204 lines
26 KiB
XML
204 lines
26 KiB
XML
<document id="7BBE5EBA42664E297E526E31E451FC06" ID-DOI="10.13133/2284-4880/545" ID-ISSN="2284-4880" ID-Zenodo-Dep="12764154" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1721242503917" checkinUser="felipe" docAuthor="Ballerio, Alberto" docDate="2021" docId="0F74755011D5FF72FCAEAD851B0794C8" docLanguage="en" docName="FragEntomol.53.2.105-282.pdf" docOrigin="Fragmenta entomologica 53 (2)" docSource="http://dx.doi.org/10.13133/2284-4880/545" docStyle="DocumentStyle:8499FF4308F512386C0E43DC290A16BD.1:FragEntomol.2021-.journal_article.0cover" docStyleId="8499FF4308F512386C0E43DC290A16BD" docStyleName="FragEntomol.2021-.journal_article.0cover" docStyleVersion="1" docTitle="Philharmostes acutus Ballerio, 2021, sp. nov." docType="treatment" docUuid="F092F52C-5DA5-48A6-82C1-01B7A7E8D3F1" docUuidSource="ZooBank" docVersion="2" lastPageNumber="249" masterDocId="F34D0D28115AFFE2FF9EAF31192A9224" masterDocTitle="The Ceratocanthinae of Madagascar and Comoro Islands: a revision of the genera Synarmostes and Goudotostes, and of the flightless Philharmostes, with description of 64 new species (Coleoptera: Scarabaeoidea, Hybosoridae)" masterLastPageNumber="282" masterPageNumber="105" pageNumber="248" updateTime="1721247602959" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-4.0">
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<mods:titleInfo id="5D067B884C39824D301D247858B72A59">
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<mods:title id="19F7A81DCD559661A701C5D905AE9E8E">The Ceratocanthinae of Madagascar and Comoro Islands: a revision of the genera Synarmostes and Goudotostes, and of the flightless Philharmostes, with description of 64 new species (Coleoptera: Scarabaeoidea, Hybosoridae)</mods:title>
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</mods:titleInfo>
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<mods:name id="F33DB859E9BA0E74EE8E633019AEE833" type="personal">
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<mods:roleTerm id="03DA69BBEA9CD6E2011B616CF6C2F2CF">Author</mods:roleTerm>
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<mods:namePart id="17924B633CE2E8DE85FD7CF0C9066A14">Ballerio, Alberto</mods:namePart>
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<mods:typeOfResource id="AA96B9DE184BB6E20272D4F1708C5263">text</mods:typeOfResource>
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<mods:title id="8BF6BFA1646EE29B91F83159D704A58A">Fragmenta entomologica</mods:title>
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</mods:titleInfo>
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<mods:part id="C38BEF7F5ECCC37789AA878315498E65">
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<mods:date id="03C5C05956F0760A9EE6A4332F2DA106">2021</mods:date>
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<mods:detail id="716D4703F9860DB5C9809DF04B6C7C81" type="pubDate">
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<mods:number id="C029554CB56C97F642714D8CA9638EDE">2021-11-30</mods:number>
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<mods:number id="33CF694D6CE3A28D4CEA24E8D6FED206">53</mods:number>
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<mods:detail id="6207D2B95E773FC1E127E60F61B03A23" type="issue">
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<mods:number id="28989886006EF180C630B2CE25A7C551">2</mods:number>
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<mods:start id="7681CFC5D31B57E5CC680341016610C5">105</mods:start>
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<mods:url id="0738EF69E5154C40416288E9BDE05845">http://dx.doi.org/10.13133/2284-4880/545</mods:url>
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</mods:location>
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<mods:classification id="049D4EA968353259C6042705750EF7D9">journal article</mods:classification>
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<mods:identifier id="4E3A0E89170DA0348A82FB05073224A4" type="DOI">10.13133/2284-4880/545</mods:identifier>
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<mods:identifier id="5B39FD05638120128A0673CCF4B484BA" type="ISSN">2284-4880</mods:identifier>
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<subSubSection id="CFC797CD11D5FF6DFCAEAD851D5990E8" box="[816,1139,692,716]" pageId="143" pageNumber="248" type="nomenclature">
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<paragraph id="8762C44611D5FF6DFCAEAD851D5990E8" blockId="143.[816,1139,692,716]" box="[816,1139,692,716]" pageId="143" pageNumber="248">
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<heading id="DC2A732A11D5FF6DFCAEAD851D5990E8" box="[816,1139,692,716]" fontSize="10" level="2" pageId="143" pageNumber="248" reason="5">
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<taxonomicName id="40DDBFC511D5FF6DFCAEAD851D3D90E8" ID-CoL="8SSR4" authorityName="Ballerio" authorityYear="2021" box="[816,1047,692,716]" class="Insecta" family="Hybosoridae" genus="Philharmostes" kingdom="Animalia" order="Coleoptera" pageId="143" pageNumber="248" phylum="Arthropoda" rank="species" species="acutus" status="sp. nov.">
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<emphasis id="B5A9185411D5FF6DFCAEAD851D3D90E8" box="[816,1047,692,716]" italics="true" pageId="143" pageNumber="248">Philharmostes acutus</emphasis>
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</taxonomicName>
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<emphasis id="B5A9185411D5FF6DFB83AD851D5990E8" bold="true" box="[1053,1139,692,716]" pageId="143" pageNumber="248">
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<taxonomicNameLabel id="AE9AA52F11D5FF6DFB83AD851D5990E8" box="[1053,1139,692,716]" pageId="143" pageNumber="248" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="CFC797CD11D5FF6DFCAEADE51D9C9108" pageId="143" pageNumber="248" type="description">
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<paragraph id="8762C44611D5FF6DFCAEADE51D9C9108" blockId="143.[816,1436,724,812]" pageId="143" pageNumber="248">
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(
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<figureCitation id="1FE6D8C311D5FF6DFCA6ADE51AA390C8" box="[824,905,724,748]" captionStart="Fig" captionStartId="157.[136,167,1858,1877]" captionTargetBox="[173,1363,184,1829]" captionTargetId="figure-4@157.[110,1403,166,1862]" captionTargetPageId="157" captionText="Fig. 82 – Philharmostes schweikerti sp. nov., a, dorsal view of distended paratype; b, ventral view of rolled up paratype; c, dorsal view of rolled up paratype; d, lateral view of rolled up paratype; Philharmostes acutus sp. nov., e, dorsal view of distended paratype; f, ventral view of rolled up paratype; g, dorsal view of rolled up paratype; h, lateral view of rolled up paratype." figureDoi="http://doi.org/10.5281/zenodo.12764353" httpUri="https://zenodo.org/record/12764353/files/figure.png" pageId="143" pageNumber="248">Figs 82</figureCitation>
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e-h,
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<quantity id="402569A311D5FF6DFC21ADE51AC590C9" box="[959,1007,724,749]" metricMagnitude="-2" metricUnit="kg" metricValue="8.5" pageId="143" pageNumber="248" unit="g" value="85.0">85 g</quantity>
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, 87 a-c, 89) http://zoobank.org/
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<uri id="F34CC84411D5FF6DFB94ADC51D9C9108" pageId="143" pageNumber="248">
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urn:lsid:zoobank.org:act:
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<uuid id="F37BFE9311D5FF6DFA86ADC51D9C9108" pageId="143" pageNumber="248">F092F52C- 5DA5-48A6-82C1-01B7A7E8D3F1</uuid>
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</uri>
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="CFC797CD11D5FF6DFCC8AC651DDB9688" pageId="143" pageNumber="248" type="materials_examined">
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<paragraph id="8762C44611D5FF6DFCC8AC651DDB9688" blockId="143.[816,1438,852,1932]" pageId="143" pageNumber="248">
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<emphasis id="B5A9185411D5FF6DFCC8AC651AF49148" bold="true" box="[854,990,852,876]" pageId="143" pageNumber="248">Type series.</emphasis>
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<materialsCitation id="37B5CE1B11D5FF6DFC78AC651C099648" collectingDate="2005-11-14" collectorName="San Francisco, CA & Brian L. Fisher" country="Madagascar" county="Academy of Sciences" determinerName="A. Ballerio" elevation="225" latitude="-16.455" location="Mananara-Nord" longLatPrecision="21" longitude="49.7875" municipality="Parc" pageId="143" pageNumber="248" specimenCount="1" stateProvince="Toamasina" typeStatus="holotype">
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<typeStatus id="58667AE411D5FF6DFC78AC651D669148" box="[998,1100,852,876]" pageId="143" pageNumber="248" type="holotype">Holotype</typeStatus>
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, sex unstated, deposited in the collection of the California
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<collectingCounty id="6E03BCCA11D5FF6DFBF0AC451C4E91A8" box="[1134,1380,884,908]" pageId="143" pageNumber="248">Academy of Sciences</collectingCounty>
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||
,
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<collectorName id="2A28A19011D5FF6DFAEAAC451AF99188" pageId="143" pageNumber="248">San Francisco, CA</collectorName>
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, USA, labelled as follows: / CASTYPE 20288 / CASENT 8135322 /
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<collectingCountry id="FFCA84D611D5FF6DFB0CAC851C3D91E8" box="[1170,1303,948,972]" name="Madagascar" pageId="143" pageNumber="248">Madagascar</collectingCountry>
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:
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<collectingRegion id="45190AA411D5FF6DFAB7AC851A4391C8" country="Madagascar" name="Toamasina" pageId="143" pageNumber="248">Toamasina Prov.</collectingRegion>
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<collectingMunicipality id="67065E3C11D5FF6DFCECACE41A8B91C8" box="[882,929,981,1004]" pageId="143" pageNumber="248">Parc</collectingMunicipality>
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national
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<location id="8202929D11D5FF6DFB93ACE51D9091C8" LSID="urn:lsid:plazi:treatment:0F74755011D5FF72FCAEAD851B0794C8:8202929D11D5FF6DFB93ACE51D9091C8" box="[1037,1210,980,1004]" country="Madagascar" county="Academy of Sciences" latitude="-16.455" longLatPrecision="21" longitude="49.7875" municipality="Parc" name="Mananara-Nord" pageId="143" pageNumber="248" stateProvince="Toamasina">Mananara-Nord</location>
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,
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||
<quantity id="402569A311D5FF6DFB56ACE51C2691C8" box="[1224,1292,980,1004]" metricMagnitude="3" metricUnit="m" metricValue="7.1" pageId="143" pageNumber="248" unit="km" value="7.1">7.1km</quantity>
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261°
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||
<location id="8202929D11D5FF6DFACCACE51A409628" LSID="urn:lsid:plazi:treatment:0F74755011D5FF72FCAEAD851B0794C8:8202929D11D5FF6DFACCACE51A409628" country="Madagascar" county="Academy of Sciences" latitude="-16.455" longLatPrecision="21" longitude="49.7875" municipality="Parc" name="Antanambe" pageId="143" pageNumber="248" stateProvince="Toamasina">Antanambe</location>
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||
elev
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||
<quantity id="402569A311D5FF6DFC3AACC51AC29628" box="[932,1000,1012,1036]" metricMagnitude="2" metricUnit="m" metricValue="2.25" pageId="143" pageNumber="248" unit="m" value="225.0">
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<elevation id="0CF0237511D5FF6DFC3AACC51AC29628" box="[932,1000,1012,1036]" metricMagnitude="2" metricUnit="m" metricValue="2.25" pageId="143" pageNumber="248" unit="m" value="225.0">225 m</elevation>
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</quantity>
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||
<date id="F363E28611D5FF6DFC71ACC51D979628" box="[1007,1213,1012,1036]" pageId="143" pageNumber="248" value="2005-11-14">
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||
<collectingDate id="E3271B6E11D5FF6DFC71ACC51D979628" box="[1007,1213,1012,1036]" pageId="143" pageNumber="248" value="2005-11-14">14 November 2005</collectingDate>
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||
</date>
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||
/
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||
<geoCoordinate id="E2E9A28111D5FF6DFB4CACC51C499628" box="[1234,1379,1012,1036]" degrees="16" direction="south" minutes="27" orientation="latitude" pageId="143" pageNumber="248" precision="15" seconds="18" value="-16.455">16° 27’ 18” S</geoCoordinate>
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<geoCoordinate id="E2E9A28111D5FF6DFAF4ACC51AB79608" degrees="049" direction="east" minutes="47" orientation="longitude" pageId="143" pageNumber="248" precision="15" seconds="15" value="49.7875">049° 47’ 15” E</geoCoordinate>
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||
/
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||
<location id="8202929D11D5FF6DFC24AB251C379608" LSID="urn:lsid:plazi:treatment:0F74755011D5FF72FCAEAD851B0794C8:8202929D11D5FF6DFC24AB251C379608" box="[954,1309,1044,1068]" country="Madagascar" county="Academy of Sciences" latitude="-16.455" longLatPrecision="21" longitude="49.7875" municipality="Parc" name="California Academy of Sciences" pageId="143" pageNumber="248" stateProvince="Toamasina">California Academy of Sciences</location>
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||
coll.
