237 lines
28 KiB
XML
237 lines
28 KiB
XML
<document id="5968F60740AF04502D82BC19D63ABD05" ID-DOI="10.13133/2284-4880/545" ID-ISSN="2284-4880" ID-Zenodo-Dep="12764154" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1721242503917" checkinUser="felipe" docAuthor="Ballerio, Alberto" docDate="2021" docId="0F747550116DFFDAFCAEAD2418199788" docLanguage="en" docName="FragEntomol.53.2.105-282.pdf" docOrigin="Fragmenta entomologica 53 (2)" docSource="http://dx.doi.org/10.13133/2284-4880/545" docStyle="DocumentStyle:8499FF4308F512386C0E43DC290A16BD.1:FragEntomol.2021-.journal_article.0cover" docStyleId="8499FF4308F512386C0E43DC290A16BD" docStyleName="FragEntomol.2021-.journal_article.0cover" docStyleVersion="1" docTitle="Synarmostes monoculus Ballerio, 2021, sp. nov." docType="treatment" docUuid="0F5561B7-1A77-4314-9C16-EA6A8F0551EF" docUuidSource="ZooBank" docVersion="2" lastPageNumber="161" masterDocId="F34D0D28115AFFE2FF9EAF31192A9224" masterDocTitle="The Ceratocanthinae of Madagascar and Comoro Islands: a revision of the genera Synarmostes and Goudotostes, and of the flightless Philharmostes, with description of 64 new species (Coleoptera: Scarabaeoidea, Hybosoridae)" masterLastPageNumber="282" masterPageNumber="105" pageNumber="160" updateTime="1721247602959" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-4.0">
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<mods:title id="8A81C2309AEC6E531252C9CFE01EF077">The Ceratocanthinae of Madagascar and Comoro Islands: a revision of the genera Synarmostes and Goudotostes, and of the flightless Philharmostes, with description of 64 new species (Coleoptera: Scarabaeoidea, Hybosoridae)</mods:title>
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<mods:roleTerm id="B83ECA5463D449D5A74411EB3B3F91CD">Author</mods:roleTerm>
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<mods:namePart id="A36BFBCCACA3EDAABEFFE73ECE5CB7D9">Ballerio, Alberto</mods:namePart>
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<mods:typeOfResource id="CF36351A368D439BD4F0612238E2DB62">text</mods:typeOfResource>
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<mods:title id="0BB64FA7F7011ABC757830BB34F31463">Fragmenta entomologica</mods:title>
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</mods:titleInfo>
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<mods:part id="1021E85BC26E27424623B2803C5825C5">
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<mods:date id="A54033435A42E97899EF85E360C3D444">2021</mods:date>
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<mods:number id="15D9F2FB82A56D55C2C0F9FFD73D6928">2021-11-30</mods:number>
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<mods:number id="BD67D15B004A53B9F68F5BC188B12C61">53</mods:number>
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<mods:start id="1E2B70484FEE68E93CF9E96CDA080D05">105</mods:start>
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<mods:url id="E31C7F404D5FC1A4AC661474A3512DE3">http://dx.doi.org/10.13133/2284-4880/545</mods:url>
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<mods:classification id="93FA666FC1C6C19D5604A00CBF6C10D6">journal article</mods:classification>
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<mods:identifier id="4445C4734DE852ED109EDECD708D20E7" type="DOI">10.13133/2284-4880/545</mods:identifier>
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<mods:identifier id="7C5A07F0595391A13282DB9E30D36CF3" type="ISSN">2284-4880</mods:identifier>
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<mods:identifier id="4D1039D39DCB0935A54DFEC04E68945E" type="ZooBank">B3F75A2C-7603-4027-B8E1-F4A8B5A0CDD9</mods:identifier>
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<treatment id="0F747550116DFFDAFCAEAD2418199788" LSID="urn:lsid:zoobank.org:act:0F5561B7-1A77-4314-9C16-EA6A8F0551EF" httpUri="http://treatment.plazi.org/id/0F747550116DFFDAFCAEAD2418199788" lastPageId="56" lastPageNumber="161" pageId="55" pageNumber="160">
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<subSubSection id="CFC797CD116DFFD5FCAEAD241DA49008" box="[816,1166,532,556]" pageId="55" pageNumber="160" type="nomenclature">
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<paragraph id="8762C446116DFFD5FCAEAD241DA49008" blockId="55.[816,1166,532,556]" box="[816,1166,532,556]" pageId="55" pageNumber="160">
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<heading id="DC2A732A116DFFD5FCAEAD241DA49008" box="[816,1166,532,556]" fontSize="10" level="2" pageId="55" pageNumber="160" reason="5">
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<taxonomicName id="40DDBFC5116DFFD5FCAEAD241D189008" ID-CoL="8SSSC" authorityName="Ballerio" authorityYear="2021" box="[816,1074,532,556]" class="Insecta" family="Hybosoridae" genus="Synarmostes" kingdom="Animalia" order="Coleoptera" pageId="55" pageNumber="160" phylum="Arthropoda" rank="species" species="monoculus" status="sp. nov.">
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<emphasis id="B5A91854116DFFD5FCAEAD241D189008" box="[816,1074,532,556]" italics="true" pageId="55" pageNumber="160">Synarmostes monoculus</emphasis>
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</taxonomicName>
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||
<emphasis id="B5A91854116DFFD5FBA6AD251DA49008" bold="true" box="[1080,1166,532,556]" pageId="55" pageNumber="160">
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<taxonomicNameLabel id="AE9AA52F116DFFD5FBA6AD251DA49008" box="[1080,1166,532,556]" pageId="55" pageNumber="160" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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||
</heading>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="CFC797CD116DFFD5FCAEAD051D8190A8" pageId="55" pageNumber="160" type="description">
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<paragraph id="8762C446116DFFD5FCAEAD051D8190A8" blockId="55.[816,1435,564,652]" pageId="55" pageNumber="160">
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(
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<figureCitation id="1FE6D8C3116DFFD5FCA6AD051AA39068" box="[824,905,564,588]" captionStart="Fig" captionStartId="30.[151,182,1858,1877]" captionTargetBox="[155,1445,178,1828]" captionTargetId="figure-13@30.[148,1454,912,1832]" captionTargetPageId="30" captionText="Fig. 18 – Synarmostes caterinae sp. nov., a, dorsal view of distended holotype; b, ventral view of rolled up paratype; c, dorsal view of rolled up paratype; d, lateral view of rolled up paratype; Synarmostes monoculus sp. nov., e, dorsal view of distended holotype; f, ventral view of rolled up paratype; g, dorsal view of rolled up paratype; h, lateral view of rolled up paratype." figureDoi="http://doi.org/10.5281/zenodo.12764205" httpUri="https://zenodo.org/record/12764205/files/figure.png" pageId="55" pageNumber="160">Figs 18</figureCitation>
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e-h, 31 m-o, 35) http://zoobank.org/
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<uri id="F34CC844116DFFD5FB93AD651D8190A8" pageId="55" pageNumber="160">
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||
urn:lsid:zoobank.org:act:
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||
<uuid id="F37BFE93116DFFD5FA81AD651D8190A8" pageId="55" pageNumber="160">0F5561B7- 1A77-4314-9C16-EA6A8F0551EF</uuid>
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</uri>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="CFC797CD116DFFD5FCC8AD851ACF9608" pageId="55" pageNumber="160" type="materials_examined">
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<paragraph id="8762C446116DFFD5FCC8AD851ACF9608" blockId="55.[816,1437,692,1932]" pageId="55" pageNumber="160">
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<emphasis id="B5A91854116DFFD5FCC8AD851AF690E8" bold="true" box="[854,988,692,716]" pageId="55" pageNumber="160">Type series.</emphasis>
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<materialsCitation id="37B5CE1B116DFFD5FC7BAD851C7D9188" collectingDate="2009-10-24" collectingDateMax="2009-10-27" collectingDateMin="2009-10-24" collectorName="San Francisco, CA & B. L. Fisher" country="Madagascar" county="California Acad. of Sciences" determinerName="A. Ballerio" elevation="300" latitude="-18.062222" location="Academy of Sciences" longLatPrecision="21" longitude="44.525833" municipality="Km" pageId="55" pageNumber="160" specimenCount="1" specimenCount-male="1" stateProvince="California" typeStatus="holotype">
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<typeStatus id="58667AE4116DFFD5FC7BAD851D6190E8" box="[997,1099,692,716]" pageId="55" pageNumber="160" type="holotype">Holotype</typeStatus>
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,
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<specimenCount id="91DB0FCF116DFFD5FBC6AD841D4490E8" box="[1112,1134,693,716]" count="1" pageId="55" pageNumber="160" type="male">♂</specimenCount>
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, deposited in the collection of the
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<collectingRegion id="45190AA4116DFFD5FCE1ADE51AC190C8" box="[895,1003,724,748]" country="United States of America" name="California" pageId="55" pageNumber="160">California</collectingRegion>
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<location id="8202929D116DFFD5FC68ADE51DC090C8" LSID="urn:lsid:plazi:treatment:0F747550116DFFDAFCAEAD2418199788:8202929D116DFFD5FC68ADE51DC090C8" box="[1014,1258,724,748]" country="Madagascar" county="California Acad. of Sciences" latitude="-18.062222" longLatPrecision="21" longitude="44.525833" municipality="Km" name="Academy of Sciences" pageId="55" pageNumber="160" stateProvince="California">Academy of Sciences</location>
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,
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<collectorName id="2A28A190116DFFD5FB64ADE51A729128" pageId="55" pageNumber="160">San Francisco, CA</collectorName>
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, USA, labelled as follows: / CASTYPE 20257 /
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<collectingCountry id="FFCA84D6116DFFD5FAFCADC51AAE9109" name="Madagascar" pageId="55" pageNumber="160">Madagascar</collectingCountry>
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:
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<location id="8202929D116DFFD5FC13AC251D629108" LSID="urn:lsid:plazi:treatment:0F747550116DFFDAFCAEAD2418199788:8202929D116DFFD5FC13AC251D629108" box="[909,1096,788,812]" country="Madagascar" county="California Acad. of Sciences" latitude="-18.062222" longLatPrecision="21" longitude="44.525833" municipality="Km" name="Majunga Reserve" pageId="55" pageNumber="160" stateProvince="California">Majunga Réserve</location>
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forestière
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<location id="8202929D116DFFD5FB26AC251C229108" LSID="urn:lsid:plazi:treatment:0F747550116DFFDAFCAEAD2418199788:8202929D116DFFD5FB26AC251C229108" box="[1208,1288,788,812]" country="Madagascar" county="California Acad. of Sciences" latitude="-18.062222" longLatPrecision="21" longitude="44.525833" municipality="Km" name="Beanka" pageId="55" pageNumber="160" stateProvince="California">Beanka</location>
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52.7
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<collectingMunicipality id="67065E3C116DFFD5FADCAC241C409108" box="[1346,1386,789,812]" pageId="55" pageNumber="160">Km</collectingMunicipality>
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<locationDeviation id="96D0DD64116DFFD5FAEEAC241A8D9168" location="Maintirano" pageId="55" pageNumber="160">E of Maintirano</locationDeviation>
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el.
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<quantity id="402569A3116DFFD5FC50AC051D3B9168" box="[974,1041,820,844]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="55" pageNumber="160" unit="m" value="300.0">
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||
<elevation id="0CF02375116DFFD5FC50AC051D3B9168" box="[974,1041,820,844]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="55" pageNumber="160" unit="m" value="300.0">300 m</elevation>
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</quantity>
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<date id="F363E286116DFFD5FB86AC051DEF9168" box="[1048,1221,820,844]" pageId="55" pageNumber="160" value="2009-10-24" valueMax="2009-10-27" valueMin="2009-10-24">
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<collectingDate id="E3271B6E116DFFD5FB86AC051DEF9168" box="[1048,1221,820,844]" pageId="55" pageNumber="160" value="2009-10-24" valueMax="2009-10-27" valueMin="2009-10-24">24-27 Oct. 2009</collectingDate>
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||
</date>
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/
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||
<geoCoordinate id="E2E9A281116DFFD5FB4BAC051C499168" box="[1237,1379,820,844]" degrees="18" direction="south" minutes="03" orientation="latitude" pageId="55" pageNumber="160" precision="15" seconds="44" value="-18.062222">18° 03’ 44” S</geoCoordinate>
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||
<geoCoordinate id="E2E9A281116DFFD5FAF4AC051AB09148" degrees="044" direction="east" minutes="31" orientation="longitude" pageId="55" pageNumber="160" precision="15" seconds="33" value="44.525833">044° 31’ 33” E</geoCoordinate>
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<collectingCounty id="6E03BCCA116DFFD5FC3CAC651DF39148" box="[930,1241,852,876]" pageId="55" pageNumber="160">California Acad. of Sciences</collectingCounty>
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coll.
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<collectorName id="2A28A190116DFFD5FA87AC641CB79148" box="[1305,1437,852,876]" pageId="55" pageNumber="160">B. L. Fisher</collectorName>
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et al. tropical dry forest on tsingy sifted litter BLF22925/
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<taxonomicName id="40DDBFC5116DFFD5FCAEACA41D189188" authorityName="Ballerio" authorityYear="2021" box="[816,1074,916,940]" class="Insecta" family="Hybosoridae" genus="Synarmostes" kingdom="Animalia" order="Coleoptera" pageId="55" pageNumber="160" phylum="Arthropoda" rank="species" species="monoculus" status="sp. nov.">
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<emphasis id="B5A91854116DFFD5FCAEACA41D189188" box="[816,1074,916,940]" italics="true" pageId="55" pageNumber="160">Synarmostes monoculus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="AE9AA52F116DFFD5FBA6ACA41D589188" box="[1080,1138,917,940]" pageId="55" pageNumber="160" rank="species">n. sp.</taxonomicNameLabel>
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det.
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<determinerName id="72BE2882116DFFD5FB3BACA51C319188" box="[1189,1307,916,940]" pageId="55" pageNumber="160">A. Ballerio</determinerName>
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2020
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</materialsCitation>
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<typeStatus id="58667AE4116DFFD5FAC0ACA51A4091E9" pageId="55" pageNumber="160" type="holotype">Holotypus</typeStatus>
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<specimenCount id="91DB0FCF116DFFD5FCECAC841AAC91E8" box="[882,902,949,972]" count="1" pageId="55" pageNumber="160" type="male">♂</specimenCount>
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/ [distended, glued on card, dissected, male genitalia glued in DMHF resin on a separate card, same pin].
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<materialsCitation id="37B5CE1B116DFFD5FCAFACC41ACB9608" collectingDate="2009-10-24" collectingDateMax="2009-10-27" collectingDateMin="2009-10-24" collectionCode="CASC" collectorName="San Francisco, CA & B. L. Fisher" country="Madagascar" county="California Acad. of Sciences" determinerName="A. Ballerio" elevation="300" latitude="-18.062222" location="Academy of Sciences" longLatPrecision="21" longitude="44.525833" municipality="Km" pageId="55" pageNumber="160" specimenCount="2" specimenCount-male="2" stateProvince="California" typeStatus="paratype">
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<typeStatus id="58667AE4116DFFD5FCAFACC41AB39629" box="[817,921,1013,1037]" pageId="55" pageNumber="160" type="paratype">Paratypes</typeStatus>
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[
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<specimenCount id="91DB0FCF116DFFD5FC35ACC51AFF9628" box="[939,981,1012,1036]" count="1" pageId="55" pageNumber="160" type="male">1 ♂</specimenCount>
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dissected]:
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<specimenCount id="91DB0FCF116DFFD5FBC7ACC51DAE9628" box="[1113,1156,1012,1036]" count="1" pageId="55" pageNumber="160" type="male">1 ♂</specimenCount>
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||
(
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<collectionCode id="E1CC5C83116DFFD5FB0AACC51DF59628" box="[1172,1247,1012,1036]" pageId="55" pageNumber="160">CASC</collectionCode>
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), same collecting data as holotype
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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||
<subSubSection id="CFC797CD116DFFD5FCC8AB051DCA9488" pageId="55" pageNumber="160" type="diagnosis">
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<paragraph id="8762C446116DFFD5FCC8AB051DCA9488" blockId="55.[816,1437,692,1932]" pageId="55" pageNumber="160">
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<emphasis id="B5A91854116DFFD5FCC8AB051AE69668" bold="true" box="[854,972,1076,1100]" pageId="55" pageNumber="160">Diagnosis.</emphasis>
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<taxonomicName id="40DDBFC5116DFFD5FC49AB041DCA9668" authorityName="Ballerio" authorityYear="2021" box="[983,1248,1076,1100]" class="Insecta" family="Hybosoridae" genus="Synarmostes" kingdom="Animalia" order="Coleoptera" pageId="55" pageNumber="160" phylum="Arthropoda" rank="species" species="monoculus" status="sp. nov.">
|
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<emphasis id="B5A91854116DFFD5FC49AB041DCA9668" box="[983,1248,1076,1100]" italics="true" pageId="55" pageNumber="160">Synarmostes monoculus</emphasis>
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</taxonomicName>
|
||
<emphasis id="B5A91854116DFFD5FB75AB051C6D9668" bold="true" box="[1259,1351,1076,1100]" pageId="55" pageNumber="160">
|
||
<taxonomicNameLabel id="AE9AA52F116DFFD5FB75AB051C6D9668" box="[1259,1351,1076,1100]" pageId="55" pageNumber="160" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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can be differentiated from all other species of
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<taxonomicName id="40DDBFC5116DFFD5FB70AB641C5C9648" authorityName="Germar" authorityYear="1843" box="[1262,1398,1109,1132]" class="Insecta" family="Hybosoridae" genus="Synarmostes" kingdom="Animalia" order="Coleoptera" pageId="55" pageNumber="160" phylum="Arthropoda" rank="genus">
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||
<emphasis id="B5A91854116DFFD5FB70AB641C5C9648" box="[1262,1398,1109,1132]" italics="true" pageId="55" pageNumber="160">Synarmostes</emphasis>
|
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</taxonomicName>
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by the following combination of characters: a) small size, b) ventral ocular area small-sized, c) dorsal ocular area vestigial, d) dorsal setation absent (20×), e) flightlessness, f) carinae of elytral apical third regular (without tubercles or other irregularities), g) scutellum with transverse comma-shaped punctation, h) pronotal punctation consisting of transverse comma-shaped punctures, i) pronotum without an anteromedian area smooth or with very sparse punctation, j) head with dense and impressed simple punctation, k) elytra with small transverse sparse comma-shaped punctures mixed to some horseshoe-shaped punctures.
|
||
<taxonomicName id="40DDBFC5116DFFD5FC32AAE41D9F97C8" authorityName="Ballerio" authorityYear="2021" box="[940,1205,1492,1516]" class="Insecta" family="Hybosoridae" genus="Synarmostes" kingdom="Animalia" order="Coleoptera" pageId="55" pageNumber="160" phylum="Arthropoda" rank="species" species="monoculus" status="sp. nov.">
|
||
<emphasis id="B5A91854116DFFD5FC32AAE41D9F97C8" box="[940,1205,1492,1516]" italics="true" pageId="55" pageNumber="160">Synarmostes monoculus</emphasis>
|
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</taxonomicName>
|
||
<emphasis id="B5A91854116DFFD5FB5FAAE51C3697C8" bold="true" box="[1217,1308,1492,1516]" pageId="55" pageNumber="160">
|
||
<taxonomicNameLabel id="AE9AA52F116DFFD5FB5FAAE51C3697C8" box="[1217,1308,1492,1516]" pageId="55" pageNumber="160" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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belongs to the
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<taxonomicName id="40DDBFC5116DFFD5FCC2AAC41A959428" box="[860,959,1525,1548]" class="Insecta" family="Hybosoridae" genus="Synarmostes" kingdom="Animalia" order="Coleoptera" pageId="55" pageNumber="160" phylum="Arthropoda" rank="species" species="antsingyi">
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<emphasis id="B5A91854116DFFD5FCC2AAC41A959428" box="[860,959,1525,1548]" italics="true" pageId="55" pageNumber="160">antsingyi</emphasis>
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</taxonomicName>
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group of species and is very similar to
|
||
<taxonomicName id="40DDBFC5116DFFD5FA16AAC41AA29408" authorityName="Ballerio" authorityYear="2021" class="Insecta" family="Hybosoridae" genus="Synarmostes" kingdom="Animalia" order="Coleoptera" pageId="55" pageNumber="160" phylum="Arthropoda" rank="species" species="antingyi" status="sp. nov.">
|
||
<emphasis id="B5A91854116DFFD5FA16AAC41AA29408" italics="true" pageId="55" pageNumber="160">S. antingyi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B5A91854116DFFD5FC0CA9251AC19408" bold="true" box="[914,1003,1556,1580]" pageId="55" pageNumber="160">
|
||
<taxonomicNameLabel id="AE9AA52F116DFFD5FC0CA9251AC19408" box="[914,1003,1556,1580]" pageId="55" pageNumber="160" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, from which differs mainly because of the presence of uniform punctation on pronotal disc and the vestigial dorsal ocular area and to
|
||
<taxonomicName id="40DDBFC5116DFFD5FB5BA9641C109448" authorityName="Ballerio" authorityYear="2021" box="[1221,1338,1620,1644]" class="Insecta" family="Hybosoridae" genus="Synarmostes" kingdom="Animalia" order="Coleoptera" pageId="55" pageNumber="160" phylum="Arthropoda" rank="species" species="beankae" status="sp. nov.">
|
||
<emphasis id="B5A91854116DFFD5FB5BA9641C109448" box="[1221,1338,1620,1644]" italics="true" pageId="55" pageNumber="160">S. beankae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B5A91854116DFFD5FAA1A9651CBF9448" bold="true" box="[1343,1429,1620,1644]" pageId="55" pageNumber="160">
|
||
<taxonomicNameLabel id="AE9AA52F116DFFD5FAA1A9651CBF9448" box="[1343,1429,1620,1644]" pageId="55" pageNumber="160" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, from which differs because of the smaller elytral punctation and the vestigial dorsal ocular area.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="CFC797CD116DFFDAFCC8A98518D79688" lastPageId="56" lastPageNumber="161" pageId="55" pageNumber="160" type="description">
|
||
<paragraph id="8762C446116DFFDAFCC8A985186C90E8" blockId="55.[816,1437,692,1932]" lastBlockId="56.[151,772,180,1452]" lastPageId="56" lastPageNumber="161" pageId="55" pageNumber="160">
|
||
<emphasis id="B5A91854116DFFD5FCC8A9851ACA94E8" bold="true" box="[854,992,1716,1740]" pageId="55" pageNumber="160">Description.</emphasis>
|
||
Size: HL =
|
||
<quantity id="402569A3116DFFD5FBFFA9851D9C94E8" box="[1121,1206,1716,1740]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="55" pageNumber="160" unit="mm" value="0.6">0.6 mm</quantity>
|
||
; HW =
|
||
<quantity id="402569A3116DFFD5FA97A9851C7494E8" box="[1289,1374,1716,1740]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="55" pageNumber="160" unit="mm" value="1.1">1.1 mm</quantity>
|
||
; PL = 1.0 mm; PW =
|
||
<quantity id="402569A3116DFFD5FC50A9E51D0894C8" box="[974,1058,1748,1772]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="55" pageNumber="160" unit="mm" value="1.8">1.8 mm</quantity>
|
||
; EL =
|
||
<quantity id="402569A3116DFFD5FBFAA9E51D9D94C8" box="[1124,1207,1748,1772]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" pageId="55" pageNumber="160" unit="mm" value="1.9">1.9 mm</quantity>
|
||
; EW =
|
||
<quantity id="402569A3116DFFD5FA9DA9E51C7D94C8" box="[1283,1367,1748,1772]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="55" pageNumber="160" unit="mm" value="1.8">1.8 mm</quantity>
|
||
. Overall morphology as in generic diagnosis. Small-sized flightless
|
||
<taxonomicName id="40DDBFC5116DFFD5FCFFA8241ACC9508" authorityName="Germar" authorityYear="1843" box="[865,998,1813,1836]" class="Insecta" family="Hybosoridae" genus="Synarmostes" kingdom="Animalia" order="Coleoptera" pageId="55" pageNumber="160" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B5A91854116DFFD5FCFFA8241ACC9508" box="[865,998,1813,1836]" italics="true" pageId="55" pageNumber="160">Synarmostes</emphasis>
|
||
</taxonomicName>
|
||
. Body convex. Dark-brown with bronze sheen, shiny, dorsum glabrous (20×), underside, tarsi and antennae reddish-brown, setation yellowish-brown.
|
||
<emphasis id="B5A91854116DFFD5FAC5A8651CBE9548" box="[1371,1428,1876,1900]" italics="true" pageId="55" pageNumber="160">Head</emphasis>
|
||
: W/L ratio = 1.40, clypeal apex obtuse, apical projection small but distinctly protruding and sharp; dorsal ocular area absent (probably vestigial: a hint of eye visible in transap- rence only), inferior ocular area small; head dorsal surface, except from smooth vertex, uniformly covered by large impressed dense simple punctures (interpunctural distance about twice to thrice the diameter of each puncture), a distinctive deep curved line is present just below the gena on the spot where the dorsal ocular area is normally present. Fore margin with one or two irregular rows of long transverse lines. Vertex smooth with one continuous transverse impressed line.
|
||
<emphasis id="B5A918541162FFDAFED5AEC4189E9028" box="[331,436,501,524]" italics="true" pageId="56" pageNumber="161">Pronotum</emphasis>
|
||
: subrectangular (W/L ratio = 1.88), regularly convex, pronotal surface almost uniformly punctured, only sides and base almost impunctate; punctures short transverse comma-shaped, each one bearing a simple puncture internally, curvature centripetally orient- ed. Interpunctural distance once to three times longer than punctural width.
|
||
</paragraph>
|
||
<paragraph id="8762C4461162FFDAFF23ADE518D79688" blockId="56.[151,772,180,1452]" pageId="56" pageNumber="161">
|
||
<taxonomicName id="40DDBFC51162FFDAFF23ADE5180F90C8" box="[189,293,724,748]" class="Trilobita" family="Styginidae" genus="Scutellum" kingdom="Animalia" order="Corynexochida" pageId="56" pageNumber="161" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B5A918541162FFDAFF23ADE5180F90C8" box="[189,293,724,748]" italics="true" pageId="56" pageNumber="161">Scutellum</emphasis>
|
||
</taxonomicName>
|
||
: with some transverse comma-shaped punctures.
|
||
<emphasis id="B5A918541162FFDAFF42ADC518349128" box="[220,286,756,780]" italics="true" pageId="56" pageNumber="161">Elytra</emphasis>
|
||
: longer than wide (W/L ratio = 0.90); subovoidal (dorsal view); inferior sutural stria reaching humeral area although under the form of a very shallow fine line; humeral callus indistinct; elytral surface regularly convex, uniformly covered by sparse punctation. Interpunctural distance once or twice the maximum width of punctures. Punctures comma-shaped becoming horseshoe-shaped on apical third, small and transverse, each one having a simple setigerous puncture internally. Elytral distal third with eight raised longitudinal carinae (excluding elytral suture), sometimes having a longitudinal shallow line among them.
|
||
<emphasis id="B5A918541162FFDAFF09AB6419F29648" box="[151,216,1109,1132]" italics="true" pageId="56" pageNumber="161">Wings</emphasis>
|
||
: flightless (apterous).
|
||
<emphasis id="B5A918541162FFDAFE5AAB651B759648" box="[452,607,1108,1132]" italics="true" pageId="56" pageNumber="161">Male genitalia</emphasis>
|
||
: aedeagus (L =
|
||
<quantity id="402569A31162FFDAFF06AB4519C196A8" box="[152,235,1140,1164]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="56" pageNumber="161" unit="mm" value="0.8">0.8 mm</quantity>
|
||
) with parameres as in
|
||
<figureCitation id="1FE6D8C31162FFDAFE41AB451B0596A8" box="[479,559,1140,1164]" captionStart="Fig" captionStartId="43.[136,167,1885,1904]" captionTargetBox="[136,1450,199,1856]" captionTargetId="figure-4@43.[124,1461,194,1869]" captionTargetPageId="43" captionText="Fig. 31 – Aedeagi and spicula gastrales, a–c, Synarmostes antsingyi (Tsingy de Bemaraha); d–f, Synarmostes beankae sp. nov., holotype; g–i, Synarmostes bekaraokae sp. nov., holotype; j–l, Synarmostes caterinae sp. nov., holotype; m–o, Synarmostes monoculus sp. nov., holotype." figureDoi="http://doi.org/10.5281/zenodo.12764235" httpUri="https://zenodo.org/record/12764235/files/figure.png" pageId="56" pageNumber="161">Figs 31</figureCitation>
|
||
m-n, spiculum gastrale (L =
|
||
<quantity id="402569A31162FFDAFE9EABA518799688" box="[256,339,1172,1196]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="56" pageNumber="161" unit="mm" value="0.8">0.8 mm</quantity>
|
||
) as in
|
||
<figureCitation id="1FE6D8C31162FFDAFE06ABA518D39688" box="[408,505,1172,1196]" captionStart="Fig" captionStartId="43.[136,167,1885,1904]" captionTargetBox="[136,1450,199,1856]" captionTargetId="figure-4@43.[124,1461,194,1869]" captionTargetPageId="43" captionText="Fig. 31 – Aedeagi and spicula gastrales, a–c, Synarmostes antsingyi (Tsingy de Bemaraha); d–f, Synarmostes beankae sp. nov., holotype; g–i, Synarmostes bekaraokae sp. nov., holotype; j–l, Synarmostes caterinae sp. nov., holotype; m–o, Synarmostes monoculus sp. nov., holotype." figureDoi="http://doi.org/10.5281/zenodo.12764235" httpUri="https://zenodo.org/record/12764235/files/figure.png" pageId="56" pageNumber="161">Fig. 31 o</figureCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="CFC797CD1162FFDAFF23AB8518769728" pageId="56" pageNumber="161" type="etymology">
|
||
<paragraph id="8762C4461162FFDAFF23AB8518769728" blockId="56.[151,772,180,1452]" pageId="56" pageNumber="161">
|
||
<emphasis id="B5A918541162FFDAFF23AB85181496E8" bold="true" box="[189,318,1204,1228]" pageId="56" pageNumber="161">Etymology.</emphasis>
|
||
Latin adjective in the nominative singular meaning “with a single eye”, due to the absence of the dorsal ocular area.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="CFC797CD1162FFDAFF23AA2518199788" pageId="56" pageNumber="161" type="distribution">
|
||
<paragraph id="8762C4461162FFDAFF23AA2518199788" blockId="56.[151,772,180,1452]" pageId="56" pageNumber="161">
|
||
<emphasis id="B5A918541162FFDAFF23AA2518F69708" bold="true" box="[189,476,1300,1324]" pageId="56" pageNumber="161">Distribution and habitat.</emphasis>
|
||
Known only from the
|
||
<typeStatus id="58667AE41162FFDAFD4BAA241A299709" box="[725,771,1301,1325]" pageId="56" pageNumber="161">type</typeStatus>
|
||
locality in western
|
||
<collectingCountry id="FFCA84D61162FFDAFEF4AA0518C79768" box="[362,493,1332,1356]" name="Madagascar" pageId="56" pageNumber="161">Madagascar</collectingCountry>
|
||
. The
|
||
<typeStatus id="58667AE41162FFDAFDB5AA041B739769" box="[555,601,1333,1357]" pageId="56" pageNumber="161">type</typeStatus>
|
||
series was collected by sifting litter (leaf mold, rotten wood) on a Tsingy in tropical dry forest. See under
|
||
<taxonomicName id="40DDBFC51162FFDAFD9CAA441B5297A8" authorityName="Ballerio" authorityYear="2021" box="[514,632,1396,1420]" class="Insecta" family="Hybosoridae" genus="Synarmostes" kingdom="Animalia" order="Coleoptera" pageId="56" pageNumber="161" phylum="Arthropoda" rank="species" species="beankae" status="sp. nov.">
|
||
<emphasis id="B5A918541162FFDAFD9CAA441B5297A8" box="[514,632,1396,1420]" italics="true" pageId="56" pageNumber="161">S. beankae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B5A918541162FFDAFD1CAA451BF197A8" bold="true" box="[642,731,1396,1420]" pageId="56" pageNumber="161">
|
||
<taxonomicNameLabel id="AE9AA52F1162FFDAFD1CAA451BF197A8" box="[642,731,1396,1420]" pageId="56" pageNumber="161" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
for further details.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |