Crane, 1941</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="2BC84D47FFD870711D92E91BFA6B71C6" authority="Crane, 1941" authorityName="Crane" authorityYear="1941" box="[985,1282,872,896]" class="Malacostraca" family="Ocypodidae" genus="Uca" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="485" phylum="Arthropoda" rank="species" species="zacae" subGenus="Minuca">
|
||
<emphasis id="DEBCEAD6FFD870711D92E91BFC8471C6" box="[985,1005,873,896]" italics="true" pageId="14" pageNumber="485">U</emphasis>
|
||
. (
|
||
<emphasis id="DEBCEAD6FFD870711A4FE91BFB7271C6" box="[1028,1051,873,896]" italics="true" pageId="14" pageNumber="485">M</emphasis>
|
||
.)
|
||
<emphasis id="DEBCEAD6FFD870711A79E91BFB1B71C6" box="[1074,1138,873,896]" italics="true" pageId="14" pageNumber="485">zacae</emphasis>
|
||
<bibRefCitation id="88594B35FFD870711A33E91AFA6B71C6" author="Crane" box="[1144,1282,872,896]" pageId="14" pageNumber="485" refString="Crane, J. (1941) Crabs of the genus Uca from the west coast of Central America. Eastern Pacific Expeditions of the New York Zoological Society. XXVI. Zoologica, 26 (19), 145 - 208." type="journal article" year="1941">Crane, 1941</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="2BC84D47FFD870711B46E91BFEB971E2" authority="Rathbun, 1894" authorityName="Rathbun" authorityYear="1894" class="Malacostraca" family="Ocypodidae" genus="Uca" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="485" phylum="Arthropoda" rank="species" species="latimanus" subGenus="Leptuca">
|
||
<emphasis id="DEBCEAD6FFD870711B46E91BFA4871C6" box="[1293,1313,873,896]" italics="true" pageId="14" pageNumber="485">U</emphasis>
|
||
. (
|
||
<emphasis id="DEBCEAD6FFD870711B73E91BFAFB71C6" box="[1336,1426,873,896]" italics="true" pageId="14" pageNumber="485">Leptuca</emphasis>
|
||
)
|
||
<emphasis id="DEBCEAD6FFD870711EDCE9FEFE6F71E3" box="[151,262,908,933]" italics="true" pageId="14" pageNumber="485">latimanus</emphasis>
|
||
(Rathbun, 1894)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="2BC84D47FFD870711FABE9FFFCAA71E2" authority="Novak & Salmon, 1974" authorityName="Novak & Salmon" authorityYear="1974" box="[480,963,908,933]" class="Malacostraca" family="Ocypodidae" genus="Uca" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="485" phylum="Arthropoda" rank="species" species="panacea" subGenus="Leptuca">
|
||
<emphasis id="DEBCEAD6FFD870711FABE9FFFE9D71E2" box="[480,500,909,932]" italics="true" pageId="14" pageNumber="485">U</emphasis>
|
||
. (
|
||
<emphasis id="DEBCEAD6FFD870711C5AE9FFFD4971E2" box="[529,544,909,932]" italics="true" pageId="14" pageNumber="485">L</emphasis>
|
||
.)
|
||
<emphasis id="DEBCEAD6FFD870711C77E9FFFDF271E2" box="[572,667,909,932]" italics="true" pageId="14" pageNumber="485">panacea</emphasis>
|
||
<bibRefCitation id="88594B35FFD870711CECE9FEFCAA71E2" author="Novak" box="[679,963,908,933]" pageId="14" pageNumber="485" refString="Novak, A. & Salmon, M. (1974) Uca panacea, a new species of fiddler crab from the Gulf coast of the United States. Proceedings of the Biological Society of Washington, 87, 313 - 326." type="journal article" year="1974">Novak & Salmon, 1974</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="2BC84D47FFD870711D98E9FFFA5271E2" authority="Bosc, 1802" authorityName="Bosc" authorityYear="1802" box="[979,1339,908,933]" class="Malacostraca" family="Ocypodidae" genus="Uca" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="485" phylum="Arthropoda" rank="species" species="pugilator" subGenus="Leptuca">
|
||
<emphasis id="DEBCEAD6FFD870711D98E9FFFC8E71E2" box="[979,999,909,932]" italics="true" pageId="14" pageNumber="485">U</emphasis>
|
||
. (
|
||
<emphasis id="DEBCEAD6FFD870711A48E9FFFB7B71E2" box="[1027,1042,909,932]" italics="true" pageId="14" pageNumber="485">L</emphasis>
|
||
.)
|
||
<emphasis id="DEBCEAD6FFD870711A64E9FEFBFE71E3" box="[1071,1175,908,933]" italics="true" pageId="14" pageNumber="485">pugilator</emphasis>
|
||
(Bosc, 1802)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="2BC84D47FFD870711B07E9FFFE97718E" authority="Stimpson, 1859" authorityName="Stimpson" authorityYear="1859" class="Malacostraca" family="Ocypodidae" genus="Uca" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="485" phylum="Arthropoda" rank="species" species="subcylindrica" subGenus="Leptuca">
|
||
<emphasis id="DEBCEAD6FFD870711B07E9FFFA0971E2" box="[1356,1376,909,932]" italics="true" pageId="14" pageNumber="485">U</emphasis>
|
||
. (
|
||
<emphasis id="DEBCEAD6FFD870711B37E9FFFAE271E2" box="[1404,1419,909,932]" italics="true" pageId="14" pageNumber="485">L</emphasis>
|
||
.)