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<collectorName id="2A28A19011D5FF6DFAFFAB251ABF9668" pageId="143" pageNumber="248">Brian L. Fisher</collectorName>
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et al. sifted litter rainforest BLF 12556/
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<taxonomicName id="40DDBFC511D5FF6DFAF9AB051AC49648" authorityName="Ballerio" authorityYear="2021" class="Insecta" family="Hybosoridae" genus="Philharmostes" kingdom="Animalia" order="Coleoptera" pageId="143" pageNumber="248" phylum="Arthropoda" rank="species" species="acutus" status="sp. nov.">
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||
<emphasis id="B5A9185411D5FF6DFAF9AB051AC49648" italics="true" pageId="143" pageNumber="248">Philharmostes acutus</emphasis>
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||
</taxonomicName>
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||
<taxonomicNameLabel id="AE9AA52F11D5FF6DFC69AB641D199648" box="[1015,1075,1109,1132]" pageId="143" pageNumber="248" rank="species">n. sp.</taxonomicNameLabel>
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det.
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<determinerName id="72BE288211D5FF6DFBF5AB651DCF9648" box="[1131,1253,1108,1132]" pageId="143" pageNumber="248">A. Ballerio</determinerName>
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2020
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</materialsCitation>
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<typeStatus id="58667AE411D5FF6DFAB3AB651CB69648" box="[1325,1436,1108,1132]" pageId="143" pageNumber="248" type="holotype">Holotypus</typeStatus>
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/.
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<materialsCitation id="37B5CE1B11D5FF6DFCD6AB441DC79688" collectingDate="2005-11-14" collectionCode="CASC, ABCB" collectorName="San Francisco, CA & Brian L. Fisher" country="Madagascar" county="Academy of Sciences" determinerName="A. Ballerio" elevation="225" latitude="-16.455" location="Mananara-Nord" longLatPrecision="21" longitude="49.7875" municipality="Parc" pageId="143" pageNumber="248" specimenCount="1" stateProvince="Toamasina" typeStatus="paratype">
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<typeStatus id="58667AE411D5FF6DFCD6AB441A9E96A9" box="[840,948,1141,1165]" pageId="143" pageNumber="248" type="paratype">Paratypes</typeStatus>
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: 3 exx. (sex unstated) (
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<quantity id="402569A311D5FF6DFB24AB451DCD96A8" box="[1210,1255,1140,1164]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="143" pageNumber="248" unit="in" value="2.0">2 in</quantity>
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<collectionCode id="E1CC5C8311D5FF6DFB6EAB451C1C96A8" box="[1264,1334,1140,1164]" pageId="143" pageNumber="248">CASC</collectionCode>
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||
and
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||
<quantity id="402569A311D5FF6DFAEEAB451CB696A8" box="[1392,1436,1140,1164]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="143" pageNumber="248" unit="in" value="1.0">1 in</quantity>
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||
<collectionCode id="E1CC5C8311D5FF6DFCAEABA51A579688" box="[816,893,1172,1196]" pageId="143" pageNumber="248">ABCB</collectionCode>
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), same collecting data as holotype
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</materialsCitation>
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||
.
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="CFC797CD11D5FF6DFCC8AB851CBD94C8" pageId="143" pageNumber="248" type="diagnosis">
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||
<paragraph id="8762C44611D5FF6DFCC8AB851CBD94C8" blockId="143.[816,1438,852,1932]" pageId="143" pageNumber="248">
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||
<emphasis id="B5A9185411D5FF6DFCC8AB851AEF96E8" bold="true" box="[854,965,1204,1228]" pageId="143" pageNumber="248">Diagnosis.</emphasis>
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||
<taxonomicName id="40DDBFC511D5FF6DFC50AB851D8696E8" authorityName="Ballerio" authorityYear="2021" box="[974,1196,1204,1228]" class="Insecta" family="Hybosoridae" genus="Philharmostes" kingdom="Animalia" order="Coleoptera" pageId="143" pageNumber="248" phylum="Arthropoda" rank="species" species="acutus" status="sp. nov.">
|
||
<emphasis id="B5A9185411D5FF6DFC50AB851D8696E8" box="[974,1196,1204,1228]" italics="true" pageId="143" pageNumber="248">Philharmostes acutus</emphasis>
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||
</taxonomicName>
|
||
<emphasis id="B5A9185411D5FF6DFB28AB851C2096E8" bold="true" box="[1206,1290,1204,1228]" pageId="143" pageNumber="248">
|
||
<taxonomicNameLabel id="AE9AA52F11D5FF6DFB28AB851C2096E8" box="[1206,1290,1204,1228]" pageId="143" pageNumber="248" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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can be differentiated from all other species of
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<taxonomicName id="40DDBFC511D5FF6DFB0FABE51C0F96C8" authorityName="Kolbe" authorityYear="1895" box="[1169,1317,1236,1260]" class="Insecta" family="Hybosoridae" genus="Philharmostes" kingdom="Animalia" order="Coleoptera" pageId="143" pageNumber="248" phylum="Arthropoda" rank="genus">
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||
<emphasis id="B5A9185411D5FF6DFB0FABE51C0F96C8" box="[1169,1317,1236,1260]" italics="true" pageId="143" pageNumber="248">Philharmostes</emphasis>
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</taxonomicName>
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by the following combination of characters: a) flightlessness, b) dorsal ocular area small, c) head with clypeal disc covered by large impressed simple punctation, d) dorsum with brown colour, without metallic sheen, e) pronotal fore angles broadly truncate, f) pronotum with disc slightly raised with respect to sides, g) pronotal disc covered by large horseshoe-shaped punctation mixed to some large comma-shaped punctation, h) elytral lateral carina complete, h) elytral lateral carina sharp, i) elytral lateral carina slightly sinuate in lateral view, j) elytral lateral carina in correspondence with humeral area forming a sharp angle (dorsal view), k) elytral basal third with parasutural area with large horseshoe-shaped punctures with opening directed backwards or outwards, l) elytral punctation made of large horseshoe-shaped punctures, m) elytral fore angles distinct and strong, n) head with vertex having one irregular transverse row of larger impressed simple punctures.
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="CFC797CD11D5FF72FCC8A9C518429748" lastPageId="144" lastPageNumber="249" pageId="143" pageNumber="248" type="description">
|
||
<paragraph id="8762C44611D5FF6DFCC8A9C51CB095A8" blockId="143.[816,1438,852,1932]" pageId="143" pageNumber="248">
|
||
<emphasis id="B5A9185411D5FF6DFCC8A9C51ACB9528" bold="true" box="[854,993,1780,1804]" pageId="143" pageNumber="248">Description.</emphasis>
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HL:
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||
<quantity id="402569A311D5FF6DFB84A9C51D459528" box="[1050,1135,1780,1804]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="143" pageNumber="248" unit="mm" value="0.6">0.6 mm</quantity>
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||
; HW:
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<quantity id="402569A311D5FF6DFB2BA9C51C209528" box="[1205,1290,1780,1804]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="143" pageNumber="248" unit="mm" value="1.1">1.1 mm</quantity>
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||
; PL: 1.0 mm; PW:
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<quantity id="402569A311D5FF6DFCF6A8251AEA9508" box="[872,960,1812,1836]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" pageId="143" pageNumber="248" unit="mm" value="1.9">1.9 mm</quantity>
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; EL: 2.0 mm; EW 2.0; overall morphology as in generic diagnosis. Small flightless
|
||
<taxonomicName id="40DDBFC511D5FF6DFB76A8051CB39568" authorityName="Martinez" authorityYear="1968" box="[1256,1433,1844,1868]" class="Insecta" family="Hybosoridae" kingdom="Animalia" order="Coleoptera" pageId="143" pageNumber="109" phylum="Arthropoda" rank="subFamily" subFamily="Ceratocanthinae">Ceratocanthinae</taxonomicName>
|
||
. Body moderately convex. Dark brown, shiny, dorsum glabrous (40×), underside, tarsi and antennae reddish-brown.
|
||
</paragraph>
|
||
<paragraph id="8762C44611CAFF72FF23AF8518429748" blockId="144.[150,772,180,1772]" pageId="144" pageNumber="249">
|
||
<emphasis id="B5A9185411CAFF72FF23AF8519DF92E8" box="[189,245,180,204]" italics="true" pageId="144" pageNumber="249">Head</emphasis>
|
||
: W/L ratio = 1.36; apical margin triangular with angle blunt and obtuse and sides weakly curved; genae produced outwards, acute; genal canthus complete, fused with the occipital portion of head; dorsal ocular area small-sized, interocular distance about 19 times the maximum width of dorsal ocular area, disc uniformly covered by large impressed simple punctation, fore margin with some irregular impressed transverse short lines. Interpunctural distance of simple punctures once to three times larger than punctural diameter.
|
||
<emphasis id="B5A9185411CAFF72FE80AEE418AC93C8" box="[286,390,469,492]" italics="true" pageId="144" pageNumber="249">Pronotum</emphasis>
|
||
: short and transverse (W/L ratio = 1.73), weakly convex, surface irregular, with discal area slightly raised with respect to sides; base and sides broadly rounded (dorsal view); fore margin thick, distinct along median portion then less distinct and discontinuous till fore angles, fore angles gently truncate, disc covered by impressed comma-shaped punctures, sides covered by dense large relatively shallow horseshoe-shaped punctures, with opening oriented outwards. Each puncture bearing in the middle a very short simple seta. Interpunctural distance shorter than punctural diameter.
|
||
<taxonomicName id="40DDBFC511CAFF72FE77AC251B7A9108" box="[489,592,788,812]" class="Trilobita" family="Styginidae" genus="Scutellum" kingdom="Animalia" order="Corynexochida" pageId="144" pageNumber="249" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B5A9185411CAFF72FE77AC251B7A9108" box="[489,592,788,812]" italics="true" pageId="144" pageNumber="249">Scutellum</emphasis>
|
||
</taxonomicName>
|
||
: covered by several small impressed horseshoe-shaped.
|
||
<emphasis id="B5A9185411CAFF72FDA3AC051B549168" box="[573,638,820,844]" italics="true" pageId="144" pageNumber="249">Elytra</emphasis>
|
||
: longer than wide (W/L ratio =0.93); subrectangular (dorsal view); humeral callus indistinct but marked by the thick starting point of lateral carina forming a sharp angle (dorsal view); elytral surface convex, covered by dense (interpunctural distance much shorter than punctural diameter) relatively shallow punctation. Punctures on proximal and median third horseshoe-shaped with a broad opening directed outwards, distal third covered by dense horseshoe-shaped punctures with a short opening directed backwards. Lateral carina distinct, thick, sharp and complete, slightly sinuate (lateral view), and forming a sharp angle in correspondence with humeral area forming a right angle (dorsal view). Pseudoepileura covered by dense broad horseshoe-shaped shallow punctures with a broad opening directed towards lateral carina.
|
||
<emphasis id="B5A9185411CAFF72FF09AA2419FD9708" box="[151,215,1301,1324]" italics="true" pageId="144" pageNumber="249">Wings</emphasis>
|
||
: flightless.
|
||
<emphasis id="B5A9185411CAFF72FEC8AA2518D99708" box="[342,499,1300,1324]" italics="true" pageId="144" pageNumber="249">Male genitalia</emphasis>
|
||
: aedeagus (L =
|
||
<quantity id="402569A311CAFF72FD39AA251BD69708" box="[679,764,1300,1324]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="144" pageNumber="249" unit="mm" value="0.9">0.9 mm</quantity>
|
||
) with parameres as in
|
||
<figureCitation id="1FE6D8C311CAFF72FEEDAA0518EA9768" box="[371,448,1332,1356]" captionStart="Fig" captionStartId="162.[151,182,1885,1904]" captionTargetBox="[214,1388,167,1862]" captionTargetId="figure-13@162.[211,1391,163,1865]" captionTargetPageId="162" captionText="Fig. 87 – Aedeagi and spicula gastrales, a–c, Philharmostes acutus sp. nov.; d–e, Philharmostes apterus sp. nov.; f–h, Philharmostes nelsoni sp. nov. (paratype from Col de Tanatana)." figureDoi="http://doi.org/10.5281/zenodo.12764365" httpUri="https://zenodo.org/record/12764365/files/figure.png" pageId="144" pageNumber="249">Figs 87</figureCitation>
|
||
a-b, spiculum gastrale (L =
|
||
<quantity id="402569A311CAFF72FD7CAA0519E89748" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="144" pageNumber="249" unit="mm" value="0.7">0.7 mm</quantity>
|
||
) as in
|
||
<figureCitation id="1FE6D8C311CAFF72FE98AA65184E9748" box="[262,356,1364,1388]" captionStart="Fig" captionStartId="162.[151,182,1885,1904]" captionTargetBox="[214,1388,167,1862]" captionTargetId="figure-13@162.[211,1391,163,1865]" captionTargetPageId="162" captionText="Fig. 87 – Aedeagi and spicula gastrales, a–c, Philharmostes acutus sp. nov.; d–e, Philharmostes apterus sp. nov.; f–h, Philharmostes nelsoni sp. nov. (paratype from Col de Tanatana)." figureDoi="http://doi.org/10.5281/zenodo.12764365" httpUri="https://zenodo.org/record/12764365/files/figure.png" pageId="144" pageNumber="249">Fig. 87 c</figureCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="CFC797CD11CAFF72FF23AA451BDA9788" pageId="144" pageNumber="249" type="etymology">
|
||
<paragraph id="8762C44611CAFF72FF23AA451BDA9788" blockId="144.[150,772,180,1772]" pageId="144" pageNumber="249">
|
||
<emphasis id="B5A9185411CAFF72FF23AA45181497A8" bold="true" box="[189,318,1396,1420]" pageId="144" pageNumber="249">Etymology.</emphasis>
|
||
Latin adjective in the nominative singular, meaning “acute”, referring to the humeral area of elytra.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="CFC797CD11CAFF72FF23AA851B0794C8" pageId="144" pageNumber="249" type="distribution">
|
||
<paragraph id="8762C44611CAFF72FF23AA851B0794C8" blockId="144.[150,772,180,1772]" pageId="144" pageNumber="249">
|
||
<emphasis id="B5A9185411CAFF72FF23AA8518F697E8" bold="true" box="[189,476,1460,1484]" pageId="144" pageNumber="249">Distribution and habitat.</emphasis>
|
||
Known only from the
|
||
<typeStatus id="58667AE411CAFF72FD4BAA841A2997E9" box="[725,771,1461,1485]" pageId="144" pageNumber="249">type</typeStatus>
|
||
locality in Eastern
|
||
<collectingCountry id="FFCA84D611CAFF72FEFDAAE518CC97C8" box="[355,486,1492,1516]" name="Madagascar" pageId="144" pageNumber="249">Madagascar</collectingCountry>
|
||
. The area is dominated by the humid climate of the east coast, with a heavy average annual rainfall. The
|
||
<typeStatus id="58667AE411CAFF72FEE5A92418839409" box="[379,425,1557,1581]" pageId="144" pageNumber="249">type</typeStatus>
|
||
series was collected by sifting leaf litter in rainforest. The vegetation cover consists of “humid forest” (
|
||
<bibRefCitation id="E34CB9B711CAFF72FEDFA9651B3C9448" author="Moat J. & Smith P." box="[321,534,1620,1644]" pageId="144" pageNumber="249" refId="ref103114" refString="Moat J., Smith P. 2007. Atlas of the vegetation of Madagascar. Royal Botanic Garden, Kew 124 pp." type="book" year="2007">Moat & Smith 2007</bibRefCitation>
|
||
) also known as “moist evergreen forest” (
|
||
<bibRefCitation id="E34CB9B711CAFF72FEFDA9451B7894A8" author="Goodman S. M. & Raherilalao M. J. & Wohlhauser S." box="[355,594,1652,1676]" pageId="144" pageNumber="249" refId="ref102713" refString="Goodman S. M., Raherilalao M. J., Wohlhauser S. (eds). 2018. Les aires potegees terrestres de Madagascar: leur histoire, description et biote / The terrestrial protected areas of Madagascar: their history, description and biota. Association Vahatra, Antananarivo. 1716 pp." type="book" year="2018">Goodman et al. 2018</bibRefCitation>
|
||
). The collecting locality falls within the protected areas “Parc National de Mananara-Nord”, more precisely within the Verezanantso- ro Massif section of the national park.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |