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<document id="63CFA3C8D544F1C53F23CB600ADF136A" ID-DOI="10.13133/2284-4880/545" ID-ISSN="2284-4880" ID-Zenodo-Dep="12764154" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1721242503917" checkinUser="felipe" docAuthor="Ballerio, Alberto" docDate="2021" docId="0F74755011CAFF73FF09A8251B809068" docLanguage="en" docName="FragEntomol.53.2.105-282.pdf" docOrigin="Fragmenta entomologica 53 (2)" docSource="http://dx.doi.org/10.13133/2284-4880/545" docStyle="DocumentStyle:8499FF4308F512386C0E43DC290A16BD.1:FragEntomol.2021-.journal_article.0cover" docStyleId="8499FF4308F512386C0E43DC290A16BD" docStyleName="FragEntomol.2021-.journal_article.0cover" docStyleVersion="1" docTitle="Philharmostes apterus Ballerio, 2021, sp. nov." docType="treatment" docUuid="D56FBECA-8917-4959-AC3D-528CF53E48C5" docUuidSource="ZooBank" docVersion="2" lastPageNumber="250" masterDocId="F34D0D28115AFFE2FF9EAF31192A9224" masterDocTitle="The Ceratocanthinae of Madagascar and Comoro Islands: a revision of the genera Synarmostes and Goudotostes, and of the flightless Philharmostes, with description of 64 new species (Coleoptera: Scarabaeoidea, Hybosoridae)" masterLastPageNumber="282" masterPageNumber="105" pageNumber="249" updateTime="1721247602959" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-4.0">
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<mods:title id="F21169F9315530E7A07A2B461A8FAED9">The Ceratocanthinae of Madagascar and Comoro Islands: a revision of the genera Synarmostes and Goudotostes, and of the flightless Philharmostes, with description of 64 new species (Coleoptera: Scarabaeoidea, Hybosoridae)</mods:title>
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<mods:name id="1C57D6CB977732A60B2B6FBF4373DE2A" type="personal">
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<mods:namePart id="0CE1ECF529363596D56F9CD7CEBCDACB">Ballerio, Alberto</mods:namePart>
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<mods:title id="A52B98204E4D06298AC9D9D19E59104C">Fragmenta entomologica</mods:title>
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<mods:part id="CA664D8BD1BE7229040E5E759067105C">
<mods:date id="7BE7923D6A7629364B7E52B173D9B0FD">2021</mods:date>
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<mods:number id="5E2B85334246840A76CE727FAE6624C9">2021-11-30</mods:number>
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<mods:classification id="23DCE71622C48DD287DCEE265A492D3C">journal article</mods:classification>
<mods:identifier id="ABB44807791C4198EE33E29F8114A9AD" type="DOI">10.13133/2284-4880/545</mods:identifier>
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<treatment id="0F74755011CAFF73FF09A8251B809068" LSID="urn:lsid:zoobank.org:act:D56FBECA-8917-4959-AC3D-528CF53E48C5" httpUri="http://treatment.plazi.org/id/0F74755011CAFF73FF09A8251B809068" lastPageId="145" lastPageNumber="250" pageId="144" pageNumber="249">
<subSubSection id="CFC797CD11CAFF72FF09A82518CE9508" box="[151,484,1812,1836]" pageId="144" pageNumber="249" type="nomenclature">
<paragraph id="8762C44611CAFF72FF09A82518CE9508" blockId="144.[151,484,1812,1836]" box="[151,484,1812,1836]" pageId="144" pageNumber="249">
<heading id="DC2A732A11CAFF72FF09A82518CE9508" box="[151,484,1812,1836]" fontSize="10" level="2" pageId="144" pageNumber="249" reason="5">
<taxonomicName id="40DDBFC511CAFF72FF09A82518A29508" ID-CoL="8SSSB" authorityName="Ballerio" authorityYear="2021" box="[151,392,1812,1836]" class="Insecta" family="Hybosoridae" genus="Philharmostes" kingdom="Animalia" order="Coleoptera" pageId="144" pageNumber="249" phylum="Arthropoda" rank="species" species="apterus" status="sp. nov.">
<emphasis id="B5A9185411CAFF72FF09A82518A29508" box="[151,392,1812,1836]" italics="true" pageId="144" pageNumber="249">Philharmostes apterus</emphasis>
</taxonomicName>
<emphasis id="B5A9185411CAFF72FE11A82518CE9508" bold="true" box="[399,484,1812,1836]" pageId="144" pageNumber="249">
<taxonomicNameLabel id="AE9AA52F11CAFF72FE11A82518CE9508" box="[399,484,1812,1836]" pageId="144" pageNumber="249" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="CFC797CD11CAFF72FF09A8051B3A95A8" pageId="144" pageNumber="249" type="description">
<paragraph id="8762C44611CAFF72FF09A8051B3A95A8" blockId="144.[151,771,1844,1932]" pageId="144" pageNumber="249">
(
<figureCitation id="1FE6D8C311CAFF72FF01A80519DA9568" box="[159,240,1844,1868]" captionStart="Fig" captionStartId="154.[151,181,1860,1879]" captionTargetBox="[209,1396,191,1822]" captionTargetId="figure-13@154.[193,1435,180,1826]" captionTargetPageId="154" captionText="Fig. 79 Philharmostes apterus sp. nov., paratype, a, pronotum. b, ventral view of rolled up specimen; c, dorsal view of rolled up specimen; d, lateral view of rolled up specimen; Philharmostes fereocellatus sp. nov., holotype, e, dorsal view of distended specimen; f, ventral view of rolled up specimen; g, dorsal view of rolled up specimen; h, lateral view of rolled up specimen." figureDoi="http://doi.org/10.5281/zenodo.12764345" httpUri="https://zenodo.org/record/12764345/files/figure.png" pageId="144" pageNumber="249">Figs 79</figureCitation>
a-d, 85 h, 87 d-e, 89) http://zoobank.org/
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urn:lsid:zoobank.org:act:
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</paragraph>
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<subSubSection id="CFC797CD11CAFF72FCFBAF851C309028" pageId="144" pageNumber="249" type="materials_examined">
<paragraph id="8762C44611CAFF72FCFBAF851C309028" blockId="144.[831,1454,180,1932]" pageId="144" pageNumber="249">
<emphasis id="B5A9185411CAFF72FCFBAF851AC792E8" bold="true" box="[869,1005,180,204]" pageId="144" pageNumber="249">Type series.</emphasis>
<materialsCitation id="37B5CE1B11CAFF72FC68AF851A5E93C8" collectingDate="2001-03-04" collectingDateMax="2001-03-09" collectingDateMin="2001-03-04" collectorName="San Francisco, CA &amp; Fisher &amp; Griswold" country="Madagascar" county="California Acad. of Sciences" determinerName="A. Ballerio" elevation="600" latitude="-13.798612" location="Academy of Sciences" longLatPrecision="21" longitude="48.161667" municipality="Foret d'Ambilanivy" pageId="144" pageNumber="249" specimenCount="1" stateProvince="California" typeStatus="holotype">
<typeStatus id="58667AE411CAFF72FC68AF851D7692E8" box="[1014,1116,180,204]" pageId="144" pageNumber="249" type="holotype">Holotype</typeStatus>
, sex unstated, deposited in the collection of the
<collectingRegion id="45190AA411CAFF72FB98AFE51D5892C8" box="[1030,1138,212,236]" country="United States of America" name="California" pageId="144" pageNumber="249">California</collectingRegion>
<location id="8202929D11CAFF72FBE3AFE51C5992C8" LSID="urn:lsid:plazi:treatment:0F74755011CAFF73FF09A8251B809068:8202929D11CAFF72FBE3AFE51C5992C8" box="[1149,1395,212,236]" country="Madagascar" county="California Acad. of Sciences" latitude="-13.798612" longLatPrecision="21" longitude="48.161667" municipality="Foret d'Ambilanivy" name="Academy of Sciences" pageId="144" pageNumber="249" stateProvince="California">Academy of Sciences</location>
,
<collectorName id="2A28A19011CAFF72FA1DAFE51AC89328" pageId="144" pageNumber="249">San Francisco, CA</collectorName>
, USA, labelled as follows: / CASTYPE 19223 / CASENT 5503872 /
<collectingCountry id="FFCA84D611CAFF72FBE9AE251DD69308" box="[1143,1276,276,300]" name="Madagascar" pageId="144" pageNumber="249">Madagascar</collectingCountry>
:
<location id="8202929D11CAFF72FA98AE251D6B9368" LSID="urn:lsid:plazi:treatment:0F74755011CAFF73FF09A8251B809068:8202929D11CAFF72FA98AE251D6B9368" country="Madagascar" county="California Acad. of Sciences" latitude="-13.798612" longLatPrecision="21" longitude="48.161667" municipality="Foret d'Ambilanivy" name="Province d'Antisiranana Ampasindava" pageId="144" pageNumber="249" stateProvince="California">Province dAntisiranana Ampasindava</location>
,
<collectingMunicipality id="67065E3C11CAFF72FBD5AE051C0A9368" box="[1099,1312,308,332]" pageId="144" pageNumber="249">Forêt dAmbilanivy</collectingMunicipality>
,
<locationDeviation id="96D0DD6411CAFF72FAB7AE051AEA9348" location="Ambaliha" pageId="144" pageNumber="249">
<quantity id="402569A311CAFF72FAB7AE051C599368" box="[1321,1395,308,332]" metricMagnitude="3" metricUnit="m" metricValue="3.9" pageId="144" pageNumber="249" unit="km" value="3.9">3.9 km</quantity>
181° S Ambaliha
</locationDeviation>
elev.
<quantity id="402569A311CAFF72FB99AE651D659348" box="[1031,1103,340,364]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="144" pageNumber="249" unit="m" value="600.0">
<elevation id="0CF0237511CAFF72FB99AE651D659348" box="[1031,1103,340,364]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="144" pageNumber="249" unit="m" value="600.0">600 m</elevation>
</quantity>
<date id="F363E28611CAFF72FBC7AE651C269348" box="[1113,1292,340,364]" pageId="144" pageNumber="249" value="2001-03-04" valueMax="2001-03-09" valueMin="2001-03-04">
<collectingDate id="E3271B6E11CAFF72FBC7AE651C269348" box="[1113,1292,340,364]" pageId="144" pageNumber="249" value="2001-03-04" valueMax="2001-03-09" valueMin="2001-03-04">4-9 March 2001</collectingDate>
</date>
/
<geoCoordinate id="E2E9A28111CAFF72FAB7AE651A6493A8" degrees="13" direction="south" minutes="47" orientation="latitude" pageId="144" pageNumber="249" precision="15" seconds="55" value="-13.798612">13° 47 55” S</geoCoordinate>
<geoCoordinate id="E2E9A28111CAFF72FCC6AE451AD593A8" box="[856,1023,372,396]" degrees="048" direction="east" minutes="09" orientation="longitude" pageId="144" pageNumber="249" precision="15" seconds="42" value="48.161667">048° 09 42” E</geoCoordinate>
coll.
<collectorName id="2A28A19011CAFF72FBDFAE451DAF93A8" box="[1089,1157,372,396]" pageId="144" pageNumber="249">Fisher</collectorName>
,
<collectorName id="2A28A19011CAFF72FB0CAE451DDF93A8" box="[1170,1269,372,396]" pageId="144" pageNumber="249">Griswold</collectorName>
et al.
<collectingCounty id="6E03BCCA11CAFF72FAA1AE451AD79388" pageId="144" pageNumber="249">California Acad. of Sciences</collectingCounty>
sifted litter in rainforest collection code: BLF3252 /
<taxonomicName id="40DDBFC511CAFF72FC5EAE851D9E93E8" authorityName="Ballerio" authorityYear="2021" box="[960,1204,436,460]" class="Insecta" family="Hybosoridae" genus="Philharmostes" kingdom="Animalia" order="Coleoptera" pageId="144" pageNumber="249" phylum="Arthropoda" rank="species" species="apterus" status="sp. nov.">
<emphasis id="B5A9185411CAFF72FC5EAE851D9E93E8" box="[960,1204,436,460]" italics="true" pageId="144" pageNumber="249">Philharmostes apterus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="AE9AA52F11CAFF72FB23AE841DD093E8" box="[1213,1274,437,460]" pageId="144" pageNumber="249" rank="species">n. sp.</taxonomicNameLabel>
det.
<determinerName id="72BE288211CAFF72FAACAE851C8193E8" box="[1330,1451,436,460]" pageId="144" pageNumber="249">A. Ballerio</determinerName>
2020
</materialsCitation>
<typeStatus id="58667AE411CAFF72FCE2AEE51AC193C8" box="[892,1003,468,492]" pageId="144" pageNumber="249" type="holotype">Holotypus</typeStatus>
/.
<materialsCitation id="37B5CE1B11CAFF72FB98AEE41C3D9028" collectingDate="2001-03-04" collectingDateMax="2001-03-09" collectingDateMin="2001-03-04" collectionCode="CASC, ABCB" collectorName="San Francisco, CA &amp; Fisher &amp; Griswold" country="Madagascar" county="California Acad. of Sciences" determinerName="A. Ballerio" elevation="600" latitude="-13.798612" location="Academy of Sciences" longLatPrecision="21" longitude="48.161667" municipality="Foret d'Ambilanivy" pageId="144" pageNumber="249" specimenCount="1" stateProvince="California" typeStatus="paratype">
<typeStatus id="58667AE411CAFF72FB98AEE41D5893C9" box="[1030,1138,469,493]" pageId="144" pageNumber="249" type="paratype">Paratypes</typeStatus>
: 98 exx. (
<quantity id="402569A311CAFF72FB47AEE51C3B93C8" box="[1241,1297,468,492]" metricMagnitude="0" metricUnit="m" metricValue="2.1082" pageId="144" pageNumber="249" unit="in" value="83.0">83 in</quantity>
<collectionCode id="E1CC5C8311CAFF72FA89AEE51C7793C8" box="[1303,1373,468,492]" pageId="144" pageNumber="249">CASC</collectionCode>
and
<quantity id="402569A311CAFF72FA0FAEE51A7E9028" metricMagnitude="-1" metricUnit="m" metricValue="3.81" pageId="144" pageNumber="249" unit="in" value="15.0">15 in</quantity>
<collectionCode id="E1CC5C8311CAFF72FCC7AEC51A8C9028" box="[857,934,500,524]" pageId="144" pageNumber="249">ABCB</collectionCode>
), same collecting data as holotype
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.
</paragraph>
</subSubSection>
<subSubSection id="CFC797CD11CAFF72FCFBAD251D229148" pageId="144" pageNumber="249" type="diagnosis">
<paragraph id="8762C44611CAFF72FCFBAD251D229148" blockId="144.[831,1454,180,1932]" pageId="144" pageNumber="249">
<emphasis id="B5A9185411CAFF72FCFBAD251AF09008" bold="true" box="[869,986,532,556]" pageId="144" pageNumber="249">Diagnosis.</emphasis>
<taxonomicName id="40DDBFC511CAFF72FC7FAD251DF99008" authorityName="Ballerio" authorityYear="2021" box="[993,1235,532,556]" class="Insecta" family="Hybosoridae" genus="Philharmostes" kingdom="Animalia" order="Coleoptera" pageId="144" pageNumber="249" phylum="Arthropoda" rank="species" species="apterus" status="sp. nov.">
<emphasis id="B5A9185411CAFF72FC7FAD251DF99008" box="[993,1235,532,556]" italics="true" pageId="144" pageNumber="249">Philharmostes apterus</emphasis>
</taxonomicName>
<emphasis id="B5A9185411CAFF72FB45AD251C1B9008" bold="true" box="[1243,1329,532,556]" pageId="144" pageNumber="249">
<taxonomicNameLabel id="AE9AA52F11CAFF72FB45AD251C1B9008" box="[1243,1329,532,556]" pageId="144" pageNumber="249" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
can be differentiated from all other species of
<taxonomicName id="40DDBFC511CAFF72FB59AD051C4B9068" authorityName="Kolbe" authorityYear="1895" box="[1223,1377,564,588]" class="Insecta" family="Hybosoridae" genus="Philharmostes" kingdom="Animalia" order="Coleoptera" pageId="144" pageNumber="249" phylum="Arthropoda" rank="genus">
<emphasis id="B5A9185411CAFF72FB59AD051C4B9068" box="[1223,1377,564,588]" italics="true" pageId="144" pageNumber="249">Philharmostes</emphasis>
</taxonomicName>
by the following combination of characters: a) flightlessness, b) dorsal ocular area small, c) head with clypeal disc covered by sparse fine simple punctation, d) dorsum with brown colour, without metallic sheen, e) pronotal fore angles broadly truncate, f) pronotum with disc slightly raised with respect to sides, g) pronotal disc covered by sparse large ocellate punctation, h) elytral lateral carina complete, h) elytral lateral carina blunt, i) elytral punctation formed by ocellate punctures.
</paragraph>
</subSubSection>
<subSubSection id="CFC797CD11CAFF73FCFBAC451BDF9348" lastPageId="145" lastPageNumber="250" pageId="144" pageNumber="249" type="description">
<paragraph id="8762C44611CAFF73FCFBAC451BB69368" blockId="144.[831,1454,180,1932]" lastBlockId="145.[136,758,180,588]" lastPageId="145" lastPageNumber="250" pageId="144" pageNumber="249">
<emphasis id="B5A9185411CAFF72FCFBAC451AD991A8" bold="true" box="[869,1011,884,908]" pageId="144" pageNumber="249">Description.</emphasis>
HL:
<quantity id="402569A311CAFF72FBB5AC451DAA91A8" box="[1067,1152,884,908]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="144" pageNumber="249" unit="mm" value="0.7">0.7 mm</quantity>
; HW: 1.0 mm; PL:
<quantity id="402569A311CAFF72FACCAC451C8D91A8" box="[1362,1447,884,908]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="144" pageNumber="249" unit="mm" value="0.9">0.9 mm</quantity>
; PW:
<quantity id="402569A311CAFF72FCE9ACA51AE49188" box="[887,974,916,940]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" pageId="144" pageNumber="249" unit="mm" value="1.6">1.6 mm</quantity>
; EL:
<quantity id="402569A311CAFF72FB92ACA51D499188" box="[1036,1123,916,940]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" pageId="144" pageNumber="249" unit="mm" value="1.9">1.9 mm</quantity>
; EW 1.8; overall morphology as in generic diagnosis. Small flightless
<taxonomicName id="40DDBFC511CAFF72FB6AAC851C8391E8" authorityName="Martinez" authorityYear="1968" box="[1268,1449,948,972]" class="Insecta" family="Hybosoridae" kingdom="Animalia" order="Coleoptera" pageId="144" pageNumber="109" phylum="Arthropoda" rank="subFamily" subFamily="Ceratocanthinae">Ceratocanthinae</taxonomicName>
. Body convex. Reddish-brown with a slight bronze sheen, shiny, dorsum glabrous (40×), underside, tarsi and antennae reddish-brown.
<emphasis id="B5A9185411CAFF72FBA1AB251D509608" box="[1087,1146,1044,1068]" italics="true" pageId="144" pageNumber="249">Head</emphasis>
: W/L ratio = 1.29; clypeus triangular with angle blunt and obtuse and sides weakly curved; genae produced outwards, acute; genal canthus complete, fused with the occipital portion of head; dorsal ocular area medium-sized, interocular distance about eleven times the maximum width of dorsal ocular area, vertex covered by dense simple impressed punctation, disc uniformly covered by shallower and sparser (nter- punctural distance three to five times longer than punctural diameter) simple punctation, fore margin with some transverse large comma-shaped punctures and a few irregular impressed transverse short lines.
<emphasis id="B5A9185411CAFF72FB65AA441C4D97A8" box="[1275,1383,1397,1420]" italics="true" pageId="144" pageNumber="249">Pronotum</emphasis>
: short and transverse (W/L ratio = 1.84), weakly convex, surface irregular; base and sides broadly rounded (dorsal view); discal area slightly raised compared to sides; fore margin thick and limited to the median portion, not reaching the fore angles, fore angles deeply truncate, whole surface covered by impressed large horseshoe-shaped punctures, with a short opening oriented centripetally. Each puncture bearing in the middle a very short simple seta. Interpunctural distance shorter than punctural diameter.
<taxonomicName id="40DDBFC511CAFF72FAA6A9A51C899488" box="[1336,1443,1684,1708]" class="Trilobita" family="Styginidae" genus="Scutellum" kingdom="Animalia" order="Corynexochida" pageId="144" pageNumber="249" phylum="Arthropoda" rank="genus">
<emphasis id="B5A9185411CAFF72FAA6A9A51C899488" box="[1336,1443,1684,1708]" italics="true" pageId="144" pageNumber="249">Scutellum</emphasis>
</taxonomicName>
: completely covered by several ocellate punctures.
<emphasis id="B5A9185411CAFF72FAC1A9851C8994E8" box="[1375,1443,1716,1740]" italics="true" pageId="144" pageNumber="249">Elytra</emphasis>
: longer than wide (W/L ratio = 0,84); slightly rounded (dorsal view) with maximum width around median third; humeral callus indistinct but marked by the thick starting point of lateral carina; elytral surface convex, covered by dense (interpunctural distance much shorter than punctural diameter) larger horseshoe-shaped puncturs with opening directed backwards mixed to few ocellate punctures. Lateral carina distinct, thick, sharp and complete, broadly S-shaped (lateral view). Pseudoepileura covered by dense large horseshoe-shaped punctures with opening directed towards lateral carina.
<emphasis id="B5A9185411CBFF73FE7FAE041B099368" box="[481,547,309,332]" italics="true" pageId="145" pageNumber="250">Wings</emphasis>
: flightless.
</paragraph>
<paragraph id="8762C44611CBFF73FF30AE651BDF9348" blockId="145.[136,758,180,588]" box="[174,757,340,364]" pageId="145" pageNumber="250">
<emphasis id="B5A9185411CBFF73FF30AE65186A9348" box="[174,320,340,364]" italics="true" pageId="145" pageNumber="250">Male genitalia</emphasis>
: aedeagus with parameres as in
<figureCitation id="1FE6D8C311CBFF73FDE0AE651BE39348" box="[638,713,340,364]" captionStart="Fig" captionStartId="162.[151,182,1885,1904]" captionTargetBox="[214,1388,167,1862]" captionTargetId="figure-13@162.[211,1391,163,1865]" captionTargetPageId="162" captionText="Fig. 87 Aedeagi and spicula gastrales, ac, Philharmostes acutus sp. nov.; de, Philharmostes apterus sp. nov.; fh, Philharmostes nelsoni sp. nov. (paratype from Col de Tanatana)." figureDoi="http://doi.org/10.5281/zenodo.12764365" httpUri="https://zenodo.org/record/12764365/files/figure.png" pageId="145" pageNumber="250">Figs 87</figureCitation>
d-e.
</paragraph>
</subSubSection>
<subSubSection id="CFC797CD11CBFF73FF30AE451B6A93E8" pageId="145" pageNumber="250" type="etymology">
<paragraph id="8762C44611CBFF73FF30AE451B6A93E8" blockId="145.[136,758,180,588]" pageId="145" pageNumber="250">
<emphasis id="B5A9185411CBFF73FF30AE45180493A8" bold="true" box="[174,302,372,396]" pageId="145" pageNumber="250">Etymology.</emphasis>
Latinized Ancient Greek adjective in the nominative singular, meaning “without wings” due to the complete absence of metathoracic wings.
</paragraph>
</subSubSection>
<subSubSection id="CFC797CD11CBFF73FF30AEE51B809068" pageId="145" pageNumber="250" type="distribution">
<paragraph id="8762C44611CBFF73FF30AEE51B809068" blockId="145.[136,758,180,588]" pageId="145" pageNumber="250">
<emphasis id="B5A9185411CBFF73FF30AEE5189493C8" bold="true" box="[174,446,468,492]" pageId="145" pageNumber="250">Distribution and habitat.</emphasis>
Known only from the
<typeStatus id="58667AE411CBFF73FD3BAEE41BF893C9" box="[677,722,469,493]" pageId="145" pageNumber="250">type</typeStatus>
locality in northern
<collectingCountry id="FFCA84D611CBFF73FEA1AEC518949028" box="[319,446,500,524]" name="Madagascar" pageId="145" pageNumber="250">Madagascar</collectingCountry>
. The
<typeStatus id="58667AE411CBFF73FE6BAEC41B089029" box="[501,546,501,525]" pageId="145" pageNumber="250">type</typeStatus>
series was collected by sifting leaf litter in rainforest. For details on the collecting locality see under
<taxonomicName id="40DDBFC511CBFF73FEDDAD051B7A9068" authorityName="Ballerio" authorityYear="2021" box="[323,592,564,588]" class="Insecta" family="Hybosoridae" genus="Goudotostes" kingdom="Animalia" order="Coleoptera" pageId="145" pageNumber="250" phylum="Arthropoda" rank="species" species="electrimontis" status="sp. nov.">
<emphasis id="B5A9185411CBFF73FEDDAD051B7A9068" box="[323,592,564,588]" italics="true" pageId="145" pageNumber="250">Goudotostes electrimontis</emphasis>
</taxonomicName>
<emphasis id="B5A9185411CBFF73FDC9AD051B809068" bold="true" box="[599,682,564,588]" pageId="145" pageNumber="250">
<taxonomicNameLabel id="AE9AA52F11CBFF73FDC9AD051B809068" box="[599,682,564,588]" pageId="145" pageNumber="250" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
</treatment>
</document>