treatments-xml/data/0F/74/75/0F7475501169FFD1FF16AE041ACF9188.xml

265 lines
30 KiB
XML
Raw Blame History

This file contains ambiguous Unicode characters

This file contains Unicode characters that might be confused with other characters. If you think that this is intentional, you can safely ignore this warning. Use the Escape button to reveal them.

<document id="E340F5B8263E516F3F6289823753FA8E" ID-DOI="10.13133/2284-4880/545" ID-ISSN="2284-4880" ID-Zenodo-Dep="12764154" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1721242503917" checkinUser="felipe" docAuthor="Ballerio, Alberto" docDate="2021" docId="0F7475501169FFD1FF16AE041ACF9188" docLanguage="en" docName="FragEntomol.53.2.105-282.pdf" docOrigin="Fragmenta entomologica 53 (2)" docSource="http://dx.doi.org/10.13133/2284-4880/545" docStyle="DocumentStyle:8499FF4308F512386C0E43DC290A16BD.1:FragEntomol.2021-.journal_article.0cover" docStyleId="8499FF4308F512386C0E43DC290A16BD" docStyleName="FragEntomol.2021-.journal_article.0cover" docStyleVersion="1" docTitle="Synarmostes fortis Ballerio, 2021, sp. nov." docType="treatment" docUuid="5C0B54B8-92E7-4CAA-B199-D5B2D46D83C2" docUuidSource="ZooBank" docVersion="3" lastPageNumber="156" masterDocId="F34D0D28115AFFE2FF9EAF31192A9224" masterDocTitle="The Ceratocanthinae of Madagascar and Comoro Islands: a revision of the genera Synarmostes and Goudotostes, and of the flightless Philharmostes, with description of 64 new species (Coleoptera: Scarabaeoidea, Hybosoridae)" masterLastPageNumber="282" masterPageNumber="105" pageNumber="156" updateTime="1721247602959" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-4.0">
<mods:mods id="0C4DBEC26E0D46EEB7B31C92245F6BA5" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="187C355E2E92699B5A92AD37AB7099CD">
<mods:title id="4331615B16F37FA690997A47F9B3B8D2">The Ceratocanthinae of Madagascar and Comoro Islands: a revision of the genera Synarmostes and Goudotostes, and of the flightless Philharmostes, with description of 64 new species (Coleoptera: Scarabaeoidea, Hybosoridae)</mods:title>
</mods:titleInfo>
<mods:name id="C631E58923A1A4FFDE621559ECCA8955" type="personal">
<mods:role id="B3CDFD91C0789353658A3D6F874F9CBC">
<mods:roleTerm id="C07D88A5BFD11DD5706F91FEA2E73748">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="A8F6AB473F0601423E65CAE1318064BD">Ballerio, Alberto</mods:namePart>
</mods:name>
<mods:typeOfResource id="1459CC3785EF2D9708393EE5404BF3C7">text</mods:typeOfResource>
<mods:relatedItem id="646375AF6AF51657AAF85BBDEF51A77E" type="host">
<mods:titleInfo id="7FBDD17689C0A55F653A737A589FB1B9">
<mods:title id="44C70B7AA6A12395997EECE8F26C741B">Fragmenta entomologica</mods:title>
</mods:titleInfo>
<mods:part id="FD90E3EDED3AC9F8964F755C4FCE9903">
<mods:date id="DFCB1F215505B4E6FB384BDD43B7D0BF">2021</mods:date>
<mods:detail id="5DF24EA25AC0A2946FC91EB9526FE2C1" type="pubDate">
<mods:number id="D0EDA43969D83D0F082D3DDA0DBD4DA9">2021-11-30</mods:number>
</mods:detail>
<mods:detail id="CBEB1CE2C55FAF41E11A1586706A6463" type="volume">
<mods:number id="3E4256E67C2BEBD9330EFE538DDCB1AF">53</mods:number>
</mods:detail>
<mods:detail id="6185D395F8165C095B317079F2E86ECE" type="issue">
<mods:number id="54223A5B0495A3731E4291C6E52D0531">2</mods:number>
</mods:detail>
<mods:extent id="2B41702ABD17C5CEB870E6EB467FE663" unit="page">
<mods:start id="38ABFB6FAFD64201230AA9DD423D8080">105</mods:start>
<mods:end id="3BF53A16A619A5D4574DE87F12A9B903">282</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location id="38386448EF0BB4643BDA17E11C7F22B7">
<mods:url id="B21033EBBEA57902589039A8577D1706">http://dx.doi.org/10.13133/2284-4880/545</mods:url>
</mods:location>
<mods:classification id="2BAF4E172E58873262C70A8C7A89F60F">journal article</mods:classification>
<mods:identifier id="CF3938FCB16737D8323CF1F7AAAC1422" type="DOI">10.13133/2284-4880/545</mods:identifier>
<mods:identifier id="59B9BDEE5B26EAE8B52194227C12469F" type="ISSN">2284-4880</mods:identifier>
<mods:identifier id="835E149D298C3413081BF94BECBF9549" type="Zenodo-Dep">12764154</mods:identifier>
<mods:identifier id="53DF19A1A2D5604AED3035E4A0B25093" type="ZooBank">B3F75A2C-7603-4027-B8E1-F4A8B5A0CDD9</mods:identifier>
</mods:mods>
<treatment id="0F7475501169FFD1FF16AE041ACF9188" ID-DOI="http://doi.org/10.5281/zenodo.12764055" ID-Zenodo-Dep="12764055" LSID="urn:lsid:zoobank.org:act:5C0B54B8-92E7-4CAA-B199-D5B2D46D83C2" httpUri="http://treatment.plazi.org/id/0F7475501169FFD1FF16AE041ACF9188" lastPageNumber="156" pageId="51" pageNumber="156">
<subSubSection id="CFC797CD1169FFD1FF16AE0418809368" box="[136,426,308,332]" pageId="51" pageNumber="156" type="nomenclature">
<paragraph id="8762C4461169FFD1FF16AE0418809368" blockId="51.[136,426,308,332]" box="[136,426,308,332]" pageId="51" pageNumber="156">
<heading id="DC2A732A1169FFD1FF16AE0418809368" box="[136,426,308,332]" fontSize="10" level="2" pageId="51" pageNumber="156" reason="5">
<taxonomicName id="40DDBFC51169FFD1FF16AE0418649368" ID-CoL="8SSQ5" authorityName="Ballerio" authorityYear="2021" box="[136,334,308,332]" class="Insecta" family="Hybosoridae" genus="Synarmostes" kingdom="Animalia" order="Coleoptera" pageId="51" pageNumber="156" phylum="Arthropoda" rank="species" species="fortis" status="sp. nov.">
<emphasis id="B5A918541169FFD1FF16AE0418649368" box="[136,334,308,332]" italics="true" pageId="51" pageNumber="156">Synarmostes fortis</emphasis>
</taxonomicName>
<emphasis id="B5A918541169FFD1FECBAE0518809368" bold="true" box="[341,426,308,332]" pageId="51" pageNumber="156">
<taxonomicNameLabel id="AE9AA52F1169FFD1FECBAE0518809368" box="[341,426,308,332]" pageId="51" pageNumber="156" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="CFC797CD1169FFD1FF16AE651B3F9388" pageId="51" pageNumber="156" type="description">
<paragraph id="8762C4461169FFD1FF16AE651B3F9388" blockId="51.[136,755,340,428]" pageId="51" pageNumber="156">
(
<figureCitation id="1FE6D8C31169FFD1FF0EAE6519CB9348" box="[144,225,340,364]" captionStart="Fig" captionStartId="34.[151,183,1858,1877]" captionTargetBox="[190,1405,209,1823]" captionTargetId="figure-13@34.[188,1422,195,1843]" captionTargetPageId="34" captionText="Fig. 22 Synarmostes compactus sp. nov., a, dorsal view of distended paratype; b, ventral view of rolled up paratype; c, dorsal view of rolled up paratype; d, lateral view of rolled up paratype; Synarmostes fortis sp. nov., e, dorsal view of distended holotype; f, ventral view of rolled up paratype; g, dorsal view of rolled up paratype; h, lateral view of rolled up paratype." figureDoi="http://doi.org/10.5281/zenodo.12764214" httpUri="https://zenodo.org/record/12764214/files/figure.png" pageId="51" pageNumber="156">Figs 22</figureCitation>
e-h, 30 j-l, 35) http://zoobank.org/
<uri id="F34CC8441169FFD1FEFAAE451B3F9388" pageId="51" pageNumber="156">
urn:lsid:zoobank.org:act:
<uuid id="F37BFE931169FFD1FDEAAE451B3F9388" pageId="51" pageNumber="156">5C0B54B8- 92E7-4CAA-B199-D5B2D46D83C2</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="CFC797CD1169FFD1FF30AEE51BC49628" pageId="51" pageNumber="156" type="materials_examined">
<paragraph id="8762C4461169FFD1FF30AEE51BC49628" blockId="51.[135,757,468,1932]" pageId="51" pageNumber="156">
<emphasis id="B5A918541169FFD1FF30AEE5181393C8" bold="true" box="[174,313,468,492]" pageId="51" pageNumber="156">Type series.</emphasis>
<materialsCitation id="37B5CE1B1169FFD1FED9AEE518B590C8" collectingDate="2003-12-07" collectorName="San Francisco, CA &amp; de Bekaraoka &amp; B. L. Fisher" country="Madagascar" determinerName="A. Ballerio" elevation="150" latitude="-13.166667" location="California Acad. of Sciences" longLatPrecision="21" longitude="49.71" municipality="Academy of Sciences" pageId="51" pageNumber="156" specimenCount="1" specimenCount-male="1" stateProvince="Antsiranana" typeStatus="holotype">
<typeStatus id="58667AE41169FFD1FED9AEE5188693C8" box="[327,428,468,492]" pageId="51" pageNumber="156" type="holotype">Holotype</typeStatus>
,
<specimenCount id="91DB0FCF1169FFD1FE21AEE418FF93C8" box="[447,469,469,492]" count="1" pageId="51" pageNumber="156" type="male"></specimenCount>
, deposited in the collection of the California
<collectingMunicipality id="67065E3C1169FFD1FEEAAEC51B4A9028" box="[372,608,500,524]" pageId="51" pageNumber="156">Academy of Sciences</collectingMunicipality>
,
<collectorName id="2A28A1901169FFD1FDF3AEC519F39008" pageId="51" pageNumber="156">San Francisco, CA</collectorName>
, USA, labelled as follows: / CASTYPE 20252 / CASENT8135204 /
<collectingCountry id="FFCA84D61169FFD1FEFFAD0518CF9068" box="[353,485,564,588]" name="Madagascar" pageId="51" pageNumber="156">Madagascar</collectingCountry>
:
<collectingRegion id="45190AA41169FFD1FE70AD051B449068" box="[494,622,564,588]" country="Madagascar" name="Antsiranana" pageId="51" pageNumber="156">Antsiranana</collectingRegion>
forêt
<collectorName id="2A28A1901169FFD1FD33AD0519F69048" pageId="51" pageNumber="156">de Bekaraoka</collectorName>
<locationDeviation id="96D0DD641169FFD1FF7BAD6518D29048" box="[229,504,596,620]" location="Daraina" pageId="51" pageNumber="156">
<quantity id="402569A31169FFD1FF7BAD65181B9048" box="[229,305,596,620]" metricMagnitude="3" metricUnit="m" metricValue="6.8" pageId="51" pageNumber="156" unit="km" value="6.8">6.8 km</quantity>
60° ENE Daraina
</locationDeviation>
el.
<quantity id="402569A31169FFD1FDBDAD651B429048" box="[547,616,596,620]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="51" pageNumber="156" unit="m" value="150.0">
<elevation id="0CF023751169FFD1FDBDAD651B429048" box="[547,616,596,620]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="51" pageNumber="156" unit="m" value="150.0">150 m</elevation>
</quantity>
<date id="F363E2861169FFD1FDEFAD64199790A8" pageId="51" pageNumber="156" value="2003-12-07">
<collectingDate id="E3271B6E1169FFD1FDEFAD64199790A8" pageId="51" pageNumber="156" value="2003-12-07">7 December 2003</collectingDate>
</date>
/
<geoCoordinate id="E2E9A2811169FFD1FF46AD45184390A8" box="[216,361,628,652]" degrees="13" direction="south" minutes="10" orientation="latitude" pageId="51" pageNumber="156" precision="15" seconds="00" value="-13.166667">13° 10 00” S</geoCoordinate>
<geoCoordinate id="E2E9A2811169FFD1FEECAD451B3390A8" box="[370,537,628,652]" degrees="049" direction="east" minutes="42" orientation="longitude" pageId="51" pageNumber="156" precision="15" seconds="36" value="49.71">049° 42 36” E</geoCoordinate>
<location id="8202929D1169FFD1FDBDAD4519CE9088" LSID="urn:lsid:plazi:treatment:0F7475501169FFD1FF16AE041ACF9188:8202929D1169FFD1FDBDAD4519CE9088" country="Madagascar" latitude="-13.166667" longLatPrecision="21" longitude="49.71" municipality="Academy of Sciences" name="California Acad. of Sciences" pageId="51" pageNumber="156" stateProvince="Antsiranana">California Acad. of Sciences</location>
collector:
<collectorName id="2A28A1901169FFD1FECAADA418F99088" box="[340,467,660,684]" pageId="51" pageNumber="156">B. L. Fisher</collectorName>
, sifted litter (leaf mold, rotten wood) tropical dry forest BLF9872 /
<taxonomicName id="40DDBFC51169FFD1FDAFAD841BDE90E8" authorityName="Ballerio" authorityYear="2021" box="[561,756,692,716]" class="Insecta" family="Hybosoridae" genus="Synarmostes" kingdom="Animalia" order="Coleoptera" pageId="51" pageNumber="156" phylum="Arthropoda" rank="species" species="fortis" status="sp. nov.">
<emphasis id="B5A918541169FFD1FDAFAD841BDE90E8" box="[561,756,692,716]" italics="true" pageId="51" pageNumber="156">Synarmostes fortis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="AE9AA52F1169FFD1FF19ADE4199590C8" box="[135,191,725,748]" pageId="51" pageNumber="156" rank="species">n. sp.</taxonomicNameLabel>
det.
<determinerName id="72BE28821169FFD1FF6EADE5184E90C8" box="[240,356,724,748]" pageId="51" pageNumber="156">A. Ballerio</determinerName>
2020
</materialsCitation>
<typeStatus id="58667AE41169FFD1FE3AADE51B3890C8" box="[420,530,724,748]" pageId="51" pageNumber="156" type="holotype">Holotypus</typeStatus>
<specimenCount id="91DB0FCF1169FFD1FD86ADE41B0690C8" box="[536,556,725,748]" count="1" pageId="51" pageNumber="156" type="male"></specimenCount>
/ [distended, glued on card, dissected, male genitalia glued in DMHF resin on a separate card, same pin].
<materialsCitation id="37B5CE1B1169FFD1FE3CAC2419939149" collectingDate="2003-12-07" collectionCode="CASC, ABCB" collectorName="San Francisco, CA &amp; de Bekaraoka &amp; B. L. Fisher" country="Madagascar" determinerName="A. Ballerio" elevation="150" latitude="-13.166667" location="California Acad. of Sciences" longLatPrecision="21" longitude="49.71" municipality="Academy of Sciences" pageId="51" pageNumber="156" specimenCount="4" specimenCount-male="4" stateProvince="Antsiranana" typeStatus="paratype">
<typeStatus id="58667AE41169FFD1FE3CAC241B239109" box="[418,521,789,813]" pageId="51" pageNumber="156" type="paratype">Paratypes</typeStatus>
[
<specimenCount id="91DB0FCF1169FFD1FD86AC251B149108" box="[536,574,788,812]" count="1" pageId="51" pageNumber="156" type="male">1 ♂</specimenCount>
dissected]:
<specimenCount id="91DB0FCF1169FFD1FD24AC251BDE9108" box="[698,756,788,812]" count="3" pageId="51" pageNumber="156" type="male">3 ♂♂</specimenCount>
(
<quantity id="402569A31169FFD1FF0EAC0519939168" box="[144,185,820,844]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="51" pageNumber="156" unit="in" value="2.0">2 in</quantity>
<collectionCode id="E1CC5C831169FFD1FF21AC05182E9168" box="[191,260,820,844]" pageId="51" pageNumber="156">CASC</collectionCode>
and
<quantity id="402569A31169FFD1FEA9AC05184A9168" box="[311,352,820,844]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="51" pageNumber="156" unit="in" value="1.0">1 in</quantity>
<collectionCode id="E1CC5C831169FFD1FEFBAC05189B9168" box="[357,433,820,844]" pageId="51" pageNumber="156">ABCB</collectionCode>
), same collecting data as holotype
</materialsCitation>
.
<emphasis id="B5A918541169FFD1FF57AC65182C91A8" bold="true" pageId="51" pageNumber="156">Further material examined (excluded from the type series)</emphasis>
:
<materialsCitation id="37B5CE1B1169FFD1FE8DAC451BF09628" collectingDate="2003-12-03" collectionCode="CASC" collectorName="de Binara &amp; B. L. Fisher" country="Madagascar" elevation="725" latitude="-13.263333" location="California Acad. of Sciences" longLatPrecision="21" longitude="49.603333" pageId="51" pageNumber="156" specimenCode="CASENT 8135284" specimenCount="1" specimenCount-female="1" stateProvince="Antsiranana">
<specimenCount id="91DB0FCF1169FFD1FE8DAC45181091A8" box="[275,314,884,908]" count="1" pageId="51" pageNumber="156" type="female">1 ♀</specimenCount>
(
<collectionCode id="E1CC5C831169FFD1FED9AC4518B891A8" box="[327,402,884,908]" pageId="51" pageNumber="156">CASC</collectionCode>
): /
<specimenCode id="D77B6C3D1169FFD1FE2EAC451B5391A8" box="[432,633,884,908]" collectionCode="CASC" pageId="51" pageNumber="156">CASENT 8135284</specimenCode>
/
<collectingCountry id="FFCA84D61169FFD1FD12AC4519869188" name="Madagascar" pageId="51" pageNumber="156">Madagascar</collectingCountry>
:
<collectingRegion id="45190AA41169FFD1FF28ACA5181C9188" box="[182,310,916,940]" country="Madagascar" name="Antsiranana" pageId="51" pageNumber="156">Antsiranana</collectingRegion>
forêt
<collectorName id="2A28A1901169FFD1FEE8ACA518F79188" box="[374,477,916,940]" pageId="51" pageNumber="156">de Binara</collectorName>
<locationDeviation id="96D0DD641169FFD1FE7BACA51BDE9188" box="[485,756,916,940]" location="Daraina" pageId="51" pageNumber="156">
<quantity id="402569A31169FFD1FE7BACA51B1A9188" box="[485,560,916,940]" metricMagnitude="3" metricUnit="m" metricValue="9.1" pageId="51" pageNumber="156" unit="km" value="9.1">9.1 km</quantity>
233° SW Daraina
</locationDeviation>
elev.
<quantity id="402569A31169FFD1FF5EAC85180691E8" box="[192,300,948,972]" metricMagnitude="2" metricUnit="m" metricValue="7.25" metricValueMax="8.0" metricValueMin="6.5" pageId="51" pageNumber="156" unit="m" value="725.0" valueMax="800.0" valueMin="650.0">
<elevation id="0CF023751169FFD1FF5EAC85180691E8" box="[192,300,948,972]" metricMagnitude="2" metricUnit="m" metricValue="7.25" metricValueMax="8.0" metricValueMin="6.5" pageId="51" pageNumber="156" unit="m" value="725.0" valueMax="800.0" valueMin="650.0">650-800m</elevation>
</quantity>
<date id="F363E2861169FFD1FEAAAC8518DB91E8" box="[308,497,948,972]" pageId="51" pageNumber="156" value="2003-12-03">
<collectingDate id="E3271B6E1169FFD1FEAAAC8518DB91E8" box="[308,497,948,972]" pageId="51" pageNumber="156" value="2003-12-03">3 December 2003</collectingDate>
</date>
/
<geoCoordinate id="E2E9A2811169FFD1FD99AC851BB891E8" box="[519,658,948,972]" degrees="13" direction="south" minutes="15" orientation="latitude" pageId="51" pageNumber="156" precision="15" seconds="48" value="-13.263333">13° 15 48” S</geoCoordinate>
<geoCoordinate id="E2E9A2811169FFD1FD07AC8519EF91C8" degrees="049" direction="east" minutes="36" orientation="longitude" pageId="51" pageNumber="156" precision="15" seconds="12" value="49.603333">049° 36 12” E</geoCoordinate>
<location id="8202929D1169FFD1FF53ACE518D691C8" LSID="urn:lsid:plazi:treatment:0F7475501169FFD1FF16AE041ACF9188:8202929D1169FFD1FF53ACE518D691C8" box="[205,508,980,1004]" country="Madagascar" latitude="-13.263333" longLatPrecision="21" longitude="49.603333" name="California Acad. of Sciences" pageId="51" pageNumber="156" stateProvince="Antsiranana">California Acad. of Sciences</location>
collector:
<collectorName id="2A28A1901169FFD1FDF1ACE41BDA91C8" box="[623,752,980,1004]" pageId="51" pageNumber="156">B. L. Fisher</collectorName>
, sifted litter (leaf mold, rotten wood) rainforest BLF9656
</materialsCitation>
/.
</paragraph>
</subSubSection>
<subSubSection id="CFC797CD1169FFD1FF30AB251B189788" pageId="51" pageNumber="156" type="diagnosis">
<paragraph id="8762C4461169FFD1FF30AB251B189788" blockId="51.[135,757,468,1932]" pageId="51" pageNumber="156">
<emphasis id="B5A918541169FFD1FF30AB25180E9608" bold="true" box="[174,292,1044,1068]" pageId="51" pageNumber="156">Diagnosis.</emphasis>
<taxonomicName id="40DDBFC51169FFD1FEB6AB2418C59608" authorityName="Ballerio" authorityYear="2021" box="[296,495,1044,1068]" class="Insecta" family="Hybosoridae" genus="Synarmostes" kingdom="Animalia" order="Coleoptera" pageId="51" pageNumber="156" phylum="Arthropoda" rank="species" species="fortis" status="sp. nov.">
<emphasis id="B5A918541169FFD1FEB6AB2418C59608" box="[296,495,1044,1068]" italics="true" pageId="51" pageNumber="156">Synarmostes fortis</emphasis>
</taxonomicName>
<emphasis id="B5A918541169FFD1FE6AAB251B629608" bold="true" box="[500,584,1044,1068]" pageId="51" pageNumber="156">
<taxonomicNameLabel id="AE9AA52F1169FFD1FE6AAB251B629608" box="[500,584,1044,1068]" pageId="51" pageNumber="156" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
can be differentiated from all other species of
<taxonomicName id="40DDBFC51169FFD1FE4EAB041B729668" authorityName="Germar" authorityYear="1843" box="[464,600,1077,1100]" class="Insecta" family="Hybosoridae" genus="Synarmostes" kingdom="Animalia" order="Coleoptera" pageId="51" pageNumber="156" phylum="Arthropoda" rank="genus">
<emphasis id="B5A918541169FFD1FE4EAB041B729668" box="[464,600,1077,1100]" italics="true" pageId="51" pageNumber="156">Synarmostes</emphasis>
</taxonomicName>
by the following combination of characters: a) medium size, b) ventral ocular area small-sized, c) dorsal ocular area vestigial, d) dorsal setation absent (20×), e) flightlessness, f) pronotum without callosities on disc, g) pronotum weakly saddle-shaped in lateral view, h) elytral punctation consisting of irregular rows of large transverse comma-shaped punctures, i) carinae of elytral apical third irregular (with some weak tubercles or other irregularities).
<taxonomicName id="40DDBFC51169FFD1FDB2AA041BDE9768" authorityName="Ballerio" authorityYear="2021" box="[556,756,1332,1356]" class="Insecta" family="Hybosoridae" genus="Synarmostes" kingdom="Animalia" order="Coleoptera" pageId="51" pageNumber="156" phylum="Arthropoda" rank="species" species="fortis" status="sp. nov.">
<emphasis id="B5A918541169FFD1FDB2AA041BDE9768" box="[556,756,1332,1356]" italics="true" pageId="51" pageNumber="156">Synarmostes fortis</emphasis>
</taxonomicName>
<emphasis id="B5A918541169FFD1FF16AA6519F79748" bold="true" box="[136,221,1364,1388]" pageId="51" pageNumber="156">
<taxonomicNameLabel id="AE9AA52F1169FFD1FF16AA6519F79748" box="[136,221,1364,1388]" pageId="51" pageNumber="156" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
belongs to the
<taxonomicName id="40DDBFC51169FFD1FEE3AA6518E29748" box="[381,456,1364,1388]" class="Insecta" family="Hybosoridae" genus="Synarmostes" kingdom="Animalia" order="Coleoptera" pageId="51" pageNumber="156" phylum="Arthropoda" rank="species" species="tibialis">
<emphasis id="B5A918541169FFD1FEE3AA6518E29748" box="[381,456,1364,1388]" italics="true" pageId="51" pageNumber="156">tibialis</emphasis>
</taxonomicName>
group of species and can be differentiated from the other species of the group by the combination of characters b), c) and h).
</paragraph>
</subSubSection>
<subSubSection id="CFC797CD1169FFD1FF30AA851C7A90E8" pageId="51" pageNumber="156" type="description">
<paragraph id="8762C4461169FFD1FF30AA851C7A90E8" blockId="51.[135,757,468,1932]" lastBlockId="51.[816,1438,180,940]" pageId="51" pageNumber="156">
<emphasis id="B5A918541169FFD1FF30AA85181897E8" bold="true" box="[174,306,1460,1484]" pageId="51" pageNumber="156">Description.</emphasis>
Size: HL =
<quantity id="402569A31169FFD1FE28AA851B2097E8" box="[438,522,1460,1484]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="51" pageNumber="156" unit="mm" value="0.6">0.6 mm</quantity>
; HW =
<quantity id="402569A31169FFD1FDFEAA851B9E97E8" box="[608,692,1460,1484]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="51" pageNumber="156" unit="mm" value="1.2">1.2 mm</quantity>
; PL =
<quantity id="402569A31169FFD1FF16AAE519F097C8" box="[136,218,1492,1516]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="51" pageNumber="156" unit="mm" value="1.1">1.1 mm</quantity>
; PW =
<quantity id="402569A31169FFD1FEB8AAE5185297C8" box="[294,376,1492,1516]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" pageId="51" pageNumber="156" unit="mm" value="1.9">1.9 mm</quantity>
; EL = 1,
<quantity id="402569A31169FFD1FE52AAE51B2597C8" box="[460,527,1492,1516]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" pageId="51" pageNumber="156" unit="mm" value="9.0">9 mm</quantity>
; EW =
<quantity id="402569A31169FFD1FDC2AAE51B8597C8" box="[604,687,1492,1516]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" pageId="51" pageNumber="156" unit="mm" value="1.9">1.9 mm</quantity>
. Overall morphology as in generic diagnosis. Small-sized flightless
<taxonomicName id="40DDBFC51169FFD1FF16A92418239408" authorityName="Germar" authorityYear="1843" box="[136,265,1557,1580]" class="Insecta" family="Hybosoridae" genus="Synarmostes" kingdom="Animalia" order="Coleoptera" pageId="51" pageNumber="156" phylum="Arthropoda" rank="genus">
<emphasis id="B5A918541169FFD1FF16A92418239408" box="[136,265,1557,1580]" italics="true" pageId="51" pageNumber="156">Synarmostes</emphasis>
</taxonomicName>
. Body strongly convex. Black, shiny, dorsum glabrous (20×), underside, tarsi and antennae reddish-brown, setation yellowish.
<emphasis id="B5A918541169FFD1FEC9A96518A59448" box="[343,399,1620,1644]" italics="true" pageId="51" pageNumber="156">Head</emphasis>
: W/L ratio = 1.37; clypeal apex obtuse, apical projection hardly appreciable, dorsal ocular area vestigial (hardly visible in transparence only), ventral ocular area small; head dorsal surface uniformly covered by dense strongly impressed large simple punctation. Fore margin with some irregular transverse lines. Interpunctural distance of simple impressed punctures irregular: from less than to once or twice larger than the punctural diameter. Vertex smooth with one transverse impressed irregular line shortly interrupted multiple times.
<emphasis id="B5A918541169FFD1FE39A8441B2695A8" box="[423,524,1909,1932]" italics="true" pageId="51" pageNumber="156">Pronotum</emphasis>
: subrectangular (W/L ratio = 1.91), irregularly convex, slightly depressed anteromedially with fore margin slightly raised, base and sides with sparse punctation, disc covered by dense short transverse impressed comma-shaped punctures, sides of disc covered by dense longitudinal comma-shaped punctures smaller than on disc. Interpunctural distance shorter than punctural diameter.
<taxonomicName id="40DDBFC51169FFD1FCAFAE451ABF93A8" box="[817,917,372,396]" class="Trilobita" family="Styginidae" genus="Scutellum" kingdom="Animalia" order="Corynexochida" pageId="51" pageNumber="156" phylum="Arthropoda" rank="genus">
<emphasis id="B5A918541169FFD1FCAFAE451ABF93A8" box="[817,917,372,396]" italics="true" pageId="51" pageNumber="156">Scutellum</emphasis>
</taxonomicName>
: base smooth, distal third covered by a few small comma-shaped punctures.
<emphasis id="B5A918541169FFD1FBD0AEA51DA79388" box="[1102,1165,404,428]" italics="true" pageId="51" pageNumber="156">Elytra</emphasis>
: longer than wide (W/L ratio = 0.88); subovoidal (dorsal view); inferior sutural stria reaching humeral area; humeral callus absent; elytral surface regularly strongly convex, covered by small relatively sparse comma-shaped punctures with opening directed backward. Interpunctural distance irregular: once or twice the length of punctural diameter. Elytral distal third with at least eight raised longitudinal irregular carinae.
<emphasis id="B5A918541169FFD1FB2EAD441DDD90A8" box="[1200,1271,629,652]" italics="true" pageId="51" pageNumber="156">Wings:</emphasis>
flightless.
<emphasis id="B5A918541169FFD1FAF8AD451AA09088" italics="true" pageId="51" pageNumber="156">Male genitalia</emphasis>
: aedeagus (L =
<quantity id="402569A31169FFD1FBADADA51DAE9088" box="[1075,1156,660,684]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="51" pageNumber="156" unit="mm" value="0.9">0.9 mm</quantity>
) with parameres as in
<figureCitation id="1FE6D8C31169FFD1FAEEADA51A6090E8" captionStart="Fig" captionStartId="42.[147,177,1885,1904]" captionTargetBox="[171,1427,173,1853]" captionTargetId="figure-13@42.[146,1452,163,1861]" captionTargetPageId="42" captionText="Fig. 30 Aedeagi and spicula gastrales, ac, Synarmostes aes sp. nov., holotype; df, Synarmostes separatus sp. nov., holotype; gi, Synarmostes compactus sp. nov., holotype; jl, Synarmostes fortis sp. nov., holotype; mo, Synarmostes makirovanae sp. nov., holotype; pr, Synarmostes lavasoae sp. nov., holotype." figureDoi="http://doi.org/10.5281/zenodo.12764233" httpUri="https://zenodo.org/record/12764233/files/figure.png" pageId="51" pageNumber="156">Figs 30</figureCitation>
j-k, spiculum gastrale (L =
<quantity id="402569A31169FFD1FBFCAD851D9890E8" box="[1122,1202,692,716]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="51" pageNumber="156" unit="mm" value="0.8">0.8 mm</quantity>
) as in
<figureCitation id="1FE6D8C31169FFD1FB6AAD851C6690E8" box="[1268,1356,692,716]" captionStart="Fig" captionStartId="42.[147,177,1885,1904]" captionTargetBox="[171,1427,173,1853]" captionTargetId="figure-13@42.[146,1452,163,1861]" captionTargetPageId="42" captionText="Fig. 30 Aedeagi and spicula gastrales, ac, Synarmostes aes sp. nov., holotype; df, Synarmostes separatus sp. nov., holotype; gi, Synarmostes compactus sp. nov., holotype; jl, Synarmostes fortis sp. nov., holotype; mo, Synarmostes makirovanae sp. nov., holotype; pr, Synarmostes lavasoae sp. nov., holotype." figureDoi="http://doi.org/10.5281/zenodo.12764233" httpUri="https://zenodo.org/record/12764233/files/figure.png" pageId="51" pageNumber="156">
Fig.
<specimenCount id="91DB0FCF1169FFD1FABCAD851C6690E8" box="[1314,1356,692,716]" count="30" pageId="51" pageNumber="156" type="generic">30 l</specimenCount>
</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="CFC797CD1169FFD1FCC8ADE51CA79168" pageId="51" pageNumber="156" type="distribution">
<paragraph id="8762C4461169FFD1FCC8ADE51CA79168" blockId="51.[816,1438,180,940]" pageId="51" pageNumber="156">
<emphasis id="B5A918541169FFD1FCC8ADE51D4690C8" bold="true" box="[854,1132,724,748]" pageId="51" pageNumber="156">Distribution and habitat.</emphasis>
See under
<taxonomicName id="40DDBFC51169FFD1FB7CADE41C5D90C8" authorityName="Ballerio" authorityYear="2021" box="[1250,1399,724,748]" class="Insecta" family="Hybosoridae" genus="Synarmostes" kingdom="Animalia" order="Coleoptera" pageId="51" pageNumber="156" phylum="Arthropoda" rank="species" species="bekaraokae" status="sp. nov.">
<emphasis id="B5A918541169FFD1FB7CADE41C5D90C8" box="[1250,1399,724,748]" italics="true" pageId="51" pageNumber="156">S. bekaraokae</emphasis>
</taxonomicName>
<emphasis id="B5A918541169FFD1FAE3ADE51A759128" bold="true" pageId="51" pageNumber="156">
<taxonomicNameLabel id="AE9AA52F1169FFD1FAE3ADE51A759128" pageId="51" pageNumber="156" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
A female tentatively attributed to this species has been found in the Binara forest (see under
<taxonomicName id="40DDBFC51169FFD1FB31AC251A5A9168" authorityName="Ballerio" authorityYear="2021" class="Insecta" family="Hybosoridae" genus="Goudotostes" kingdom="Animalia" order="Coleoptera" pageId="51" pageNumber="156" phylum="Arthropoda" rank="species" species="rasoamananae" status="sp. nov.">
<emphasis id="B5A918541169FFD1FB31AC251A5A9168" italics="true" pageId="51" pageNumber="156">Goudotostes rasoamananae</emphasis>
</taxonomicName>
<emphasis id="B5A918541169FFD1FCE9AC051AE09168" bold="true" box="[887,970,820,844]" pageId="51" pageNumber="156">
<taxonomicNameLabel id="AE9AA52F1169FFD1FCE9AC051AE09168" box="[887,970,820,844]" pageId="51" pageNumber="156" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
for more details on the collecting locality).
</paragraph>
</subSubSection>
<subSubSection id="CFC797CD1169FFD1FCC8AC651ACF9188" pageId="51" pageNumber="156" type="etymology">
<paragraph id="8762C4461169FFD1FCC8AC651ACF9188" blockId="51.[816,1438,180,940]" pageId="51" pageNumber="156">
<emphasis id="B5A918541169FFD1FCC8AC651AFD9148" bold="true" box="[854,983,852,876]" pageId="51" pageNumber="156">Etymology.</emphasis>
Latin adjective in the nominative singular, meaning “strong”, due to the hard bodied appearance of this new species.
</paragraph>
</subSubSection>
</treatment>
</document>