|
||
<emphasis id="DEBCEAD6FFD870711EDCE9DDFE59718E" box="[151,304,943,968]" italics="true" pageId="14" pageNumber="485">subcylindrica</emphasis>
|
||
(
|
||
<bibRefCitation id="88594B35FFD870711F0BE9DDFE9E718E" author="Stimpson" box="[320,503,943,968]" pageId="14" pageNumber="485" refString="Stimpson, W. (1859) Notes on North American Crustacea, No. 1. Annals of the Lyceum of Natural History of New York, 7 (4 - 5), 49 - 93, pl. I." type="journal article" year="1859">Stimpson, 1859</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="2BC84D47FFD870711C75E9C3FCF5718E" authority="Crane, 1941" authorityName="Crane" authorityYear="1941" box="[574,924,943,968]" class="Malacostraca" family="Ocypodidae" genus="Uca" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="485" phylum="Arthropoda" rank="species" species="tenuipedis" subGenus="Leptuca">
|
||
<emphasis id="DEBCEAD6FFD870711C75E9C3FD3B718E" box="[574,594,945,968]" italics="true" pageId="14" pageNumber="485">U</emphasis>
|
||
. (
|
||
<emphasis id="DEBCEAD6FFD870711C20E9C3FD13718E" box="[619,634,945,968]" italics="true" pageId="14" pageNumber="485">L</emphasis>
|
||
.)
|
||
<emphasis id="DEBCEAD6FFD870711CDFE9DDFC6E718E" box="[660,775,943,968]" italics="true" pageId="14" pageNumber="485">tenuipedis</emphasis>
|
||
<bibRefCitation id="88594B35FFD870711D5AE9C2FCF5718E" author="Crane" box="[785,924,944,968]" pageId="14" pageNumber="485" refString="Crane, J. (1941) Crabs of the genus Uca from the west coast of Central America. Eastern Pacific Expeditions of the New York Zoological Society. XXVI. Zoologica, 26 (19), 145 - 208." type="journal article" year="1941">Crane, 1941</bibRefCitation>
|
||
</taxonomicName>
|
||
(see also
|
||
<bibRefCitation id="88594B35FFD870711A5AE9DDFBA4718E" author="Rosenberg" box="[1041,1229,943,968]" pageId="14" pageNumber="485" refString="Rosenberg, M. S. (2001) The systematics and taxonomy of fiddler crabs: a phylogeny of the genus Uca. Journal of Crustacean Biology, 21, 839 - 869. http: // dx. doi. org / 10.1163 / 20021975 - 99990176" type="journal article" year="2001">Rosenberg 2001</bibRefCitation>
|
||
). Christy & Wada (2015) noted that the claws of male
|
||
<taxonomicName id="2BC84D47FFD870711C61E9A7FDDD71AB" box="[554,692,980,1005]" class="Malacostraca" family="Ocypodidae" genus="Uca" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="485" phylum="Arthropoda" rank="species" species="pugilator">
|
||
<emphasis id="DEBCEAD6FFD870711C61E9A7FDDD71AB" box="[554,692,980,1005]" italics="true" pageId="14" pageNumber="485">U. pugilator</emphasis>
|
||
</taxonomicName>
|
||
often slip when males fight and that the relatively dry sand can be difficult for a challenger to grip. They suggested a smooth claw may allow males in their role as challengers to persist in a fight at the entrance to a burrow without being thrown by the defender, which has the positional advantage and is able to grip the burrow shaft. Similarly, a defending male
|
||
<taxonomicName id="2BC84D47FFD870711D97EE33FBEB761E" box="[988,1154,1089,1112]" class="Malacostraca" family="Ocypodidae" genus="Uca" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="485" phylum="Arthropoda" rank="species" species="panamensis">
|
||
<emphasis id="DEBCEAD6FFD870711D97EE33FBEB761E" box="[988,1154,1089,1112]" italics="true" pageId="14" pageNumber="485">U. panamensis</emphasis>
|
||
</taxonomicName>
|
||
in a burrow under stones may have a decided positional advantage over challengers attempting to grip sand or shell hash. Too little is known about possible positional asymmetries during combat in the other species to know whether selection for proficiency in combat when males are challengers may explain why they too have smooth claws.
|
||
</paragraph>
|
||
<paragraph id="EC7736C4FFD870711E8CEEBDFD167776" blockId="14.[151,1437,151,2013]" pageId="14" pageNumber="485">
|
||
The brush of long stiff setae on the finger’s tips of minor cheliped (
|
||
<figureCitation id="74F32A41FFD870711DF7EEBDFC9676AE" box="[956,1023,1231,1256]" captionStart="FIGURE 3" captionStartId="8.[151,250,1887,1909]" captionTargetBox="[161,1423,207,1857]" captionTargetId="figure@8.[151,1436,193,1866]" captionTargetPageId="8" captionText="FIGURE 3. Major and minor chelipeds of Uca panamensis (neotype, USNM 1294205, CW 16.8 mm). A, B, outer and inner views of major cheliped; C, D, outer and inner views of minor cheliped. Scale, C, D = 2 mm." httpUri="https://zenodo.org/record/235576/files/figure.png" pageId="14" pageNumber="485">Fig. 3</figureCitation>
|
||
C, D) is a very unusual character not present in other fiddler crab taxa; and is apparently an adaptation for scraping algae from the hard rock surface (
|
||
<bibRefCitation id="88594B35FFD870711ED5EF6AFE4D7776" author="Crane" box="[158,292,1304,1328]" pageId="14" pageNumber="485" refString="Crane, J. (1941) Crabs of the genus Uca from the west coast of Central America. Eastern Pacific Expeditions of the New York Zoological Society. XXVI. Zoologica, 26 (19), 145 - 208." type="journal article" year="1941">Crane 1941</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="88594B35FFD870711F7BEF65FE027776" author="Crane" box="[304,363,1303,1328]" pageId="14" pageNumber="485" refString="Crane, J. (1975) Fiddler Crabs of the World (Ocypodidae: Genus Uca). Princeton University Press, Princeton, New Jersey, xxiii + 736 pp." type="book" year="1975">1975</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="88594B35FFD870711F3CEF65FD1A7776" author="Takeda" box="[375,627,1303,1328]" pageId="14" pageNumber="485" refString="Takeda, S. & Murai, M. (2003) Morphological and behavioural adaptations to the rocky substrate by the fiddler crab Uca panamensis (Stimpson, 1859): preference for feeding substratum and feeding mechanism. Journal of Experimental Marine Biology and Ecology, 287, 179 - 191. http: // dx. doi. org / 10.1016 / S 0022 - 0981 (02) 00550 - 6" type="journal article" year="2003">Takeda & Murai 2003</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="EC7736C4FFD870711E8CEF4EFE6A7416" blockId="14.[151,1437,151,2013]" pageId="14" pageNumber="485">
|
||
The urocardiac ossicles of gastric mill in
|
||
<emphasis id="DEBCEAD6FFD870711CF7EF4FFC7C7712" box="[700,789,1341,1364]" italics="true" pageId="14" pageNumber="485">Petruca</emphasis>
|
||
<emphasis id="DEBCEAD6FFD870711D6AEF4EFCD37713" bold="true" box="[801,954,1340,1365]" pageId="14" pageNumber="485">subgen. nov.</emphasis>
|
||
are of the simple form, i.e., with fewer transverse ridges of median teeth, especially those on the posterior tooth plate (
|
||
<figureCitation id="74F32A41FFD870711A1BEF2DFBF0773E" box="[1104,1177,1375,1400]" captionStart="FIGURE 5" captionStartId="10.[151,250,1927,1949]" captionTargetBox="[159,1427,205,1895]" captionTargetId="figure@10.[151,1436,193,1906]" captionTargetPageId="10" captionText="FIGURE 5. Urocardiac ossicle of Uca panamensis. A, B, neotype, USNM 1294205 (16.8 mm); C, ZRC (CW 12.6 mm, ♂); D, ZRC (CW 13.7 mm, ♀). A, C, D, ventral view; B, lateroventral view. Scale, A, B = 1 mm; C, D = 0.5 mm." httpUri="https://zenodo.org/record/235578/files/figure.png" pageId="14" pageNumber="485">Fig. 5</figureCitation>
|
||
; Shih 2015). Another character is the swollen median part of the stem region of the gastric mill in this subgenus (
|
||
<figureCitation id="74F32A41FFD870711A8CEFF6FA6477DB" box="[1223,1293,1412,1437]" captionStart="FIGURE 5" captionStartId="10.[151,250,1927,1949]" captionTargetBox="[159,1427,205,1895]" captionTargetId="figure@10.[151,1436,193,1906]" captionTargetPageId="10" captionText="FIGURE 5. Urocardiac ossicle of Uca panamensis. A, B, neotype, USNM 1294205 (16.8 mm); C, ZRC (CW 12.6 mm, ♂); D, ZRC (CW 13.7 mm, ♀). A, C, D, ventral view; B, lateroventral view. Scale, A, B = 1 mm; C, D = 0.5 mm." httpUri="https://zenodo.org/record/235578/files/figure.png" pageId="14" pageNumber="485">Fig. 5</figureCitation>
|
||
A), which is present in the subgenera
|
||
<emphasis id="DEBCEAD6FFD870711F98EFDBFD4D7786" box="[467,548,1449,1472]" italics="true" pageId="14" pageNumber="485">Xeruca</emphasis>
|
||
,
|
||
<taxonomicName id="2BC84D47FFD870711C70EFDAFDEE7786" box="[571,647,1448,1472]" class="Malacostraca" family="Ocypodidae" genus="Afruca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="485" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="DEBCEAD6FFD870711C70EFDAFDEE7786" box="[571,647,1448,1472]" italics="true" pageId="14" pageNumber="485">Afruca</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="2BC84D47FFD870711C93EFDBFC6E7786" box="[728,775,1449,1472]" class="Malacostraca" family="Ocypodidae" genus="Uca" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="485" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="DEBCEAD6FFD870711C93EFDBFC6E7786" box="[728,775,1449,1472]" italics="true" pageId="14" pageNumber="485">Uca</emphasis>
|
||
</taxonomicName>
|
||
(Shih 2015: fig. 8A, B; Franklin Barnwell, personal communication), but not yet observed in
|
||
<taxonomicName id="2BC84D47FFD870711C2FEFBFFDD377A2" box="[612,698,1485,1508]" class="Malacostraca" family="Ocypodidae" genus="Minuca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="485" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="DEBCEAD6FFD870711C2FEFBFFDD377A2" box="[612,698,1485,1508]" italics="true" pageId="14" pageNumber="485">Minuca</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="74F32A41FFD870711C83EFBEFC6277A3" box="[712,779,1484,1509]" captionStart="FIGURE 6" captionStartId="11.[151,250,1886,1908]" captionTargetBox="[173,1413,207,1861]" captionTargetId="figure@11.[163,1423,193,1868]" captionTargetPageId="11" captionText="FIGURE 6. Urocardiac ossicle of the subgenus Minuca species. A, Uca brevifrons (ZRC 2012.0126, CW 22.1, ♂); B, U. cf. virens (NCHUZOOL 13946, CW 17.0 mm, ♂); C, U. victoriana (NCHUZOOL 13945, CW 12.2 mm, ♀); D, U. pugnax (NCHUZOOL 13941, CW 15.2 mm, ♂); E, U. burgersi (NCHUZOOL 13956, CW 13.0 mm); F, U. rapax (NCHUZOOL 13942, CW 23.4 mm, ♂); G, U. mordax (NCHUZOOL 13940, CW 18.0 mm, ♀); H, U. minax (NCHUZOOL 13957, CW 23.2 mm, ♂); I, U. vocator (NCHUZOOL 13948, CW 21.9 mm, ♀). Scale = 1 mm." httpUri="https://zenodo.org/record/235579/files/figure.png" pageId="14" pageNumber="485">Fig. 6</figureCitation>
|
||
) and
|
||
<taxonomicName id="2BC84D47FFD870711D01EFBFFCCD77A2" box="[842,932,1485,1508]" class="Malacostraca" family="Ocypodidae" genus="Leptuca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="485" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="DEBCEAD6FFD870711D01EFBFFCCD77A2" box="[842,932,1485,1508]" italics="true" pageId="14" pageNumber="485">Leptuca</emphasis>
|
||
</taxonomicName>
|
||
(except in
|
||
<taxonomicName id="2BC84D47FFD870711A6AEFBFFBCB77A2" box="[1057,1186,1485,1508]" class="Malacostraca" family="Ocypodidae" genus="Uca" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="485" phylum="Arthropoda" rank="species" species="panacea">
|
||
<emphasis id="DEBCEAD6FFD870711A6AEFBFFBCB77A2" box="[1057,1186,1485,1508]" italics="true" pageId="14" pageNumber="485">U. panacea</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<figureCitation id="74F32A41FFD870711AFBEFBEFB9D77A2" box="[1200,1268,1484,1509]" captionStart="FIGURE 7" captionStartId="12.[151,250,1882,1904]" captionTargetBox="[176,1408,203,1846]" captionTargetId="figure@12.[165,1421,193,1861]" captionTargetPageId="12" captionText="FIGURE 7. Urocardiac ossicle of the subgenus Leptuca species. A, Uca subcylindrica (NCHUZOOL 13952, CW 15.2 mm, ♂); B, U. terpsichores (NCHUZOOL 13582, CW 9.9 mm, ♂); C, U. uruguayensis (NCHUZOOL 13577, CW 14.1 mm, ♂); D, U. spinicarpa (NCHUZOOL 13947, CW 13.3 mm, ♀); E, U. panacea (NCHUZOOL 13950, CW 14.8 mm, ♂); F, U. pugilator (NCHUZOOL 13586, CW 15.9 mm, ♂); G, U. speciosa (NCHUZOOL 13951, CW 12.6 mm, ♂); H, U. umbratila (NCHUZOOL 13579, CW 16.2 mm, ♂); I, U. thayeri (NCHUZOOL 13953, CW 19.0 mm, ♂). Scale = 1 mm." httpUri="https://zenodo.org/record/235580/files/figure.png" pageId="14" pageNumber="485">Fig. 7</figureCitation>
|
||
). Although the median part of the stem region of
|
||
<taxonomicName id="2BC84D47FFD870711C52EF83FDF2744E" box="[537,667,1521,1544]" class="Malacostraca" family="Ocypodidae" genus="Uca" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="485" phylum="Arthropoda" rank="species" species="panacea">
|
||
<emphasis id="DEBCEAD6FFD870711C52EF83FDF2744E" box="[537,667,1521,1544]" italics="true" pageId="14" pageNumber="485">U. panacea</emphasis>
|
||
</taxonomicName>
|
||
is swollen, it is not wider than the median teeth in
|
||
<emphasis id="DEBCEAD6FFD870711AABEF83FA50744E" box="[1248,1337,1521,1544]" italics="true" pageId="14" pageNumber="485">Petruca</emphasis>
|
||
<emphasis id="DEBCEAD6FFD870711B0AEF9DFFA1746B" bold="true" pageId="14" pageNumber="485">subgen. nov.</emphasis>
|
||
, and the median teeth on the posterior tooth plate also are more complex in structure (with 5 transverse ridges) (
|
||
<figureCitation id="74F32A41FFD870711ED4EC45FF8A7416" box="[159,227,1591,1616]" captionStart="FIGURE 7" captionStartId="12.[151,250,1882,1904]" captionTargetBox="[176,1408,203,1846]" captionTargetId="figure@12.[165,1421,193,1861]" captionTargetPageId="12" captionText="FIGURE 7. Urocardiac ossicle of the subgenus Leptuca species. A, Uca subcylindrica (NCHUZOOL 13952, CW 15.2 mm, ♂); B, U. terpsichores (NCHUZOOL 13582, CW 9.9 mm, ♂); C, U. uruguayensis (NCHUZOOL 13577, CW 14.1 mm, ♂); D, U. spinicarpa (NCHUZOOL 13947, CW 13.3 mm, ♀); E, U. panacea (NCHUZOOL 13950, CW 14.8 mm, ♂); F, U. pugilator (NCHUZOOL 13586, CW 15.9 mm, ♂); G, U. speciosa (NCHUZOOL 13951, CW 12.6 mm, ♂); H, U. umbratila (NCHUZOOL 13579, CW 16.2 mm, ♂); I, U. thayeri (NCHUZOOL 13953, CW 19.0 mm, ♂). Scale = 1 mm." httpUri="https://zenodo.org/record/235580/files/figure.png" pageId="14" pageNumber="485">Fig. 7</figureCitation>
|
||
E).
|
||
</paragraph>
|
||
<paragraph id="EC7736C4FFD870711E8CEC2EFB1975D2" blockId="14.[151,1437,151,2013]" pageId="14" pageNumber="485">
|
||
The urocardiac ossicles of
|
||
<taxonomicName id="2BC84D47FFD870711FB6EC2FFD3B7432" box="[509,594,1629,1652]" class="Malacostraca" family="Ocypodidae" genus="Minuca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="485" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="DEBCEAD6FFD870711FB6EC2FFD3B7432" box="[509,594,1629,1652]" italics="true" pageId="14" pageNumber="485">Minuca</emphasis>
|
||
</taxonomicName>
|
||
show a more consistent pattern than those of
|
||
<taxonomicName id="2BC84D47FFD870711A20EC2FFBAC7432" box="[1131,1221,1629,1652]" class="Malacostraca" family="Ocypodidae" genus="Leptuca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="485" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="DEBCEAD6FFD870711A20EC2FFBAC7432" box="[1131,1221,1629,1652]" italics="true" pageId="14" pageNumber="485">Leptuca</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="74F32A41FFD870711A9CEC2EFA407433" box="[1239,1321,1628,1653]" captionStart="FIGURE 6" captionStartId="11.[151,250,1886,1908]" captionTargetBox="[173,1413,207,1861]" captionTargetId="figure@11.[163,1423,193,1868]" captionTargetPageId="11" captionText="FIGURE 6. Urocardiac ossicle of the subgenus Minuca species. A, Uca brevifrons (ZRC 2012.0126, CW 22.1, ♂); B, U. cf. virens (NCHUZOOL 13946, CW 17.0 mm, ♂); C, U. victoriana (NCHUZOOL 13945, CW 12.2 mm, ♀); D, U. pugnax (NCHUZOOL 13941, CW 15.2 mm, ♂); E, U. burgersi (NCHUZOOL 13956, CW 13.0 mm); F, U. rapax (NCHUZOOL 13942, CW 23.4 mm, ♂); G, U. mordax (NCHUZOOL 13940, CW 18.0 mm, ♀); H, U. minax (NCHUZOOL 13957, CW 23.2 mm, ♂); I, U. vocator (NCHUZOOL 13948, CW 21.9 mm, ♀). Scale = 1 mm." httpUri="https://zenodo.org/record/235579/files/figure.png" pageId="14" pageNumber="485">Figs. 6</figureCitation>
|
||
,
|
||
<figureCitation id="74F32A41FFD870711B7CEC2EFA2E7432" box="[1335,1351,1628,1652]" captionStart="FIGURE 7" captionStartId="12.[151,250,1882,1904]" captionTargetBox="[176,1408,203,1846]" captionTargetId="figure@12.[165,1421,193,1861]" captionTargetPageId="12" captionText="FIGURE 7. Urocardiac ossicle of the subgenus Leptuca species. A, Uca subcylindrica (NCHUZOOL 13952, CW 15.2 mm, ♂); B, U. terpsichores (NCHUZOOL 13582, CW 9.9 mm, ♂); C, U. uruguayensis (NCHUZOOL 13577, CW 14.1 mm, ♂); D, U. spinicarpa (NCHUZOOL 13947, CW 13.3 mm, ♀); E, U. panacea (NCHUZOOL 13950, CW 14.8 mm, ♂); F, U. pugilator (NCHUZOOL 13586, CW 15.9 mm, ♂); G, U. speciosa (NCHUZOOL 13951, CW 12.6 mm, ♂); H, U. umbratila (NCHUZOOL 13579, CW 16.2 mm, ♂); I, U. thayeri (NCHUZOOL 13953, CW 19.0 mm, ♂). Scale = 1 mm." httpUri="https://zenodo.org/record/235580/files/figure.png" pageId="14" pageNumber="485">7</figureCitation>
|
||
). In
|
||
<taxonomicName id="2BC84D47FFD870711BCAEC2FFF9C74DE" class="Malacostraca" family="Ocypodidae" genus="Uca" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="485" phylum="Arthropoda" rank="species" species="burgersi">
|
||
<emphasis id="DEBCEAD6FFD870711BCAEC2FFF9C74DE" italics="true" pageId="14" pageNumber="485">U. burgersi</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="2BC84D47FFD870711F4FECF3FE0274DE" box="[260,363,1665,1688]" class="Malacostraca" family="Ocypodidae" genus="Uca" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="485" phylum="Arthropoda" rank="species" species="minax">
|
||
<emphasis id="DEBCEAD6FFD870711F4FECF3FE0274DE" box="[260,363,1665,1688]" italics="true" pageId="14" pageNumber="485">U. minax</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="2BC84D47FFD870711F31ECF3FE9874DE" box="[378,497,1663,1688]" class="Malacostraca" family="Ocypodidae" genus="Uca" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="485" phylum="Arthropoda" rank="species" species="mordax">
|
||
<emphasis id="DEBCEAD6FFD870711F31ECF3FE9874DE" box="[378,497,1663,1688]" italics="true" pageId="14" pageNumber="485">U. mordax</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="2BC84D47FFD870711C4AECF3FD1E74DE" box="[513,631,1665,1688]" class="Malacostraca" family="Ocypodidae" genus="Uca" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="485" phylum="Arthropoda" rank="species" species="pugnax">
|
||
<emphasis id="DEBCEAD6FFD870711C4AECF3FD1E74DE" box="[513,631,1665,1688]" italics="true" pageId="14" pageNumber="485">U. pugnax</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="2BC84D47FFD870711CCEECF3FD8374DE" box="[645,746,1665,1688]" class="Malacostraca" family="Ocypodidae" genus="Uca" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="485" phylum="Arthropoda" rank="species" species="rapax">
|
||
<emphasis id="DEBCEAD6FFD870711CCEECF3FD8374DE" box="[645,746,1665,1688]" italics="true" pageId="14" pageNumber="485">U. rapax</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="2BC84D47FFD870711CB2ECF3FCE674DE" box="[761,911,1665,1688]" class="Malacostraca" family="Ocypodidae" genus="Uca" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="485" phylum="Arthropoda" rank="species" species="victoriana">
|
||
<emphasis id="DEBCEAD6FFD870711CB2ECF3FCE674DE" box="[761,911,1665,1688]" italics="true" pageId="14" pageNumber="485">U. victoriana</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="2BC84D47FFD870711DD5ECF3FB4074DE" box="[926,1065,1663,1688]" class="Malacostraca" family="Ocypodidae" genus="Uca" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="485" phylum="Arthropoda" rank="species" species="virens">
|
||
<emphasis id="DEBCEAD6FFD870711DD5ECF3FCD074DE" box="[926,953,1665,1688]" italics="true" pageId="14" pageNumber="485">U.</emphasis>
|
||
cf.
|
||
<emphasis id="DEBCEAD6FFD870711DADECF3FB4074DE" box="[998,1065,1665,1688]" italics="true" pageId="14" pageNumber="485">virens</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="2BC84D47FFD870711A22ECF3FB8B74DE" box="[1129,1250,1665,1688]" class="Malacostraca" family="Ocypodidae" genus="Uca" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="485" phylum="Arthropoda" rank="species" species="vocator">
|
||
<emphasis id="DEBCEAD6FFD870711A22ECF3FB8B74DE" box="[1129,1250,1665,1688]" italics="true" pageId="14" pageNumber="485">U. vocator</emphasis>
|
||
</taxonomicName>
|
||
of
|
||
<taxonomicName id="2BC84D47FFD870711B42ECF3FA3674DE" box="[1289,1375,1665,1688]" class="Malacostraca" family="Ocypodidae" genus="Minuca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="485" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="DEBCEAD6FFD870711B42ECF3FA3674DE" box="[1289,1375,1665,1688]" italics="true" pageId="14" pageNumber="485">Minuca</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="74F32A41FFD870711B25EC0DFFCE74FB" captionStart="FIGURE 6" captionStartId="11.[151,250,1886,1908]" captionTargetBox="[173,1413,207,1861]" captionTargetId="figure@11.[163,1423,193,1868]" captionTargetPageId="11" captionText="FIGURE 6. Urocardiac ossicle of the subgenus Minuca species. A, Uca brevifrons (ZRC 2012.0126, CW 22.1, ♂); B, U. cf. virens (NCHUZOOL 13946, CW 17.0 mm, ♂); C, U. victoriana (NCHUZOOL 13945, CW 12.2 mm, ♀); D, U. pugnax (NCHUZOOL 13941, CW 15.2 mm, ♂); E, U. burgersi (NCHUZOOL 13956, CW 13.0 mm); F, U. rapax (NCHUZOOL 13942, CW 23.4 mm, ♂); G, U. mordax (NCHUZOOL 13940, CW 18.0 mm, ♀); H, U. minax (NCHUZOOL 13957, CW 23.2 mm, ♂); I, U. vocator (NCHUZOOL 13948, CW 21.9 mm, ♀). Scale = 1 mm." httpUri="https://zenodo.org/record/235579/files/figure.png" pageId="14" pageNumber="485">Fig. 6</figureCitation>
|
||
), there are 6–9 transverse ridges on the median teeth of the posterior tooth plate and 2–4 weak pairs of cusps on the stem region; except for
|
||
<taxonomicName id="2BC84D47FFD870711F83ECBBFD3274A6" box="[456,603,1735,1760]" class="Malacostraca" family="Ocypodidae" genus="Uca" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="485" phylum="Arthropoda" rank="species" species="brevifrons">
|
||
<emphasis id="DEBCEAD6FFD870711F83ECBBFD3274A6" box="[456,603,1735,1760]" italics="true" pageId="14" pageNumber="485">U. brevifrons</emphasis>
|
||
</taxonomicName>
|
||
, which has only four ridges on the posterior tooth plate and a pair of cusps on the stem region. Species of
|
||
<taxonomicName id="2BC84D47FFD870711FB1EC9FFD3C7542" box="[506,597,1773,1796]" class="Malacostraca" family="Ocypodidae" genus="Leptuca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="485" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="DEBCEAD6FFD870711FB1EC9FFD3C7542" box="[506,597,1773,1796]" italics="true" pageId="14" pageNumber="485">Leptuca</emphasis>
|
||
</taxonomicName>
|
||
show a more diverse pattern of the urocardiac ossicles (
|
||
<figureCitation id="74F32A41FFD870711A91EC9EFA497542" box="[1242,1312,1772,1797]" captionStart="FIGURE 7" captionStartId="12.[151,250,1882,1904]" captionTargetBox="[176,1408,203,1846]" captionTargetId="figure@12.[165,1421,193,1861]" captionTargetPageId="12" captionText="FIGURE 7. Urocardiac ossicle of the subgenus Leptuca species. A, Uca subcylindrica (NCHUZOOL 13952, CW 15.2 mm, ♂); B, U. terpsichores (NCHUZOOL 13582, CW 9.9 mm, ♂); C, U. uruguayensis (NCHUZOOL 13577, CW 14.1 mm, ♂); D, U. spinicarpa (NCHUZOOL 13947, CW 13.3 mm, ♀); E, U. panacea (NCHUZOOL 13950, CW 14.8 mm, ♂); F, U. pugilator (NCHUZOOL 13586, CW 15.9 mm, ♂); G, U. speciosa (NCHUZOOL 13951, CW 12.6 mm, ♂); H, U. umbratila (NCHUZOOL 13579, CW 16.2 mm, ♂); I, U. thayeri (NCHUZOOL 13953, CW 19.0 mm, ♂). Scale = 1 mm." httpUri="https://zenodo.org/record/235580/files/figure.png" pageId="14" pageNumber="485">Fig. 7</figureCitation>
|
||
), with 3–8 transverse ridges on the posterior tooth plate and 1–10 pairs of cusps on the stem region, e.g., 3 and 2 (= three ridges on posterior tooth plate and two pairs of cusps on stem region, hereinafter the same) of
|
||
<taxonomicName id="2BC84D47FFD870711A88ED47FA17750B" box="[1219,1406,1844,1869]" class="Malacostraca" family="Ocypodidae" genus="Uca" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="485" phylum="Arthropoda" rank="species" species="subcylindrica">
|
||
<emphasis id="DEBCEAD6FFD870711A88ED47FA17750B" box="[1219,1406,1844,1869]" italics="true" pageId="14" pageNumber="485">U. subcylindrica</emphasis>
|
||
</taxonomicName>
|
||
; 4 and 1 of
|
||
<taxonomicName id="2BC84D47FFD870711EB1ED2BFECC7536" box="[250,421,1879,1904]" class="Malacostraca" family="Ocypodidae" genus="Uca" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="485" phylum="Arthropoda" rank="species" species="terpsichores">
|
||
<emphasis id="DEBCEAD6FFD870711EB1ED2BFECC7536" box="[250,421,1879,1904]" italics="true" pageId="14" pageNumber="485">U. terpsichores</emphasis>
|
||
</taxonomicName>
|
||
; 4 and 2 of
|
||
<taxonomicName id="2BC84D47FFD870711C66ED2BFD8D7536" box="[557,740,1881,1904]" class="Malacostraca" family="Ocypodidae" genus="Uca" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="485" phylum="Arthropoda" rank="species" species="uruguayensis">
|
||
<emphasis id="DEBCEAD6FFD870711C66ED2BFD8D7536" box="[557,740,1881,1904]" italics="true" pageId="14" pageNumber="485">U. uruguayensis</emphasis>
|
||
</taxonomicName>
|
||
; 5 and 1 of
|
||
<taxonomicName id="2BC84D47FFD870711D27ED2BFB6F7536" box="[876,1030,1881,1904]" class="Malacostraca" family="Ocypodidae" genus="Uca" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="485" phylum="Arthropoda" rank="species" species="spinicarpa">
|
||
<emphasis id="DEBCEAD6FFD870711D27ED2BFB6F7536" box="[876,1030,1881,1904]" italics="true" pageId="14" pageNumber="485">U. spinicarpa</emphasis>
|
||
</taxonomicName>
|
||
; 5 and 2 of
|
||
<taxonomicName id="2BC84D47FFD870711AC7ED2BFA677536" box="[1164,1294,1881,1904]" class="Malacostraca" family="Ocypodidae" genus="Uca" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="485" phylum="Arthropoda" rank="species" species="panacea">
|
||
<emphasis id="DEBCEAD6FFD870711AC7ED2BFA677536" box="[1164,1294,1881,1904]" italics="true" pageId="14" pageNumber="485">U. panacea</emphasis>
|
||
</taxonomicName>
|
||
; 7 and 10 of
|
||
<taxonomicName id="2BC84D47FFD870711EDCED0FFE4875D3" box="[151,289,1916,1941]" class="Malacostraca" family="Ocypodidae" genus="Uca" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="485" phylum="Arthropoda" rank="species" species="pugilator">
|
||
<emphasis id="DEBCEAD6FFD870711EDCED0FFE4875D3" box="[151,289,1916,1941]" italics="true" pageId="14" pageNumber="485">U. pugilator</emphasis>
|
||
</taxonomicName>
|
||
; 8 and 3 of
|
||
<taxonomicName id="2BC84D47FFD870711FE3ED0FFD4375D2" box="[424,554,1917,1940]" class="Malacostraca" family="Ocypodidae" genus="Uca" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="485" phylum="Arthropoda" rank="species" species="speciosa">
|
||
<emphasis id="DEBCEAD6FFD870711FE3ED0FFD4375D2" box="[424,554,1917,1940]" italics="true" pageId="14" pageNumber="485">U. speciosa</emphasis>
|
||
</taxonomicName>
|
||
; 8 and 4 of
|
||
<taxonomicName id="2BC84D47FFD870711CFAED0FFC2875D3" box="[689,833,1916,1941]" class="Malacostraca" family="Ocypodidae" genus="Uca" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="485" phylum="Arthropoda" rank="species" species="umbratila">
|
||
<emphasis id="DEBCEAD6FFD870711CFAED0FFC2875D3" box="[689,833,1916,1941]" italics="true" pageId="14" pageNumber="485">U. umbratila</emphasis>
|
||
</taxonomicName>
|
||
; and 8 and 6 of
|
||
<taxonomicName id="2BC84D47FFD870711DBCED0FFB0075D3" box="[1015,1129,1916,1941]" class="Malacostraca" family="Ocypodidae" genus="Uca" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="485" phylum="Arthropoda" rank="species" species="thayeri">
|
||
<emphasis id="DEBCEAD6FFD870711DBCED0FFB0075D3" box="[1015,1129,1916,1941]" italics="true" pageId="14" pageNumber="485">U. thayeri</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="EC7736C4FFD870711E8CEDD2FAF5759A" blockId="14.[151,1437,151,2013]" pageId="14" pageNumber="485">
|
||
<bibRefCitation id="88594B35FFD870711E8CEDD2FE3675FF" author="Crane" box="[199,351,1952,1977]" pageId="14" pageNumber="485" refString="Crane, J. (1975) Fiddler Crabs of the World (Ocypodidae: Genus Uca). Princeton University Press, Princeton, New Jersey, xxiii + 736 pp." type="book" year="1975">Crane (1975)</bibRefCitation>
|
||
treated
|
||
<taxonomicName id="2BC84D47FFD870711F8BEDD3FD0375FE" box="[448,618,1953,1976]" class="Malacostraca" family="Ocypodidae" genus="Uca" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="485" phylum="Arthropoda" rank="species" species="panamensis">
|
||
<emphasis id="DEBCEAD6FFD870711F8BEDD3FD0375FE" box="[448,618,1953,1976]" italics="true" pageId="14" pageNumber="485">U. panamensis</emphasis>
|
||
</taxonomicName>
|
||
as a member of
|
||
<taxonomicName id="2BC84D47FFD870711D7EEDD3FCE375FE" box="[821,906,1953,1976]" class="Malacostraca" family="Ocypodidae" genus="Minuca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="485" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="DEBCEAD6FFD870711D7EEDD3FCE375FE" box="[821,906,1953,1976]" italics="true" pageId="14" pageNumber="485">Minuca</emphasis>
|
||
</taxonomicName>
|
||
, in part because of the similarity of their G1 structures, especially with
|
||
<taxonomicName id="2BC84D47FFD870711F87EDB7FD2F759A" box="[460,582,1989,2012]" class="Malacostraca" family="Ocypodidae" genus="Uca" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="485" phylum="Arthropoda" rank="species" species="vocator">
|
||
<emphasis id="DEBCEAD6FFD870711F87EDB7FD2F759A" box="[460,582,1989,2012]" italics="true" pageId="14" pageNumber="485">U. vocator</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="2BC84D47FFD870711C13EDB7FC70759B" box="[600,793,1988,2013]" class="Malacostraca" family="Ocypodidae" genus="Uca" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="485" phylum="Arthropoda" rank="species" species="ecuadoriensis">
|
||
<emphasis id="DEBCEAD6FFD870711C13EDB7FC70759B" box="[600,793,1988,2013]" italics="true" pageId="14" pageNumber="485">U. ecuadoriensis</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="2BC84D47FFD870711D15EDB7FC80759A" box="[862,1001,1989,2012]" class="Malacostraca" family="Ocypodidae" genus="Uca" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="485" phylum="Arthropoda" rank="species" species="pygmaea">
|
||
<emphasis id="DEBCEAD6FFD870711D15EDB7FC80759A" box="[862,1001,1989,2012]" italics="true" pageId="14" pageNumber="485">U. pygmaea</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="88594B35FFD870711DB0EDB6FB8C759B" author="Crane" box="[1019,1253,1988,2013]" pageId="14" pageNumber="485" refString="Crane, J. (1975) Fiddler Crabs of the World (Ocypodidae: Genus Uca). Princeton University Press, Princeton, New Jersey, xxiii + 736 pp." type="book" year="1975">Crane 1975: fig. 66</bibRefCitation>
|
||
). The G1 of
|
||
<emphasis id="DEBCEAD6FFD870711BCAEDB7FAF5759A" box="[1409,1436,1989,2012]" italics="true" pageId="14" pageNumber="485">U.</emphasis>
|
||
</paragraph>
|
||
<paragraph id="EC7736C4FFD970701EDCEAEBFB527293" blockId="15.[151,1436,151,213]" pageId="15" pageNumber="486">
|
||
<taxonomicName id="2BC84D47FFD970701EDCEAEBFE7272F6" box="[151,283,153,176]" class="Malacostraca" family="Ocypodidae" genus="Uca" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="486" phylum="Arthropoda" rank="species" species="panamensis">
|
||
<emphasis id="DEBCEAD6FFD970701EDCEAEBFE7272F6" box="[151,283,153,176]" italics="true" pageId="15" pageNumber="486">panamensis</emphasis>
|
||
</taxonomicName>
|
||
is still unusual, with the large inner process bent at right angles to cover the flanges (
|
||
<figureCitation id="74F32A41FFD970701B50EAE5FA0D72F6" box="[1307,1380,151,176]" captionStart="FIGURE 4" captionStartId="9.[151,250,1917,1939]" captionTargetBox="[159,1422,203,1887]" captionTargetId="figure@9.[151,1436,193,1896]" captionTargetPageId="9" captionText="FIGURE 4. Uca panamensis, right G 1 (neotype, USNM 1294205, CW 16.8 mm). A, dorsal view; B, dorsal view of terminal segment; C, D, ventral view of terminal segment. Scale, A = 2 mm; B, C = 0.5 mm." httpUri="https://zenodo.org/record/235577/files/figure.png" pageId="15" pageNumber="486">Fig. 4</figureCitation>
|
||
B), a character not seen in any
|
||
<taxonomicName id="2BC84D47FFD970701FFFEACFFD637292" box="[436,522,189,212]" class="Malacostraca" family="Ocypodidae" genus="Minuca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="486" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="DEBCEAD6FFD970701FFFEACFFD637292" box="[436,522,189,212]" italics="true" pageId="15" pageNumber="486">Minuca</emphasis>
|
||
</taxonomicName>
|
||
or
|
||
<taxonomicName id="2BC84D47FFD970701C64EACFFDE07292" box="[559,649,189,212]" class="Malacostraca" family="Ocypodidae" genus="Leptuca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="486" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="DEBCEAD6FFD970701C64EACFFDE07292" box="[559,649,189,212]" italics="true" pageId="15" pageNumber="486">Leptuca</emphasis>
|
||
</taxonomicName>
|
||
species (see
|
||
<bibRefCitation id="88594B35FFD970701D57EACEFCCB7293" author="Crane" box="[796,930,188,213]" pageId="15" pageNumber="486" refString="Crane, J. (1975) Fiddler Crabs of the World (Ocypodidae: Genus Uca). Princeton University Press, Princeton, New Jersey, xxiii + 736 pp." type="book" year="1975">Crane 1975</bibRefCitation>
|
||
: figs. 66–71).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |