238 lines
32 KiB
XML
238 lines
32 KiB
XML
<document id="B92066A3008027A24604D941553B14B5" ID-DOI="10.13133/2284-4880/545" ID-ISSN="2284-4880" ID-Zenodo-Dep="12764154" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1721242503917" checkinUser="felipe" docAuthor="Ballerio, Alberto" docDate="2021" docId="0F747550110EFFB7FF09ADA519CB9068" docLanguage="en" docName="FragEntomol.53.2.105-282.pdf" docOrigin="Fragmenta entomologica 53 (2)" docSource="http://dx.doi.org/10.13133/2284-4880/545" docStyle="DocumentStyle:8499FF4308F512386C0E43DC290A16BD.1:FragEntomol.2021-.journal_article.0cover" docStyleId="8499FF4308F512386C0E43DC290A16BD" docStyleName="FragEntomol.2021-.journal_article.0cover" docStyleVersion="1" docTitle="Goudotostes phantasticus Ballerio, 2021, sp. nov." docType="treatment" docUuid="28B3C8F3-E83C-4479-AD62-D5C6C7642AC9" docUuidSource="ZooBank" docVersion="2" lastPageNumber="190" masterDocId="F34D0D28115AFFE2FF9EAF31192A9224" masterDocTitle="The Ceratocanthinae of Madagascar and Comoro Islands: a revision of the genera Synarmostes and Goudotostes, and of the flightless Philharmostes, with description of 64 new species (Coleoptera: Scarabaeoidea, Hybosoridae)" masterLastPageNumber="282" masterPageNumber="105" pageNumber="189" updateTime="1721247602959" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-4.0">
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<mods:title id="1CFAA33F104AD7598FB8D6ED3559A3AE">The Ceratocanthinae of Madagascar and Comoro Islands: a revision of the genera Synarmostes and Goudotostes, and of the flightless Philharmostes, with description of 64 new species (Coleoptera: Scarabaeoidea, Hybosoridae)</mods:title>
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<mods:namePart id="9F17323BA2C4CF59A8236A4EE29DCFF5">Ballerio, Alberto</mods:namePart>
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<mods:title id="9FD939EA04AF3D73814723FD5E8A79F5">Fragmenta entomologica</mods:title>
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<mods:part id="405D11EA386DC91183D73B45F8474AC6">
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<mods:date id="648CFCDBA9ECF3F32B9CCD4D3EAF5622">2021</mods:date>
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<mods:number id="87DE8CC72EBFE69A3931CFF38B881E69">2021-11-30</mods:number>
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<mods:number id="9AEC1B389B3899F8A7FA14AD6DECC6D1">53</mods:number>
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<mods:start id="61A50C53BFA23FEA99DF948897FEB17D">105</mods:start>
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<mods:url id="F9752E51305C74ADE0532D54A0D408EC">http://dx.doi.org/10.13133/2284-4880/545</mods:url>
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<mods:classification id="E0834C9BF9C6B05CEE385405FE257900">journal article</mods:classification>
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<mods:identifier id="6762934B4728396877AA631DF2FDEC84" type="DOI">10.13133/2284-4880/545</mods:identifier>
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<treatment id="0F747550110EFFB7FF09ADA519CB9068" LSID="urn:lsid:zoobank.org:act:28B3C8F3-E83C-4479-AD62-D5C6C7642AC9" httpUri="http://treatment.plazi.org/id/0F747550110EFFB7FF09ADA519CB9068" lastPageId="85" lastPageNumber="190" pageId="84" pageNumber="189">
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<subSubSection id="CFC797CD110EFFB6FF09ADA51B2C9088" box="[151,518,660,684]" pageId="84" pageNumber="189" type="nomenclature">
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<paragraph id="8762C446110EFFB6FF09ADA51B2C9088" blockId="84.[151,518,660,684]" box="[151,518,660,684]" pageId="84" pageNumber="189">
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<heading id="DC2A732A110EFFB6FF09ADA51B2C9088" box="[151,518,660,684]" fontSize="10" level="2" pageId="84" pageNumber="189" reason="5">
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<taxonomicName id="40DDBFC5110EFFB6FF09ADA518809088" ID-CoL="8SSS6" authorityName="Ballerio" authorityYear="2021" box="[151,426,660,684]" class="Insecta" family="Hybosoridae" genus="Goudotostes" kingdom="Animalia" order="Coleoptera" pageId="84" pageNumber="189" phylum="Arthropoda" rank="species" species="phantasticus" status="sp. nov.">
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<emphasis id="B5A91854110EFFB6FF09ADA518809088" box="[151,426,660,684]" italics="true" pageId="84" pageNumber="189">Goudotostes phantasticus</emphasis>
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</taxonomicName>
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<emphasis id="B5A91854110EFFB6FE2FADA51B2C9088" bold="true" box="[433,518,660,684]" pageId="84" pageNumber="189">
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<taxonomicNameLabel id="AE9AA52F110EFFB6FE2FADA51B2C9088" box="[433,518,660,684]" pageId="84" pageNumber="189" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="CFC797CD110EFFB6FF09AD851B349128" pageId="84" pageNumber="189" type="description">
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<paragraph id="8762C446110EFFB6FF09AD851B349128" blockId="84.[151,771,692,780]" pageId="84" pageNumber="189">
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(
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<figureCitation id="1FE6D8C3110EFFB6FF01AD8519DD90E8" box="[159,247,692,716]" captionStart="Fig" captionStartId="10.[151,182,1858,1877]" captionTargetBox="[202,1343,183,1827]" captionTargetId="figure-13@10.[172,1427,160,1833]" captionTargetPageId="10" captionText="Fig. 6 – Posterior view of elytral apex. a, Goudotostes antsahabensis sp. nov.; b, Goudotostes randrianirinai sp. nov.; c, Goudotostes phantasticus sp. nov.; d, Synarmostes sp.; e, Synarmostes sp.; f, Synarmostes separatus sp. nov.; g, Synarmostes tibialis; h, Philharmostes scapulatus sp. nov.; i, Pseudosynarmostes mitsinjo." figureDoi="http://doi.org/10.5281/zenodo.12764176" httpUri="https://zenodo.org/record/12764176/files/figure.png" pageId="84" pageNumber="189">Figs 6 c</figureCitation>
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,
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<figureCitation id="1FE6D8C3110EFFB6FE9CAD85183790E8" box="[258,285,692,716]" captionStart="Fig" captionStartId="137.[136,168,1858,1877]" captionTargetBox="[156,1411,186,1827]" captionTargetId="figure-4@137.[146,1429,177,1849]" captionTargetPageId="137" captionText="Fig. 72 – Goudotostes phantasticus sp. nov., a, paratype, dorsal view of distended specimen; b, paratype, ventral view of rolled up specimen; c, paratype, dorsal view of rolled up specimen; d, paratype, lateral view of rolled up specimen; e, paratype, ventral view of rolled up specimen; f, paratype, pronotum of distended specimen; g–h, holotype, dorsal and ventral view of spiculum gastrale; i–m, holotype, five views of aedeagus." figureDoi="http://doi.org/10.5281/zenodo.12764331" httpUri="https://zenodo.org/record/12764331/files/figure.png" pageId="84" pageNumber="189">72</figureCitation>
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a-m, 77) http://zoobank.org/
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<uri id="F34CC844110EFFB6FEEAADE51B349128" pageId="84" pageNumber="189">
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urn:lsid:zoobank.org:act:
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<uuid id="F37BFE93110EFFB6FD19ADE51B349128" pageId="84" pageNumber="189">28B3C8F3- E83C-4479-AD62-D5C6C7642AC9</uuid>
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</uri>
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</paragraph>
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||
</subSubSection>
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<subSubSection id="CFC797CD110EFFB6FF23AC051BC09688" pageId="84" pageNumber="189" type="materials_examined">
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<paragraph id="8762C446110EFFB6FF23AC051BC09688" blockId="84.[150,772,820,1932]" pageId="84" pageNumber="189">
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<emphasis id="B5A91854110EFFB6FF23AC05186F9168" bold="true" box="[189,325,820,844]" pageId="84" pageNumber="189">Type series.</emphasis>
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<materialsCitation id="37B5CE1B110EFFB6FECEAC051B2F9668" collectingDate="2005-11-28" collectorName="San Francisco, CA & Brian L. Fisher" country="Madagascar" county="Academy of Sciences" determinerName="A. Ballerio" elevation="390" latitude="-17.92389" location="California Acad. of Sciences" longLatPrecision="21" longitude="49.199722" municipality="Reserve Betampona Camp Rendrirendry" pageId="84" pageNumber="189" specimenCount="1" specimenCount-male="1" stateProvince="Toamasina" typeStatus="holotype">
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||
<typeStatus id="58667AE4110EFFB6FECEAC0518999168" box="[336,435,820,844]" pageId="84" pageNumber="189" type="holotype">Holotype</typeStatus>
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,
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<specimenCount id="91DB0FCF110EFFB6FE5CAC0418FD9168" box="[450,471,821,844]" count="1" pageId="84" pageNumber="189" type="male">♂</specimenCount>
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||
, deposited in the collection of the California
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<collectingCounty id="6E03BCCA110EFFB6FED6AC651B079148" box="[328,557,852,876]" pageId="84" pageNumber="189">Academy of Sciences</collectingCounty>
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||
,
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<collectorName id="2A28A190110EFFB6FDA6AC651BD59148" box="[568,767,852,876]" pageId="84" pageNumber="189">San Francisco, CA</collectorName>
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||
, USA, labelled as follows: / CASTYPE 20275 / CASENT 8135435 /
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<collectingCountry id="FFCA84D6110EFFB6FE9AACA518AF9188" box="[260,389,916,940]" name="Madagascar" pageId="84" pageNumber="189">Madagascar</collectingCountry>
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:
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<collectingRegion id="45190AA4110EFFB6FE13ACA51B119188" box="[397,571,916,940]" country="Madagascar" name="Toamasina" pageId="84" pageNumber="189">Toamasina Prov.</collectingRegion>
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,
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<collectingMunicipality id="67065E3C110EFFB6FDDDACA418A291E8" pageId="84" pageNumber="189">Reserve Betampona Camp Rendrirendry</collectingMunicipality>
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||
<quantity id="402569A3110EFFB6FE11AC8518CD91E8" box="[399,487,948,972]" metricMagnitude="4" metricUnit="m" metricValue="3.41" pageId="84" pageNumber="189" unit="km" value="34.1">34.1 km</quantity>
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332°
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||
<collectingRegion id="45190AA4110EFFB6FDB9AC851BBD91E8" box="[551,663,948,972]" country="Madagascar" name="Toamasina" pageId="84" pageNumber="189">Toamasina</collectingRegion>
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el.
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||
<quantity id="402569A3110EFFB6FD21AC851A2991E8" box="[703,771,948,972]" metricMagnitude="2" metricUnit="m" metricValue="3.9" pageId="84" pageNumber="189" unit="m" value="390.0">
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||
<elevation id="0CF02375110EFFB6FD21AC851A2991E8" box="[703,771,948,972]" metricMagnitude="2" metricUnit="m" metricValue="3.9" pageId="84" pageNumber="189" unit="m" value="390.0">390 m</elevation>
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</quantity>
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||
<date id="F363E286110EFFB6FF09ACE5184B91C8" box="[151,353,980,1004]" pageId="84" pageNumber="189" value="2005-11-28">
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||
<collectingDate id="E3271B6E110EFFB6FF09ACE5184B91C8" box="[151,353,980,1004]" pageId="84" pageNumber="189" value="2005-11-28">28 November 2005</collectingDate>
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||
</date>
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||
/
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||
<geoCoordinate id="E2E9A281110EFFB6FEE9ACE51B2D91C8" box="[375,519,980,1004]" degrees="17" direction="south" minutes="55" orientation="latitude" pageId="84" pageNumber="189" precision="15" seconds="26" value="-17.92389">17° 55’ 26” S</geoCoordinate>
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<geoCoordinate id="E2E9A281110EFFB6FD90ACE51B8091C8" box="[526,682,980,1004]" degrees="049" direction="east" minutes="11" orientation="longitude" pageId="84" pageNumber="189" precision="15" seconds="59" value="49.199722">049° 11’ 59” E</geoCoordinate>
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<location id="8202929D110EFFB6FD2CACE5185D9628" LSID="urn:lsid:plazi:treatment:0F747550110EFFB7FF09ADA519CB9068:8202929D110EFFB6FD2CACE5185D9628" country="Madagascar" county="Academy of Sciences" latitude="-17.92389" longLatPrecision="21" longitude="49.199722" municipality="Reserve Betampona Camp Rendrirendry" name="California Acad. of Sciences" pageId="84" pageNumber="189" stateProvince="Toamasina">California Acad. of Sciences</location>
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||
coll.
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||
<collectorName id="2A28A190110EFFB6FE2FACC51B659628" box="[433,591,1012,1036]" pageId="84" pageNumber="189">Brian L. Fisher</collectorName>
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||
et al. sifted litter, rainforest collection code: BLF 13125/
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||
<taxonomicName id="40DDBFC5110EFFB6FDA2AB2519CC9668" authorityName="Ballerio" authorityYear="2021" class="Insecta" family="Hybosoridae" genus="Goudotostes" kingdom="Animalia" order="Coleoptera" pageId="84" pageNumber="189" phylum="Arthropoda" rank="species" species="phantasticus" status="sp. nov.">
|
||
<emphasis id="B5A91854110EFFB6FDA2AB2519CC9668" italics="true" pageId="84" pageNumber="189">Goudotostes phantasticus</emphasis>
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||
</taxonomicName>
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||
<taxonomicNameLabel id="AE9AA52F110EFFB6FF73AB04180F9668" box="[237,293,1077,1100]" pageId="84" pageNumber="189" rank="species">n. sp.</taxonomicNameLabel>
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||
det.
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||
<determinerName id="72BE2882110EFFB6FEC9AB0518E09668" box="[343,458,1076,1100]" pageId="84" pageNumber="189">A. Ballerio</determinerName>
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||
2020
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||
</materialsCitation>
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||
<typeStatus id="58667AE4110EFFB6FD92AB051B529668" box="[524,632,1076,1100]" pageId="84" pageNumber="189" type="holotype">Holotypus</typeStatus>
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<specimenCount id="91DB0FCF110EFFB6FDE0AB041BB89668" box="[638,658,1077,1100]" count="1" pageId="84" pageNumber="189" type="male">♂</specimenCount>
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/ [distend- ed, glued on card, dissected, male genitalia glued in DMHF resin on a separate card, same pin].
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<materialsCitation id="37B5CE1B110EFFB6FD86AB441BCC9688" collectingDate="2005-11-28" collectionCode="CASC, ABCB" collectorName="San Francisco, CA & Brian L. Fisher" country="Madagascar" county="Academy of Sciences" determinerName="A. Ballerio" elevation="390" latitude="-17.92389" location="California Acad. of Sciences" longLatPrecision="21" longitude="49.199722" municipality="Reserve Betampona Camp Rendrirendry" pageId="84" pageNumber="189" specimenCount="6" specimenCount-female="6" stateProvince="Toamasina" typeStatus="paratype">
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||
<typeStatus id="58667AE4110EFFB6FD86AB441BAA96A9" box="[536,640,1141,1165]" pageId="84" pageNumber="189" type="paratype">Paratypes</typeStatus>
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||
:
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||
<specimenCount id="91DB0FCF110EFFB6FD15AB451BE396A8" box="[651,713,1140,1164]" count="6" pageId="84" pageNumber="189" type="female">6 ♀♀</specimenCount>
|
||
(
|
||
<quantity id="402569A3110EFFB6FD47AB451A2E96A8" box="[729,772,1140,1164]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="84" pageNumber="189" unit="in" value="4.0">4 in</quantity>
|
||
<collectionCode id="E1CC5C83110EFFB6FF09ABA519F19688" box="[151,219,1172,1196]" pageId="84" pageNumber="189">CASC</collectionCode>
|
||
and
|
||
<quantity id="402569A3110EFFB6FE93ABA5181E9688" box="[269,308,1172,1196]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="84" pageNumber="189" unit="in" value="2.0">2 in</quantity>
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||
<collectionCode id="E1CC5C83110EFFB6FEA7ABA518AE9688" box="[313,388,1172,1196]" pageId="84" pageNumber="189">ABCB</collectionCode>
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||
), same collecting data as holotype
|
||
</materialsCitation>
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||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="CFC797CD110EFFB6FF23AB851CBF92C8" pageId="84" pageNumber="189" type="diagnosis">
|
||
<paragraph id="8762C446110EFFB6FF23AB851CBF92C8" blockId="84.[150,772,820,1932]" lastBlockId="84.[830,1452,180,1932]" pageId="84" pageNumber="189">
|
||
<emphasis id="B5A91854110EFFB6FF23AB85181896E8" bold="true" box="[189,306,1204,1228]" pageId="84" pageNumber="189">Diagnosis.</emphasis>
|
||
<taxonomicName id="40DDBFC5110EFFB6FEA5AB851B7B96E8" authorityName="Ballerio" authorityYear="2021" box="[315,593,1204,1228]" class="Insecta" family="Hybosoridae" genus="Goudotostes" kingdom="Animalia" order="Coleoptera" pageId="84" pageNumber="189" phylum="Arthropoda" rank="species" species="phantasticus" status="sp. nov.">
|
||
<emphasis id="B5A91854110EFFB6FEA5AB851B7B96E8" box="[315,593,1204,1228]" italics="true" pageId="84" pageNumber="189">Goudotostes phantasticus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B5A91854110EFFB6FDC4AB851B9896E8" bold="true" box="[602,690,1204,1228]" pageId="84" pageNumber="189">
|
||
<taxonomicNameLabel id="AE9AA52F110EFFB6FDC4AB851B9896E8" box="[602,690,1204,1228]" pageId="84" pageNumber="189" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
can be differentiated from all other species of
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||
<taxonomicName id="40DDBFC5110EFFB6FDA8ABE51B9196C8" authorityName=", sensu Ballerio" authorityYear="2008" box="[566,699,1236,1260]" class="Insecta" family="Hybosoridae" genus="Goudotostes" kingdom="Animalia" order="Coleoptera" pageId="84" pageNumber="189" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B5A91854110EFFB6FDA8ABE51B9196C8" box="[566,699,1236,1260]" italics="true" pageId="84" pageNumber="189">Goudotostes</emphasis>
|
||
</taxonomicName>
|
||
by the following combination of characters: a) small size, b) dorsal ocular area absent, c) clypeal apical projection pointed, d) clypeal punctation made of dense impressed large simple punctures, e) pronotal anterior carina monolobate, f) pronotal basal carina monolobate, g) pronotal disc covered by sparse (interpunctural distance about as long as punctural diameter) large simple impressed punctures, h) pronotum with base raised to form a short inverted T-shaped carina (dorsal view), i) elytral dorsal area covered by five (including sutural one) longitudinal continuous blunt carinae occupying about three quarters of elytral length, j) parameres strongly asymmetrical, long less than one third the length of phallobase, as in
|
||
<figureCitation id="1FE6D8C3110EFFB6FE5FA9451B2594A8" box="[449,527,1652,1676]" captionStart="Fig" captionStartId="137.[136,168,1858,1877]" captionTargetBox="[156,1411,186,1827]" captionTargetId="figure-4@137.[146,1429,177,1849]" captionTargetPageId="137" captionText="Fig. 72 – Goudotostes phantasticus sp. nov., a, paratype, dorsal view of distended specimen; b, paratype, ventral view of rolled up specimen; c, paratype, dorsal view of rolled up specimen; d, paratype, lateral view of rolled up specimen; e, paratype, ventral view of rolled up specimen; f, paratype, pronotum of distended specimen; g–h, holotype, dorsal and ventral view of spiculum gastrale; i–m, holotype, five views of aedeagus." figureDoi="http://doi.org/10.5281/zenodo.12764331" httpUri="https://zenodo.org/record/12764331/files/figure.png" pageId="84" pageNumber="189">Fig. 72</figureCitation>
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||
i-m, k) spiculum gastrale with strongly developed ninth hemisternite, shaped as in
|
||
<figureCitation id="1FE6D8C3110EFFB6FF4FA985183494E8" box="[209,286,1716,1740]" captionStart="Fig" captionStartId="137.[136,168,1858,1877]" captionTargetBox="[156,1411,186,1827]" captionTargetId="figure-4@137.[146,1429,177,1849]" captionTargetPageId="137" captionText="Fig. 72 – Goudotostes phantasticus sp. nov., a, paratype, dorsal view of distended specimen; b, paratype, ventral view of rolled up specimen; c, paratype, dorsal view of rolled up specimen; d, paratype, lateral view of rolled up specimen; e, paratype, ventral view of rolled up specimen; f, paratype, pronotum of distended specimen; g–h, holotype, dorsal and ventral view of spiculum gastrale; i–m, holotype, five views of aedeagus." figureDoi="http://doi.org/10.5281/zenodo.12764331" httpUri="https://zenodo.org/record/12764331/files/figure.png" pageId="84" pageNumber="189">Fig. 72</figureCitation>
|
||
g-h, l) apical spur of protibiae very long and thin, distinctly longer than in all other known
|
||
<taxonomicName id="40DDBFC5110EFFB6FD0CA9E5199F9528" authorityName=", sensu Ballerio" authorityYear="2008" class="Insecta" family="Hybosoridae" genus="Goudotostes" kingdom="Animalia" order="Coleoptera" pageId="84" pageNumber="189" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B5A91854110EFFB6FD0CA9E5199F9528" italics="true" pageId="84" pageNumber="189">Goudotostes</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<taxonomicName id="40DDBFC5110EFFB6FF5FA9C518FE9528" authorityName="Ballerio" authorityYear="2021" box="[193,468,1780,1804]" class="Insecta" family="Hybosoridae" genus="Goudotostes" kingdom="Animalia" order="Coleoptera" pageId="84" pageNumber="189" phylum="Arthropoda" rank="species" species="phantasticus" status="sp. nov.">
|
||
<emphasis id="B5A91854110EFFB6FF5FA9C518FE9528" box="[193,468,1780,1804]" italics="true" pageId="84" pageNumber="189">Goudotostes phantasticus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B5A91854110EFFB6FE47A9C51B049528" bold="true" box="[473,558,1780,1804]" pageId="84" pageNumber="189">
|
||
<taxonomicNameLabel id="AE9AA52F110EFFB6FE47A9C51B049528" box="[473,558,1780,1804]" pageId="84" pageNumber="189" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
because of its small size could be mistaken for a species belonging to the
|
||
<taxonomicName id="40DDBFC5110EFFB6FD5BA8251A299508" authorityName="Ballerio" authorityYear="2021" box="[709,771,1812,1836]" class="Insecta" family="Hybosoridae" genus="Goudotostes" kingdom="Animalia" order="Coleoptera" pageId="84" pageNumber="189" phylum="Arthropoda" rank="species" species="laevis">
|
||
<emphasis id="B5A91854110EFFB6FD5BA8251A299508" box="[709,771,1812,1836]" italics="true" pageId="84" pageNumber="189">laevis</emphasis>
|
||
</taxonomicName>
|
||
group of species, however, due to the monolobate pronotal basal and anterior carinae, to the presence of four basal elytral carinae and to the parameres lacking the distinctive invagination found in the
|
||
<taxonomicName id="40DDBFC5110EFFB6FBD1AF851DA792E8" authorityName="Ballerio" authorityYear="2021" box="[1103,1165,180,204]" class="Insecta" family="Hybosoridae" genus="Goudotostes" kingdom="Animalia" order="Coleoptera" pageId="84" pageNumber="189" phylum="Arthropoda" rank="species" species="laevis">
|
||
<emphasis id="B5A91854110EFFB6FBD1AF851DA792E8" box="[1103,1165,180,204]" italics="true" pageId="84" pageNumber="189">laevis</emphasis>
|
||
</taxonomicName>
|
||
group, it represents an isolate species, whose closest relationships remain unclear.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="CFC797CD110EFFB7FCFBAFC51BB29348" lastPageId="85" lastPageNumber="190" pageId="84" pageNumber="189" type="description">
|
||
<paragraph id="8762C446110EFFB7FCFBAFC51BB29348" blockId="84.[830,1452,180,1932]" lastBlockId="85.[135,757,180,588]" lastPageId="85" lastPageNumber="190" pageId="84" pageNumber="189">
|
||
<emphasis id="B5A91854110EFFB6FCFBAFC51AC59328" bold="true" box="[869,1007,244,268]" pageId="84" pageNumber="189">Description.</emphasis>
|
||
Size: HL =
|
||
<quantity id="402569A3110EFFB6FBE7AFC51DFA9328" box="[1145,1232,244,268]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="84" pageNumber="189" unit="mm" value="0.7">0.7 mm</quantity>
|
||
; HW =
|
||
<quantity id="402569A3110EFFB6FAB4AFC51CAB9328" box="[1322,1409,244,268]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="84" pageNumber="189" unit="mm" value="1.3">1.3 mm</quantity>
|
||
; PL =
|
||
<quantity id="402569A3110EFFB6FCC9AE251A849308" box="[855,942,276,300]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="84" pageNumber="189" unit="mm" value="1.2">1.2 mm</quantity>
|
||
; PW =
|
||
<quantity id="402569A3110EFFB6FB9CAE251D739308" box="[1026,1113,276,300]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" pageId="84" pageNumber="189" unit="mm" value="2.1">2.1 mm</quantity>
|
||
; EL =
|
||
<quantity id="402569A3110EFFB6FB3BAE251DD69308" box="[1189,1276,276,300]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" pageId="84" pageNumber="189" unit="mm" value="2.2">2.2 mm</quantity>
|
||
; EW =
|
||
<quantity id="402569A3110EFFB6FACCAE251C829308" box="[1362,1448,276,300]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" pageId="84" pageNumber="189" unit="mm" value="2.1">2.1 mm</quantity>
|
||
. Overall morphology as in generic diagnosis. Small-sized
|
||
<taxonomicName id="40DDBFC5110EFFB6FCA1AE651AE99348" authorityName=", sensu Ballerio" authorityYear="2008" box="[831,963,340,364]" class="Insecta" family="Hybosoridae" genus="Goudotostes" kingdom="Animalia" order="Coleoptera" pageId="84" pageNumber="189" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B5A91854110EFFB6FCA1AE651AE99348" box="[831,963,340,364]" italics="true" pageId="84" pageNumber="189">Goudotostes</emphasis>
|
||
</taxonomicName>
|
||
. Body strongly convex. Dark brown with bronze sheen especially on carinae, shiny, dorsum with extremely short fine setation (50×), underside, tarsi and antennae dark reddish-brown, setation yellowish-brown.
|
||
<emphasis id="B5A91854110EFFB6FCA1AEE51A5293C8" box="[831,888,468,492]" italics="true" pageId="84" pageNumber="189">Head</emphasis>
|
||
: W/L ratio = 1.79; clypeal apex obtuse, apical projection strongly developed and sexually dimorphic, sides relatively rounded with slightly unaligned genae; dorsal ocular area absent, genal canthus indistinct, ventral ocular area small-sized, genae acute, distinctly protruding outwards, genal suture indistinct; head dorsal surface covered by large impressed simple punctures on vertex becoming transversely comma-shaped on frons and clypeus (clearly as a result of the fusion of two or more simple punctures). Interpunctural distance equal to or inferior than punctural diameter. Frons bearing a longitudinal irregular swelling extending to the proximal portion of clypeus.
|
||
<emphasis id="B5A91854110EFFB6FAA5AC041C8E9168" box="[1339,1444,821,844]" italics="true" pageId="84" pageNumber="189">Pronotum</emphasis>
|
||
: subrectangular (dorsal view) (W/L ratio = 1.87), convex, fore angles broadly subtruncate; pronotal disc covered by short large impressed simple punctures becoming shallow- er and horseshoe-shaped with opening directed outwards punctures at sides and base, each one bearing a fine setigerous pore inside, interpunctural distance about equal to punctural diameter; pronotal bead present only along fore margin; pronotal base distinctly raised in correspondence with scutellum; fore margin distinctly raised in correspondence with head insertion; pronotal sculpturing made of large and raised tubercles and carinae as follows: monolobate basal carina smooth inverted T-shaped in dorsal view, with branches of T being longer than the main axis, base of disc with two raised tubercles, each side with two raised transverse carinae, monolobate anterior carina distinctly raised to form one small weakly raised tubercle with a coarse U- or V-shaped apex (frontal view).
|
||
<taxonomicName id="40DDBFC5110EFFB6FA99AA651C459748" box="[1287,1391,1364,1388]" class="Trilobita" family="Styginidae" genus="Scutellum" kingdom="Animalia" order="Corynexochida" pageId="84" pageNumber="189" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B5A91854110EFFB6FA99AA651C459748" box="[1287,1391,1364,1388]" italics="true" pageId="84" pageNumber="189">Scutellum</emphasis>
|
||
</taxonomicName>
|
||
: base smooth, median area covered by impressed transverse short comma-shaped punctures, apical third and sides smooth.
|
||
<emphasis id="B5A91854110EFFB6FCA1AA851AAB97E8" box="[831,897,1460,1484]" italics="true" pageId="84" pageNumber="189">Elytra</emphasis>
|
||
: longer than wide (W/L ratio = 0.95), subovoidal (dorsal view), strongly convex; humeral callus marked by a tubercle; elytral surface uniformly covered by small narrow horseshoe-shaped punctures with opening directed backwards and having a setigerous pore inside, interpunctural distance about equal to punctural diameter; inferior sutural stria weakly developed, striated articular area hardly visible in lateral view, relatively narrow and short, marginal area medium-sized; elytral articular process well developed, smooth and shiny. Elytral sculpturing as follows: base with two strongly raised large oblique carinae near apex of scutellum, next to them a humeral oblique longer carina; elytral suture marked by a longitudinal line almost reaching the scutellum, elytral dorsally with five longitudinal continuous carinae occupying about three quarters of elytral length, elytral sides with irregular rows of short longitudinal carinae. Protibiae with apical spur thin and distinctly longer than in all other
|
||
<taxonomicName id="40DDBFC5110FFFB7FE37AFC51B079328" authorityName=", sensu Ballerio" authorityYear="2008" box="[425,557,244,268]" class="Insecta" family="Hybosoridae" genus="Goudotostes" kingdom="Animalia" order="Coleoptera" pageId="85" pageNumber="190" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B5A91854110FFFB7FE37AFC51B079328" box="[425,557,244,268]" italics="true" pageId="85" pageNumber="190">Goudotostes</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<emphasis id="B5A91854110FFFB7FDA3AFC41B549328" box="[573,638,245,268]" italics="true" pageId="85" pageNumber="190">Wings</emphasis>
|
||
: flightless.
|
||
<emphasis id="B5A91854110FFFB7FF16AE25180E9308" box="[136,292,276,300]" italics="true" pageId="85" pageNumber="190">Male genitalia</emphasis>
|
||
: aedeagus (L =
|
||
<quantity id="402569A3110FFFB7FE4FAE251B0F9308" box="[465,549,276,300]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="85" pageNumber="190" unit="mm" value="1.4">1.4 mm</quantity>
|
||
) with parameres as in
|
||
<figureCitation id="1FE6D8C3110FFFB7FF3AAE0519DF9368" box="[164,245,308,332]" captionStart="Fig" captionStartId="137.[136,168,1858,1877]" captionTargetBox="[156,1411,186,1827]" captionTargetId="figure-4@137.[146,1429,177,1849]" captionTargetPageId="137" captionText="Fig. 72 – Goudotostes phantasticus sp. nov., a, paratype, dorsal view of distended specimen; b, paratype, ventral view of rolled up specimen; c, paratype, dorsal view of rolled up specimen; d, paratype, lateral view of rolled up specimen; e, paratype, ventral view of rolled up specimen; f, paratype, pronotum of distended specimen; g–h, holotype, dorsal and ventral view of spiculum gastrale; i–m, holotype, five views of aedeagus." figureDoi="http://doi.org/10.5281/zenodo.12764331" httpUri="https://zenodo.org/record/12764331/files/figure.png" pageId="85" pageNumber="190">Figs 72</figureCitation>
|
||
i-m, spiculum gastrale with strongly developed ninth hemisternite (L =
|
||
<quantity id="402569A3110FFFB7FE1FAE6518FE9348" box="[385,468,340,364]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" pageId="85" pageNumber="190" unit="mm" value="1.6">1.6 mm</quantity>
|
||
) as in
|
||
<figureCitation id="1FE6D8C3110FFFB7FD87AE651B429348" box="[537,616,340,364]" captionStart="Fig" captionStartId="137.[136,168,1858,1877]" captionTargetBox="[156,1411,186,1827]" captionTargetId="figure-4@137.[146,1429,177,1849]" captionTargetPageId="137" captionText="Fig. 72 – Goudotostes phantasticus sp. nov., a, paratype, dorsal view of distended specimen; b, paratype, ventral view of rolled up specimen; c, paratype, dorsal view of rolled up specimen; d, paratype, lateral view of rolled up specimen; e, paratype, ventral view of rolled up specimen; f, paratype, pronotum of distended specimen; g–h, holotype, dorsal and ventral view of spiculum gastrale; i–m, holotype, five views of aedeagus." figureDoi="http://doi.org/10.5281/zenodo.12764331" httpUri="https://zenodo.org/record/12764331/files/figure.png" pageId="85" pageNumber="190">Figs 72</figureCitation>
|
||
g-h.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="CFC797CD110FFFB7FF30AE451B1693E8" pageId="85" pageNumber="190" type="etymology">
|
||
<paragraph id="8762C446110FFFB7FF30AE451B1693E8" blockId="85.[135,757,180,588]" pageId="85" pageNumber="190">
|
||
<emphasis id="B5A91854110FFFB7FF30AE45180493A8" bold="true" box="[174,302,372,396]" pageId="85" pageNumber="190">Etymology.</emphasis>
|
||
Latin adjective in the nominative singular, meaning “amazing” or “admirable”, due to the extremely unusual and beautiful dorsal sculpturing.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="CFC797CD110FFFB7FF30AEE519CB9068" pageId="85" pageNumber="190" type="distribution">
|
||
<paragraph id="8762C446110FFFB7FF30AEE519CB9068" blockId="85.[135,757,180,588]" pageId="85" pageNumber="190">
|
||
<emphasis id="B5A91854110FFFB7FF30AEE518E293C8" bold="true" box="[174,456,468,492]" pageId="85" pageNumber="190">Distribution and habitat.</emphasis>
|
||
Known only from the
|
||
<typeStatus id="58667AE4110FFFB7FD59AEE41BDE93C9" box="[711,756,469,493]" pageId="85" pageNumber="190">type</typeStatus>
|
||
locality. Collected by sifting litter in rainforest. For details on the collecting localities see under
|
||
<taxonomicName id="40DDBFC5110FFFB7FE62AD241BD99008" authorityName="Ballerio" authorityYear="2021" box="[508,755,533,556]" class="Insecta" family="Hybosoridae" genus="Synarmostes" kingdom="Animalia" order="Coleoptera" pageId="85" pageNumber="190" phylum="Arthropoda" rank="species" species="compactus" status="sp. nov.">
|
||
<emphasis id="B5A91854110FFFB7FE62AD241BD99008" box="[508,755,533,556]" italics="true" pageId="85" pageNumber="190">Synarmostes compactus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B5A91854110FFFB7FF19AD0519F39068" bold="true" box="[135,217,564,588]" pageId="85" pageNumber="190">
|
||
<taxonomicNameLabel id="AE9AA52F110FFFB7FF19AD0519F39068" box="[135,217,564,588]" pageId="85" pageNumber="190" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |