286 lines
36 KiB
XML
286 lines
36 KiB
XML
<document id="041F1BDF57B159B00CBCBCAED6ECAAF8" ID-DOI="10.13133/2284-4880/545" ID-ISSN="2284-4880" ID-Zenodo-Dep="12764154" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1721242503917" checkinUser="felipe" docAuthor="Ballerio, Alberto" docDate="2021" docId="0F7475501109FFB6FF16AD251BDF9048" docLanguage="en" docName="FragEntomol.53.2.105-282.pdf" docOrigin="Fragmenta entomologica 53 (2)" docSource="http://dx.doi.org/10.13133/2284-4880/545" docStyle="DocumentStyle:8499FF4308F512386C0E43DC290A16BD.1:FragEntomol.2021-.journal_article.0cover" docStyleId="8499FF4308F512386C0E43DC290A16BD" docStyleName="FragEntomol.2021-.journal_article.0cover" docStyleVersion="1" docTitle="Goudotostes parvus Ballerio, 2021, sp. nov." docType="treatment" docUuid="275E01F2-4136-497C-8C12-4948B81AC029" docUuidSource="ZooBank" docVersion="2" lastPageNumber="189" masterDocId="F34D0D28115AFFE2FF9EAF31192A9224" masterDocTitle="The Ceratocanthinae of Madagascar and Comoro Islands: a revision of the genera Synarmostes and Goudotostes, and of the flightless Philharmostes, with description of 64 new species (Coleoptera: Scarabaeoidea, Hybosoridae)" masterLastPageNumber="282" masterPageNumber="105" pageNumber="188" updateTime="1721247602959" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-4.0">
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<mods:mods id="D933468D1F83058367DB3D7673049674" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="264B54B6C4695DE0B7CC708A9892405E">
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<mods:title id="D415BD40091B07CE5ACA1490E09F3F07">The Ceratocanthinae of Madagascar and Comoro Islands: a revision of the genera Synarmostes and Goudotostes, and of the flightless Philharmostes, with description of 64 new species (Coleoptera: Scarabaeoidea, Hybosoridae)</mods:title>
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</mods:titleInfo>
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<mods:name id="01C5A35A4B4C0DE1076D43960CBA1886" type="personal">
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<mods:role id="C11DAC5B2FF8ED234E32EE25D98EF9C4">
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<mods:roleTerm id="E533793628A5080F37B08814D4ED394C">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="0D446E1F9EB2C9D51CE0437065AC0ACE">Ballerio, Alberto</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="1F51507B4641037F4557A718EBDEBB77">text</mods:typeOfResource>
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<mods:titleInfo id="32147F40A843F974D43EAC5D21E97521">
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<mods:title id="017EAE324C352807F91C0CB4B1853622">Fragmenta entomologica</mods:title>
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</mods:titleInfo>
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<mods:part id="59346442F5F5D485BB64B0F3F1CE5117">
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<mods:date id="B0363B993FE9B8DB698D69F124E276C6">2021</mods:date>
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<mods:detail id="05197A3027AA7A7CE79581B146BC7828" type="pubDate">
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<mods:number id="F97653BAD34638AABCA7C9A994F9864D">2021-11-30</mods:number>
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</mods:detail>
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<mods:detail id="2B291E95A29A45E685C31E920C97B16E" type="volume">
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<mods:number id="C337130A0C83575F2153F291666D9497">53</mods:number>
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</mods:detail>
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<mods:detail id="8C8319484BC84457A5F9EFD2A3819E83" type="issue">
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<mods:number id="46BC7FD5AC03E6DF7D1D2EF58A7413AF">2</mods:number>
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</mods:detail>
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<mods:extent id="5415317E5AE7CE5FC7F2CDE78FB2B8B1" unit="page">
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<mods:start id="52B926EDFEA23BC08BFDECF782250F52">105</mods:start>
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<mods:end id="AD165D68F6484E3E025C3952FE853411">282</mods:end>
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</mods:extent>
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<mods:location id="AE609179157D5F2CCB4B352B1B93B4E9">
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<mods:url id="32ED5966E0C74A3AB5B75F3EF9D52DA3">http://dx.doi.org/10.13133/2284-4880/545</mods:url>
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</mods:location>
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<mods:classification id="9E5600CFCC48C6FB40CE347BE1A410F9">journal article</mods:classification>
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<mods:identifier id="5B71B3CFC2C207BEFBDDC7F2629AC957" type="DOI">10.13133/2284-4880/545</mods:identifier>
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<mods:identifier id="CC8D1D6B472389046EF7566A77B54706" type="ISSN">2284-4880</mods:identifier>
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<mods:identifier id="3C969A4CD70C5C1A4529CC1F0BC62507" type="Zenodo-Dep">12764154</mods:identifier>
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<mods:identifier id="13F985EED0F33E9E073DACA759F4C8EB" type="ZooBank">B3F75A2C-7603-4027-B8E1-F4A8B5A0CDD9</mods:identifier>
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</mods:mods>
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<treatment id="0F7475501109FFB6FF16AD251BDF9048" LSID="urn:lsid:zoobank.org:act:275E01F2-4136-497C-8C12-4948B81AC029" httpUri="http://treatment.plazi.org/id/0F7475501109FFB6FF16AD251BDF9048" lastPageId="84" lastPageNumber="189" pageId="83" pageNumber="188">
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<subSubSection id="CFC797CD1109FFB1FF16AD2518939008" box="[136,441,532,556]" pageId="83" pageNumber="188" type="nomenclature">
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<paragraph id="8762C4461109FFB1FF16AD2518939008" blockId="83.[136,441,532,556]" box="[136,441,532,556]" pageId="83" pageNumber="188">
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<heading id="DC2A732A1109FFB1FF16AD2518939008" box="[136,441,532,556]" fontSize="10" level="2" pageId="83" pageNumber="188" reason="5">
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<taxonomicName id="40DDBFC51109FFB1FF16AD2518779008" ID-CoL="8SSNL" authorityName="Ballerio" authorityYear="2021" box="[136,349,532,556]" class="Insecta" family="Hybosoridae" genus="Goudotostes" kingdom="Animalia" order="Coleoptera" pageId="83" pageNumber="188" phylum="Arthropoda" rank="species" species="parvus" status="sp. nov.">
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<emphasis id="B5A918541109FFB1FF16AD2518779008" box="[136,349,532,556]" italics="true" pageId="83" pageNumber="188">Goudotostes parvus</emphasis>
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</taxonomicName>
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<emphasis id="B5A918541109FFB1FEFDAD2518939008" bold="true" box="[355,441,532,556]" pageId="83" pageNumber="188">
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<taxonomicNameLabel id="AE9AA52F1109FFB1FEFDAD2518939008" box="[355,441,532,556]" pageId="83" pageNumber="188" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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||
</paragraph>
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</subSubSection>
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<subSubSection id="CFC797CD1109FFB1FF16AD0518D190A8" pageId="83" pageNumber="188" type="description">
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<paragraph id="8762C4461109FFB1FF16AD0518D190A8" blockId="83.[136,755,564,652]" pageId="83" pageNumber="188">
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(
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<figureCitation id="1FE6D8C31109FFB1FF0EAD0519CB9068" box="[144,225,564,588]" captionStart="Fig" captionStartId="131.[136,167,1858,1877]" captionTargetBox="[135,1442,172,1817]" captionTargetId="figure-4@131.[135,1450,166,1833]" captionTargetPageId="131" captionText="Fig. 66 – Goudotostes parvus sp. nov., a, holotype, dorsal view of distended specimen; b, paratype, dorsal view of rolled up specimen; c, paratype, ventral view of rolled up specimen; d, paratype, ventral view of rolled up specimen; e, paratype, lateral view of rolled up specimen; f, holotype, spiculum gastrale; g–k, holotype, four views of aedeagus." figureDoi="http://doi.org/10.5281/zenodo.12764315" httpUri="https://zenodo.org/record/12764315/files/figure.png" pageId="83" pageNumber="188">Figs 66</figureCitation>
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a-k, 77) http://zoobank.org/
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<uri id="F34CC8441109FFB1FEFBAD6518D190A8" pageId="83" pageNumber="188">
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||
urn:lsid:zoobank.org:act:
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<uuid id="F37BFE931109FFB1FDE9AD6518D190A8" pageId="83" pageNumber="188">275E01F2- 4136-497C-8C12-4948B81AC029</uuid>
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</uri>
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||
</paragraph>
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||
</subSubSection>
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<subSubSection id="CFC797CD1109FFB1FF30AD851B4E9708" pageId="83" pageNumber="188" type="materials_examined">
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<paragraph id="8762C4461109FFB1FF30AD851B4E9708" blockId="83.[136,756,692,1932]" pageId="83" pageNumber="188">
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<emphasis id="B5A918541109FFB1FF30AD85181A90E8" bold="true" box="[174,304,692,716]" pageId="83" pageNumber="188">Type series.</emphasis>
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<materialsCitation id="37B5CE1B1109FFB1FEABAD8518FF91C8" collectingDate="2003-11-15" collectingDateMax="2003-11-18" collectingDateMin="2003-11-15" collectorName="San Francisco, CA & B. L. Fisher" country="Madagascar" county="Academy of Sciences" determinerName="A. Ballerio" elevation="775" latitude="-14.435" location="California Acad. of Sciences" longLatPrecision="21" longitude="49.76" municipality="Parc National de Marojejy Manantenina River" pageId="83" pageNumber="188" specimenCount="1" specimenCount-male="1" stateProvince="Antsiranana" typeStatus="holotype">
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<typeStatus id="58667AE41109FFB1FEABAD8518B190E8" box="[309,411,692,716]" pageId="83" pageNumber="188" type="holotype">Holotype</typeStatus>
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,
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<specimenCount id="91DB0FCF1109FFB1FE3AAD84189090E8" box="[420,442,693,716]" count="1" pageId="83" pageNumber="188" type="male">♂</specimenCount>
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, deposited in the collection of the California
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<collectingCounty id="6E03BCCA1109FFB1FEBBADE51B3990C8" box="[293,531,724,748]" pageId="83" pageNumber="188">Academy of Sciences</collectingCounty>
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,
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<collectorName id="2A28A1901109FFB1FDBEADE51BDA90C8" box="[544,752,724,748]" pageId="83" pageNumber="188">San Francisco, CA</collectorName>
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, USA, labelled as follows: / CASTYPE 20277 / CASENT 8135384 /
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<collectingCountry id="FFCA84D61109FFB1FF64AC2518559108" box="[250,383,788,812]" name="Madagascar" pageId="83" pageNumber="188">Madagascar</collectingCountry>
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:
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<collectingRegion id="45190AA41109FFB1FE16AC251B239108" box="[392,521,788,812]" country="Madagascar" name="Antsiranana" pageId="83" pageNumber="188">Antsiranana</collectingRegion>
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<collectingMunicipality id="67065E3C1109FFB1FD91AC2418899168" pageId="83" pageNumber="188">Parc National de Marojejy Manantenina River</collectingMunicipality>
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<locationDeviation id="96D0DD641109FFB1FE31AC051BED9168" box="[431,711,820,844]" location="Andapa" pageId="83" pageNumber="188">
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<quantity id="402569A31109FFB1FE31AC051B219168" box="[431,523,820,844]" metricMagnitude="4" metricUnit="m" metricValue="2.76" pageId="83" pageNumber="188" unit="km" value="27.6">27.6 km</quantity>
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35° NE Andapa
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</locationDeviation>
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||
<locationDeviation id="96D0DD641109FFB1FD4DAC0518969148" location="Manantenina" pageId="83" pageNumber="188">
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||
<quantity id="402569A31109FFB1FD4DAC0519809148" metricMagnitude="3" metricUnit="m" metricValue="9.6" pageId="83" pageNumber="188" unit="km" value="9.6">9.6 km</quantity>
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327° NNW Manantenina
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||
</locationDeviation>
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||
el.
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||
<quantity id="402569A31109FFB1FE7BAC651B039148" box="[485,553,852,876]" metricMagnitude="2" metricUnit="m" metricValue="7.75" pageId="83" pageNumber="188" unit="m" value="775.0">
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||
<elevation id="0CF023751109FFB1FE7BAC651B039148" box="[485,553,852,876]" metricMagnitude="2" metricUnit="m" metricValue="7.75" pageId="83" pageNumber="188" unit="m" value="775.0">775 m</elevation>
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||
</quantity>
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||
/
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||
<date id="F363E2861109FFB1FDA0AC65199791A8" pageId="83" pageNumber="188" value="2003-11-15" valueMax="2003-11-18" valueMin="2003-11-15">
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||
<collectingDate id="E3271B6E1109FFB1FDA0AC65199791A8" pageId="83" pageNumber="188" value="2003-11-15" valueMax="2003-11-18" valueMin="2003-11-15">15-18 November 2003</collectingDate>
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||
</date>
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||
<geoCoordinate id="E2E9A2811109FFB1FF56AC45184891A8" box="[200,354,884,908]" degrees="14" direction="south" minutes="26" orientation="latitude" pageId="83" pageNumber="188" precision="15" seconds="06" value="-14.435">14° 26’ 06” S</geoCoordinate>
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<geoCoordinate id="E2E9A2811109FFB1FEF2AC451B3F91A8" box="[364,533,884,908]" degrees="049" direction="east" minutes="45" orientation="longitude" pageId="83" pageNumber="188" precision="15" seconds="36" value="49.76">049° 45’ 36” E</geoCoordinate>
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<location id="8202929D1109FFB1FDBEAC4519CF9188" LSID="urn:lsid:plazi:treatment:0F7475501109FFB6FF16AD251BDF9048:8202929D1109FFB1FDBEAC4519CF9188" country="Madagascar" county="Academy of Sciences" latitude="-14.435" longLatPrecision="21" longitude="49.76" municipality="Parc National de Marojejy Manantenina River" name="California Acad. of Sciences" pageId="83" pageNumber="188" stateProvince="Antsiranana">California Acad. of Sciences</location>
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||
coll.
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||
<collectorName id="2A28A1901109FFB1FEB6ACA418859188" box="[296,431,916,940]" pageId="83" pageNumber="188">B. L. Fisher</collectorName>
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||
et al. sifted litter (leaf mold, rotten wood) rainforest code: BLF 8872/
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||
<taxonomicName id="40DDBFC51109FFB1FDA5AC85198691C8" authorityName="Ballerio" authorityYear="2021" class="Insecta" family="Hybosoridae" genus="Goudotostes" kingdom="Animalia" order="Coleoptera" pageId="83" pageNumber="188" phylum="Arthropoda" rank="species" species="parvus" status="sp. nov.">
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||
<emphasis id="B5A918541109FFB1FDA5AC85198691C8" italics="true" pageId="83" pageNumber="188">Goudotostes parvus</emphasis>
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||
</taxonomicName>
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||
<taxonomicNameLabel id="AE9AA52F1109FFB1FF2DACE419C791C8" box="[179,237,981,1004]" pageId="83" pageNumber="188" rank="species">n. sp.</taxonomicNameLabel>
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det.
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<determinerName id="72BE28821109FFB1FEBCACE518B391C8" box="[290,409,980,1004]" pageId="83" pageNumber="188">A. Ballerio</determinerName>
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2020
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</materialsCitation>
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<typeStatus id="58667AE41109FFB1FE43ACE51B6691C8" box="[477,588,980,1004]" pageId="83" pageNumber="188" type="holotype">Holotypus</typeStatus>
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<specimenCount id="91DB0FCF1109FFB1FDCAACE41B4291C8" box="[596,616,981,1004]" count="1" pageId="83" pageNumber="188" type="male">♂</specimenCount>
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/ [distended, glued on card, dissected, male genitalia glued in DMHF resin on a separate card, same pin].
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<materialsCitation id="37B5CE1B1109FFB1FD90AB241B659708" collectingDate="2003-11-18" collectingDateMax="2003-11-18" collectingDateMin="2003-11-15" collectionCode="CASC, ABCB" collectorName="B. L. Fisher" country="Madagascar" county="Academy of Sciences" determinerName="A. Ballerio" elevation="1325" latitude="-14.443334" location="California Acad. of Sciences" longLatPrecision="21" longitude="49.743332" municipality="Parc National de Marojejy Antranohofa" pageId="83" pageNumber="188" specimenCount="7" specimenCount-male="6" stateProvince="Antsiranana" typeStatus="paratype">
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||
<typeStatus id="58667AE41109FFB1FD90AB241B5C9609" box="[526,630,1045,1069]" pageId="83" pageNumber="188" type="paratype">Paratypes</typeStatus>
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||
[
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||
<specimenCount id="91DB0FCF1109FFB1FD18AB251BEE9608" box="[646,708,1044,1068]" count="3" pageId="83" pageNumber="188" type="male">3 ♂♂</specimenCount>
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dissected]:
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<specimenCount id="91DB0FCF1109FFB1FF7FAB0518369668" box="[225,284,1076,1100]" count="3" pageId="83" pageNumber="188" type="male">3 ♂♂</specimenCount>
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||
and 19 exx. sex unstated (
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||
<quantity id="402569A31109FFB1FDADAB051B439668" box="[563,617,1076,1100]" metricMagnitude="-1" metricUnit="m" metricValue="4.572" pageId="83" pageNumber="188" unit="in" value="18.0">18 in</quantity>
|
||
<collectionCode id="E1CC5C831109FFB1FDF1AB051B9F9668" box="[623,693,1076,1100]" pageId="83" pageNumber="188">CASC</collectionCode>
|
||
and
|
||
<quantity id="402569A31109FFB1FD79AB0519B79648" metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="83" pageNumber="188" unit="in" value="4.0">4 in</quantity>
|
||
<collectionCode id="E1CC5C831109FFB1FF3DAB6519DA9648" box="[163,240,1108,1132]" pageId="83" pageNumber="188">ABCB</collectionCode>
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||
), same collecting data as holotype.
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||
<specimenCount id="91DB0FCF1109FFB1FDF3AB651B889648" box="[621,674,1108,1132]" count="1" pageId="83" pageNumber="188" type="generic">1 ex.</specimenCount>
|
||
sex us- tated (
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||
<collectionCode id="E1CC5C831109FFB1FF57AB45183F96A8" box="[201,277,1140,1164]" pageId="83" pageNumber="188">CASC</collectionCode>
|
||
): /
|
||
<collectingCountry id="FFCA84D61109FFB1FEACAB45189D96A8" box="[306,439,1140,1164]" name="Madagascar" pageId="83" pageNumber="188">Madagascar</collectingCountry>
|
||
:
|
||
<collectingRegion id="45190AA41109FFB1FE20AB451B1596A8" box="[446,575,1140,1164]" country="Madagascar" name="Antsiranana" pageId="83" pageNumber="188">Antsiranana</collectingRegion>
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||
<collectingMunicipality id="67065E3C1109FFB1FDDBAB44185C9688" pageId="83" pageNumber="188">Parc National de Marojejy Antranohofa</collectingMunicipality>
|
||
<locationDeviation id="96D0DD641109FFB1FEE3ABA51BBC9688" box="[381,662,1172,1196]" location="Andapa" pageId="83" pageNumber="188">
|
||
<quantity id="402569A31109FFB1FEE3ABA518FE9688" box="[381,468,1172,1196]" metricMagnitude="4" metricUnit="m" metricValue="2.66" pageId="83" pageNumber="188" unit="km" value="26.6">26.6 km</quantity>
|
||
31° NNE Andapa
|
||
</locationDeviation>
|
||
<locationDeviation id="96D0DD641109FFB1FD02ABA518BE96E8" location="Manantenina" pageId="83" pageNumber="188">
|
||
<quantity id="402569A31109FFB1FD02ABA51BDE9688" box="[668,756,1172,1196]" metricMagnitude="4" metricUnit="m" metricValue="1.0699999999999998" pageId="83" pageNumber="188" unit="km" value="10.7">10.7 km</quantity>
|
||
318° NNW Manantenina
|
||
</locationDeviation>
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||
el.
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||
<quantity id="402569A31109FFB1FE23AB851B2596E8" box="[445,527,1204,1228]" metricMagnitude="3" metricUnit="m" metricValue="1.325" pageId="83" pageNumber="188" unit="m" value="1325.0">
|
||
<elevation id="0CF023751109FFB1FE23AB851B2596E8" box="[445,527,1204,1228]" metricMagnitude="3" metricUnit="m" metricValue="1.325" pageId="83" pageNumber="188" unit="m" value="1325.0">1325 m</elevation>
|
||
</quantity>
|
||
/
|
||
<date id="F363E2861109FFB1FDBBAB851BDE96E8" box="[549,756,1204,1228]" pageId="83" pageNumber="188" value="2003-11-18">
|
||
<collectingDate id="E3271B6E1109FFB1FDBBAB851BDE96E8" box="[549,756,1204,1228]" pageId="83" pageNumber="188" value="2003-11-18">18 November 2003</collectingDate>
|
||
</date>
|
||
<geoCoordinate id="E2E9A2811109FFB1FF16ABE5183296C8" box="[136,280,1236,1260]" degrees="14" direction="south" minutes="26" orientation="latitude" pageId="83" pageNumber="188" precision="15" seconds="36" value="-14.443334">14° 26’ 36” S</geoCoordinate>
|
||
<geoCoordinate id="E2E9A2811109FFB1FE80ABE5189796C8" box="[286,445,1236,1260]" degrees="049" direction="east" minutes="44" orientation="longitude" pageId="83" pageNumber="188" precision="15" seconds="36" value="49.743332">049° 44’ 36” E</geoCoordinate>
|
||
<location id="8202929D1109FFB1FE5DABE51BDE96C8" LSID="urn:lsid:plazi:treatment:0F7475501109FFB6FF16AD251BDF9048:8202929D1109FFB1FE5DABE51BDE96C8" box="[451,756,1236,1260]" country="Madagascar" latitude="-14.443334" longLatPrecision="21" longitude="49.743332" municipality="Parc National de Marojejy Antranohofa" name="California Acad. of Sciences" pageId="83" pageNumber="188" stateProvince="Antsiranana">California Acad. of Sciences</location>
|
||
collector
|
||
<collectorName id="2A28A1901109FFB1FF70ABC4185B9728" box="[238,369,1268,1292]" pageId="83" pageNumber="188">B. L. Fisher</collectorName>
|
||
et al. sifted litter (leaf mold, rotten wood) montane rainforest code: BLF 9080
|
||
</materialsCitation>
|
||
/.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="CFC797CD1109FFB1FF30AA051CB99328" pageId="83" pageNumber="188" type="diagnosis">
|
||
<paragraph id="8762C4461109FFB1FF30AA051CB99328" blockId="83.[136,756,692,1932]" lastBlockId="83.[816,1437,180,1932]" pageId="83" pageNumber="188">
|
||
<emphasis id="B5A918541109FFB1FF30AA05180B9768" bold="true" box="[174,289,1332,1356]" pageId="83" pageNumber="188">Diagnosis.</emphasis>
|
||
<taxonomicName id="40DDBFC51109FFB1FEB4AA0518D59768" authorityName="Ballerio" authorityYear="2021" box="[298,511,1332,1356]" class="Insecta" family="Hybosoridae" genus="Goudotostes" kingdom="Animalia" order="Coleoptera" pageId="83" pageNumber="188" phylum="Arthropoda" rank="species" species="parvus" status="sp. nov.">
|
||
<emphasis id="B5A918541109FFB1FEB4AA0518D59768" box="[298,511,1332,1356]" italics="true" pageId="83" pageNumber="188">Goudotostes parvus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B5A918541109FFB1FD99AA051B749768" bold="true" box="[519,606,1332,1356]" pageId="83" pageNumber="188">
|
||
<taxonomicNameLabel id="AE9AA52F1109FFB1FD99AA051B749768" box="[519,606,1332,1356]" pageId="83" pageNumber="188" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
can be easily differentiated from all other species of
|
||
<taxonomicName id="40DDBFC51109FFB1FDB9AA651B819748" authorityName=", sensu Ballerio" authorityYear="2008" box="[551,683,1364,1388]" class="Insecta" family="Hybosoridae" genus="Goudotostes" kingdom="Animalia" order="Coleoptera" pageId="83" pageNumber="188" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B5A918541109FFB1FDB9AA651B819748" box="[551,683,1364,1388]" italics="true" pageId="83" pageNumber="188">Goudotostes</emphasis>
|
||
</taxonomicName>
|
||
by the following combination of characters: a) small size, b) dorsal ocular area absent, c) clypeal apical projection rounded, d) clypeal punctation made of dense large deeply impressed simple punctures, e) fronto-clypeal tubercle present, f) pronotal anterior carina bilobate, g) pronotal basal carina bilobate, h) pronotal disc covered by relatively sparse large simple deeply impressed punctures, i) elytral dorsal area covered by two longitudinal irregular rows of blunt very sharp carinae, j) sides of elytra covered by several distinctly protruding tubercles, k) elytral first interstria (area between sutural stria and first longitudinal carina) with dense punctation (interpunctural distance inferior than punctural diameter), l) left paramere with ventral margin invaginated (a character which places this species in the
|
||
<taxonomicName id="40DDBFC51109FFB1FDF1A8251B879508" authorityName="Ballerio" authorityYear="2021" box="[623,685,1812,1836]" class="Insecta" family="Hybosoridae" genus="Goudotostes" kingdom="Animalia" order="Coleoptera" pageId="83" pageNumber="188" phylum="Arthropoda" rank="species" species="laevis">
|
||
<emphasis id="B5A918541109FFB1FDF1A8251B879508" box="[623,685,1812,1836]" italics="true" pageId="83" pageNumber="188">laevis</emphasis>
|
||
</taxonomicName>
|
||
group of species), m) the distinctive shape of parameres (as in
|
||
<figureCitation id="1FE6D8C31109FFB1FF16A86519FD9548" box="[136,215,1876,1900]" captionStart="Fig" captionStartId="131.[136,167,1858,1877]" captionTargetBox="[135,1442,172,1817]" captionTargetId="figure-4@131.[135,1450,166,1833]" captionTargetPageId="131" captionText="Fig. 66 – Goudotostes parvus sp. nov., a, holotype, dorsal view of distended specimen; b, paratype, dorsal view of rolled up specimen; c, paratype, ventral view of rolled up specimen; d, paratype, ventral view of rolled up specimen; e, paratype, lateral view of rolled up specimen; f, holotype, spiculum gastrale; g–k, holotype, four views of aedeagus." figureDoi="http://doi.org/10.5281/zenodo.12764315" httpUri="https://zenodo.org/record/12764315/files/figure.png" pageId="83" pageNumber="188">Figs 66</figureCitation>
|
||
g-k). Within the
|
||
<taxonomicName id="40DDBFC51109FFB1FE13A86518E19548" authorityName="Ballerio" authorityYear="2021" box="[397,459,1876,1900]" class="Insecta" family="Hybosoridae" genus="Goudotostes" kingdom="Animalia" order="Coleoptera" pageId="83" pageNumber="188" phylum="Arthropoda" rank="species" species="laevis">
|
||
<emphasis id="B5A918541109FFB1FE13A86518E19548" box="[397,459,1876,1900]" italics="true" pageId="83" pageNumber="188">laevis</emphasis>
|
||
</taxonomicName>
|
||
species group,
|
||
<taxonomicName id="40DDBFC51109FFB1FDEEA86519FA95A8" authorityName="Ballerio" authorityYear="2021" class="Insecta" family="Hybosoridae" genus="Goudotostes" kingdom="Animalia" order="Coleoptera" pageId="83" pageNumber="188" phylum="Arthropoda" rank="species" species="parvus" status="sp. nov.">
|
||
<emphasis id="B5A918541109FFB1FDEEA86519FA95A8" italics="true" pageId="83" pageNumber="188">Goudotostes parvus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B5A918541109FFB1FF4BA845180395A8" bold="true" box="[213,297,1908,1932]" pageId="83" pageNumber="188">
|
||
<taxonomicNameLabel id="AE9AA52F1109FFB1FF4BA845180395A8" box="[213,297,1908,1932]" pageId="83" pageNumber="188" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is similar to
|
||
<taxonomicName id="40DDBFC51109FFB1FE32A8451BB095A8" authorityName="Ballerio" authorityYear="2021" box="[428,666,1908,1932]" class="Insecta" family="Hybosoridae" genus="Goudotostes" kingdom="Animalia" order="Coleoptera" pageId="83" pageNumber="188" phylum="Arthropoda" rank="species" species="masoalae" status="sp. nov.">
|
||
<emphasis id="B5A918541109FFB1FE32A8451BB095A8" box="[428,666,1908,1932]" italics="true" pageId="83" pageNumber="188">Goudotostes masoalae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B5A918541109FFB1FD3FA8451BDE95A8" bold="true" box="[673,756,1908,1932]" pageId="83" pageNumber="188">
|
||
<taxonomicNameLabel id="AE9AA52F1109FFB1FD3FA8451BDE95A8" box="[673,756,1908,1932]" pageId="83" pageNumber="188" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
due to the sculpturing of elytra but
|
||
<taxonomicName id="40DDBFC51109FFB1FB02AF841C2992E8" authorityName="Ballerio" authorityYear="2021" box="[1180,1283,181,204]" class="Insecta" family="Hybosoridae" genus="Goudotostes" kingdom="Animalia" order="Coleoptera" pageId="83" pageNumber="188" phylum="Arthropoda" rank="species" species="parvus" status="sp. nov.">
|
||
<emphasis id="B5A918541109FFB1FB02AF841C2992E8" box="[1180,1283,181,204]" italics="true" pageId="83" pageNumber="188">G. parvus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B5A918541109FFB1FA97AF851C7792E8" bold="true" box="[1289,1373,180,204]" pageId="83" pageNumber="188">
|
||
<taxonomicNameLabel id="AE9AA52F1109FFB1FA97AF851C7792E8" box="[1289,1373,180,204]" pageId="83" pageNumber="188" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is unmistakable because of the rounded apical projection of clypeus. It’s also the smallest among all known
|
||
<taxonomicName id="40DDBFC51109FFB1FA96AFC51CA69328" authorityName=", sensu Ballerio" authorityYear="2008" box="[1288,1420,244,268]" class="Insecta" family="Hybosoridae" genus="Goudotostes" kingdom="Animalia" order="Coleoptera" pageId="83" pageNumber="188" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B5A918541109FFB1FA96AFC51CA69328" box="[1288,1420,244,268]" italics="true" pageId="83" pageNumber="188">Goudotostes</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="CFC797CD1109FFB6FCC8AE251BFE9328" lastPageId="84" lastPageNumber="189" pageId="83" pageNumber="188" type="description">
|
||
<paragraph id="8762C4461109FFB6FCC8AE251BFE9328" blockId="83.[816,1437,180,1932]" lastBlockId="84.[151,772,180,620]" lastPageId="84" lastPageNumber="189" pageId="83" pageNumber="188">
|
||
<emphasis id="B5A918541109FFB1FCC8AE251ACA9308" bold="true" box="[854,992,276,300]" pageId="83" pageNumber="188">Description.</emphasis>
|
||
Size: HL =
|
||
<quantity id="402569A31109FFB1FBF4AE251DEB9308" box="[1130,1217,276,300]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="83" pageNumber="188" unit="mm" value="0.8">0.8 mm</quantity>
|
||
; HW =
|
||
<quantity id="402569A31109FFB1FA84AE251C589308" box="[1306,1394,276,300]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="83" pageNumber="188" unit="mm" value="1.2">1.2 mm</quantity>
|
||
; PL =
|
||
<quantity id="402569A31109FFB1FCD6AE051AB59368" box="[840,927,308,332]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="83" pageNumber="188" unit="mm" value="1.1">1.1 mm</quantity>
|
||
; PW =
|
||
<quantity id="402569A31109FFB1FC6DAE051D609368" box="[1011,1098,308,332]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" pageId="83" pageNumber="188" unit="mm" value="1.7">1.7 mm</quantity>
|
||
; EL =
|
||
<quantity id="402569A31109FFB1FB08AE051DC79368" box="[1174,1261,308,332]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="83" pageNumber="188" unit="mm" value="1.8">1.8 mm</quantity>
|
||
; EW =
|
||
<quantity id="402569A31109FFB1FADCAE051CB39368" box="[1346,1433,308,332]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" pageId="83" pageNumber="188" unit="mm" value="1.7">1.7 mm</quantity>
|
||
. Overall morphology as in generic diagnosis. Small-sized
|
||
<taxonomicName id="40DDBFC51109FFB1FCAEAE451A9E93A8" authorityName=", sensu Ballerio" authorityYear="2008" box="[816,948,372,396]" class="Insecta" family="Hybosoridae" genus="Goudotostes" kingdom="Animalia" order="Coleoptera" pageId="83" pageNumber="188" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B5A918541109FFB1FCAEAE451A9E93A8" box="[816,948,372,396]" italics="true" pageId="83" pageNumber="188">Goudotostes</emphasis>
|
||
</taxonomicName>
|
||
. Body strongly convex. Black, shiny, dorsum with extremely short fine setation (50×), underside, tarsi and antennae dark reddish-brown, setation yellowish-brown.
|
||
<emphasis id="B5A918541109FFB1FC35AEE51ACE93C8" box="[939,996,468,492]" italics="true" pageId="83" pageNumber="188">Head</emphasis>
|
||
: W/L ratio = 1.47; clypeal apex obtuse, apical projection strongly developed and sexually dimorphic, sides almost rectilinear with slightly unaligned genae in correspondence with the genal suture; dorsal ocular area absent, genal canthus indistinct, ventral ocular area small-sized, genae acute, distinctly protruding outwards, genal suture indistinct; head dorsal surface uniformly covered by dense large impressed simple punctures mixed to large impressed transverse comma-shaped punctures, fore margin marked by few irregular transverse lines; fronto-clypeal tubercle weakly raised. Interpunctural distance of punctures inferior than punctural diameter on clypeal disc.
|
||
<emphasis id="B5A918541109FFB1FAB2AC041CBF9168" box="[1324,1429,821,844]" italics="true" pageId="83" pageNumber="188">Pronotum</emphasis>
|
||
: subrectangular (dorsal view) (W/L ratio = 1.68), with a complete longitudinal row of tubercles, convex and unevenly sculptured (discal area gently raised with respect to sides), fore angles broadly subtruncate; pronotal surface covered by dense large deep simple punctures becoming sparser, smaller and comma-shaped at sides, area near fore angles with few irregular lines, interpunctural distance inferior than punctural diameter (equal to punctural diameter only on disc), pronotal bead complete; pronotal base raised in correspondence with scutellum; fore margin raised in correspondence with head insertion; pronotal sculpturing made of large and gently raised tubercles and carinae as follows: bilobate basal carina smooth and weakly raised, sides of disc with one gently raised transverse short carina and another distinctly raised transverse longer carina reaching outer margin, bilobate anterior carina weakly raised.
|
||
<taxonomicName id="40DDBFC51109FFB1FCAEAA651AB29748" box="[816,920,1364,1388]" class="Trilobita" family="Styginidae" genus="Scutellum" kingdom="Animalia" order="Corynexochida" pageId="83" pageNumber="188" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B5A918541109FFB1FCAEAA651AB29748" box="[816,920,1364,1388]" italics="true" pageId="83" pageNumber="188">Scutellum</emphasis>
|
||
</taxonomicName>
|
||
: basal area and sides smooth, remaining surface covered by impressed transverse large comma-shaped punctures.
|
||
<emphasis id="B5A918541109FFB1FC37AAA51AC19788" box="[937,1003,1428,1452]" italics="true" pageId="83" pageNumber="188">Elytra</emphasis>
|
||
: longer than wide (W/L ratio = 0.90), subovoidal (dorsal view), strongly convex; humeral callus marked by a tubercle; elytral surface covered by small deep sparse horseshoe-shaped punctures with opening directed backwards, inwards or outwards, interpunctural distance inferior than punctural diameter. Striated articular area hardly visible in lateral view, relatively narrow and short; inferior sutural stria visible complete; marginal area large-sized; elytral articular process well developed, smooth and shiny, isolated from remaining elytral surface by a deep furrow. Elytral sculpturing as follows: base with one raised large tubercle below articular process with apex directed backwards (parasutural tubercle), next to it an oblique larg- er raised tubercle with apex directed backwards (humeral tubercle), dorsal area of elytra with a longitudinal row of short irregular and asymmetrical carinae, sides and apical third with denser raised tubercles.
|
||
<emphasis id="B5A91854110EFFB6FD93AF841B6492E8" box="[525,590,181,204]" italics="true" pageId="84" pageNumber="189">Wings</emphasis>
|
||
: flightless.
|
||
<emphasis id="B5A91854110EFFB6FD52AF8519DF92C8" italics="true" pageId="84" pageNumber="189">Male genitalia</emphasis>
|
||
: aedeagus (L =
|
||
<quantity id="402569A3110EFFB6FE05AFE518C692C8" box="[411,492,212,236]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="84" pageNumber="189" unit="mm" value="1.1">1.1 mm</quantity>
|
||
) with parameres as in
|
||
<figureCitation id="1FE6D8C3110EFFB6FD48AFE519989328" captionStart="Fig" captionStartId="131.[136,167,1858,1877]" captionTargetBox="[135,1442,172,1817]" captionTargetId="figure-4@131.[135,1450,166,1833]" captionTargetPageId="131" captionText="Fig. 66 – Goudotostes parvus sp. nov., a, holotype, dorsal view of distended specimen; b, paratype, dorsal view of rolled up specimen; c, paratype, ventral view of rolled up specimen; d, paratype, ventral view of rolled up specimen; e, paratype, lateral view of rolled up specimen; f, holotype, spiculum gastrale; g–k, holotype, four views of aedeagus." figureDoi="http://doi.org/10.5281/zenodo.12764315" httpUri="https://zenodo.org/record/12764315/files/figure.png" pageId="84" pageNumber="189">Figs 66</figureCitation>
|
||
g-k, spiculum gastrale (L = 1.0 mm) as in
|
||
<figureCitation id="1FE6D8C3110EFFB6FDEDAFC51BFA9328" box="[627,720,244,268]" captionStart="Fig" captionStartId="131.[136,167,1858,1877]" captionTargetBox="[135,1442,172,1817]" captionTargetId="figure-4@131.[135,1450,166,1833]" captionTargetPageId="131" captionText="Fig. 66 – Goudotostes parvus sp. nov., a, holotype, dorsal view of distended specimen; b, paratype, dorsal view of rolled up specimen; c, paratype, ventral view of rolled up specimen; d, paratype, ventral view of rolled up specimen; e, paratype, lateral view of rolled up specimen; f, holotype, spiculum gastrale; g–k, holotype, four views of aedeagus." figureDoi="http://doi.org/10.5281/zenodo.12764315" httpUri="https://zenodo.org/record/12764315/files/figure.png" pageId="84" pageNumber="189">
|
||
Fig.
|
||
<specimenCount id="91DB0FCF110EFFB6FD3DAFC51BFA9328" box="[675,720,244,268]" count="66" pageId="84" pageNumber="189" type="generic">66 f</specimenCount>
|
||
</figureCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="CFC797CD110EFFB6FF23AE251B389348" pageId="84" pageNumber="189" type="etymology">
|
||
<paragraph id="8762C446110EFFB6FF23AE251B389348" blockId="84.[151,772,180,620]" pageId="84" pageNumber="189">
|
||
<emphasis id="B5A91854110EFFB6FF23AE2518179308" bold="true" box="[189,317,276,300]" pageId="84" pageNumber="189">Etymology.</emphasis>
|
||
Latin adjective in the nominative singular meaning “little”, since this is the smallest known repre- sentative of the genus
|
||
<taxonomicName id="40DDBFC5110EFFB6FE18AE651B219348" authorityName=", sensu Ballerio" authorityYear="2008" box="[390,523,340,364]" class="Insecta" family="Hybosoridae" genus="Goudotostes" kingdom="Animalia" order="Coleoptera" pageId="84" pageNumber="189" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B5A91854110EFFB6FE18AE651B219348" box="[390,523,340,364]" italics="true" pageId="84" pageNumber="189">Goudotostes</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="CFC797CD110EFFB6FF23AE451BDF9048" pageId="84" pageNumber="189" type="distribution">
|
||
<paragraph id="8762C446110EFFB6FF23AE451BDF9048" blockId="84.[151,772,180,620]" pageId="84" pageNumber="189">
|
||
<emphasis id="B5A91854110EFFB6FF23AE4518F393A8" bold="true" box="[189,473,372,396]" pageId="84" pageNumber="189">Distribution and habitat.</emphasis>
|
||
Known only from the
|
||
<typeStatus id="58667AE4110EFFB6FD48AE441A2993A9" box="[726,771,373,397]" pageId="84" pageNumber="189">type</typeStatus>
|
||
locality in Eastern
|
||
<collectingCountry id="FFCA84D6110EFFB6FEF6AEA518C29388" box="[360,488,404,428]" name="Madagascar" pageId="84" pageNumber="189">Madagascar</collectingCountry>
|
||
. The
|
||
<typeStatus id="58667AE4110EFFB6FDB7AEA41B7C9389" box="[553,598,405,429]" pageId="84" pageNumber="189">type</typeStatus>
|
||
series was collected by sifting leaf litter and rotten wood in rainforest. The vegetation cover consists of “moist evergreen forest”, and the collecting sites fall within the “forêt ombrophile de montagne” between 800 and
|
||
<quantity id="402569A3110EFFB6FE5CAD251B3A9008" box="[450,528,532,556]" metricMagnitude="3" metricUnit="m" metricValue="1.45" pageId="84" pageNumber="189" unit="m" value="1450.0">1450 m</quantity>
|
||
(
|
||
<bibRefCitation id="E34CB9B7110EFFB6FD85AD251BD29008" author="Goodman S. M. & Raherilalao M. J. & Wohlhauser S." box="[539,760,532,556]" pageId="84" pageNumber="189" refId="ref102713" refString="Goodman S. M., Raherilalao M. J., Wohlhauser S. (eds). 2018. Les aires potegees terrestres de Madagascar: leur histoire, description et biote / The terrestrial protected areas of Madagascar: their history, description and biota. Association Vahatra, Antananarivo. 1716 pp." type="book" year="2018">Goodman et al. 2018</bibRefCitation>
|
||
). The collecting locality falls within the protected area “Parc National de Morojejy (Forêts humides de l’Antsiranana)”.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |