264 lines
33 KiB
XML
264 lines
33 KiB
XML
<document id="8BF5448032EDA3F88502568502F9E777" ID-DOI="10.13133/2284-4880/545" ID-ISSN="2284-4880" ID-Zenodo-Dep="12764154" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1721242503917" checkinUser="felipe" docAuthor="Ballerio, Alberto" docDate="2021" docId="0F7475501115FFB2FF16A825184294A8" docLanguage="en" docName="FragEntomol.53.2.105-282.pdf" docOrigin="Fragmenta entomologica 53 (2)" docSource="http://dx.doi.org/10.13133/2284-4880/545" docStyle="DocumentStyle:8499FF4308F512386C0E43DC290A16BD.1:FragEntomol.2021-.journal_article.0cover" docStyleId="8499FF4308F512386C0E43DC290A16BD" docStyleName="FragEntomol.2021-.journal_article.0cover" docStyleVersion="1" docTitle="Goudotostes masoalae Ballerio, 2021, sp. nov." docType="treatment" docUuid="0D95A1D6-93B5-4DBE-BA5E-A14E892DC50C" docUuidSource="ZooBank" docVersion="2" lastPageNumber="185" masterDocId="F34D0D28115AFFE2FF9EAF31192A9224" masterDocTitle="The Ceratocanthinae of Madagascar and Comoro Islands: a revision of the genera Synarmostes and Goudotostes, and of the flightless Philharmostes, with description of 64 new species (Coleoptera: Scarabaeoidea, Hybosoridae)" masterLastPageNumber="282" masterPageNumber="105" pageNumber="184" updateTime="1721247602959" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-4.0">
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<mods:titleInfo id="87A0BC56CA261BA9BE8F36B832B62B9E">
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<mods:title id="59922237F4A7EE8D4CAF72972D94E960">The Ceratocanthinae of Madagascar and Comoro Islands: a revision of the genera Synarmostes and Goudotostes, and of the flightless Philharmostes, with description of 64 new species (Coleoptera: Scarabaeoidea, Hybosoridae)</mods:title>
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</mods:titleInfo>
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<mods:name id="493426C9C60A6788E7C31AB757BDF8C5" type="personal">
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<mods:role id="93CC33FE0FADB0E044721CF5C761D5F7">
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<mods:roleTerm id="61D91DA71240D741B40550963227D146">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="B45475CC8CA93471CA701BBE9133330F">Ballerio, Alberto</mods:namePart>
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<mods:typeOfResource id="BB7127B50BD7BB741C42F853A938F54D">text</mods:typeOfResource>
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<mods:titleInfo id="7CDF5034FDE35DA664B9C5430FE5CEF3">
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<mods:title id="14EC9BBB7E3152F5D1A58B6B2674BD1E">Fragmenta entomologica</mods:title>
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</mods:titleInfo>
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<mods:part id="D2B3D4EA331202D569672B47CA96E04D">
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<mods:date id="691E879FBDE28B293F18CA262E506CEF">2021</mods:date>
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<mods:detail id="7D3FB7C9E5510DD0221D94E42F2E9877" type="pubDate">
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<mods:number id="B0479996774E98FA27A51C5B255CE641">2021-11-30</mods:number>
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<mods:detail id="450A4125946F96348F327F24B857B486" type="volume">
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<mods:number id="CD787B76AD1969899286B3CE127DBD21">53</mods:number>
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</mods:detail>
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<mods:detail id="B6A6BF838CC2092C7FE09852E516117C" type="issue">
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<mods:number id="FC524BD42C131D8ED36EA47D8EA713E9">2</mods:number>
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<mods:start id="39D5068A9EC84C041740B9F759AC7FF9">105</mods:start>
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<mods:end id="40BDCA0E8910FF6694EDFAF436630BF2">282</mods:end>
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</mods:extent>
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<mods:location id="BF166355DF907AFDBCDD900A11DBD3E9">
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<mods:url id="1CBEF26FF47D9B00D64AD74E6A8235B6">http://dx.doi.org/10.13133/2284-4880/545</mods:url>
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<mods:classification id="CECED7893224C3B6CB226BBD475E142C">journal article</mods:classification>
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<mods:identifier id="250CC6DCE56CA736E56DA99CE6574371" type="DOI">10.13133/2284-4880/545</mods:identifier>
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<mods:identifier id="56DEA7D44DBCD6D414FC91A7645B1B26" type="ISSN">2284-4880</mods:identifier>
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<mods:identifier id="84E6EAF0913C394B62AAB432BB678DFC" type="Zenodo-Dep">12764154</mods:identifier>
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<mods:identifier id="26A9371A60A7D5E1DCAF169122DAB1B7" type="ZooBank">B3F75A2C-7603-4027-B8E1-F4A8B5A0CDD9</mods:identifier>
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</mods:mods>
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<treatment id="0F7475501115FFB2FF16A825184294A8" LSID="urn:lsid:zoobank.org:act:0D95A1D6-93B5-4DBE-BA5E-A14E892DC50C" httpUri="http://treatment.plazi.org/id/0F7475501115FFB2FF16A825184294A8" lastPageId="80" lastPageNumber="185" pageId="79" pageNumber="184">
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<subSubSection id="CFC797CD1115FFADFF16A82518FC9508" box="[136,470,1812,1836]" pageId="79" pageNumber="184" type="nomenclature">
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<paragraph id="8762C4461115FFADFF16A82518FC9508" blockId="79.[136,470,1812,1836]" box="[136,470,1812,1836]" pageId="79" pageNumber="184">
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<heading id="DC2A732A1115FFADFF16A82518FC9508" box="[136,470,1812,1836]" fontSize="10" level="2" pageId="79" pageNumber="184" reason="5">
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<taxonomicName id="40DDBFC51115FFADFF16A82518509508" ID-CoL="8SSS7" authorityName="Ballerio" authorityYear="2021" box="[136,378,1812,1836]" class="Insecta" family="Hybosoridae" genus="Goudotostes" kingdom="Animalia" order="Coleoptera" pageId="79" pageNumber="184" phylum="Arthropoda" rank="species" species="masoalae" status="sp. nov.">
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<emphasis id="B5A918541115FFADFF16A82518509508" box="[136,378,1812,1836]" italics="true" pageId="79" pageNumber="184">Goudotostes masoalae</emphasis>
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</taxonomicName>
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<emphasis id="B5A918541115FFADFE1FA82518FC9508" bold="true" box="[385,470,1812,1836]" pageId="79" pageNumber="184">
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<taxonomicNameLabel id="AE9AA52F1115FFADFE1FA82518FC9508" box="[385,470,1812,1836]" pageId="79" pageNumber="184" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="CFC797CD1115FFADFF16A8051B3395A8" pageId="79" pageNumber="184" type="description">
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<paragraph id="8762C4461115FFADFF16A8051B3395A8" blockId="79.[136,756,1844,1932]" pageId="79" pageNumber="184">
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(
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<figureCitation id="1FE6D8C31115FFADFF0EA80519CB9568" box="[144,225,1844,1868]" captionStart="Fig" captionStartId="135.[136,167,1858,1877]" captionTargetBox="[198,1404,184,1825]" captionTargetId="figure-4@135.[160,1428,166,1830]" captionTargetPageId="135" captionText="Fig. 70 – Goudotostes masoalae sp. nov., a, female paratype, dorsal view of distended specimen; b, paratype, ventral view of rolled up specimen; c, paratype, dorsal view of rolled up specimen; d, paratype, ventral view of rolled up specimen; e, paratype, lateral view of rolled up specimen; f, paratype, pronotum of rolled up specimen; g, holotype, spiculum gastrale; h–k, holotype, four views of aedeagus." figureDoi="http://doi.org/10.5281/zenodo.12764325" httpUri="https://zenodo.org/record/12764325/files/figure.png" pageId="79" pageNumber="184">Figs 70</figureCitation>
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a-k, 77) http://zoobank.org/
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<uri id="F34CC8441115FFADFEFCA8651B3395A8" pageId="79" pageNumber="184">
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||
urn:lsid:zoobank.org:act:
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<uuid id="F37BFE931115FFADFDEFA8651B3395A8" pageId="79" pageNumber="184">0D95A1D6- 93B5-4DBE-BA5E-A14E892DC50C</uuid>
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||
</uri>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="CFC797CD1115FFADFCC8AF851C6890A8" pageId="79" pageNumber="184" type="materials_examined">
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<paragraph id="8762C4461115FFADFCC8AF851C6890A8" blockId="79.[816,1439,180,1932]" pageId="79" pageNumber="184">
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<emphasis id="B5A918541115FFADFCC8AF851AF092E8" bold="true" box="[854,986,180,204]" pageId="79" pageNumber="184">Type series.</emphasis>
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<materialsCitation id="37B5CE1B1115FFADFC7EAF851CB79348" collectingDate="1993-12-02" collectorName="Chicago, IL & B. L. Fisher" country="Madagascar" county="Ambohitstondroina" determinerName="A. Ballerio" location="Field Museum of Natural" municipality="Tamatave" pageId="79" pageNumber="184" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
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<typeStatus id="58667AE41115FFADFC7EAF851D6E92E8" box="[992,1092,180,204]" pageId="79" pageNumber="184" type="holotype">Holotype</typeStatus>
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,
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<specimenCount id="91DB0FCF1115FFADFBD0AF841D4E92E8" box="[1102,1124,181,204]" count="1" pageId="79" pageNumber="184" type="male">♂</specimenCount>
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, deposited in the collection of the
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<location id="8202929D1115FFADFCC8AFE51D7792C8" LSID="urn:lsid:plazi:treatment:0F7475501115FFB2FF16A825184294A8:8202929D1115FFADFCC8AFE51D7792C8" box="[854,1117,212,236]" country="Madagascar" county="Ambohitstondroina" municipality="Tamatave" name="Field Museum of Natural" pageId="79" pageNumber="184">Field Museum of Natural</location>
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history,
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<collectorName id="2A28A1901115FFADFB29AFE51C1F92C8" box="[1207,1333,212,236]" pageId="79" pageNumber="184">Chicago, IL</collectorName>
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||
, USA, labelled as follows:
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<emphasis id="B5A918541115FFADFC71AFC51ADC9328" bold="true" box="[1007,1014,244,268]" pageId="79" pageNumber="184">/</emphasis>
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||
<collectingCountry id="FFCA84D61115FFADFC63AFC51DAA9328" box="[1021,1152,244,268]" name="Madagascar" pageId="79" pageNumber="184">Madagascar</collectingCountry>
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||
:
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<collectingMunicipality id="67065E3C1115FFADFB16AFC41DC79328" box="[1160,1261,245,268]" pageId="79" pageNumber="184">Tamatave</collectingMunicipality>
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||
<locationDeviation id="96D0DD641115FFADFB6AAFC51AA59308" location="Ambanizana" pageId="79" pageNumber="184">
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||
<quantity id="402569A31115FFADFB6AAFC51C179328" box="[1268,1341,244,268]" metricMagnitude="3" metricUnit="m" metricValue="6.9" pageId="79" pageNumber="184" unit="km" value="6.9">6.9 km</quantity>
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NE Ambanizana
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</locationDeviation>
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,
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<collectingCounty id="6E03BCCA1115FFADFC07AE251D4D9308" box="[921,1127,276,300]" pageId="79" pageNumber="184">Ambohitstondroina</collectingCounty>
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,
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<date id="F363E2861115FFADFBECAE251DCD9308" box="[1138,1255,276,300]" pageId="79" pageNumber="184" value="1993-12-02">
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||
<collectingDate id="E3271B6E1115FFADFBECAE251DCD9308" box="[1138,1255,276,300]" pageId="79" pageNumber="184" value="1993-12-02">2.XII.1993</collectingDate>
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</date>
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, FMHD#93-338, rainforest, sifted litter (leaf mold, rotten wood)
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<collectorName id="2A28A1901115FFADFA85AE041CB29368" box="[1307,1432,308,332]" pageId="79" pageNumber="184">B. L. Fisher</collectorName>
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, #976 /
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<taxonomicName id="40DDBFC51115FFADFCE0AE651D479348" authorityName="Ballerio" authorityYear="2021" box="[894,1133,340,364]" class="Insecta" family="Hybosoridae" genus="Goudotostes" kingdom="Animalia" order="Coleoptera" pageId="79" pageNumber="184" phylum="Arthropoda" rank="species" species="masoalae" status="sp. nov.">
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||
<emphasis id="B5A918541115FFADFCE0AE651D479348" box="[894,1133,340,364]" italics="true" pageId="79" pageNumber="184">Goudotostes masoalae</emphasis>
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||
</taxonomicName>
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||
<taxonomicNameLabel id="AE9AA52F1115FFADFBE9AE641D989348" box="[1143,1202,341,364]" pageId="79" pageNumber="184" rank="species">n. sp.</taxonomicNameLabel>
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||
det.
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||
<determinerName id="72BE28821115FFADFB76AE651C759348" box="[1256,1375,340,364]" pageId="79" pageNumber="184">A. Ballerio</determinerName>
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||
2020
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||
</materialsCitation>
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||
<typeStatus id="58667AE41115FFADFCAEAE451AB793A8" box="[816,925,372,396]" pageId="79" pageNumber="184" type="holotype">Holotypus</typeStatus>
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||
<specimenCount id="91DB0FCF1115FFADFC39AE441A9193A8" box="[935,955,373,396]" count="1" pageId="79" pageNumber="184" type="male">♂</specimenCount>
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||
/ [distended, glued on card, dissected, male genitalia glued in DMHF resin on a separate card, same pin].
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<materialsCitation id="37B5CE1B1115FFADFCF8AE841C0490A8" collectingDate="2003-02-26" collectingDateMax="2003-03-06" collectingDateMin="2003-02-26" collectionCode="FMNH, ABCB, CASC" collectorName="D. Andriamalala & Andriamalala, D" country="Madagascar" county="Ambanizana" determinerName="A. Ballerio" elevation="900" latitude="-15.571667" location="California Acad. of Sciences" longLatPrecision="21" longitude="50.00611" municipality="Parc National Masoala" pageId="79" pageNumber="184" specimenCount="32" specimenCount-female="19" specimenCount-male="13" stateProvince="Toamasina" typeStatus="paratype">
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<typeStatus id="58667AE41115FFADFCF8AE841AE693E9" box="[870,972,437,461]" pageId="79" pageNumber="184" type="paratype">Paratypes</typeStatus>
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[12 dissected
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<specimenCount id="91DB0FCF1115FFADFBC1AE841DA693E8" box="[1119,1164,437,460]" count="1" pageId="79" pageNumber="184" type="male">♂♂</specimenCount>
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]:
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<specimenCount id="91DB0FCF1115FFADFB05AE851DC893E8" box="[1179,1250,436,460]" count="11" pageId="79" pageNumber="184" type="male">11 ♂♂</specimenCount>
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and
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<specimenCount id="91DB0FCF1115FFADFA8DAE851C7093E8" box="[1299,1370,436,460]" count="17" pageId="79" pageNumber="184" type="female">17 ♀♀</specimenCount>
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||
(
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<quantity id="402569A31115FFADFAF9AE851CB793E8" box="[1383,1437,436,460]" metricMagnitude="-1" metricUnit="m" metricValue="5.334" pageId="79" pageNumber="184" unit="in" value="21.0">21 in</quantity>
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||
<collectionCode id="E1CC5C831115FFADFCAEAEE41A5693C8" box="[816,892,469,492]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="79" pageNumber="184" type="Museum">FMNH</collectionCode>
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||
and
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||
<quantity id="402569A31115FFADFC33AEE41AFE93C8" box="[941,980,468,492]" metricMagnitude="-1" metricUnit="m" metricValue="1.778" pageId="79" pageNumber="184" unit="in" value="7.0">7 in</quantity>
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||
<collectionCode id="E1CC5C831115FFADFC46AEE51D0993C8" box="[984,1059,468,492]" pageId="79" pageNumber="184">ABCB</collectionCode>
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||
), same collecting data as holotype.
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||
<specimenCount id="91DB0FCF1115FFADFA11AEE51A6E9028" count="1" pageId="79" pageNumber="184" type="male">1 ♂</specimenCount>
|
||
and
|
||
<specimenCount id="91DB0FCF1115FFADFCEBAEC51A9A9028" box="[885,944,500,524]" count="2" pageId="79" pageNumber="184" type="female">2 ♀♀</specimenCount>
|
||
(
|
||
<collectionCode id="E1CC5C831115FFADFC23AEC51D229028" box="[957,1032,500,524]" pageId="79" pageNumber="184">CASC</collectionCode>
|
||
):
|
||
<collectingCountry id="FFCA84D61115FFADFB86AEC51DBC9028" box="[1048,1174,500,524]" name="Madagascar" pageId="79" pageNumber="184">Madagascar</collectingCountry>
|
||
<collectingRegion id="45190AA41115FFADFB02AEC51C279028" box="[1180,1293,500,524]" country="Madagascar" name="Toamasina" pageId="79" pageNumber="184">Toamasina</collectingRegion>
|
||
<collectingCounty id="6E03BCCA1115FFADFA8FAEC51CB29028" box="[1297,1432,500,524]" pageId="79" pageNumber="184">Ambanizana</collectingCounty>
|
||
,
|
||
<collectingMunicipality id="67065E3C1115FFADFCAEAD241D0B9008" box="[816,1057,532,556]" pageId="79" pageNumber="184">Parc National Masoala</collectingMunicipality>
|
||
elev.
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||
<quantity id="402569A31115FFADFBFFAD251D8C9008" box="[1121,1190,532,556]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="79" pageNumber="184" unit="m" value="900.0">
|
||
<elevation id="0CF023751115FFADFBFFAD251D8C9008" box="[1121,1190,532,556]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="79" pageNumber="184" unit="m" value="900.0">900 m</elevation>
|
||
</quantity>
|
||
<date id="F363E2861115FFADFB33AD251CB69008" box="[1197,1436,532,556]" pageId="79" pageNumber="184" value="2003-02-26" valueMax="2003-03-06" valueMin="2003-02-26">
|
||
<collectingDate id="E3271B6E1115FFADFB33AD251CB69008" box="[1197,1436,532,556]" pageId="79" pageNumber="184" value="2003-02-26" valueMax="2003-03-06" valueMin="2003-02-26">26 Feb.-6 March 2003</collectingDate>
|
||
</date>
|
||
<geoCoordinate id="E2E9A2811115FFADFCACAD051A959068" box="[818,959,564,588]" degrees="15" direction="south" minutes="34" orientation="latitude" pageId="79" pageNumber="184" precision="15" seconds="18" value="-15.571667">15° 34’ 18” S</geoCoordinate>
|
||
<geoCoordinate id="E2E9A2811115FFADFC5BAD051D4A9068" box="[965,1120,564,588]" degrees="050" direction="east" minutes="00" orientation="longitude" pageId="79" pageNumber="184" precision="15" seconds="22" value="50.00611">050° 00’ 22” E</geoCoordinate>
|
||
/
|
||
<location id="8202929D1115FFADFBEAAD051CB49068" LSID="urn:lsid:plazi:treatment:0F7475501115FFB2FF16A825184294A8:8202929D1115FFADFBEAAD051CB49068" box="[1140,1438,564,588]" country="Madagascar" county="Ambanizana" latitude="-15.571667" longLatPrecision="21" longitude="50.00611" municipality="Parc National Masoala" name="California Acad. of Sciences" pageId="79" pageNumber="184" stateProvince="Toamasina">California Acad. of Sciences</location>
|
||
collectors:
|
||
<collectorName id="2A28A1901115FFADFC3DAD641D799048" box="[931,1107,596,620]" pageId="79" pageNumber="184">D. Andriamalala</collectorName>
|
||
, D. Silva et al. sifted litter (leaf mold, rotten wood) montane rainforest BLF8651
|
||
</materialsCitation>
|
||
/.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="CFC797CD1115FFADFCC8ADA51DB69708" pageId="79" pageNumber="184" type="diagnosis">
|
||
<paragraph id="8762C4461115FFADFCC8ADA51DB69708" blockId="79.[816,1439,180,1932]" pageId="79" pageNumber="184">
|
||
<emphasis id="B5A918541115FFADFCC8ADA51AE19088" bold="true" box="[854,971,660,684]" pageId="79" pageNumber="184">Diagnosis.</emphasis>
|
||
<taxonomicName id="40DDBFC51115FFADFC4FADA51DE99088" authorityName="Ballerio" authorityYear="2021" box="[977,1219,660,684]" class="Insecta" family="Hybosoridae" genus="Goudotostes" kingdom="Animalia" order="Coleoptera" pageId="79" pageNumber="184" phylum="Arthropoda" rank="species" species="masoalae" status="sp. nov.">
|
||
<emphasis id="B5A918541115FFADFC4FADA51DE99088" box="[977,1219,660,684]" italics="true" pageId="79" pageNumber="184">Goudotostes masoalae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B5A918541115FFADFB54ADA51C359088" bold="true" box="[1226,1311,660,684]" pageId="79" pageNumber="184">
|
||
<taxonomicNameLabel id="AE9AA52F1115FFADFB54ADA51C359088" box="[1226,1311,660,684]" pageId="79" pageNumber="184" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
can be easily differentiated from all other species of
|
||
<taxonomicName id="40DDBFC51115FFADFA89AD851CB690E8" authorityName=", sensu Ballerio" authorityYear="2008" box="[1303,1436,692,716]" class="Insecta" family="Hybosoridae" genus="Goudotostes" kingdom="Animalia" order="Coleoptera" pageId="79" pageNumber="184" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B5A918541115FFADFA89AD851CB690E8" box="[1303,1436,692,716]" italics="true" pageId="79" pageNumber="184">Goudotostes</emphasis>
|
||
</taxonomicName>
|
||
by the following combination of characters: a) small size, b) dorsal ocular area absent, c) clypeal apical projection pointed, d) clypeal punctation made of dense small impressed comma-shaped punctures, e) fronto-clypeal tubercle present, f) pronotal anterior carina bilobate, g) pronotal basal carina bilobate, h) pronotal disc covered by dense small simple deeply impressed punctures, i) elytral dorsal area covered by longitudinal irregular rows of blunt weakly pronounced carinae and small tubercles, j) sides of elytra covered by several small weakly pronounced tubercles, k) left paramere with ventral margin invaginated (a character which places this species in the
|
||
<taxonomicName id="40DDBFC51115FFADFA89AB051C7F9668" authorityName="Ballerio" authorityYear="2021" box="[1303,1365,1076,1100]" class="Insecta" family="Hybosoridae" genus="Goudotostes" kingdom="Animalia" order="Coleoptera" pageId="79" pageNumber="184" phylum="Arthropoda" rank="species" species="laevis">
|
||
<emphasis id="B5A918541115FFADFA89AB051C7F9668" box="[1303,1365,1076,1100]" italics="true" pageId="79" pageNumber="184">laevis</emphasis>
|
||
</taxonomicName>
|
||
group of species), l) the distinctive shape of parameres as in
|
||
<figureCitation id="1FE6D8C31115FFADFAEAAB651A6196A8" captionStart="Fig" captionStartId="135.[136,167,1858,1877]" captionTargetBox="[198,1404,184,1825]" captionTargetId="figure-4@135.[160,1428,166,1830]" captionTargetPageId="135" captionText="Fig. 70 – Goudotostes masoalae sp. nov., a, female paratype, dorsal view of distended specimen; b, paratype, ventral view of rolled up specimen; c, paratype, dorsal view of rolled up specimen; d, paratype, ventral view of rolled up specimen; e, paratype, lateral view of rolled up specimen; f, paratype, pronotum of rolled up specimen; g, holotype, spiculum gastrale; h–k, holotype, four views of aedeagus." figureDoi="http://doi.org/10.5281/zenodo.12764325" httpUri="https://zenodo.org/record/12764325/files/figure.png" pageId="79" pageNumber="184">Fig. 70</figureCitation>
|
||
h-k. Within the
|
||
<taxonomicName id="40DDBFC51115FFADFC67AB451D1D96A8" authorityName="Ballerio" authorityYear="2021" box="[1017,1079,1140,1164]" class="Insecta" family="Hybosoridae" genus="Goudotostes" kingdom="Animalia" order="Coleoptera" pageId="79" pageNumber="184" phylum="Arthropoda" rank="species" species="laevis">
|
||
<emphasis id="B5A918541115FFADFC67AB451D1D96A8" box="[1017,1079,1140,1164]" italics="true" pageId="79" pageNumber="184">laevis</emphasis>
|
||
</taxonomicName>
|
||
species group,
|
||
<taxonomicName id="40DDBFC51115FFADFB42AB451A419688" authorityName="Ballerio" authorityYear="2021" class="Insecta" family="Hybosoridae" genus="Goudotostes" kingdom="Animalia" order="Coleoptera" pageId="79" pageNumber="184" phylum="Arthropoda" rank="species" species="masoalae" status="sp. nov.">
|
||
<emphasis id="B5A918541115FFADFB42AB451A419688" italics="true" pageId="79" pageNumber="184">Goudotostes masoalae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B5A918541115FFADFCEBABA51AE79688" bold="true" box="[885,973,1172,1196]" pageId="79" pageNumber="184">
|
||
<taxonomicNameLabel id="AE9AA52F1115FFADFCEBABA51AE79688" box="[885,973,1172,1196]" pageId="79" pageNumber="184" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is similar to
|
||
<taxonomicName id="40DDBFC51115FFADFBFDABA51C119688" authorityName="Ballerio" authorityYear="2021" box="[1123,1339,1172,1196]" class="Insecta" family="Hybosoridae" genus="Goudotostes" kingdom="Animalia" order="Coleoptera" pageId="79" pageNumber="184" phylum="Arthropoda" rank="species" species="parvus" status="sp. nov.">
|
||
<emphasis id="B5A918541115FFADFBFDABA51C119688" box="[1123,1339,1172,1196]" italics="true" pageId="79" pageNumber="184">Goudotostes parvus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B5A918541115FFADFADAABA51CB79688" bold="true" box="[1348,1437,1172,1196]" pageId="79" pageNumber="184">
|
||
<taxonomicNameLabel id="AE9AA52F1115FFADFADAABA51CB79688" box="[1348,1437,1172,1196]" pageId="79" pageNumber="184" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
due to the sculpturing of elytra but
|
||
<taxonomicName id="40DDBFC51115FFADFB2AAB841C1796E8" authorityName="Ballerio" authorityYear="2021" box="[1204,1341,1204,1228]" class="Insecta" family="Hybosoridae" genus="Goudotostes" kingdom="Animalia" order="Coleoptera" pageId="79" pageNumber="184" phylum="Arthropoda" rank="species" species="masoalae" status="sp. nov.">
|
||
<emphasis id="B5A918541115FFADFB2AAB841C1796E8" box="[1204,1341,1204,1228]" italics="true" pageId="79" pageNumber="184">G. masoalae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B5A918541115FFADFADBAB851CB796E8" bold="true" box="[1349,1437,1204,1228]" pageId="79" pageNumber="184">
|
||
<taxonomicNameLabel id="AE9AA52F1115FFADFADBAB851CB796E8" box="[1349,1437,1204,1228]" pageId="79" pageNumber="184" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
has pointed clypeal apical projection as well as denser and smaller pronotal punctation and denser and smaller comma-shaped punctation on clypeus.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="CFC797CD1115FFB2FCC8AA051BC49769" lastPageId="80" lastPageNumber="185" pageId="79" pageNumber="184" type="description">
|
||
<paragraph id="8762C4461115FFB2FCC8AA051BC49769" blockId="79.[816,1439,180,1932]" lastBlockId="80.[150,772,180,1676]" lastPageId="80" lastPageNumber="185" pageId="79" pageNumber="184">
|
||
<emphasis id="B5A918541115FFADFCC8AA051ACD9768" bold="true" box="[854,999,1332,1356]" pageId="79" pageNumber="184">Description.</emphasis>
|
||
Size: HL =
|
||
<quantity id="402569A31115FFADFBE0AA051DF09768" box="[1150,1242,1332,1356]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="79" pageNumber="184" unit="mm" value="0.9">0.9 mm</quantity>
|
||
; HW =
|
||
<quantity id="402569A31115FFADFAA2AA051CBD9768" box="[1340,1431,1332,1356]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="79" pageNumber="184" unit="mm" value="1.2">1.2 mm</quantity>
|
||
; PL =
|
||
<quantity id="402569A31115FFADFCE6AA651AFE9748" box="[888,980,1364,1388]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="79" pageNumber="184" unit="mm" value="1.1">1.1 mm</quantity>
|
||
; PW =
|
||
<quantity id="402569A31115FFADFBABAA651DB89748" box="[1077,1170,1364,1388]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" pageId="79" pageNumber="184" unit="mm" value="1.9">1.9 mm</quantity>
|
||
; EL = 2.0 mm; EW = 2.0 mm. Overall morphology as in generic diagnosis. Small-sized
|
||
<taxonomicName id="40DDBFC51115FFADFC25AAA51D6C9788" authorityName=", sensu Ballerio" authorityYear="2008" box="[955,1094,1428,1452]" class="Insecta" family="Hybosoridae" genus="Goudotostes" kingdom="Animalia" order="Coleoptera" pageId="79" pageNumber="184" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B5A918541115FFADFC25AAA51D6C9788" box="[955,1094,1428,1452]" italics="true" pageId="79" pageNumber="184">Goudotostes</emphasis>
|
||
</taxonomicName>
|
||
. Body strongly convex. Black, shiny, dorsum with extremely short fine setation (50×), underside, tarsi and antennae dark reddish-brown, setation yellowish-brown.
|
||
<emphasis id="B5A918541115FFADFBAFAAC51D469428" box="[1073,1132,1524,1548]" italics="true" pageId="79" pageNumber="184">Head</emphasis>
|
||
: W/L ratio = 1.60; clypeal apex obtuse, apical projection strongly developed and sexually dimorphic, sides almost rectilinear with slightly unaligned genae in correspondence with the genal suture; dorsal ocular area absent, genal canthus indistinct, ventral ocular area small-sized, genae rounded, distinctly protruding outwards, genal suture indistinct; frons covered by large simple impressed punctures, interpunctural distance equal to or slightly inferior than punctural diameter, clypeal dorsal surface covered by medium-sized dense impressed comma-shaped punctures, fore margin marked by some irregular transverse lines. Interpunctural distance of clypeal punctures inferior than punctural diameter. Fronto-clypeal tubercle raised pointed and smooth.
|
||
<emphasis id="B5A91854110AFFB2FE70AFE41B7192C8" box="[494,603,213,236]" italics="true" pageId="80" pageNumber="185">Pronotum</emphasis>
|
||
: subrectangular (dorsal view) (W/L ratio = 1.90), convex, fore angles distinctly subtruncate; pronotal surface covered by dense large deep simple punctures becoming comma-shaped at sides, area near fore angles with few irregular lines, interpunctural distance mostly inferior than or equal to punctural diameter; pronotal bead present only along sides and fore angles; posterior carina bilobate and raised; anterior carina bilobate and raised; remaining pronotal sculpturing made of large and gently raised tubercles and carinae as follows: sides of disc with two gently raised transverse short carinae, each side with one gently raised tubercle and one transverse carina reaching outer margin.
|
||
<taxonomicName id="40DDBFC5110AFFB2FE33AD451B3090A8" box="[429,538,628,652]" class="Trilobita" family="Styginidae" genus="Scutellum" kingdom="Animalia" order="Corynexochida" pageId="80" pageNumber="185" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B5A91854110AFFB2FE33AD451B3090A8" box="[429,538,628,652]" italics="true" pageId="80" pageNumber="185">Scutellum</emphasis>
|
||
</taxonomicName>
|
||
: covered by sparse impressed transverse large comma-shaped punctures.
|
||
<emphasis id="B5A91854110AFFB2FF09AD8519F690E8" box="[151,220,692,716]" italics="true" pageId="80" pageNumber="185">Elytra</emphasis>
|
||
: longer than wide (W/L ratio = 1.07), subovoidal (dorsal view), strongly convex; humeral callus marked by a tubercle; elytral surface uniformly covered by large sparse deep comma-shaped punctures with opening directed backwards mixed to large impressed simple punctures, interpunctural distance about equal to punctural diameter. Striated articular area hardly visible in lateral view, relatively narrow and short; inferior sutural stria visible and complete; marginal area medium-sized; elytral articular process well developed, smooth and shiny, isolated from remaining elytral surface by a deep furrow. Elytral sculpturing as follows: base with one raised large tubercle below articular process (parasutural tubercle), next to it a humeral oblique large raised tubercle (humeral tubercle), dorsal area of elytra with two irregular and asymmetrical rows of weakly raised smooth short tubercles or, in some specimens, short carinae, sides and apical third with some more raised tubercles and longitudinal short carinae.
|
||
<emphasis id="B5A91854110AFFB2FEB0ABC4185A9728" box="[302,368,1269,1292]" italics="true" pageId="80" pageNumber="185">Wings</emphasis>
|
||
: flightless (micropterous).
|
||
<emphasis id="B5A91854110AFFB2FD08ABC519FB9708" italics="true" pageId="80" pageNumber="185">Male genitalia</emphasis>
|
||
: aedeagus (L =
|
||
<quantity id="402569A3110AFFB2FE1AAA2518F09708" box="[388,474,1300,1324]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="80" pageNumber="185" unit="mm" value="1.2">1.2 mm</quantity>
|
||
) with parameres as in
|
||
<figureCitation id="1FE6D8C3110AFFB2FD4AAA2519989768" captionStart="Fig" captionStartId="135.[136,167,1858,1877]" captionTargetBox="[198,1404,184,1825]" captionTargetId="figure-4@135.[160,1428,166,1830]" captionTargetPageId="135" captionText="Fig. 70 – Goudotostes masoalae sp. nov., a, female paratype, dorsal view of distended specimen; b, paratype, ventral view of rolled up specimen; c, paratype, dorsal view of rolled up specimen; d, paratype, ventral view of rolled up specimen; e, paratype, lateral view of rolled up specimen; f, paratype, pronotum of rolled up specimen; g, holotype, spiculum gastrale; h–k, holotype, four views of aedeagus." figureDoi="http://doi.org/10.5281/zenodo.12764325" httpUri="https://zenodo.org/record/12764325/files/figure.png" pageId="80" pageNumber="185">Figs 70</figureCitation>
|
||
h-k, spiculum gastrale (L = 1.0 mm) as in
|
||
<figureCitation id="1FE6D8C3110AFFB2FD18AA051BC09769" box="[646,746,1332,1357]" captionStart="Fig" captionStartId="135.[136,167,1858,1877]" captionTargetBox="[198,1404,184,1825]" captionTargetId="figure-4@135.[160,1428,166,1830]" captionTargetPageId="135" captionText="Fig. 70 – Goudotostes masoalae sp. nov., a, female paratype, dorsal view of distended specimen; b, paratype, ventral view of rolled up specimen; c, paratype, dorsal view of rolled up specimen; d, paratype, ventral view of rolled up specimen; e, paratype, lateral view of rolled up specimen; f, paratype, pronotum of rolled up specimen; g, holotype, spiculum gastrale; h–k, holotype, four views of aedeagus." figureDoi="http://doi.org/10.5281/zenodo.12764325" httpUri="https://zenodo.org/record/12764325/files/figure.png" pageId="80" pageNumber="185">
|
||
Fig.
|
||
<quantity id="402569A3110AFFB2FD26AA051BC09769" box="[696,746,1332,1357]" metricMagnitude="-2" metricUnit="kg" metricValue="7.0" pageId="80" pageNumber="185" unit="g" value="70.0">70 g</quantity>
|
||
</figureCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="CFC797CD110AFFB2FF23AA65180897A8" pageId="80" pageNumber="185" type="etymology">
|
||
<paragraph id="8762C446110AFFB2FF23AA65180897A8" blockId="80.[150,772,180,1676]" pageId="80" pageNumber="185">
|
||
<emphasis id="B5A91854110AFFB2FF23AA6518149748" bold="true" box="[189,318,1364,1388]" pageId="80" pageNumber="185">Etymology.</emphasis>
|
||
Noun in the genitive case. Named after the
|
||
<typeStatus id="58667AE4110AFFB2FF09AA4419EF97A9" box="[151,197,1397,1421]" pageId="80" pageNumber="185">type</typeStatus>
|
||
locality.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="CFC797CD110AFFB2FF23AAA5184294A8" pageId="80" pageNumber="185" type="distribution">
|
||
<paragraph id="8762C446110AFFB2FF23AAA5184294A8" blockId="80.[150,772,180,1676]" pageId="80" pageNumber="185">
|
||
<emphasis id="B5A91854110AFFB2FF23AAA518F69788" bold="true" box="[189,476,1428,1452]" pageId="80" pageNumber="185">Distribution and habitat.</emphasis>
|
||
Known only from the
|
||
<typeStatus id="58667AE4110AFFB2FD4BAAA41A299789" box="[725,771,1429,1453]" pageId="80" pageNumber="185">type</typeStatus>
|
||
locality in Eastern
|
||
<collectingCountry id="FFCA84D6110AFFB2FEF6AA8518C197E8" box="[360,491,1460,1484]" name="Madagascar" pageId="80" pageNumber="185">Madagascar</collectingCountry>
|
||
. The
|
||
<typeStatus id="58667AE4110AFFB2FDB7AA841B7D97E9" box="[553,599,1461,1485]" pageId="80" pageNumber="185">type</typeStatus>
|
||
series was collected by sifting leaf litter and rotten wood in montane rainforest. The vegetation cover consists of “medium altitude moist evergreen forest”, (
|
||
<bibRefCitation id="E34CB9B7110AFFB2FE7FA9251BE39408" author="Goodman S. M. & Raherilalao M. J. & Wohlhauser S." box="[481,713,1556,1580]" pageId="80" pageNumber="185" refId="ref102713" refString="Goodman S. M., Raherilalao M. J., Wohlhauser S. (eds). 2018. Les aires potegees terrestres de Madagascar: leur histoire, description et biote / The terrestrial protected areas of Madagascar: their history, description and biota. Association Vahatra, Antananarivo. 1716 pp." type="book" year="2018">Goodman et al. 2018</bibRefCitation>
|
||
) also known as “humid forest” (
|
||
<bibRefCitation id="E34CB9B7110AFFB2FE2BA9051BBC9468" author="Moat J. & Smith P." box="[437,662,1588,1612]" pageId="80" pageNumber="185" refId="ref103114" refString="Moat J., Smith P. 2007. Atlas of the vegetation of Madagascar. Royal Botanic Garden, Kew 124 pp." type="book" year="2007">Moat & Smith 2007</bibRefCitation>
|
||
). The collecting locality falls within the protected area “Parc National de Masoala”.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |