313 lines
38 KiB
XML
313 lines
38 KiB
XML
<document id="84936B39166FAB4AD83605F41ECF92C6" ID-DOI="10.13133/2284-4880/545" ID-ISSN="2284-4880" ID-Zenodo-Dep="12764154" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1721242503917" checkinUser="felipe" docAuthor="Ballerio, Alberto" docDate="2021" docId="0F747550110DFFBAFF16ACC5182C9688" docLanguage="en" docName="FragEntomol.53.2.105-282.pdf" docOrigin="Fragmenta entomologica 53 (2)" docSource="http://dx.doi.org/10.13133/2284-4880/545" docStyle="DocumentStyle:8499FF4308F512386C0E43DC290A16BD.1:FragEntomol.2021-.journal_article.0cover" docStyleId="8499FF4308F512386C0E43DC290A16BD" docStyleName="FragEntomol.2021-.journal_article.0cover" docStyleVersion="1" docTitle="Goudotostes rajemisonae Ballerio, 2021, sp. nov." docType="treatment" docUuid="60D90FF0-2D8D-44F0-BB37-3F924E9836C2" docUuidSource="ZooBank" docVersion="2" lastPageNumber="193" masterDocId="F34D0D28115AFFE2FF9EAF31192A9224" masterDocTitle="The Ceratocanthinae of Madagascar and Comoro Islands: a revision of the genera Synarmostes and Goudotostes, and of the flightless Philharmostes, with description of 64 new species (Coleoptera: Scarabaeoidea, Hybosoridae)" masterLastPageNumber="282" masterPageNumber="105" pageNumber="192" updateTime="1721247602959" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-4.0">
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<mods:mods id="5ABD7D8854CD2D6C0A0BA9CD1915264A" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="5D33E83BA67FA49B701CABE6A0F21571">
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<mods:title id="D07904E4F2825F3C68F43AB52EE27FAE">The Ceratocanthinae of Madagascar and Comoro Islands: a revision of the genera Synarmostes and Goudotostes, and of the flightless Philharmostes, with description of 64 new species (Coleoptera: Scarabaeoidea, Hybosoridae)</mods:title>
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</mods:titleInfo>
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<mods:name id="E092689B150543DDCF19BDA94FD3F548" type="personal">
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<mods:role id="95507A1B66D904549053F92785C5A4C2">
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<mods:roleTerm id="FB4057CDC2A72A46C17176A73EC89D71">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="0688E46C912C5366680B794A8CA85823">Ballerio, Alberto</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="3F632E4F99D5B2EA2213B87C7FD429F9">text</mods:typeOfResource>
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<mods:titleInfo id="76B8EA31257EB08F2DDE322E08088C9F">
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<mods:title id="666A06244C4CC9C38DCA2F93D0F2615E">Fragmenta entomologica</mods:title>
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</mods:titleInfo>
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<mods:part id="FFFEF932FBC78938748BFE36A41BB692">
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<mods:date id="3D8487FF7E2FF04C4F626CADE3E05CF0">2021</mods:date>
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<mods:detail id="98C5216D85C8845E183B145A9658EC23" type="pubDate">
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<mods:number id="DDB6587D49F1A10723D1445A83EA32DE">2021-11-30</mods:number>
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</mods:detail>
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<mods:detail id="749B35A16D4960E24CA82C62B6BFA767" type="volume">
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<mods:number id="77B622B1E87473A8EADAC4515FC79229">53</mods:number>
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</mods:detail>
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<mods:detail id="C2A5A204398B87B146FEEE992E78AED4" type="issue">
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<mods:number id="0BD2D65D6460E419FCAA3CD4F8942C0B">2</mods:number>
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</mods:detail>
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<mods:extent id="02971EB3E407701F0F528F29277E44CA" unit="page">
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<mods:start id="5EF6AE048A52AB844A83804CC10329DB">105</mods:start>
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<mods:end id="CA0D14B06E420AEDE4C231450996951B">282</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location id="889E26187686C29F428B9142AE582B6A">
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<mods:url id="DB548277C197490F2536AF2A1D87C578">http://dx.doi.org/10.13133/2284-4880/545</mods:url>
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</mods:location>
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<mods:classification id="1AA952332907C22587C2FBAB341AD2D3">journal article</mods:classification>
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<mods:identifier id="AE876D21620D131FE9C8A3F8AF9A4DE3" type="DOI">10.13133/2284-4880/545</mods:identifier>
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<mods:identifier id="240F3D6DC96A9683CF195685E39B22BF" type="ISSN">2284-4880</mods:identifier>
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<mods:identifier id="CE1E2B29BD2E4684C8C9D4550A547E04" type="Zenodo-Dep">12764154</mods:identifier>
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<mods:identifier id="8F7E42BE8301BBBE72ACAACA28343418" type="ZooBank">B3F75A2C-7603-4027-B8E1-F4A8B5A0CDD9</mods:identifier>
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</mods:mods>
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<treatment id="0F747550110DFFBAFF16ACC5182C9688" LSID="urn:lsid:zoobank.org:act:60D90FF0-2D8D-44F0-BB37-3F924E9836C2" httpUri="http://treatment.plazi.org/id/0F747550110DFFBAFF16ACC5182C9688" lastPageId="88" lastPageNumber="193" pageId="87" pageNumber="192">
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<subSubSection id="CFC797CD110DFFB5FF16ACC518DE9628" box="[136,500,1012,1036]" pageId="87" pageNumber="192" type="nomenclature">
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<paragraph id="8762C446110DFFB5FF16ACC518DE9628" blockId="87.[136,500,1012,1036]" box="[136,500,1012,1036]" pageId="87" pageNumber="192">
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<heading id="DC2A732A110DFFB5FF16ACC518DE9628" box="[136,500,1012,1036]" fontSize="10" level="2" pageId="87" pageNumber="192" reason="5">
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<taxonomicName id="40DDBFC5110DFFB5FF16ACC518B29628" ID-CoL="8SSRR" authorityName="Ballerio" authorityYear="2021" box="[136,408,1012,1036]" class="Insecta" family="Hybosoridae" genus="Goudotostes" kingdom="Animalia" order="Coleoptera" pageId="87" pageNumber="192" phylum="Arthropoda" rank="species" species="rajemisonae" status="sp. nov.">
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<emphasis id="B5A91854110DFFB5FF16ACC518B29628" box="[136,408,1012,1036]" italics="true" pageId="87" pageNumber="192">Goudotostes rajemisonae</emphasis>
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</taxonomicName>
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<emphasis id="B5A91854110DFFB5FE01ACC518DE9628" bold="true" box="[415,500,1012,1036]" pageId="87" pageNumber="192">
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||
<taxonomicNameLabel id="AE9AA52F110DFFB5FE01ACC518DE9628" box="[415,500,1012,1036]" pageId="87" pageNumber="192" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="CFC797CD110DFFB5FF16AB251B289648" pageId="87" pageNumber="192" type="description">
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||
<paragraph id="8762C446110DFFB5FF16AB251B289648" blockId="87.[136,755,1044,1132]" pageId="87" pageNumber="192">
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||
(
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<figureCitation id="1FE6D8C3110DFFB5FF0EAB2519F99608" box="[144,211,1044,1068]" captionStart="Fig" captionStartId="12.[151,182,1858,1877]" captionTargetBox="[151,1451,166,1830]" captionTargetId="figure-13@12.[148,1454,163,1834]" captionTargetPageId="12" captionText="Fig. 8 – a, Cryptosphaeroides hystrix, antenna (ped: pedicellus); b, Synarmostes benaraensis sp. nov., antenna; c, Pseudosynarmostes mitsinjo, antenna, antenna; d, Goudotostes mayottensis sp. nov.; e, Philharmostes scapulatus sp. nov., antenna; f, Goudotostes rajemisonae sp. nov., aedeagus showing invaginated left paramere (pi: paramere invagination); g, Goudotostes rajemisonae sp. nov., detail of circled area of Fig. f showing sensilla." figureDoi="http://doi.org/10.5281/zenodo.12764180" httpUri="https://zenodo.org/record/12764180/files/figure.png" pageId="87" pageNumber="192">Figs 8</figureCitation>
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f-g, 67 a-j, 77) http://zoobank.org/
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<uri id="F34CC844110DFFB5FEFCAB051B289648" pageId="87" pageNumber="192">
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||
urn:lsid:zoobank.org:act:
|
||
<uuid id="F37BFE93110DFFB5FDEEAB051B289648" pageId="87" pageNumber="192">60D90FF0- 2D8D-44F0-BB37-3F924E9836C2</uuid>
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||
</uri>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="CFC797CD110DFFB5FF30ABA518659528" pageId="87" pageNumber="192" type="materials_examined">
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<paragraph id="8762C446110DFFB5FF30ABA518659528" blockId="87.[136,758,1172,1932]" pageId="87" pageNumber="192">
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<emphasis id="B5A91854110DFFB5FF30ABA5181C9688" bold="true" box="[174,310,1172,1196]" pageId="87" pageNumber="192">Type series.</emphasis>
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<materialsCitation id="37B5CE1B110DFFB5FEA3ABA5185397E8" collectingDate="2003-03-08" collectingDateMax="2003-03-12" collectingDateMin="2003-03-08" collectorName="San Francisco, CA & Fisher & Griswold" country="Madagascar" county="Academy of Sciences" determinerName="A. Ballerio" elevation="470" latitude="-15.1883335" location="Ambinanitelo" longLatPrecision="21" longitude="49.614998" municipality="Montagne d'Anjanaharibe" pageId="87" pageNumber="192" specimenCount="1" specimenCount-male="1" stateProvince="Toamasina" typeStatus="holotype">
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||
<typeStatus id="58667AE4110DFFB5FEA3ABA5188F9688" box="[317,421,1172,1196]" pageId="87" pageNumber="192" type="holotype">Holotype</typeStatus>
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||
,
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<specimenCount id="91DB0FCF110DFFB5FE2FABA418ED9688" box="[433,455,1173,1196]" count="1" pageId="87" pageNumber="192" type="male">♂</specimenCount>
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, deposited in the collection of the California
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<collectingCounty id="6E03BCCA110DFFB5FED3AB851B6896E8" box="[333,578,1204,1228]" pageId="87" pageNumber="192">Academy of Sciences</collectingCounty>
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||
,
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<collectorName id="2A28A190110DFFB5FDCEAB85199A96C8" pageId="87" pageNumber="192">San Francisco, CA</collectorName>
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||
, USA, labelled as follows: / CASTYPE 20278
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<emphasis id="B5A91854110DFFB5FD25ABE51BE896C8" bold="true" box="[699,706,1236,1260]" pageId="87" pageNumber="192">/</emphasis>
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||
HT: CASENT 8058272 /
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<collectingCountry id="FFCA84D6110DFFB5FEEDABC518D19728" box="[371,507,1268,1292]" name="Madagascar" pageId="87" pageNumber="192">Madagascar</collectingCountry>
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||
:
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<collectingRegion id="45190AA4110DFFB5FD99ABC51B569728" box="[519,636,1268,1292]" country="Madagascar" name="Toamasina" pageId="87" pageNumber="192">Toamasina</collectingRegion>
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<collectingMunicipality id="67065E3C110DFFB5FD18ABC4181B9708" pageId="87" pageNumber="192">Montagne d’Anjanaharibe</collectingMunicipality>
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||
18.0 km 21° NNE
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||
<location id="8202929D110DFFB5FD81AA251B9F9708" LSID="urn:lsid:plazi:treatment:0F747550110DFFBAFF16ACC5182C9688:8202929D110DFFB5FD81AA251B9F9708" box="[543,693,1300,1324]" country="Madagascar" county="Academy of Sciences" latitude="-15.1883335" longLatPrecision="21" longitude="49.614998" municipality="Montagne d'Anjanaharibe" name="Ambinanitelo" pageId="87" pageNumber="192" stateProvince="Toamasina">Ambinanitelo</location>
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elev.
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||
<quantity id="402569A3110DFFB5FF16AA0519FA9768" box="[136,208,1332,1356]" metricMagnitude="2" metricUnit="m" metricValue="4.7" pageId="87" pageNumber="192" unit="m" value="470.0">
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||
<elevation id="0CF02375110DFFB5FF16AA0519FA9768" box="[136,208,1332,1356]" metricMagnitude="2" metricUnit="m" metricValue="4.7" pageId="87" pageNumber="192" unit="m" value="470.0">470 m</elevation>
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||
</quantity>
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||
<date id="F363E286110DFFB5FF44AA0518A29768" box="[218,392,1332,1356]" pageId="87" pageNumber="192" value="2003-03-08" valueMax="2003-03-12" valueMin="2003-03-08">
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||
<collectingDate id="E3271B6E110DFFB5FF44AA0518A29768" box="[218,392,1332,1356]" pageId="87" pageNumber="192" value="2003-03-08" valueMax="2003-03-12" valueMin="2003-03-08">8-12 Mar. 2003</collectingDate>
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||
</date>
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||
<geoCoordinate id="E2E9A281110DFFB5FE0CAA051B049768" box="[402,558,1332,1356]" degrees="15" direction="south" minutes="11" orientation="latitude" pageId="87" pageNumber="192" precision="15" seconds="18" value="-15.1883335">15° 11’ 18” S</geoCoordinate>
|
||
<geoCoordinate id="E2E9A281110DFFB5FDA6AA051BC89768" box="[568,738,1332,1356]" degrees="049" direction="east" minutes="36" orientation="longitude" pageId="87" pageNumber="192" precision="15" seconds="54" value="49.614998">049° 36’ 54” E</geoCoordinate>
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||
/
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||
<location id="8202929D110DFFB5FF16AA6518E89748" LSID="urn:lsid:plazi:treatment:0F747550110DFFBAFF16ACC5182C9688:8202929D110DFFB5FF16AA6518E89748" box="[136,450,1364,1388]" country="Madagascar" county="Academy of Sciences" latitude="-15.1883335" longLatPrecision="21" longitude="49.614998" municipality="Montagne d'Anjanaharibe" name="California Acad. of Sciences" pageId="87" pageNumber="192" stateProvince="Toamasina">California Acad. of Sciences</location>
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||
coll.
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||
<collectorName id="2A28A190110DFFB5FD9FAA651B6D9748" box="[513,583,1364,1388]" pageId="87" pageNumber="192">Fisher</collectorName>
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||
,
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||
<collectorName id="2A28A190110DFFB5FDCCAA651B9D9748" box="[594,695,1364,1388]" pageId="87" pageNumber="192">Griswold</collectorName>
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et al. sifted litter (leaf mold, rotten wood) in rainforest collection code: BLF 8002/
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||
<taxonomicName id="40DDBFC5110DFFB5FE13AAA51B839788" authorityName="Ballerio" authorityYear="2021" box="[397,681,1428,1452]" class="Insecta" family="Hybosoridae" genus="Goudotostes" kingdom="Animalia" order="Coleoptera" pageId="87" pageNumber="192" phylum="Arthropoda" rank="species" species="rajemisonae" status="sp. nov.">
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||
<emphasis id="B5A91854110DFFB5FE13AAA51B839788" box="[397,681,1428,1452]" italics="true" pageId="87" pageNumber="192">Goudotostes rajemisonae</emphasis>
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||
</taxonomicName>
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||
<taxonomicNameLabel id="AE9AA52F110DFFB5FD2AAAA41BDE9788" box="[692,756,1429,1452]" pageId="87" pageNumber="192" rank="species">n. sp.</taxonomicNameLabel>
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||
det.
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||
<determinerName id="72BE2882110DFFB5FF24AA85181297E8" box="[186,312,1460,1484]" pageId="87" pageNumber="192">A. Ballerio</determinerName>
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||
2020
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||
</materialsCitation>
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||
<typeStatus id="58667AE4110DFFB5FE1BAA8518DC97E8" box="[389,502,1460,1484]" pageId="87" pageNumber="192" type="holotype">Holotypus</typeStatus>
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||
<specimenCount id="91DB0FCF110DFFB5FD9FAA841B3F97E8" box="[513,533,1461,1484]" count="1" pageId="87" pageNumber="192" type="male">♂</specimenCount>
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/ [distended, glued on card, dissected, male genitalia glued in DMHF resin on a separate card, same pin].
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<materialsCitation id="37B5CE1B110DFFB5FE43AAC418139528" collectingDate="2003-03-17" collectingDateMax="2003-03-21" collectingDateMin="2003-03-17" collectionCode="CASC, ABCB" collectorName="Fisher & Griswold" country="Madagascar" county="Academy of Sciences" determinerName="A. Ballerio" elevation="600" latitude="-15.288334" location="California Acad. of Sciences" longLatPrecision="21" longitude="49.548332" municipality="Montagne d'Akirindro" pageId="87" pageNumber="192" specimenCount="27" specimenCount-female="12" specimenCount-male="15" stateProvince="Toamasina" typeStatus="paratype">
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||
<typeStatus id="58667AE4110DFFB5FE43AAC41B6D9429" box="[477,583,1525,1549]" pageId="87" pageNumber="192" type="paratype">Paratypes</typeStatus>
|
||
[
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||
<specimenCount id="91DB0FCF110DFFB5FDC6AAC51BBD9428" box="[600,663,1524,1548]" count="3" pageId="87" pageNumber="192" type="male">3 ♂♂</specimenCount>
|
||
dissect- ed]:
|
||
<specimenCount id="91DB0FCF110DFFB5FF22A92519CC9408" box="[188,230,1556,1580]" count="1" pageId="87" pageNumber="192" type="male">1 ♂</specimenCount>
|
||
and
|
||
<specimenCount id="91DB0FCF110DFFB5FEBEA92518749408" box="[288,350,1556,1580]" count="2" pageId="87" pageNumber="192" type="female">2 ♀♀</specimenCount>
|
||
(
|
||
<quantity id="402569A3110DFFB5FEEEA92518B79408" box="[368,413,1556,1580]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="87" pageNumber="192" unit="in" value="2.0">2 in</quantity>
|
||
<collectionCode id="E1CC5C83110DFFB5FE38A92518C69408" box="[422,492,1556,1580]" pageId="87" pageNumber="192">CASC</collectionCode>
|
||
and
|
||
<quantity id="402569A3110DFFB5FDB8A9251B789408" box="[550,594,1556,1580]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="87" pageNumber="192" unit="in" value="1.0">1 in</quantity>
|
||
<collectionCode id="E1CC5C83110DFFB5FDC4A9251B829408" box="[602,680,1556,1580]" pageId="87" pageNumber="192">ABCB</collectionCode>
|
||
), same collecting data as holotype.
|
||
<specimenCount id="91DB0FCF110DFFB5FE55A9051B329468" box="[459,536,1588,1612]" count="11" pageId="87" pageNumber="192" type="male">11 ♂♂</specimenCount>
|
||
and
|
||
<specimenCount id="91DB0FCF110DFFB5FDC8A9051B8E9468" box="[598,676,1588,1612]" count="10" pageId="87" pageNumber="192" type="female">10 ♀♀</specimenCount>
|
||
(
|
||
<quantity id="402569A3110DFFB5FD26A9051BDE9468" box="[696,756,1588,1612]" metricMagnitude="-1" metricUnit="m" metricValue="4.064" pageId="87" pageNumber="192" unit="in" value="16.0">16 in</quantity>
|
||
<collectionCode id="E1CC5C83110DFFB5FF16A96519E49448" box="[136,206,1620,1644]" pageId="87" pageNumber="192">CASC</collectionCode>
|
||
and
|
||
<quantity id="402569A3110DFFB5FE98A965181A9448" box="[262,304,1620,1644]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="87" pageNumber="192" unit="in" value="5.0">5 in</quantity>
|
||
<collectionCode id="E1CC5C83110DFFB5FEA9A96518AC9448" box="[311,390,1620,1644]" pageId="87" pageNumber="192">ABCB</collectionCode>
|
||
): /
|
||
<collectingCountry id="FFCA84D6110DFFB5FE36A9651B1A9448" box="[424,560,1620,1644]" name="Madagascar" pageId="87" pageNumber="192">Madagascar</collectingCountry>
|
||
:
|
||
<collectingRegion id="45190AA4110DFFB5FDA5A9651B9A9448" box="[571,688,1620,1644]" country="Madagascar" name="Toamasina" pageId="87" pageNumber="192">Toamasina</collectingRegion>
|
||
<collectingMunicipality id="67065E3C110DFFB5FD26A964186694A8" pageId="87" pageNumber="192">Montagne d’Akirindro</collectingMunicipality>
|
||
<locationDeviation id="96D0DD64110DFFB5FECDA9451B9194A8" box="[339,699,1652,1676]" location="Ambinanitelo" pageId="87" pageNumber="192">
|
||
<quantity id="402569A3110DFFB5FECDA94518B494A8" box="[339,414,1652,1676]" metricMagnitude="3" metricUnit="m" metricValue="7.6" pageId="87" pageNumber="192" unit="km" value="7.6">7.6 km</quantity>
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||
341° NNW Ambinanitelo
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||
</locationDeviation>
|
||
elev.
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||
<quantity id="402569A3110DFFB5FF16A9A519E49488" box="[136,206,1684,1708]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="87" pageNumber="192" unit="m" value="600.0">
|
||
<elevation id="0CF02375110DFFB5FF16A9A519E49488" box="[136,206,1684,1708]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="87" pageNumber="192" unit="m" value="600.0">600 m</elevation>
|
||
</quantity>
|
||
<date id="F363E286110DFFB5FF4BA9A518A79488" box="[213,397,1684,1708]" pageId="87" pageNumber="192" value="2003-03-17" valueMax="2003-03-21" valueMin="2003-03-17">
|
||
<collectingDate id="E3271B6E110DFFB5FF4BA9A518A79488" box="[213,397,1684,1708]" pageId="87" pageNumber="192" value="2003-03-17" valueMax="2003-03-21" valueMin="2003-03-17">17-21 Mar. 2003</collectingDate>
|
||
</date>
|
||
/
|
||
<geoCoordinate id="E2E9A281110DFFB5FE3AA9A51B139488" box="[420,569,1684,1708]" degrees="15" direction="south" minutes="17" orientation="latitude" pageId="87" pageNumber="192" precision="15" seconds="18" value="-15.288334">15° 17’ 18” S</geoCoordinate>
|
||
<geoCoordinate id="E2E9A281110DFFB5FDDFA9A51BCF9488" box="[577,741,1684,1708]" degrees="049" direction="east" minutes="32" orientation="longitude" pageId="87" pageNumber="192" precision="15" seconds="54" value="49.548332">049° 32’ 54” E</geoCoordinate>
|
||
/
|
||
<location id="8202929D110DFFB5FF16A98518E894E8" LSID="urn:lsid:plazi:treatment:0F747550110DFFBAFF16ACC5182C9688:8202929D110DFFB5FF16A98518E894E8" box="[136,450,1716,1740]" country="Madagascar" latitude="-15.288334" longLatPrecision="21" longitude="49.548332" municipality="Montagne d'Akirindro" name="California Acad. of Sciences" pageId="87" pageNumber="192" stateProvince="Toamasina">California Acad. of Sciences</location>
|
||
coll.
|
||
<collectorName id="2A28A190110DFFB5FD9FA9851B6D94E8" box="[513,583,1716,1740]" pageId="87" pageNumber="192">Fisher</collectorName>
|
||
,
|
||
<collectorName id="2A28A190110DFFB5FDCCA9851B9D94E8" box="[594,695,1716,1740]" pageId="87" pageNumber="192">Griswold</collectorName>
|
||
et al. sifted litter (leaf mold, rotten wood) rainforest collection code: BLF 8250
|
||
</materialsCitation>
|
||
/.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="CFC797CD110DFFB5FF30A8251DE69188" pageId="87" pageNumber="192" type="diagnosis">
|
||
<paragraph id="8762C446110DFFB5FF30A8251DE69188" blockId="87.[136,758,1172,1932]" lastBlockId="87.[816,1438,180,1932]" pageId="87" pageNumber="192">
|
||
<emphasis id="B5A91854110DFFB5FF30A825180E9508" bold="true" box="[174,292,1812,1836]" pageId="87" pageNumber="192">Diagnosis.</emphasis>
|
||
<taxonomicName id="40DDBFC5110DFFB5FEB3A8251B6B9508" authorityName="Ballerio" authorityYear="2021" box="[301,577,1812,1836]" class="Insecta" family="Hybosoridae" genus="Goudotostes" kingdom="Animalia" order="Coleoptera" pageId="87" pageNumber="192" phylum="Arthropoda" rank="species" species="rajemisonae" status="sp. nov.">
|
||
<emphasis id="B5A91854110DFFB5FEB3A8251B6B9508" box="[301,577,1812,1836]" italics="true" pageId="87" pageNumber="192">Goudotostes rajemisonae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B5A91854110DFFB5FDD5A8251B899508" bold="true" box="[587,675,1812,1836]" pageId="87" pageNumber="192">
|
||
<taxonomicNameLabel id="AE9AA52F110DFFB5FDD5A8251B899508" box="[587,675,1812,1836]" pageId="87" pageNumber="192" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
can be differentiated from all other species of
|
||
<taxonomicName id="40DDBFC5110DFFB5FDD9A8051BE49568" authorityName=", sensu Ballerio" authorityYear="2008" box="[583,718,1844,1868]" class="Insecta" family="Hybosoridae" genus="Goudotostes" kingdom="Animalia" order="Coleoptera" pageId="87" pageNumber="192" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B5A91854110DFFB5FDD9A8051BE49568" box="[583,718,1844,1868]" italics="true" pageId="87" pageNumber="192">Goudotostes</emphasis>
|
||
</taxonomicName>
|
||
by the following combination of characters: a) small size, b) dorsal ocular area absent, c) clypeal apical projection pointed, d) clypeal punctation made of dense large deeply impressed simple punctures, e) fronto-clypeal tubercle present, f) pronotal anterior carina bilobate, g) pronotal basal carina bilobate, h) pronotal disc covered by relatively sparse (mainly on disc) large simple deeply impressed punctures, i) elytral dorsal area covered by three longitudinal carinae shortly interrupted multiple times, j) elytral first interstria (area between sutural stria and first longitudinal carina) with dense punctation (interpunctural distance inferior than punctural diameter), k) left paramere with ventral margin invaginated (a character which places this species in the
|
||
<taxonomicName id="40DDBFC5110DFFB5FBD2AD251DA19008" authorityName="Ballerio" authorityYear="2021" box="[1100,1163,532,556]" class="Insecta" family="Hybosoridae" genus="Goudotostes" kingdom="Animalia" order="Coleoptera" pageId="87" pageNumber="192" phylum="Arthropoda" rank="species" species="laevis">
|
||
<emphasis id="B5A91854110DFFB5FBD2AD251DA19008" box="[1100,1163,532,556]" italics="true" pageId="87" pageNumber="192">laevis</emphasis>
|
||
</taxonomicName>
|
||
group of species), l) the distinctive shape of parameres (as in
|
||
<figureCitation id="1FE6D8C3110DFFB5FB53AD051C0A9068" box="[1229,1312,564,588]" captionStart="Fig" captionStartId="132.[151,182,1885,1904]" captionTargetBox="[161,1420,173,1837]" captionTargetId="figure-13@132.[145,1443,142,1858]" captionTargetPageId="132" captionText="Fig. 67 – Goudotostes rajemisonae sp. nov., holotype,a, dorsal view of distended specimen; b, ventral view of rolled up specimen; c, dorsal view of rolled up specimen; d, ventral view of rolled up specimen; e, lateral view of rolled up specimen; f, spiculum gastrale; g–j, four views of aedeagus." figureDoi="http://doi.org/10.5281/zenodo.12764317" httpUri="https://zenodo.org/record/12764317/files/figure.png" pageId="87" pageNumber="192">Figs 67</figureCitation>
|
||
g-j). Within the
|
||
<taxonomicName id="40DDBFC5110DFFB5FCE9AD651A9C9048" authorityName="Ballerio" authorityYear="2021" box="[887,950,596,620]" class="Insecta" family="Hybosoridae" genus="Goudotostes" kingdom="Animalia" order="Coleoptera" pageId="87" pageNumber="192" phylum="Arthropoda" rank="species" species="laevis">
|
||
<emphasis id="B5A91854110DFFB5FCE9AD651A9C9048" box="[887,950,596,620]" italics="true" pageId="87" pageNumber="192">laevis</emphasis>
|
||
</taxonomicName>
|
||
species group,
|
||
<taxonomicName id="40DDBFC5110DFFB5FBFFAD651C5F9048" authorityName="Ballerio" authorityYear="2021" box="[1121,1397,596,620]" class="Insecta" family="Hybosoridae" genus="Goudotostes" kingdom="Animalia" order="Coleoptera" pageId="87" pageNumber="192" phylum="Arthropoda" rank="species" species="rajemisonae" status="sp. nov.">
|
||
<emphasis id="B5A91854110DFFB5FBFFAD651C5F9048" box="[1121,1397,596,620]" italics="true" pageId="87" pageNumber="192">Goudotostes rajemisonae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B5A91854110DFFB5FAE3AD651A4A90A8" bold="true" pageId="87" pageNumber="192">
|
||
<taxonomicNameLabel id="AE9AA52F110DFFB5FAE3AD651A4A90A8" pageId="87" pageNumber="192" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
can be confused with
|
||
<taxonomicName id="40DDBFC5110DFFB5FBD0AD451CB690A8" authorityName="Ballerio" authorityYear="2021" box="[1102,1436,628,652]" class="Insecta" family="Hybosoridae" genus="Goudotostes" kingdom="Animalia" order="Coleoptera" pageId="87" pageNumber="192" phylum="Arthropoda" rank="species" species="rafanomezantsoai" status="sp. nov.">
|
||
<emphasis id="B5A91854110DFFB5FBD0AD451CB690A8" box="[1102,1436,628,652]" italics="true" pageId="87" pageNumber="192">Goudotostes rafanomezantsoai</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B5A91854110DFFB5FCAEADA51AAD9088" bold="true" box="[816,903,660,684]" pageId="87" pageNumber="192">
|
||
<taxonomicNameLabel id="AE9AA52F110DFFB5FCAEADA51AAD9088" box="[816,903,660,684]" pageId="87" pageNumber="192" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, from which differs mainly because of the elytral sculpturing: in
|
||
<taxonomicName id="40DDBFC5110DFFB5FC4DAD841D5390E8" authorityName="Ballerio" authorityYear="2021" box="[979,1145,693,716]" class="Insecta" family="Hybosoridae" genus="Goudotostes" kingdom="Animalia" order="Coleoptera" pageId="87" pageNumber="192" phylum="Arthropoda" rank="species" species="rajemisonae" status="sp. nov.">
|
||
<emphasis id="B5A91854110DFFB5FC4DAD841D5390E8" box="[979,1145,693,716]" italics="true" pageId="87" pageNumber="192">G. rajemisonae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B5A91854110DFFB5FBE1AD851DFF90E8" bold="true" box="[1151,1237,692,716]" pageId="87" pageNumber="192">
|
||
<taxonomicNameLabel id="AE9AA52F110DFFB5FBE1AD851DFF90E8" box="[1151,1237,692,716]" pageId="87" pageNumber="192" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
elytra are covered by at least three rows of longitudinal carinae, sometimes shortly interrupted, whereas in
|
||
<taxonomicName id="40DDBFC5110DFFB5FB10ADC51C599128" authorityName="Ballerio" authorityYear="2021" box="[1166,1395,756,780]" class="Insecta" family="Hybosoridae" genus="Goudotostes" kingdom="Animalia" order="Coleoptera" pageId="87" pageNumber="192" phylum="Arthropoda" rank="species" species="rafanomezantsoai" status="sp. nov.">
|
||
G.
|
||
<emphasis id="B5A91854110DFFB5FB2CADC51C599128" box="[1202,1395,756,780]" italics="true" pageId="87" pageNumber="192">rafanomezantsoai</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B5A91854110DFFB5FAE0ADC51A74910F" bold="true" italics="true" pageId="87" pageNumber="192">
|
||
<taxonomicNameLabel id="AE9AA52F110DFFB5FAE0ADC51A74910F" pageId="87" pageNumber="192" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
the longitudinal carinae are replaced by sparse irregular tubercles mixed to few very short carinae. The shape of paremeres is different too, with right paramere of
|
||
<taxonomicName id="40DDBFC5110DFFB5FA1CAC641ADB91A8" authorityName="Ballerio" authorityYear="2021" class="Insecta" family="Hybosoridae" genus="Goudotostes" kingdom="Animalia" order="Coleoptera" pageId="87" pageNumber="192" phylum="Arthropoda" rank="species" species="rafanomezantsoai" status="sp. nov.">
|
||
<emphasis id="B5A91854110DFFB5FA1CAC641ADB91A8" italics="true" pageId="87" pageNumber="192">G. rafanomezantsoai</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B5A91854110DFFB5FC66AC451D6591A8" bold="true" box="[1016,1103,884,908]" pageId="87" pageNumber="192">
|
||
<taxonomicNameLabel id="AE9AA52F110DFFB5FC66AC451D6591A8" box="[1016,1103,884,908]" pageId="87" pageNumber="192" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
subtruncate, whereas in
|
||
<taxonomicName id="40DDBFC5110DFFB5FAC5AC441AB79188" authorityName="Ballerio" authorityYear="2021" class="Insecta" family="Hybosoridae" genus="Goudotostes" kingdom="Animalia" order="Coleoptera" pageId="87" pageNumber="192" phylum="Arthropoda" rank="species" species="rajemisonae" status="sp. nov.">
|
||
<emphasis id="B5A91854110DFFB5FAC5AC441AB79188" italics="true" pageId="87" pageNumber="192">G. rajemisonae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B5A91854110DFFB5FC3BACA51AD19188" bold="true" box="[933,1019,916,940]" pageId="87" pageNumber="192">
|
||
<taxonomicNameLabel id="AE9AA52F110DFFB5FC3BACA51AD19188" box="[933,1019,916,940]" pageId="87" pageNumber="192" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is broadly pointed.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="CFC797CD110DFFBAFCC8AC851B879168" lastPageId="88" lastPageNumber="193" pageId="87" pageNumber="192" type="description">
|
||
<paragraph id="8762C446110DFFB5FCC8AC851DB795A8" blockId="87.[816,1438,180,1932]" pageId="87" pageNumber="192">
|
||
<emphasis id="B5A91854110DFFB5FCC8AC851ACB91E8" bold="true" box="[854,993,948,972]" pageId="87" pageNumber="192">Description.</emphasis>
|
||
Size: HL =
|
||
<quantity id="402569A3110DFFB5FBF4AC851DEB91E8" box="[1130,1217,948,972]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="87" pageNumber="192" unit="mm" value="0.9">0.9 mm</quantity>
|
||
; HW =
|
||
<quantity id="402569A3110DFFB5FA84AC851C5891E8" box="[1306,1394,948,972]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="87" pageNumber="192" unit="mm" value="1.3">1.3 mm</quantity>
|
||
; PL =
|
||
<quantity id="402569A3110DFFB5FCD6ACE51AB591C8" box="[840,927,980,1004]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="87" pageNumber="192" unit="mm" value="1.2">1.2 mm</quantity>
|
||
; PW =
|
||
<quantity id="402569A3110DFFB5FC6DACE51D6091C8" box="[1011,1098,980,1004]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" pageId="87" pageNumber="192" unit="mm" value="2.2">2.2 mm</quantity>
|
||
; EL =
|
||
<quantity id="402569A3110DFFB5FB08ACE51DC791C8" box="[1174,1261,980,1004]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" pageId="87" pageNumber="192" unit="mm" value="2.2">2.2 mm</quantity>
|
||
; EW =
|
||
<quantity id="402569A3110DFFB5FADCACE51CB391C8" box="[1346,1433,980,1004]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" pageId="87" pageNumber="192" unit="mm" value="2.1">2.1 mm</quantity>
|
||
. Overall morphology as in generic diagnosis. Small-sized
|
||
<taxonomicName id="40DDBFC5110DFFB5FCAEAB251A9F9608" authorityName=", sensu Ballerio" authorityYear="2008" box="[816,949,1044,1068]" class="Insecta" family="Hybosoridae" genus="Goudotostes" kingdom="Animalia" order="Coleoptera" pageId="87" pageNumber="192" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B5A91854110DFFB5FCAEAB251A9F9608" box="[816,949,1044,1068]" italics="true" pageId="87" pageNumber="192">Goudotostes</emphasis>
|
||
</taxonomicName>
|
||
. Body strongly convex. Black, shiny, dorsum with extremely short fine setation (50×), underside, tarsi and antennae dark reddish-brown, setation yellowish-brown.
|
||
<emphasis id="B5A91854110DFFB5FC32AB451ACC96A8" box="[940,998,1140,1164]" italics="true" pageId="87" pageNumber="192">Head</emphasis>
|
||
: W/L ratio = 1.39; clypeal apex obtuse, apical projection strongly developed and sexually dimorphic, sides almost rectilinear with slightly unaligned genae in correspondence with the genal suture; dorsal ocular area absent, genal canthus indistinct, ventral ocular area small-sized, genae acute, distinctly protruding outwards, genal suture indistinct; head dorsal surface uniformly covered by dense large impressed simple punctures, fore margin marked by some irregular transverse lines. Interpunctural distance inferior than punctural diameter. Fronto-clypeal tubercle very weakly raised, smooth and blunt.
|
||
<emphasis id="B5A91854110DFFB5FAB5AA841CBF97E8" box="[1323,1429,1461,1484]" italics="true" pageId="87" pageNumber="192">Pronotum</emphasis>
|
||
: subrectangular (dorsal view) (W/L ratio = 2.00), convex, fore angles gently subtruncate; pronotal surface covered by dense large deep simple punctures becoming sparser smaller and comma-shaped at sides, area near fore angles with few irregular lines, interpunctural distance mostly as long as punctural diameter; pronotal bead complete although not visible from above along base; bilobate basal carina raised; bilobate anterior carina raised; pronotal sculpturing made of large and gently raised tubercles and carinae as follows: sides of disc with one gently raised transverse carina reaching outer margin.
|
||
<taxonomicName id="40DDBFC5110DFFB5FB60A8251C4D9508" box="[1278,1383,1812,1836]" class="Trilobita" family="Styginidae" genus="Scutellum" kingdom="Animalia" order="Corynexochida" pageId="87" pageNumber="192" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B5A91854110DFFB5FB60A8251C4D9508" box="[1278,1383,1812,1836]" italics="true" pageId="87" pageNumber="192">Scutellum</emphasis>
|
||
</taxonomicName>
|
||
: basal area smooth, remaining surface covered by impressed transverse large comma-shaped punctures mixed to some large impressed simple punctures.
|
||
</paragraph>
|
||
<paragraph id="8762C4461102FFBAFF21AF851B879168" blockId="88.[151,772,180,1196]" pageId="88" pageNumber="193">
|
||
<emphasis id="B5A918541102FFBAFF21AF8519D592E8" box="[191,255,180,204]" italics="true" pageId="88" pageNumber="193">Elytra</emphasis>
|
||
: longer than wide (W/L ratio = 0.86), subovoidal (dorsal view), strongly convex; humeral callus marked by a tubercle; elytral surface covered by large deep horseshoe-shaped punctures with opening directed backwards mixed to large impressed comma-shaped and simple punctures, interpunctural distance almost equal to or inferior than punctural diameter. Striated articular area hardly visible in lateral view, relatively narrow and short; inferior sutural stria visible and complete; marginal area large-sized; elytral articular process well developed, smooth and shiny, isolated from remaining elytral surface by a deep furrow. Elytral sculpturing as follows: base with one raised large tubercle below articular process with apex directed backwards (parasutural tubercle), next to it a humeral oblique large raised tubercle with apex directed backwards (humeral tubercle), dorsal area of elytra with three to four irregular and asymmetrical rows of weakly raised smooth carinae, sides and apical third with some more raised tubercles and longitudinal short carinae.
|
||
<emphasis id="B5A918541102FFBAFE27ADC418D09128" box="[441,506,757,780]" italics="true" pageId="88" pageNumber="193">Wings</emphasis>
|
||
: flightless.
|
||
<emphasis id="B5A918541102FFBAFDE7ADC519999108" italics="true" pageId="88" pageNumber="193">Male genitalia</emphasis>
|
||
: aedeagus (L =
|
||
<quantity id="402569A31102FFBAFEFBAC2518909108" box="[357,442,788,812]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="88" pageNumber="193" unit="mm" value="0.7">0.7 mm</quantity>
|
||
) with parameres as in
|
||
<figureCitation id="1FE6D8C31102FFBAFD2CAC251A2E9108" box="[690,772,788,812]" captionStart="Fig" captionStartId="132.[151,182,1885,1904]" captionTargetBox="[161,1420,173,1837]" captionTargetId="figure-13@132.[145,1443,142,1858]" captionTargetPageId="132" captionText="Fig. 67 – Goudotostes rajemisonae sp. nov., holotype,a, dorsal view of distended specimen; b, ventral view of rolled up specimen; c, dorsal view of rolled up specimen; d, ventral view of rolled up specimen; e, lateral view of rolled up specimen; f, spiculum gastrale; g–j, four views of aedeagus." figureDoi="http://doi.org/10.5281/zenodo.12764317" httpUri="https://zenodo.org/record/12764317/files/figure.png" pageId="88" pageNumber="193">Figs 67</figureCitation>
|
||
g-j, spiculum gastrale (L =
|
||
<quantity id="402569A31102FFBAFE2BAC051B2D9168" box="[437,519,820,844]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="88" pageNumber="193" unit="mm" value="0.6">0.6 mm</quantity>
|
||
) as in
|
||
<figureCitation id="1FE6D8C31102FFBAFDD2AC051B809168" box="[588,682,820,844]" captionStart="Fig" captionStartId="132.[151,182,1885,1904]" captionTargetBox="[161,1420,173,1837]" captionTargetId="figure-13@132.[145,1443,142,1858]" captionTargetPageId="132" captionText="Fig. 67 – Goudotostes rajemisonae sp. nov., holotype,a, dorsal view of distended specimen; b, ventral view of rolled up specimen; c, dorsal view of rolled up specimen; d, ventral view of rolled up specimen; e, lateral view of rolled up specimen; f, spiculum gastrale; g–j, four views of aedeagus." figureDoi="http://doi.org/10.5281/zenodo.12764317" httpUri="https://zenodo.org/record/12764317/files/figure.png" pageId="88" pageNumber="193">
|
||
Fig.
|
||
<specimenCount id="91DB0FCF1102FFBAFDE3AC051B809168" box="[637,682,820,844]" count="67" pageId="88" pageNumber="193" type="generic">67 f</specimenCount>
|
||
</figureCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="CFC797CD1102FFBAFF23AC651A2E9188" pageId="88" pageNumber="193" type="etymology">
|
||
<paragraph id="8762C4461102FFBAFF23AC651A2E9188" blockId="88.[151,772,180,1196]" pageId="88" pageNumber="193">
|
||
<emphasis id="B5A918541102FFBAFF23AC6518179148" bold="true" box="[189,317,852,876]" pageId="88" pageNumber="193">Etymology.</emphasis>
|
||
Noun in the genitive case. Named after Balsama Rajemison (
|
||
<collectingCountry id="FFCA84D61102FFBAFE10AC451B3B91A8" box="[398,529,884,908]" name="Madagascar" pageId="88" pageNumber="193">Madagascar</collectingCountry>
|
||
Biodiversity Center,
|
||
<collectingRegion id="45190AA41102FFBAFF09ACA518079188" box="[151,301,916,940]" country="Madagascar" name="Antananarivo" pageId="88" pageNumber="193">Antananarivo</collectingRegion>
|
||
), member of the Malagasy lab team of CAS.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="CFC797CD1102FFBAFF23AC85182C9688" pageId="88" pageNumber="193" type="distribution">
|
||
<paragraph id="8762C4461102FFBAFF23AC85182C9688" blockId="88.[151,772,180,1196]" pageId="88" pageNumber="193">
|
||
<emphasis id="B5A918541102FFBAFF23AC8518F691E8" bold="true" box="[189,476,948,972]" pageId="88" pageNumber="193">Distribution and habitat.</emphasis>
|
||
Known only from the
|
||
<typeStatus id="58667AE41102FFBAFD4BAC841A2991E9" box="[725,771,949,973]" pageId="88" pageNumber="193">type</typeStatus>
|
||
locality in Eastern
|
||
<collectingCountry id="FFCA84D61102FFBAFEF6ACE518C191C8" box="[360,491,980,1004]" name="Madagascar" pageId="88" pageNumber="193">Madagascar</collectingCountry>
|
||
. The
|
||
<typeStatus id="58667AE41102FFBAFDB7ACE41B7D91C9" box="[553,599,981,1005]" pageId="88" pageNumber="193">type</typeStatus>
|
||
series was collected by sifting leaf litter and rotten wood in rainforest. The vegetation cover consists of “lowland moist evergreen forest”, (
|
||
<bibRefCitation id="E34CB9B71102FFBAFF6AAB0518F29668" author="Goodman S. M. & Raherilalao M. J. & Wohlhauser S." box="[244,472,1076,1100]" pageId="88" pageNumber="193" refId="ref102713" refString="Goodman S. M., Raherilalao M. J., Wohlhauser S. (eds). 2018. Les aires potegees terrestres de Madagascar: leur histoire, description et biote / The terrestrial protected areas of Madagascar: their history, description and biota. Association Vahatra, Antananarivo. 1716 pp." type="book" year="2018">Goodman et al. 2018</bibRefCitation>
|
||
) also known as “humid forest” (
|
||
<bibRefCitation id="E34CB9B71102FFBAFF4DAB6518969648" author="Moat J. & Smith P." box="[211,444,1108,1132]" pageId="88" pageNumber="193" refId="ref103114" refString="Moat J., Smith P. 2007. Atlas of the vegetation of Madagascar. Royal Botanic Garden, Kew 124 pp." type="book" year="2007">Moat & Smith 2007</bibRefCitation>
|
||
). The collecting locality falls within the protected area “Réserve Spéciale d’Anjanaharibe-Sud”.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |